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Lichenology

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disciple of Linnaeus, who is today considered the father of lichenology, starting the taxonomy of lichens with his pioneering study of Swedish lichens in Lichenographiae Suecicae Prodromus of 1798 or in his Synopsis Methodica Lichenum, Sistens omnes hujus Ordinis Naturalis of 1814. These studies and
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grouped them into their own genus. He adopted the Latin term lichen, which had already been used by Pliny who had imported it from Theophrastus but up until then this term had not been widely employed. The original meaning of the Greek word λειχήν (leichen) was moss that in its turn derives from the
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where the age of an exposed surface can be found by studying the age of lichens growing on them. Age dating in this way can be absolute or relative because the growth of these organisms can be arrested under various conditions. The technique provides an average age of the older individual lichens
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During the 20th century botany and mycology were still attempting to solve the two main problems surrounding lichens. On the one hand the definition of lichens and the relationship between the two symbionts and the taxonomic position of these organisms within the plant and fungal kingdoms. There
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considered the definition imperfect because it is impossible to determine which one thallus is of a specific structure since thalli changed depending upon the substrate and conditions in which they developed. This researcher represents one of the main trends among lichenologists who consider it
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demonstrated that all lichens were the result of fungal attack on the cells of algal cells and that all these algae also exist free in nature. This researcher was the first to recognise the dual nature of lichens as a result of the capture of the algal component by the fungal component. In 1873
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Over the years research shed new light into the nature of these organisms still classified as plants. A controversial issue surrounding lichens since the early 19th century is their reproduction. In these years a group of researchers faithful to the tenets of Linnaeus considered that lichens
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classifications are the cornerstone of subsequent investigations. In these early years of structuring the new discipline various works of outstanding scientific importance appeared such as Lichenographia Europaea Reformata published in 1831 by
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Gollapudi, S. R.; Telikepalli, H; Jampani, H. B.; Mirhom, Y. W.; Drake, S. D.; Bhattiprolu, K. R.; Vander Velde, D; Mitscher, L. A. (1994). "Alectosarmentin, a new antimicrobial dibenzofuranoid lactol from the lichen, Alectoria sarmentosa".
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Lichens as a group have received less attention in classical treatises on botany than other groups although the relationship between humans and some species has been documented from early times. Several species have appeared in the works of
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were identified in the lichen structure, which heightened the contradictions. At first the presence of algae was taken as being due to contamination due to collection of samples in damp conditions and they were not considered as being in a
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But these works suffer from being superficial and mere lists of species without further physiological studies. It took until the middle of the 19th century for research to catch up using biochemical and physiological methods. In Germany
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in 1950 and is especially useful in areas exposed for less than 1000 years. Growth is greatest in the first 20 to 100 years with 15–50 mm growth per year and less in the following years with average growth of 2–4 mm per year.
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Despite the above studies the dual nature of lichens remained no more than a theory until in 1939 the Swiss researcher Eugen A Thomas was able to reproduce in the laboratory the phenotype of the lichen
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reproduced sexually and had sexual reproductive organs, as in other plants, independent of whether asexual reproduction also occurred. Other researchers only considered asexual reproduction by means of
1941: 546: 432: 644:. The definition finally adopted was that lichen could be considered as the association between a fungus and a photosynthetic symbiont resulting in a thallus of specific structure. 566:, A new study on the Origin of Organisms", which aims to explain a new theory of Symbiogenesis by lichens and other organisms as evidenced by his earlier work "Nature and Origin of 659:
Today studies in lichenology are not restricted to the description and taxonomy of lichens but have application in various scientific fields. Especially important are studies on
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and their reproductive mechanisms were totally ignored. For centuries naturalists had included lichens in diverse groups until in the early 18th century a French researcher
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The difficulty of giving a definition applicable to every known lichen has been debated since lichenologists first recognised the dual nature of lichens. In 1982 the
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in 1867 showed no dependence of the algal component upon the lichen thallus and that the algal component could live independently of the thallus. It was in 1869 that
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convened a meeting to adopt a single definition of lichen drawing on the proposals of a committee. The chairman of this committee was the renowned researcher
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Mike Crockett, Stacie Kageyama, Delfina Homen, Carrie Lewis, Jane Osborn, Logan Sander (2003). "Antibacterial Properties of four Pacific North West Lichens".
