227:
213:
185:
199:
31:
51:
244:
Adults and larvae occur in leaf litter, decaying fruits, moss, and under bark of dead trees. Adults of several species and larvae of a few occur in flowers. Adults and larvae of many genera and species are believed to be predatory (they feed on freshly killed small insects), though a few seem to be
313:
Newton, A. F., Jr., M. K. Thayer, J. S. Ashe, and D. S. Chandler. 2001. 22. Staphylinidae
Latreille, 1802. p. 272–418. In: R. H. Arnett, Jr., and M. C. Thomas (eds.). American beetles, Volume 1. CRC Press; Boca Raton, Florida. ix + 443
166:
near the base of the head, and in a few, the elytra cover the entire abdomen. The maxillary mala of larvae are strap-shaped, but not as long as in the
291:
281:
276:
271:
2417:
286:
175:
2241:
2391:
2396:
354:
226:
150:
Typical adults are 1.5 to 6 mm long, somewhat broader in shape than are most
Staphylinidae, with somewhat longer
212:
2357:
2453:
2422:
2370:
184:
2404:
198:
2486:
2303:
171:
50:
2458:
2432:
347:
2445:
2256:
253:
The
Omaliinae subfamily is large (comprising over 100 genera), and is divided into these seven
2440:
1493:
8:
2294:
1712:
340:
2481:
45:
2339:
1981:
1398:
155:
30:
2344:
2078:
1857:
1852:
1775:
1533:
549:
245:
phytophagous (they damage flowers) or saprophagous (they feed on decaying fruits).
159:
2308:
1767:
1720:
1565:
1549:
1225:
985:
954:
869:
787:
678:
646:
254:
2409:
2365:
2352:
2279:
2170:
2121:
1976:
1783:
1451:
1435:
1374:
1269:
1240:
1168:
1132:
1116:
1032:
977:
882:
779:
628:
590:
570:
466:
2475:
2152:
1956:
1947:
1688:
1659:
1651:
1643:
1617:
1592:
1488:
1475:
1414:
1332:
1235:
1230:
1111:
1045:
1016:
1000:
964:
934:
887:
841:
808:
724:
716:
687:
525:
332:
139:
2129:
2053:
1799:
1791:
1746:
1730:
1680:
1601:
1557:
1509:
1427:
1340:
1209:
1196:
1176:
1137:
1037:
1024:
995:
929:
917:
813:
770:
753:
745:
695:
533:
503:
434:
423:
407:
261:
2378:
2288:
2215:
2186:
2160:
2147:
2087:
2045:
2029:
2021:
2013:
1997:
1989:
1925:
1883:
1875:
1844:
1759:
1738:
1704:
1696:
1609:
1541:
1525:
1517:
1480:
1467:
1443:
1361:
1353:
1303:
1295:
1253:
1217:
1188:
1160:
1155:
1071:
1066:
1058:
1008:
944:
939:
853:
818:
803:
795:
711:
703:
520:
167:
112:
2318:
2331:
2142:
2103:
2095:
2037:
1900:
1862:
1815:
1807:
1627:
1575:
1459:
1390:
1261:
1245:
1201:
1142:
1106:
1086:
1050:
959:
949:
823:
557:
541:
2383:
328:
2207:
2199:
2191:
2178:
2134:
1933:
1908:
1836:
1664:
1635:
1319:
1150:
1124:
1076:
990:
892:
861:
833:
666:
641:
636:
617:
582:
471:
454:
446:
395:
383:
82:
62:
2250:
2326:
2273:
2111:
2061:
2005:
1964:
1913:
1570:
1501:
1382:
1094:
900:
874:
732:
565:
512:
491:
401:
266:
1828:
1820:
1672:
1406:
1327:
1311:
297:
In North
America, 55 genera and more than 200 species are found.
