1275:
827:
1623:
931:
998: oz), and the average length is 150 mm (6 in). At the time of birth, the scales are soft and white. After several days, they harden and darken to resemble those of an adult pangolin. During the vulnerable stage, the mother stays with her offspring in the burrow, nursing it, and wraps her body around it if she senses danger. The young cling to the mother's tail as she moves about, although, in burrowing species, they remain in the burrow for the first 2–4 weeks of life. At one month, they first leave the burrow riding on the mother's back.
816:
1585:
171:
721:
705:
877:. Most of their diet consists of various species of ants and termites, and may be supplemented by other insects, especially larvae. They are somewhat particular and tend to consume only one or two species of insects, even when many species are available to them. A pangolin can consume 140 to 200 g (5 to 7 oz) of insects per day. Pangolins are an important regulator of termite populations in their natural habitats.
270:
138:
892:, and they have other physical characteristics to help them eat ants and termites. Their skeletal structure is sturdy and they have strong front legs that are used for tearing into termite mounds. They use their powerful front claws to dig into trees, soil, and vegetation to find prey, then proceed to use their long tongues to probe inside the insect tunnels and to retrieve their prey.
2371:
1633:
Many attempts have been made to breed pangolins in captivity, but due to their reliance on wide-ranging habitats and very particular diets, these attempts are often unsuccessful. Pangolins have significantly decreased immune responses due to a genetic dysfunction, making them extremely fragile. They
5218:
Aitken-Palmer, Copper; deMaar, Thomas W.; Johnson, James G.; Langan, Jennifer; Bergmann, Jonathan; Chinnadurai, Sathya; Guerra, Hector; Carboni, Deborah A.; Adkesson, Michael J. (September 2019). "Complications
Associated with Pregnancy and Parturition in African White-bellied Pangolins (Phataginus
4444:
Xiao, Kangpeng; Zhai, Junqiong; Feng, Yaoyu; Zhou, Niu; Zhang, Xu; Zou, Jie-Jian; Li, Na; Guo, Yaqiong; Li, Xiaobing; Shen, Xuejuan; Zhang, Zhipeng; Shu, Fanfan; Huang, Wangyi; Li, Yu; Zhang, Ziding; Chen, Rui-Ai; Wu, Ya-jiang; Peng, Shi-Ming; Huang, Mian; Xie, Wei-Jun; Cai, Qin-Hui; Hou, Fang-Hui;
1649:
The idea of farming pangolins to reduce the number being illegally trafficked is being explored with little success. The third
Saturday in February is promoted as World Pangolin Day by the conservation NPO Annamiticus. World Pangolin Day has been noted for its effectiveness in generating awareness
1571:
Pangolin parts are also used for medicinal purposes in other Asian countries such as India, Nepal and
Pakistan. In some parts of India and Nepal, locals believe that wearing the scales of a pangolin can help prevent pneumonia. Pangolin scales have also been used for medicinal purposes in Malaysia,
1567:
of the People's
Republic of China included Chinese pangolin scales as an ingredient in TCM formulations. Pangolins were removed from the pharmacopoeia starting from the first half of 2020. Although pangolin scales have been removed from the list of raw ingredients, the scales are still listed as a
962:
Pangolins are solitary and meet only to mate, with mating typically taking place at night after the male and female pangolin meet near a watering hole. Males are larger than females, weighing up to 40% more. While the mating season is not defined, they typically mate once each year, usually during
3468:
Gaubert, Philippe; Antunes, Agostinho; Meng, Hao; Miao, Lin; Peigné, Stéphane; Justy, Fabienne; Njiokou, Flobert; Dufour, Sylvain; Danquah, Emmanuel; Alahakoon, Jayanthi; Verheyen, Erik (11 May 2018). "The
Complete Phylogeny of Pangolins: Scaling Up Resources for the Molecular Tracing of the Most
1699:
became important communities for protecting pangolins and their habitats and has greatly improved the survival of pangolins. These centers work with local aboriginal tribes and forest police in the
National Police Agency to prevent poaching, trafficking, and smuggling of pangolins, especially to
1614:
of the animal while educating communities on its importance. Another suggested approach to fighting pangolin (and general wildlife) trafficking consists in "following the money" rather than "the animal", which aims to disrupt smugglers' profits by interrupting money flows. Financial intelligence
1604:
As a result of increasing threats to pangolins, mainly in the form of illegal, international trade in pangolin skin, scales, and meat, these species have received increasing conservation attention in recent years. As of
January 2020, the IUCN considered all eight species of pangolin on its
1609:
as threatened. The IUCN SSC Pangolin
Specialist Group launched a global action plan to conserve pangolins, dubbed "Scaling up Pangolin Conservation", in July 2014. This action plan aims to improve all aspects of pangolin conservation with an added emphasis on combating poaching and
2387:
1615:
gathering could thus become a key tool in protecting these animals, although this opportunity is often overlooked. In 2018, a
Chinese NGO launched the Counting Pangolins movement, calling for joint efforts to save the mammals from trafficking. Wildlife conservation group
963:
the summer or autumn. Rather than the males seeking out the females, males mark their location with urine or feces and the females find them. If competition over a female occurs, the males use their tails as clubs to fight for the opportunity to mate with her.
1354:. In the past decade, numerous seizures of illegally trafficked pangolin and pangolin meat have taken place in Asia. In one such incident in April 2013, 10,000 kg (22,000 pounds) of pangolin meat were seized from a Chinese vessel that ran aground in the
1525:, a recipe for expelling evil spirits with a formulation of scales, herbs, and minerals appeared in 682, and in 752 CE the idea that pangolin scales could also stimulate milk secretion in lactating women, one of the main uses today, was recommended in the
969:
periods differ by species, ranging from roughly 70 to 140 days. African pangolin females usually give birth to a single offspring at a time, but the
Asiatic species may give birth to from one to three. Weight at birth is 80 to 450 g
2090:"A Mystery Species Was Discovered in Trafficked Pangolin Scales - Researchers believed there were eight species of the strange mammals. But a ninth was identified genetically, although no one knew it was a separate species in the wild"
4858:
Zhang, Fang; Kong, Lin-lin; Zhang, Yi-ye; Li, Shu-Chuen (2012). "Evaluation of Impact on Health-Related Quality of Life and Cost Effectiveness of Traditional Chinese Medicine: A Systematic Review of Randomized Clinical Trials".
5311:
Challender, Daniel W. S.; Sas-Rolfes, Michael't; Ades, Gary W. J.; Chin, Jason S. C.; Ching-Min Sun, Nick; Chong, Ju lian; Connelly, Ellen; Hywood, Lisa; Luz, Sonja; Mohapatra, Rajesh K.; de Ornellas, Paul (1 October 2019).
3454:
Sean P. Heighton, Rémi Allio, Jérôme Murienne, Jordi Salmona, Hao Meng, Céline Scornavacca, Armanda D. S. Bastos, Flobert Njiokou, Darren W. Pietersen, Marie-Ka Tilak, Shu-Jin Luo, Frédéric Delsuc, Philippe Gaubert (2023.)
511:, which they capture using their long tongues. They tend to be solitary animals, meeting only to mate and produce a litter of one to three offspring, which they raise for about two years. Pangolins superficially resemble
1335:
China had been the main destination country for pangolins until 2018, where it was surpassed by Vietnam. In 2019, Vietnam was reported to have seized the largest volumes of pangolin scales, surpassing Nigeria that year.
1521:("Variorum of Shennong's Classic of Materia Medica", 500 CE), which recommends pangolin scales for protection against ant bites; burning the scales as a cure for people crying hysterically during the night. During the
7209:
1429:
share only up to 92% of their RNA. Ecologists worried that the early speculation about pangolins being the source may have led to mass slaughters, endangering them further, which was similar to what happened to
3838:
Ingram, Daniel J.; Coad, Lauren; Abernethy, Katharine A.; Maisels, Fiona; Stokes, Emma J.; Bobo, Kadiri S.; Breuer, Thomas; Gandiwa, Edson; Ghiurghi, Andrea; Greengrass, Elizabeth; Holmern, Tomas (March 2018).
1854:
Kenneth E. Kinman (1994) "The Kinman System: Toward a Stable Cladisto-Eclectic Classification of Organisms: Living and Extinct, 48 Phyla, 269 Classes, 1,719 Orders", Hays, Kan. (P. O. Box 1377, Hays 67601), 88
1700:
black markets in China. These centers have also helped to reveal the causes of death and injury among Taiwan's pangolin population. Today, Taiwan has the highest population density of pangolins in the world.
1682:
is one of the few conservation grounds for pangolins in the world after the country enacted the 1989 Wildlife Conservation Act. The introduction of Wildlife Rehabilitation Centers in places like Luanshan
4223:
2121:
D'Cruze, Neil; Assou, Délagnon; Coulthard, Emma; Norrey, John; Megson, David; Macdonald, David W.; Harrington, Lauren A.; Ronfot, Delphine; Segniagbeto, Gabriel H.; Auliya, Mark (5 November 2020).
1510:
for animal body parts, despite concerns about toxicity, transmission of diseases from animals to humans, and species extermination. The ongoing demand for parts as ingredients continues to fuel
4981:
3744:
1346:
Though pangolins are protected by an international ban on their trade, populations have suffered from illegal trafficking due to beliefs in East Asia that their ground-up scales can stimulate
1302:. Their meat is also considered a delicacy. 100,000 are estimated to be trafficked a year to China and Vietnam, amounting to over one million over the past decade. This makes them the most
1002:
takes place around three months of age, when the young begin to eat insects in addition to nursing. At two years of age, the offspring are sexually mature and are abandoned by the mother.
728:
The physical appearance of a pangolin is marked by large, hardened, overlapping, plate-like scales, which are soft on newborn pangolins, but harden as the animal matures. They are made of
1358:. In another case in August 2016, an Indonesian man was arrested after police raided his home and found over 650 pangolins in freezers on his property. The same threat is reported in
4081:
4913:
4196:
899:
is sticky, causing ants and termites to stick to their long tongues when they are hunting through insect tunnels. Without teeth, pangolins also lack the ability to chew; but while
1642:
in captivity, complications that can lead to an early death. In addition, pangolins rescued from illegal trade often have a higher chance of being infected with parasites such as
1417:, China, hypothesized that SARS-CoV-2 had originated in bats, and prior to infecting humans, was circulating among pangolins. The illicit Chinese trade of pangolins for use in
6528:
3253:
Amrine-Madsen, H.; Koepfli, K.P.; Wayne, R.K.; Springer, M.S. (2003). "A new phylogenetic marker, apolipoprotein B, provides compelling evidence for eutherian relationships".
4104:
1476:. To refer to both sampling site and hosts, they were named Dongyang pangolin virus (DYPV) and Lishui pangolin virus (LSPV). The DYPV pestivirus was also identified in
5474:
Archer, Lucy J.; Papworth, Sarah K.; Apale, Charity M.; Corona, Darlyn B.; Gacilos, Josefa T.; Amada, Ronald L.; Waterman, Carly; Turvey, Samuel T. (1 December 2020).
3713:
2527:
Wang, Bin; Yang, Wen; Sherman, Vincent R.; Meyers, Marc A. (2016). "Pangolin armor: Overlapping, structure, and mechanical properties of the keratinous scales".
4219:
4122:
3360:
Du Toit, Z.; Grobler, J. P.; Kotzé, A.; Jansen, R.; Brettschneider, H.; Dalton, D. L. (2014). "The complete mitochondrial genome of Temminck's ground pangolin (
4846:
The vacuum created by China's failure to adequately support a disciplined scientific approach to traditional Chinese medicine has been filled by pseudoscience
3991:
495:
scales, similar in material to fingernails and toenails, covering their skin; they are the only known mammals with this feature. They live in hollow trees or
4171:
861:
Some pangolins walk with their front claws bent under the foot pad, although they use the entire foot pad on their rear limbs. Furthermore, some exhibit a
743:
are made, and are structurally and compositionally very different from the scales of reptiles. The pangolin's scaled body is comparable in appearance to a
1446:
and its binding to receptors in pangolins had minimal effect from the virus, and therefore were not likely mechanisms for COVID-19 infections in humans.
5681:
3946:
4790:
4989:
3752:
2884:
Coulson, Ian M; Heath, Martha E (December 1997). "Foraging behaviour and ecology of the Cape pangolin (Manis temminckii) in north-western Zimbabwe".
