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desert, and coast to caves, mammalian burrows, and vegetation. They live in accordance with where their prey lives. One division of the
Geobiote feeds on maggots and eggs that are found in forest vegetation or in carrion. Naturally, these are found in heavily forested areas. The second and third divisions hunt for arthropods that feed on dead plant matter. Thus, these types of Geobiotes are found in the sand and burrowed in soil. The fourth type of Geobiotes feed on fly eggs that grow on fresh
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fly breeding resources, and they can be distinguished from fly larvae by the following: Beetle larvae possess a hard, head capsule, often brown in colour. Fly larvae lack a head capsule, instead having distinct, internal, black mouth hooks (cephalopharyngeal skeleton of mouthparts) at the anterior end of their body." Understanding how long ago the eggs were laid and the time period of the developmental stages is important for determining the time of death.
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899:. If the Histeridae beetles are present, the investigator can assume that some of the other insects have been eaten by the hister beetles. Due to their importance in forensic entomology, research is continually being conducted on this family's life cycle and development, their prevalence in specific locations, and their geographical distribution.
535:, dead vegetation, sandy areas, under tree bark, mammal burrows, and ant/termite colonies. Charactertistics of the hister beetle are dependent upon its habitat. For example, the flat hister beetles are found under bark, while the cylindrical beetles are not. The shape of the beetle will vary from species to species.
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cement. This makes their outer shell harder and protects them during this vulnerable stage. While they pupate, they breathe through spiracles on the abdomen. The beetle is non-feeding and immobile in this stage, as their internal structure is breaking down and rebuilding to its adult form. Under good
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The hister beetles will hide under the dead body in the soil during the day and come out at night to feed. For this reason, the dead body must be examined at different times of the day. After collecting the hister beetles from a body, isolate them because they are predacious beetles and eat anything
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are destructive and when they hatch they eat the maggots in the dead body. The stage of the beetle and other insects in the body at the time of investigation helps to determine the time of death. "Beetle larvae are ... helpful when determining post-mortem time interval. Beetle larvae often reside in
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The family
Histeridae is very diverse, so different species can be found on the body at different times because of their various feeding habits. This needs to be noted when examining a dead body. They are predacious beetles and arrive when there is material to feed on, like other beetles or maggots.
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Because members of the family
Histeridae can be found on carrion, they have proven to be important in certain forensic investigations. The predacious hister beetles will feed on the various insects on the body, primarily Diptera. To estimate a person's time of death, forensic investigators must look
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This subclade is the most diverse and the largest of the hister predator subclasses. The
Geobites' body structures are generally circular, and they are known for their digging tendencies. This subclass is separated into five more divisions. The members of this subclass live anywhere from the soil,
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This division consists of those
Histeridae that live in close proximity with social colonies of arthropods such as ants and termites. Histeridae that live near ants can live in a harmonious or hostile relationship. The hostile hister beetles feed on the ants. The harmonious hister beetles eat the
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This subclade is the smallest of the four. They live on plant litter and feed on the tiny arthropods found there. Microhisterids, like all other
Histeridae, become specialized to hunt their prey and live in their habitats. Like the fifth division of Geobiotes, some Microhisterids are known to be
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The larval stage of the beetle typically goes through two instars, and the second instar is the longest stage of its entire development, taking up 39% of overall development time. It takes 5.1±0.1 days on average for the first instar to develop into the second. The larval form of the insect will
553:. As new food sources are introduced into an environment, the predators of that food source will soon follow. Histeridae live in areas where their prey feed. Some examples are mammalian nests where other arthropods scavenge for food or carrion where maggots will arrive.
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Because the
Histeridae are predacious, they can be utilized as control agents, but must be stored in isolation. The hister beetles have proven useful in both the control of pest flies in poultry houses and pastures, and against pest beetles of stored food products.
914:. Forty species of Coleoptera, including the hister beetles, were observed on twelve pig carcasses over a one-year period. The beetles’ distribution and their time of colonization in the various decomposition stages over the year were examined and recorded.
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Histeridae is a large and diverse family. There are more than 410 genera and 4,800 described species in
Histeridae worldwide, with more than 500 species in North America. The beetles range in size, shape and color. Many of the adults are predacious.
