2274:
2322:
4100:
3609:
3452:
331:. This, combined with the differences in fauna between the two formations, has led some scientists, most notably Scott Sampson, to conclude that there was some barrier separating northern and southern Laramidia at this time. However, preliminary re-calibration of late Cretaceous formation correlations suggests that the upper part of the Kaiparowits, where many of the unique species are found, is actually younger than the Dinosaur Park, and that some Kaiparowits species may simply be the descendants of Dinosaur Park species. However, new dates reveal that this is simply an artifact of inaccurate
1553:
2262:
1609:
425:
1565:
1589:
241:
1541:
1577:
2298:
1601:
1142:
117:
1154:
1130:
2286:
223:
406:. The jungle theory would also support why almost all the animals in the Kaiparowits Formation were new species, and why the deposits were so plentiful. Without the need for herbivores to migrate to find food, and theropods to migrate after herbivores, a whole ecosystem could evolve secluded from interbreeding. The theory also supported why the dinosaurs adorned such features like the 15 horns of
2310:
2577:
quadrate, frontals, parietals, braincase, ectopterygoids, epipterygoid, pterygoids, angulars, surangulars, prearticular, articular, atlas, postaxial cervical vertebrae, cervical ribs, dorsal vertebrae, dorsal ribs, sacral vertebrae, caudal vertebrae, chevrons, portions of ilia, pubes, ischia, complete femur, tibia, fibula, pedal phalanx, and ungual.
319:, with most specimens from the lower half of the formation, but exploration is only comparatively recent, with most work being done since 1982. It has been estimated that less than 10% of the Kaiparowits formation has been explored for fossils. The Natural History Museum of Utah has conducted most fieldwork.
2462:
partial pelvises, tibiae, partial axial column, caudal vertebrae, carpus, antebrachium, femora, partial metatarsals, an incomplete manus, incomplete sets of phalanges, incomplete dorsal ribs, partially damaged cranial material, a nearly complete hind limb, a partial foot, and an isolated articulated
1761:
A mostly complete skull, a partial subadult skull, a partial skull roof and partial braincase, a partially articulated skeleton consisting of a portion of the skull and lower jaws, most of the dorsal, sacral, and caudal vertebral series, fragmentary ribs, scapulae, coracoid, humerus, and the entire
1635:
A partial left mandible, a disarticulated forelimb consisting of a fused scapulocoracoid, humerus, radius, ulna, metacarpals, manual phalanx and an ungual, a scapula, coracoid, cervical vertebrae, caudal vertebrae, dorsal centra vertebrae, sacral vertebrae, chevrons, a complete cervical half ring,
1523:
A complete skull, both mandibles, predentary, dorsal vertebrae, dorsosacral vertebrae, sacral vertebrae, caudosacral vertebrae, caudal vertebrae, dorsal ribs, a complete tail club, both scapulae, coracoid, humerus, ulna, partial ilium, femur, tibia, fibula, phalanx, partial cervical osteoderm half
397:
is today. It is extremely fossil rich, with thousands of plants and animal fossils being preserved in amongst its sandstone and mudstone deposits. Based on plants remains including multiple vines, leaves, and branches, It was assumed by paleontologists Scott
Sampson and his colleagues that Utah in
2576:
A lacrimal, jugal, frontal, squamosal, otoccipital and proötic, otoccipital, basisphenoid, quadrates, jugal, maxilla, dentary, articular, cervical vertebra, scapula, coracoid, humerus, ulna, femur and a subadult specimen consisting of a maxilla, lacrimals, postorbitals, squamosal, quadratojugal,
2631:
Multiple adult specimens consisting of femora, tibiae, fibula, a complete astragalus and calcaneum, two metatarsal III, complete pedal PIII-1 and PIII-2, pedal phalanxes, pedal unguals, caudal vertebrae, isolated fused parietals, a partial dentary, an isolated humerus, an isolated lacrimal, an
4331:
Thomas D. Carr; Thomas E. Williamson; Brooks B. Britt; Ken
Stadtman (2011). "Evidence for high taxonomic and morphologic tyrannosauroid diversity in the Late Cretaceous (Late Campanian) of the American Southwest and a new short-skulled tyrannosaurid from the Kaiparowits formation of Utah".
2093:
partially complete skulls, a partial postorbital consisting of the nearly complete supraorbital horncore, an isolated rostrum, premaxilla, nasal fragment, jugal, squamosal and parietal, an associated post cranium, and two immature specimens consisting of partial postorbitals with complete
2606:
An isolated frontal, isolated caudal vertebra, a proximal tibia, fragmentary metatarsals, pedal phalanges, and pedal unguals, in addition to fragmentary cranial remains, including the basioccipital, fused parietals, portions of both squamosals, and isolated teeth.
2382:
Fragmentary distal left radius, complete left carpus including the semilunate and radiale, and left manus with complete digit I and III, complete digit II, fragmentary distal metatarsals and pedal phalanges, and articulated distal portion of pedal digit II.
2813:
Beveridge, Tegan L.; Roberts, Eric M.; Titus, Alan L. (October 2020). "Volcaniclastic member of the richly fossiliferous
Kaiparowits Formation reveals new insights for regional correlation and tectonics in southern Utah during the latest Campanian".
2427:
Cervical vertebrae, thoracic vertebrae, pygostyle, phalanges, a complete humerus, femur, tarsometatarsus, a partial scapula, coracoid, furcula, tibiotarsus, and fragments of the sternum, radius, ulna, carpometacarpus, and manual phalanges.
2959:
Eaton, Jeffrey G.; Cifelli, Richard L.; Hutchinson, J. Howard; Kirkland, James I.; Parrish, J. Michael (1999). "Cretaceous vertebrate faunas from the
Kaiparowits Plateau, south-central Utah". In Gillete, David D. (ed.).
1731:
Right side of an articulated partial skull, including the ventral lacrimal, jugal, epijugal, quadratojugal, postorbital, maxilla, squamosal, and rostrolateral parietal in the region of the supratemporal fenestra.
4503:
2025:
isolated partial skulls, a partial skeleton consisting of mostly the pelvic region, fragmentary associated elements, and a juvenile specimen consisting of a partial skull and articulated skeleton.
3284:
3834:
3129:
4383:
Titus, Alan L.; Knoll, Katja; Sertich, Joseph J.W.; Yamamura, Daigo; Suarez, Celina A.; Glasspool, Ian J.; Ginouves, Jonathan E.; Lukacic, Abigail K.; Roberts, Eric M. (2021-04-19).
1929:
An almost complete skull, a disarticulated adult skull, an isolated squamosal, a syncervical, fragmentary dorsal vertebrae, associated left forelimb and fragmentary right forelimb.
4577:
4496:
2675:"Revised geochronology, correlation, and dinosaur stratigraphic ranges of the Santonian-Maastrichtian (Late Cretaceous) formations of the Western Interior of North America"
354:
The timeline below follows the re-calibrated timeline of Fowler (2017), showing species from the
Kaiparowits Formation in green, and related species from Alberta in blue.
361:
4385:"Geology and taphonomy of a unique tyrannosaurid bonebed from the upper Campanian Kaiparowits Formation of southern Utah: implications for tyrannosaurid gregariousness"
4244:
2910:"Geology and taphonomy of a unique tyrannosaurid bonebed from the upper Campanian Kaiparowits Formation of southern Utah: implications for tyrannosaurid gregariousness"
3558:
Gates, Terry A.; Lund, Eric K.; Boyd, C. A.; DeBlieux, Donald D.; Titus, Alan L.; Evans, David C.; Getty, Michael A.; Kirkland, James I.; Eaton, Jeffrey G. (2013).
4704:
4489:
1762:
pelvis, and a partial skeleton consisting of dorsal, sacral, and caudal vertebral series, a number of ribs, scapula, humerus, radius, ulna, and a complete pelvis.
358:
3942:
381:
2850:
176:
3560:"Ornithopod Dinosaurs from the Grand Staircase–Escalante National Monument Region, Utah, and Their Role in Paleobiogeographic and Macroevolutionary Studies"
3092:
Hutchison, J. Howard; Michael J. Knell; Donald B. Brinkman (2013). "Turtles from the
Kaiparowits Formation, Utah". In Alan L. Titus, Mark A. Loewen (ed.).
