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Nautilus (genus)

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specimens have demonstrated that prolonged exposure to temperatures over 25 degrees Celsius will eventually result in death after several days. However, individuals have been documented to experience temperatures higher than this, and survive, as long as they are not exposed to these temperatures for
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eggs are laid in capsules, usually 3–4 cm long, which gradually harden when exposed to seawater. It is not yet known how exactly the juveniles break out of these capsules, yet it has been hypothesized that they are able to chew their way out, using their beak. The genus exhibits a skewed sex
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shell in an aquarium resulted in the shell floating for over two years, and one recovered shell was revealed to have been afloat for a period of 11 years. Furthermore, shells have been demonstrated to drift considerable distances in this time, contributing to their extensive distribution across
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participate in routine vertical migration, in which they ascend to shallow areas of reefs, between 100 and 150 metres deep, during the night to feed, and later descend to depths of 250–350 metres during the day, however, these depths may vary depending on local geographic characteristics.
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are generally fed a diet of whole shrimp, fish, crab, and lobster moults. Several aquaria around the world host specimens of the genus, however, there have not yet been any successful attempts of breeding in captivity, despite viable eggs being produced at several locations. Two
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In addition to defining species, genetic studies have also provided evidence for the evolution of the genus over time. Mitochondrial DNA studies have indicated that the genus is currently undergoing evolutionary radiation in the Indo-Pacific. The divergence between the genus
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have been observed to spend days in deeper areas around coral reefs, to avoid predation from turtles and carnivorous fish, and ascend to shallow areas of the reef during nights. Here, they engage in scavenging activity, seeking out animal remains, and the moults of
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species usually travel and feed alone. Nautilus return to deeper areas following daybreak and also lay eggs in these locations, which take approximately one year to hatch. This behaviour may have ensured their survival during the
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have been noted to exhibit an extensive range of depth, close to 500 metres, however, they were demonstrated to be at risk of implosion when exceeding their depth and pressure limits. Depending on the species, the shells of live
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shells can float to the surface, where they can remain for a considerable time period, however the buoyancy of shells after death was found to be dependent on several factors, such as the rate of decay. An experiment with a
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Various authors claim that the genus consists of between four and seven extant species; this remains the subject of debate. Nautiloids are typically found in shallow ocean waters in tropical seas, mainly within the
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ratio, biased towards male individuals. This phenomenon has been observed at several locations around the globe, with population samples consisting of up to 95% males. The reason for this is currently unknown.
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in a trap. Additionally, many drift shells exhibit small holes which match the patterns produced by octopus boring into shell to feed. Teleosts, such as triggerfish, have also been observed to feed on
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studies, utilising two gene regions, also have led to the notion that many of the morphological differences between differeont Nautilus populations are simply localised variations within the single
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Dunstan, A.; Alanis, O.; Marshall, J. (November 2010). "Nautilus pompilius fishing and population decline in the Philippines: A comparison with an unexploited Australian Nautilus population".
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Landman, Neil H.; Cochran, J. Kirk; Rye, Danny M.; Tanabe, Kazushige; Arnold, John M. (1994). "Early life history of Nautilus: evidence from isotopic analyses of aquarium-reared specimens".
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Bonacum, James; Landman, Neil H.; Mapes, Royal H.; White, Matthew M.; White, Alicia-Jeannette; Irlam, Justin (March 2011). "Evolutionary Radiation of Present-DayNautilusandAllonautilus".
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are localised to the Indo-Pacific, specifically the tropical seas within this area, however the full extent of their geographic distribution has yet to be recorded. The movements of
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are opportunistic scavengers and feed on a variety of crustaceans, including their moults, and fish, however, they have been observed to feed on chicken and bat bait. Initially,
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Combosch, David J.; Lemer, Sarah; Ward, Peter D.; Landman, Neil H.; Giribet, Gonzalo (2017-10-07). "Genomic signatures of evolution in Nautilus —An endangered living fossil".
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can occasionally be found closer to the surface than 100 metres, however, the minimum depth they can reach is determined by factors such as water temperature and season. All
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were validated by the genetic study, two undescribed, but genetically distinct, species were discovered in the South Pacific. One of these cryptic species was recorded from
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In the years following this conclusion, fossil species were still sometimes assigned to the genus; however, many other authors insisted that these be excluded. In
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includes several species represented in the fossil record; however, these have also been contentious in their placement, and some are only provisionally accepted.
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Many shells recovered from areas of the world have not yet been identified down to the level of species, however, are still identifiable as members of the genus
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Zoological Results Based on Material from New Britain, New Guinea, Loyalty Islands and Elsewhere: The anatomy and development of Peripatus novae-britanniae
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were thought to actively hunt certain prey, however, this activity has only been recorded in traps, where prey species are confined in close proximity to
1155:. This study resulted in the description of three new species in 2023, recognizing the species from Fiji and American Samoa as two taxa rather than one. 2385:
Chamberlain, John A.; Ward, Peter D.; Weaver, J. Scott (1981). "Post-mortem ascent of Nautilus shells: implications for cephalopod paleobiogeography".
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There has been much debate over the validity of species within the genus, and several identified species have since been reclassified or determined as
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The feeding behaviour of the genus has been identified from observation of captive individuals, as well as the stomach contents of wild specimens.
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Wray, Charles G.; Landman, Neil H.; Saunders, W. Bruce; Bonacum, James (1995). "Genetic divergence and geographic diversification in Nautilus".
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carries shells from the Philippines to areas such as Japan, and the Equatorial current is responsible for many of the shells recovered from the
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exist along a spectrum, with a range of individuals displaying a combination of characteristics, further invalidating them as separate species.
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Ward, Peter; Dooley, Frederick; Barord, Gregory Jeff (2016-02-29). "Nautilus: biology, systematics, and paleobiology as viewed from 2015".
