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Organelle

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1403: 1381: 311: 2985: 201:(also called membrane-bounded organelles) or are spatially distinct functional units without a surrounding lipid bilayer (non-membrane bounded organelles). Although most organelles are functional units within cells, some function units that extend outside of cells are often termed organelles, such as 700:
There are also occasional exceptions to the number of membranes surrounding organelles, listed in the tables below (e.g., some that are listed as double-membrane are sometimes found with single or triple membranes). In addition, the number of individual organelles of each type found in a given cell
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The first, broader conception of organelles is that they are membrane-bounded structures. However, even by using this definition, some parts of the cell that have been shown to be distinct functional units do not qualify as organelles. Therefore, the use of organelle to also refer to non-membrane
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WĂ€hrend die Fortpflanzungszellen der vielzelligen Tiere unthĂ€tig fortleben bis sie sich loslösen, wandern und entwickeln, treten die einzelligen Tiere auch durch die an der Fortpflanzung beteiligten Leibesmasse in Verkehr mit der Außenwelt und viele bilden sich dafĂŒr auch besondere Organula".
460:). In a footnote, which was published as a correction in the next issue of the journal, he justified his suggestion to call organs of unicellular organisms "organella" since they are only differently formed parts of one cell, in contrast to multicellular organs of multicellular organisms. 1466:
has revealed a number of compartmentalization features. The Planctomycetota cell plan includes intracytoplasmic membranes that separates the cytoplasm into paryphoplasm (an outer ribosome-free space) and pirellulosome (or riboplasm, an inner ribosome-containing space). Membrane-bounded
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It may possibly be of advantage to use the word organula here instead of organ, following a suggestion by Möbius. Functionally differentiated multicellular aggregates in multicellular forms or metazoa are in this sense organs, while, for functionally differentiated portions of
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Footnote on p. 448: "Die Organe der Heteroplastiden bestehen aus vereinigten Zellen. Da die Organe der Monoplastiden nur verschieden ausgebildete Teile e i n e r Zelle sind schlage ich vor, sie „Organula" zu nennen
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Not all eukaryotic cells have each of the organelles listed below. Exceptional organisms have cells that do not include some organelles (such as mitochondria) that might otherwise be considered universal to eukaryotes. The several
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that carry out particular and specialized functions, but they lack membrane boundaries. Many of these are referred to as "proteinaceous organelles" as their main structure is made of proteins. Such cell structures include:
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Lindsay, M. R.; Webb, R. I.; Strous, M; Jetten, M. S.; Butler, M. K.; Forde, R. J.; Fuerst, J. A. (2001). "Cell compartmentalisation in planctomycetes: Novel types of structural organisation for the bacterial cell".
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Die Vacuolen sind demnach in strengem Sinne keine bestĂ€ndigen Organe oder O r g a n u l a (wie Möbius die Organe der Einzelligen im Gegensatz zu denen der Vielzelligen zu nennen
1432:; but slowly, details are emerging about prokaryotic internal structures that overturn these assumptions. An early false turn was the idea developed in the 1970s that bacteria might contain 779:
rough endoplasmic reticulum is covered with ribosomes (which are bound to the ribosome membrane), has folds that are flat sacs; smooth endoplasmic reticulum has folds that are tubular
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Dr. H. G. Bronn's Klassen u. Ordnungen des Thier-Reichs wissenschaftlich dargestellt in Wort und Bild. Erster Band. Protozoa. Dritte Abtheilung: Infusoria und System der Radiolaria
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Scheffel A, Gruska M, Faivre D, Linaroudis A, Plitzko JM, SchĂŒler D (March 2006). "An acidic protein aligns magnetosomes along a filamentous structure in magnetotactic bacteria".
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of bodily organs to microscopic cellular substructures is obvious, as from even early works, authors of respective textbooks rarely elaborate on the distinction between the two.
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cells are structurally complex, and by definition are organized, in part, by interior compartments that are themselves enclosed by lipid membranes that resemble the outermost
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However, there is increasing evidence of compartmentalization in at least some prokaryotes. Recent research has revealed that at least some prokaryotes have
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Kerfeld CA, Sawaya MR, Tanaka S, Nguyen CV, Phillips M, Beeby M, Yeates TO (August 2005). "Protein structures forming the shell of primitive organelles".
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Using this definition, there would only be two broad classes of organelles (i.e. those that contain their own DNA, and have originated from endosymbiotic
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Imanian B, Carpenter KJ, Keeling PJ (March–April 2007). "Mitochondrial genome of a tertiary endosymbiont retains genes for electron transport proteins".
476:, a space often bounded by one or two lipid bilayers, some cell biologists choose to limit the term to include only those cell compartments that contain 1933: 3456: 2179:
Nott TJ, Petsalaki E, Farber P, Jervis D, Fussner E, Plochowietz A, Craggs TD, Bazett-Jones DP, Pawson T, Forman-Kay JD, Baldwin AJ (March 2015).
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Fahey RC, Newton GL, Arrick B, Overdank-Bogart T, Aley SB (April 1984). "Entamoeba histolytica: a eukaryote without glutathione metabolism".
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The mechanisms by which such non-membrane bounded organelles form and retain their spatial integrity have been likened to liquid-liquid
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Other organelles are also suggested to have endosymbiotic origins, but do not contain their own DNA (notably the flagellum – see
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Badano JL, Mitsuma N, Beales PL, Katsanis N (September 2006). "The ciliopathies: an emerging class of human genetic disorders".
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Jetten, Mike S. M.; Niftrik, Laura van; Strous, Marc; Kartal, Boran; Keltjens, Jan T.; Op den Camp, Huub J. M. (2009-06-01).
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translation and folding of new proteins (rough endoplasmic reticulum), expression of lipids (smooth endoplasmic reticulum)
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cis-face (convex) nearest to rough endoplasmic reticulum; trans-face (concave) farthest from rough endoplasmic reticulum
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or for such differentiated portions of the unicellular germ-elements of metazoa, the diminutive organula is appropriate.
