31:
52:
2544:
2501:
2069:, suggest an origin for this group in the Permian or later. In contrast, total-evidence dating paints a more ancient picture, tracing the complex thalloid liverworts back to the Silurian-Devonian boundary, highlighting a narrative of morphological stability across epochs. Thus, the complex thalloid liverworts emerge as significant players in the ongoing saga of plant evolution, their history intertwined with the deep complexities of geological time.
1067:(Townrow) H. M. Anderson (Middle Triassic), for instance, is a Marchantia-like fossil whose detailed morphological characters (e.g., thallus with midrib, reduced air chambers, rhizoids and ventral scales) suggest a nested position within Marchantiales. Some Riccia-like fossils have even been assigned to families based on their overall morphology and branching patterns, such as the case of
1071:
Li & Sun (Middle
Jurassic). The first phylogenetic analyses that include both extinct and extant Marchantiales have further clarified the relationships among these taxa and have revealed new relationships among families. Likewise, the inclusion of fossils in total-evidence analyses implied that
1060:
Extinct
Marchantiales - which commonly date back to the Mesozoic - can be grouped in Marchantia-like and Riccia-like fossils according to their overall morphology. While the phylogenetic relationships among many extinct and extant Marchantiales remain equivocal, it has been suggested that some
1056:
Extinct complex thalloid liverworts are often represented by coalified compressions that preserve superficial morphological traits and do not allow exhaustively analysing their fine anatomy; though, in exceptional cases, fossils might preserve cell details.
2061:
in the Early
Permian of Russia offers the first clear evidence of the Marchantiales in the Paleozoic. However, rosette-shaped fossils that resemble Ricciaceae are as old as the Early Devonian – suggesting a much older origin for the group.
2746:"Defying death: incorporating fossils into the phylogeny of the complex thalloid liverworts (Marchantiidae, Marchantiophyta) confirms high order clades but reveals discrepancies in family-level relationships"
2050:, spanning Carboniferous strata of various regions, presents a puzzling case as its affiliation with the complex thalloid liverworts dependen on its individual species. Similarly, the Carboniferous
2793:"Dating the evolution of the complex thalloid liverworts (Marchantiopsida): total-evidence dating analysis supports a Late Silurian-Early Devonian origin and post-Mesozoic morphological stasis"
198:
are more numerous. The majority of genera are characterized by the presence of (a) special stalked vertical branches called archegoniophores or carpocephala, and (b) sterile cells called
1157:
1483:
1437:
1240:
2710:"Ricciopsis sandaolingensis sp. nov., a new fossil bryophyte from the Middle Jurassic Xishanyao Formation in the Turpan-Hami Basin, Xinjiang, Northwest China"
3817:
3538:
2791:
Flores, Jorge R.; Bippus, Alexander C.; de
Ullivarri, Carmen Fernández; Suárez, Guillermo M.; Hyvönen, Jaakko; Tomescu, Alexandru M. F. (December 2023).
4014:
3847:
3908:
3898:
3094:
3994:
3765:
3415:
2265:
1534:
251:
4186:
3999:
3979:
3410:
3362:
2245:
1855:
854:
4009:
3308:
1203:
4225:
3299:
2593:"Divergence times and the evolution of morphological complexity in an early land plant lineage (Marchantiopsida) with a slow molecular rate"
2274:
2254:
2174:
1812:
861:
499:
258:
2154:
532:
2234:
2224:
1175:
929:
888:
2043:
Liverworts, possess a compelling narrative of evolutionary intricacies, particularly within the realm of complex thalloid liverworts.
3136:
4160:
3179:
2500:
4199:
3004:"Combined phylogenetic analysis of the subclass Marchantiidae (Marchantiophyta): towards a robustly diagnosed classification"
2656:
4204:
3099:
3104:
3084:
Grolle, Riclef (1983). "Nomina generica
Hepaticarum; references, types and synonymies". Acta Botanica Fennica 121, 1-62.
3065:
Crandall-Stotler, Barbara J. & Stotler, Raymond E. "Morphology and classification of the
Marchantiophyta". page 63
3078:
2885:
2551:
4230:
2054:
raises questions about its relationship with the
Blasiales, the sister group to Marchantiales. The appearance of
4276:
2709:
2081:. The order Lunulariales, proposed by Long 2006, has been recently re-included in Marchantiales as a family.
4212:
3158:
3129:
3184:
3174:
51:
4113:
4108:
4095:
4289:
4343:
4240:
2988:
2636:
3169:
3122:
2637:"Why Are Bryophytes So Rare in the Fossil Record? A Spotlight on Taphonomy and Fossil Preservation"
4086:
2679:"A review of the Bryophyta from the Upper Triassic Molteno Formation, Karroo Basin, South Africa"
2314:
912:
4338:
4266:
4048:
2635:
Tomescu, Alexandru M.F.; Bomfleur, Benjamin; Bippus, Alexander C.; Savoretti, Adolfina (2018),
4261:
2975:
2966:
171:
145:
4253:
4191:
3003:
2792:
4134:
3949:
2124:
1086:
381:
8:
4019:
3857:
30:
4310:
3739:
3164:
3049:
2862:
2839:
2773:
2648:
2513:
151:
46:
4248:
4121:
4004:
3944:
3893:
3883:
3513:
3463:
3074:
3041:
3033:
2938:
2867:
2820:
2812:
2777:
2765:
2745:
2652:
2614:
2464:
1134:
778:
3114:
3053:
3002:
Flores, Jorge R.; Catalano, Santiago A.; Muñoz, Jesus; Suárez, Guillermo M. (2018).
2917:"New Higher Taxa of Complex Thalloid Liverworts (Marchantiophyta – Marchantiopsida)"
1072:
some groups of complex thalloid liverworts might be older than previously inferred.
4315:
3989:
3783:
3775:
3729:
3596:
3591:
3543:
3533:
3372:
3326:
3194:
3023:
3015:
2961:
2928:
2857:
2847:
2804:
2757:
2744:
Flores, Jorge R; Bippus, Alexander C; Suárez, Guillermo M; Hyvönen, Jaakko (2020).
2721:
2690:
2644:
2604:
1113:
155:, a common and often troublesome weed in moist, temperate gardens and greenhouses.