163:
1419:
1279:
912:
389:
377:
364:
151:
102:
92:
72:
154:(without serial punctures), the head with a broad neck, the
735:(furniture beetles, death watch beetles, spider beetles)
162:
of five articles. In almost all genera is a pair of
158:which are only slightly broader at the apex, and
2473:
1612:(jewel beetles, or metallic wood-boring beetles)
1335:(true weevils, bark beetles, ambrosia beetles)
502:
362:
348:
1578:(ironclad beetles, cylindrical bark beetles)
248:
355:
341:
29:
2242:List of subgroups of the order Coleoptera
1322:(straight snout weevils, New York weevil)
421:
1749:(cicada beetle, cicada parasite beetles)
2474:
1839:(false fireflies, long-lipped beetles)
664:
615:
2255:
2254:
663:
614:
489:
488:
420:
336:
2433:b70939b1-0562-4780-ba51-7b29565b7e7a
2371:1459adc9-bb18-4879-8798-b0e618731439
1865:(false metallic wood-boring beetles)
13:
14:
2498:
321:
1476:Perimylopidae, or Promecheilidae
1061:(minute brown scavenger beetles)
225:
211:
197:
183:
49:
1691:(variegated mud-loving beetles)
307:
1:
1520:(narrow-waisted bark beetles)
1212:(parasitic flat bark beetles)
1163:(short-winged flower beetles)
1019:(lady beetles, or God's cows)
756:(tooth-necked fungus beetles)
300:
2032:(sand-loving scarab beetles)
1422:(minute tree-fungus beetles)
1248:(silvanid flat bark beetles)
7:
2181:(primitive carrion beetles)
2137:(minute mud-loving beetles)
2000:(bumble bee scarab beetles)
1770:(soft-bodied plant beetles)
1741:(soft bodied plant beetles)
544:(predaceous diving beetles)
327:Omaliinae at Bugguide.net.
10:
2503:
2016:(scavenger scarab beetles)
2008:(enigmatic scarab beetles)
1181:Lamingtonium binnaberrense
895:(soft-wing flower beetles)
239:
145:
2263:
2239:
2169:
2155:(water scavenger beetles)
2120:
2086:
2077:
1955:
1946:
1899:
1888:Rhinorhipus tamborinensis
1874:
1758:
1729:
1667:(long-toed water beetles)
1626:
1600:
1591:
1560:(polypore fungus beetles)
1462:(palm and flower beetles)
1446:(tumbling flower beetles)
1393:(ant-like flower beetles)
1373:
1352:
1278:
1171:(lined flat bark beetles)
1145:(pleasing fungus beetles)
1085:
1040:(handsome fungus beetles)
976:
832:
778:
769:
744:
686:
677:
673:
659:
624:
610:
585:(burrowing water beetles)
498:
484:
430:
416:
372:
249:Systematics and evolution
46:Scientific classification
44:
37:
28:
23:
2202:(feather-winged beetles)
1536:(false longhorn beetles)
1483:(jugular-horned beetles)
1430:(false darkling beetles)
1401:(cryptic fungus beetles)
1220:(shining flower beetles)
560:(crawling water beetles)
459:Sikhotealinia zhiltzovae
1683:(forest stream beetles)
1544:(synchroa bark beetles)
1504:(dead log bark beetles)
1496:(fire-coloured beetles)
1385:(ant-like leaf beetles)
1119:(silken fungus beetles)
1053:(well polished beetles)
1027:(minute fungus beetles)
846:Acanthocnemus nigricans
233:Phloeonomus laesicollis
138:are a subfamily of the
2194:(round fungus beetles)
1528:(false flower beetles)
1512:(wedge-shaped beetles)
1470:(false blister beetle)
1454:(hairy fungus beetles)
1409:(conifer bark beetles)
1306:(leaf-rolling weevils)
967:(bark-gnawing beetles)
727:(wounded-tree beetles)
593:(false ground beetles)
515:(trout-stream beetles)
2441:Paleobiology Database
2106:(false clown beetles)
1928:(plate-thigh beetles)
1810:(false click beetles)
1715:(water-penny beetles)
1364:(ship-timber beetles)
1343:(pine flower weevils)