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are not as structurally complex as eukaryotes, and were once thought to have little internal organization, and lack
3442: 2414: 1407: 144: 217:(these could be referred to as membrane bound in the sense that they are attached to (or bound to) the membrane). 3089: 2959: 334: 2660:
Ryter A (January–February 1988). "Contribution of new cryomethods to a better knowledge of bacterial anatomy".
1524:; studies have revealed that the cell membrane and the thylakoid membranes are not continuous with each other. 3191: 2418: 2181:"Phase transition of a disordered nuage protein generates environmentally responsive membraneless organelles" 1782: 575: 532: 427: 132: 2306:
Brangwynne CP, Eckmann CR, Courson DS, Rybarska A, Hoege C, Gharakhani J, JĂŒlicher F, Hyman AA (June 2009).
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bounded structures such as ribosomes is common and accepted. This has led many texts to delineate between
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The second, more restrictive definition of organelle includes only those cell compartments that contain
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Alberts B, Johnson A, Lewis J, Morgan D, Raff MC, Roberts K, Walter P, Wilson JH, Hunt T (2014-11-18).
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varies depending upon the function of that cell.The cell membrane and cell wall are not organelles.
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Tsai Y, Sawaya MR, Cannon GC, Cai F, Williams EB, Heinhorst S, Kerfeld CA, Yeates TO (June 2007).
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theory which said that microorganisms have the same organs of multicellular animals, only minor.
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Journal de l'anatomie et de la physiologie normales et pathologiques de l'homme et des animaux
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Adhesion to other cells for conjugation or to a solid substrate to create motile forces.
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a hair-like appendage sticking out (though partially embedded into) the plasma membrane
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which protrudes outside the cell, and its motor, as well as the largely extracellular
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Fuerst JA (October 13, 2005). "Intracellular compartmentation in planctomycetes".
1972: 3505: 3074: 2627: 2279: 2196: 2050:"Das Sterben der einzelligen und der vielzelligen Tiere. Vergleichend betrachtet" 1757: 1586: 1574: 1509: 1489: 1463: 802: 524: 407: 344: 245: 178: 113: 3233: 2973: 1900: 3513: 3037: 2587: 2574:
Anderson P, Kedersha N (March 2008). "Stress granules: the Tao of RNA triage".
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movement in or of external medium; "critical developmental signaling pathway".
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energy production from the oxidation of glucose substances and the release of
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have been discovered in five Planctomycetota "anammox" genera, which perform
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Nuclear import of histone fold motif containing heterodimers by importin 13
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vesicle that sequesters cytoplasmic material and organelles for degradation
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bounded organelles. The non-membrane bounded organelles, also called large
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Alberts B, Johnson A, Lewis J, Raff M, Roberts K, Walter P (2002).
1708: 1656: 1437: 1274: 1242: 1187: 1132: 1100: 1054: 1003: 918: 674: 552: 510: 412: 384: 328: 1380: 441:(i.e., little organ) for cellular structures was German zoologist 3385: 3285: 3238: 3114: 2995: 1692: 1472: 1354: 1157: 872: 735: 690: 643: 416: 370: 365: 261: 253: 177:
comes from the idea that these structures are parts of cells, as
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breakdown of large molecules (e.g., proteins + polysaccharides)
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Robin C, Pouchet G, Duval MM, Retterrer E, Tourneux F (1891).
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Murat, Dorothee; Byrne, Meghan; Komeili, Arash (2010-10-01).
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sorting, packaging, processing and modification of proteins
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degradation of unneeded or damaged proteins by proteolysis
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DNA maintenance, controls all activities of the cell, RNA