4100:
4126:
3969:
3959:
3862:
3852:
3749:
3691:
3686:
3664:
3566:
3555:
3523:
3502:
3492:
3445:
3430:
3400:
3354:
3318:
3243:
3211:
3146:
2475:
2385:
2285:
2195:
1713:
1651:
1614:
1390:
1361:
1322:
1291:
1099:
827:
791:
630:
140:
136:
83:
73:
4217:
3954:
3888:
3842:
3420:
3203:
2521:
2345:
2294:
1468:
451:
224:
2933:
2916:
2852:
4332:
4071:
3984:
3974:
3964:
3929:
3878:
3837:
3832:
3701:
3650:
3627:
3586:
3440:
3344:
3289:
3275:
3037:
2957:
2942:
2816:
2455:
2325:
2104:
1636:
1420:
1348:
771:
292:
93:
2958:"Supplemental Information 1: Bryophyte Phylogeny Poster 2019 - full A0 size"
4302:
3939:
3934:
3913:
3903:
3827:
3734:
3711:
3637:
3619:
3528:
3425:
3405:
3336:
3045:
2871:
2824:
2769:
2618:
2444:
2405:
2394:
2365:
2215:
2204:
1870:
1777:
1751:
1698:
1258:
1216:
874:
834:
700:
666:
637:
610:
186:
is often used to typify the order, although there are also many species of
2893:
2543:
4173:
4080:
3822:
3721:
3681:
3576:
3390:
3229:
2484:
2304:
2085:
1556:
946:
798:
278:
163:
143:(also known as "complex thalloid liverworts") that includes species like
3028:
3706:
3696:
3673:
3645:
3487:
3435:
3395:
3367:
3269:
3221:
2956:
Cole, Theodor C H; Hilger, Hartmut H; Goffinet, Bernard (24 May 2019).
2694:
2528:
2425:
2354:
2145:
1763:
1585:
1502:
686:
437:
231:
182:
167:
4178:
3019:
2808:
2761:
2609:
2592:
4147:
3801:
3793:
3479:
3261:
3252:
2334:
2094:
1336:
340:
188:
159:
4042:
2678:
4297:
4165:
4065:
3109:
2414:
2374:
2184:
2134:
2077:
Taxonomy based on work by Söderström et al. 2016 and synonyms from
673:
617:
549:
364:
4152:
2726:
2535:
2419:
Bischoff ex
Lindenberg 1829 non (Blume) Hooker & Thomson 1855
2708:
Li, Ruiyun; Li, Xiaoqiang; Wang, Hongshan; Sun, Bainian (2019).
2434:
2164:
2114:
1794:
1051:
717:
482:
316:
199:
194:
2790:
2634:
4284:
4139:
178:
63:
2890:
Collection of genus-group names in a systematic arrangement
2079:
Collection of genus-group names in a systematic arrangement
3001:
2743:
2129:
Lehmann & Lindenberg 1832 nom. cons. non Pomel 1883
2065:
Molecular analyses, calibrated with the
Triassic fossil
2573:
Limpricht, G. (1877). "Lebermoose". In Cohn, F. (ed.).
1075:
Summary tree based on the work by Flores et al. 2020:
3144:
2319:
1061:
fossils are closely related to extant Marchantiales.
149:, a widespread plant often found beside rivers, and
205:
2955:
4330:
3073:. (Cambridge: Cambridge University Press:2000).
3069:A. Jonathan Shaw & Bernard Goffinet (Eds.),
16:Order of non-vascular plants known as liverworts
2886:"Part 2- Plantae (starting with Chlorophycota)"
2554:Cross section through a marchantialian thallus.
3130:
2840:"World checklist of hornworts and liverworts"
1052:Phylogeny (extant and extinct Marchantiales)
210:Based on the work by Villarreal et al. 2015
2707:
3137:
3123:
2837:
2590:
2542:
29:
3027:
2965:
2932:
2861:
2851:
2725:
2608:
2572:
3260:
2676:
2038:
4331:
2967:10.7287/peerj.preprints.27571v3/supp-1
2526:
170:existing as a short-lived part of the
4047:
4046:
3118:
2533:
2519:
4290:E648C0AC-4086-B022-8C88-5CCC7B208D98
4241:d61dab13-7429-4d9c-9215-4a6d2e0c775d
2914:
2739:
2737:
2630:
2628:
2586:
2584:
2511:
2506:
2249:Stotler & Crandall-Stotler 2000
122:
13:
2649:10.1016/b978-0-12-813012-4.00016-4
174:, dependent upon the gametophyte.
14:
4355:
3088:
2734:
2625:
2581:
2072:
166:generation is dominant, with the
2838:Söderström; et al. (2016).
2591:Villarreal; et al. (2015).
2577:. Vol. 1. pp. 225–352.
2499:
2469:Linnaeus 1753 non non Hesse 1923
206:Phylogeny (extant Marchantiales)
50:
2995:
2949:
2908:
2575:Kryptogamen-Flora von Schlesien
1158:Pallaviciniites sandaolingensis
2878:
2831:
2784:
2701:
2670:
2643:, Elsevier, pp. 375–416,
2566:
1:
2559:
3105:Additional photos of species
7:
2921:Edinburgh Journal of Botany
10:
4360:
2714:Palaeontologia Electronica
2641:Transformative Paleobotany
2099:Palisot De Beauvisage 1805
1069:Ricciopsis sandaolingensis
4055:
3922:
3871:
3810:
3792:
3774:
3757:
3748:
3720:
3672:
3663:
3636:
3618:
3611:
3554:
3501:
3478:
3471:
3462:
3381:
3353:
3335:
3317:
3297:
3288:
3251:
3242:
3220:
3202:
3193:
3153:
3100:Photos of various species
2934:10.1017/S0960428606000606
2915:Long, D. G. (July 2006).
2853:10.3897/phytokeys.59.6261
2067:Marchantites cyathodoides
1868:
1853:
1846:
1808:
1791:
1774:
1767:
1749:
1742:
1735:
1711:
1696:
1689:
1682:
1649:
1634:
1627:
1612:
1605:
1598:
1583:
1576:
1569:
1554:
1547:
1532:
1525:
1499:
1484:Marchantites huolinhensis
1479:
1472:
1462:
1438:Marchantites cyathodoides
1433:
1418:
1411:
1404:
1388:
1381:
1374:
1359:
1352:
1334:
1327:
1289:
1256:
1241:Metzgeriothallus sharonae
1236:
1229:
1214:
1207:
1197:
1173:
1153:
1146:
1139:
1127:
1110:
1103:
1083:
1065:Marchantites cyathodoides
943:
926:
909:
902:
885:
878:
858:
848:
831:
821:
795:
775:
765:
758:
714:
697:
690:
670:
660:
634:
614:
604:
597:
590:
546:
529:
522:
496:
479:
472:
465:
448:
441:
431:
378:
361:
354:
337:
330:
313:
306:
289:
282:
272:
255:
245:
228:
218:
192:and species of the genus
118:
113:
47:Scientific classification
45:
37:
28:
23:
2677:Anderson, Heidi (1976).
2109:Austin ex Underwood 1884
2697:– via WireDSpace.
2683:Palaeontologia Africana
202:inside the sporangium.
41:- a thallose liverwort
2983:Cite journal requires
4262:Paleobiology Database
2409:MĂĽller ex Grolle 1972
2289:MĂĽller ex Grolle 1972
2259:Kunze ex Lehmann 1834
2199:MĂĽller ex Grolle 1972
2139:Raddi 1818 nom. cons.
2119:Corda 1829 nom. cons.