374:
371:
364:
3062:
Herrero, Lucia; Farke, Andrew A (2010). "Hadrosaurid
Dinosaur Skin Impressions from the Upper Cretaceous Kaiparowits Formation of Southern Utah, USA".
4674:
561:
are also known from the
Kaiparowits, including an excellently preserved hadrosaur skin impression known from a recent analysis by Herrero and Farke.
375:
360:
2540:
Dorsal vertebrae, sacral vertebrae, caudal vertebrae, ulna, additional forelimb fragments, a partial pelvis, and partial left and right hind limbs.
363:
4245:
A preliminary report on the theropod dinosaur fauna of the late
Campanian Kaiparowits Formation, Grand Staircase-Escalante National Monument, Utah.
3292:
3981:"Description and rediagnosis of the crested hadrosaurid (Ornithopoda) dinosaur Parasaurolophus cyrtocristatus on the basis of new cranial remains"
3846:
367:
4512:
4050:
Taxonomic revision of latest
Cretaceous North american basal neonithischian taxa and a phylogenetic analysis of basal ornithischian relationships
390:
270:
3313:
2876:
937:
379:
376:
369:
368:
372:
3027:
Zanno, Lindsay E.; Sampson, Scott D. (2005). "A new oviraptorosaur (Theropoda; Maniraptora) from the Late Cretaceous (Campanian) of Utah".
378:
815:
373:
359:
342:
stratigraphic data, the fossil-bearing portion of the Kaiparowits Formation dates from about 77.24 to 75.02 million years ago, with the
3774:
Sampson, S. D.; Loewen, M. A.; Farke, A. A.; Roberts, E. M.; Forster, C. A.; Smith, J. A.; Titus, A. L. (2010). Stepanova, Anna (ed.).
1224:
A new species yet to be described. Reached around 2 meters in length and is known from skull bones and a partial juvenile skeleton.
693:
377:
4140:
Zanno, L. E.; Sampson, S. D. (2005). "A new oviraptorosaur (Theropoda; Maniraptora) from the Late Cretaceous (Campanian) of Utah".
2651:
370:
389:
The Kaiparowits Formation is a muddy bed that was deposited between about 77.3 to 72.8 million years ago, in the area where the
4099:
Zanno, Lindsay E.; Loewen, Mark A.; Farke, Andrew A.; Kim, Gy-Su; Claessens, Leon P. A. M.; McGarrity, Christopher T. (2013).
3835:"Evidence for a second lambeosaur from the Upper Campanian Kaiparowits Formation (Grand Staircase–Escalante National Monument"
2747:"Calibrating the zenith of dinosaur diversity in the Campanian of the Western Interior Basin by CA-ID-TIMS U–Pb geochronology"
3195:
3101:
2969:
2588:
known from immature specimens. It was the largest carnivore in the area, as well as the most common tyrannosaur in the area.
362:
4628:
3724:"A new species of Gryposaurus (Dinosauria: Hadrosauridae) from the late Campanian Kaiparowits Formation, southern Utah, USA"
4669:
4592:
3285:"Crocodyliforms from the late cretaceous of grand staircase–escalante national monument and vicinity, Southern Utah, U.S.A"
4261:
Hartman, Scott; Mortimer, Mickey; Wahl, William R.; Lomax, Dean R.; Lippincott, Jessica; Lovelace, David M. (2019-07-10).
4679:
3347:"Leidyosuchus (Crocodylia: Alligatoroidea) from the Upper Cretaceous Kaiparowits Formation (late Campanian) of Utah, USA"
1980:
disarticulated specimens and articulated specimens, including articulated left and right feet from a single individual.
1660:
An isolated partial skull lacking the left lateral-most portion and much of the anterodorsal and posteroventral palate.
2632:
isolated jugal and a juvenile specimen consisting of fused parietals, a partial unfused frontal, and partial dentary.
4467:
4115:
3958:
3624:
3608:
Loewen, Mark A.; Farke, Andrew A.; Sampson, Scott D.; Getty, Michael A.; Lund, Eric K.; O'Connor, Patrick M. (2013).
3571:
3467:
2862:
2273:
366:
365:
4153:
3040:
4664:
3187:
516:
3130:"A new species of baenid turtle from the Kaiparowits Formation (Upper Cretaceous, Campanian) of southern Utah"
2745:
Ramezani, Jahandar; Beveridge, Tegan L.; Rogers, Raymond R.; Eberth, David A.; Roberts, Eric M. (2022-09-26).
1804:
as the only difference is size and stratigraphy, the Kaiparowits specimens being much larger and more recent.
4699:
4689:
4263:"A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight"
327:
Traditionally, the Kaiparowits Formation has been considered to be roughly equivalent in age to the northern
3880:"A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia"
2321:
4684:
169:
3497:
The evolution and biogeography of ankylosaurid dinosaurs from the late cretaceous of western north america
3451:
Loewen, Mark A.; Burns, Michael E.; Getty, Michael A.; Kirkland, James I.; Vickaryous, Matthew K. (2013).
3658:
4694:
4607:
4186:"The most complete enantiornithine from North America and a phylogenetic analysis of the Avisauridae"
3321:
2884:
3659:
Lindsay E. Zanno; David J. Varricchio; Patrick M. O'Connor; Alan L. Titus; Michael J. Knell (2011).
4567:
1893:
3931:"3.1 Utah, United States; 12. Kaiparowits Formation," in Weishampel, et al. (2004). Pages 579-580.
3453:"Review of Late Cretaceous Ankylosaurian Dinosaurs from the Grand Staircase Region, Southern Utah"
3184:
Non-dinosaurian lower vertebrates across the Cretaceous-Tertiary boundary in northeastern Montana
403:
328:
4633:
2958:
4481:
4640:
4330:
3947:
from the Upper Cretaceous Kaiparowits Formation of Southern Utah based on a series of skulls"
2326:
2261:
2964:. Miscellaneous Publication 99-1. Salt Lake City: Utah Geological Survey. pp. 345–353.
1797:
Several specimens including a virtually complete articulated skull and a fragmentary skull.
1552:
4341:
3787:
3676:
3249:
3141:
2758:
2686:
262:
130:
8:
4602:
4445:
2506:
551:
339:
266:
4453:
4345:
4048:
3791:
3776:"New Horned Dinosaurs from Utah Provide Evidence for Intracontinental Dinosaur Endemism"
3680:
3253:
3145:
2908:
Titus AL, Knoll K, Sertich JJ, Yamamura D, Suarez CA, Glasspool IJ, et al. (2021).
2762:
2690:
4419:
4384:
4365:
4297:
4262:
4212:
4185:
4157:
4015:
3980:
3941:
Gates, Terry; Evans, David C.; Birthisel, Tylor; Bourke, Jason; Zanno, Lindsay (2017).
3904:
3879:
3810:
3775:
3699:
3660:
3495:
3409:
3383:
gen. et sp. nov., from the upper Campanian Kaiparowits Formation of southern Utah, USA"
3378:
3345:
Xu, Heendong A.; Woodward, Samuel J.; Henn, Madison M.; Farke, Andrew A. (2014-01-30).
3265:
3157:
3044:
3009:
2936:
2909:
2787:
2746:
2717:
2674:
1672:
1608:
926:
4183:
1331:
A new genus that exceeded 3 meters in length. Has a thin snout suited for piscivory.
1124:
proper, but a close cousin. It would extend the family's time range to the Campanian.
4612:
4463:
4424:
4406:
4357:
4302:
4284:
4217:
4161:
4111:
4020:
4002:
3909:
3815:
3740:
3723:
3704:
3620:
3567:
3559:
3463:
3414:
3269:
3213:
3191:
3097:
3048:
3013:
2992:(Dinosauria: Hadrosauridae) from the Upper Campanian Kaiparowits Formation of Utah".
2965:
2941:
2831:
2792:
2774:
2722:
2704:
2489:
2302:
1531:
4369:
3833:
Armour Smith, Elliot; Sertich, Joseph; Alger-Meyer, Evan; Sartin, Catherine (2018).