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Some
Histeridae species live in an integrated nest with ants and termites. Some species have been found to be fed by the ants, while others simply feed on leftover insect larvae that the ants do not want. In contrast, in their adult stage
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Hister beetles are found throughout the world in various habitats. Histeridae have been located in North
America, Central America, Europe, Asia, and Australia, but each hister beetle occupies certain niches. The beetles live in dung,
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468:. Histeridae has two common names, the “clown” beetle and the “hister” beetle. There have been several theories explaining the origin of these common names. One theory for the “hister” nickname comes from the work of
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The larvae and adult forms of Histeridae have been known to feed on dung, carrion, decomposing vegetation, other insects, larvae, and pupae. Hister beetles are able to locate both dung and carrion through
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Dictionary of word roots and combining forms compiled from the Greek, Latin, and other languages, with special reference to biological terms and scientific names. Mountain View, Calif: Mayfield Pub. Co.,
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The average time of development from egg to adult at 30 °C is 20.5 days. The eggs of most species are off-white and oval in shape. The egg takes on average 3.8±0.02 days to hatch into the first
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eggs, are found near tree bark. The latter also feed on insects and prefer to live in forested areas. Species of the cylindrical Dendrobites usually hunt prey that is unique for that species.
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and their mandibular mouth parts can cause excess damage to the body. Insects can also help determine the season the body died in. The hister beetle is more prevalent in spring and summer.
472:, a Roman poet. Juvenal used the word “hister” to mean a dirty, lowly being. Another theory for the origin of this beetle's name stems from the fact that in Latin, “hister” means actor.
602:. When found on dung, carrion and vegetation, the hister beetles will feed on the fly larvae found there. The predacious hister beetle will feed on soft-bodied insect eggs and larvae,
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They have two common body shapes. One is flattened while the other has a more cylindrical appearance. The former usually lives near tree barks. This is because the prey they feed on,
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same food as the ants, however they may not be in direct competition for the food. These beetles have an excretory organ that produces an odor telling the ants they mean no harm.
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are located on the abdomen and are the tracheal system of the beetle. Oxygen is brought into the body by spiracles and tiny sacs exchange the oxygen like a mammal's lung.
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and forewings are normally well developed in beetles, but in Histeridae the elytra are shortened and rectangular. The shortened elytra expose the final two of the seven
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Arnett, R. H. 1968. Histeridae. In: The Beetles of the United States (A manual for identification). The American Entomological Institute, Ann Arbor, xii + 1112 pp.
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takes place, the male leaves enough sperm in the female to fertilize all the eggs in the female's ovaries. The excess sperm is kept in a special structure called
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Davies, A. "Family Histeridae." Checklist of beetles of Canada and Alaska. Canadian National Collection of Insects, Arachnids and Nematodes. 11 Mar. 2009 <
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is the study of insects related to crime scenes. Insects can be very helpful at a crime scene and give people an idea of what happened and when it happened.
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The pupal form of the beetle is similar in appearance to the adult form. They have outer cells produced in the larval stages that are reinforced with
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Myers, P., R. Espinosa, and C. S. Parr. "Family Histeridae." Animal Diversity Web. 1998. University of Michigan Museum of Zoology. 22 Mar. 2009 <
862:. This division of Geobiotes is found near animal homes such as nests and burrows. The last division of Geobiotes lives in caves. They feed on the
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Stephens, Stephanie A. "An Overview of the Coleopteran Family Histeridae and its Significance to Forensic Entomology." 2003. 13 Mar. 2009 <
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1658:(Coleoptera: Histeridae)." Journal of Entomological Science. CABI. Vet. Toxicology & Entomology Res. Lab., College station. 20 Mar. 2009.
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Brundage, Adrienne. "Importance of Beetles Part 2." Ento 431 Class Period. Texas A&M University: Kleberg, College Station. 11 Mar. 2009.
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and other arthropods that occupy the vegetation and fungi found there. This type of Geobiote is known to include species that become blind.
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which is typically shiny black or metallic green. The two main shapes for this family are oval and flat. The elytra is shorter than the
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pests that infest dung or to control house flies. Histeridae are most active at night and they will play dead if they feel threatened.
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Brundage, Adrienne. "Development and Orders." Ento 431 Class Period. Texas A&M University: Kleberg, College Station. 11 Mar. 2009.
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feeders are most active at night and will fake death if they feel threatened. This family of beetles will occupy almost any kind of
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Zhou, Yu-Lingzi; Ślipiński, Adam; Ren, Dong; Parker, Joseph (2019-04-16). Long, John A; Tautz, Diethard; Thayer, Margaret (eds.).