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do not have an elaborate courtship process. Males have been observed to attempt to mate with any object the same size and shape as another
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in captivity is relatively poor, primarily due to the stress that individuals are subjected to during transportation. As many as 50-80% of
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are unable to easily move across areas deeper than 800 metres, and most of their activity occurs at a depth of 100–300 metres deep.
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die during transportation, and this percentage can be higher if individuals are exposed to high temperatures. In captivity,
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has been contentious for decades, and the genus has been reconfigured and redefined several times throughout its history.
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species lack ocular muscles and instead move via a stalk, which contains both muscle and connective tissue. Additionally,
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based on both genetic and morphological data. One study, sampling Nautiluses in 2012, demonstrated that the features of
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Gregory J. Barord, David J. Combosch, Gonzalo Giribet, Neil Landman, Sarah Lemer, Job Veloso et Peter D. Ward, « 
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in aquaria, however, specific care is necessary in order to ensure their survival in captivity. The survival rate of
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have unique sensory organs, which differ from related genera in several ways. Unlike other cephalopods, the eyes of
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Barber, V. C.; Wright, D. E. (1969). "The fine structure of the sense organs of the cephalopod mollusc Nautilus".
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are able to ascend at speeds of approximately 2 metres per minute and descend at speeds of 3 metres per minute.
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split diverged further, involving migrations east to Vanuatu, Fiji, and American Samoa, as well as west, to the
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shell from Broome, Australia. The majority of recovered shells, like this example, are found as drift shells.
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species, however these did not exactly correlate to the described species of the genus. Whilst the status of
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locate these food sources by using their tentacles, which have chemosensory functions, as well as by sight.
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also perform several sensory functions. Their ocular and preocular tentacles feature cilia, and operate as
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drift shells are able to make their way across oceans via currents. Following the death of an individual,
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overfishing in the Philippines, which resulted in an 80% decline in the population from 1980 to 2010.
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temperature thresholds is also supplemented by the study of captive individuals in aquaria. Captive
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longer than 10 hours. Optimal temperatures for the genus tend to range from 9-21 degrees Celsius.
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should be placed in the genus, despite many fossil species having already been assigned to it.
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coastal areas. Several ocean currents have been identified to contribute to this process. The
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reproduction comes from captive species in aquaria. From these specimens, it appears that
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was well studied in the 19th century, it was not until 1996 that soft tissues of
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Miller, A. K. (September 1949). "The Last Surge of the Nautiloid Cephalopods".
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Convention on International Trade in Endangered Species of Wild Fauna and Flora
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Linnaeus, 1758 (Mollusca, Cephalopoda) from the Coral Sea and South Pacific