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has shown some antiviral activity with some lichen substances. In 1989 K Hirabayashi presented his investigations on inhibitory lichen polysaccharides in
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Francisco Javier Toledo Marante, Ana Garcia Costellano, Francisco Leon Oyola and Jaime Bermejo Barrera "Ecologia Quimica en Hongos y Liquenes" (Spa)
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that are made through the interaction of lichens with their environment. Lichen is extremely sensitive to various air pollutants, especially to
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providing a minimum age of the medium being studied. Lichenometry relies upon the fact that the maximum diameter of the largest thallus of an
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lichen growing on a substrate is directly proportional to the time from first exposure of the area to the environment as seen in studies by
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definition soon brought criticism from various lichenologists and there soon emerged reviews and suggestions for amendments. For example,
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Greek verb λείχω (liekho) to suck because of the great ability of these organisms to absorb water. In its original use the term signified
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Miguel Mateo Garcia, The Growth Curve of Rhizocarpon geographicum in the Valley of Madriu Andorra, Biophysical Processes in Cold Media
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although the studies are not very deep. During the first centuries of the modern age they were usually put forward as examples of
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it was not until the early 20th century that modern science became interested in them. The discovery of various substances with
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relation with the fungal part of the thallus. That the algae continued to multiply showed that they were not mere contaminants.
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A Textbook of General Lichenology with Descriptions and Figures of the Genera Occurring in the North Eastern United States
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action in lichen thalli was essential for scientists to become aware of the possible importance of these organisms to
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Marshak, A.; Barry, G. T.; Craig, L. C. (1947). "Antibiotic Compound Isolated from the Lichen Ramalina reticulata".
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Index of Collectors in Knowles the Lichens of Ireland (1929) and Porter's Supplement: with a Conspectus of Lichen
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Lichenology and Bryology in the Galapagos Islands with Checklists of the Lichens and Bryophytes thus far Reported
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concluded form studying many different lichen species that the relationship between fungi and algae was purely
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Biodeterioration of Stone Surfaces: Lichens and Biofilms as Weathering Agents of Rocks and Cultural Heritage
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in an article of 1852 "Memorie pour servir a l'Histoire des Lichens Organographique et Physiologique" by
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Antibacterial action of substances produced by lichens is related to their ability to disrupt bacterial
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Fortschritte der Chemie organischer Naturstoffe/ Progress in the Chemistry of Organic Natural Products
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who first suggested in 1865 that lichens were merely the result of parasitism of various fungi of the
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Irish Lichenology 1858–1880: Selected Letters of Isaac Carroll, Theobald Jones, Charles Larbalestier
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Fink, Bruce (1913). "The Nature and Classification of Lichens: II. The Lichen and its Algal Host".
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Honegger, Rosmarie (2000). "Simon Schwendener (1829–1919) and The Dual Hypothesis of Lichens".
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in the Plant Kingdom". These new ideas can be studied today under the title of the Theory of
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impossible to give a single definition to lichens since they are a unique type of organism.
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Scientific publications settled many unknown facts about lichens. In the French publication
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Bustinza, Francisco (1948) "Contribution to the Study of Antibiotics Produced by Lichens".
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David L. Hawksworth (1989) "Interactions Fungus and Alga in Lichen Symbiosis liquenoides"
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Brodo, Irwin M; Laurie-Bourque, Susan; Sharnoff, Sylvia Duran; Sharnoff, Stephen (2016).
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Interest in the potential of substances synthesised by lichens increased with the end of
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made the first division of the group created by Tournefort separating the sub-families
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and the compounds identified as responsible for bacterial inhibition were shown to be
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Hirabayashi, K; Iwata, S; Ito, M; Shigeta, S; Narui, T; Mori, T; Shibata, S (1989).
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The Journal of the Hattori Botanical Laboratory: Devoted to Bryology and Lichenology
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Bustinza, Francisco (1951) "Contribution to the Study of Antibacterial Activity in
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Studies in Lichenology with Emphasis on Chemotaxonomy, Geography and Phytochemistry
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These new discoveries were becoming increasingly contradictory for scientists. The
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Lichenology in the British Isles, 1568–1975: An Historical and Biographical Survey
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appeared numerous renowned researchers within the field of lichenology such as
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Neli Kika Honda & Wagner Vilegas (1998) "The Chemistry of Lichens" (Port)
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Flechten Follmann: Contributions to Lichenology in Honour of Gerhard Follmann
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Synopsis Methodica Lichenum: Systens omnes hujus ordinis naturalis detectas
1240: 1205: 957:, Gerhard Follmann, F. J. A. Daniels, Margot Schultz and Jorge Peine (1995) 845: 766: 620: 218: 188: 160: 95: 83: 2564: 2252: 2150: 724:. Studies showed that the substance that inhibited growth of bacteria was 2912: 2029:
http://lichens.science.oregonstate.edu/antibiotics/lichen_antibiotics.htm
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with eight divisions within the group according to the morphology of the
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Williams, Thomas A. (1856). "The Status of the Algo-Lichen Hypothesis".