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The hister beetles lay their eggs in dead bodies and over a short period of time they develop into various stages. The Histeridae
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https://web.archive.org/web/20110714075700/http://resources.metapress.com/pdf-preview.axd?code=362v3k1361r2x385&size=largest
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Byrd, Dr. J. H. "Definition and Information." Forensic Entomology: Insects in Legal Investigations. 1998. 21 Mar. 2009 <
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Dillon, Elizabeth S., and Lawrence R. Dillon. A Manual of Common Beetles of Eastern North America. Peterson and Co., 1961.
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Fuller, John. "What do bugs have to do with forensic science?" How Stuff Works? University of Georgia. 21 Mar. 2009 <
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arrive at the dead body within a few hours. Certain species of the hister beetles follow shortly behind and prey on the
1466:Ă–zdemir, Senem & Sert, Osman (2009). "Determination of Coleoptera fauna on carcasses in Ankara province, Turkey".
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546:(termitophiles). The hister beetles can live in harmony with the ants or prey on the ants, depending on the species.
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range in length from three millimeters to several centimeters. They have a membranous body with a limited amount of
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583:, around 99 million years old, including those belonging to the extant subfamily Haeteriinae, and the living genus
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1255:"Beetle - MSN Encarta." MSN Encarta : Online Encyclopedia, Dictionary, Atlas, and Homework. 07 Apr. 2009 <
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1640:"Hister beetles - Family Histeridae." Decomposition: Hister Beetles. 2003. Australian Museum. 11 Mar. 2009 <
1434:"Hister beetles - Family Histeridae." Decomposition: Hister Beetles. 2003. Australian Museum. 11 Mar. 2009 <
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Gillott, C. 1995. The Remaining Endopterygote Orders. In: Entomology. Plenum Press, New York, xviii + 798 pp.
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arrive at the body first, when it is fresh to late (beginning of decomposition to advanced decomposition).
651:. The larval form does not resemble the adult form and the pupa, non-feeding and has internally developing
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514:, also known as a body cavity. They have a tube shaped heart that spans the length of the body, and use
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Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011).
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Dadour, Dr. Ian, and Dr. David Cook. "Collecting From a Corpse." Web Ring. 2001. 23 Mar. 2009 <
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to all living members of the family. The oldest crown group representatives are known from the
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arrive to feed when the carrion is bloat to dry (full of maggots to no maggots). In contrast,
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A remarkable ability of some hister beetles is their capacity to live in close proximity with
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around the body and it is horizontally segmented. The legs are short and do not help much in
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as blood. This blood does not contain any oxygen, but carries nutrients throughout the body.
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temperature conditions, the hister beetle will stay in the pupal stage for about a week.
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1633:(Erichson) (Coleoptera:Histeridae)." BioControl (2005). 14 Mar. 2005. 20 Mar. 2009 <
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Summerlin, J. W., and G. T. Fincher. "Laboratory observations on the life cycle of
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http://animaldiversity.ummz.umich.edu/site/accounts/classification/Histeridae.html
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499:. A distinctive characteristic is that the antennae are elbowed and contain three
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https://web.archive.org/web/20090330011619/http://www.beetlelady.com/?page_id=7
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http://encarta.msn.com/encnet/refpages/RefArticle.aspx?refid=761555773&pn=2
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in particular. Some species of Histeridae will even feed on other Histeridae.
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and will even eat other hister beetles, they must be isolated when collected.
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Jiang, Rixin; Song, Weicai; Yang, Huiyu; Shi, Chao; Wang, Shuo (July 2020).
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The male and female reproductive organs are hidden underneath the last few
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exposed. Hister beetles have specialized heads that can retract into their
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http://www.deathonline.net/decomposition/corpse_fauna/beetles/hister.htm
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http://www.deathonline.net/decomposition/corpse_fauna/beetles/hister.htm
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Mazur, Slawomir. "Ecology and Habitats of Histeridae." 13 Mar. 2009 <
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Achiano, K. A., and J. H. Giliomee. "Biology of the house fly predator
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while the male's is adapted as a copulatory structure. The female has
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http://www.museum.hokudai.ac.jp/exhibition/kikaku03/6sympoMazur.html
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Peter W. Kovarik & Michael S. Caterino (2001). "Histeridae". In
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The majority of Histeridae species prefer dried, decaying habitats.