3665:
gen. et sp. nov., from the Upper Cretaceous Western Interior Basin of North America"
3161:
424:
4562:
4414:
4396:
4349:
4292:
4274:
4207:
4197:
4149:
4010:
3992:
3899:
3891:
3805:
3795:
3735:
3694:
3684:
3404:
3394:
3261:
3257:
3149:
3036:
3005:
3001:
2931:
2921:
2823:
2782:
2766:
2712:
2694:
2510:
1189:
957:
547:
525:
436:
413:
286:
3153:
2827:
4582:
4531:
4457:
3800:
3689:
2699:
2636:
2619:
2436:
2352:
2336:
2254:
2251:
2007:
1593:
1564:
494:
343:
4243:
Zanno, L.E., Weirsma, J.P., Loewen, M.A., Sampson, S.D. and Getty, M.A. (2010).
4587:
2770:
2582:
2520:
2446:
2388:
2314:
1907:
1876:
1826:
1717:
1693:
1645:
1623:
1581:
1545:
1498:
864:
445:
4353:
3186:. Volume 134 of University of California publications in geological sciences.
2877:"FIRST REPORT OF THE HYPER-GIANT CRETACEOUS CROCODYLIAN DEINOSUCHUS FROM UTAH"
1735:
Might be a distinct taxon based on its episquamosal and squamosal morphology.
1588:
4658:
4410:
4288:
4006:
2835:
2778:
2708:
2585:
2556:
2396:
2266:
2234:
1997:
1991:
1934:
1860:
1855:
A nearly complete skull and a disarticulated skull of a subadult individual.
1836:
1809:
1569:
1464:
1201:
1146:
487:
430:
408:
308:
278:
246:
192:
178:
68:
27:
3346:
4572:
4546:
4526:
4449:
4428:
4361:
4306:
4221:
4024:
3913:
3895:
3819:
3708:
3418:
3091:
2945:
2796:
2726:
2611:
2594:
2545:
2391:
2074:
2043:
1960:
1943:
1770:
1767:
1613:
1339:
1134:
1120:
1095:
1061:
727:
558:
508:
503:
498:
453:
346:
Upper Valley Member estimated to date from 73.8 to 72.8 million years ago.
332:
1576:
1540:
4541:
4536:
2433:
2297:
2166:
2064:
2061:
1968:
1742:
1600:
1557:
1430:
1257:
1158:
1141:
1027:
788:
756:
625:
538:
533:
512:
482:
449:
259:
43:
4401:
3997:
2926:
1249:
Known from a lower jaw fragment. Is the oldest known true caiman found.
116:
4279:
4202:
3564:
At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah
3399:
3289:
At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah
3094:
At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah
2152:
1985:
1863:
1416:
1153:
1013:
611:
477:
465:
293:
88:
53:
4108:
At the Top of the Grand Staircase-The Late Cretaceous of Southern Utah
3979:
Gates, Terry A.; Evans, David C.; Sertich, Joseph J. W. (2021-01-25).
3617:
At the Top of the Grand Staircase-The Late Cretaceous of Southern Utah
3460:
At the Top of the Grand Staircase-The Late Cretaceous of Southern Utah
929:. By far the largest native turtle at about 78 centimeters in length.
2903:
2901:
2477:
2362:
2278:
2098:
A chasmosaurine ceratopsid known from adult and juvenile specimens.
2057:
A partial disarticulated skull and fragmentary postcranial material.
2033:
then believed to be a new species, then tentatively referred back to
1867:
1773:
904:
842:
529:
457:
399:
305:
301:
282:
157:
93:
37:
31:
3832:
1639:
An indeterminate ankylosaurid known from various partial specimens.
4184:
Jessie Atterholt; J. Howard Hutchison; Jingmai K. O’Connor (2018).
2200:
1688:
The youngest example of brachylophosaurin material from Laramidia.
894:
715:
490:
312:
290:
153:
83:
78:
63:
58:
48:
3878:
Sampson, SD; Lund, EK; Loewen, MA; Farke, AA; Clayton, KE (2013).
2898:
1129:
2408:
2290:
2285:
2159:
1636:
dorsal ribs, a partial tail club, numerous osteoderms and teeth.
1423:
1020:
659:
618:
98:
73:
3610:"Ceratopsid Dinosaurs from the Grand Staircase of Southern Utah"
3376:
4511:
1280:
A very large alligatoroid, almost or over 10 meters in length.
874:
521:
473:
469:
316:
357:
2744:
2130:
1394:
991:
589:
297:
121:
Fluvial deposits of the Kaiparowits Formation at "The Blues".
4462:, Berkeley: University of California Press, pp. 1–880,
1709:
Also known as "Kaiparowits Centrosaurine B." Different from
4154:
10.1671/0272-4634(2005)025[0897:anotmf]2.0.co;2
3041:
10.1671/0272-4634(2005)025[0897:ANOTMF]2.0.CO;2
461:
394:
274:
228:
4249:
Learning from the Land Symposium: Geology and Paleontology
1825:
An indeterminate hadrosaurine, initially interpreted as a
4382:
3940:
3773:
2907:
4444:
4260:
3450:
3314:"First Report of the Hyper-Giant Cretaceous Crocodylian
2309:
3607:
2988:
Gates, Terry; Sampson, Scott (2006). "A new species of
536:. Recent finds include large specimens of the duckbill
3877:
3235:
3233:
3231:
3229:
3227:
3225:
3223:
3221:
3026:
2348:
An isolated pedal phalanx PII-I and isolated unguals.
1780:
is the most common hadrosaur found in this formation.
4101:"Late Cretaceous Theropod Dinosaurs of Southern Utah"
3557:
3209:
3207:
3182:
Bryant, Laurie J. (1989). "Systematic Paleontology".
2812:
1663:
Described in a thesis, different from the other two
748:
A baenid, notable for its atypical nasal structure.
4098:
3722:Gates, Terry A.; Sampson, Scott D. (October 2007).
3344:
3340:
3338:
3218:
2987:
1822:An articulated tail, partial pelvis, and left leg.
3214:http://archive.li.suu.edu/docs/ms130/AR/eatonj.pdf
3204:
1956:A cervical spine, numerous osteoderms, and teeth.
3978:
3087:
3085:
3083:
3081:
3079:
3077:
1983:One specimen shares similar foot morphology with
1706:A partial skull and an isolated right squamosal.
4656:
4239:
4237:
4235:
4233:
4231:
3873:
3871:
3869:
3867:
3865:
3863:
3335:
773:middle unit of the upper Kaiparowits Formation
4110:. Indiana University Press. pp. 504–525.
3769:
3767:
3765:
3763:
3761:
3759:
3757:
3755:
3753:
3751:
3619:. Indiana University Press. pp. 488–503.
3566:. Indiana University Press. pp. 463–481.
3489:
3487:
3485:
3483:
3481:
3479:
3462:. Indiana University Press. pp. 445–462.
3372:
3370:
3368:
3366:
3364:
3096:. Indiana University Press. pp. 295–318.
3074:
1937:ceratopsid with rounded horns above its eyes.
1302:A very large alligatoroid, similar in size to
281:. It is over 2800 feet (850 m) thick, and is
4497:
4228:
4106:. In Titus, Alan L.; Loewen, Mark A. (eds.).
3860:
3615:. In Titus, Alan L.; Loewen, Mark A. (eds.).
3562:. In Titus, Alan L.; Loewen, Mark A. (eds.).
3458:. In Titus, Alan L.; Loewen, Mark A. (eds.).
3446:
3444:
3442:
3440:
3438:
3436:
3434:
3432:
3430:
3428:
3177:
3175:
3173:
3171:
3064:PalArch's Journal of Vertebrate Palaeontology
2513:dromaeosaurid represented by isolated teeth.
4251:. Washington, DC: Bureau of Land Management.
4139:
4135:
4133:
4131:
4129:
4127:
4053:(Thesis). The University of Texas at Austin.
3721:
3061:
2195:
2125:
1459:
1389:
1362:A new species known from a partial skeleton
1056:
986:
654:
584:
4705:Grand Staircase–Escalante National Monument
4513:Grand Staircase–Escalante National Monument
4326:
4324:
4322:
4320:
4318:
4316:
4042:
4040:
4038:
4036:
4034:
3748:
3476:
3377:Jelle P. Wiersma; Randall B. Irmis (2018).
3361:
391:Grand Staircase–Escalante National Monument
315:beds represent floodplain deposits. It is
271:Grand Staircase–Escalante National Monument
4504:
4490:
4179:
4177:
4175:
4173:
4171:
3425:
3168:
1524:rings, and dorsal and lateral osteoderms.