2099:
7204:
6885:
6874:
4147:
7086:
6695:
6665:
6657:
6607:
6591:
3653:
3089:
6914:
6813:
6745:
6703:
599:
484:
include two species each, all found in sub-Saharan Africa. These species range in size from 30 to 100 cm (12 to 39 in). A number of
7125:
6859:
2431:"Combined proteomics and transcriptomics reveal the genetic basis underlying the differentiation of skin appendages and immunity in pangolin"
5476:"Scaling up local ecological knowledge to prioritise areas for protection: Determining Philippine pangolin distribution, status and threats"
4071:
1366:. The overexploitation comes from hunting pangolins for game meat and the reduction of their forest habitats due to deforestation caused by
6922:
6615:
6599:
5551:
Pei, Curtis Jai-Chyi; Chen, Chen-Chih; Chi, Meng-Jou; Lin, Wen-Chi; Lin, Jing-Shiun; Arora, Bharti; Sun, Nick Ching-Min (6 February 2019).
4186:
5672:
4904:
7199:
6821:
6623:
5377:
Chua, Marcus A.H.; Tan, Audrey; Carrasco, Luis Roman (2021). "Species awareness days: Do people care or are we preaching to the choir?".
5177:
5150:
5011:
3202:
Murphy WJ, Eizirik E, et al. (14 December 2001). "Resolution of the Early Placental Mammal Radiation Using Bayesian Phylogenetics".
3063:
2769:
4833:
1983:
5448:
6214:
5690:
4702:
2496:
2026:
1274:
5162:
1332:
which prohibits international trade, except when the product is intended for non-commercial purposes and a permit has been granted.
7060:
1868:
Zoological names. A list of phyla, classes, and orders, prepared for section F, American Association for the Advancement of Science
1438:
outbreak. It was later proved that the testing which suggested that pangolins were a potential host for the virus was flawed, when
2783:
Ma, Jing-E; Li, Lin-Miao; Jiang, Hai-Ying; Zhang, Xiu-Juan; Li, Juan; Li, Guan-Yu; Yuan, Li-Hong; Wu, Jun; Chen, Jin-Ping (2017).
2601:
7099:
5649:
3808:
1426:
911:, and it is also covered in keratinous spines. These spines further aid in the grinding up and digestion of the pangolin's prey.
7179:
5644:
5524:"Taiwan's Path to Pangolin Conservation : How a Mega Pangolin Leather Exporter Transformed into a Conservation Specialist"
4096:
3556:
7117:
5085:
5053:
4827:
3186:
3046:
2960:
2930:
2765:
2324:
2009:
1316:
have become commercially extinct in certain ranges as a result of overhunting. In November 2010, pangolins were added to the
7104:
4011:
Boakye, Maxwell Kwame; Pietersen, Darren William; Kotzé, Antoinette; Dalton, Desiré-Lee; Jansen, Raymond (20 January 2015).
1320:'s list of evolutionarily distinct and endangered mammals. All eight species of pangolin are assessed as threatened by the
614:
797:. Large pangolins can extend their tongues as much as 40 cm (16 in), with a diameter of only about 0.5 cm (
2251:
2123:"Snake oil and pangolin scales: insights into wild animal use at "Marché des Fétiches" traditional medicine market, Togo"
751:, while it protects its face by tucking it under its tail. The scales are sharp, providing extra defense from predators.
3721:
3506:"Mitochondrial genomes of African pangolins and insights into evolutionary patterns and phylogeny of the family Manidae"
2785:"Transcriptomic analysis identifies genes and pathways related to myrmecophagy in the Malayan pangolin (Manis javanica)"
2213:
3405:
2860:
2674:
2293:
858:
pangolins live in hollow trees, whereas the ground-dwelling species dig tunnels to a depth of 3.5 m (11 ft).
2027:"The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis"
1866:
1667:
In December 2020, a study found that it is "not too late" to establish conservation efforts for Philippine pangolins (
1576:, Philippines, have held the belief that elders could avoid prostate illnesses by wearing belts made with the scales.
1537:("Formulas from Benevolent Sages Compiled During the Era of Peace and Tranquility"), compiled by Wang Huaiyin in 992.
5744:
4957:
4118:
1999:
5688:
The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis
848:
is also active by day, while other species of pangolins spend most of the daytime sleeping, curled up into a ball ("
826:
4161:
2727:
2043:
1616:
7184:
7130:
1299:
5657:
5523:
2381:
2344:
2089:
2034:
895:
The structure of their tongue and stomach is key to aiding pangolins in obtaining and digesting insects. Their
595:
5711:
2246:
6207:
5614:
2356:
4780:
3841:"Assessing Africa-Wide Pangolin Exploitation by Scaling Local Data: Assessing African pangolin exploitation"
3629:
The Illegal Wildlife Trade: Inside the World of Poachers, Smugglers and Traders (Studies of Organized Crime)
1553:
1495:
1418:
1317:
1295:
1279:
4366:"Viral Metagenomics Revealed Sendai Virus and Coronavirus Infection of Malayan Pangolins (Manis javanica)"
1622:
7112:
7008:
4905:"'If we don't buy, they don't die': Tackling the global demand that's driving the illegal wildlife trade"
4137:
907:), which accumulate in their stomachs to help to grind up ants. This part of their stomach is called the
3114:
Hua, Liushuai; Gong, Shiping; Wang, Fumin; Li, Weiye; Ge, Yan; Li, Xiaonan; Hou, Fanghui (8 June 2015).
4637:
Gao, Wen-Hua; Lin, Xian-Dan; Chen, Yan-Mei; Xie, Chun-Gang; Tan, Zhi-Zhou; Zhou, Jia-Jun; Chen, Shuai;
3093:
3038:
7013:
3675:
Challender, D.; Willcox, D.H.A.; Panjang, E.; Lim, N.; Nash, H.; Heinrich, S. & Chong, J. (2019).
3643:
2370:
Vergara, Benito S.; Idowu, Panna Melizah H.; Sumangil, Julia H.; Gonzales, Juan Carlos; Dans, Andres.
170:
5682:
Proceedings of the Workshop on Trade and Conservation of Pangolins Native to South and Southeast Asia
5420:
3908:
Challender, D.; Wu, S.; Kaspal, P.; Khatiwada, A.; Ghose, A.; Ching-Min Su, N.; Suwal, Laxmi (2019).
1733:
1443:
1394:
31:
17:
5103:"Small NGOs and Agenda-Setting in Global Conservation Governance: The Case of Pangolin Conservation"
7194:
6200:
3302:
2570:
922:
tails to hang from tree branches and strip away bark from the trunk, exposing insect nests inside.
5702:
2978:"Loss of teeth and enamel in tetrapods: fossil record, genetic data and morphological adaptations"
1908:
1882:
4551:
4166:
2626:
2505:
2316:
1895:
1499:
865:
stance for some behaviour, and may walk a few steps bipedally. Pangolins are also good swimmers.
584:
4416:
3457:"Pangolin genomes offer key insights and resources for the world’s most trafficked wild mammals"
3178:
3171:
7156:
6948:
4785:
2597:
2416:
778:
250:
5666:
5174:
5069:
5037:
4945:
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3032:
2946:
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1804:
1619:
has identified 159 smuggling routes used by pangolin traffickers and aims to shut these down.
7151:
5737:
5528:
4813:
4325:
Cyranoski, David (7 February 2020). "Did pangolins spread the China coronavirus to people?".
2920:
1709:
1422:
1390:
1325:
1249:
939:
930:
7143:
7091:
5701:
Bromley, Victoria (Director/Producer), Young, Nora (Narrator/Host), Diekmann, Maria (2018).
3504:
du Toit, Z.; du Plessis, M.; Dalton, D. L.; Jansen, R.; Paul Grobler, J.; Kotzé, A. (2017).
2615:
Chan, Lap-Ki (1995). "Extrinsic Lingual Musculature of Two Pangolins (Pholidota: Manidae)".
1237:. Within the latter, a split separates the large terrestrial African pangolins of the genus
7034:
6995:
5568:
5487:
5386:
5325:
4743:
4710:
4507:
4458:
4024:
3418:
3262:
3211:
3123:
2991:
2893:
2617:
2442:
2184:
1745:
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for human transmission. However, whole-genome comparison found that the pangolin and human
1248:
Asian and African pangolins are thought to have diverged about 41.37 Ma ago. Moreover, the
845:
670:
566:
527:
516:
5687:
2063:
1922:
8:
6986:
6137:(Rats, guinea pigs, squirrels, beavers, chinchillas, porcupines, capybaras and relatives)
6116:(Pigs, camels, hippos, deer, buffalo, gazelles, giraffes, whales, dolphins and relatives)
3414:
2905:
2127:
1503:
1478:
1370:. The pangolin are hunted as game meat for both medicinal purposes and food consumption.
935:
713:
572:
539:
5572:
5491:
5390:
5329:
5193:
4747:
4511:
4462:
4028:
3422:
3266:
3215:
3127:
2897:
2852:
2446:
2429:
Li, Hui-Ming; Liu, Ping; Zhang, Xiu-Juan; Li, Lin-Miao; et al. (3 September 2020).
2351:. Vol. II. Mytilene, Greece: Salavopoulos & Kinderlis. p. 567 – via
2188:
2141:
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1977:
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7189:
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5715:
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5402:
5288:
5263:
5244:
5132:
4944:
Xing, S.; Bonebrake, T. C.; Cheng, W.; Zhang, M.; Ades, G.; Shaw, D.; Zhou, Y. (2019).
4671:
4646:
4616:
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4528:
4495:
4392:
4365:
4346:
4076:
4047:
4012:
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3583:
3532:
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3330:
3297:
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Beck, Robin; Bininda-Emonds, Olaf; Cardillo, Marcel; Liu, Fu-Guo; Purvis, Andy (2006).
3252:
3235:
3146:
3115:
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2982:
2977:
2952:
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2817:
2784:
2707:
2638:
2585:
2513:
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2430:
2218:
2154:
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2055:
2004:(3rd ed.). Baltimore, Maryland, USA: Johns Hopkins University Press. p. 530.
1950:
1837:
1384:
794:
590:
554:
543:
535:
282:
165:
5314:"Evaluating the feasibility of pangolin farming and its potential conservation impact"
4013:"Knowledge and uses of African pangolins as a source of traditional medicine in Ghana"
3909:
3816:
3676:
3274:
1339:
Pangolins are also hunted and eaten in Ghana and are one of the more popular types of
7138:
7021:
6105:
5596:
5505:
5406:
5293:
5248:
5236:
5151:
Spotlight: Pangolin conservationists call for ban on illegal trade of mammal products
5136:
5124:
5081:
5049:
4953:
4884:
4876:
4823:
4761:
4676:
4620:
4598:
4533:
4476:
4397:
4350:
4338:
4052:
3613:
3601:
3537:
3486:
3381:
3335:
3278:
3227:
3182:
3151:
3042:
3013:
2999:
2956:
2926:
2856:
2822:
2798:
2670:
2548:
2476:
2458:
2408:
2377:
2320:
2158:
2146:
2005:
1954:
1942:
1410:
1189:
are even closer relatives to pangolins, being classified with pangolins in the clade
881:
689:
662:
548:
38:
3438:
3239:
2870:
1841:
1646:, further lessening their chance for rehabilitation and reintroduction to the wild.
747:. It can curl up into a ball when threatened, with its overlapping scales acting as
538:
mammals in the world. As of January 2020, there are eight species of pangolin whose
7026:
6836:
6794:
6192:
6126:
5867:
5804:
5730:
5586:
5576:
5495:
5394:
5343:
5333:
5283:
5275:
5228:
5114:
4868:
4751:
4666:
4658:
4638:
4606:
4590:
4523:
4515:
4466:
4387:
4377:
4330:
4302:
4191:
4042:
4032:
3862:
3852:
3591:
3527:
3517:
3478:
3426:
3399:
Kondrashov, Peter; Agadjanian, Alexandre K. (2012). "A nearly complete skeleton of
3373:
3325:
3315:
3270:
3219:
3141:
3131:
3003:
2995:
2901:
2848:
2812:
2802:
2740:
2703:
2630:
2540:
2509:
2466:
2450:
2192:
2136:
2059:
2047:
1972:
1934:
1829:
1757:
1753:
1684:
1643:
1439:
1431:
1363:
1312:
748:
578:
411:
7000:
1584:
1193:. The split between carnivorans and pangolins is estimated to have occurred 79.47
6638:
6568:
6277:
6266:
6224:
6019:
5920:
5888:
5694:
5676:
5661:
5581:
5398:
5181:
5163:
World Pangolin Day: Conservationists demand greater protection to stop extinction
4819:
4037:
3430:
2736:
2723:"Behavioural descriptions of Indian pangolins (Manis crassicaudata) in captivity"
2544:
1968:
1714:
1692:
1593:
1552:
secrete milk. The supposed health effects of pangolin meat and scales claimed by
1310:, has led to a large decrease in the numbers of pangolins. Some species, such as
1256:
within Pholidota suggests the group originated in Eurasia, consistent with their
1190:
1148:
830:
819:
815:
627:
235:
5500:
5475:
5338:
5313:
3992:"First ivory, now pangolin scales: Vietnam remains wildlife trafficking hotspot"
1938:
1173:), but recent genetic evidence indicates their closest living relatives are the
488:
pangolin species are also known. In September 2023, nine species were reported.