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species from mid-Cretaceous Burmese amber (Coleoptera: Histeridae)"
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http://www.canacoll.org/Coleo/Checklist/PDF%20files/HISTERIDAE.pdf
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Histeridae live in widespread habitats because they feed on other
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stages of other insects. Certain species are also used to control
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investigations to help in time of death estimation. Also, certain
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found worldwide. They can be easily identified by their shortened
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1314:"A Mesozoic clown beetle myrmecophile (Coleoptera: Histeridae)"
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present. Insects that feed on dead bodies increase the rate of
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throughout the world. Hister beetles have proved useful during
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which holds the sperm until the eggs are fully developed.
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http://science.howstuffworks.com/forensic-entomology2.htm
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to the copulatory structure, which stays concealed until
729:) side. The female's structure is modified to work as an
2130:(furniture beetles, death watch beetles, spider beetles)
1230:
416:
The hister beetles are easily identified by their shiny
382:
are used in the control of livestock pests that infest
1311:
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350:. This very diverse group of beetles contains 3,900
1282:
679:it can look pale brown and be leathery in texture.
1545:http://agspsrv34.agric.wa.gov.au/ento/forensic.htm
895:at the insects on the body and determine time of
700:. They move mostly through muscular contraction.
3982:
3007:(jewel beetles, or metallic wood-boring beetles)
1623:Further references (not necessarily used inline)
1502:http://www.forensicentomology.com/definition.htm
1370:
835:worth mentioning. These subclades are known as:
831:There are four subclades of Histeridae that are
2730:(true weevils, bark beetles, ambrosia beetles)
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1196:
1194:
1192:
1190:
1188:
1132:
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745:. The males also have a duct that carries the
1743:
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671:is shiny and smooth. In certain species like
2973:(ironclad beetles, cylindrical bark beetles)
1139:"Family-group names in Coleoptera (Insecta)"
1465:
1185:
1127:
1750:
1736:
1722:Lectures on Forensically Important Insects
1512:http://www.forensicentomology.com/info.htm
917:
100:
3637:List of subgroups of the order Coleoptera
2717:(straight snout weevils, New York weevil)
1816:
1347:
1329:
1168:
1158:
819:Learn how and when to remove this message
635:, meaning it goes through four stages of
3144:(cicada beetle, cicada parasite beetles)
403:
362:exposed, and their geniculate (elbowed)
1569:Integrated Taxonomic Information System
737:that carry the developed eggs from the
556:
3983:
3234:(false fireflies, long-lipped beetles)
2059:
2010:
440:, hister beetles commonly feed on the
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2058:
2009:
1884:
1883:
1815:
1731:
979:
3945:B718293D-FFCE-F20F-FF0D-FC9EBD8DF65D
3906:c9c5cba9-392d-4db4-818b-e6d3d8b48861
3818:6a08aaed-bb27-42ac-8a42-576a697dd7bf
3260:(false metallic wood-boring beetles)
1581:
1557:
801:adding citations to reliable sources
772:
575:of China, which was resolved as the
889:
561:The oldest record of the family is
111:with larvae and anatomical details
13:
619:will actually feed on adult ants.
397:
14:
4017:
1663:
948:They do not actually feed on the
592:
4006:Taxa named by Leonard Gyllenhaal
2871:Perimylopidae, or Promecheilidae
2456:(minute brown scavenger beetles)
1670:
1091:
1077:
1063:
1049:
1035:
1021:
1007:
993:
777:
488:. The head has compound eyes, a
424:with typically two of the seven
120:
32:
3086:(variegated mud-loving beetles)
1605:
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1494:
1480:10.1016/j.forsciint.2008.09.018
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788:needs additional citations for
692:around the head. There is some
1468:Forensic Science International
1305:
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1221:
626:
109:Central European clown beetles
1:
2915:(narrow-waisted bark beetles)
2607:(parasitic flat bark beetles)
2558:(short-winged flower beetles)
2414:(lady beetles, or God's cows)
2151:(tooth-necked fungus beetles)
1401:10.1016/j.cretres.2020.104443
1120:
503:that form a club at the end.