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has own DNA; theorized to be engulfed by the ancestral
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Critical Reviews in Biochemistry and Molecular Biology
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Komeili A, Li Z, Newman DK, Jensen GJ (January 2006).
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do not possess eukaryotic organelles, some do contain
228:. There are many types of organelles, particularly in 2694: 2149: 1996:"Embryology: The Structure of the Human Spermatozoon" 1961:. Gottingen: Vandenhoeck & Ruprecht. p. 218. 63: 1925: 1147:
a few unicellular eukaryotes that lack mitochondria
69: 2608: 2227: 959:, organizes cell division by forming spindle fibers 697:are distributed among some but not all eukaryotes. 411:are defined as confined functional units within an 60: 2444:"The Genetic Systems of Mitochondria and Plastids" 2278: 2228:Banani SF, Lee HO, Hyman AA, Rosen MK (May 2017). 2151: 1612:light harvesting complex attached to cell membrane 1882: 1484:Compartmentalization is a feature of prokaryotic 1394:. (B) Image of intact carboxysomes isolated from 3543: 2573: 2518:(Sixth ed.). Garland Science. p. 679. 2092: 2802: 2800: 1269:all eukaryotes, all archaea, and some bacteria 897:Minor eukaryotic organelles and cell components 3450: 3021: 2040: 2038: 468:While most cell biologists consider the term 2797: 2541:Annual Review of Genomics and Human Genetics 1878: 1876: 1874: 1872: 1870: 1868: 1866: 1192:detects light and possibly shapes, allowing 2662:Annales de l'Institut Pasteur. Microbiology 2150:Campbell NA, Reece JB, Mitchell LG (2002). 1953: 646:(e.g. in plants, algae, and some protists). 269: 232:. They include structures that make up the 169:is a specialized subunit, usually within a 3457: 3443: 3028: 3014: 2532: 2035: 1889:Cold Spring Harbor Perspectives in Biology 1530:Prokaryotic organelles and cell components 283: 2884: 2636: 2626: 2331: 2253: 2204: 2118: 1908: 1863: 1830: 1375: 173:, that has a specific function. The name 2095:"Organelle evolution: what's in a name?" 1971: 1931: 1885:"Cell Biology of Prokaryotic Organelles" 1401: 1379: 1231:breakdown of metabolic hydrogen peroxide 877:storage, transportation, helps maintain 657: 268:, which are thought to act as primitive 2405: 2276: 2093:Keeling PJ, Archibald JM (April 2008). 14: 3544: 2936:10.1146/annurev.micro.59.030804.121258 2921: 2365:The Journal of Eukaryotic Microbiology 2074: 2044: 1804: 1802: 1496:, which are reaction centers found in 3464: 3438: 3009: 2659: 2553:10.1146/annurev.genom.7.080505.115610 2411:The Biogenesis of Cellular Organelles 2234:Nature Reviews Molecular Cell Biology 1993: 669:. The larger organelles, such as the 315:Components of a typical animal cell: 280:, are often spoken of as organelles. 1959:: Ernst Haeckel and Scientific Faith 2158:(6th ed.). Benjamin Cummings. 1799: 24: 25: 3568: 2953: 1739:photosystem proteins and pigments 1646:inorganic crystal, lipid membrane 1475:. In the Planctomycetota species 929:most animals (including sponges) 565:large DNA and protein complexes: 509:large RNA and protein complexes: 27:Specialized subunit within a cell 2996:Tree of Life project: Eukaryotes 2983: 2377:10.1111/j.1550-7408.2007.00245.x 1512:found bonded to cell membranes. 1089:energy & hydrogen production 923:helps spermatozoa fuse with ovum 309: 248:), and other structures such as 145:Anatomical terms of microanatomy 47: 2915: 2852: 2746: 2688: 2653: 2602: 2567: 2507: 2464: 2435: 2415:Springer Science+Business Media 2399: 2356: 2299: 2270: 2221: 2172: 2143: 1994:Ryder JA, ed. (February 1889). 1934:"Mastering the Parts of a Cell" 1526: 1459:in bacteria, reported in 2006. 893: 703: 433:Credited as the first to use a 2576:Trends in Biochemical Sciences 2068: 2018: 1987: 1965: 1947: 1522:light-dependent photosynthesis 987:animals, protists, few plants 677:, are easily visible with the 551:; membrane protein complexes: 224:, and can also be purified by 13: 1: 3192:Microtubule organizing center 2924:Annual Review of Microbiology 2516:Molecular biology of the cell 2448:Molecular Biology of the Cell 2419:National Institutes of Health 1932:Peterson L (April 17, 2010). 