6971:
6902:
6734:
6517:
6387:
6065:
6011:
5912:
5896:
5077:
5045:
4642:
4334:
4245:
3969:
Convention on International Trade in Endangered Species of Wild Fauna and Flora
3964:
3674:
3377:
3311:
2454:
1696:
1688:
1511:
1303:
1269:
1257:
1089:
951:
774:
650:
634:
560:
523:
4519:
4496:"Probable Pangolin Origin of SARS-CoV-2 Associated with the COVID-19 Outbreak"
4471:
4446:
4307:
4290:
3522:
3116:"Captive breeding of pangolins: current status, problems and future prospects"
2494:
Spearman, R.I.C. (2008). "On the nature of the horny scales of the pangolin".
2051:
7173:
6851:
6554:
6355:
6056:
5928:
5619:
5509:
5128:
4880:
4765:
4602:
3918:
3883:
3685:
3605:
3596:
3571:
3136:
2462:
2150:
1606:
1564:
1560:
are based on their consumption of ants, long tongues, and protective scales.
1307:
1186:
1080:
915:
611:
531:
150:
90:
3482:
3223:
1771:
1343:, while local healers use the pangolin as a source of traditional medicine.
6477:
6466:
6169:(Lorises, lemurs, tarsiers, monkeys, apes (including humans) and relatives)
6073:
6003:
5904:
5859:
5600:
5297:
5240:
4888:
4680:
4594:
4537:
4480:
4401:
4342:
4187:"Chinese vessel on Philippine coral reef caught with illegal pangolin meat"
4142:
4056:
3541:
3490:
3385:
3339:
3320:
3282:
3231:
3155:
3017:
2826:
2552:
2480:
1946:
1639:
1589:
1530:
1522:
1507:
1226:
1154:
1040:
844:
animals which use their well-developed sense of smell to find insects. The
834:
762:. They have short legs, with sharp claws which they use for burrowing into
720:
158:
5279:
4872:
4138:"'Noah's Ark' of 5,000 rare animals found floating off the coast of China"
3778:
3364:; Smuts, 1832) and phylogenetic position of the Pholidota (Weber, 1904)".
2976:
Davit-Béal, Tiphaine; Tucker, Abigail S.; Sire, Jean-Yves (1 April 2009).
2745:
2722:
1401:
taken from pangolins was found to be a 99% match with SARS coronavirus 2 (
7210:
Species endangered by human consumption for medicinal or magical purposes
7073:
6980:
6779:
6768:
6711:
6374:
6366:
6303:
6113:
6035:
5953:
5425:
5119:
5102:
4662:
4420:
4272:
3572:"The Use and Prescription of Pangolin in Traditional Vietnamese Medicine"
1654:
1611:
1533:, the notion of penetrating and clearing blockages was emphasized in the
1398:
1355:
1050:
904:
874:
733:
65:
5615:"Taiwanese Researchers Collaborate With Locals In Pangolin Conservation"
5449:"It's not too late – yet – to save the Philippine pangolin, study finds"
6399:
6142:
5995:
5838:
5795:
5779:
5722:
5012:"Did China really ban the pangolin trade? Not quite, investigators say"
4220:"Boat Filled With 22,000 Pounds Of Pangolin Hits Endangered Coral Reef"
3857:
3840:
2807:
2642:
1597:
1545:
1465:
1461:
1402:
1194:
1097:
1037:
919:
862:
462:
447:
110:
75:
5363:
5348:
5232:
4382:
3867:
1278:
Confiscated black market pangolin scales, which are in high demand in
7078:
7047:
6753:
6574:
6504:
6150:
6097:
5974:
5966:
5846:
5812:
5264:"Pangolin genomes and the evolution of mammalian scales and immunity"
4982:"Pangolins receive surprising lifeline with new protections in China"
4756:
4731:
3996:
3403:(Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis".
2845:
Xenarthra and Pholidota (Armadillos, Anteaters, Sloths and Pangolins)
2197:
2172:
1635:
1549:
1502:
exists for the efficacy of those practices, and they have no logical
1414:
1347:
1206:
1174:
1170:
1158:
1027:
966:
885:
849:
841:
744:
740:
704:
515:, though the two are not closely related; they have merely undergone
512:
500:
182:
115:
59:
6942:
5654:
4815:
Global Technological Change: From Hard Technology to Soft Technology
4447:"Isolation of SARS-CoV-2-related coronavirus from Malayan pangolins"
3777:
Haenlein, Alexandria; Reid, Cathy; Keatinge, Tom (10 October 2018).
3570:
Sexton, Rebecca; Nguyen, Trang; Roberts, David L. (1 January 2021).
2634:
1785:
7065:
7052:
6965:
6316:
6242:
5945:
5773:
3648:
3557:"Change the List: The Most Trafficked Mammal You've Never Heard Of"
2536:
1833:
1671:), a species that is only found on the island province of Palawan.
1557:
1541:
1406:
1340:
1283:
1162:
1061:
900:
855:
773:
The tongues of pangolins are extremely long, and like those of the
737:
709:
508:
202:
105:
100:
85:
80:
70:
1626:
Pangolins (in rectangular cages) in an illegal wildlife market in
1294:
because their scales are believed to have medicinal properties in
6443:
6432:
6418:
6329:
6166:
6043:
5875:
5785:
4246:"Indonesian man arrested as 650 pangolins found dead in freezers"
3745:"Tiger Eyes, Crocodile Penis: It's What's For Dinner in Malaysia"
3587:
2001:
Mammal Species of the World: A Taxonomic and Geographic Reference
1659:
1627:
1573:
1367:
1359:
1291:
1122:
1112:
1071:
999:
955:
947:
943:
908:
889:
790:
767:
729:
492:
485:
468:
451:
401:
154:
146:
120:
95:
5217:
4647:"Newly identified viral genomes in pangolins with fatal disease"
4417:"1st known case of coronavirus traced back to November in China"
4291:"Pangolins in south-west Nigeria – current status and prognosis"
1786:"Scientific names of mammal orders: from descriptive to uniform"
1328:. All pangolin species are currently listed under Appendix I of
6248:
6236:
6158:
6134:
5982:
5829:
5767:
5754:
5650:
Pangolin: Wildlife summary from the African Wildlife Foundation
2281:. Washington, D.C.: Smithsonian Institution Press. p. 604.
2120:
1998:
Schlitter, Duane A. (2005). Wilson, D.E.; Reeder, D.M. (eds.).
1976:
1883:"The Zoologist: a monthly journal of natural history. (Vol. 1)"
1679:
1351:
896:
786:
496:
269:
212:
192:
127:
3201:
1572:
Indonesia and northern Myanmar. Indigenous people in southern
1494:
Pangolin scales and flesh are used as ingredients for various
7039:
6257:
6027:
4439:
4437:
3503:
3295:
2883:
2789:
2352:
1329:
1182:
1178:
1166:
782:
759:
755:
456:
222:
5310:
2925:. Pinetown, South Africa: 30° South Publishers. p. 36.
2922:
The South African Bushveld: A Field Guide from the Waterberg
137:
3907:
2256:
1483:
1435:
1321:
950:
island group. It is threatened by illegal poaching for the
504:
438:
432:
5070:"Meat and medicine: historic and contemporary use in Asia"
5038:"Meat and medicine: historic and contemporary use in Asia"
4946:"Meat and medicine: historic and contemporary use in Asia"
4434:
4010:
3837:
2690:
Wilson, Amelia E. (January 1994). "Husbandry of pangolins
1540:
In the 21st century, the main uses of pangolin scales are
417:
6081:
5473:
4289:
Sodeinde, Olufemi A.; Adedipe, Segun R. (24 April 2009).
4100:
3947:"'Asian unicorn' and scaly anteater make endangered list"
3644:"Over a million pangolins slaughtered in the last decade"
3359:
763:
423:
3392:
2566:
2369:
1529:("Arcane Essentials from the Imperial Library"). In the
1362:, where the animal is on the verge of extinction due to
4943:
4589:(7). Centers for Disease Control and Prevention: 1810.
4494:
Zhang, Tao; Wu, Qunfu; Zhang, Zhigang (19 March 2020).
3884:"The world's most-trafficked mammal – and the scaliest"
3714:"Crime Blotter: Pangolin Scales, Tiger Skins, and More"
2419:, translated A. F. Scholfield 1959 vol. 3, pp. 266-268.
1544:
practices based on unproven claims the scales dissolve
1241:
from the small arboreal African pangolins of the genus
3467:
958:, where it is regarded as a luxury medicinal delicacy.
5553:"Mortality and morbidity in wild Taiwanese pangolin (
5175:
How China is combating wildlife trafficking in Africa
4861:
The Journal of Alternative and Complementary Medicine
3169:
Dickman, Christopher R. (1984). MacDonald, D. (ed.).
3064:"All hope is not lost for vanishing Palawan pangolin"
2526:
1820:
Cope, E. D. (1889). "The Edentata of North America".
414:
6222:
4948:. In Challender, D.; Nash, H.; Waterman, C. (eds.).
3944:
3776:
3398:
2975:
1898:, London, Day and Son, Ltd., XVI + 420 pp., 101 maps
1460:
In 2020, two novel RNA viruses distantly related to
754:
Pangolins can emit a noxious-smelling chemical from
435:
429:
420:
4973:
4072:"No Species Is Safe From Burgeoning Wildlife Trade"
3569:
3298:"A higher-level MRP supertree of placental mammals"
3092:. University of Wisconsin–La Crosse. Archived from
1290:Pangolins are in high demand in southern China and
426:
6108:(Horses, zebras, donkeys, rhinoceroses and tapirs)
4903:
4445:Chen, Wu; Xiao, Lihua; Shen, Yongyi (7 May 2020).
4097:"The Most Trafficked Mammal You've Never Heard Of"
3170:
2376:. Washington, D.C.: Island Publishing House, Inc.
1153:The order Pholidota was long considered to be the
4703:"Quackery and superstition: species pay the cost"
4320:
4318:
3779:"What's the secret to saving this rare creature?"
7171:
6427:
5261:
4857:
4700:
4443:
4284:
4282:
3772:
3770:
3548:
3246:
3034:The Princeton Field Guide to Prehistoric Mammals
1909:"A manual of the anatomy of vertebrated animals"
1662:: Jackie Chan & Pangolins (Kung Fu Pangolin)
781:, the root of the tongue is not attached to the
37:"Pholidota" redirects here. For the orchid, see
5803:
5376:
4952:(First ed.). Academic Press. p. 233.
4288:
3450:
3448:
3289:
2669:. New York City: Arch Cape Press. p. 252.
2241:
2239:
2237:
1005:
4778:
4315:
3355:
3353:
3351:
3349:
2939:
2912:
2782:
2721:Mohapatra, Rajesh K.; Panda, Sudarsen (2014).
2720:
1967:
1809:(in German). Vol. T.3. Berlin: G. Reimer.
1772:"A corrected classification of the edentates."
1517:The first record of pangolin scales occurs in
600:International Union for Conservation of Nature
526:(for their meat and scales, which are used in
6208:
5738:
5550:
5366:. Pangolins.org/Annamiticus. 26 October 2011.
5194:"Pangolins – Species we work with at TRAFFIC"
4636:
4279:
3803:
3801:
3799:
3767:
3113:
2292:Liddell, Henry George; Scott, Robert (1940).