2015: 1361: 915: 659: 513: 503: 433: 379:, in Washington State and from Late-Eocene to Early Oligocene sediments in 371: 249: 2328: 3652: 3572: 3317: 3044: 2924: 2900: 2831: 2811: 2793: 2730: 2705: 2213: 1325: 1324:. Shells have been found across a wide range of coastal areas, including 1197: 394: 57: 3644: 3349: 3275: 2992: 2661: 2312: 2179: 2007: 1687: 1649: 1403: 1329: 1247: 486: 469: 380: 355: 342: 190: 102: 67: 3657: 2441:
Dunstan, Andrew J.; Ward, Peter D.; Marshall, N. Justin (2011-02-22).
2077: 1902: 1877: 1608:(Nautilidae, Cephalopoda) from Pangasinan, northwestern Philippines". 1143:. Whilst this study recorded five species, its results suggested that 1052:
Several genetic studies have also been conducted on select species of
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female has two reproductive organs whose functions are unknown, the
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Arthur K. Miller provided a detailed description of the shell of
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Nautilus : the biology and paleobiology of a living fossil
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Goedert, James L.; Kiel, Steffen; Tsai, Cheng-Hsiu (2022).
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In addition to observing wild specimens, our knowledge of
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Zeitschrift für Zellforschung und Mikroskopische Anatomie
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Wani, R.; et al. (2008). "First discovery of fossil
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species are likely endangered, based on information from
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10.2517/1342-8144(2008)12[89:FDOFNP]2.0.CO;2
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from the genus. Furthermore, some biologists claim that
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species. This same 2011 study, however, suggested that
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are from Early Pleistocene sediments off the coast of
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A 2017 study determined that there were likely five
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(2010). 520:, he determined that only extant species of 2298: 2148:"Notes on the Anatomy of Nautilus Pompilius" 1804:: CS1 maint: multiple names: authors list ( 1647: 662:; also biggest and most widespread species) 1932:Teichert, Curt; Matsumoto, Tatsuro (1987), 1599: 1597: 1595: 1273: 1188:likely occurred around New Guinea, and the 1000:species have been recognised, specifically 983: 383:. The oldest fossils of the modern species 2605: 2591: 2557: 2022: 1866: 123: 2484: 2466: 1901: 1016:, and three more were described in 2023. 1592: 1541:The majority of our knowledge regarding 1483: 1307: 1162: 1082: 2145: 3787: 2047:, vol. 1143, 2023, p. 51-69. 1977: 1479: 416:, including all species in the genera 3539: 3538: 2586: 2508: 2506: 2504: 2436: 2434: 2432: 2294: 2292: 2290: 2195: 2193: 2191: 2189: 2146:Griffin, Lawrence E. (October 1897). 2117: 2115: 2113: 2111: 2059: 2057: 2055: 2053: 1767: 1765: 1763: 1761: 1759: 1757: 1755: 1753: 1751: 1749: 1747: 1745: 1743: 1741: 1739: 1648:Ward, P. D.; Saunders, W. B. (1997). 974:Nomen Dubium: likely a giant form of 460:The classification of species within 3772:FDF3E501-41CD-403C-93F5-595762F38B76 3736:18BE1A77-7D47-D6B3-4636-3A6B5569704E 1927: 1925: 1923: 1921: 1825: 1823: 1821: 1819: 1817: 1815: 1737: 1735: 1733: 1731: 1729: 1727: 1725: 1723: 1721: 1719: 1643: 1641: 1639: 1603: 1566:male has a reproductive organ named 362:in this genus differ significantly, 2612: 2570:. University Press. pp. 778–9. 1492:at the Aquarium Finisterrae, Spain. 476:, originally defined as any coiled- 13: 2501: 2429: 2287: 2186: 2108: 2050: 1934:The Ancestry of the Genus Nautilus 1047: 598:Meek, 1862 (provisional placement) 424:, is listed on Appendix II of the 14: 3811: 2578: 1918: 1812: 1716: 1636: 1215: 455: 849: 831: 813: 795: 777: 759: 723:(provisional placement) – Taiwan 147: 55: 19:This article is about the genus 2378: 2343: 2236: 2202:American Malacological Bulletin 2139: 1536: 1413:Cretaceous-Paleogene extinction 1167:The first and oldest fossil of 408:, and more specifically to the 1971: 1832:Swiss Journal of Palaeontology 1702: 1116:was a species synonymous with 412:species. The entire family of 1: 2364:10.1016/j.fishres.2010.06.015 1585: 2468:10.1371/journal.pone.0016311 1710:"Checklist of CITES species" 1476:withdraw into their shells. 