1793: 1417:and intracytoplasmic membrane 1387:Halothiobacillus neapolitanus 1095:a few unicellular eukaryotes 1073:conversion of fat into sugars 576:microtubule-organizing center 533:DNA polymerase III holoenzyme 220:Organelles are identified by 2674:10.1016/0769-2609(88)90095-6 2628:10.1371/journal.pbio.0050144 2197:10.1016/j.molcel.2015.01.013 1778:Membrane vesicle trafficking 1557:anaerobic ammonium oxidation 1473:anaerobic ammonium oxidation 838:constituting element of the 744:, traps energy from sunlight 537:RNA polymerase II holoenzyme 356:Smooth endoplasmic reticulum 7: 1901:10.1101/cshperspect.a000422 1788:Vesiculo-vacuolar organelle 1751: 1627:movement in external medium 1508:, which are photosynthetic 1384:(A) Electron micrograph of 1341:single-membrane compartment 1250:single-membrane compartment 1234:single-membrane compartment 1199:double-membrane compartment 1144:double-membrane compartment 1124:single-membrane compartment 1108:single-membrane compartment 1092:double-membrane compartment 1076:single-membrane compartment 1050:single-membrane compartment 942:double-membrane compartment 926:single-membrane compartment 883:single-membrane compartment 858:double-membrane compartment 832:double-membrane compartment 810:single-membrane compartment 773:single-membrane compartment 747:double-membrane compartment 707:Major eukaryotic organelles 340:Rough endoplasmic reticulum 266:bacterial microcompartments 10: 3573: 2588:10.1016/j.tibs.2007.12.003 1593:bacterial microcompartment 1350:Other related structures: 1266:very large protein complex 640:(in almost all eukaryotes) 607:neuronal transport granule 500:, are large assemblies of 445:(1884), who used the term 3472: 3411: 3346: 3271: 3162: 3048: 2974:Resources in your library 2877:10.1080/10409230902722783 2281:Introduction to Histology 2120:10.1016/j.cub.2008.02.065 2054:Biologisches Centralblatt 1783:Host–pathogen interaction 1137:probably plays a role in 308: 299: 290: 143: 131: 119: 107: 95: 90: 80: 42: 37: 32: 3187:Prokaryotic cytoskeleton 2810:Archives of Microbiology 2075:Walker, Patrick (2009). 2060:(13, 14): 389–392, 448. 1537:Organelle/macromolecule 1398:. Scale bars are 100 nm. 1390:cells, arrows highlight 1015:detects light, allowing 904:Organelle/Macromolecule 463: 377:; with which, comprises 2724:10.1126/science.1123231 2493:10.1126/science.6322306 2333:10.1126/science.1172046 1954:Di Gregorio MA (2005). 1841:10.1126/science.1113397 1322:mRNA encoding proteins 1176:pre-ribosome production 752:kleptoplastic organisms 284:History and terminology 1650:magnetotactic bacteria 1500:of the cell membrane. 1418: 1399: 1376:Prokaryotic organelles 1202:members of the family 1027:and other unicellular 828:adenosine triphosphate 498:biomolecular complexes 472:to be synonymous with 270:prokaryotic organelles 3177:Intermediate filament 3070:Endoplasmic reticulum 2830:10.1007/s002030100280 2011:unicellular organisms 1957:From Here to Eternity 1723:bacteria and archaea 1616:green sulfur bacteria 1502:Green sulfur bacteria 1478:Gemmata obscuriglobus 1462:The bacterial phylum 1426:cellular compartments 1411:Brocadia anammoxidans 1405: 1383: 766:endoplasmic reticulum 760:cell (endosymbiosis) 658:Eukaryotic organelles 652:evolution of flagella 620:deoxyribonucleic acid 478:deoxyribonucleic acid 373:(fluid that contains 242:endoplasmic reticulum 3522:Biomolecular complex 3424:Extracellular matrix 2992:at Wikimedia Commons 1773:Organelle biogenesis 1768:Endosymbiotic theory 1717:of RNA into proteins 1643:magnetic orientation 1283:of RNA into proteins 1000:coiled hollow tubule 750:plants, algae, rare 453:, the diminutive of 3127:Cytoplasmic granule 3001:Organelle Databases 2822:2001ArMic.175..413L 2775:10.1038/nature04382 2767:2006Natur.440..110S 2716:2006Sci...311..242K 2485:1984Sci...224...70F 2324:2009Sci...324.1729B 2277:Cormack DH (1984). 2111:2008CBio...18.R345K 2000:American Naturalist 1823:2005Sci...309..936K 1533: 1442:electron microscopy 1360:endomembrane system 1139:Iron–sulfur cluster 900: 789:locomotion, sensory 710: 331:(dots as part of 5) 302:Animal cell diagram 234:endomembrane system 3152:Weibel–Palade body 3036:Structures of the 2246:10.1038/nrm.2017.7 2048:(September 1884). 1687:prokaryotic cells 1573:" bacteria within 1527: 1419: 1400: 1338:material transport 1031:organisms such as 894: 704: 539:, symmetric viral 443:Karl August Möbius 226:cell fractionation 126:H1.00.01.0.00009 3539: 3538: 3466:Hierarchy of life 3432: 3431: 3212:Spindle pole body 2988:Media related to 2960:Library resources 2457:978-0-8153-3218-3 2428:978-0-306-47990-8 2318:(5935): 1729–32. 2292:978-0-397-52114-2 2165:978-0-8053-6624-2 1749: 1748: 1661:DNA maintenance, 1633:some prokaryotes 1510:antenna complexes 1449:microcompartments 1348: 1347: 1218:not characterized 1215:not characterized 1163:bundled filaments 1011:eyespot apparatus 892: 891: 864:contains bulk of 525:protein complexes 403: 402: 159: 158: 154: 16:(Redirected from 3564: 3532: 3459: 3452: 3445: 3436: 3435: 3030: 3023: 3016: 3007: 3006: 2987: 2948: 2947: 2919: 2913: 2912: 2910: 2909: 2888: 2856: 2850: 2849: 2804: 2795: 2794: 2750: 2744: 2743: 2701: 2692: 2686: 2685: 2657: 2651: 2650: 2640: 2630: 2606: 2600: 2599: 2571: 2565: 2564: 2536: 2530: 2529: 2511: 2505: 2504: 2468: 2462: 2461: 2450:(4th ed.). 