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Edwards, Howell G.M.; Newton, Emma M.; Wynn-Williams, David D. (2003).
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substances. In 1947 antibacterial action was identified in extracts of
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type algae and others. Successive studies such as those carried out by
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Erik Acharius (1757–1819), Swedish botanist, the father of lichenology
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presented a research paper "The Theory of two Plasms as the basis of
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Lichenologists may also study the growth and growth rate of lichens,
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Environmental Lichenology: Biomonitoring Trace Element Air Pollution
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antitumour activity by substances identified in two common lichens
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Guide to using a Lichen-Based Index to Assess Nitrogen Air Quality
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Current biophysical processes in cold environments: recent studies
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Synopsis of lichen Methods, systems of this natural order detected
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The Present Status and Potentialities of the Lichenology in China
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and his new system of classification lichens are retained in the
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Procesos biofísicos actuales en medios fríos: estudios recientes
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in response to the morphological characteristics of the lichen
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Schneider, Albert (1895). "The Biological Status of Lichens".
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who demonstrated antibacterial action of lichens of the genus
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made up of an intimate symbiotic association of a microscopic
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Central European Bryological and Lichenological Society (Ger)
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Identifying North American Lichens a Guide to the Literature
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10.1639/0007-2745(2000)103[0307:ssatdh]2.0.co;2
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Study of lichens draws knowledge from several disciplines:
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French botanist (Jean-Baptiste) Édouard Bornet (1828–1911)
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Methods for species identification include reference to
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Netherlands Bryological and Lichenological Society (nl)
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10.1641/0006-3568(2003)053[1008:mm]2.0.co;2
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capacity. This is possible due to the action of lichen
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began to publish works of great scientific importance.
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Keys to Lichens of North America: Revised and Expanded
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Contemporary Lichenology and Lichens of Western Oregon
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Against this background appeared the Swedish botanist