1455: 1393: 1368:at these sites, however, as 1250:, which are small sacs with 1158: 1096:Nautilus attracted to bait, 7: 1942:10.1007/978-1-4899-5040-6_2 10: 3816: 1173:Philippine National Museum 1042:Allonautilus scrobiculatus 1022:Allonautilus scrobiculatus 538: 33: 18: 3547: 3523: 3492: 3458: 3421: 3371: 3348: 3339: 3308: 3274: 3239: 3230: 3180: 3150: 3107:Black-foot opihi/Hawaiian 3097: 3069: 3000: 2991: 2935: 2856: 2802: 2759: 2627: 2618: 2535:10.1017/s009483730001112x 2407:10.1017/s0094837300025549 2265:10.1017/s009483730001321x 1844:10.1007/s13358-016-0112-7 1680:10.1017/S0022336000036039 1278:Species within the genus 868:Other described species: 279: 274: 255: 248: 144:Scientific classification 142: 131: 122: 44: 2978:Placopecten magellanicus 2822:New Zealand green-lipped 1610:Paleontological Research 1274:Habitat and distribution 1135:, whilst the other from 984:Controversy over species 400:The commonly used term ' 3483:Acanthopleura granulata 3052:(group of four species) 1944:(inactive 2024-04-15), 1659:Journal of Paleontology 1496:It is possible to keep 1184:, and its sister taxon 600:- Western North America 3476:Acanthopleura echinata 3386:Atlantic white-spotted 2837:Asian/Philippine green 2564:Arthur Willey (1902). 1580:Owen's laminated organ 1568:Van der Hoeven's organ 1493: 1317: 1176: 1101: 1018:Nautilus scrobiculatus 680:Nautilus praepompilius 3718:Paleobiology Database 3412:Amphioctopus fangsiao 3056:South African abalone 2928:("true oyster" genus) 2694:California butterclam 2037:Three new species of 1576:Organ of Valenciennes 1517:eggs were hatched at 1487: 1311: 1166: 1095: 1078:Nautilus stenomphalus 728:Nautilus vanuatuensis 709:G.B. Sowerby II, 1849 705:Nautilus stenomphalus 645:G.B. Sowerby II, 1849 641:Nautilus macromphalus 618:Nautilus geelongensis 559:Nautilus balcombensis 446:South Pacific islands 319:Nautilus vanuatuensis 312:Nautilus stenomphalus 291:Nautilus macromphalus 3137:Ribbed Mediterranean 2652:Grooved carpet shell 2214:10.4003/006.029.0221 1044:had been dissected. 711:– Great Barrier Reef 480:species with simple 377:Hoko River Formation 23:. For the family of 3132:Rayed Mediterranean 2726:Japanese littleneck 2527:1994Pbio...20...40L 2459:2011PLoSO...616311D 2399:1981Pbio....7..494C 2257:1995Pbio...21..220W 2152:Zoological Bulletin 1672:1997JPal...71.1054W 1480:Nautilus in aquaria 1336:, the Philippines, 1151:were synonyms with 690:Nautilus samoaensis 582:Nautilus butonensis 570:Nautilus belauensis 305:Nautilus samoaensis 284:Nautilus belauensis 3505:Land snail farming 2919:Gillardeau oysters 2637:Atlantic jackknife 2352:Fisheries Research 2313:10.1007/bf00335442 1606:Nautilus pompilius 1494: 1490:Nautilus pompilius 1318: 1242:Below their eyes, 1190:Great Barrier Reef 1177: 1169:chambered nautilus 1102: 1074:Nautilus pompilius 990:taxonomic synonyms 874:Described species 743:Nautilus vitiensis 652:Nautilus pompilius 594:Nautilus campbelli 547:Nautilus altifrons 444:to the open water 410:Nautilus pompilius 386:Nautilus pompilius 326:Nautilus vitiensis 298:Nautilus pompilius 135:Nautilus pompilius 36:Chambered nautilus 3782: 3781: 3705:Open Tree of Life 3541:Taxon identifiers 3532: 3531: 3510:Gastropod anatomy 3469:Chiton magnificus 3454: 3453: 3431:New Zealand arrow 3335: 3334: 3331: 3330: 3206:Kelletia lischkei 3171:Littorina sitkana 3117:Yellow-foot opihi 2987: 2986: 2881:Colchester native 2127:Discover Magazine 2078:10.1111/mec.14344 2072:(21): 5923–5938. 2066:Molecular Ecology 1951:978-1-4899-5042-0 1903:10.1111/iar.12442 1783:978-90-481-3298-0 1106:mitochondrial DNA 1093: 981: 980: 863: 842: 824: 806: 788: 770: 752: 737: 722: 717:Nautilus taiwanus 710: 699: 685: 657: 646: 635: 623: 611: 606:Nautilus cookanus 599: 587: 575: 564: 552: 516:of the genus. In 442:Strait of Malacca 334: 333: 244: 16:Genus of molluscs 3807: 3800:Nautiloid genera 3775: 3774: 3762: 3761: 3749: 3748: 3739: 3738: 3726: 3725: 3713: 3712: 3700: 3699: 3687: 3686: 3674: 3673: 3661: 3660: 3648: 3647: 3635: 3634: 3622: 3621: 3609: 3608: 3596: 3595: 3583: 3582: 3581: 3568: 3567: 3566: 3536: 3535: 3346: 3345: 3260:Chorus giganteus 3237: 3236: 2998: 2997: 2961:Pecten jacobaeus 2668:Mactra stultorum 2625: 2624: 2607: 2600: 2593: 2584: 2583: 2572: 2571: 2561: 2555: 2554: 2510: 2499: 2498: 2488: 2470: 2438: 2427: 2426: 2382: 2376: 2375: 2347: 2341: 2340: 2296: 2285: 2284: 2240: 2234: 2233: 2197: 2184: 2183: 2143: 2137: 2136: 