2439: 2433: 2432: 2403: 2397: 2396: 2360: 2354: 2353: 2335: 2303: 2297: 2296: 2284: 2274: 2268: 2267: 2257: 2225: 2219: 2218: 2208: 2176: 2170: 2169: 2157: 2147: 2141: 2140: 2122: 2090: 2081: 2080: 2072: 2066: 2065: 2042: 2033: 2032: 2022: 2016: 2015: 1991: 1985: 1984: 1979:. p. 1412. 1969: 1963: 1962: 1951: 1945: 1944: 1942: 1941: 1929: 1923: 1922: 1912: 1880: 1861: 1860: 1834: 1806: 1630:protein filament 1534: 1494:"chromatophores" 1312:most eukaryotes 1247:secretory portal 1182:most eukaryotes 901: 795:some eukaryotes 711: 679:light microscope 613:phase separation 490:membrane-bounded 474:cell compartment 347:(or, Golgi body) 313: 303: 294: 288: 287: 238:nuclear envelope 230:eukaryotic cells 151:edit on Wikidata 148: 76: 75: 72: 71: 68: 65: 62: 59: 56: 53: 30: 29: 21: 3572: 3571: 3567: 3566: 3565: 3563: 3562: 3561: 3542: 3541: 3540: 3535: 3476: 3468: 3463: 3433: 3428: 3407: 3342: 3267: 3158: 3075:Golgi apparatus 3051: 3044: 3034: 2980: 2979: 2978: 2968: 2967: 2963: 2956: 2951: 2920: 2916: 2907: 2905: 2857: 2853: 2805: 2798: 2761:(7080): 110–4. 2751: 2747: 2710:(5758): 242–5. 2699: 2693: 2689: 2658: 2654: 2607: 2603: 2572: 2568: 2537: 2533: 2526: 2512: 2508: 2469: 2465: 2458: 2440: 2436: 2429: 2404: 2400: 2361: 2357: 2304: 2300: 2293: 2275: 2271: 2226: 2222: 2177: 2173: 2166: 2148: 2144: 2099:Current Biology 2091: 2084: 2073: 2069: 2043: 2036: 2023: 2019: 1992: 1988: 1970: 1966: 1952: 1948: 1939: 1937: 1936:. Lesson Planet 1930: 1926: 1895:(10): a000422. 1881: 1864: 1832:10.1.1.1026.896 1817:(5736): 936–8. 1807: 1800: 1796: 1758:CoRR hypothesis 1754: 1587:carbon fixation 1575:Planctomycetota 1490:Purple bacteria 1464:Planctomycetota 1396:H. neapolitanus 1378: 1344:all eukaryotes 1328:most organisms 1289:all eukaryotes 1253:all eukaryotes 1237:all eukaryotes 1179:protein–DNA–RNA 1141:(Fe–S) assembly 1121:pigment storage 945:all eukaryotes 835:most eukaryotes 803:Golgi apparatus 660: 466: 399: 345:Golgi apparatus 301: 292: 286: 246:Golgi apparatus 155: 50: 46: 28: 23: 22: 15: 12: 11: 5: 3570: 3560: 3559: 3554: 3537: 3536: 3534: 3533: 3473: 3470: 3469: 3462: 3461: 3454: 3447: 3439: 3430: 3429: 3427: 3426: 3421: 3415: 3413: 3409: 3408: 3406: 3405: 3400: 3395: 3394: 3393: 3388: 3378: 3377: 3376: 3371: 3366: 3356: 3350: 3348: 3347:Other internal 3344: 3343: 3341: 3340: 3335: 3334: 3333: 3328: 3323: 3318: 3313: 3308: 3303: 3298: 3293: 3283: 3277: 3275: 3269: 3268: 3266: 3265: 3264: 3263: 3258: 3248: 3247: 3246: 3241: 3236: 3231: 3221: 3216: 3215: 3214: 3209: 3204: 3199: 3189: 3184: 3179: 3174: 3168: 3166: 3160: 3159: 3157: 3156: 3155: 3154: 3149: 3144: 3139: 3134: 3124: 3123: 3122: 3117: 3112: 3107: 3102: 3097: 3087: 3082: 3077: 3072: 3067: 3062: 3056: 3054: 3046: 3045: 3033: 3032: 3025: 3018: 3010: 3004: 3003: 2998: 2993: 2977: 2976: 2970: 2969: 2958: 2957: 2955: 2954:External links 2952: 2950: 2949: 2914: 2871:(2–3): 65–84. 2851: 2796: 2745: 2687: 2652: 2601: 2566: 2531: 2525:978-0815345244 2524: 2506: 2479:(4644): 70–2. 2463: 2456: 2434: 2427: 2398: 2355: 2298: 2291: 2285:. Lippincott. 2269: 2240:(5): 285–298. 2220: 2191:(5): 936–947. 2185:Molecular Cell 2171: 2164: 2142: 2082: 2067: 2034: 2017: 1986: 1964: 1946: 1924: 1862: 1797: 1795: 1792: 1791: 1790: 1785: 1780: 1775: 1770: 1765: 1760: 1753: 1750: 1747: 1746: 1740: 1737: 1735:photosynthesis 1732: 1725: 1724: 1721: 1718: 1712: 1705: 1704: 1703:some bacteria 1701: 1698: 1695: 1689: 1688: 1685: 1682: 1679: 1673: 1672: 1669: 1666: 1659: 1653: 1652: 1647: 1644: 1641: 1635: 1634: 1631: 1628: 1625: 1619: 1618: 1613: 1610: 1608:photosynthesis 1605: 1599: 1598: 1597:some bacteria 1595: 1591:protein-shell 1589: 1584: 1578: 1577: 1565: 1564:lipid membrane 1559: 1554: 1548: 1547: 1544: 1541: 1540:Main function 1538: 1520:membranes for 1516:have internal 1486:photosynthetic 1377: 1374: 1373: 1372: 1367: 1362: 1357: 1346: 1345: 1342: 1339: 1336: 1330: 1329: 1326: 1323: 1320: 1314: 1313: 1310: 1300: 1297: 1295:stress granule 1291: 1290: 1287: 1284: 1278: 1271: 1270: 1267: 1264: 1261: 1255: 1254: 1251: 1248: 1245: 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521: 502:macromolecules 465: 462: 424:FĂ©lix Dujardin 422:In the 1830s, 401: 400: 398: 397: 392: 387: 382: 368: 363: 358: 353: 348: 342: 337: 332: 326: 321: 314: 306: 305: 297: 296: 285: 282: 199:lipid bilayers 157: 156: 147: 141: 140: 135: 129: 128: 123: 117: 116: 111: 105: 104: 99: 93: 92: 88: 87: 82: 78: 77: 44: 40: 39: 35: 34: 26: 9: 6: 4: 3: 2: 3569: 3558: 3555: 3553: 3550: 3549: 3547: 3531: 3527: 3526:Macromolecule 3523: 3519: 3515: 