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Keys to Lichens of North America: Revised and Expanded
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Contributions to Lichenology: In Honour of A. Henssen
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Diccionario de Liquenologia/Dictionary of Lichenology
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Although several species of lichen have been used in
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Later lichenologists include the American scientists
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of lichens was first intensively investigated by the
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LichenPortal.org - The Consortium of Lichen Herbaria
2426:"Lichens - The William & Lynda Steere Herbarium" 1912:] (in Spanish). Edicions Universitat Barcelona. 2560:
Belgium, Luxembourg and Northern France, Lichens of
1901: 404:or Enumeratio Critico Lichenum Europaeorum 1850 by 2719:Swiss Bryological and Lichenological Society (Ger) 2585:Czech Bryological and Lichenological Society (Cze) 2408:"Botany collections | Natural History Museum" 2383:"Fungi & Lichens | National Museum Wales" 2011: 2009: 867:, D. L. Hawksworth and M. R. D. Seaward (Dec 1977) 678:K S Merezhkovsky Russian Lichenologist (1855–1921) 2054: 995:Les Lichens - Morphologie, Biologie, Systematique 3071: 2480:"University of Michigan Herbarium - Collections" 728:. Something similar occurred with the substance 2555:American Bryological and Lichenological Society 2006: 1026:Swedish Lichenology: Dedicated to Roland Moberg 892:, Volume II: v.2, Margalith Galun (30 Sep 1988) 2318:"Collections | Canadian Museum of Nature" 1843: 1452: 148:. Some components of certain lichens may also 144:which can be characteristic of a given lichen 140:can be used to detect the presence of certain 2749: 1986:Schaetzl Randall, J; Sharon Anderson (2005). 1761:"Harvard University Herbaria & Libraries" 1699:"Harvard University Herbaria & Libraries" 1685:"Harvard University Herbaria & Libraries" 1671:"Harvard University Herbaria & Libraries" 1040:, Larry St. Clair and Mark Seaward (Oct 2011) 2282:"Herbaria | The British Lichen Society" 884:Lichenology in Indian Subcontinent 1966–1977 584:by combining its two identified components. 2575:Checklists of Lichens and Lichenolous Fungi 1992:. Cambridge University Press. p. 562. 2756: 2742: 2714:Swedish Lichens Lief & Anita Stridvall 1509: 682: 611:Lichenology has found applications beyond 2605:International Association for Lichenology 2242: 1861: 1634: 1062: 638:International Association for Lichenology 109:on lichens. An example reference work is 2763: 2116:Annals of the Botanical Garden of Madrid 2099:Annals of the Botanical Garden of Madrid 2017:Annals of the Botanical Garden of Madrid 1808: 1711: 1584: 673: 544: 410: 341: 333: 94:. Study of lichens is conducted by both 29: 769:along with the growing interest in all 90:. Scholars of lichenology are known as 27:Branch of mycology that studies lichens 14: 3072: 2231:Chemical & Pharmaceutical Bulletin 1971:: CS1 maint: archived copy as title ( 1463:. Yale University Press. p. 424. 825:research. These studies revealed some 2737: 2632:Japanese Lichenological Society (Eng) 2627:Japanese Lichenological Society (Eng) 1637:Bulletin of the Torrey Botanical Club 1334:National Botanical Research Institute 1307: 943:Lichens to Biomonitor the Environment 3040: 2622:Italian Lichenological Society (Ita) 1773: 1748:A Popular History of British Lichens 1623:A Popular History of British Lichens 1573:A Popular History of British Lichens 1560:A Popular History of British Lichens 1538:Joseph Pitton de Tournefort (1700). 