1191:(root-eating beetles)
1011:(minute bark beetles)
922:Phloiophilus edwardsi
449:(reticulated beetles)
219:Deinopteroloma pictum
2366:Fauna Europaea (new)
1984:(false stag beetles)
1969:Belohina inexpectata
1831:(net-winged beetles)
1794:(rare click beetles)
1707:(travertine beetles)
1699:(minute mud beetles)
1264:(dry-fungus beetles)
1099:Agapytho foveicollis
856:(false skin beetles)
719:(Jacobson's beetles)
439:Crowsoniella relicta
16:Subfamily of beetles
1982:Diphyllostomatidae
1314:(primitive weevils)
1127:(flat bark beetles)
877:(checkered beetles)
864:(fruitworm beetles)
552:(whirligig beetles)
191:Acidota subcarinata
39:Acidota subcarinata
2487:Beetle subfamilies
1847:(glowworm beetles)
1552:(darkling beetles)
1399:Archeocrypticidae
1256:(palmetto beetles)
1003:(dry bark beetles)
790:(longhorn beetles)
2469:
2468:
2257:Taxon identifiers
2248:
2247:
2235:
2234:
2231:
2230:
2227:
2226:
2219:
2211:
2210:(carrion beetles)
2203:
2195:
2182:
2156:
2138:
2107:
2099:
2073:
2072:
2065:
2057:
2049:
2041:
2033:
2025:
2017:
2009:
2001:
1993:
1985:
1972:
1942:
1941:
1929:
1921:
1891:
1866:
1848:
1840:
1832:
1824:
1811:
1803:
1795:
1787:
1786:(soldier beetles)
1779:
1776:Brachypsectridae
1771:
1750:
1742:
1716:
1708:
1700:
1692:
1684:
1676:
1668:
1655:
1647:
1639:
1613:
1587:
1586:
1579:
1561:
1553:
1545:
1537:
1534:Stenotrachelidae
1529:
1521:
1513:
1505:
1497:
1484:
1471:
1463:
1455:
1447:
1439:
1438:(blister beetles)
1431:
1423:
1410:
1402:
1394:
1386:
1365:
1344:
1336:
1323:
1315:
1307:
1299:
1289:
1265:
1257:
1249:
1221:
1213:
1205:
1192:
1184:
1172:
1164:
1146:
1128:
1120:
1102:
1062:
1054:
1041:
1028:
1020:
1012:
1004:
986:Akalyptoischiidae
968:
925:
908:
896:
878:
865:
857:
849:
799:
791:
765:
764:
757:
736:
728:
720:
707:
699:
655:
654:
632:
606:
605:
602:
601:
594:
586:
578:
561:
553:
545:
537:
529:
516:
480:
479:
462:
450:
442:
205:Brathinus nitidus
132:
131:
128:
2494:
2462:
2461:
2449:
2448:
2436:
2435:
2426:
2425:
2413:
2412:
2410:NHMSYS0001719277
2400:
2399:
2387:
2386:
2374:
2373:
2361:
2360:
2348:
2347:
2335:
2334:
2322:
2321:
2312:
2311:
2299:
2298:
2297:
2284:
2283:
2282:
2252:
2251:
2217:
2209:
2201:
2193:
2180:
2154:
2136:
2105:
2097:
2084:
2083:
2079:Staphyliniformia
2063:
2055:
2047:
2039:
2031:
2023:
2015:
2007:
1999:
1991:
1983:
1966:
1953:
1952:
1927:
1918:Declinia relicta
1915:
1885:
1864:
1858:Sinopyrophoridae
1853:Rhagophthalmidae
1846:
1838:
1830:
1822:
1809:
1801:
1793:
1785:
1777:
1769:
1768:Artematopodidae
1748:
1740:
1714:
1706:
1698:
1690:
1682:
1675:(riffle beetles)
1674:
1666:
1654:(turtle beetles)
1653:
1645:
1637:
1611:
1598:
1597:
1577:
1559:
1551:
1543:
1535:
1527:
1519:
1511:
1503:
1495:
1482:
1469:
1461:
1453:
1445:
1437:
1429:
1421:
1408:
1400:
1392:
1384:
1363:
1342:
1334:
1321:
1313:
1305:
1298:(fungus weevils)
1297:
1288:
1287:
1283:
1270:Tasmosalpingidae
1263:
1255:
1247:
1219:
1211:
1203:
1190:
1178:
1170:
1162:
1144:
1126:
1118:
1096:
1060:
1052:
1039:
1026:
1018:
1010:
1002:
966:
919:
902:
894:
876:
863:
855:
843:
842:Acanthocnemidae
797:
789:
776:
775:
755:
734:
726:
718:
705:
697:
684:
683:
675:
674:
661:
660:
630:
612:
611:
592:
584:
572:
559:
551:
543:
535:
528:(ground beetles)
527:
514:
500:
499:
486:
485:
456:
448:
436:
435:Crowsoniellidae
418:
417:
357:
350:
343:
334:
333:
315:
311:
229:
215:
201:
187:
142:, rove beetles.