1592:from India, presented in 1875–76 to the then
1393:of a specific receptor-binding domain of the
1209:of extant Manidae separates Asian pangolins (
884:. They also lack teeth. They rely heavily on
724:Schematic drawing of pangolin scale histology
5074:Pangolins: Science, Society and Conservation
5042:Pangolins: Science, Society and Conservation
4950:Pangolins: Science, Society and Conservation
4811:
4493:
3445:
3024:
2838:
2836:
2660:
2658:
2656:
2654:
2652:
2487:
2428:
2338:
2336:
2291:
2234:
2081:
2018:
1449:
534:of their natural habitats, and are the most
476:comprises four species found in Asia, while
4772:
4363:
3497:
3461:
3346:
3109:
3107:
3105:
3103:
3087:
2759:
1783:
618:
6215:
6201:
5745:
5731:
5413:
4632:
4630:
4184:
3796:
3090:"Rollin' With the Pangolin – Reproduction"
2918:
2842:
1657:made a public service announcement called
1468:have been detected in the genomes of dead
499:, depending on the species. Pangolins are
268:
136:
5590:
5580:
5499:
5347:
5337:
5287:
5118:
4939:
4937:
4935:
4933:
4931:
4851:
4755:
4670:
4610:
4527:
4470:
4391:
4381:
4324:
4306:
4119:"23 tonnes of pangolins seized in a week"
4046:
4036:
3945:Agence France-Presse (19 November 2010).
3866:
3856:
3595:
3531:
3521:
3329:
3319:
3145:
3135:
3007:
2833:
2816:
2806:
2744:
2714:
2664:
2649:
2497:Zoological Journal of the Linnean Society
2470:
2342:
2333:
2196:
2140:
2087:
1997:
758:near the anus, similar to the spray of a
6076:(Hedgehogs, shrews, moles and relatives)
5752:
5100:
4069:
3881:
3668:
3100:
3062:Fabro, Keith Anthony S. (10 June 2019).
3030:
2520:
2493:
2310:
1911:, New York, d, Appleton and Co., 431 pp.
1896:"The geographic distribution of mammals"
1621:
1583:
1282:, set to be destroyed by authorities in
1273:
929:
825:
814:
719:
703:
7205:Species endangered by human consumption
4805:
4781:"What Is Traditional Chinese Medicine?"
4627:
4364:Liu, P.; Chen, W.; Chen, J.-P. (2019).
4217:
3626:
3168:
2583:
2363:
2315:(10th ed.). Oxfordshire, England:
2214:"A Struggle to Save the Scaly Pangolin"
2088:Incorvala, Darren (25 September 2023).
1802:
1774:Science, new ser., vol. 31, pp. 913-914
610:The name of order Pholidota comes from
14:
7172:
5712:Coronavirus: Revenge of the Pangolins?
5704:Nature: The World's Most Wanted Animal
4928:
4696:
4694:
4692:
4690:
4572:
4566:
4414:
4174:from the original on 22 February 2015.
4125:from the original on 26 November 2014.
4094:
4084:from the original on 22 February 2015.
3742:
3711:
3554:
2772:from the original on 23 December 2014.
2689:
2604:from the original on 21 December 2014.
2573:from the original on 22 February 2015.
2349:A Malay-English dictionary (romanised)
2102:from the original on 25 September 2023
2024:
1864:
1806:Systematische Phylogenie: Wirbelthiere
1512:pangolin poaching, hunting and trading
6947:
6946:
6196:
5726:
4979:
4901:
4135:
4107:from the original on 2 February 2015.
3957:
3641:
3255:Molecular Phylogenetics and Evolution
3061:
3055:
2276:
2211:
1590:armour made of gilded pangolin scales
1568:key ingredient in various medicines.
1548:, promote blood circulation, or help
1378:
732:, the same material from which human
5221:Journal of Zoo and Wildlife Medicine
4408:
4276:(Nigeria), Saturday 18 February 2017
4185:Carrington, Damian (15 April 2013).
4150:from the original on 3 October 2014.
3882:Fletcher, Martin (5 February 2015).
3656:from the original on 8 December 2014
3177:. New York: Facts on File. pp.
2906:10.1111/j.1365-2028.1997.101-89101.x
2614:
2304:
1920:
1871:. Duke University Press. p. 24.
1819:
1634:are susceptible to diseases such as
1506:, their popularity still drives the
564:) are critically endangered, three (
503:, and their diet consists of mainly
4732:"Chinese Medicine and the Pangolin"
4687:
4552:"Civet Cat Slaughter To Fight SARS"
4070:Wassener, Bettina (12 March 2013).
3989:
3919:IUCN Red List of Threatened Species
3686:IUCN Red List of Threatened Species
3162:
2853:10.1002/9780470015902.a0001556.pub2
2843:Rose, K. D.; Gaudin, T. J. (2010).
2279:The Composition of Scientific Words
2270:
2252:IUCN Red List of Threatened Species
2173:"Chinese Medicine and the Pangolin"
2142:10.3897/natureconservation.39.47879
1142:
633:The name "pangolin" comes from the
24:
7200:Extant Paleocene first appearances
5421:"Jackie Chan fights for pangolins"
4836:from the original on 20 March 2017
4793:from the original on 15 April 2014
4779:Steven Novella (25 January 2012).
4701:Mariëtte Le Roux (25 March 2018).
4226:from the original on 18 April 2013
4199:from the original on 16 April 2013
3406:Journal of Vertebrate Paleontology
2762:"Rollin' With the Pangolin – Diet"
2708:10.1111/j.1748-1090.1994.tb03578.x
2514:10.1111/j.1096-3642.1967.tb00508.x
2025:Gaudin, Timothy (28 August 2009).
1738:Journal of Vertebrate Paleontology
25:
7221:
5638:
5567:(2). Pingtung, Taiwan: e0198230.
5262:Choo, S. W.; et al. (2016).
5080:. 23 November 2019. p. 231.
5048:. 23 November 2019. p. 228.
3712:Actman, Jani (20 December 2015).
2766:University of Wisconsin–La Crosse
2343:Wilkinson, Richard James (1932).
2313:Concise Oxford English Dictionary
1923:"Classification of the Edentates"
1306:in the world. This, coupled with
491:Pangolins have large, protective
5607:
5544:
5516:
5467:
5441:
5370:
5356:
5304:
5255:
5211:
5186:
5167:
5155:
5143:
5094:
4980:Maron, Dina Fine (9 June 2020).
4916:from the original on 24 May 2022
4415:Bryner, Jeanna (15 March 2020).
4095:Sutter, John D. (3 April 2014).
3000:10.1111/j.1469-7580.2009.01060.x
2728:International Journal of Zoology
2667:Great Book of the Animal Kingdom
2665:Mondadori, Arnoldo, ed. (1988).
1489:
1324:, while three are classified as
1200:
641:meaning "one who rolls up" from
410:
169:
63:
5555:Manis pentadactyla pentadactyla
5480:Global Ecology and Conservation
5318:Global Ecology and Conservation
5149:Xinhua News (21 November 2018)
5062:
5030:
5004:
4895:
4724:
4544:
4487:
4357:
4264:
4238:
4218:Molland, Judy (16 April 2013).
4211:
4178:
4154:
4136:Watts, Jonathan (25 May 2007).
4129:
4111:
4088:
4063:
4004:
3983:
3938:
3901:
3875:
3831:
3736:
3705:
3635:
3620:
3563:
3195:
3081:
2969:
2877:
2776:
2753:
2683:
2608:
2577:
2559:
2422:
2402:
2285:
2212:Goode, Emilia (31 March 2015).
2205:
2165:
2114:
2044:Springer Science+Business Media
1991:
1961:
1914:
1901:
1579:
925:
688:In ancient India, according to
657:). However, the modern name is
649:"to roll"; it was used for the
5161:China Plus (18 February 2017)
4902:Boyle, Louise (30 June 2020).
4121:. Traffic.org. 17 March 2008.
3743:Cruise, Adam (18 April 2015).
3642:Hance, Jeremy (29 July 2014).
3555:Sutter, John D. (April 2014).
3469:Trafficked Mammals on Earth".
2847:. John Wiley & Sons, Ltd.
2373:Interesting Philippine Animals
2035:Journal of Mammalian Evolution
1888:
1875:
1865:Pearse, Arthur Sperry (1936).
1858:
1848:
1813:
1796:
1777:
1764:
1758:10.1080/02724634.1994.10011592
1726:
1607:Red List of Threatened Species
1117:Gaudin, Emry & Wible, 2009
1066:Gaudin, Emry & Wible, 2009
1055:Gaudin, Emry & Wible, 2009
1032:Gaudin, Emry & Wible, 2009
699:
596:Red List of Threatened Species
27:Mammals of the order Pholidota
13:
1:
7180:Critically endangered animals
5107:Global Environmental Politics
3576:Tropical Conservation Science
3275:10.1016/S1055-7903(03)00118-0
2504:(310). Oxfordshire, England:
2357:National Library of Australia
2277:Brown, Roland Wilbur (1956).
1720:
5582:10.1371/journal.pone.0198230
5399:10.1016/j.biocon.2021.109002
5364:"World Pangolin Day – About"
4742:(3558): 72. 1 January 1938.
4583:Emerging Infectious Diseases
4038:10.1371/journal.pone.0117199
3431:10.1080/02724634.2012.694319
3031:Prothero, Donald R. (2016).
2797:. Corte Madera, California:
2545:10.1016/j.actbio.2016.05.028
2311:Pearsall, Judy, ed. (2002).
2183:(3558): 72. 1 January 1938.
1907:Huxley, Thomas Henry (1872)
1496:traditional Chinese medicine
1419:traditional Chinese medicine
1318:Zoological Society of London
1280:traditional Chinese medicine
1137:
1006:Classification and phylogeny
903:, they ingest small stones (
810:
605:
7:
5501:10.1016/j.gecco.2020.e01395
5453:Mongabay Environmental News
5339:10.1016/j.gecco.2019.e00714
5016:Mongabay Environmental News
3173:The Encyclopedia of Mammals
3088:Grosshuesch, Craig (2012).
2760:Grosshuesch, Craig (2012).
2625:(2). Oxfordshire, England:
1939:10.1126/science.32.810.56.a
1770:Lane, Henry Higgins (1910)
1703:
1409:and is responsible for the
1373:
1205:The first dichotomy in the
1010:
10:
7226:
6145:(Rabbits, hares and pikas)
6100:(Dogs, cats and relatives)
6006:(Tenrecs and golden moles)
5645:ZSL Pangolin Conservation
5173:Xinhua News (8 June 2018)
4554:. CBSNews. 11 January 2004
4335:10.1038/d41586-020-00364-2
3378:10.1016/j.gene.2014.08.040
3039:Princeton University Press
2886:African Journal of Ecology
2696:International Zoo Yearbook
2455:10.1038/s41598-020-71513-w
2300:. Perseus Digital Library.
2042:(4). Heidelberg, Germany:
1486:from a diseased pangolin.
1405:), the virus which causes
1382:
1267:
1263:
1213:) from African pangolins (
1146:
1084:Szalay & Schrenk 1998
1044:Szalay & Schrenk, 1994
914:Some species, such as the
582:) are endangered and two (
299:Szalay & Schrenk, 1994
36:
29:
6955:
6899:
6869:
6831:
6807:
6789:
6763:
6729:
6675:
6651:
6633:
6585:
6563:
6549:
6514:
6498:
6487:
6461:
6413:
6384:
6350:
6325:
6312:
6299:
6286:
6275:
6231:
6125:
6064:
6055:
5994:
5965:
5944:
5931:(Kangaroos and relatives)
5887:
5858:
5837:
5828:
5794:
5762:
5693:25 September 2015 at the
5655:Tree of Life of Pholidota
5101:Shibaike, Takumi (2021).
4520:10.1016/j.cub.2020.03.022
4472:10.1038/s41586-020-2313-x
4308:10.1017/S0030605300028283
3951:The Sydney Morning Herald
3523:10.1186/s12864-017-4140-5
3037:. Princeton, New Jersey:
2052:10.1007/s10914-009-9119-9
1790:Visnyk of Lviv University
1674:
1177:, with which they form a
880:Pangolins have very poor
833:defending itself against
770:mounds and for climbing.
685:(with the same meaning).
619:
288:
281:
277:Ranges of living species
276:
267:
263:
258:
166:Scientific classification
164:
144:
135:
51:
32:Pangolin (disambiguation)
6841:(paraphyletic subfamily)
6643:(paraphyletic subfamily)
5923:(Bilbies and bandicoots)
5180:30 November 2020 at the
3990:Sen (18 February 2020).
3597:10.1177/1940082920985755
3413:(5). Bethesda Maryland:
3303:BMC Evolutionary Biology
3137:10.3897/zookeys.507.6970
2919:Gutteridge, Lee (2008).
2764:. La Crosse, Wisconsin:
2535:. Oxfordshire, England:
1784:Zagorodniuk, I. (2008).