146:Marchantia polymorpha
3950:Blepharidophyllaceae
2209:Hill 1773 nom. cons.
2039:Origin and evolution
1087:Takakia ceratophylla
39:Conocephalum conicum
4020:Trichotemnomataceae
3858:Pseudolepicoleaceae
3818:Blepharostomataceae
3740:Neotrichocoleaceae
3539:Pseudomoerckiaceae
3510:Phyllothalliineae
3145:Classification of
3110:Morphobank project
152:Lunularia cruciata
4326:
4325:
4249:Open Tree of Life
4049:Taxon identifiers
4040:
4039:
4036:
4035:
4032:
4031:
4028:
4027:
4005:Solenostomataceae
3945:Balantiopsidaceae
3894:Cephaloziellaceae
3884:Anastrophyllaceae
3659:
3658:
3607:
3606:
3583:Fossombroniineae
3520:Pallaviciniineae
3514:Phyllothalliaceae
3464:Jungermanniopsida
3458:
3457:
3454:
3453:
3284:
3283:
3238:
3237:
3195:Haplomitriopsida
3071:Bryophyte Biology
3020:10.1111/cla.12225
2896:on 6 October 2016
2809:10.1111/nph.19254
2762:10.1111/cla.12442
2658:978-0-12-813012-4
2610:10.1111/nph.13716
2490:
2480:
2470:
2460:
2450:
2440:
2430:
2420:
2410:
2400:
2390:
2380:
2370:
2360:
2350:
2340:
2330:
2320:
2310:
2300:
2290:
2280:
2270:
2260:
2250:
2240:
2230:
2220:
2210:
2200:
2190:
2180:
2170:
2160:
2150:
2140:
2130:
2120:
2110:
2100:
2090:
2052:Blasiites lobatus
2035:
2034:
2025:
2024:
2016:
2015:
2007:
2006:
1998:
1997:
1989:
1988:
1980:
1979:
1971:
1970:
1962:
1961:
1953:
1952:
1944:
1943:
1935:
1934:
1926:
1925:
1917:
1916:
1908:
1907:
1899:
1898:
1890:
1889:
1881:
1880:
1835:
1834:
1826:
1825:
1724:
1723:
1671:
1670:
1662:
1661:
1514:
1513:
1451:
1450:
1311:
1310:
1302:
1301:
1278:
1277:
1269:
1268:
1186:
1185:
1135:Jungermanniopsida
1048:
1047:
1039:
1038:
1030:
1029:
1021:
1020:
1012:
1011:
1003:
1002:
994:
993:
985:
984:
976:
975:
967:
966:
958:
957:
810:
809:
747:
746:
738:
737:
729:
728:
649:
648:
579:
578:
570:
569:
561:
560:
511:
510:
420:
419:
411:
410:
402:
401:
393:
392:
130:
129:
109:
4351:
4344:Liverwort orders
4319:
4318:
4306:
4305:
4293:
4292:
4280:
4279:
4270:
4269:
4257:
4256:
4244:
4243:
4234:
4233:
4221:
4220:
4218:NHMSYS0000066340
4208:
4207:
4195:
4194:
4182:
4181:
4169:
4168:
4156:
4155:
4143:
4142:
4130:
4129:
4117:
4116:
4104:
4103:
4091:
4090:
4089:
4076:
4075:
4074:
4044:
4043:
4015:Stephaniellaceae
3990:Jungermanniaceae
3923:Jungermanniineae
3848:Mastigophoraceae
3784:Schistochilaceae
3776:Perssoniellineae
3755:
3754:
3730:Herzogianthaceae
3670:
3669:
3616:
3615:
3597:Petalophyllaceae
3592:Fossombroniaceae
3563:Calyculariineae
3544:Sandeothallaceae
3534:Pallaviciniaceae
3476:
3475:
3469:
3468:
3373:Sphaerocarpaceae
3327:Neohodgsoniaceae
3295:
3294:
3258:
3257:
3249:
3248:
3200:
3199:
3180:Acrogymnospermae
3139:
3132:
3125:
3116:
3115:
3058:
3057:
3031:
2999:
2993:
2992:
2986:
2981:
2979:
2971:
2969:
2953:
2947:
2946:
2936:
2927:(2–3): 257–262.
2912:
2906:
2905:
2903:
2901:
2892:. Archived from
2882:
2876:
2875:
2865:
2855:
2835:
2829:
2828:
2803:(5): 2137–2150.
2788:
2782:
2781:
2741:
2732:
2731:
2729:
2705:
2699:
2698:
2674:
2668:
2667:
2666:
2665:
2632:
2623:
2622:
2612:
2588:
2579:
2578:
2570:
2546:
2538:
2531:
2524:
2517:
2509:
2503:
2488:
2478:
2468:
2458:
2448:
2438:
2429:Reichenbach 1828
2428:
2418:
2408:
2398:
2388:
2378:
2368:
2358:
2348:
2338:
2328:
2318:
2308:
2298:
2288:
2278:
2268:
2258:
2248:
2238:
2228:
2218:
2208:
2198:
2188:
2178:
2168:
2158:
2148:
2138:
2128:
2118:
2108:
2098:
2088:
2029:
2028:
1849:
1848:
1770:
1769:
1745:
1744:
1738:
1737:
1692:
1691:
1685:
1684:
1630:
1629:
1608:
1607:
1601:
1600:
1579:
1578:
1572:
1571:
1550:
1549:
1528:
1527:
1475:
1474:
1465:
1464:
1414:
1413:
1407:
1406:
1384:
1383:
1377:
1376:
1355:
1354:
1330:
1329:
1232:
1231:
1210:
1209:
1200:
1199:
1149:
1148:
1142:
1141:
1130:
1129:
1114:Haplomitriopsida
1106:
1105:
1079:
1078:
905:
904:
881:
880:
851:
850:
824:
823:
768:
767:
761:
760:
693:
692:
663:
662:
607:
606:
600:
599:
593:
592:
525:
524:
475:
474:
468:
467:
444:
443:
434:
433:
357:
356:
333:
332:
309:
308:
285:
284:
275:
274:
248:
247:
221:
220:
214:
213:
107:
55:
54:
33:
21:
20:
4359:
4358:
4354:
4353:
4352:
4350:
4349:
4348:
4329:
4328:
4327:
4322:
4314:
4309:
4301:
4296:
4288:
4283:
4275:
4273:
4265:
4260:
4252:
4247:
4239:
4237:
4229:
4224:
4216:
4211:
4203:
4198:
4190:
4185:
4177:
4172:
4164:
4159:
4151:
4146:
4138:
4133:
4125:
4120:
4112:
4107:
4099:
4094:
4085:
4084:
4079:
4070:
4069:
4064:
4051:
4041:
4024:
3970:Gymnomitriaceae
3960:Endogemmataceae
3918:
3909:Obtusifoliaceae
3899:Hygrobiellaceae
3867:
3863:Trichocoleaceae
3853:Plagiochilaceae
3806:
3788:
3770:
3750:Jungermanniales
3744:
3716:
3692:Lepidolaenaceae
3687:Goebeliellaceae
3665:Jungermanniidae
3655:
3632:
3603:
3567:Calyculariaceae
3556:Fossombroniales
3550:
3524:Hymenophytaceae
3503:Pallaviciniales
3497:
3493:Noterocladaceae
3450:
3446:Wiesnerellaceae
3431:Monosoleniaceae
3401:Conocephalaceae
3377:
3355:Sphaerocarpales
3349:
3331:
3319:Neohodgsoniales
3313:
3280:
3244:Marchantiopsida
3234:
3216:
3212:Haplomitriaceae
3189:
3170:Marchantiophyta
3149:
3147:Marchantiophyta
3143:
3091:
3062:
3061:
3000:
2996:
2984:
2982:
2973:
2972:
2954:
2950:
2913:
2909:
2899:
2897:
2884:
2883:
2879:
2836:
2832:
2797:New Phytologist
2789:
2785:
2742:
2735:
2706:
2702:
2675:
2671:
2663:
2661:
2659:
2633:
2626:
2597:New Phytologist
2589:
2582:
2571:
2567:
2562:
2557:
2556:
2555:
2553:
2548:
2547:
2540:
2539:
2534:
2532:
2527:
2525:
2520:
2518:
2512:
2510:
2507:
2504:
2476:Wiesnerellaceae
2386:Monosoleniaceae
2329:Klingrräff 1858
2315:Stephensoniella
2286:Exormothecaceae
2196:Conocephalaceae
2075:
2041:
2036:
2026:
2017:
2008:
1999:
1990:
1981:
1972:
1963:
1954:
1945:
1936:
1927:
1918:
1909:
1900:
1891:
1882:
1836:
1827:
1725:
1714:Conocephalaceae
1672:
1663:
1652:Monosoleniaceae
1615:Wiesnerellaceae
1515:
1452:
1391:Sphaerocarpales
1362:Neohodgsoniales
1323:Marchantiopsida
1312:
1303:
1292:Jungermanniidae
1279:
1270:
1187:
1100:Marchantiophyta
1054:
1049:
1040:
1031:
1022:
1013:
1004:
995:
986:
977:
968:
959:
913:Stephensoniella
828:Monosoleniaceae
811:
792:Wiesnerellaceae
748:
739:
730:
650:
631:Conocephalaceae
580:
571:
562:
512:
421:
412:
403:
394:
208:
106:
84:Marchantiopsida
74:Marchantiophyta
49:
17:
12:
11:
5:
4357:
4347:
4346:
4341:
4324:
4323:
4321:
4320:
4316:wfo-9000000315
4307:
4294:
4281:
4271:
4258:
4245:
4235:
4222:
4209:
4196:
4183:
4170:
4157:
4144:
4131:
4118:
4105:
4092:
4077:
4061:
4059:
4053:
4052:
4038:
4037:
4034:
4033:
4030:
4029:
4026:
4025:
4023:
4022:
4017:
4012:
4007:
4002:
3997:
3995:Notoscyphaceae
3992:
3987:
3982:
3977:
3972:
3967:
3962:
3957:
3955:Calypogeiaceae
3952:
3947:
3942:
3937:
3932:
3926:
3924:
3920:
3919:
3917:
3916:
3911:
3906:
3901:
3896:
3891:
3889:Cephaloziaceae
3886:
3881:
3875:
3873:
3872:Cephaloziineae
3869:
3868:
3866:
3865:
3860:
3855:
3850:
3845:
3843:Lophocoleaceae
3840:
3835:
3830:
3825:
3820:
3814:
3812:
3811:Lophocoleineae
3808:
3807:
3805:
3804:
3798:
3796:
3790:
3789:
3787:
3786:
3780:
3778:
3772:
3771:
3769:
3768:
3766:Diettertiaceae
3761:
3759:
3758:Diettertiineae
3752:
3746:
3745:
3743:
3742:
3737:
3732:
3726:
3724:
3718:
3717:
3715:
3714:
3709:
3704:
3699:
3694:
3689:
3684:
3678:
3676:
3667:
3661:
3660:
3657:
3656:
3654:
3653:
3648:
3642:
3640:
3634:
3633:
3631:
3630:
3624:
3622:
3613:
3609:
3608:
3605:
3604:
3602:
3601:
3600:
3599:
3594:
3589:
3581:
3580:
3579:
3571:
3570:
3569:
3560:
3558:
3552:
3551:
3549:
3548:
3547:
3546:
3541:
3536:
3531:
3526:
3518:
3517:
3516:
3507:
3505:
3499:
3498:
3496:
3495:
3490:
3484:
3482:
3473:
3466:
3460:
3459:
3456:
3455:
3452:
3451:
3449:
3448:
3443:
3438:
3433:
3428:
3423:
3421:Marchantiaceae
3418:
3416:Dumortieraceae
3413:
3408:
3403:
3398:
3393:
3387:
3385:
3379:
3378:
3376:
3375:
3370:
3365:
3359:
3357:
3351:
3350:
3348:
3347:
3341:
3339:
3333:
3332:
3330:
3329:
3323:
3321:
3315:
3314:
3312:
3311:
3304:
3302:
3292:
3286:
3285:
3282:
3281:
3279:
3278:
3272:
3266:
3264:
3255:
3246:
3240:
3239:
3236:
3235:
3233:
3232:
3226:
3224:
3218:
3217:
3215:
3214:
3208:
3206:
3204:Haplomitriales
3197:
3191:
3190:
3188:
3187:
3182:
3177:
3175:Polypodiophyta
3172:
3167:
3162:
3154:
3151:
3150:
3142:
3141:
3134:
3127:
3119:
3113:
3112:
3107:
3102:
3097:
3090:
3089:External links
3087:
3086:
3085:
3082:
3060:
3059:
3014:(5): 517–541.
2994:
2985:|journal=
2948:
2907:
2877:
2830:
2783:
2756:(3): 231–247.
2733:
2700:
2669:
2657:
2624:
2603:(4): 1734–46.