432:and two geniculate (elbowed)
358:that leaves two of the seven
3427:(sand-loving scarab beetles)
2817:(minute tree-fungus beetles)
2643:(silvanid flat bark beetles)
768:
459:
7:
3576:(primitive carrion beetles)
3532:(minute mud-loving beetles)
3395:(bumble bee scarab beetles)
3165:(soft-bodied plant beetles)
3136:(soft bodied plant beetles)
1939:(predaceous diving beetles)
1108:
506:Clown beetles have an open
10:
4022:
3411:(scavenger scarab beetles)
3403:(enigmatic scarab beetles)
2576:Lamingtonium binnaberrense
2290:(soft-wing flower beetles)
1099:Paromalus parallelepipedus
717:Male and female structures
633:holometabolous development
525:
475:
3658:
3634:
3564:
3550:(water scavenger beetles)
3515:
3481:
3472:
3350:
3341:
3294:
3283:Rhinorhipus tamborinensis
3269:
3153:
3124:
3062:(long-toed water beetles)
3021:
2995:
2986:
2955:(polypore fungus beetles)
2857:(palm and flower beetles)
2841:(tumbling flower beetles)
2788:(ant-like flower beetles)
2768:
2747:
2673:
2566:(lined flat bark beetles)
2540:(pleasing fungus beetles)
2480:
2435:(handsome fungus beetles)
2371:
2227:
2173:
2164:
2139:
2081:
2072:
2068:
2054:
2019:
2005:
1980:(burrowing water beetles)
1893:
1879:
1825:
1811:
1767:
1217:. Vol. 1. CRC Press.
1115:List of Histeridae genera
902:A study was conducted at
682:
366:with clubbed ends. These
322:
315:
221:
216:
117:Scientific classification
115:
108:
99:
23:
3597:(feather-winged beetles)
2931:(false longhorn beetles)
2878:(jugular-horned beetles)
2825:(false darkling beetles)
2796:(cryptic fungus beetles)
2615:(shining flower beetles)
1955:(crawling water beetles)
1854:Sikhotealinia zhiltzovae
1373:"Discovery of the first
703:
631:Histeridae goes through
464:Histeridae was named by
30:Early Cretaceous–Present
3078:(forest stream beetles)
2939:(synchroa bark beetles)
2899:(dead log bark beetles)
2891:(fire-coloured beetles)
2780:(ant-like leaf beetles)
2514:(silken fungus beetles)
2448:(well polished beetles)
2422:(minute fungus beetles)
2241:Acanthocnemus nigricans
962:Saprinus pennsylvanicus
918:Importance in forensics
3589:(round fungus beetles)
2923:(false flower beetles)
2907:(wedge-shaped beetles)
2865:(false blister beetle)
2849:(hairy fungus beetles)
2804:(conifer bark beetles)
2701:(leaf-rolling weevils)
2362:(bark-gnawing beetles)
2122:(wounded-tree beetles)
1988:(false ground beetles)
1910:(trout-stream beetles)
1702:Overview of Histeridae
1160:10.3897/zookeys.88.807
1085:Pachylister inaequalis
1057:Hister quadrimaculatus
658:
436:with clubbed ends. As
411:
3927:Paleobiology Database
3501:(false clown beetles)
3323:(plate-thigh beetles)
3205:(false click beetles)
3110:(water-penny beetles)
2759:(ship-timber beetles)
2738:(pine flower weevils)
2586:(root-eating beetles)
2406:(minute bark beetles)
2317:Phloiophilus edwardsi
1844:(reticulated beetles)
1043:Haeterius ferrugineus
407:
3813:Fauna Europaea (new)
3379:(false stag beetles)
3364:Belohina inexpectata
3226:(net-winged beetles)
3189:(rare click beetles)
3102:(travertine beetles)
3094:(minute mud beetles)
2659:(dry-fungus beetles)
2494:Agapytho foveicollis
2251:(false skin beetles)
2114:(Jacobson's beetles)
1834:Crowsoniella relicta
1694:Histeridae of Europe
1071:Onthophilus striatus
954:Hister quadrinotatus
904:Hacettepe University
797:improve this article
725:on the mesosternal (
557:Evolutionary history
542:(myrmecophiles) and
3377:Diphyllostomatidae
2709:(primitive weevils)
2522:(flat bark beetles)
2272:(checkered beetles)
2259:(fruitworm beetles)
1947:(whirligig beetles)
1712:Forensic Entomology
1565:"Histeridae Report"
1393:2020CrRes.11104443J
1381:Cretaceous Research
1331:10.7554/eLife.44985
1205:Arnett, Ross H. Jr.