1498:A Popular History of British Lichens 1485:A Popular History of British Lichens 1329:Centraalbureau voor Schimmelcultures 844:Recent work in the field of applied 802:with a subsequent loss of bacterial 167:, the ecological role of lichens in 129:by the same three authors joined by 3052: 2652:Links to Lichens and Lichenologists 2642:Lichen Herbarium University of Oslo 2590:French Lichenological Society (Fre) 2518:. December 19, 2023. Archived from 2500:. December 19, 2023. Archived from 1016:, H. M. Jahns and A. Henssen (1990) 473:reproductive organ being unique to 165:role of lichens in nutrient cycling 24: 2647:Lichenland Oregon State University 2482:. February 2, 2014. Archived from 2446:. February 1, 2014. Archived from 2358:"Herbarium: Iowa State University" 874:Lichenology in Indian Subcontinent 817:From the 1950s the lichen product 558:In 1909 the Russian lichenologist 25: 3111: 2548: 2201:Química Nova 22(1) ISSN 0100-4042 1989:Soils Genesis & Geomorphology 1902:Antonio Gómez Ortiz, ed. (1998). 1319:Botanische Staatssammlung München 1095:Ferdinand Christian Gustav Arnold 985:Lichens from West of Hudson's Bay 908:Bibliography of Irish Lichenology 896:A Textbook of General Lichenology 203: 3051: 3039: 3028: 3027: 1889:"Pixie Cups (Cladonia pyxidata)" 1746:Lauder Lindsay, William (1856). 1621:Lauder Lindsay, William (1856). 1558:Lauder Lindsay, William (1856). 1496:Lauder Lindsay, William (1856). 1483:Lauder Lindsay, William (1856). 1375:University of Michigan Herbarium 991:Vol. 1), John W. Thompson (1953) 898:, Albert Schneider (24 May 2013) 481:organisms. With improvements in 310:Lichenographiae Suecia prodromus 125:and that book's 2016 expansion, 2526: 2508: 2490: 2472: 2464:. April 6, 2013. Archived from 2454: 2436: 2418: 2400: 2375: 2350: 2328: 2310: 2304:www.botanischestaatssammlung.de 2292: 2286:www.britishlichensociety.org.uk 2274: 2259: 2218: 2204: 2192: 2157: 2121: 2104: 2091: 2048: 2034: 2022: 1979: 1934: 1895: 1881: 1870: 1837: 1802: 1767: 1753: 1740: 1705: 1691: 1677: 1663: 1628: 1615: 1597: 1384:University of Wisconsin-Madison 1364:Royal Botanic Garden, Edinburgh 1004:), David J. Galloway (Jul 1988) 855: 671:and prevents water absorption. 553: 460:Annales des Sciences Naturelles 390: 281:Kingdom forming a single group 2637:Lichenological Resources (Rus) 2269:28 ISSN 0370-3908 pp. 509–528 2170:Journal of Molecular Structure 1578: 1565: 1552: 1548:] (in Latin). Vol. 1. 1531: 1503: 1490: 1477: 1424: 1354:Natural History Museum, London 375:, as well as amateurs such as 183:topics including the study of 13: 1: 2684:National Biodiversity Gateway 2186:10.1016/S0022-2860(02)00626-9 1445: 1105:Friedrich August Georg Bitter 1046:, Vernon Ahmadjian (Aug 1993) 933:, Albert Schneider (Mar 2010) 916:, Kenneth Allen Hornak (1998) 598:Antonina Georgievna Borissova 2336:":: Welcome to NBRI ::" 2267:Columbian Academy of Science 2077:10.1126/science.106.2756.394 1417: 1120:Jean-Baptiste Édouard Bornet 532:Jean-Baptiste Edouard Bornet 7: 2689:Nordic Lichen Society (Eng) 2674:Microscopy of Lichens (Ger) 2131:Journal of Natural Products 1593:] (in Latin). Svanborg. 1541:Institutiones rei herbariae 1399: 1256:Joseph Pitton de Tournefort 1090:Johannes Müller Argoviensis 1052:, Thomas H. Nash (Jan 2008) 969:, Zennosuke Iwatsuki (1983) 910:, M. E. Mitchell (Nov 1972) 890:CRC Handbook of Lichenology 328:Synopsis methodica lichenum 231:Institutiones Rei Herbariae 227:Joseph Pitton de Tournefort 10: 3116: 2657:Lichens of Ireland Project 1844:Cavalier-Smith, T (2003). 1510:Eckel, P. M. (2010–2021). 