115:
4418:
4400:
4296:
4278:
4211:
4201:
4124:
4014:
3996:
3903:
3809:
3799:
3739:
3728:Zoological Journal of the Linnean Society
3698:
3688:
3408:
3398:
3379:"A new southern Laramidian ankylosaurid,
3123:
3121:
3119:
3117:
3115:
3113:
2935:
2925:
2786:
2716:
2698:
546:, and the first described remains of the
4675:Upper Cretaceous Series of North America
4313:
4031:
3654:
3652:
3650:
3648:
3646:
3644:
3642:
3640:
3638:
3636:
2652:List of dinosaur-bearing rock formations
2320:
2308:
2296:
2284:
2272:
2260:
2198:
1607:
1599:
1587:
1575:
1563:
1551:
1539:
1462:
1152:
1140:
1128:
1059:
657:
423:
335:, and both formations had similar ages.
4168:
3927:
3925:
3923:
3493:
2203:reported from the Kaiparowits Formation
1467:reported from the Kaiparowits Formation
1064:reported from the Kaiparowits Formation
662:reported from the Kaiparowits Formation
434:disturbed from its rest by a wandering
4657:
4094:
4092:
4090:
4088:
4086:
4084:
4082:
4080:
3603:
3553:
3551:
3549:
3547:
3545:
3543:
3541:
3539:
3537:
3535:
3533:
3531:
3529:
3527:
3181:
3127:
3110:
2983:
2981:
2952:
2848:
2672:
1529:A species of ankylosaurine related to
4485:
4078:
4076:
4074:
4072:
4070:
4068:
4066:
4064:
4062:
4060:
3633:
3601:
3599:
3597:
3595:
3593:
3591:
3589:
3587:
3585:
3583:
3525:
3523:
3521:
3519:
3517:
3515:
3513:
3511:
3509:
3507:
3282:
2842:
2673:Fowler, Denver Warwick (2017-11-22).
4046:
3920:
2808:
2806:
2740:
2738:
2736:
2668:
2666:
2257:only represented by isolated teeth.
3957:(Supplement 1): 117. Archived from
3020:
2978:
2463:foot with an associated limb bone.
2182:Uncertain or tentative taxa are in
1446:Uncertain or tentative taxa are in
1043:Uncertain or tentative taxa are in
641:Uncertain or tentative taxa are in
16:Rock formation in the United States
13:
4142:Journal of Vertebrate Paleontology
4057:
3951:Journal of Vertebrate Paleontology
3839:Journal of Vertebrate Paleontology
3580:
3504:
3283:Irmis, Randall B. (January 2013).
3242:Journal of Vertebrate Paleontology
3134:Journal of Vertebrate Paleontology
3029:Journal of Vertebrate Paleontology
2994:Journal of Vertebrate Paleontology
2570:"The Blues"; UMNH VP Locality 597
2439:represented by an adult specimen.
349:
14:
4716:
2803:
2733:
2663:
2142:Taxon falsely reported as present
1406:Taxon falsely reported as present
1369:
1003:Taxon falsely reported as present
882:indet., possibly several species
601:Taxon falsely reported as present
402:was a dense jungle bordering the
3741:10.1111/j.1096-3642.2007.00349.x
2548:known from a subadult specimen.
1513:Horse Mountain Gryposaur Quarry
239:
221:
41:
4438:
4376:
4254:
3972:
3934:
3826:
3715:
3306:
3276:
3055:
2962:Vertebrate Paleontology in Utah
2147:Dubious taxon or junior synonym
1411:Dubious taxon or junior synonym
1008:Dubious taxon or junior synonym
606:Dubious taxon or junior synonym
311:are the deposit of rivers, and
4629:DomĂnguez–Escalante expedition
3494:Wiersma, Jelle Pouwel (2016).
3262:10.1080/02724634.2004.10010643
3188:University of California Press
3006:10.1080/02724634.2006.10010069
2869:
966:
517:Parasaurolophus cyrtocristatus
1:
3500:(Thesis). University of Utah.
3154:10.1080/02724634.2015.1009084
2828:10.1016/j.cretres.2020.104527
2657:
419:
3801:10.1371/journal.pone.0012292
3690:10.1371/journal.pone.0024487
3661:"A new troodontid theropod,
2700:10.1371/journal.pone.0188426
2105:
7:
4670:Geologic formations of Utah
4459:The Dinosauria, 2nd edition
4047:Boyd, Clint Aaroen (2012).
2645:
2109:
1538:
1373:
970:
568:
10:
4721:
4680:Cretaceous geology of Utah
3128:Lively, Joshua R. (2015).
2861:(5): 60–79. Archived from
2851:"Digging Utah's Dinosaurs"
2771:10.1038/s41598-022-19896-w
2467:Probably not referable to
2259:
1127:
720:
564:
384:
289:formation was formed from
273:, in the southern part of
140:790 m (2,600 ft)
4621:
4608:Straight Cliffs Formation
4555:
4519:
4354:10.1007/s00114-011-0762-7
3291:: 424–444. Archived from
2116:
2112:
2029:Originally identified as
1739:
1380:
1376:
1254:
977:
973:
575:
571:
520:, and a variety of early
234:
216:
168:
163:
149:
144:
136:
126:
114:
26:
21:
4568:Canyons of the Escalante
2114:
2094:supraorbital horncores.
1894:Hypacrosaurus altispinus
1378:
975:
573:
542:, including the species
444:Animals present include
3381:Akainacephalus johnsoni
3240:"Abstracts of Papers".
2849:Miller, P. (May 2014).
2218:Stratigraphic position
2067:described in a thesis.
1482:Stratigraphic position
1079:Stratigraphic position
677:Stratigraphic position
404:Western Interior Seaway
329:Dinosaur Park Formation
4634:Hole in the Rock Trail
3896:10.1098/rspb.2013.1186
2537:Middle of middle unit
2379:Middle of middle unit
2330:
2318:
2306:
2294:
2282:
2270:
2190:taxa are discredited.
1888:Two partial skeletons
1654:UMNH VP locality 1506
1617:
1605:
1597:
1585:
1573:
1561:
1549:
1454:taxa are discredited.
1187:Is similar in form to
1162:
1150:
1138:
1051:taxa are discredited.
649:taxa are discredited.
441:
322:
193:37.40000°N 111.68333°W
4665:Kaiparowits Formation
4641:Xylorhiza cronquistii
4598:Kaiparowits Formation
2473:"Ornithomimus" sedens
2471:. Closely related to
2422:UCMP locality V93097
2394:, closely related to
2327:Teratophoneus curriei
2324:
2312:
2300:
2288:
2276:
2264:
2090:Lower to middle unit
2054:Lower to middle unit
2022:Lower to middle unit
1921:UMNH VP Locality 940
1611:
1603:
1591:
1579:
1567:
1555:
1543:
1156:
1144:
1132:
925:A possible member of
427:
256:Kaiparowits Formation
204:Kaiparowits Formation
22:Kaiparowits Formation
4700:Paleontology in Utah
4690:Sandstone formations
4446:Weishampel, David B.
3845:: 83. Archived from
2865:on December 7, 2014.
1870:on the hind margin.
1800:May be referable to
300:in coastal southern
198:37.40000; -111.68333
131:Geological formation
4685:Mudstone formations
4603:Kaiparowits Plateau
4402:10.7717/peerj.11013
4346:2011NW.....98..241C
4334:Naturwissenschaften
3998:10.7717/peerj.10669
3792:2010PLoSO...512292S
3681:2011PLoSO...624487Z
3254:2004JVPal..24S...1.
3146:2015JVPal..35E9084L
2927:10.7717/peerj.11013
2855:National Geographic
2816:Cretaceous Research
2763:2022NatSR..1216026R
2691:2017PLoSO..1288426F
1866:with ten hook-like
1685:An isolated jugal.
552:Hagryphus giganteus
448:(sharks and rays),
267:Kaiparowits Plateau
188: /
28:Stratigraphic range
4280:10.7717/peerj.7247
4203:10.7717/peerj.5910
3943:"A new species of
3890:(1766): 20131186.
3400:10.7717/peerj.5016
3190:. pp. 10–58.
2751:Scientific Reports
2507:saurornitholestine
2331:
2319:
2307:
2295:
2283:
2271:
2137:Reclassified taxon
2035:P. cyrtocristatus.