2134: 2133: 2119: 2106: 2105: 2061: 2048: 2033: 2020: 2019: 1975: 1969: 1968: 1962: 1954: 1929: 1916: 1915: 1905: 1873: 1864: 1863: 1827: 1810: 1809: 1803: 1795: 1769: 1714: 1713: 1706: 1700: 1699: 1645: 1634: 1633: 1601: 1519:Waikiki Aquarium 1388:Marshall Islands 1384:Kuroshio Current 1268:mechanoreceptors 1118:A. scrobiculatus 1094: 1058:N. scrobiculatus 961:N. scrobiculatus 948:N. scrobiculatus 935:N. scrobiculatus 925:N. umbiliculatus 913:Reclassified as 908:N. scrobiculatus 871: 870: 856: 853: 840: 835: 822: 817: 804: 799: 786: 781: 768: 763: 746: 731: 720: 708: 693: 683: 655: 644: 633: 630:Nautilus javanus 621: 609: 597: 585: 573: 562: 550: 236: 152: 151: 127: 117: 54: 50:Temporal range: 42: 41: 3815: 3814: 3810: 3809: 3808: 3806: 3805: 3804: 3785: 3784: 3783: 3778: 3770: 3765: 3757: 3752: 3744: 3742: 3734: 3729: 3721: 3716: 3708: 3703: 3695: 3690: 3682: 3677: 3669: 3664: 3656: 3651: 3643: 3638: 3630: 3625: 3617: 3612: 3604: 3599: 3591: 3586: 3577: 3576: 3571: 3562: 3561: 3556: 3543: 3533: 3528: 3519: 3515:Bivalve anatomy 3488: 3450: 3436:Japanese flying 3417: 3367: 3327: 3304: 3270: 3226: 3176: 3146: 3127:Common European 3093: 3065: 2983: 2967:Peruvian calico 2931: 2852: 2849:(mussel family) 2798: 2755: 2700:Senilia senilis 2614: 2613:Edible mollusks 2611: 2581: 2576: 2575: 2562: 2558: 2511: 2502: 2439: 2430: 2383: 2379: 2348: 2344: 2297: 2288: 2241: 2237: 2198: 2187: 2164:10.2307/1535418 2144: 2140: 2131: 2129: 2123:"Coils of Time" 2121: 2120: 2109: 2062: 2051: 2034: 2023: 1992:10.2307/2405560 1976: 1972: 1956: 1955: 1952: 1930: 1919: 1874: 1867: 1828: 1813: 1797: 1796: 1784: 1770: 1717: 1708: 1707: 1703: 1646: 1637: 1602: 1593: 1588: 1539: 1482: 1458: 1396: 1276: 1218: 1161: 1114:N. macromphalus 1083: 1068:are members of 1062:N. stenomphalus 1050: 1048:Genetic studies 1010:N. stenomphalus 1006:N. macromphalus 986: 864: 854: 845: 841:N. stenomphalus 836: 827: 823:N. vanuatuensis 818: 809: 800: 791: 787:N. macromphalus 782: 773: 764: 684:Shimansky, 1957 671:N. p. suluensis 666:N. p. pompilius 647:– New Caledonia 612:- North America 610:Whitfield, 1892 563:(Chapman, 1915) 551:(Chapman, 1915) 541: 512:, becoming the 458: 364:morphologically 275:Living species 270: 261: 235: 146: 138:in an aquarium 118: 116: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 52: 51: 48: 38: 32: 17: 12: 11: 5: 3813: 3803: 3802: 3797: 3780: 3779: 3777: 3776: 3763: 3750: 3740: 3727: 3714: 3701: 3688: 3675: 3662: 3649: 3636: 3623: 3610: 3597: 3584: 3569: 3553: 3551: 3545: 3544: 3530: 3529: 3524: 3521: 3520: 3518: 3517: 3512: 3507: 3502: 3500:Oyster farming 3496: 3494: 3493:Related topics 3490: 3489: 3487: 3486: 3479: 3472: 3464: 3462: 3456: 3455: 3452: 3451: 3449: 3448: 3443: 3438: 3433: 3427: 3425: 3419: 3418: 3416: 3415: 3408: 3403: 3398: 3393: 3388: 3383: 3377: 3375: 3369: 3368: 3366: 3365: 3360: 3354: 3352: 3343: 3337: 3336: 3333: 3332: 3329: 3328: 3326: 3325: 3320: 3314: 3312: 3306: 3305: 3303: 3302: 3295: 3288: 3285:Cornu aspersum 3280: 3278: 3272: 3271: 3269: 3268: 3263: 3256: 3251: 3245: 3243: 3234: 3228: 3227: 3225: 3224: 3219: 3214: 3209: 3202: 3197: 3192: 3186: 3184: 3178: 3177: 3175: 3174: 3167: 3162: 3156: 3154: 3148: 3147: 3145: 3144: 3139: 3134: 3129: 3124: 3119: 3114: 3109: 3103: 3101: 3095: 3094: 3092: 3091: 3086: 3081: 3075: 3073: 3067: 3066: 3064: 3063: 3058: 3053: 3047: 3042: 3037: 3032: 3027: 3022: 3017: 3012: 3006: 3004: 2995: 2989: 2988: 2985: 2984: 2982: 2981: 2974: 2969: 2964: 2957: 2952: 2947: 2941: 2939: 2933: 2932: 2930: 2929: 2921: 2916: 2908: 2903: 2898: 2893: 2888: 2883: 2878: 2873: 2868: 2862: 2860: 2854: 2853: 2851: 2850: 2844: 2839: 2834: 2829: 2824: 2819: 2814: 2808: 2806: 2800: 2799: 2797: 2796: 2794:Giant Atlantic 2791: 2786: 2781: 2776: 2771: 2765: 2763: 2757: 2756: 2754: 2753: 2746: 2738: 2733: 2728: 2723: 2718: 2716:Triangle shell 2713: 2708: 2703: 2696: 2691: 2686: 2681: 2676: 2671: 2664: 2659: 2654: 2649: 2644: 2639: 2633: 2631: 2622: 2616: 2615: 2610: 2609: 2602: 2595: 2587: 2580: 2579:External links 2577: 2574: 2573: 2556: 2500: 2428: 2393:(4): 494–509. 2377: 2358:(2): 239–247. 2342: 2307:(3): 293–312. 2286: 2251:(2): 220–228. 2235: 2208:(1–2): 77–93. 2185: 2158:(3): 147–161. 2138: 2107: 2049: 2021: 1986:(3): 231–238. 1970: 1950: 1917: 1865: 1838:(1): 169–185. 1811: 1782: 1715: 1701: 1666:(6): 1054–64. 