1369:Royal Botanic Gardens, Kew 1130:François Fulgis Chevallier 1070:Henry Nicollon des Abbayes 975:, W. Clayton Fraser (1968) 939:, Liu Hua Jie (1 Jan 2000) 732:synthesised by the lichen 590:Henry Nicollon des Abbayes 322:Lichenographia universalis 198: 3023: 3015:List of mycology journals 2992: 2961: 2921: 2841: 2834: 2788: 2775: 1571:Lauder Lindsay, William. 1517:Missouri Botanical Garden 1413:, an award in lichenology 1359:New York Botanical Garden 1324:Canadian Museum of Nature 989:Lichens of Arctic America 951:, William A. Weber (1966) 921:Bibliotheca Lichenologica 451:and in the United States 246:. Some forty years later 1342:, Ada Hayden Herbarium, 1216:Konstantin Mereschkowski 1211:William Allport Leighton 1201:Georgij Karlovich Kreyer 619:in a technique known as 560:Konstantin Mereschkowski 449:William Allport Leighton 269:After the revolution in 111:Lichens of North America 2699:Paleo-Lichenology (Ger) 1877:Species:Eugen A. Thomas 1714:The American Naturalist 1585:Acharius, Erik (1814). 1349:National Museum Cardiff 1236:Charles Christian Plitt 1196:August von Krempelhuber 1181:David Leslie Hawksworth 1145:William Louis Culberson 997:, Fernand Moreau (1927) 963:, Joyce E. Sloof (1993) 902:Horizons in Lichenology 821:was the object of most 781:d-protolichosteric acid 683:Lichens in pharmacology 615:itself in the field of 373:Konstantin Merezhkovsky 2694:North American Lichens 2565:British Lichen Society 2444:"RBGE Lichen Taxonomy" 2118:(10) ISSN pp. 144–149. 1546:Institutions of botany 1314:British Lichen Society 1301:Alexander Zahlbruckner 1150:Johann Jacob Dillenius 1100:Heinrich Anton de Bary 1063:Notable lichenologists 832:Peltigera leucophlebia 740:Gram negative bacteria 679: 550: 416: 370:evolutionary biologist 354: 339: 223:spontaneous generation 177:anatomy and physiology 169:biological soil crusts 35: 3005:List of fungal orders 2704:Russian Lichens (Rus) 2580:Chilean Lichens (Spa) 2534:"National Museums NI" 2101:(7) ISSN pp. 511–548. 1340:Iowa State University 1175:Carolyn Wilson Harris 1058:, S. Hunek (Oct 2013) 704:Rufus Paul Burkholder 677: 661:environmental quality 548: 414: 345: 337: 173:morphology of lichens 33: 3090:Branches of mycology 3000:Glossary of mycology 1291:William Alfred Weber 1276:Edvard August Vainio 1186:Georg Franz Hoffmann 1140:Chicita F. Culberson 1044:The Lichen Symbiosis 878:Dharani Dhar Awasthi 689:traditional medicine 594:William Alfred Weber 477:but absent in other 131:Susan Laurie-Bourque 3085:Branches of biology 3010:List of mycologists 2244:10.1248/cpb.37.2410 2178:2003JMoSt.651...27E 2143:10.1021/np50109a009 2069:1947Sci...106..394M 1846:"Microbial Muddles" 1379:Ann Arbor, Michigan 1191:Peter Wilfred James 886:, Ajay Singh (1980) 735:Ramalina reticulata 653:David L. Hawksworth 606:George Albert Llano 505:who specialised in 191:and its own set of 2782:Outline of lichens 2662:2014-12-20 at the 2615:2014-12-20 at the 2172:. 651–653: 27–37. 2112:Cetraria islandica 1406:Outline of lichens 1388:Madison, Wisconsin 1308:Lichen collections 1271:Dalip Kumar Upreti 1165:Elias Magnus Fries 1115:Antonina Borissova 925:Elisabeth Peveling 904:D. H. Dalby (1988) 776:Cetraria islandica 680: 551: 417: 355: 340: 258:, Coralloides and 107:single-access keys 36: 3067: 3066: 2957: 2956: 2853:teleomorph/sexual 2430:sweetgum.nybg.org 2063:(2756): 394–395. 