2031:P. cyrtocristatus,
1925:Lower middle unit
1852:Lower middle unit
1673:Brachylophosaurini
1657:Lower middle unit
1618:
1606:
1598:
1586:
1574:
1562:
1550:
1518:Lower middle unit
1401:Reclassified taxon
1163:
1151:
1139:
998:Reclassified taxon
927:Nanhsiungchelyidae
596:Reclassified taxon
504:Ornithomimus velox
442:
4652:
4651:
4613:Wahweap Formation
4454:OsmĂłlska, Halszka
3197:978-0-520-09735-3
3103:978-0-253-00896-1
3000:(3, Suppl): 65A.
2971:978-1-55791-634-1
2643:
2642:
2490:Saurornitholestes
2303:Saurornitholestes
2194:
2193:
2185:
2173:
2172:
2169:
2162:
2155:
2148:
2143:
2138:
2133:
2103:
2102:
2015:P. cyrtocristatus
1897:; different from
1532:Nodocephalosaurus
1458:
1457:
1449:
1437:
1436:
1433:
1426:
1419:
1412:
1407:
1402:
1397:
1367:
1366:
1055:
1054:
1046:
1034:
1033:
1030:
1023:
1016:
1009:
1004:
999:
994:
964:
963:
653:
652:
644:
632:
631:
628:
621:
614:
607:
602:
597:
592:
548:oviraptorosaurian
526:multituberculates
509:armored dinosaurs
338:According to new
252:
251:
4712:
4695:Fluvial deposits
4645:
4563:Aquarius Plateau
4506:
4499:
4492:
4483:
4482:
4478:
4477:
4476:
4433:
4432:
4422:
4404:
4380:
4374:
4373:
4328:
4311:
4310:
4300:
4282:
4258:
4252:
4241:
4226:
4225:
4215:
4205:
4181:
4166:
4165:
4137:
4122:
4121:
4105:
4096:
4055:
4054:
4044:
4029:
4028:
4018:
4000:
3976:
3970:
3969:
3967:
3966:
3938:
3932:
3929:
3918:
3917:
3907:
3875:
3858:
3857:
3855:
3854:
3830:
3824:
3823:
3813:
3803:
3771:
3746:
3745:
3743:
3719:
3713:
3712:
3702:
3692:
3656:
3631:
3630:
3614:
3605:
3578:
3577:
3555:
3502:
3501:
3491:
3474:
3473:
3457:
3448:
3423:
3422:
3412:
3402:
3374:
3359:
3358:
3342:
3333:
3332:
3330:
3329:
3320:. Archived from
3310:
3304:
3303:
3301:
3300:
3280:
3274:
3273:
3237:
3216:
3211:
3202:
3201:
3179:
3166:
3165:
3125:
3108:
3107:
3089:
3072:
3071:
3059:
3053:
3052:
3024:
3018:
3017:
2985:
2976:
2975:
2956:
2950:
2949:
2939:
2929:
2905:
2896:
2895:
2893:
2892:
2883:. Archived from
2873:
2867:
2866:
2846:
2840:
2839:
2810:
2801:
2800:
2790:
2742:
2731:
2730:
2720:
2702:
2685:(11): e0188426.
2670:
2196:
2183:
2165:
2158:
2151:
2146:
2141:
2136:
2129:
2126:
2110:
1751:G. monumentensis
1460:
1447:
1429:
1422:
1415:
1410:
1405:
1400:
1393:
1390:
1374:
1289:D. riograndensis
1190:Allognathosuchus
1057:
1044:
1026:
1019:
1012:
1007:
1002:
997:
990:
987:
971:
958:softshell turtle
655:
642:
624:
617:
610:
605:
600:
595:
588:
585:
569:
544:G. monumentensis
414:sexual selection
412:, they were for
287:Upper Cretaceous
245:
243:
242:
227:
225:
224:
212:
211:
209:
208:
207:
205:
200:
199:
194:
189:
186:
185:
184:
181:
119:
110:
40:
19:
18:
4720:
4719:
4715:
4714:
4713:
4711:
4710:
4709:
4655:
4654:
4653:
4648:
4644:(plant species)
4638:
4617:
4583:Grand Staircase
4551:
4532:Escalante River
4520:Bodies of water
4515:
4510:
4474:
4472:
4470:
4441:
4436:
4381:
4377:
4329:
4314:
4259:
4255:
4242:
4229:
4182:
4169:
4138:
4125:
4118:
4103:
4097:
4058:
4045:
4032:
3977:
3973:
3964:
3962:
3945:Parasaurolophus
3939:
3935:
3930:
3921:
3876:
3861:
3852:
3850:
3831:
3827:
3772:
3749:
3720:
3716:
3657:
3634:
3627:
3612:
3606:
3581:
3574:
3556:
3505:
3492:
3477:
3470:
3455:
3449:
3426:
3375:
3362:
3343:
3336:
3327:
3325:
3312:
3311:
3307:
3298:
3296:
3281:
3277:
3248:: 1–139. 2004.
3239:
3238:
3219:
3212:
3205:
3198:
3180:
3169:
3140:(6): e1009084.
3126:
3111:
3104:
3090:
3075:
3060:
3056:
3025:
3021:
2986:
2979:
2972:
2957:
2953:
2906:
2899:
2890:
2888:
2875:
2874:
2870:
2847:
2843:
2811:
2804:
2743:
2734:
2671:
2664:
2660:
2648:
2620:Tyrannosauridae
2511:velociraptorine
2502:Isolated teeth
2437:enantiornithine
2337:Dromaeosauridae
2247:Isolated teeth
2181:
2124:
2108:
2008:Parasaurolophus
1899:Parasaurolophus
1711:Nasutoceratops.
1594:Parasaurolophus
1445:
1388:
1372:
1042:
985:
969:
865:paracryptodiran
640:
583:
567:
446:chondrichthyans
422:
387:
382:
380:
352:
350:Biostratigraphy
325:
277:in the western
240:
238:
222:
220:
203:
201:
197:
195:
191:
190:
187:
182:
179:
177:
175:
174:
122:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
36:77.3–72.8
35:
34:
17:
12:
11:
5:
4718:
4708:
4707:
4702:
4697:
4692:
4687:
4682:
4677:
4672:
4667:
4650:
4649:
4647:
4646:
4636:
4631:
4625:
4623:
4619:
4618:
4616:
4615:
4610:
4605:
4600:
4595:
4590:
4588:Grosvenor Arch
4585:
4580:
4575:
4570:
4565:
4559:
4557:
4553:
4552:
4550:
4549:
4544:
4539:
4534:
4529:
4523:
4521:
4517:
4516:
4509:
4508:
4501:
4494:
4486:
4480:
4479:
4468:
4440:
4437:
4435:
4434:
4375:
4340:(3): 241–246.
4312:
4253:
4227:
4167:
4148:(4): 897–904.
4123:
4116:
4056:
4030:
3971:
3933:
3919:
3859:
3825:
3747:
3734:(2): 351–376.
3714:
3663:Talos sampsoni
3632:
3625:
3579:
3572:
3503:
3475:
3468:
3424:
3360:
3334:
3305:
3275:
3217:
3203:
3196:
3167:
3109:
3102:
3073:
3054:
3035:(4): 897–904.