1635: 1590: 1589: 1587: 1584: 1538: 1535: 1481: 1478: 1457: 1454: 1395: 1392: 1275: 1272: 1256:chemoreceptors 1231:eyes lack any 1217: 1216:Sensory organs 1214: 1208:, and western 1160: 1157: 1141:American Samoa 1104:Additionally, 1098:American Samoa 1049: 1046: 1026:N. umbiculatus 985: 982: 979: 978: 972: 964: 963: 957: 951: 950: 944: 938: 937: 928: 920: 919: 911: 903: 902: 899: 891: 890: 887: 879: 878: 875: 866: 865: 855: 848: 846: 837: 830: 828: 819: 812: 810: 801: 794: 792: 783: 776: 774: 765: 758: 755: 754: 739: 724: 712: 701: 686: 675: 674: 673: 668: 656:Linnaeus, 1758 648: 637: 625: 613: 601: 589: 577: 574:Saunders, 1981 566: 554: 540: 537: 504:paleobiologist 472:of the family 457: 456:Classification 454: 438:Coral Triangle 332: 331: 330: 329: 322: 315: 308: 301: 294: 287: 277: 276: 272: 271: 262: 253: 252: 246: 245: 228: 224: 223: 218: 214: 213: 208: 204: 203: 198: 194: 193: 188: 184: 183: 178: 174: 173: 168: 164: 163: 158: 154: 153: 140: 139: 129: 128: 120: 119: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 49: 15: 9: 6: 4: 3: 2: 3812: 3801: 3798: 3796: 3793: 3792: 3790: 3773: 3768: 3764: 3760: 3755: 3751: 3747: 3741: 3737: 3732: 3728: 3724: 3719: 3715: 3711: 3706: 3702: 3698: 3693: 3689: 3685: 3680: 3676: 3672: 3667: 3663: 3659: 3654: 3650: 3646: 3641: 3637: 3633: 3628: 3624: 3620: 3615: 3611: 3607: 3602: 3598: 3594: 3589: 3585: 3580: 3574: 3570: 3565: 3559: 3555: 3554: 3552: 3550: 3546: 3542: 3537: 3527: 3522: 3516: 3513: 3511: 3508: 3506: 3503: 3501: 3498: 3497: 3495: 3491: 3485: 3484: 3480: 3478: 3477: 3473: 3471: 3470: 3466: 3465: 3463: 3461: 3457: 3447: 3444: 3442: 3439: 3437: 3434: 3432: 3429: 3428: 3426: 3424: 3420: 3414: 3413: 3409: 3407: 3404: 3402: 3399: 3397: 3396:Pacific giant 3394: 3392: 3389: 3387: 3384: 3382: 3379: 3378: 3376: 3374: 3370: 3364: 3361: 3359: 3356: 3355: 3353: 3351: 3347: 3344: 3342: 3338: 3324: 3321: 3319: 3316: 3315: 3313: 3311: 3307: 3301: 3300: 3299:Helix pomatia 3296: 3294: 3293: 3292:Helix lucorum 3289: 3287: 3286: 3282: 3281: 3279: 3277: 3273: 3267: 3264: 3262: 3261: 3257: 3255: 3252: 3250: 3247: 3246: 3244: 3242: 3238: 3235: 3233: 3229: 3223: 3220: 3218: 3215: 3213: 3210: 3208: 3207: 3203: 3201: 3198: 3196: 3193: 3191: 3188: 3187: 3185: 3183: 3179: 3173: 3172: 3168: 3166: 3163: 3161: 3158: 3157: 3155: 3153: 3149: 3143: 3140: 3138: 3135: 3133: 3130: 3128: 3125: 3123: 3120: 3118: 3115: 3113: 3110: 3108: 3105: 3104: 3102: 3100: 3096: 3090: 3087: 3085: 3082: 3080: 3077: 3076: 3074: 3072: 3068: 3062: 3059: 3057: 3054: 3051: 3048: 3046: 3043: 3041: 3038: 3036: 3033: 3031: 3028: 3026: 3023: 3021: 3018: 3016: 3013: 3011: 3008: 3007: 3005: 3003: 2999: 2996: 2994: 2990: 2980: 2979: 2975: 2973: 2970: 2968: 2965: 2963: 2962: 2958: 2956: 2953: 2951: 2948: 2946: 2943: 2942: 2940: 2938: 2934: 2927: 2926: 2922: 2920: 2917: 2915: 2913: 2912:Ostra chilena 2909: 2907: 2904: 2902: 2899: 2897: 2894: 2892: 2889: 2887: 2884: 2882: 2879: 2877: 2874: 2872: 2869: 2867: 2864: 2863: 2861: 2859: 2855: 2848: 2845: 2843: 2840: 2838: 2835: 2833: 2830: 2828: 2825: 2823: 2820: 2818: 2817:Mediterranean 2815: 2813: 2810: 2809: 2807: 2805: 2801: 2795: 2792: 2790: 2787: 2785: 2782: 2780: 2777: 2775: 2772: 2770: 2767: 2766: 2764: 2762: 2758: 2752: 2751: 2747: 2745:(razor genus) 2744: 2743: 2739: 2737: 2734: 2732: 2729: 2727: 2724: 2722: 2719: 2717: 2714: 2712: 2709: 2707: 2704: 2702: 2701: 2697: 2695: 2692: 2690: 2687: 2685: 2684:Pacific razor 2682: 2680: 2677: 2675: 2672: 2670: 2669: 2665: 2663: 2660: 2658: 2655: 2653: 2650: 2648: 2645: 2643: 2642:Atlantic surf 2640: 2638: 2635: 2634: 2632: 2630: 2626: 2623: 2621: 2617: 2608: 2603: 2601: 2596: 2594: 2589: 2588: 2585: 2569: 2568: 2560: 2552: 2548: 2544: 2540: 2536: 2532: 2528: 2524: 2520: 2516: 2509: 2507: 2505: 2496: 2492: 2487: 2482: 2478: 2474: 2469: 2464: 2460: 2456: 2453:(2): e16311. 2452: 2448: 2444: 2437: 2435: 2433: 2424: 2420: 2416: 2412: 2408: 2404: 2400: 2396: 2392: 2388: 2381: 2373: 2369: 2365: 2361: 2357: 2353: 2346: 2338: 2334: 2330: 2326: 2322: 2318: 2314: 2310: 2306: 2302: 2295: 2293: 2291: 2282: 2278: 2274: 2270: 2266: 2262: 2258: 2254: 2250: 2246: 2239: 2231: 2227: 2223: 2219: 2215: 2211: 2207: 2203: 2196: 2194: 2192: 2190: 2181: 2177: 2173: 2169: 2165: 2161: 2157: 2153: 2149: 2142: 2128: 2124: 2118: 2116: 2114: 2112: 2103: 2099: 2095: 2091: 2087: 2083: 2079: 2075: 2071: 2067: 2060: 2058: 2056: 2054: 2046: 2042: 2040: 2032: 2030: 2028: 2026: 2017: 2013: 2009: 2005: 2001: 1997: 1993: 1989: 1985: 1981: 1974: 1966: 1960: 1953: 1947: 1943: 1939: 1935: 1928: 1926: 1924: 1922: 1913: 1909: 1904: 1899: 1895: 1891: 1887: 1885: 1881: 1872: 1870: 1861: 1857: 1853: 1849: 1845: 1841: 1837: 