3511: 3507: 3503: 3499: 3495: 3491: 3487: 3483: 3479: 3475: 3474: 3471: 3467: 3460: 3455: 3453: 3448: 3446: 3441: 3440: 3437: 3425: 3422: 3420: 3417: 3416: 3414: 3410: 3404: 3401: 3399: 3396: 3392: 3389: 3387: 3384: 3383: 3382: 3379: 3375: 3372: 3370: 3367: 3365: 3362: 3361: 3360: 3357: 3355: 3352: 3351: 3349: 3345: 3339: 3336: 3332: 3329: 3327: 3324: 3322: 3321:Proteinoplast 3319: 3317: 3314: 3312: 3309: 3307: 3304: 3302: 3299: 3297: 3294: 3292: 3289: 3288: 3287: 3284: 3282: 3281:Mitochondrion 3279: 3278: 3276: 3274: 3273:Endosymbionts 3270: 3262: 3259: 3257: 3256:Lamellipodium 3254: 3253: 3252: 3249: 3245: 3242: 3240: 3237: 3235: 3232: 3230: 3227: 3226: 3225: 3222: 3220: 3217: 3213: 3210: 3208: 3205: 3203: 3200: 3198: 3195: 3194: 3193: 3190: 3188: 3185: 3183: 3180: 3178: 3175: 3173: 3172:Microfilament 3170: 3169: 3167: 3165: 3161: 3153: 3150: 3148: 3145: 3143: 3140: 3138: 3135: 3133: 3130: 3129: 3128: 3125: 3121: 3118: 3116: 3113: 3111: 3108: 3106: 3103: 3101: 3098: 3096: 3093: 3092: 3091: 3088: 3086: 3085:Autophagosome 3083: 3081: 3078: 3076: 3073: 3071: 3068: 3066: 3063: 3061: 3060:Cell membrane 3058: 3057: 3055: 3053: 3050:Endomembrane 3047: 3043: 3039: 3031: 3026: 3024: 3019: 3017: 3012: 3011: 3008: 3002: 2999: 2997: 2994: 2991: 2986: 2982: 2981: 2975: 2972: 2971: 2966: 2961: 2945: 2941: 2937: 2933: 2929: 2925: 2918: 2904: 2900: 2896: 2892: 2887: 2882: 2878: 2874: 2870: 2866: 2862: 2855: 2847: 2843: 2839: 2835: 2831: 2827: 2823: 2819: 2816:(6): 413–29. 2815: 2811: 2803: 2801: 2792: 2788: 2784: 2780: 2776: 2772: 2768: 2764: 2760: 2756: 2749: 2741: 2737: 2733: 2729: 2725: 2721: 2717: 2713: 2709: 2705: 2698: 2691: 2683: 2679: 2675: 2671: 2667: 2663: 2656: 2648: 2644: 2639: 2634: 2629: 2624: 2620: 2616: 2612: 2605: 2597: 2593: 2589: 2585: 2582:(3): 141–50. 2581: 2577: 2570: 2562: 2558: 2554: 2550: 2546: 2542: 2535: 2527: 2521: 2517: 2510: 2502: 2498: 2494: 2490: 2486: 2482: 2478: 2474: 2467: 2459: 2453: 2449: 2445: 2438: 2430: 2424: 2420: 2416: 2412: 2408: 2402: 2394: 2390: 2386: 2382: 2378: 2374: 2371:(2): 146–53. 2370: 2366: 2359: 2351: 2347: 2343: 2339: 2334: 2329: 2325: 2321: 2317: 2313: 2309: 2302: 2294: 2288: 2283: 2282: 2273: 2265: 2261: 2256: 2251: 2247: 2243: 2239: 2235: 2231: 2224: 2216: 2212: 2207: 2202: 2198: 2194: 2190: 2186: 2182: 2175: 2167: 2161: 2156: 2155: 2146: 2138: 2134: 2130: 2126: 2121: 2116: 2112: 2108: 2105:(8): R345-7. 2104: 2100: 2096: 2089: 2087: 2078: 2071: 2064: 2059: 2055: 2051: 2047: 2041: 2039: 2030: 2029: 2021: 2014: 2012: 2005: 2001: 1997: 1990: 1983: 1978: 1974: 1968: 1960: 1956: 1950: 1935: 1928: 1920: 1916: 1911: 1906: 1902: 1898: 1894: 1890: 1886: 1879: 1877: 1875: 1873: 1871: 1869: 1867: 1858: 1854: 1850: 1846: 1842: 1838: 1833: 1828: 1824: 1820: 1816: 1812: 1805: 1803: 1798: 1789: 1786: 1784: 1781: 1779: 1776: 1774: 1771: 1769: 1766: 1764: 1761: 1759: 1756: 1755: 1745: 1744:cyanobacteria 1741: 1738: 1736: 1733: 1730: 1727: 1726: 1722: 1719: 1716: 1713: 1710: 1707: 1706: 1702: 1699: 1696: 1694: 1691: 1690: 1686: 1683: 1680: 1678: 1675: 1674: 1670: 1667: 1664: 1663:transcription 1660: 1658: 1655: 1654: 1651: 1648: 1645: 1642: 1640: 1637: 1636: 1632: 1629: 1626: 1624: 1621: 1620: 1617: 1614: 1611: 1609: 1606: 1604: 1601: 1600: 1596: 1594: 1590: 1588: 1585: 1583: 1580: 1579: 1576: 1572: 1571: 1566: 1563: 1560: 1558: 1555: 1553: 1550: 1549: 1545: 1542: 1539: 1536: 1535: 1532: 1531: 1525: 1523: 1519: 1515: 1514:Cyanobacteria 1511: 1507: 1503: 1499: 1498:invaginations 1495: 1491: 1487: 1482: 1480: 1479: 1474: 1470: 1469:anammoxosomes 1465: 1460: 1458: 1454: 1450: 1445: 1443: 1439: 1436:folds termed 1435: 1434:cell membrane 1431: 1428:and internal 1427: 1423: 1416: 1413:, showing an 1412: 1410: 1406:Structure of 1404: 1397: 1393: 1389: 1388: 1382: 1371: 1368: 1366: 1363: 1361: 1358: 1356: 1353: 1352: 1351: 1343: 1340: 1337: 1335: 1332: 1331: 1327: 1325:membraneless 1324: 1321: 1319: 1316: 1315: 1311: 1309: 1307: 1302:membraneless 1301: 1299:mRNA storage 1298: 1296: 1293: 1292: 1288: 1285: 1282: 1279: 1276: 1273: 1272: 1268: 1265: 1262: 1260: 1257: 1256: 1252: 1249: 1246: 1244: 1241: 1240: 1236: 1233: 1230: 1228: 1225: 1224: 1220: 1217: 1214: 1212: 1209: 1208: 1205: 1201: 1198: 1196:to take place 1195: 1191: 1189: 1186: 1185: 1181: 1178: 1175: 1173: 1170: 1169: 1165: 1162: 1159: 1156: 1154: 1151: 1150: 1146: 1143: 1140: 1136: 1134: 1131: 1130: 1126: 1123: 1120: 1118: 1115: 1114: 1110: 1107: 1104: 1102: 1099: 1098: 1094: 1091: 1088: 1086: 1085:hydrogenosome 1083: 1082: 1078: 1075: 1072: 1070: 1067: 1066: 1062: 1061: 1056: 1052: 1049: 1047: 1043: 1041: 1038: 1037: 1034: 1030: 1026: 1023: 1021: 1019:to take place 1018: 1014: 1012: 1009: 1008: 1005: 1002: 999: 996: 