126:
54:
53:
33:
21:
20:
2502:
2501:
2497:
2496:
2495:
2493:
2492:
2491:
2472:
2471:
2470:
2465:
2457:
2452:
2444:
2439:
2431:
2429:
2421:
2416:
2408:
2403:
2395:
2390:
2382:
2377:
2369:
2364:
2356:
2351:
2343:
2338:
2330:
2325:
2317:
2315:
2307:
2302:
2293:
2292:
2287:
2278:
2277:
2272:
2259:
2249:
2244:
2223:
2165:
2116:
2098:(clown beetles)
2069:
2040:(betsy beetles)
1948:Scarabaeiformia
1938:
1895:
1870:
1802:(click beetles)
1778:(Texas beetles)
1754:
1725:
1721:Ptilodactylidae
1646:(cedar beetles)
1622:
1583:
1566:Trictenotomidae
1452:Mycetophagidae
1369:
1348:
1285:
1284:
1282:
1274:
1226:Phloeostichidae
1177:Lamingtoniidae
1169:Laemophloeidae
1117:Cryptophagidae
1081:
972:
955:Thanerocleridae
918:Phloiophilidae
905:Metaxina ornata
870:Chaetosomatidae
828:
761:
740:
698:(auger beetles)
679:Bostrichiformia
669:
651:
647:Torridincolidae
631:(skiff beetles)
629:Hydroscaphidae
620:
598:
591:Trachypachidae
536:(tiger beetles)
494:
476:
426:
412:
368:
361:
324:
319:
318:
312:
308:
303:
292:Aphaenostemmini
251:
242:
235:
230:
221:
216:
207:
202:
193:
188:
148:
125:
48:
17:
12:
11:
5:
2500:
2490:
2489:
2484:
2467:
2466:
2464:
2463:
2450:
2437:
2427:
2414:
2401:
2388:
2375:
2362:
2353:Fauna Europaea
2349:
2336:
2323:
2313:
2300:
2285:
2269:
2267:
2261:
2260:
2246:
2245:
2240:
2237:
2236:
2233:
2232:
2229:
2228:
2225:
2224:
2222:
2221:
2218:(rove beetles)
2216:Staphylinidae
2213:
2205:
2197:
2189:
2184:
2175:
2173:
2171:Staphylinoidea
2167:
2166:
2164:
2163:
2158:
2153:Hydrophilidae
2150:
2145:
2140:
2132:
2126:
2124:
2122:Hydrophiloidea
2118:
2117:
2115:
2114:
2109:
2101:
2092:
2090:
2081:
2075:
2074:
2071:
2070:
2068:
2067:
2064:(hide beetles)
2059:
2051:
2048:(rain beetles)
2043:
2035:
2027:
2024:(stag beetles)
2019:
2011:
2003:
1995:
1987:
1979:
1977:Bolboceratidae
1974:
1961:
1959:
1950:
1944:
1943:
1940:
1939:
1937:
1936:
1931:
1923:
1911:
1905:
1903:
1897:
1896:
1894:
1893:
1884:Rhinorhipidae
1880:
1878:
1876:Rhinorhipoidea
1872:
1871:
1869:
1868:
1860:
1855:
1850:
1842:
1834:
1826:
1818:
1813:
1805:
1797:
1789:
1781:
1773:
1764:
1762:
1756:
1755:
1753:
1752:
1744:
1735:
1733:
1727:
1726:
1724:
1723:
1718:
1710:
1702:
1694:
1689:Heteroceridae
1686:
1678:
1670:
1662:
1657:
1652:Chelonariidae
1649:
1644:Callirhipidae
1641:
1638:(pill beetles)
1632:
1630:
1624:
1623:
1621:
1620:
1615:
1606:
1604:
1595:
1589:
1588:
1585:
1584:
1582:
1581:
1573:
1568:
1563:
1555:
1550:Tenebrionidae
1547:
1539:
1531:
1523:
1515:
1507:
1499:
1491:
1486:
1478:
1473:
1465:
1457:
1449:
1441:
1433:
1425:
1417:
1412:
1404:
1396:
1388:
1379:
1377:
1375:Tenebrionoidea
1371:
1370:
1368:
1367:
1358:
1356:
1350:
1349:
1347:
1346:
1338:
1333:Curculionidae
1330:
1325:
1317:
1309:
1301:
1292:
1290:
1280:Curculionoidea
1276:
1275:
1273:
1272:
1267:
1259:
1251:
1243:
1241:Protocucujidae
1238:
1233:
1228:
1223:
1215:
1207:
1199:
1194:
1186:
1174:
1166:
1158:
1153:
1148:
1140:
1135:
1133:Cybocephalidae
1130:
1122:
1114:
1109:
1104:
1091:
1089:
1083:
1082:
1080:
1079:
1074:
1069:
1064:
1056:
1048:
1043:
1035:
1033:Discolomatidae
1030:
1022:
1017:Coccinellidae
1014:
1006:
1001:Bothrideridae
998:
993:
988:
982:
980:
978:Coccinelloidea
974:
973:
971:
970:
965:Trogossitidae
962:
957:
952:
947:
942:
937:
932:
927:
915:
910:
898:
890:
885:
883:Lophocateridae
880:
872:
867:
859:
851:
838:
836:
830:
829:
827:
826:
821:
816:
811:
806:
801:
798:(leaf beetles)
796:Chrysomelidae
793:
784:
782:
780:Chrysomeloidea
773:
767:
766:
763:
762:
760:
759:
750:
748:
742:
741:
739:
738:
730:
725:Nosodendridae
722:
717:Jacobsoniidae
714:
709:
706:(skin beetles)
701:
692:
690:
681:
671:
670:
657:
656:
653:
652:
650:
649:
644:
639:
634:
625:
622:
621:
608:
607:
604:
603:
600:
599:
597:
596:
588:
580:
575:Meru phyllisae
568:
563:
555:
547:
539:
531:
523:
518:
509:
507:
496:
495:
482:
481:
478:
477:
475:
474:
469:
467:Micromalthidae
464:
452:
444:
431:
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1554:
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1510:Ripiphoridae
1508:
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743:
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702:
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696:Bostrichidae
694:
693:
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623:
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581:
579:
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569:
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556:
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534:Cicindelidae
532:
530:
524:
522:
519:
517:
511:
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497:
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366:
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335:
329:
326:
325:
310:
306:
298:
293:
290:
288:
285:
283:
282:Corneolabiini
280:
278:
277:Hadrognathini
275:
273:
270:
268:
265:
263:
260:
259:
258:
256:
246:
234:
228:
223:
220:
214:
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192:
186:
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180:
179:
177:
173:
169:
165:
161:
157:
153:
143:
141:
140:Staphylinidae
137:
127:MacLeay, 1825
124:
121:
118:
117:
114:
113:Staphylinidae
111:
108:
107:
104:
101:
98:
97:
94:
91:
88:
87:
84:
81:
78:
77:
74:
71:
68:
67:
64:
61:
58:
57:
52:
47:
43:
40:
36:
32:
27:
22:
19:
2264:
2143:Helophoridae
2135:Georissidae
2130:Epimetopidae
2046:Pleocomidae
2030:Ochodaeidae
2014:Hybosoridae
1998:Glaphyridae
1990:Geotrupidae
1968:
1965:Belohinidae
1926:Eucinetidae
1917:
1914:Decliniidae
1887:
1845:Phengodidae
1784:Cantharidae
1739:Dascillidae
1731:Dascilloidea
1713:Psephenidae
1705:Lutrochidae
1697:Limnichidae
1610:Buprestidae
1602:Buprestoidea
1542:Synchroidae
1526:Scraptiidae
1518:Salpingidae
1481:Prostomidae
1468:Oedemeridae
1444:Mordellidae
1362:Lymexylidae
1354:Lymexyloidea
1304:Attelabidae
1296:Anthribidae
1254:Smicripidae
1218:Phalacridae
1202:Nitidulidae
1197:Myraboliidae
1189:Monotomidae
1180:
1161:Kateretidae