617:– "clad in scales" from
594:) are vulnerable on the
145:Pangolins from families
5815:(Platypus and echidnas)
5675:23 October 2018 at the
5379:Biological Conservation
4167:The Wall Street Journal
3224:10.1126/science.1067179
2627:Oxford University Press
2596:. Ann Arbor, Michigan:
2584:Andrews, James (2011).
2569:. Pangolins.org. 2015.
2506:Oxford University Press
2317:Oxford University Press
2298:A Greek-English Lexicon
2247:"Manidae Family search"
1984:Encyclopædia Britannica
1885:, London, J. Van Voorst
1803:Haeckel, Ernst (1895).
1638:and the development of
1498:preparations. While no
868:
585:Phataginus tetradactyla
7185:Myrmecophagous mammals
6046:(Dugongs and manatees)
5985:(Anteaters and sloths)
4786:Science-Based Medicine
4595:10.3201/eid2707.et2707
4573:Partin, Clyde (2021).
3965:"The CITES Appendices"
3627:van Uhm, D.P. (2016).
3321:10.1186/1471-2148-6-93
3310:(1). London, England:
2598:University of Michigan
2417:Loeb Classical Library
1894:Murray, Andrew (1866)
1792:. Biology (48): 33–43.
1630:
1601:
1287:
1229:to a clade comprising
1221:). Within the former,
959:
938:pup and its mother, a
837:
823:
779:tube-lipped nectar bat
725:
717:
692:, it was known as the
524:threatened by poaching
7152:Paleobiology Database
6799:(paraphyletic family)
5915:(Quolls and dunnarts)
5707:. United States: PBS.
5660:29 April 2021 at the
5529:The Reporter (Taiwan)
5280:10.1101/gr.203521.115
4873:10.1089/acm.2011.0315
4812:Zhouying Jin (2005).
3631:. New York: Springer.
3483:10.1093/jhered/esx097
2992:John Wiley & Sons
2955:. 2015. p. 215.
2948:Wildlife of the World
1881:Edward Newman (1843)
1734:"Abstracts of Papers"
1710:Mammal classification
1625:
1587:
1535:Taiping sheng hui fan
1391:nucleic acid sequence
1326:critically endangered
1277:
1197:(million years) ago.
940:critically endangered
933:
829:
818:
723:
707:
396:, sometimes known as
5120:10.1162/glep_a_00623
4711:Agence France-Presse
3926:: e.T12764A123585318
3845:Conservation Letters
3693:: e.T12763A123584856
3590:: 1940082920985755.
3415:Taylor & Francis
3096:on 23 December 2014.
2990:(4). New York City:
2618:Journal of Mammalogy
2594:Animal Diversity Web
2567:"Meet the Pangolin!"
2069:on 25 September 2015
1921:Gill, Theo. (1910).
918:, use their strong,
846:long-tailed pangolin
822:in defensive posture
785:bone, but is in the
671:Philippine languages
567:Phataginus tricuspis
528:traditional medicine
517:convergent evolution
454:, has three genera:
30:For other uses, see
6875:Propalaeanodontinae
5670:video of a pangolin
5668:National Geographic
5623:. 25 September 2019
5573:2019PLoSO..1498230S
5492:2020GEcoC..2401395A
5391:2021BCons.25509002C
5330:2019GEcoC..2000714C
4986:National Geographic
4748:1938Natur.141R..72.
4512:2020CBio...30E1346Z
4506:(7): 1346–1351.e2.
4463:2020Natur.583..286X
4029:2015PLoSO..1017199B
3755:on 20 December 2015
3749:National Geographic
3724:on 22 December 2015
3718:National Geographic
3471:Journal of Heredity
3423:2012JVPal..32..983K
3267:2003MolPE..28..225A
3216:2001Sci...294.2348M
3210:(5550): 2348–2351.
3128:2015ZooK..507...99H
2951:. London, England:
2898:1997AfJEc..35..361R
2746:10.1155/2014/795062
2735:. London, England:
2447:2020NatSR..1014566L
2189:1938Natur.141R..72.
2128:Nature Conservation
1822:American Naturalist
1750:1994JVPal..14S...1.
1504:mechanism of action
1500:scientific evidence
1479:Amblyomma javanense
1421:was suggested as a
1300:Vietnamese medicine
1296:traditional Chinese
936:Philippine pangolin
840:Most pangolins are
714:Museum of Osteology
573:Manis crassicaudata
540:conservation status
5899:(Monito del monte)
5716:The New York Times
5076:(First ed.).
4663:10.1093/ve/veaa020
4645:(1 January 2020).
4162:"Asia in Pictures"
4077:The New York Times
3912:Manis pentadactyla
3858:10.1111/conl.12389
3813:www.pangolinsg.org
3584:University of Kent
3362:Smutsia temminckii
2983:Journal of Anatomy
2953:Dorling Kindersley
2908:– via EBSCO.
2808:10.7717/peerj.4140
2529:Acta Biomaterialia
2435:Scientific Reports
2219:The New York Times
2095:The New York Times
1631:
1602:
1556:practitioners and
1474:Manis pentadactyla
1385:Origin of COVID-19
1379:COVID-19 infection
1350:or cure cancer or
1313:Manis pentadactyla
1288:
1223:Manis pentadactyla
960:
838:
824:
726:
718:
591:Smutsia temminckii
7167:
7166:
7139:Open Tree of Life
6949:Taxon identifiers
6940:
6939:
6936:
6935:
6932:
6931:
6895:
6894:
6842:
6800:
6725:
6724:
6721:
6720:
6644:
6578:
6545:
6544:
6541:
6540:
6457:
6456:
6453:
6452:
6409:
6408:
6338:See in template:
6223:Mammals of clade
6190:
6189:
6186:
6185:
6182:
6181:
6178:
6177:
6170:
6162:
6154:
6146:
6138:
6117:
6109:
6101:
6093:
6085:
6077:
6047:
6039:
6031:
6023:
6015:
6014:(Elephant shrews)
6007:
5986:
5978:
5959:
5940:
5939:
5932:
5924:
5916:
5908:
5907:(Marsupial moles)
5905:Notoryctemorphia
5900:
5879:
5871:
5868:Paucituberculata
5852:
5824:
5823:
5816:
5455:. 27 January 2021
5274:(10): 1312–1322.
5233:10.1638/2019-0019
5087:978-0-12-815506-6
5055:978-0-12-815506-6
4829:978-1-84150-124-6
4457:(7815): 286–289.
4383:10.3390/v11110979
4222:. London: Care2.
3188:978-0-87196-871-5
3048:978-1-4008-8445-2
2962:978-1-4654-4959-7
2932:978-1-920143-13-8
2326:978-0-19-860572-0
2011:978-0-8018-8221-0
1973:Lydekker, Richard
1669:Manis culionensis
1650:about pangolins.
1519:Ben Cao Jinji Zhu
1432:Asian palm civets
1413:. Researchers in
1411:COVID-19 pandemic
1368:timber harvesting
1304:trafficked animal
1127:
1118:
1103:
1085:
1076:
1067:
1056:
1045:
1033:
1022:
549:Manis culionensis
542:is listed in the
391:
390:
317:Zagorodniuk, 2008
292:list of synonyms:
254:
16:(Redirected from
7217:
7160:
7159:
7147:
7146:
7134:
7133:
7121:
7120:
7108:
7107:
7095:
7094:
7082:
7081:
7069:
7068:
7056:
7055:
7043:
7042:
7030:
7029:
7017:
7016:
7004:
7003:
6991:
6990:
6989:
6976:
6975:
6974:
6944:
6943:
6840:
6837:Metacheiromyinae
6805:
6804:
6798:
6795:Metacheiromyidae
6769:Escavadodontidae
6649:
6648:
6642:
6583:
6582:
6572:
6561:
6560:
6496:
6495:
6425:
6424:
6323:
6322:
6310:
6309:
6297:
6296:
6284:
6283:
6217:
6210:
6203:
6194:
6193:
6168:
6160:
6152:
6144:
6136:
6127:Euarchontoglires
6115:
6107:
6099:
6091:
6083:
6075:
6062:
6061:
6045:
6037:
6029:
6021:
6013:
6005:
5984:
5976:
5963:
5962:
5958:
5957:
5949:
5930:
5922:
5921:Peramelemorphia
5914:
5906:
5898:
5877:
5876:Didelphimorphia
5870:(Shrew opossums)
5869:
5856:
5855:
5851:
5850:
5842:
5835:
5834:
5814:
5805:Australosphenida
5801:
5800:
5747:
5740:
5733:
5724:
5723:
5708:
5633:
5632:
5630:
5628:
5611:
5605:
5604:
5594:
5584:
5548:
5542:
5541:
5539:
5537:
5520:
5514:
5513:
5503:
5471:
5465:
5464:
5462:
5460:
5445:
5439:
5438:
5436:
5434:
5429:. 23 August 2017
5417:
5411:
5410:
5374:
5368:
5367:
5360:
5354:
5353:
5351:
5341:
5308:
5302:
5301:
5291:
5259:
5253:
5252:
5215:
5209:
5208:
5206:
5204:
5190:
5184:
5171:
5165:
5159:
5153:
5147:
5141:
5140:
5122:
5098:
5092:
5091:
5066:
5060:
5059:
5034:
5028:
5027:
5025:
5023:
5008:
5002:
5001:
4999:
4997:
4988:. Archived from
4977:
4971:
4970:
4968:
4966:
4941:
4926:
4925:
4923:
4921:
4907:
4899:
4893:
4892:
4855:
4849:
4848:
4843:
4841:
4809:
4803:
4802:
4800:
4798:
4776:
4770:
4769:
4759:
4757:10.1038/141072b0
4728:
4722:
4721:
4719:
4717:
4707:The Jakarta Post
4698:
4685:
4684:
4674:
4643:Zhang, Yong-Zhen
4639:Holmes, Edward C
4634:
4625:
4624:
4614:
4570:
4564:
4563:
4561:
4559:
4548:
4542:
4541:
4531:
4491:
4485:
4484:
4474:
4441:
4432:
4431:
4429:
4427:
4412:
4406:
4405:
4395:
4385:
4361:
4355:
4354:
4322:
4313:
4312:
4310:
4286:
4277:
4268:
4262:
4261:
4259:
4257:
4252:. 26 August 2016
4242:
4236:
4235:
4233:
4231:
4215:
4209:
4208:
4206:
4204:
4192:Associated Press
4182:
4176:
4175:
4158:
4152:
4151:
4133:
4127:
4126:
4115:
4109:
4108:
4092:
4086:
4085:
4067:
4061:
4060:
4050:
4040:
4008:
4002:
4001:
3987:
3981:
3980:
3978:
3976:
3961:
3955:
3954:
3942:
3936:
3935:
3933:
3931:
3905:
3899:
3898:
3896:
3894:
3879:
3873:
3872:
3870:
3860:
3835:
3829:
3828:
3826:
3824:
3815:. Archived from
3805:
3794:
3793:
3791:
3789:
3774:
3765:
3764:
3762:
3760:
3751:. Archived from
3740:
3734:
3733:
3731:
3729:
3720:. Archived from
3709:
3703:
3702:
3700:
3698:
3672:
3666:
3665:
3663:
3661:
3639:
3633:
3632:
3624:
3618:
3617:
3599:
3567:
3561:
3560:
3552:
3546:
3545:
3535:
3525:
3501:
3495:
3494:
3465:
3459:
3452:
3443:
3442:
3396:
3390:
3389:
3357:
3344:
3343:
3333:
3323:
3293:
3287:
3286:
3250:
3244:
3243:
3199:
3193:
3192:
3176:
3166:
3160:
3159:
3149:
3139:
3111:
3098:
3097:
3085:
3079:
3078:
3076:
3074:
3059:
3053:
3052:
3028:
3022:
3021:
3011:
2973:
2967:
2966:
2943:
2937:
2936:
2916:
2910:
2909:
2881:
2875:
2874:
2840:
2831:
2830:
2820:
2810:
2780:
2774:
2773:
2757:
2751:
2750:
2748:
2718:
2712:
2711:
2687:
2681:
2680:
2662:
2647:
2646:
2612:
2606:
2605:
2590:: tree pangolin"
2581:
2575:
2574:
2563:
2557:
2556:
2524:
2518:
2517:
2491:
2485:
2484:
2474:
2426:
2420:
2406:
2400:
2399:
2397:
2395:
2390:on 27 March 2020
2386:. Archived from
2367:
2361:
2360:
2340:
2331:
2330:
2319:. p. 1030.