3019:
2977:
2970:
2951:
2897:
2881:gsa.confex.com
2868:
2841:
2802:
2732:
2661:
2659:
2656:
2655:
2654:
2647:
2644:
2641:
2640:
2635:Indeterminate
2633:
2629:
2627:
2625:
2624:Indeterminate
2622:
2616:
2615:
2610:Indeterminate
2608:
2604:
2602:
2600:
2599:Indeterminate
2597:
2591:
2590:
2583:tyrannosaurine
2578:
2574:
2571:
2568:
2561:
2551:
2550:
2541:
2538:
2535:
2532:
2525:
2515:
2514:
2503:
2500:
2498:
2496:
2495:Indeterminate
2493:
2484:
2483:
2464:
2460:
2457:
2455:
2453:Indeterminate
2450:
2447:Ornithomimidae
2442:
2441:
2429:
2425:
2423:
2420:
2413:
2403:
2402:
2384:
2380:
2377:
2374:
2367:
2357:
2356:
2351:Indeterminate
2349:
2346:
2344:
2342:
2341:Indeterminate
2339:
2333:
2332:
2315:Talos sampsoni
2258:
2252:dromaeosaurine
2248:
2245:
2243:
2241:
2240:Indeterminate
2238:
2229:
2228:
2225:
2222:
2219:
2216:
2213:
2210:
2206:
2205:
2192:
2191:
2176:
2174:
2171:
2170:
2163:
2156:
2149:
2144:
2139:
2134:
2118:
2117:
2115:
2113:
2107:
2104:
2101:
2100:
2095:
2091:
2088:
2086:
2079:
2069:
2068:
2058:
2055:
2052:
2050:
2049:"S. goldenii"
2047:
2039:
2038:
2026:
2023:
2020:
2018:
2011:
2003:
2002:
1981:
1978:
1976:
1974:
1973:Indeterminate
1971:
1965:
1964:
1959:Indeterminate
1957:
1954:
1951:
1949:
1948:Indeterminate
1946:
1940:
1939:
1930:
1927:
1922:
1919:
1912:
1908:Nasutoceratops
1902:
1901:
1889:
1886:
1884:
1882:
1881:Indeterminate
1879:
1877:Lambeosaurinae
1873:
1872:
1856:
1853:
1850:
1848:
1845:K. richardsoni
1841:
1831:
1830:
1823:
1820:
1817:
1815:
1814:Indeterminate
1812:
1806:
1805:
1798:
1795:
1792:
1790:
1783:
1782:
1763:
1759:
1756:
1754:
1747:
1737:
1736:
1733:
1729:
1726:
1723:
1722:Indeterminate
1720:
1718:Chasmosaurinae
1714:
1713:
1707:
1704:
1701:
1699:
1698:Indeterminate
1696:
1694:Centrosaurinae
1690:
1689:
1686:
1683:
1680:
1678:
1677:Indeterminate
1675:
1669:
1668:
1665:Anodontosaurus
1661:
1658:
1655:
1652:
1649:
1646:Anodontosaurus
1641:
1640:
1637:
1633:
1631:
1629:
1628:Indeterminate
1626:
1624:Ankylosauridae
1620:
1619:
1604:"Skaladromeus"
1582:Nasutoceratops
1546:Akainacephalus
1537:
1526:
1520:
1515:
1510:
1503:
1499:Akainacephalus
1493:
1492:
1489:
1486:
1483:
1480:
1477:
1474:
1470:
1469:
1465:Ornithischians
1456:
1455:
1440:
1438:
1435:
1434:
1427:
1420:
1413:
1408:
1403:
1398:
1382:
1381:
1379:
1377:
1371:
1370:Ornithischians
1368:
1365:
1364:
1359:
1357:
1355:
1353:
1352:
1351:
1344:
1334:
1333:
1328:
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1315:
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1299:
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1238:
1233:
1227:
1226:
1221:
1219:
1217:
1215:
1214:
1213:
1206:
1196:
1195:
1184:
1182:
1180:
1178:
1177:
1176:
1171:
1165:
1164:
1126:
1115:
1113:
1111:
1109:
1108:
1107:
1100:
1090:
1089:
1086:
1083:
1080:
1077:
1074:
1071:
1067:
1066:
1053:
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1037:
1035:
1032:
1031:
1024:
1017:
1010:
1005:
1000:
995:
979:
978:
976:
974:
968:
965:
962:
961:
953:
951:
949:
947:
942:
932:
931:
922:
920:
918:
916:
909:
899:
898:
890:
888:
886:
884:
879:
869:
868:
860:
858:
856:
854:
847:
837:
836:
831:
829:
827:
825:
820:
810:
809:
804:
802:
800:
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793:
783:
782:
777:
775:
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768:
761:
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750:
745:
743:
741:
739:
732:
722:
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719:
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688:
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684:
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650:
635:
633:
630:
629:
622:
615:
608:
603:
598:
593:
577:
576:
574:
572:
566:
563:
495:dromaeosaurids
488:coelurosaurian
421:
418:
386:
383:
356:
351:
348:
344:volcaniclastic
324:
321:
285:in age. This
263:rock formation
250:
249:
236:
232:
231:
218:
214:
213:
172:
166:
165:
161:
160:
151:
147:
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97:
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62:
57:
52:
47:
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24:
23:
15:
9:
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4:
3:
2:
4717:
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4578:Devils Garden
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4487:
4484:
4471:
4469:0-520-24209-2
4465:
4461:
4460:
4455:
4451:
4450:Dodson, Peter
4447:
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4430:
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4119:
4117:9780253008961
4113:
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4026:
4022:
4017:
4012:
4008:
4004:
3999:
3994:
3990:
3986:
3982:
3975:
3961:on 2017-11-07
3960:
3956:
3952:
3948:
3946:
3937:
3928:
3926:
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3915:
3911:
3906:
3901:
3897:
3893:
3889:
3885:
3881:
3874:
3872:
3870:
3868:
3866:
3864:
3849:on 2018-10-22
3848:
3844:
3840:
3836:
3829:
3821:
3817:
3812:
3807:
3802:
3797:
3793:
3789:
3786:(9): e12292.
3785:
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3696:
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3682:
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3675:(9): e24487.
3674:
3670:
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3655:
3653:
3651:
3649:
3647:
3645:
3643:
3641:
3639:
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3628:
3626:9780253008961
3622:
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3573:9780253008961
3569:
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3499:
3498:
3490:
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3480:
3471:
3469:9780253008961
3465:
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3406:
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3384:
3382:
3373:
3371:
3369:
3367:
3365:
3356:
3352:
3348:
3341:
3339:
3324:on 2017-07-29
3323:
3319:
3317:
3309:
3295:on 2017-11-07
3294:
3290:
3286:
3279:
3271:
3267:
3263:
3259:
3255:
3251:
3247:
3243:
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3208:
3199:
3193:
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3105:
3099:
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3080:
3078:
3069:
3065:
3058:
3050:
3046:
3042:
3038:
3034:
3030:
3023:
3015:
3011:
3007:
3003:
2999:
2995:
2991:
2984:
2982:
2973:
2967:
2963:
2955:
2947:
2943:
2938:
2933:
2928:
2923:
2919:
2915:
2911:
2904:
2902:
2887:on 2017-07-29
2886:
2882:
2878:
2872:
2864:
2860:
2856:
2852:
2845:
2837:
2833:
2829:
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2728:
2724:
2719:
2714:
2710:
2706:
2701:
2696:
2692:
2688:
2684:
2680:
2676:
2669:
2667:
2662:
2653:
2650:
2649:
2638:
2637:tyrannosaurid
2634:
2630:
2628:
2626:
2623:
2621:
2618:
2617:
2613:
2609:
2605:
2603:
2601:
2598:
2596:
2593:
2592:
2589:
2587:
2586:tyrannosaurid
2584:
2579:
2575:
2572:
2569:
2567:
2566:
2562:
2560:
2559:
2558:
2557:Teratophoneus
2553:
2552:
2549:
2547:
2542:
2539:
2536:
2533:
2531:
2530:
2526:
2524:
2523:
2522:
2517:
2516:
2512:
2508:
2504:
2501:
2499:
2497:
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2492:
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2470:
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2430:
2426:
2424:
2421:
2419:
2418:
2414:
2412:
2411:
2410:
2405:
2404:
2401:
2399:
2398:
2397:Chirostenotes
2393:
2390:
2389:caenagnathine
2385:
2381:
2378:
2375:
2373:
2372:
2368:
2366:
2365:
2364:
2359:
2358:
2354:
2353:dromaeosaurid
2350:
2347:
2345:
2343:
2340:
2338:
2335:
2334:
2329:
2328:
2323:
2317:
2316:
2311:
2305:
2304:
2299:
2293:
2292:
2287:
2281:
2280:
2275:
2269:
2268:
2267:Dromaeosaurus
2263:
2256:
2255:dromaeosaurid
2253:
2249:
2246:
2244:
2242:
2239:
2237:
2236:
2235:Dromaeosaurus
2231:
2230:
2226:
2223:
2220:
2217:
2214:
2211:
2208:
2207:
2204:
2202:
2197:
2189:
2180:
2177:
2175:
2168:
2164:
2161:
2157:
2154:
2150:
2145:
2140:
2135:
2132:
2128:
2127:
2123:
2120:
2119:
2111:
2099:
2096:
2092:
2089:
2087:
2085:
2084:
2080:
2078:
2077:
2076:
2071:
2070:
2066:
2063:
2059:
2056:
2053:
2051:
2048:
2045:
2041:
2040:
2037:
2036:
2032:
2027:
2024:
2021:
2019:
2017:
2016:
2012:
2010:
2009:
2005:
2004:
2000:
1999:
1998:Zephyrosaurus
1994:
1993:
1992:Oryctodromeus
1988:
1987:
1982:
1979:
1977:
1975:
1972:
1970:
1967:
1966:
1962:
1958:
1955:
1952:
1950:
1947:
1945:
1942:
1941:
1938:
1936:
1935:centrosaurine
1931:
1928:
1926:
1923:
1920:
1918:
1917:
1913:
1911:
1910:
1909:
1904:
1903:
1900:
1896:
1895:
1890:
1887:
1885:
1883:
1880:
1878:
1875:
1874:
1871:
1869:
1865:
1862:
1861:chasmosaurine
1857:
1854:
1851:
1849:
1847:
1846:
1842:
1840:
1839:
1838:
1837:Kosmoceratops
1833:
1832:
1828:
1827:lambeosaurine
1824:
1821:
1818:
1816:
1813:
1811:
1810:Hadrosaurinae
1808:
1807:
1803:
1799:
1796:
1793:
1791:
1788:
1785:
1784:
1781:
1779:
1775:
1772:
1769:
1764:
1760:
1757:
1755:
1753:
1752:
1748:
1746:
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1744:
1738:
1734:
1730:
1727:
1724:
1721:
1719:
1716:
1715:
1712:
1708:
1705:
1702:
1700:
1697:
1695:
1692:
1691:
1687:
1684:
1681:
1679:
1676:
1674:
1671:
1670:
1666:
1662:
1659:
1656:
1653:
1650:
1648:
1647:
1643:
1642:
1638:
1634:
1632:
1630:
1627:
1625:
1622:
1621:
1616:
1615:
1610:
1602:
1596:
1595:
1590:
1584:
1583:
1578:
1572:
1571:
1570:Kosmoceratops
1566:
1560:
1559:
1554:
1548:
1547:
1542:
1536:
1534:
1533:
1527:
1525:
1521:
1519:
1516:
1514:
1511:
1509:
1508:
1504:
1502:
1501:
1500:
1495:
1494:
1490:
1487:
1484:
1481:
1478:
1475:
1472:
1471:
1468:
1466:
1461:
1453:
1444:
1441:
1439:
1432:
1428:
1425:
1421:
1418:
1414:
1409:
1404:
1399:
1396:
1392:
1391:
1387:
1384:
1383:
1375:
1363:
1360:
1358:
1356:
1354:
1350:
1347:
1346:
1345:
1343:
1342:
1341:
1336:
1335:
1332:
1329:
1327:
1325:
1323:
1318:
1317:
1316:
1314:
1311:
1310:
1307:
1305:
1300:
1298:
1296:
1294:
1290:
1287:
1286:
1285:
1284:
1281:
1278:
1276:
1274:
1272:
1268:
1265:
1264:
1263:
1261:
1260:
1259:
1253:
1250:
1247:
1245:
1243:
1241:
1236:
1235:
1234:
1232:
1229:
1228:
1225:
1222:
1220:
1218:
1216:
1212:
1209:
1208:
1207:
1205:
1204:
1203:
1202:Brachychampsa
1198:
1197:
1194:
1192:
1191:
1185:
1183:
1181:
1179:
1174:
1173:
1172:
1170:
1169:Alligatoroid
1167:
1166:
1161:
1160:
1155:
1149:
1148:
1147:Brachychampsa
1143:
1137:
1136:
1131:
1125:
1123:
1122:
1116:
1114:
1112:
1110:
1106:
1103:
1102:
1101:
1099:
1098:
1097:
1092:
1091:
1087:
1084:
1081:
1078:
1075:
1072:
1069:
1068:
1065:
1063:
1058:
1050:
1041:
1038:
1036:
1029:
1025:
1022:
1018:
1015:
1011:
1006:
1001:
996:
993:
989:
988:
984:
981:
980:
972:
960:
959:
954:
952:
950:
948:
946:
943:
941:
940:
939:
934:
933:
930:
928:
923:
921:
919:
917:
915:
914:
910:
908:
907:
906:
901:
900:
897:
896:
891:
889:
887:
885:
883:
880:
878:
877:
876:
871:
870:
867:
866:
861:
859:
857:
855:
853:
852:
848:
846:
845:
844:
839:
838:
835:
832:
830:
828:
826:
824:
821:
819:
818:
817:
812:
811:
808:
805:
803:
801:
799:
797:
794:
792:
791:
790:
785:
784:
781:
778:
776:
774:
771:
769:
767:
766:
762:
760:
759:
758:
753:
752:
749:
746:
744:
742:
740:
738:
737:
733:
731:
730:
729:
724:
723:
718:
717:
712:
710:
708:
706:
704:
703:
699:
697:
696:
695:
690:
689:
685:
682:
679:
676:
673:
670:
667:
666:
663:
661:
656:
648:
639:
636:
634:
627:
623:
620:
616:
613:
609:
604:
599:
594:
591:
587:
586:
582:
579:
578:
570:
562:
560:
559:Trace fossils
556:
554:
553:
549:
545:
541:
540:
535:
534:insectivorans
531:
527:
523:
519:
518:
514:
510:
506:
505:
500:
496:
492:
489:
485:
484:
479:
475:
471:
467:
463:
459:
455:
451:
447:
439:
438:
433:
432:
431:Kosmoceratops
426:
417:
415:
411:
410:
409:Kosmoceratops
405:
401:
396:
392:
355:
347:
345:
341:
336:
334:
330:
320:
318:
317:fossiliferous
314:
310:
307:
303:
299:
295:
292:
288:
284:
280:
279:United States
276:
272:
268:
265:found in the
264:
261:
257:
248:
237:
233:
230:
219:
215:
210:
173:
171:
167:
162:
159:
155:
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139:
135:
132:
129:
125:
118:
113:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
33:
29:
25:
20:
4639:
4597:
4593:Horse Canyon
4573:Death Hollow
4547:Willis Creek
4527:Coyote Gulch
4473:, retrieved
4458:
4439:Bibliography
4392:
4388:
4378:
4337:
4333:
4270:
4266:
4256:
4248:
4193:
4189:
4145:
4141:
4107:
4049:
3988:
3984:
3974:
3963:. Retrieved
3959:the original
3954:
3950:
3944:
3936:
3887:
3884:Proc R Soc B
3883:
3851:. Retrieved
3847:the original
3842:
3838:
3828:
3783:
3779:
3731:
3727:
3717:
3672:
3668:
3662:
3616:
3563:
3496:
3459:
3390:
3386:
3380:
3354:
3350:
3326:. Retrieved
3322:the original
3315:
3308:
3297:. Retrieved
3293:the original
3288:
3278:
3245:
3241:
3183:
3137:
3133:
3093:
3067:
3063:
3057:
3032:
3028:
3022:
2997:
2993:
2989:
2961:
2954:
2917:
2913:
2889:. Retrieved
2885:the original
2880:
2871:
2863:the original
2858:
2854:
2844:
2819:
2815:
2757:(1): 16026.