1251:Simon Schwendener 1155:Alexander Elenkin 838:Collema flaccidum 715:Bacillus subtilis 581:Cladonia pyxidata 527:Simon Schwendener 433:Johann Bayrhoffer 422:Hermann Itzigsohn 351:Evernia prunastri 316:Methodus lichenum 252:Historia Muscorum 193:other terminology 154:ultraviolet light 45:that studies the 41:is the branch of 16:(Redirected from 3107: 3095:Fungi and humans 3055: 3054: 3043: 3042: 3031: 3030: 2888:anamorph/asexual 2839: 2838: 2778:Outline of fungi 2758: 2751: 2744: 2735: 2734: 2724:Tropical Lichens 2709:Scottish Lichens 2542: 2541: 2530: 2524: 2523: 2512: 2506: 2505: 2494: 2488: 2487: 2476: 2470: 2469: 2458: 2452: 2451: 2440: 2434: 2433: 2422: 2416: 2415: 2404: 2398: 2397: 2395: 2394: 2385:. Archived from 2379: 2373: 2372: 2370: 2369: 2360:. Archived from 2354: 2348: 2347: 2342:. Archived from 2332: 2326: 2325: 2314: 2308: 2307: 2296: 2290: 2289: 2278: 2272: 2263: 2257: 2256: 2246: 2222: 2216: 2215: 2208: 2202: 2196: 2190: 2189: 2161: 2155: 2154: 2125: 2119: 2108: 2102: 2095: 2089: 2088: 2052: 2046: 2045: 2038: 2032: 2026: 2020: 2013: 2004: 2003: 1983: 1977: 1976: 1970: 1962: 1960: 1959: 1953: 1947:. Archived from 1946: 1938: 1932: 1923: 1899: 1893: 1892: 1885: 1879: 1874: 1868: 1867: 1865: 1841: 1835: 1834: 1806: 1800: 1799: 1771: 1765: 1764: 1757: 1751: 1744: 1738: 1737: 1709: 1703: 1702: 1695: 1689: 1688: 1681: 1675: 1674: 1667: 1661: 1660: 1632: 1626: 1619: 1613: 1612: 1609:plants.jstor.org 1601: 1595: 1594: 1582: 1576: 1569: 1563: 1556: 1550: 1549: 1535: 1529: 1528: 1526: 1524: 1507: 1501: 1494: 1488: 1481: 1475: 1474: 1456: 1439: 1434:", according to 1428: 1261:Edward Tuckerman 1231:William Nylander 1221:Camille Montagne 1160:Andrei Famintsyn 1110:Alphonse Boistel 1080:Vernon Ahmadjian 1002:Botany Bulletins 745:Escherichia coli 642:Vernon Ahmadjian 519:Andrei Famintsyn 453:Edward Tuckerman 441:Camille Montagne 430: 363:Edward Tuckerman 359:Vernon Ahmadjian 181:ethnolichenology 123:Stephen Sharnoff 102:lichenologists. 21: 3115: 3114: 3110: 3109: 3108: 3106: 3105: 3104: 3070: 3069: 3068: 3063: 3019: 2988: 2969:Dimorphic fungi 2962:Growth patterns 2953: 2917: 2830: 2811:Medicinal fungi 2784: 2771: 2762: 2664:Wayback Machine 2617:Wayback Machine 2551: 2546: 2545: 2532: 2531: 2527: 2516:"- Collections" 2514: 2513: 2509: 2498:"- Collections" 2496: 2495: 2491: 2478: 2477: 2473: 2460: 2459: 2455: 2442: 2441: 2437: 2424: 2423: 2419: 2406: 2405: 2401: 2392: 2390: 2381: 2380: 2376: 2367: 2365: 2356: 2355: 2351: 2340:www.nbri.res.in 2334: 2333: 2329: 2316: 2315: 2311: 2298: 2297: 2293: 2280: 2279: 2275: 2264: 2260: 2223: 2219: 2210: 2209: 2205: 2197: 2193: 2162: 2158: 2126: 2122: 2109: 2105: 2096: 2092: 2053: 2049: 2040: 2039: 2035: 2027: 2023: 2014: 2007: 2000: 1984: 1980: 1964: 1963: 1957: 1955: 1951: 1944: 1942:"Archived copy" 1940: 1939: 1935: 1920: 1900: 1896: 1887: 1886: 1882: 1875: 1871: 1842: 1838: 1807: 1803: 1788:10.2307/3753090 1772: 1768: 1759: 1758: 1754: 1745: 1741: 1710: 1706: 1697: 1696: 1692: 1683: 1682: 1678: 1669: 1668: 1664: 1649:10.2307/2478161 1633: 1629: 1620: 1616: 1603: 1602: 1598: 1583: 1579: 1570: 1566: 1557: 1553: 1536: 1532: 1522: 1520: 1508: 1504: 1495: 1491: 1482: 1478: 1471: 1457: 1453: 1448: 1443: 1442: 1429: 1425: 1420: 1402: 1310: 1305: 1286:Heinrich Walter 1226:Sanjeeva Nayaka 1135:Louisa Collings 1065: 858: 789:Alectosarmentin 685: 667:, which causes 665:sulphur dioxide 556: 424: 406:Ludwig Schaerer 393: 377:Louisa Collings 338:Lichen on rocks 215:Pliny the Elder 206: 201: 142:lichen products 119:Sylvia Sharnoff 28: 23: 22: 15: 12: 11: 5: 3113: 3103: 3102: 3097: 3092: 3087: 3082: 3065: 3064: 3062: 3061: 3049: 3037: 3024: 3021: 3020: 3018: 3017: 3012: 3007: 3002: 2996: 2994: 2990: 2989: 2987: 2986: 2981: 2976: 2971: 2965: 2963: 2959: 2958: 2955: 2954: 2952: 2951: 2946: 2941: 2936: 2931: 2925: 2923: 2919: 2918: 2916: 2915: 2910: 2905: 2900: 2895: 2883: 2882: 2869: 2860: 2848: 2846: 2836: 2832: 2831: 2829: 2828: 2826:Mycotoxicology 2823: 2818: 2813: 2808: 2803: 2798: 2792: 2790: 2786: 2785: 2776: 2773: 2772: 2761: 2760: 2753: 2746: 2738: 2732: 2731: 2726: 2721: 2716: 2711: 2706: 2701: 2696: 2691: 2686: 2681: 2676: 2671: 2666: 2654: 2649: 2644: 2639: 2634: 2629: 2624: 2619: 2607: 2602: 2597: 2592: 2587: 2582: 2577: 2572: 2567: 2562: 2557: 2550: 2549:External links 2547: 2544: 2543: 2525: 2522:on 2017-07-06. 