1138:Cyclaxyridae
1098:
1095:Agapythidae
1059:Latridiidae
1009:Cerylonidae
996:Anamorphidae
930:Phycosecidae
921:
904:
901:Metaxinidae
854:Biphyllidae
845:
814:Orsodacnidae
771:Cucujiformia
704:Dermestidae
574:
513:Amphizoidae
458:
438:
424:Archostemata
408:Holometabola
406:Superorder:
400:Infraclass:
309:
296:
272:Eusphalerini
262:Anthophagini
252:
243:
232:
218:
204:
190:
149:
135:
133:
122:
38:
18:
2379:iNaturalist
2289:Wikispecies
2187:Hydraenidae
2161:Spercheidae
2148:Hydrochidae
2112:Synteliidae
2096:Histeridae
2088:Histeroidea
2038:Passalidae
2006:Glaresidae
1863:Throscidae
1823:(fireflies)
1821:Lampyridae
1808:Eucnemidae
1800:Elateridae
1760:Elateroidea
1576:Zopheridae
1460:Mycteridae
1391:Anthicidae
1262:Sphindidae
1246:Silvanidae
1156:Hobartiidae
1143:Erotylidae
1072:Mycetaeidae
1067:Murmidiidae
1051:Euxestidae
945:Rentoniidae
819:Oxypeltidae
804:Disteniidae
637:Lepiceridae
566:Hygrobiidae
558:Haliplidae
542:Dytiscidae
521:Aspidytidae
168:Proteininae
119:Subfamily:
2476:Categories
2208:Silphidae
2200:Ptiliidae
2192:Leiodidae
2179:Agyrtidae
2022:Lucanidae
1901:Scirtoidea
1837:Omethidae
1816:Jurasaidae
1665:Dryopidae
1636:Byrrhidae
1628:Byrrhoidea
1320:Brentidae
1125:Cucujidae
1107:Boganiidae
1087:Cucujoidea
960:Thymalidae
950:Rhadalidae
893:Melyridae
862:Byturidae
824:Vesperidae
733:Ptiniidae
583:Noteridae
550:Gyrinidae
526:Carabidae
455:Jurodidae
447:Cupedidae
394:Subclass:
384:Arthropoda
365:Coleoptera
301:References
287:Coryphiini
170:, and the
103:Coleoptera
83:Arthropoda
24:Omaliinae
2482:Omaliinae
2309:Omaliinae
2295:Omaliinae
2265:Omaliinae
2062:Trogidae
2056:(scarabs)
1934:Scirtidae
1909:Clambidae
1502:Pythidae
1436:Meloidae
1383:Aderidae
1286:(weevils)
1151:Helotidae
1077:Teredidae
991:Alexiidae
875:Cleridae
834:Cleroidea
667:Polyphaga
665:Suborder
618:Myxophaga
616:Suborder
571:Meruidae
490:Suborder
472:Ommatidae
422:Suborder
396:Pterygota
376:Kingdom:
176:prostheca
136:Omaliinae
123:Omaliinae
69:Kingdom:
63:Eukaryota
2327:BugGuide
2316:BioLib:
2280:Q1946931
2274:Wikidata
1829:Lycidae
1673:Elmidae
1571:Ulodidae
1407:Boridae
1312:Belidae
913:Peltidae
506:families
492:Adephaga
402:Neoptera
382:Phylum:
378:Animalia
367:families
267:Omaliini
174:lacks a
172:mandible
156:antennae
109:Family:
79:Phylum:
73:Animalia
59:Domain:
1420:Ciidae
1328:Caridae
390:Insecta
388:Class:
363:Extant
240:Ecology
146:Anatomy
99:Order:
93:Insecta
89:Class:
2459:493957
2430:NZOR:
2397:678457
2384:359912
2358:263922
504:Extant
255:tribes
164:ocelli
152:elytra
2454:WoRMS
2446:69966
2423:66542
2345:8VR5R
2332:38166
160:tarsi
2418:NCBI
2392:ITIS
2319:5747
134:The
2405:NBN
2340:CoL
2304:AFD
2478::
2456::
2443::
2420::
2407::
2394::
2381::
2368::
2355::
2342::
2329::
2306::
2291::
2276::
314:p.
257::
178:.
1971:)
1967:(
1920:)
1916:(
1890:)
1886:(
1183:)
1179:(
1101:)
1097:(
924:)
920:(
907:)
903:(
848:)
844:(
577:)
573:(
461:)
457:(
441:)
437:(
356:e
349:t
342:v
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