994: 991: 990: 986: 983: 980: 977: 975: 972: 971: 967: 964: 961: 958: 954: 952: 949: 948: 944: 941: 938: 936: 935:autophagosome 933: 932: 928: 925: 922: 920: 917: 916: 912: 909: 906: 903: 902: 899: 898: 888: 885: 882: 880: 876: 874: 871: 870: 867: 863: 860: 857: 855: 854:transcription 851: 849: 846: 845: 841: 837: 834: 831: 829: 825: 823: 822:mitochondrion 820: 819: 815: 812: 809: 806: 804: 801: 800: 797: 794: 791: 788: 786: 783: 782: 778: 775: 772: 769: 767: 764: 763: 759: 758:archaeplastid 755: 753: 749: 746: 743: 740: 737: 733: 730: 729: 725: 722: 719: 716: 713: 712: 709: 708: 702: 698: 696: 692: 686: 684: 680: 676: 672: 668: 667:cell membrane 664: 655: 653: 645: 642: 639: 636: 635: 634: 632: 627: 625: 624:endosymbiosis 621: 616: 614: 606: 603: 600: 598: 595: 593: 590: 588: 585: 583: 580: 577: 573: 570: 568: 564: 562: 558: 557:photosystem I 554: 550: 546: 543:, complex of 542: 538: 534: 530: 526: 522: 520: 516: 512: 508: 507: 506: 503: 499: 495: 491: 485: 483: 482:endosymbiosis 479: 475: 471: 461: 459: 456: 452: 448: 444: 440: 436: 431: 429: 425: 420: 418: 414: 410: 409: 396: 395:Cell membrane 393: 391: 388: 386: 383: 380: 376: 372: 369: 367: 364: 362: 361:Mitochondrion 359: 357: 354: 352: 349: 346: 343: 341: 338: 336: 333: 330: 327: 325: 322: 320: 317: 316: 312: 307: 304: 298: 295: 289: 281: 279: 275: 271: 267: 263: 259: 255: 251: 247: 243: 239: 236:(such as the 235: 231: 227: 223: 218: 216: 212: 208: 204: 200: 196: 192: 188: 184: 180: 176: 172: 168: 164: 152: 146: 142: 139: 136: 134: 130: 127: 124: 122: 118: 115: 112: 110: 106: 103: 100: 98: 94: 89: 86: 83: 79: 74: 45: 43:Pronunciation 41: 36: 31: 19: 3557:Cell anatomy 3528: > 3524: > 3520: > 3517: 3516: > 3512: > 3508: > 3504: > 3502:Organ system 3500: > 3496: > 3492: > 3488: > 3484: > 3480: > 3301:Gerontoplast 3251:Pseudopodium 3244:Radial spoke 3224:Undulipodium 3164:Cytoskeleton 3080:Parenthesome 3041: 2964: 2927: 2923: 2917: 2906:. 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Retrieved 1927: 1892: 1888: 1814: 1810: 1700:circular DNA 1697:DNA exchange 1671:prokaryotes 1568: 1552:anammoxosome 1529: 1528: 1488:structures. 1483: 1476: 1461: 1457:magnetosomes 1453:carboxysomes 1446: 1420: 1415:anammoxosome 1408: 1395: 1392:carboxysomes 1385: 1349: 1318:TIGER domain 1303: 1211:parenthesome 1204:Warnowiaceae 1060:Trypanosomes 1058: 1044:carries out 957:cytoskeleton 896: 895: 706: 705: 699: 695:chloroplasts 687: 661: 649: 638:mitochondria 628: 617: 610: 582:cytoskeleton 561:ATP synthase 494:non-membrane 493: 489: 486: 469: 467: 457: 450: 446: 438: 432: 421: 406: 405:In biology, 404: 374: 351:Cytoskeleton 293:Cell biology 250:mitochondria 219: 190: 186: 174: 166: 163:cell biology 160: 101: 3530:Biomolecule 3490:Biocoenosis 3403:Magnetosome 3369:Spliceosome 3296:Chromoplast 3291:Chloroplast 3182:Microtubule 2930:: 299–328. 2621:(6): e144. 2031:. F. Alcan. 1982:vorschlug). 1720:RNA-protein 1715:translation 1668:DNA-protein 1639:magnetosome 1582:carboxysome 1506:chlorosomes 1422:Prokaryotes 1370:microtubule 1308:complexes) 1286:RNA-protein 1281:translation 1160:contraction 1025:green algae 982:Microtubule 963:Microtubule 955:anchor for 879:homeostasis 732:chloroplast 515:spliceosome 449:(plural of 258:prokaryotes 189:the suffix 181:are to the 91:Identifiers 3552:Organelles 3546:Categories 3494:Population 3398:Proteasome 3391:Inclusions 3338:Nitroplast 3331:Apicoplast 3316:Elaioplast 3311:Amyloplast 3306:Leucoplast 3261:Filopodium 3207:Basal body 3197:Centrosome 3147:Peroxisome 3142:Glyoxysome 3132:Melanosome 3042:organelles 2990:Organelles 2908:2020-08-03 2886:2066/75127 2547:: 125–48. 1973:BĂŒtschli O 1940:2010-04-19 1794:References 1763:Ejectosome 1603:chlorosome 1570:Candidatus 1546:Organisms 1543:Structure 1451:, such as 1409:Candidatus 1365:nucleosome 1259:proteasome 1227:peroxisome 1194:phototaxis 1117:melanosome 1069:glyoxysome 1057:, such as 1046:glycolysis 1017:phototaxis 1004:cnidarians 913:Organisms 910:Structure 840:chondriome 723:Organisms 720:Structure 714:Organelle 693:including 683:microscope 663:Eukaryotic 567:nucleosome 529:proteasome 435:diminutive 390:Centrosome 375:organelles 222:microscopy 215:trichocyst 213:, and the 211:archaellum 195:diminutive 187:organelle, 18:Organellar 3518:Organelle 3486:Ecosystem 3478:Biosphere 3419:Cell wall 3381:Cytoplasm 3354:Nucleolus 3326:Tannosome 3234:Flagellum 3219:Myofibril 3202:Centriole 3137:Microbody 3110:Phagosome 2965:Organelle 2903:205694872 2407:Mullins C 1827:CiteSeerX 1731:membranes 1729:thylakoid 1623:flagellum 1562:ladderane 1518:thylakoid 1438:mesosomes 1430:membranes 1172:nucleolus 1153:myofibril 1040:glycosome 1033:euglenids 993:cnidocyst 951:centriole 785:flagellum 602:germ cell 592:nucleolus 587:flagellum 572:centriole 470:organelle 451:organulum 428:Ehrenberg 379:cytoplasm 319:Nucleolus 274:flagellum 264:-shelled 207:flagellum 175:organelle 167:organelle 102:organella 33:Organelle 3498:Organism 3412:External 3364:Ribosome 3120:Acrosome 3105:Endosome 3100:Lysosome 2944:15910279 2895:19247843 2846:21970703 2838:11491082 2783:16299495 2740:36909813 2732:16373532 2647:17518518 2596:18291657 2561:16722803 2393:20393495 2385:17403155 2350:42229928 2342:19460965 2264:28225081 2215:25747659 2137:11520942 2129:18430636 2046:Möbius K 1975:(1888). 