1833: 1826: 1824: 1822: 1820: 1818: 1816: 1807: 1801: 1793: 1789: 1785: 1779: 1775: 1768: 1766: 1764: 1762: 1760: 1758: 1756: 1754: 1752: 1750: 1748: 1746: 1744: 1742: 1740: 1738: 1736: 1734: 1732: 1730: 1728: 1726: 1724: 1722: 1720: 1711: 1705: 1697: 1693: 1689: 1685: 1681: 1677: 1673: 1669: 1665: 1661: 1660: 1655: 1653: 1644: 1642: 1640: 1631: 1627: 1623: 1619: 1615: 1611: 1607: 1600: 1598: 1596: 1591: 1583: 1581: 1577: 1573: 1569: 1565: 1561: 1558: 1554: 1552: 1548: 1544: 1534: 1531: 1527: 1522: 1520: 1516: 1511: 1507: 1503: 1499: 1491: 1486: 1477: 1475: 1471: 1467: 1463: 1453: 1451: 1446: 1442: 1438: 1434: 1430: 1425: 1423: 1418: 1414: 1409: 1405: 1400: 1391: 1389: 1385: 1380: 1375: 1371: 1367: 1363: 1359: 1355: 1351: 1347: 1343: 1339: 1335: 1331: 1328:, Australia, 1327: 1323: 1315: 1310: 1306: 1304: 1301: 1297: 1293: 1289: 1285: 1281: 1271: 1269: 1265: 1261: 1257: 1253: 1249: 1246:also feature 1245: 1240: 1238: 1234: 1230: 1226: 1222: 1213: 1211: 1207: 1203: 1199: 1195: 1191: 1187: 1183: 1174: 1171:displayed at 1170: 1165: 1156: 1154: 1150: 1146: 1145:N. belauensis 1142: 1138: 1134: 1130: 1126: 1121: 1119: 1115: 1111: 1107: 1099: 1081: 1079: 1075: 1071: 1067: 1066:N. belauensis 1063: 1059: 1055: 1045: 1043: 1039: 1035: 1034:N. texturatus 1031: 1030:N. perforatus 1027: 1023: 1019: 1015: 1014:N. belauensis 1011: 1007: 1003: 999: 995: 991: 977: 973: 971: 970: 966: 965: 962: 958: 956: 955:N. texturatus 953: 952: 949: 945: 943: 942:N. perforatus 940: 939: 936: 932: 929: 927: 926: 922: 921: 918: 917: 912: 910: 909: 905: 904: 901:Nomen Dubium 900: 898: 897: 893: 892: 889:Nomen Dubium 888: 886: 885: 881: 880: 876: 873: 872: 869: 861: 860: 852: 847: 844: 843: 834: 829: 826: 825: 816: 811: 808: 807: 798: 793: 790: 789: 780: 775: 772: 771: 769:N. belauensis 762: 757: 756: 750: 745: 744: 740: 735: 730: 729: 725: 719: 718: 713: 707: 706: 702: 697: 692: 691: 687: 682: 681: 676: 672: 669: 667: 664: 663: 661: 658:– Indonesia ( 654: 653: 649: 643: 642: 638: 632: 631: 626: 622:(Foord, 1891) 620: 619: 614: 608: 607: 602: 596: 595: 590: 584: 583: 578: 572: 571: 567: 561: 560: 555: 549: 548: 543: 542: 536: 534: 530: 525: 523: 519: 515: 511: 510: 505: 501: 497: 496:Carl Linnaeus 494:inception by 493: 489: 488: 483: 479: 475: 471: 467: 463: 453: 451: 447: 443: 439: 435: 429: 427: 423: 419: 415: 411: 407: 403: 398: 396: 392: 388: 387: 382: 378: 374: 373: 369: 365: 361: 357: 354: 351: 347: 344: 340: 339: 328: 327: 323: 321: 320: 316: 314: 313: 309: 307: 306: 302: 300: 299: 295: 293: 292: 288: 286: 285: 281: 280: 278: 273: 269: 265: 260: 259: 254: 251: 247: 243: 239: 234: 233: 229: 226: 225: 222: 219: 216: 215: 212: 209: 206: 205: 202: 199: 196: 195: 192: 189: 186: 185: 182: 179: 176: 175: 172: 169: 166: 165: 162: 159: 156: 155: 150: 145: 141: 137: 136: 130: 126: 121: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 53:Eocene–Recent 47: 43: 40: 37: 30: 26: 22: 3548: 3481: 3474: 3467: 3410: 3401:Southern red 3297: 3290: 3283: 3258: 3232:Other snails 3204: 3169: 2976: 2959: 2945:Atlantic bay 2923: 2911: 2876:Southern mud 2748: 2740: 2698: 2679:Ocean quahog 2666: 2566: 2559: 2521:(1): 40–51. 2518: 2515:Paleobiology 2514: 2450: 2446: 2390: 2387:Paleobiology 2386: 2380: 2355: 2351: 2345: 2304: 2300: 2248: 2245:Paleobiology 2244: 2238: 2205: 2201: 2155: 2151: 2141: 2130:. Retrieved 2126: 2069: 2065: 2044: 2038: 1983: 1979: 1973: 1933: 1893: 1889: 1883: 1879: 1835: 1831: 1776:. Springer. 1773: 1704: 1663: 1657: 1652:Allonautilus 1651: 1616:(1): 89–95. 1613: 1609: 1605: 1571: 1563: 1562: 1556: 1555: 1550: 1546: 1542: 1540: 1537:Reproduction 1529: 1525: 1523: 1514: 1509: 1505: 1501: 1497: 1495: 1489: 1473: 1469: 1465: 1461: 1459: 1449: 1444: 1440: 1436: 1432: 1428: 1426: 1421: 1416: 1407: 1398: 1397: 1378: 1373: 1369: 1365: 1362:South Africa 1321: 1319: 1313: 1302: 1299: 1295: 1291: 1287: 1283: 1279: 1277: 1263: 1243: 1241: 1228: 1224: 1220: 1219: 1193: 1186:Allonautilus 1185: 1181: 1178: 1153:N. pompilius 1152: 1148: 1144: 1128: 1124: 1122: 1117: 1113: 1109: 1103: 1077: 1073: 1070:N. pompilius 1069: 1065: 1061: 1057: 1053: 1051: 1041: 1038:N. pompilius 1037: 1033: 1029: 1025: 1021: 1017: 1013: 1009: 1005: 1002:N. pompilius 1001: 997: 994:nomen dubium 993: 987: 976:N. pompilius 975: 967: 960: 954: 947: 941: 934: 923: 916:Allonautilus 914: 906: 894: 882: 867: 857: 838: 820: 805:N. pompilius 802: 784: 766: 748: 741: 733: 726: 715: 703: 695: 688: 678: 670: 665: 650: 639: 634:Martin, 1879 628: 616: 604: 592: 586:Martin, 