2308:
2302:
2301:
2289:
2283:
2282:
2274:
2268:
2267:
2265:
2263:
2243:
2232:
2231:
2229:
2227:
2209:
2203:
2202:
2200:
2198:10.1038/141072b0
2169:
2163:
2162:
2144:
2118:
2112:
2111:
2109:
2107:
2085:
2079:
2078:
2076:
2074:
2068:
2062:. Archived from
2031:
2022:
2016:
2015:
1995:
1989:
1988:
1987:(11th ed.).
1980:
1978:"Pangolin"
1969:Thomas, Oldfield
1965:
1959:
1958:
1918:
1912:
1905:
1899:
1892:
1886:
1879:
1873:
1872:
1862:
1856:
1852:
1846:
1845:
1828:(272): 657–664.
1817:
1811:
1810:
1800:
1794:
1793:
1781:
1775:
1768:
1762:
1761:
1730:
1685:Yanping Township
1644:intestinal worms
1442:showed that the
1440:genetic analysis
1364:overexploitation
1231:M. crassicaudata
1143:Among placentals
1125:
1116:
1101:
1083:
1074:
1065:
1054:
1043:
1031:
1020:
997:
996:
992:
989:
983:
982:
978:
975:
806:
805:
801:
622:
621:
579:Smutsia gigantea
445:
444:
441:
440:
437:
434:
431:
428:
425:
422:
419:
416:
384:
378:
372:
366:
360:
354:
348:
345:Pholidotiformes
342:
336:
330:
324:
318:
312:
306:
300:
272:
249:
174:
173:
140:
125:
62:
55:Temporal range:
49:
48:
21:
7225:
7224:
7220:
7219:
7218:
7216:
7215:
7214:
7195:Rolling animals
7170:
7169:
7168:
7163:
7155:
7150:
7142:
7137:
7129:
7124:
7116:
7111:
7103:
7098:
7090:
7085:
7077:
7072:
7064:
7059:
7051:
7046:
7038:
7033:
7025:
7020:
7012:
7007:
6999:
6994:
6985:
6984:
6979:
6970:
6969:
6964:
6951:
6941:
6928:
6906:
6891:
6877:
6872:
6865:
6843:
6839:
6834:
6827:
6801:
6797:
6792:
6785:
6771:
6766:
6759:
6737:
6732:
6717:
6687:
6685:
6671:
6645:
6641:
6639:Epoicotheriinae
6636:
6629:
6579:
6571:
6569:Epoicotheriidae
6566:
6557:
6552:
6537:
6529:Pholidota sp. (
6521:
6510:
6492:
6491:representatives
6490:
6483:
6469:
6467:Eurotamanduidae
6464:
6449:
6435:
6430:
6421:
6416:
6405:
6391:
6380:
6358:
6353:
6346:
6332:
6328:
6319:
6315:
6306:
6302:
6293:
6289:
6280:
6278:Pholidotamorpha
6271:
6267:Pholidotamorpha
6227:
6225:Pholidotamorpha
6221:
6191:
6174:
6121:
6106:Perissodactyla
6051:
5990:
5951:
5950:
5948:
5936:
5913:Dasyuromorphia
5897:Microbiotheria
5889:Australidelphia
5883:
5844:
5843:
5841:
5820:
5790:
5758:
5751:
5719:, 6 March 2020.
5700:
5695:Wayback Machine
5677:Wayback Machine
5662:Wayback Machine
5641:
5636:
5626:
5624:
5613:
5612:
5608:
5549:
5545:
5535:
5533:
5522:
5521:
5517:
5472:
5468:
5458:
5456:
5447:
5446:
5442:
5432:
5430:
5419:
5418:
5414:
5375:
5371:
5362:
5361:
5357:
5309:
5305:
5268:Genome Research
5260:
5256:
5216:
5212:
5202:
5200:
5198:www.traffic.org
5192:
5191:
5187:
5182:Wayback Machine
5172:
5168:
5160:
5156:
5148:
5144:
5099:
5095:
5088:
5068:
5067:
5063:
5056:
5036:
5035:
5031:
5021:
5019:
5010:
5009:
5005:
4995:
4993:
4992:on 10 June 2020
4978:
4974:
4964:
4962:
4960:
4942:
4929:
4919:
4917:
4910:The Independent
4900:
4896:
4867:(12): 1108–20.
4856:
4852:
4839:
4837:
4830:
4820:Intellect Books
4810:
4806:
4796:
4794:
4777:
4773:
4730:
4729:
4725:
4715:
4713:
4699:
4688:
4651:Virus Evolution
4635:
4628:
4571:
4567:
4557:
4555:
4550:
4549:
4545:
4500:Current Biology
4492:
4488:
4442:
4435:
4425:
4423:
4413:
4409:
4362:
4358:
4323:
4316:
4287:
4280:
4269:
4265:
4255:
4253:
4244:
4243:
4239:
4229:
4227:
4216:
4212:
4202:
4200:
4183:
4179:
4170:. 27 May 2012.
4160:
4159:
4155:
4134:
4130:
4117:
4116:
4112:
4093:
4089:
4068:
4064:
4023:(1): e0117199.
4009:
4005:
3988:
3984:
3974:
3972:
3963:
3962:
3958:
3943:
3939:
3929:
3927:
3906:
3902:
3892:
3890:
3880:
3876:
3836:
3832:
3822:
3820:
3819:on 10 July 2017
3807:
3806:
3797:
3787:
3785:
3775:
3768:
3758:
3756:
3741:
3737:
3727:
3725:
3710:
3706:
3696:
3694:
3673:
3669:
3659:
3657:
3640:
3636:
3625:
3621:
3568:
3564:
3553:
3549:
3502:
3498:
3466:
3462:
3453:
3446:
3397:
3393:
3358:
3347:
3294:
3290:
3251:
3247:
3200:
3196:
3189:
3167:
3163:
3122:(507): 99–114.
3112:
3101:
3086:
3082:
3072:
3070:
3060:
3056:
3049:
3041:. p. 118.
3029:
3025:
2974:
2970:
2963:
2945:
2944:
2940:
2933:
2917:
2913:
2882:
2878:
2863:
2841:
2834:
2781:
2777:
2758:
2754:
2737:Wiley-Blackwell
2719:
2715:
2688:
2684:
2677:
2663:
2650:
2635:10.2307/1382356
2613:
2609:
2588:Manis tricuspis
2582:
2578:
2565:
2564:
2560:
2525:
2521:
2492:
2488:
2427:
2423:
2407:
2403:
2393:
2391:
2384:
2368:
2364:
2341:
2334:
2327:
2309:
2305:
2290:
2286:
2275:
2271:
2261:
2259:
2245:
2244:
2235:
2225:
2223:
2222:. New York City
2210:
2206:
2171:
2170:
2166:
2119:
2115:
2105:
2103:
2086:
2082:
2072:
2070:
2066:
2029:
2023:
2019:
2012:
1996:
1992:
1966:
1962:
1919:
1915:
1906:
1902:
1893:
1889:
1880:
1876:
1863:
1859:
1853:
1849:
1818:
1814:
1801:
1797:
1782:
1778:
1769:
1765:
1732:
1731:
1727:
1723:
1715:Pholidotamorpha
1706:
1677:
1594:Prince of Wales
1582:
1550:lactating women
1492:
1458:
1387:
1381:
1376:
1272:
1266:
1258:laurasiatherian
1203:
1191:Pholidotamorpha
1151:
1149:Pholidotamorpha
1145:
1140:
1041:Eurotamanduidae
1013:
1008:
994:
990:
987:
985:
980:
976:
973:
971:
928:
871:
831:Indian pangolin
820:Ground pangolin
813:
803:
799:
798:
702:
608:
555:M. pentadactyla
413:
409:
398:scaly anteaters
387:
382:
376:
370:
364:
358:
352:
346:
340:
334:
328:
322:
316:
310:
304:
298:
294:
293:
248:
236:Pholidotamorpha
168:
159:Eurotamanduidae
131:
124:
123:
118:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
57:
56:
53:
45:
35:
28:
23:
22:
15:
12:
11:
5:
7223:
7213:
7212:
7207:
7202:
7197:
7192:
7187:
7182:
7165:
7164:
7162:
7161:
7148:
7135:
7122:
7109:
7096:
7083:
7070:
7057:
7044:
7031:
7018:
7005:
6992:
6977:
6961:
6959:
6953:
6952:
6938:
6937:
6934:
6933:
6930:
6929:
6927:
6926:
6918:
6909:
6907:
6903:Incertae sedis
6900:
6897:
6896:
6893:
6892:
6890:
6889:
6886:Propalaeanodon
6880:
6878:
6870:
6867:
6866:
6864:
6863:
6855:
6846:
6844:
6832:
6829:
6828:
6826:
6825:
6817:
6808:
6802:
6790:
6787:
6786:
6784:
6783:
6774:
6772:
6764:
6761:
6760:
6758:
6757:
6749:
6740:
6738:
6735:Ernanodontidae
6730:
6727:
6726:
6723:
6722:
6719:
6718:
6716:
6715:
6707:
6699:
6690:
6688:
6676:
6673:
6672:
6670:
6669:
6661:
6652:
6646:
6634:
6631:
6630:
6628:
6627:
6619:
6611:
6603:
6595:
6586:
6580:
6564:
6558:
6550:
6547:
6546:
6543:
6542:
6539:
6538:
6536:
6535:
6524:
6522:
6518:Incertae sedis
6515:
6512:
6511:
6509:
6508:
6499:
6493:
6488:
6485:
6484:
6482:
6481:
6472:
6470:
6462:
6459:
6458:
6455:
6454:
6451:
6450:
6448:
6447:
6438:
6436:
6428:
6422:
6414:
6411:
6410:
6407:
6406:
6404:
6403:
6394:
6392:
6388:Incertae sedis
6385:
6382:
6381:
6379:
6378:
6370:
6361:
6359:
6351:
6348:
6347:
6345:
6344:
6335:
6333:
6326:
6320:
6313:
6307:
6300:
6294:
6287:
6281:
6276:
6273:
6272:
6270:
6269:
6260:
6251:
6245:
6239:
6232:
6229:
6228:
6220:
6219:
6212:
6205:
6197:
6188:
6187:
6184:
6183:
6180:
6179:
6176:
6175:
6173:
6172:
6164:
6156:
6148:
6140:
6131:
6129:
6123:
6122:
6120:
6119:
6111:
6103:
6095:
6087:
6079:
6070:
6068:
6066:Laurasiatheria
6059:
6053:
6052:
6050:
6049:
6041:
6033:
6025:
6020:Tubulidentata
6017:
6012:Macroscelidea
6009:
6000:
5998:
5992:
5991:
5989:
5988:
5980:
5971:
5969:
5960:
5942:
5941:
5938:
5937:
5935:
5934:
5929:Diprotodontia
5926:
5918:
5910:
5902:
5893:
5891:
5885:
5884:
5882:
5881:
5873:
5864:
5862:
5853:
5832:
5826:
5825:
5822:
5821:
5819:
5818:
5809:
5807:
5798:
5792:
5791:
5789:
5788:
5782:
5776:
5770:
5763:
5760:
5759:
5750:
5749:
5742:
5735:
5727:
5721:
5720:
5709:
5698:
5685:
5679:
5664:
5652:
5647:
5640:
5639:External links
5637:
5635:
5634:
5606:
5543:
5532:. 23 June 2019
5515:
5466:
5440:
5412:
5369:
5355:
5303:
5254:
5227:(3): 678–687.
5210:
5185:
5166:
5154:
5142:
5093:
5086:
5078:Academic Press
5061:
5054:
5046:Academic Press
5029:
5003:
4972:
4958:
4927:
4894:
4850:
4828:
4822:. p. 36.
4804:
4771:
4723:
4686:
4657:(1): veaa020.
4626:
4577:Sunda pangolin
4565:
4543:
4486:
4433:
4407:
4356:
4314:
4278:
4263:
4237:
4210:
4177:
4153:
4128:
4110:
4087:
4062:
4003:
3982:
3956:
3937:
3900:
3874:
3830:
3795:
3766:
3735:
3704:
3679:Manis javanica
3667:
3634:
3619:
3562:
3547:
3496:
3477:(4): 347–359.
3460:
3444:
3391:
3345:
3312:BioMed Central
3288:
3261:(2): 225–240.
3245:
3194:
3187:
3161:
3099:
3080:
3054:
3047:
3023:
2968:
2961:
2938:
2931:
2911:
2892:(4): 361–369.
2876:
2862:978-0470015902
2861:
2832:
2799:O'Reilly Media
2775:
2752:
2713:
2702:(1): 248–251.