1151:2011ZooK...88....1B
1029:Atholus bimaculatus
1015:Asolenus julietteae
923:Forensic entomology
390:. Because they are
324:at least 410 genera
3242:(glowworm beetles)
2947:(darkling beetles)
2794:Archeocrypticidae
2651:(palmetto beetles)
2398:(dry bark beetles)
2185:(longhorn beetles)
1550:2006-08-19 at the
1517:2018-02-15 at the
1507:2018-02-14 at the
1298:2008-02-03 at the
1262:2003-08-21 at the
1209:Thomas, Michael C.
980:Genera and species
508:circulatory system
466:Leonard Gyllenhaal
412:
342:commonly known as
3978:
3977:
3914:Open Tree of Life
3652:Taxon identifiers
3643:
3642:
3630:
3629:
3626:
3625:
3622:
3621:
3614:
3606:
3605:(carrion beetles)
3598:
3590:
3577:
3551:
3533:
3502:
3494:
3468:
3467:
3460:
3452:
3444:
3436:
3428:
3420:
3412:
3404:
3396:
3388:
3380:
3367:
3337:
3336:
3324:
3316:
3286:
3261:
3243:
3235:
3227:
3219:
3206:
3198:
3190:
3182:
3181:(soldier beetles)
3174:
3171:Brachypsectridae
3166:
3145:
3137:
3111:
3103:
3095:
3087:
3079:
3071:
3063:
3050:
3042:
3034:
3008:
2982:
2981:
2974:
2956:
2948:
2940:
2932:
2929:Stenotrachelidae
2924:
2916:
2908:
2900:
2892:
2879:
2866:
2858:
2850:
2842:
2834:
2833:(blister beetles)
2826:
2818:
2805:
2797:
2789:
2781:
2760:
2739:
2731:
2718:
2710:
2702:
2694:
2684:
2660:
2652:
2644:
2616:
2608:
2600:
2587:
2579:
2567:
2559:
2541:
2523:
2515:
2497:
2457:
2449:
2436:
2423:
2415:
2407:
2399:
2381:Akalyptoischiidae
2363:
2320:
2303:
2291:
2273:
2260:
2252:
2244:
2194:
2186:
2160:
2159:
2152:
2131:
2123:
2115:
2102:
2094:
2050:
2049:
2027:
2001:
2000:
1997:
1996:
1989:
1981:
1973:
1956:
1948:
1940:
1932:
1924:
1911:
1875:
1874:
1857:
1845:
1837:
1631:Carcinops pumilio
829:
828:
821:
329:
328:
309:
301:
293:
285:
277:
269:
261:
253:
245:
237:
229:
212:
16:Family of beetles
4013:
3971:
3970:
3958:
3957:
3948:
3947:
3935:
3934:
3922:
3921:
3909:
3908:
3899:
3898:
3886:
3885:
3883:NHMSYS0001718459
3873:
3872:
3860:
3859:
3847:
3846:
3834:
3833:
3821:
3820:
3808:
3807:
3795:
3794:
3782:
3781:
3769:
3768:
3756:
3755:
3743:
3742:
3730:
3729:
3720:
3719:
3707:
3706:
3694:
3693:
3692:
3679:
3678:
3677:
3647:
3646:
3612:
3604:
3596:
3588:
3575:
3549:
3531:
3500:
3492:
3479:
3478:
3474:Staphyliniformia
3458:
3450:
3442:
3434:
3426:
3418:
3410:
3402:
3394:
3386:
3378:
3361:
3348:
3347:
3322:
3313:Declinia relicta
3310:
3280:
3259:
3253:Sinopyrophoridae
3248:Rhagophthalmidae
3241:
3233:
3225:
3217:
3204:
3196:
3188:
3180:
3172:
3164:
3163:Artematopodidae
3143:
3135:
3109:
3101:
3093:
3085:
3077:
3070:(riffle beetles)
3069:
3061:
3049:(turtle beetles)
3048:
3040:
3032:
3006:
2993:
2992:
2972:
2954:
2946:
2938:
2930:
2922:
2914:
2906:
2898:
2890:
2877:
2864:
2856:
2848:
2840:
2832:
2824:
2816:
2803:
2795:
2787:
2779:
2758:
2737:
2729:
2716:
2708:
2700:
2693:(fungus weevils)
2692:
2683:
2682:
2678:
2665:Tasmosalpingidae
2658:
2650:
2642:
2614:
2606:
2598:
2585:
2573:
2565:
2557:
2539:
2521:
2513:
2491:
2455:
2447:
2434:
2421:
2413:
2405:
2397:
2361:
2314:
2297:
2289:
2271:
2258:
2250:
2238:
2237:Acanthocnemidae
2192:
2184:
2171:
2170:
2150:
2129:
2121:
2113:
2100:
2092:
2079:
2078:
2070:
2069:
2056:
2055:
2025:
2007:
2006:
1987:
1979:
1967:
1954:
1946:
1938:
1930:
1923:(ground beetles)
1922:
1909:
1895:
1894:
1881:
1880:
1851:
1843:
1831:
1830:Crowsoniellidae
1813:
1812:
1752:
1745:
1738:
1729:
1728:
1698:
1675:Data related to
1674:
1616:
1609:
1603:
1602:
1600:
1599:
1585:
1579:
1578:
1576:
1575:
1561:
1555:
1541:
1535:
1528:
1522:
1498:
1492:
1491:
1463:
1457:
1454:
1448:
1445:
1439:
1432:
1426:
1419:
1413:
1412:
1368:
1362:
1361:
1351:
1333:
1309:
1303:
1289:
1280:
1277:
1271:
1253:
1247:
1243:
1237:
1234:
1228:
1225:
1219:
1218:
1214:American Beetles
1200:
1183:
1182:
1172:
1162:
1134:
1095:
1081:
1067:
1053:
1039:
1025:
1011:
997:
890:Current research
824:
817:
813:
810:
804:
781:
773:
573:Yixian Formation
495:, and shortened
307:
299:
291:
283:
275:
267:
259:
251:
243:
235:
227:
207:
184:Staphyliniformia
125:
124:
104:
94:
31:
27:Temporal range:
21:
20:
4021:
4020:
4016:
4015:
4014:
4012:
4011:
4010:
4001:Beetle families
3981:
3980:
3979:
3974:
3966:
3961:
3953:
3951:
3943:
3938:
3930:
3925:
3917:
3912:
3904:
3902:
3894:
3889:
3881:
3876:
3868:
3863:
3855:
3850:
3842:
3837:
3829:
3824:
3816:
3811:
3803:
3798:
3790:
3785:
3777:
3772:
3764:
3759:
3751:
3746:
3738:
3733:
3725:
3723:
3715:
3710:
3702:
3697:
3688:
3687:
3682:
3673:
3672:
3667:
3654:
3644:
3639:
3618:
3560:
3511:
3493:(clown beetles)
3464:
3435:(betsy beetles)
3343:Scarabaeiformia
3333:
3290:
3265:
3197:(click beetles)
3173:(Texas beetles)
3149:
3120:
3116:Ptilodactylidae
3041:(cedar beetles)
3017:
2978:
2961:Trictenotomidae
2847:Mycetophagidae
2764:
2743:
2680:
2679:
2677:
2669:
2621:Phloeostichidae
2572:Lamingtoniidae
2564:Laemophloeidae
2512:Cryptophagidae
2476:
2367:
2350:Thanerocleridae
2313:Phloiophilidae
2300:Metaxina ornata
2265:Chaetosomatidae
2223:
2156:
2135:
2093:(auger beetles)
2074:Bostrichiformia
2064:
2046:
2042:Torridincolidae
2026:(skiff beetles)
2024:Hydroscaphidae
2015:
1993:
1986:Trachypachidae
1931:(tiger beetles)
1889:
1871:
1821:
1807:
1763:
1756:
1726:
1696:
1666:
1661:
1625:
1620:
1619:
1610:
1606:
1597:
1595:
1587:
1586:
1582:
1573:
1571:
1563:
1562:
1558:
1552:Wayback Machine
1542:
1538:
1529:
1525:
1519:Wayback Machine
1509:Wayback Machine
1499:
1495:
1464:
1460:
1455:
1451:
1446:
1442:
1433:
1429:
1420:
1416:
1369:
1365:
1310:
1306:
1300:Wayback Machine
1290:
1283:
1278:
1274:
1264:Wayback Machine
1254:
1250:
1244:
1240:
1235:
1231:
1226:
1222:
1201:
1186:
1135:
1128:
1123:
1111:
1106:
1105:
1104:
1101:
1096:
1087:
1082:
1073:
1068:
1059:
1054:
1045:
1040:
1031:
1026:
1017:
1012:
1003:
1001:Acritus minutus
998:
982:
958:Hister sedakovi
952:. For example,
920:
892:
876:blind as well.