1919:20739411 1857:24561197 1849:16081736 1752:See also 1709:ribosome 1657:nucleoid 1275:ribosome 1243:porosome 1188:ocelloid 1166:animals 1133:mitosome 1127:animals 1111:animals 1101:lysosome 1055:protozoa 997:stinging 968:animals 919:acrosome 792:protein 691:plastids 675:vacuoles 644:plastids 631:bacteria 553:porosome 511:ribosome 447:organula 426:refuted 413:organism 385:Lysosome 329:Ribosome 256:. While 254:plastids 193:being a 185:, hence 3386:Cytosol 3286:Plastid 3239:Axoneme 3115:Vacuole 3095:Exosome 3090:Vesicle 3065:Nucleus 2818:Bibcode 2791:4372846 2763:Bibcode 2712:Bibcode 2704:Science 2682:3289587 2638:1872035 2501:6322306 2481:Bibcode 2473:Science 2320:Bibcode 2312:Science 2255:7434221 2206:4352761 2154:Biology 2107:Bibcode 2006:: 184. 1910:2944366 1819:Bibcode 1811:Science 1742:mostly 1693:plasmid 1355:cytosol 1334:vesicle 1158:myocyte 1079:plants 984:protein 965:protein 873:vacuole 848:nucleus 736:plastid 671:nucleus 604:granule 541:capsids 458:organum 417:analogy 371:Cytosol 366:Vacuole 335:Vesicle 324:Nucleus 262:protein 114:D015388 81:Part of 38:Details 3510:Tissue 3229:Cilium 3052:system 2962:about 2942:  2901:  2893:  2844:  2836:  2789:  2781:  2755:Nature 2738:  2730:  2680:  2645:  2635:  2594:  2559:  2522:  2499:  2454:  2425:  2391:  2383:  2348:  2340:  2289:  2262:  2252:  2213:  2203:  2162:  2135:  2127:  1917:  1907:  1855:  1847:  1829:  1665:to RNA 1221:fungi 974:cilium 866:genome 726:Notes 578:(MTOC) 523:large 415:. The 408:organs 244:, and 205:, the 179:organs 3506:Organ 3482:Biome 3374:Vault 2899:S2CID 2842:S2CID 2787:S2CID 2736:S2CID 2700:(PDF) 2389:S2CID 2346:S2CID 2133:S2CID 1853:S2CID 1711:(70S) 1677:pilus 1504:have 1492:have 1277:(80S) 1053:Some 549:GroES 545:GroEL 519:vault 464:Types 455:Latin 439:organ 278:pilus 203:cilia 191:-elle 165:, an 149:[ 138:63832 97:Latin 3514:Cell 3038:cell 2940:PMID 2891:PMID 2834:PMID 2779:PMID 2728:PMID 2678:PMID 2643:PMID 2592:PMID 2557:PMID 2520:ISBN 2497:PMID 2452:ISBN 2423:ISBN 2381:PMID 2338:PMID 2287:ISBN 2260:PMID 2211:PMID 2160:ISBN 2125:PMID 1915:PMID 1845:PMID 1306:mRNP 673:and 574:and 547:and 492:and 252:and 209:and 183:body 171:cell 109:MeSH 85:Cell 3359:RNA 2932:doi 2881:hdl 2873:doi 2826:doi 2814:175 2771:doi 2759:440 2720:doi 2708:311 2670:doi 2666:139 2633:PMC 2623:doi 2584:doi 2549:doi 2489:doi 2477:224 2373:doi 2328:doi 2316:324 2250:PMC 2242:doi 2201:PMC 2193:doi 2115:doi 1905:PMC 1897:doi 1837:doi 1815:309 654:). 633:): 437:of 161:In 133:FMA 52:ɔːr 3548:: 3040:/ 2938:. 2928:59 2926:. 2897:. 2889:. 2879:. 2869:44 2867:. 2863:. 2840:. 2832:. 2824:. 2812:. 2799:^ 2785:. 2777:. 2769:. 2757:. 2734:. 2726:. 2718:. 2706:. 2702:. 2676:. 2664:. 2641:. 2631:. 2617:. 2613:. 2590:. 2580:33 2578:. 2555:. 2543:. 2495:. 2487:. 2475:. 2446:. 2421:. 2417:, 2413:. 2387:. 2379:. 2369:54 2367:. 2344:. 2336:. 2326:. 2314:. 2310:. 2258:. 2248:. 2238:18 2236:. 2232:. 2209:. 2199:. 2189:57 2187:. 2183:. 2131:. 2123:. 2113:. 2103:18 2101:. 2097:. 2085:^ 2056:. 2052:. 2037:^ 2004:23 2002:. 1998:. 1913:. 1903:. 1891:. 1887:. 1865:^ 1851:. 1843:. 1835:. 1825:. 1813:. 1801:^ 1444:. 1063:. 685:. 626:. 615:. 559:, 555:, 535:, 531:, 527:: 517:, 513:, 484:. 240:, 121:TH 3458:e 3451:t 3444:v 3029:e 3022:t 3015:v 2946:. 2934:: 2911:. 2883:: 2875:: 2848:. 2828:: 2820:: 2793:. 2773:: 2765:: 2742:. 2722:: 2714:: 2684:. 2672:: 2649:. 2625:: 2619:5 2598:. 2586:: 2563:. 2551:: 2545:7 2528:. 2503:. 2491:: 2483:: 2460:. 2431:. 2395:. 2375:: 2352:. 2330:: 2322:: 2295:. 2266:. 2244:: 2217:. 2195:: 2168:. 2139:. 2117:: 2109:: 2058:4 1943:. 1921:. 1899:: 1893:2 1859:. 1839:: 1821:: 1567:" 1304:( 738:) 734:( 381:) 153:] 73:/ 70:l 67:ɛ 64:n 61:ˈ 58:ə 55:ÉĄ 49:/ 20:)

Index

Organellar
/ɔːrɡəˈnɛl/
Cell
Latin
MeSH
D015388
TH
H1.00.01.0.00009
FMA
63832
Anatomical terms of microanatomy
edit on Wikidata
cell biology
cell
organs
body
diminutive
lipid bilayers
cilia
flagellum
archaellum
trichocyst
microscopy
cell fractionation
eukaryotic cells
endomembrane system
nuclear envelope
endoplasmic reticulum
Golgi apparatus
mitochondria

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