2754:
2750:
2682:
2678:
2595:Troodontidae
2580:
2573:Middle unit
2564:
2563:
2555:
2554:
2543:
2534:"The Blues"
2528:
2527:
2519:
2518:
2487:
2476:
2472:
2469:Ornithomimus
2468:
2466:
2459:Middle unit
2452:
2445:
2431:
2416:
2415:
2407:
2406:
2395:
2392:caenagnathid
2386:
2376:"The Blues"
2371:H. giganteus
2370:
2369:
2361:
2360:
2325:
2313:
2301:
2289:
2277:
2265:
2232:
2199:
2187:
2178:
2121:
2097:
2082:
2081:
2075:Utahceratops
2073:
2072:
2044:Skaladromeus
2034:
2030:
2028:
2014:
2013:
2006:
1996:
1990:
1984:
1953:Middle unit
1944:Nodosauridae
1932:
1924:
1915:
1914:
1906:
1905:
1898:
1892:
1858:
1844:
1843:
1835:
1834:
1802:G. notabilis
1801:
1786:
1777:
1771:saurolophine
1765:
1758:Middle unit
1750:
1749:
1741:
1740:
1710:
1703:Middle unit
1664:
1644:
1614:Utahceratops
1612:
1592:
1580:
1568:
1556:
1544:
1530:
1528:
1522:
1517:
1512:
1506:
1505:
1497:
1496:
1463:
1451:
1442:
1385:
1361:
1348:
1340:Leidyosuchus
1338:
1337:
1330:
1313:Goniopholid
1312:
1303:
1301:
1288:
1279:
1266:
1256:
1255:
1248:
1230:
1223:
1210:
1200:
1199:
1188:
1186:
1168:
1157:
1145:
1135:Bernissartia
1133:
1121:Bernissartia
1119:
1117:
1104:
1096:Bernissartia
1094:
1093:
1062:Crocodilians
1060:
1048:
1039:
982:
955:
944:
938:Helopanoplia
936:
935:
924:
912:
911:
903:
902:
892:
881:
873:
872:
862:
850:
849:
841:
840:
833:
822:
814:
813:
806:
796:Two species
795:
787:
786:
779:
772:
764:
763:
755:
754:
747:
735:
734:
728:Arvinachelys
726:
725:
713:
701:
700:
692:
691:
658:
646:
637:
580:
557:
550:
543:
537:
515:
502:
481:
478:crocodilians
443:
440:in Laramidia
435:
429:
407:
388:
353:
340:Uranium-Lead
337:
333:Ar-Ar dating
326:
255:
253:
4542:Paria River
4537:Harris Wash
3316:Deinosuchus
2990:Gryposaurus
2529:T. sampsoni
2188:crossed out
2167:Morphotaxon
1969:Ornithopoda
1891:Similar to
1819:Lower unit
1794:Lower unit
1778:Gryposaurus
1768:kritosaurin
1743:Gryposaurus
1728:Lower unit
1682:Lower unit
1558:Gryposaurus
1507:A. johnsoni
1452:crossed out
1431:Morphotaxon
1304:D. hatcheri
1267:D. hatcheri
1258:Deinosuchus
1159:Deinosuchus
1118:May not be
1049:crossed out
1028:Morphotaxon
967:Neosuchians
789:Neurankylus
757:Denazinemys
647:crossed out
626:Morphotaxon
539:Gryposaurus
499:troodontids
483:Deinosuchus
480:(including
466:salamanders
294:floodplains
260:sedimentary
196: /
170:Coordinates
4659:Categories
4475:2019-02-21
4395:: e11013.
3991:: e10669.
3965:2017-11-03
3853:2020-12-21
3328:2016-03-27
3318:from Utah"
3299:2017-11-03
2920:: e11013.
2891:2016-03-27
2822:: 104527.
2658:References
2612:troodontid
2565:T. curriei
2546:troodontid
2184:small text
2153:Ichnotaxon
2065:ornithopod
2062:orodromine
1986:Orodromeus
1961:nodosaurid
1864:ceratopsid
1448:small text
1417:Ichnotaxon
1231:Caimanine
1045:small text
1014:Ichnotaxon
913:B. nobilis
765:D. nodosoa
736:A. goldeni
702:B. grandis
643:small text
612:Ichnotaxon
530:marsupials
524:including
420:Paleofauna
202: (
183:111°41′0″W
4411:2167-8359
4289:2167-8359
4273:: e7247.
4196:: e5910.
4162:131302174
4007:2167-8359
3393:: e5016.
3351:PaleoBios
3270:220415208
3070:(2): 1–7.
3049:131302174
3014:220413406
2836:0195-6671
2779:2045-2322
2709:1932-6203
2639:remains.
2614:remains.
2478:Rativates
2434:avisaurid
2417:M. eatoni
2363:Hagryphus
2355:remains.
2279:Hagryphus
2221:Material
2215:Location
2201:Theropods
2122:Color key
2106:Theropods
2083:U. gettyi
1963:remains.
1916:N. titusi
1868:processes
1774:hadrosaur
1725:Dog Flat
1667:species.
1651:sp. nov.
1485:Material
1479:Location
1386:Color key
1319:nov. gen.
1237:nov. gen.
1175:nov. gen.
1082:Material
1076:Location
983:Color key
905:Basilemys
843:Compsemys
834:A baenid
816:Thescelus
807:A baenid
780:A baenid
680:Material
674:Location
581:Color key
491:theropods
458:sturgeons
400:Campanian
306:sandstone
302:Laramidia
296:of large
283:Campanian
180:37°24′0″N
158:sandstone
145:Lithology
137:Thickness
32:Campanian
4456:(2004),
4429:33976955
4370:13261338
4362:21253683
4307:31333906
4222:30479894
4025:33552721
3914:23864598
3820:20877459
3780:PLOS ONE
3709:21949721
3669:PLOS ONE
3419:30065856
3162:83687743
2946:33976955
2797:36163377
2727:29166406
2679:PLOS ONE
2646:See also
2212:Species
1476:Species
1073:Species
851:C. victa
671:Species
513:duckbill
493:such as
313:mudstone
291:alluvial
164:Location
154:Mudstone
4556:Geology
4420:8061582
4342:Bibcode
4298:6626525
4213:6238772
4016:7842145
3905:3730592
3811:2929175
3788:Bibcode
3700:3176273
3677:Bibcode
3410:6063217
3250:Bibcode
3142:Bibcode
2937:8061582
2788:9512893
2759:Bibcode
2718:5699823
2687:Bibcode
2409:Mirarce
2291:Mirarce
2227:Images
2160:Ootaxon
1491:Images
1424:Ootaxon
1088:Images
1021:Ootaxon
694:Boremys
686:Images
660:Turtles
619:Ootaxon
565:Turtles
522:mammals
474:lizards
470:turtles
385:Habitat
235:Country
150:Primary
105:↓
4466:
4427:
4417:
4409:
4368:
4360:
4305:
4295:
4287:
4220:
4210:
4160:
4114:
4023:
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3818:
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3707:
3697:
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3407:
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3160:
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3047:
3012:
2968:
2944:
2934:
2834:
2795:
2785:
2777:
2725:
2715:
2707:
2224:Notes
2209:Genus
1488:Notes
1473:Genus
1085:Notes
1070:Genus
945:indt.
895:adocid
875:Adocus
716:baenid
683:Notes
668:Genus
532:, and
511:, the
501:, and
454:bowfin
298:rivers
244:
226:
217:Region
4622:Other
4389:PeerJ
4366:S2CID
4267:PeerJ
4190:PeerJ
4158:S2CID
4104:(PDF)
3985:PeerJ
3613:(PDF)
3456:(PDF)
3387:PeerJ
3266:S2CID
3158:S2CID
3045:S2CID
3010:S2CID
2914:PeerJ
2521:Talos
2179:Notes
2131:Taxon
1443:Notes
1395:Taxon
1349:L. sp
1211:B. sp
1105:B. sp
1040:Notes
992:Taxon
823:int.
638:Notes
590:Taxon
462:frogs
437:Talos
258:is a
4464:ISBN
4425:PMID
4407:ISSN
4358:PMID
4303:PMID
4285:ISSN
4218:PMID
4112:ISBN
4021:PMID
4003:ISSN
3910:PMID
3816:PMID
3705:PMID
3621:ISBN
3568:ISBN
3464:ISBN
3415:PMID
3357:(3).
3192:ISBN
3098:ISBN
2966:ISBN
2942:PMID
2832:ISSN
2793:PMID
2775:ISSN
2723:PMID
2705:ISSN
2488:cf.
2475:and
2233:cf.
1995:and
1789:sp.
450:gars
398:the
395:Utah
309:beds
275:Utah
254:The
229:Utah
127:Type
44:Preęž’
4415:PMC
4397:doi
4350:doi
4293:PMC
4275:doi
4208:PMC
4198:doi
4150:doi
4011:PMC
3993:doi
3900:PMC
3892:doi
3888:280
3806:PMC
3796:doi
3736:doi
3732:151
3695:PMC
3685:doi
3405:PMC
3395:doi
3258:doi
3150:doi
3037:doi
3002:doi
2932:PMC
2922:doi
2859:225
2824:doi
2820:114
2783:PMC
2767:doi
2713:PMC
2695:doi
2509:or
2432:An
2060:An
893:An
486:),
393:of
323:Age
269:in
247:USA
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