2682:
2676:978-0517667910
2675:
2648:
2607:
2576:
2558:
2519:
2486:
2421:
2401:
2382:
2362:
2332:
2325:
2303:
2284:
2269:
2233:
2204:
2164:
2113:
2080:
2017:
2010:
1990:
1960:
1913:
1900:
1887:
1874:
1857:
1847:
1834:10.1086/274985
1812:
1795:
1776:
1763:
1744:: 1–58. 1994.
1724:
1722:
1719:
1718:
1717:
1712:
1705:
1702:
1676:
1673:
1581:
1578:
1527:Wai Tai Mi Yao
1491:
1488:
1470:Manis javanica
1457:
1448:
1380:
1377:
1375:
1372:
1270:Pangolin trade
1265:
1262:
1202:
1199:
1144:
1141:
1139:
1136:
1135:
1134:
1133:
1132:
1131:
1130:
1129:
1128:
1110:Superfamily: †
1108:
1107:
1106:
1105:
1104:
1090:Incertae sedis
1086:
1077:
1046:
1034:
1012:
1009:
1007:
1004:
952:pangolin trade
927:
924:
873:Pangolins are
870:
867:
812:
809:
775:giant anteater
701:
698:
655:Manis javanica
651:Sunda pangolin
607:
604:
522:Pangolins are
389:
388:
386:
385:
379:
373:
367:
361:
355:
349:
343:
339:Pholidotheria
337:
331:
325:
319:
313:
307:
301:
291:
290:
289:
286:
285:
279:
278:
274:
273:
265:
264:
261:
260:
256:
255:
243:
239:
238:
233:
226:
225:
220:
216:
215:
210:
206:
205:
200:
196:
195:
190:
186:
185:
180:
176:
175:
162:
161:
142:
141:
133:
132:
119:
114:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
54:
26:
9:
6:
4:
3:
2:
7222:
7211:
7208:
7206:
7203:
7201:
7198:
7196:
7193:
7191:
7188:
7186:
7183:
7181:
7178:
7177:
7175:
7158:
7153:
7149:
7145:
7140:
7136:
7132:
7127:
7123:
7119:
7114:
7110:
7106:
7101:
7097:
7093:
7088:
7084:
7080:
7075:
7071:
7067:
7062:
7058:
7054:
7049:
7045:
7041:
7036:
7032:
7028:
7023:
7019:
7015:
7010:
7006:
7002:
6997:
6993:
6988:
6982:
6978:
6973:
6967:
6963:
6962:
6960:
6958:
6954:
6950:
6945:
6925:
6924:
6919:
6917:
6916:
6911:
6910:
6908:
6905:
6904:
6898:
6888:
6887:
6882:
6881:
6879:
6876:
6868:
6862:
6861:
6856:
6854:
6853:
6852:Metacheiromys
6848:
6847:
6845:
6838:
6830:
6824:
6823:
6818:
6816:
6815:
6810:
6809:
6806:
6803:
6796:
6788:
6782:
6781:
6776:
6775:
6773:
6770:
6762:
6756:
6755:
6750:
6748:
6747:
6742:
6741:
6739:
6736:
6728:
6714:
6713:
6708:
6706:
6705:
6700:
6698:
6697:
6696:Epoicotherium
6692:
6691:
6689:
6684:
6680:
6679:Epoicotherium
6674:
6668:
6667:
6666:Tetrapassalus
6662:
6660:
6659:
6658:Pentapassalus
6654:
6653:
6650:
6647:
6640:
6632:
6626:
6625:
6620:
6618:
6617:
6612:
6610:
6609:
6608:Auroratherium
6604:
6602:
6601:
6596:
6594:
6593:
6592:Alocodontulum
6588:
6587:
6584:
6581:
6576:
6570:
6562:
6559:
6556:
6555:Palaeanodonta
6548:
6534:
6532:
6526:
6525:
6523:
6520:
6519:
6513:
6507:
6506:
6501:
6500:
6497:
6494:
6486:
6480:
6479:
6474:
6473:
6471:
6468:
6460:
6446:
6445:
6440:
6439:
6437:
6434:
6426:
6423:
6420:
6412:
6402:
6401:
6396:
6395:
6393:
6390:
6389:
6383:
6377:
6376:
6371:
6369:
6368:
6363:
6362:
6360:
6357:
6356:Patriomanidae
6349:
6343:
6342:
6337:
6336:
6334:
6331:
6324:
6321:
6318:
6311:
6308:
6305:
6298:
6295:
6292:
6285:
6282:
6279:
6274:
6268:
6264:
6261:
6259:
6255:
6252:
6250:
6246:
6244:
6240:
6238:
6234:
6233:
6230:
6226:
6218:
6213:
6211:
6206:
6204:
6199:
6198:
6195:
6171:
6165:
6163:
6157:
6155:
6149:
6147:
6141:
6139:
6133:
6132:
6130:
6128:
6124:
6118:
6114:Artiodactyla
6112:
6110:
6104:
6102:
6096:
6094:
6088:
6086:
6080:
6078:
6074:Eulipotyphla
6072:
6071:
6069:
6067:
6063:
6060:
6058:
6057:Boreoeutheria
6054:
6048:
6042:
6040:
6034:
6032:
6026:
6024:
6018:
6016:
6010:
6008:
6004:Afrosoricida
6002:
6001:
5999:
5997:
5993:
5987:
5981:
5979:
5973:
5972:
5970:
5968:
5964:
5961:
5955:
5947:
5943:
5933:
5927:
5925:
5919:
5917:
5911:
5909:
5903:
5901:
5895:
5894:
5892:
5890:
5886:
5880:
5874:
5872:
5866:
5865:
5863:
5861:
5857:
5854:
5848:
5840:
5836:
5833:
5831:
5827:
5817:
5811:
5810:
5808:
5806:
5802:
5799:
5797:
5793:
5787:
5783:
5781:
5777:
5775:
5771:
5769:
5765:
5764:
5761:
5756:
5748:
5743:
5741:
5736:
5734:
5729:
5728:
5725:
5718:
5717:
5713:
5710:
5706:
5705:
5699:
5696:
5692:
5689:
5686:
5683:
5680:
5678:
5674:
5671:
5669:
5665:
5663:
5659:
5656:
5653:
5651:
5648:
5646:
5643:
5642:
5622:
5621:
5620:The News Lens
5616:
5610:
5602:
5598:
5593:
5588:
5583:
5578:
5574:
5570:
5566:
5562:
5558:
5556:
5547:
5531:
5530:
5525:
5519:
5511:
5507:
5502:
5497:
5493:
5489:
5485:
5481:
5477:
5470:
5454:
5450:
5444:
5428:
5427:
5422:
5416:
5408:
5404:
5400:
5396:
5392:
5388:
5384:
5380:
5373:
5365:
5359:
5350:
5345:
5340:
5335:
5331:
5327:
5323:
5319:
5315:
5307:
5299:
5295:
5290:
5285:
5281:
5277:
5273:
5269:
5265:
5258:
5250:
5246:
5242:
5238:
5234:
5230:
5226:
5222:
5219:Tricuspis)".
5214:
5199:
5195:
5189:
5183:
5179:
5176:
5170:
5164:
5158:
5152:
5146:
5138:
5134:
5130:
5126:
5121:
5116:
5112:
5108:
5104:
5097:
5089:
5083:
5079:
5075:
5071:
5065:
5057:
5051:
5047:
5043:
5039:
5033:
5017:
5013:
5007:
4991:
4987:
4983:
4976:
4961:
4959:9780128155073
4955:
4951:
4947:
4940:
4938:
4936:
4934:
4932:
4915:
4911:
4906:
4898:
4890:
4886:
4882:
4878:
4874:
4870:
4866:
4862:
4854:
4847:
4835:
4831:
4825:
4821:
4817:
4816:
4808:
4792:
4788:
4787:
4782:
4775:
4767:
4763:
4758:
4753:
4749:
4745:
4741:
4737:
4733:
4727:
4712:
4708:
4704:
4697:
4695:
4693:
4691:
4682:
4678:
4673:
4668:
4664:
4660:
4656:
4652:
4648:
4644:
4640:
4633:
4631:
4622:
4618:
4613:
4608:
4604:
4600:
4596:
4592:
4588:
4584:
4580:
4578:
4575:"Etymologia:
4569:
4553:
4547:
4539:
4535:
4530:
4525:
4521:
4517:
4513:
4509:
4505:
4501:
4497:
4490:
4482:
4478:
4473:
4468:
4464:
4460:
4456:
4452:
4448:
4440:
4438:
4422:
4418:
4411:
4403:
4399:
4394:
4389:
4384:
4379:
4375:
4371:
4367:
4360:
4352:
4348:
4344:
4340:
4336:
4332:
4328:
4321:
4319:
4309:
4304:
4300:
4296:
4292:
4285:
4283:
4275:
4274:
4267:
4251:
4247:
4241:
4225:
4221:
4214:
4198:
4194:
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4188:
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4169:
4168:
4163:
4157:
4149:
4145:
4144:
4139:
4132:
4124:
4120:
4114:
4106:
4102:
4098:
4091:
4083:
4079:
4078:
4073:
4066:
4058:
4054:
4049:
4044:
4039:
4034:
4030:
4026:
4022:
4018:
4014:
4007:
3999:
3998:
3993:
3986:
3970:
3966:
3960:
3952:
3948:
3941:
3925:
3921:
3920:
3915:
3913:
3904:
3889:
3885:
3878:
3869:
3864:
3859:
3854:
3851:(2): e12389.