2507: 2504:on 2023-09-21. 2489: 2486:on 2014-02-02. 2471: 2468:on 2013-04-06. 2453: 2450:on 2014-02-01. 2435: 2417: 2399: 2374: 2349: 2346:on 2015-01-14. 2327: 2309: 2291: 2273: 2258: 2217: 2203: 2191: 2156: 2120: 2103: 2090: 2047: 2033: 2021: 2005: 1999:978-0521812016 1998: 1978: 1933: 1918: 1894: 1880: 1869: 1836: 1817:(2): 307–313. 1811:The Bryologist 1801: 1766: 1752: 1739: 1726:10.1086/274846 1704: 1690: 1676: 1662: 1643:(5): 189–198. 1627: 1614: 1596: 1577: 1564: 1551: 1530: 1502: 1489: 1476: 1470:978-0300195736 1469: 1450: 1449: 1447: 1444: 1441: 1440: 1422: 1421: 1419: 1416: 1415: 1414: 1411:Acharius Medal 1408: 1401: 1398: 1397: 1396: 1390: 1381: 1372: 1366: 1361: 1356: 1351: 1346: 1337: 1331: 1326: 1321: 1316: 1309: 1306: 1304: 1303: 1298: 1296:Francis Wilson 1293: 1288: 1283: 1278: 1273: 1268: 1266:Edmond Tulasne 1263: 1258: 1253: 1248: 1246:Rolf Santesson 1243: 1238: 1233: 1228: 1223: 1218: 1213: 1208: 1203: 1198: 1193: 1188: 1183: 1178: 1172: 1170:Nina Golubkova 1167: 1162: 1157: 1152: 1147: 1142: 1137: 1132: 1127: 1125:Irwin M. Brodo 1122: 1117: 1112: 1107: 1102: 1097: 1092: 1087: 1082: 1077: 1072: 1066: 1064: 1061: 1060: 1059: 1053: 1050:Lichen Biology 1047: 1041: 1035: 1029: 1023: 1017: 1011: 1008: 1005: 998: 992: 982: 976: 970: 964: 958: 952: 946: 940: 934: 928: 917: 911: 905: 899: 893: 887: 881: 871: 868: 862: 857: 854: 785:d-1-usnic acid 701:microbiologist 684: 681: 647:Such a simple 630:Roland Beschel 602:Irwin M. Brodo 568:Chromatophores 555: 552: 507:phytopathology 479:photosynthetic 464:Edmond Tulasne 437:Edmond Tulasne 392: 389: 332: 331: 325: 319: 313: 273:brought in by 205: 204:The beginnings 202: 200: 197: 185:edible lichens 115:Irwin M. Brodo 92:lichenologists 62:cyanobacterium 26: 9: 6: 4: 3: 2: 3112: 3101: 3098: 3096: 3093: 3091: 3088: 3086: 3083: 3081: 3078: 3077: 3075: 3060: 3059: 3050: 3048: 3047: 3038: 3036: 3035: 3026: 3025: 3022: 3016: 3013: 3011: 3008: 3006: 3003: 3001: 2998: 2997: 2995: 2991: 2985: 2982: 2980: 2977: 2975: 2972: 2970: 2967: 2966: 2964: 2960: 2950: 2947: 2945: 2942: 2940: 2937: 2935: 2932: 2930: 2927: 2926: 2924: 2920: 2914: 2911: 2909: 2906: 2904: 2903:Chlamydospore 2901: 2899: 2896: 2894: 2891: 2889: 2885: 2884: 2881: 2877: 2873: 2870: 2868: 2864: 2861: 2859: 2856: 2854: 2850: 2849: 2847: 2845: 2842:Reproduction/ 2840: 2837: 2833: 2827: 2824: 2822: 2819: 2817: 2814: 2812: 2809: 2807: 2804: 2802: 2799: 2797: 2796:Ethnomycology 2794: 2793: 2791: 2787: 2783: 2779: 2774: 2770: 2766: 2759: 2754: 2752: 2747: 2745: 2740: 2739: 2736: 2730: 2727: 2725: 2722: 2720: 2717: 2715: 2712: 2710: 2707: 2705: 2702: 2700: 2697: 2695: 2692: 2690: 2687: 2685: 2682: 2680: 2677: 2675: 2672: 2670: 2667: 2665: 2661: 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Index

Lichenologist

mycology
lichens
symbiotic
organisms
alga
cyanobacterium
filamentous
fungus
mycology
phycology
microbiology
botany
lichenologists
professional
amateur
single-access keys
Irwin M. Brodo
Sylvia Sharnoff
Stephen Sharnoff
Susan Laurie-Bourque
spot test
lichen products
species
fluoresce
ultraviolet light
lichenometry
role of lichens in nutrient cycling
biological soil crusts

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