2580:
2564:
2563:
2561:
2558:
2550:
2549:
2541:
2522:Palisade layer
2505:
2498:
2497:
2496:
2495:
2494:
2493:
2492:
2491:
2489:Schiffner 1896
2473:
2472:
2471:
2459:Dumortier 1829
2453:
2452:
2451:
2441:
2423:
2422:
2421:
2403:
2402:
2401:
2383:
2382:
2381:
2363:
2362:
2361:
2346:Marchantiaceae
2343:
2342:
2341:
2323:
2322:
2321:
2311:
2301:
2295:Aitchisoniella
2283:
2282:
2281:
2266:Dumortieraceae
2263:
2262:
2261:
2243:
2242:
2241:
2231:
2213:
2212:
2211:
2193:
2192:
2191:
2181:
2171:
2161:
2143:
2142:
2141:
2131:
2121:
2111:
2101:
2074:
2073:Classification
2071:
2040:
2037:
2033:
2032:
2027:
2023:
2022:
2019:
2018:
2014:
2013:
2010:
2009:
2005:
2004:
2001:
2000:
1996:
1995:
1992:
1991:
1987:
1986:
1983:
1982:
1978:
1977:
1974:
1973:
1969:
1968:
1965:
1964:
1960:
1959:
1956:
1955:
1951:
1950:
1947:
1946:
1942:
1941:
1938:
1937:
1933:
1932:
1929:
1928:
1924:
1923:
1920:
1919:
1915:
1914:
1911:
1910:
1906:
1905:
1902:
1901:
1897:
1896:
1893:
1892:
1888:
1887:
1884:
1883:
1879:
1878:
1875:
1874:
1867:
1864:
1863:
1860:
1859:
1852:
1847:
1845:
1842:
1841:
1838:
1837:
1833:
1832:
1829:
1828:
1824:
1823:
1820:
1819:
1807:
1804:
1803:
1800:
1799:
1790:
1787:
1786:
1783:
1782:
1773:
1768:
1766:
1760:
1759:
1756:
1755:
1748:
1743:
1741:
1736:
1734:
1731:
1730:
1727:
1726:
1722:
1721:
1718:
1717:
1710:
1707:
1706:
1703:
1702:
1695:
1690:
1688:
1683:
1681:
1678:
1677:
1674:
1673:
1669:
1668:
1665:
1664:
1660:
1659:
1656:
1655:
1648:
1645:
1644:
1641:
1640:
1633:
1628:
1626:
1623:
1622:
1619:
1618:
1611:
1606:
1604:
1599:
1597:
1594:
1593:
1590:
1589:
1582:
1577:
1575:
1570:
1568:
1565:
1564:
1561:
1560:
1553:
1548:
1546:
1543:
1542:
1539:
1538:
1535:Dumortieraceae
1531:
1526:
1524:
1521:
1520:
1517:
1516:
1512:
1511:
1508:
1507:
1498:
1495:
1494:
1491:
1490:
1478:
1473:
1471:
1469:Marchantiaceae
1463:
1461:
1458:
1457:
1454:
1453:
1449:
1448:
1445:
1444:
1432:
1429:
1428:
1425:
1424:
1417:
1412:
1410:
1405:
1403:
1402:Marchantiales
1399:
1398:
1395:
1394:
1387:
1382:
1380:
1375:
1373:
1370:
1369:
1366:
1365:
1358:
1353:
1351:
1345:
1344:
1341:
1340:
1333:
1328:
1326:
1318:
1317:
1314:
1313:
1309:
1308:
1305:
1304:
1300:
1299:
1296:
1295:
1288:
1285:
1284:
1281:
1280:
1276:
1275:
1272:
1271:
1267:
1266:
1263:
1262:
1255:
1252:
1251:
1248:
1247:
1235:
1230:
1228:
1225:
1224:
1221:
1220:
1213:
1208:
1206:
1198:
1196:
1193:
1192:
1189:
1188:
1184:
1183:
1180:
1179:
1172:
1169:
1168:
1165:
1164:
1152:
1147:
1145:
1140:
1138:
1128:
1126:
1123:
1122:
1119:
1118:
1109:
1104:
1102:
1096:
1095:
1092:
1091:
1082:
1077:
1053:
1050:
1046:
1045:
1042:
1041:
1037:
1036:
1033:
1032:
1028:
1027:
1024:
1023:
1019:
1018:
1015:
1014:
1010:
1009:
1006:
1005:
1001:
1000:
997:
996:
992:
991:
988:
987:
983:
982:
979:
978:
974:
973:
970:
969:
965:
964:
961:
960:
956:
955:
952:
951:
942:
939:
938:
935:
934:
925:
922:
921:
918:
917:
908:
903:
901:
898:
897:
894:
893:
884:
879:
877:
871:
870:
867:
866:
857:
849:
847:
844:
843:
840:
839:
830:
822:
820:
817:
816:
813:
812:
808:
807:
804:
803:
794:
788:
787:
784:
783:
774:
766:
764:
759:
757:
754:
753:
750:
749:
745:
744:
741:
740:
736:
735:
732:
731:
727:
726:
723:
722:
713:
710:
709:
706:
705:
696:
691:
689:
683:
682:
679:
678:
669:
661:
659:
656:
655:
652:
651:
647:
646:
643:
642:
633:
627:
626:
623:
622:
613:
605:
603:
598:
596:
591:
589:
586:
585:
582:
581:
577:
576:
573:
572:
568:
567:
564:
563:
559:
558:
555:
554:
545:
542:
541:
538:
537:
528:
523:
521:
518:
517:
514:
513:
509:
508:
505:
504:
495:
492:
491:
488:
487:
478:
473:
471:
466:
464:
461:
460:
457:
456:
452:Aitchisoniella
447:
442:
440:
432:
430:
427:
426:
423:
422:
418:
417:
414:
413:
409:
408:
405:
404:
400:
399:
396:
395:
391:
390:
387:
386:
377:
374:
373:
370:
369:
360:
355:
353:
350:
349:
346:
345:
336:
331:
329:
326:
325:
322:
321:
312:
307:
305:
302:
301:
298:
297:
288:
283:
281:
273:
271:
268:
267:
264:
263:
254:
252:Dumortieraceae
246:
244:
241:
240:
237:
236:
227:
225:Marchantiaceae
219:
217:
212:
207:
204:
128:
127:
123:Classification
116:
115:
111:
110:
101:
97:
96:
91:
87:
86:
81:
77:
76:
71:
67:
66:
61:
57:
56:
43:
42:
35:
34:
26:
25:
24:Marchantiales
15:
9:
6:
4:
3:
2:
4356:
4345:
4342:
4340:
4339:Marchantiales
4337:
4336:
4334:
4317:
4312:
4308:
4304:
4299:
4295:
4291:
4286:
4282:
4278:
4277:marchantiales
4272:
4268:
4263:
4259:
4255:
4250:
4246:
4242:
4236:
4232:
4227:
4223:
4219:
4214:
4210:
4206:
4201:
4197:
4193:
4188:
4184:
4180:
4175:
4171:
4167:
4162:
4158:
4154:
4149:
4145:
4141:
4136:
4132:
4128:
4123:
4119:
4115:
4110:
4106:
4102:
4097:
4093:
4088:
4087:Marchantiales
4082:
4078:
4073:
4067:
4063:
4062:
4060:
4058:
4057:Marchantiales
4054:
4050:
4045:
4021:
4018:
4016:
4013:
4011:
4008:
4006:
4003:
4001:
4000:Saccogynaceae
3998:
3996:
3993:
3991:
3988:
3986:
3985:Jackiellaceae
3983:
3981:
3980:Harpanthaceae
3978:
3976:
3975:Gyrothyraceae
3973:
3971:
3968:
3966:
3965:Geocalycaceae
3963:
3961:
3958:
3956:
3953:
3951:
3948:
3946:
3943:
3941:
3938:
3936:
3933:
3931:
3930:Acrobolbaceae
3928:
3927:
3925:
3921:
3915:
3912:
3910:
3907:
3905:
3902:
3900:
3897:
3895:
3892:
3890:
3887:
3885:
3882:
3880:
3879:Adelanthaceae
3877:
3876:
3874:
3870:
3864:
3861:
3859:
3856:
3854:
3851:
3849:
3846:
3844:
3841:
3839:
3838:Lepidoziaceae
3836:
3834:
3833:Lepicoleaceae
3831:
3829:
3826:
3824:
3821:
3819:
3816:
3815:
3813:
3809:
3803:
3800:
3799:
3797:
3795:
3791:
3785:
3782:
3781:
3779:
3777:
3773:
3767:
3763:
3762:
3760:
3756:
3753:
3751:
3747:
3741:
3738:
3736:
3733:
3731:
3728:
3727:
3725:
3723:
3719:
3713:
3710:
3708:
3705:
3703:
3702:Frullaniaceae
3700:
3698:
3695:
3693:
3690:
3688:
3685:
3683:
3680:
3679:
3677:
3675:
3671:
3668:
3666:
3662:
3652:
3651:Metzgeriaceae
3649:
3647:
3644:
3643:
3641:
3639:
3635:
3629:
3628:Pleuroziaceae
3626:
3625:
3623:
3621:
3617:
3614:
3610:
3598:
3595:
3593:
3590:
3588:
3587:Allisoniaceae
3585:
3584:
3582:
3578:
3575:
3574:
3572:
3568:
3565:
3564:
3562:
3561:
3559:
3557:
3553:
3545:
3542:
3540:
3537:
3535:
3532:
3530:
3527:
3525:
3522:
3521:
3519:
3515:
3512:
3511:
3509:
3508:
3506:
3504:
3500:
3494:
3491:
3489:
3486:
3485:
3483:
3481:
3477:
3474:
3470:
3467:
3465:
3461:
3447:
3444:
3442:
3441:Targioniaceae
3439:
3437:
3434:
3432:
3429:
3427:
3424:
3422:
3419:
3417:
3414:
3412:
3411:Cyathodiaceae
3409:
3407:
3404:
3402:
3399:
3397:
3394:
3392:
3389:
3388:
3386:
3384:
3383:Marchantiales
3380:
3374:
3371:
3369:
3366:
3364:
3363:Monocarpaceae
3361:
3360:
3358:
3356:
3352:
3346:
3345:Lunulariaceae
3343:
3342:
3340:
3338:
3334:
3328:
3325:
3324:
3322:
3320:
3316:
3310:
3306:
3305:
3303:
3301:
3296:
3293:
3291:
3290:Marchantiidae
3287:
3277:
3276:Treubiitaceae
3273:
3271:
3268:
3267:
3265:
3263:
3259:
3256:
3254:
3250:
3247:
3245:
3241:
3231:
3228:
3227:
3225:
3223:
3219:
3213:
3210:
3209:
3207:
3205:
3201:
3198:
3196:
3192:
3186:
3183:
3181:
3178:
3176:
3173:
3171:
3168:
3166:
3163:
3160:
3156:
3155:
3152:
3148:
3140:
3135:
3133:
3128:
3126:
3121:
3120:
3117:
3111:
3108:
3106:
3103:
3101:
3098:
3096:
3093:
3092:
3083:
3080:
3079:0-521-66097-1
3076:
3072:
3068:
3064:
3063:
3055:
3051:
3047:
3043:
3039:
3035:
3030:
3025:
3021:
3017:
3013:
3009:
3005:
2998:
2990:
2977:
2968:
2963:
2959:
2952:
2944:
2940:
2935:
2930:
2926:
2922:
2918:
2911:
2895:
2891:
2887:
2881:
2873:
2869:
2864:
2859:
2854:
2849:
2846:(59): 1–826.
2845:
2841:
2834:
2826:
2822:
2818:
2814:
2810:
2806:
2802:
2798:
2794:
2787:
2779:
2775:
2771:
2767:
2763:
2759:
2755:
2751:
2747:
2740:
2738:
2728:
2723:
2719:
2715:
2711:
2704:
2696:
2692:
2688:
2684:
2680:
2673:
2660:
2654:
2650:
2646:
2642:
2638:
2631:
2629:
2620:
2616:
2611:
2606:
2602:
2598:
2594:
2587:
2585:
2576:
2569:
2565:
2552:
2545:
2537:
2530:
2523:
2515:
2502:
2487:
2486:
2482:
2481:
2477:
2474:
2467:
2466:
2462:
2461:
2457:
2456:Targioniaceae
2454:
2447:
2446:
2442:
2439:Linnaeus 1753
2437:
2436:
2432:
2431:
2427:
2424:
2417:
2416:
2412:
2411:
2407:
2404:
2399:Griffith 1849
2397:
2396:
2392:
2391:
2387:
2384:
2377:
2376:
2372:
2371:
2367:
2364:
2359:Linnaeus 1753
2357:
2356:
2352:
2351:
2347:
2344:
2337:
2336:
2332:
2331:
2327:
2326:Lunulariaceae
2324:
2317:
2316:
2312:
2307:
2306:
2302:
2297:
2296:
2292:
2291:
2287:
2284:
2277:
2276:
2272:
2271:
2267:
2264:
2257:
2256:
2252:
2251:
2247:
2246:Cyathodiaceae
2244:
2239:Berkeley 1857
2237:
2236:
2232:
2227:
2226:
2222:
2221:
2217:
2214:
2207:
2206:
2202:
2201:
2197:
2194:
2187:
2186:
2182:
2179:Lindberg 1876
2177:
2176:
2172:
2169:Lindberg 1868
2167:
2166:
2162:
2159:Falconer 1848
2157:
2156:
2152:
2151:
2147:
2144:
2137:
2136:
2132:
2127:
2126:
2122:
2117:
2116:
2112:
2107:
2106:
2105:Cryptomitrium
2102:
2097:
2096:
2092:
2091:
2087:
2084:
2083:
2082:
2080:
2070:
2068:
2063:
2060:
2057:
2053:
2049:
2044:
2031:
2030:
2021:
2020:
2012:
2011:
2003:
2002:
1994:
1993:
1985:
1984:
1976:
1975:
1967:
1966:
1958:
1957:
1949:
1948:
1940:
1939:
1931:
1930:
1922:
1921:
1913:
1912:
1904:
1903:
1895:
1894:
1886:
1885:
1877:
1876:
1873:
1872:
1866:
1865:
1862:
1861:
1858:
1857:
1856:Cyathodiaceae
1851:
1850:
1844:
1843:
1840:
1839:
1831:
1830:
1822:
1821:
1818:
1817:
1815:
1814:
1806:
1805:
1802:
1801:
1798:
1797:
1796:
1789:
1788:
1785:
1784:
1781:
1780:
1779:
1772:
1771:
1765:
1762:
1761:
1758:
1757:
1754:
1753:
1747:
1746:
1740:
1739:
1733:
1732:
1729:
1728:
1720:
1719:
1716:
1715:
1709:
1708:
1705:
1704:
1701:
1700:
1694:
1693:
1687:
1686:
1680:
1679:
1676:
1675:
1667:
1666:
1658:
1657:
1654:
1653:
1647:
1646:
1643:
1642:
1639:
1638:
1637:Targioniaceae
1632:
1631:
1625:
1624:
1621:
1620:
1617:
1616:
1610:
1609:
1603:
1602:
1596:
1595:
1592:
1591:
1588:
1587:
1581:
1580:
1574:
1573:
1567:
1566:
1563:
1562:
1559:
1558:
1552:
1551:
1545:
1544:
1541:
1540:
1537:
1536:
1530:
1529:
1523:
1522:
1519:
1518:
1510:
1509:
1506:
1505:
1504:
1497:
1496:
1493:
1492:
1489:
1488:
1486:
1485:
1477:
1476:
1470:
1467:
1466:
1460:
1459:
1456:
1455:
1447:
1446:
1443:
1442:
1440:
1439:
1431:
1430:
1427:
1426:
1423:
1422:
1421:Lunulariaceae
1416:
1415:
1409:
1408:
1401:
1400:
1397:
1396:
1393:
1392:
1386:
1385:
1379:
1378:
1372:
1371:
1368:
1367:
1364:
1363:
1357:
1356:
1350:
1349:Marchantiidae
1347:
1346:
1343:
1342:
1339:
1338:
1332:
1331:
1325:
1324:
1320:
1319:
1316:
1315:
1307:
1306:
1298:
1297:
1294:
1293:
1287:
1286:
1283:
1282:
1274:
1273:
1265:
1264:
1261:
1260:
1254:
1253:
1250:
1249:
1246:
1245:
1243:
1242:
1234:
1233:
1227:
1226:
1223:
1222:
1219:
1218:
1212:
1211:
1205:
1202:
1201:
1195:
1194:
1191:
1190:
1182:
1181:
1178:
1177:
1171:
1170:
1167:
1166:
1163:
1162:
1160:
1159:
1151:
1150:
1144:
1143:
1137:
1136:
1132:
1131:
1125:
1124:
1121:
1120:
1117:
1116:
1115:
1108:
1107:
1101:
1098:
1097:
1094:
1093:
1090:
1089:
1088:
1081:
1080:
1076:
1073:
1070:
1066:
1062:
1058:
1044:
1043:
1035:
1034:
1026:
1025:
1017:
1016:
1008:
1007:
999:
998:
990:
989:
981:
980:
972:
971:
963:
962:
954:
953:
950:
949:
948:
941:
940:
937:
936:
933:
932:
931:
924:
923:
920:
919:
916:
915:
914:
907:
906:
900:
899:
896:
895:
892:
891:
890:
883:
882:
876:
873:
872:
869:
868:
865:
864:
863:
856:
855:Cyathodiaceae
853:
852:
846:
845:
842:
841:
838:
837:
836:
829:
826:
825:
819:
818:
815:
814:
806:
805:
802:
801:
800:
793:
790:
789:
786:
785:
782:
781:
780:
773:
772:Targioniaceae
770:
769:
763:
762:
756:
755:
752:
751:
743:
742:
734:
733:
725:
724:
721:
720:
719:
712:
711:
708:
707:
704:
703:
702:
695:
694:
688:
685:
684:
681:
680:
677:
676:
675:
668:
665:
664:
658:
657:
654:
653:
645:
644:
641:
640:
639:
632:
629:
628:
625:
624:
621:
620:
619:
612:
609:
608:
602:
601:
595:
594:
588:
587:
584:
583:
575:
574:
566:
565:
557:
556:
553:
552:
551:
544:
543:
540:
539:
536:
535:
534:
527:
526:
520:
519:
516:
515:
507:
506:
503:
502:
501:
494:
493:
490:
489:
486:
485:
484:
477:
476:
470:
469:
463:
462:
459:
458:
455:
454:
453:
446:
445:
439:
436:
435:
429:
428:
425:
424:
416:
415:
407:
406:
398:
397:
389:
388:
385:
384:
383:
376:
375:
372:
371:
368:
367:
366:
359:
358:
352:
351:
348:
347:
344:
343:
342:
335:
334:
328:
327:
324:
323:
320:
319:
318:
311:
310:
304:
303:
300:
299:
296:
295:
294:
293:Cryptomitrium
287:
286:
280:
277:
276:
270:
269:
266:
265:
262:
261:
260:
253:
250:
249:
243:
242:
239:
238:
235:
234:
233:
226:
223:
222:
216:
215:
211:
203:
201:
197:
196:
191:
190:
185:
184:
180:
175:
173:
169:
165:
161:
156:
154:
153:
148:
147:
142:
138:
134:
133:Marchantiales
126:
125:
124:
117:
112:
105:
104:Marchantiales
102:
99:
98:
95:
94:Marchantiidae
92:
89:
88:
85:
82:
79:
78:
75:
72:
69:
68:
65:
62:
59:
58:
53:
48:
44:
40:
36:
32:
27:
22:
19:
4056:
4010:Southbyaceae
3940:Arnelliaceae
3935:Antheliaceae
3914:Scapaniaceae
3904:Lophoziaceae
3828:Herbertaceae
3735:Ptilidiaceae
3712:Lejeuneaceae
3638:Metzgeriales
3620:Pleuroziales
3612:Metzgeriidae
3573:Makinoineae
3529:Moerckiaceae
3426:Monocleaceae
3406:Corsiniaceae
3382:
3337:Lunulariales
3309:Naiaditaceae
3185:Angiospermae
3070:
3066:
3029:10261/248464
3011:
3007:
2997:
2976:cite journal
2951:
2924:
2920:
2910:
2898:. Retrieved
2894:the original
2889:
2880:
2843:
2833:
2800:
2796:
2786:
2753:
2749:
2727:10.26879/917
2717:
2713:
2703:
2686:
2682:
2672:
2662:, retrieved
2640:
2600:
2596:
2574:
2568:
2483:
2463:
2445:Ricciocarpos
2443:
2433:
2413:
2406:Oxymitraceae
2395:Monosolenium
2393:
2373:
2366:Monocleaceae
2353:
2349:Lindley 1836
2339:Adanson 1763
2333:
2313:
2303:
2299:Kashyap 1914
2293:
2273:
2253:
2233:
2223:
2216:Corsiniaceae
2205:Conocephalum
2203:
2183:
2173:
2163:
2153:
2133:
2125:Plagiochasma
2123:
2113:
2103:
2093:
2078:
2076:
2066:
2064:
2058:
2056:Marchantites
2055:
2051:
2047:
2045:
2042:
1871:Corsiniaceae
1869:
1854:
1811:
1810:
1809:
1793:
1792:
1778:Ricciocarpos
1776:
1775:
1752:Oxymitraceae
1750:
1712:
1699:Monocleaceae
1697:
1650:
1635:
1613:
1584:
1555:
1533:
1501:
1500:
1482:
1481:
1480:
1436:
1435:
1434:
1419:
1389:
1360:
1335:
1321:
1290:
1259:Metzgeriales
1257:
1239:
1238:
1237:
1217:Pleuroziales
1215:
1204:Metzgeriidae
1174:
1156:
1155:
1154:
1133:
1112:
1111:
1085:
1084:
1074:
1068:
1064:
1063:
1059:
1055:
945:
944:
928:
927:
911:
910:
887:
886:
875:Corsiniaceae
860:
859:
835:Monosolenium
833:
832:
797:
796:
777:
776:
716:
715:
701:Ricciocarpos
699:
698:
672:
671:
667:Oxymitraceae
638:Conocephalum
636:
635:
616:
615:
611:Monocleaceae
548:
547:
531:
530:
498:
497:
481:
480:
450:
449:
382:Plagiochasma
380:
379:
363:
362:
339:
338:
315:
314:
291:
290:
257:
256:
230:
229:
209:
193:
187:
181:
176:
158:As in other
157:
150:
144:
139:of thallose
132:
131:
120:
119:
108:Limpr., 1877
103:
38:
18:
4174:iNaturalist
4081:Wikispecies
3823:Grolleaceae
3722:Ptilidiales
3682:Porellaceae
3577:Makinoaceae
3391:Aytoniaceae
3300:Naiaditales
3230:Treubiaceae
3157:Supergroup
2695:10539/16189
2508:100 ÎĽm
2485:Wiesnerella
2379:Hooker 1820
2369:Frank 1877
2309:Mitten 1870
2305:Exormotheca
2219:Engler 1892
2149:Cavers 1911
2089:Cavers 1911
2086:Aytoniaceae
2048:Hepaticites
1557:Aytoniaceae
947:Exormotheca
799:Wiesnerella
279:Aytoniaceae
164:gametophyte
4333:Categories
3707:Jubulaceae
3697:Radulaceae
3674:Porellales
3646:Aneuraceae
3488:Pelliaceae
3436:Ricciaceae
3396:Cleveaceae
3368:Riellaceae
3270:Blasiaceae
3222:Treubiales
3095:Life cycle
3008:Cladistics
2750:Cladistics
2664:2020-11-02
2560:References
2529:Parenchyma
2479:Inoue 1976
2449:Corda 1829
2426:Ricciaceae
2389:Inoue 1966
2355:Marchantia
2275:Dumortiera
2255:Cyathodium
2229:Raddi 1818
2175:Peltolepis
2146:Cleveaceae
2046:The genus
1813:Ricciopsis
1764:Ricciaceae
1586:Cleveaceae
1503:Marchantia
862:Cyathodium
687:Ricciaceae
500:Peltolepis
438:Cleveaceae
259:Dumortiera
232:Marchantia
183:Marchantia
172:life cycle
168:sporophyte
160:bryophytes
141:liverworts
90:Subclass:
70:Division:
3802:Myliaceae
3794:Myliineae
3480:Pelliales
3472:Pelliidae
3262:Blasiales
3253:Blasiidae
3165:Bryophyta
3038:1096-0031
2943:0960-4286
2844:PhytoKeys
2817:0028-646X
2778:225165843
2689:: 21–30.
2514:Epidermis
2465:Targionia
2335:Lunularia
2279:Nees 1824
2269:Long 2006
2189:Nees 1838
2155:Athalamia
2095:Asterella
1337:Blasiidae
779:Targionia
533:Athalamia
341:Asterella
189:Asterella
114:Families
60:Kingdom:
4303:35206120
4298:Tropicos
4066:Wikidata
3054:52831959
3046:34706484
2872:26929706
2825:37697646
2770:34478198
2619:26505145
2536:Rhizoids
2415:Oxymitra
2375:Monoclea
2235:Cronisia
2225:Corsinia
2185:Sauteria
2135:Reboulia
1176:Pellidae
930:Cronisia
889:Corsinia
674:Oxymitra
618:Monoclea
550:Sauteria
365:Reboulia
4072:Q139099
2900:30 June
2863:4758082
2516: ↓
200:elaters
100:Order:
80:Class:
64:Plantae
4254:927429
4238:NZOR:
4179:152699
4114:176416
4101:155304
3077:
3052:
3044:
3036:
2941:
2870:
2860:
2823:
2815:
2776:
2768:
2655:
2617:
2435:Riccia
2165:Clevea
2115:Mannia
2059:loreus
1795:Riccia
718:Riccia
483:Clevea
317:Mannia
195:Riccia
162:, the
135:is an
4285:Plazi
4274:PPE:
4267:55125
4231:28908
4205:15520
4192:11563
4187:IRMNG
4153:1MHTO
3159:Plant
3050:S2CID
2774:S2CID
2720:(2).
179:genus
137:order
4226:NCBI
4200:ITIS
4161:GBIF
4148:EPPO
4140:4135
4109:BOLD
4096:APNI
3075:ISBN
3042:PMID
3034:ISSN
2989:help
2939:ISSN
2902:2016
2868:PMID
2821:PMID
2813:ISSN
2766:PMID
2653:ISBN
2615:PMID
177:The
121:See
4311:WFO
4213:NBN
4166:376
4135:EoL
4127:3HV
4122:CoL
3024:hdl
3016:doi
2962:doi
2929:doi
2858:PMC
2848:doi
2805:doi
2801:240
2758:doi
2722:doi
2691:hdl
2645:doi
2605:doi
2601:209
4335::
4313::
4300::
4287::
4264::
4251::
4228::
4215::
4202::
4189::
4176::
4163::
4150::
4137::
4124::
4111::
4098::
4083::
4068::
3067:in
3048:.
3040:.
3032:.
3022:.
3012:34
3010:.
3006:.
2980::
2978:}}
2974:{{
2960:.
2937:.
2925:63
2923:.
2919:.
2888:.
2866:.
2856:.
2842:.
2819:.
2811:.
2799:.
2795:.
2772:.
2764:.
2754:16
2752:.
2748:.
2736:^
2718:22
2716:.
2712:.
2687:30
2685:.
2681:.
2651:,
2639:,
2627:^
2613:.
2599:.
2595:.
2583:^
1816:+
1487:+
1441:+
1244:+
1161:+
3764:â€
3307:â€
3298:â€
3274:â€
3161::
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3081:.
3056:.
3026::
3018::
2991:)
2987:(
2970:.
2964::
2945:.
2931::
2904:.
2874:.
2850::
2827:.
2807::
2780:.
2760::
2730:.
2724::
2693::
2647::
2621:.
2607::
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