1933 580: 568: 557: 545: 532: 526: 521: 514:type species 509:N. pompilius 507: 485: 465: 461: 459: 449: 448:. The genus 434:Indo-Pacific 430: 422:Allonautilus 421: 417: 413: 409: 405: 399: 384: 372:Allonautilus 370: 341:is a marine 337: 336: 335: 324: 317: 310: 303: 296: 289: 282: 258:N. pompilius 256: 250:Type species 231: 230: 133: 45: 39: 24: 20: 3653:iNaturalist 3573:Wikispecies 3446:Neon flying 3152:Periwinkles 3112:Turtle/talc 3045:Green ormer 2955:New Zealand 2925:Crassostrea 2906:Sydney rock 2784:New Zealand 2706:Smooth clam 2674:Blunt gaper 1404:crustaceans 1248:rhinophores 1198:Philippines 1149:N. repertus 969:N. repertus 959:Synonym of 946:Synonym of 884:N. ambiguus 859:N. taiwanus 721:Huang, 2002 636:- Indonesia 624:- Australia 588:- Indonesia 565:- Australia 553:- Australia 436:, from the 395:Philippines 201:Nautiloidea 191:Cephalopoda 3795:Nautiluses 3789:Categories 3363:Bottletail 3350:Cuttlefish 3310:Freshwater 3254:Korean mud 3217:Pear whelk 2993:Gastropods 2950:Great/king 2896:Windowpane 2891:Portuguese 2827:California 2731:Razor clam 2711:Soft-shell 2132:2020-05-12 1890:Island Arc 1586:References 1330:Seychelles 896:N. alumnus 474:Nautilidae 470:type genus 414:Nautilidae 406:Nautilidae 381:Kazakhstan 356:Nautilidae 343:cephalopod 221:Nautilidae 197:Subclass: 34:See also: 25:Nautilidae 3358:Spineless 3222:Lightning 3190:Channeled 2847:Mytilidae 2736:Pod razor 2657:Hard clam 2543:0094-8373 2477:1932-6203 2415:0094-8373 2372:0165-7836 2321:0302-766X 2273:0094-8373 2222:0740-2783 2172:0898-1051 2086:0962-1083 2043: », 2000:0014-3820 1980:Evolution 1912:247532223 1878:"Miocene 1852:1664-2376 1800:cite book 1792:613896438 1630:129517075 1456:Predation 1394:Behaviour 1354:Sri Lanka 1334:Mauritius 1303:Pompilius 1260:tentacles 1210:Australia 1206:Indonesia 1159:Evolution 738:– Vanuatu 487:Ammonites 428:(CITES). 211:Nautilida 167:Kingdom: 161:Eukaryota 3593:Nautilus 3579:Nautilus 3564:Q1324463 3558:Wikidata 3549:Nautilus 3526:Category 3441:Humboldt 3391:Big blue 3249:Mud-flat 3200:Kellet's 3040:Greenlip 3035:Blacklip 2937:Scallops 2620:Bivalves 2551:89164986 2495:21364981 2447:PLOS ONE 2423:89107860 2337:19565604 2281:85659002 2230:86014620 2094:28872211 2039:Nautilus 2016:18138384 1959:citation 1884:Nautilus 1880:Nautilus 1860:87025055 1696:87319305 1572:Nautilus 1564:Nautilus 1557:Nautilus 1551:Nautilus 1547:Nautilus 1543:Nautilus 1530:Nautilus 1526:Nautilus 1515:Nautilus 1510:Nautilus 1506:Nautilus 1502:Nautilus 1498:Nautilus 1474:Nautilus 1470:Nautilus 1466:Nautilus 1462:Nautilus 1450:Nautilus 1445:Nautilus 1441:Nautilus 1437:Nautilus 1433:Nautilus 1429:Nautilus 1422:Nautilus 1417:Nautilus 1408:Nautilus 1399:Nautilus 1379:Nautilus 1374:Nautilus 1370:Nautilus 1366:Nautilus 1346:Thailand 1322:Nautilus 1314:Nautilus 1300:Nautilus 1296:Nautilus 1292:Nautilus 1288:Nautilus 1284:Nautilus 1280:Nautilus 1264:Nautilus 1244:Nautilus 1229:Nautilus 1225:Nautilus 1221:Nautilus 1194:Nautilus 1182:Nautilus 1125:Nautilus 1110:Nautilus 1054:Nautilus 998:Nautilus 533:Nautilus 522:Nautilus 466:Nautilus 462:Nautilus 450:Nautilus 418:Nautilus 402:nautilus 338:Nautilus 264:Linnaeus 238:Linnaeus 232:Nautilus 217:Family: 181:Mollusca 177:Phylum: 171:Animalia 157:Domain: 46:Nautilus 29:Nautilus 21:Nautilus 3767:ZooBank 3746:8329633 3671:1286821 3645:2289060 3460:Chitons 3373:Octopus 3341:Inkfish 3323:Nerites 3212:Knobbed 3099:Limpets 3084:Elegant 3071:Conches 3061:Chilean 3002:Abalone 2886:Pacific 2871:Olympia 2866:Eastern 2858:Oysters 2804:Mussels 2761:Cockles 2750:Paphies 2647:Geoduck 2523:Bibcode 2486:3043052 2455:Bibcode 2395:Bibcode 2329:4903946 2253:Bibcode 2180:1535418 2102:2153267 2045:ZooKeys 2008:2405560 1688:1306604 1668:Bibcode 1262:of the 1133:Vanuatu 931:Synonym 877:Status 747:Barord 732:Barord 700:– Samoa 694:Barord 576:– Palau 539:Species 502:, when 482:sutures 468:is the 393:in the 360:Species 350:mollusk 348:in the 227:Genus: 207:Order: 187:Class: 132:A live 3759:204549 3743:uBio: 3710:616358 3684:555825 3658:123387 3381:Common 3266:Bailer 3195:Common 3182:Whelks 3165:Banded 3160:Common 3142:Rustic 2914:/Bluff 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Index

Nautilus
Chambered nautilus
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Nautilus pompilius
Scientific classification
Edit this classification
Eukaryota
Animalia
Mollusca
Cephalopoda
Nautiloidea
Nautilida
Nautilidae
Nautilus
Linnaeus
1758
Type species
N. pompilius

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