3850:
3846:
3842:
3834:
3818:
3814:
3810:
3809:"Action Plan"
3804:
3802:
3800:
3784:
3780:
3773:
3771:
3754:
3750:
3746:
3739:
3723:
3719:
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3682:
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3671:
3655:
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3645:
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3630:
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3615:
3611:
3607:
3603:
3598:
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3585:
3581:
3577:
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3566:
3558:
3551:
3543:
3539:
3534:
3529:
3524:
3519:
3515:
3511:
3507:
3500:
3492:
3488:
3484:
3480:
3476:
3472:
3464:
3458:
3451:
3449:
3440:
3436:
3432:
3428:
3424:
3420:
3416:
3412:
3408:
3407:
3402:
3395:
3387:
3383:
3379:
3375:
3371:
3367:
3363:
3356:
3354:
3352:
3350:
3341:
3337:
3332:
3327:
3322:
3317:
3313:
3309:
3305:
3304:
3299:
3292:
3284:
3280:
3276:
3272:
3268:
3264:
3260:
3256:
3249:
3241:
3237:
3233:
3229:
3225:
3221:
3217:
3213:
3209:
3205:
3198:
3190:
3184:
3180:
3175:
3174:
3165:
3157:
3153:
3148:
3143:
3138:
3133:
3129:
3125:
3121:
3117:
3110:
3108:
3106:
3104:
3095:
3091:
3084:
3069:
3065:
3058:
3050:
3044:
3040:
3036:
3035:
3027:
3019:
3015:
3010:
3005:
3001:
2997:
2993:
2989:
2985:
2984:
2979:
2972:
2964:
2958:
2954:
2950:
2949:
2942:
2934:
2928:
2924:
2923:
2915:
2907:
2903:
2899:
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2891:
2887:
2880:
2872:
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2828:
2824:
2819:
2814:
2809:
2804:
2800:
2796:
2792:
2791:
2786:
2779:
2771:
2767:
2763:
2756:
2747:
2742:
2738:
2734:
2730:
2729:
2724:
2717:
2709:
2705:
2701:
2697:
2693:
2686:
2678:
2672:
2668:
2661:
2659:
2657:
2655:
2653:
2644:
2640:
2636:
2632:
2628:
2624:
2620:
2619:
2611:
2603:
2599:
2595:
2591:
2589:
2580:
2572:
2568:
2562:
2554:
2550:
2546:
2542:
2538:
2534:
2530:
2523:
2515:
2511:
2507:
2503:
2499:
2498:
2490:
2482:
2478:
2473:
2468:
2464:
2460:
2456:
2452:
2448:
2444:
2440:
2436:
2432:
2425:
2418:
2414:
2410:
2405:
2389:
2385:
2379:
2375:
2374:
2366:
2358:
2354:
2350:
2346:
2339:
2337:
2328:
2322:
2318:
2314:
2307:
2299:
2295:
2288:
2280:
2273:
2258:
2254:
2253:
2248:
2242:
2240:
2238:
2221:
2220:
2215:
2208:
2199:
2194:
2190:
2186:
2182:
2178:
2174:
2168:
2160:
2156:
2152:
2148:
2143:
2138:
2134:
2130:
2129:
2124:
2117:
2101:
2097:
2096:
2091:
2084:
2065:
2061:
2057:
2053:
2049:
2045:
2041:
2037:
2036:
2028:
2021:
2013:
2007:
2003:
2002:
1994:
1986:
1985:
1979:
1974:
1970:
1964:
1956:
1952:
1948:
1944:
1940:
1936:
1932:
1928:
1924:
1917:
1910:
1904:
1897:
1891:
1884:
1878:
1870:
1869:
1861:
1851:
1843:
1839:
1835:
1831:
1827:
1823:
1816:
1808:
1807:
1799:
1791:
1787:
1780:
1773:
1767:
1759:
1755:
1751:
1747:
1743:
1739:
1735:
1729:
1725:
1716:
1713:
1711:
1708:
1707:
1701:
1698:
1695:townships in
1694:
1690:
1686:
1681:
1672:
1670:
1665:
1663:
1661:
1656:
1651:
1647:
1645:
1641:
1637:
1629:
1624:
1620:
1618:
1613:
1608:
1599:
1595:
1591:
1586:
1577:
1575:
1569:
1566:
1565:pharmacopoeia
1563:The official
1561:
1559:
1555:
1554:folk medicine
1551:
1547:
1543:
1538:
1536:
1532:
1528:
1524:
1520:
1515:
1513:
1509:
1505:
1501:
1497:
1490:Folk medicine
1487:
1485:
1481:
1480:
1475:
1471:
1467:
1463:
1456:
1452:
1447:
1445:
1444:spike protein
1441:
1437:
1433:
1428:
1427:coronaviruses
1424:
1420:
1416:
1412:
1408:
1404:
1400:
1399:coronaviruses
1397:belonging to
1396:
1395:spike protein
1392:
1386:
1371:
1369:
1365:
1361:
1357:
1353:
1349:
1344:
1342:
1337:
1333:
1331:
1327:
1323:
1319:
1315:
1314:
1309:
1308:deforestation
1305:
1301:
1297:
1293:
1285:
1281:
1276:
1271:
1261:
1259:
1255:
1251:
1246:
1244:
1240:
1236:
1232:
1228:
1224:
1220:
1216:
1212:
1208:
1201:Among Manidae
1198:
1196:
1192:
1188:
1187:Palaeanodonts
1184:
1180:
1176:
1172:
1168:
1164:
1161:(neotropical
1160:
1156:
1150:
1124:
1120:
1119:
1115:
1114:
1109:
1100:
1099:
1094:
1093:
1092:
1091:
1087:
1082:
1081:Patriomanidae
1078:
1073:
1069:
1068:
1064:
1063:
1059:Superfamily:
1058:
1057:
1053:
1052:
1047:
1042:
1039:
1035:
1030:
1029:
1024:
1023:
1019:
1015:
1014:
1003:
1001:
968:
964:
957:
954:to China and
953:
949:
945:
941:
937:
932:
923:
921:
917:
916:tree pangolin
912:
910:
906:
902:
898:
893:
891:
887:
883:
878:
876:
875:insectivorous
866:
864:
859:
857:
853:
851:
847:
843:
836:
835:Asiatic lions
832:
828:
821:
817:
808:
796:
792:
788:
784:
780:
776:
771:
769:
765:
761:
757:
752:
750:
746:
742:
739:
735:
731:
722:
715:
711:
706:
697:
695:
691:
686:
684:
680:
676:
672:
669:; and in the
668:
664:
660:
656:
652:
648:
644:
640:
636:
631:
629:
625:
616:
613:
612:Ancient Greek
603:
601:
597:
593:
592:
587:
586:
581:
580:
575:
574:
569:
568:
563:
562:
557:
556:
551:
550:
546:tier. Three (
545:
541:
537:
533:
532:deforestation
529:
525:
520:
518:
514:
510:
506:
502:
498:
494:
489:
487:
483:
479:
475:
471:
470:
465:
464:
459:
458:
453:
449:
443:
407:
404:of the order
403:
399:
395:
383:Haeckel, 1895
380:
374:
368:
362:
356:
350:
344:
341:Haeckel, 1895
338:
332:
329:Haeckel, 1895
326:
323:Haeckel, 1895
320:
314:
311:Gervais, 1854
308:
302:
296:
295:
287:
284:
280:
275:
271:
266:
262:
257:
252:
247:
244:
241:
240:
237:
234:
231:
228:
227:
224:
221:
218:
217:
214:
211:
208:
207:
204:
201:
198:
197:
194:
191:
188:
187:
184:
181:
178:
177:
172:
167:
163:
160:
156:
152:
151:Patriomanidae
148:
143:
139:
134:
129:
122:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
61:
50:
47:
43:
41:
33:
19:
6956:
6921:
6915:Arcticanodon
6913:
6901:
6884:
6858:
6850:
6820:
6814:Brachianodon
6812:
6778:
6752:
6746:Asiabradypus
6744:
6710:
6704:Molaetherium
6702:
6694:
6682:
6678:
6664:
6656:
6622:
6614:
6606:
6598:
6590:
6575:paraphyletic
6530:
6516:
6503:
6478:Eurotamandua
6476:
6442:
6415:Superfamily:
6398:
6386:
6373:
6365:
6339:
6314:Superfamily:
6290:
6262:
6253:
6153:(Treeshrews)
6089:
6036:Proboscidea
5977:(Armadillos)
5860:Ameridelphia
5813:Monotremata
5714:
5703:
5667:
5625:. Retrieved
5618:
5609:
5564:
5560:
5554:
5546:
5534:. Retrieved
5527:
5518:
5483:
5479:
5469:
5457:. Retrieved
5452:
5443:
5431:. Retrieved
5424:
5415:
5382:
5378:
5372:
5358:
5321:
5317:
5306:
5271:
5267:
5257:
5224:
5220:
5213:
5201:. Retrieved
5197:
5188:
5169:
5157:
5145:
5113:(2): 45–69.
5110:
5106:
5096:
5073:
5064:
5041:
5032:
5020:. Retrieved
5015:
5006:
4994:. Retrieved
4990:the original
4985:
4975:
4963:. Retrieved
4949:
4918:. Retrieved
4909:
4897:
4864:
4860:
4853:
4845:
4838:. Retrieved
4814:
4807:
4795:. Retrieved
4784:
4774:
4739:
4735:
4726:
4714:. Retrieved
4706:
4654:
4650:
4586:
4582:
4576:
4568:
4556:. Retrieved
4546:
4503:
4499:
4489:
4454:
4450:
4424:. Retrieved
4410:
4373:
4369:
4359:
4326:
4301:(1): 43–50.
4298:
4294:
4271:
4266:
4254:. Retrieved
4249:
4240:
4228:. Retrieved
4213:
4201:. Retrieved
4190:
4180:
4165:
4156:
4143:The Guardian
4141:
4131:
4113:
4090:
4075:
4065:
4020:
4016:
4006:
3995:
3985:
3973:. Retrieved
3968:
3959:
3950:
3940:
3928:. Retrieved
3923:
3917:
3911:
3903:
3891:. Retrieved
3887:
3877:
3848:
3844:
3833:
3823:15 September
3821:. Retrieved
3817:the original
3812:
3786:. Retrieved
3782:
3757:. Retrieved
3753:the original
3748:
3738:
3726:. Retrieved
3722:the original
3717:
3707:
3695:. Retrieved
3690:
3684:
3678:
3670:
3658:. Retrieved
3647:
3637:
3628:
3622:
3579:
3575:
3565:
3550:
3513:
3510:BMC Genomics
3509:
3499:
3474:
3470:
3463:
3417:: 983–1001.
3410:
3404:
3400:
3394:
3372:(1): 49–54.
3369:
3365:
3361:
3307:
3301:
3291:
3258:
3254:
3248:
3207:
3203:
3197:
3172:
3164:
3119:
3094:the original
3083:
3071:. Retrieved
3067:
3057:
3033:
3026:
2987:
2981:
2971:
2947:
2941:
2921:
2914:
2889:
2885:
2879:
2844:
2794:
2788:
2778:
2755:
2732:
2726:
2716:
2699:
2695:
2691:
2685:
2666:
2622:
2616:
2610:
2593:
2587:
2579:
2561:
2532:
2528:
2522:
2501:
2495:
2489:
2441:(1): 14566.
2438:
2434:
2424:
2412:
2404:
2392:. Retrieved
2388:the original
2372:
2365:
2348:
2345:"tĕnggiling"
2312:
2306:
2297:
2287:
2278:
2272:
2260:. Retrieved
2250:
2224:. Retrieved
2217:
2207:
2180:
2176:
2167:
2132:
2126:
2116:
2106:26 September
2104:. Retrieved
2093:
2083:
2071:. Retrieved
2064:the original
2039:
2033:
2020:
2000:
1993:
1982:
1963:
1930:
1926:
1916:
1903:
1890:
1877:
1867:
1860:
1850:
1825:
1821:
1815:
1805:
1798:
1789:
1779:
1766:
1741:
1737:
1728:
1678:
1668:
1666:
1658:
1652:
1648:
1632:
1603:
1596:, the later
1580:Conservation
1570:
1562:
1539:
1534:
1531:Song dynasty
1526:
1523:Tang dynasty
1518:
1516:
1508:black market
1493:
1477:
1473:
1469:
1466:coltiviruses
1462:pestiviruses
1459:
1454:
1450:
1388:
1345:
1338:
1334:
1311:
1289:
1253:
1252:position of
1247:
1242:
1238:
1234:
1230:
1227:sister group
1222:
1218:
1214:
1210:
1204:
1155:sister taxon
1152:
1126:Storch, 2003
1111:
1102:Filhol, 1893
1096:
1088:
1060:
1049:
1026:
1017:
965:
961:
926:Reproduction
913:
894:
879:
872:
860:
854:
839:
789:between the
772:
753:
727:
696:(φαττάγης).
693:
687:
682:
678:
674:
667:terenggiling
666:
658:
654:
646:
642:
638:
632:
623:
609:
589:
583:
577:
571:
565:
559:
553:
547:
530:) and heavy
521:
490:
481:
477:
473:
467:
461:
455:
450:family, the
405:
397:
393:
392:
371:Huxley, 1872
365:Murray, 1866
359:Newman, 1843
353:Pearse, 1936
351:Pholidotina
347:Kinman, 1994
297:Afredentata
245:
229:
58:47.8–0
46:
39:
7074:iNaturalist
6981:Wikispecies
6860:Palaeanodon
6780:Escavadodon
6712:Xenocranium
6683:Xenocranium
6375:Patriomanis
6367:Cryptomanis
6304:Eupholidota
6159:Dermoptera
6151:Scandentia
6143:Lagomorpha
6092:(Pangolins)
6082:Chiroptera
6038:(Elephants)
6028:Hyracoidea
6022:(Aardvarks)
5784:(unranked)
5426:China Daily
4840:18 February
4558:28 February
4421:LiveScience
4376:(11): 979.
4273:Daily Trust
3073:14 December
2994:: 477–501.
2629:: 472–480.
2508:: 267–273.
2394:14 December
2046:: 235–305.
1933:(810): 56.
1655:Jackie Chan
1612:trafficking
1546:blood clots
1434:during the
1356:Philippines
1260:phylogeny.
1235:M. javanica
1175:carnivorans
1051:Eupholidota
1021:Weber, 1904
905:gastroliths
807: in).
734:fingernails
700:Description
561:M. javanica
446:). The one
375:Squamigera
321:Manitheria
315:Maniformes
7174:Categories
6923:Melaniella
6871:Subfamily:
6833:Subfamily:
6635:Subfamily:
6616:Dipassalus
6600:Amelotabes
6419:Eomanoidea
6400:Necromanis
6098:Carnivora
6090:Pholidota
5996:Afrotheria
5975:Cingulata
5956:inclusive)
5878:(Opossums)
5849:inclusive)
5839:Metatheria
5796:Yinotheria
5780:Vertebrata
5778:Subphylum
5627:8 February
5536:8 February
5486:: e01395.
5459:27 January
5385:: 109002.
5349:2263/75894
5324:: e00714.
5203:10 January
4195:. London.
4146:. London.
3975:28 January
3930:30 January
3893:10 October
3868:1893/25549
3788:10 October
3697:30 January
3516:(1): 746.
2413:On Animals
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536:trafficked
513:armadillos
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463:Phataginus
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1121:Family: †
1079:Family: †
1028:Euromanis
1018:Pholidota
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606:Etymology
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369:Squamata
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