825:
814:
808:
805:
794:
782:
771:
719:
706:
685:
661:
629:
595:
559:
528:
478:
462:
414:
413:
409:Hister unicolor
400:
398:Characteristics
386:and to control
308:Bickhardt, 1913
292:Bickhardt, 1914
284:Blanchard, 1845
260:Gyllenhal, 1808
236:Bickhardt, 1914
206:
119:
110:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
4019:
4009:
4008:
4003:
3998:
3993:
3976:
3975:
3973:
3972:
3959:
3949:
3936:
3923:
3910:
3900:
3887:
3874:
3861:
3848:
3835:
3822:
3809:
3800:Fauna Europaea
3796:
3783:
3770:
3757:
3744:
3731:
3721:
3708:
3695:
3680:
3664:
3662:
3656:
3655:
3641:
3640:
3635:
3632:
3631:
3628:
3627:
3624:
3623:
3620:
3619:
3617:
3616:
3613:(rove beetles)
3611:Staphylinidae
3608:
3600:
3592:
3584:
3579:
3570:
3568:
3566:Staphylinoidea
3562:
3561:
3559:
3558:
3553:
3548:Hydrophilidae
3545:
3540:
3535:
3527:
3521:
3519:
3517:Hydrophiloidea
3513:
3512:
3510:
3509:
3504:
3496:
3487:
3485:
3476:
3470:
3469:
3466:
3465:
3463:
3462:
3459:(hide beetles)
3454:
3446:
3443:(rain beetles)
3438:
3430:
3422:
3419:(stag beetles)
3414:
3406:
3398:
3390:
3382:
3374:
3372:Bolboceratidae
3369:
3356:
3354:
3345:
3339:
3338:
3335:
3334:
3332:
3331:
3326:
3318:
3306:
3300:
3298:
3292:
3291:
3289:
3288:
3279:Rhinorhipidae
3275:
3273:
3271:Rhinorhipoidea
3267:
3266:
3264:
3263:
3255:
3250:
3245:
3237:
3229:
3221:
3213:
3208:
3200:
3192:
3184:
3176:
3168:
3159:
3157:
3151:
3150:
3148:
3147:
3139:
3130:
3128:
3122:
3121:
3119:
3118:
3113:
3105:
3097:
3089:
3084:Heteroceridae
3081:
3073:
3065:
3057:
3052:
3047:Chelonariidae
3044:
3039:Callirhipidae
3036:
3033:(pill beetles)
3027:
3025:
3019:
3018:
3016:
3015:
3010:
3001:
2999:
2990:
2984:
2983:
2980:
2979:
2977:
2976:
2968:
2963:
2958:
2950:
2945:Tenebrionidae
2942:
2934:
2926:
2918:
2910:
2902:
2894:
2886:
2881:
2873:
2868:
2860:
2852:
2844:
2836:
2828:
2820:
2812:
2807:
2799:
2791:
2783:
2774:
2772:
2770:Tenebrionoidea
2766:
2765:
2763:
2762:
2753:
2751:
2745:
2744:
2742:
2741:
2733:
2728:Curculionidae
2725:
2720:
2712:
2704:
2696:
2687:
2685:
2675:Curculionoidea
2671:
2670:
2668:
2667:
2662:
2654:
2646:
2638:
2636:Protocucujidae
2633:
2628:
2623:
2618:
2610:
2602:
2594:
2589:
2581:
2569:
2561:
2553:
2548:
2543:
2535:
2530:
2528:Cybocephalidae
2525:
2517:
2509:
2504:
2499:
2486:
2484:
2478:
2477:
2475:
2474:
2469:
2464:
2459:
2451:
2443:
2438:
2430:
2428:Discolomatidae
2425:
2417:
2412:Coccinellidae
2409:
2401:
2396:Bothrideridae
2393:
2388:
2383:
2377:
2375:
2373:Coccinelloidea
2369:
2368:
2366:
2365:
2360:Trogossitidae
2357:
2352:
2347:
2342:
2337:
2332:
2327:
2322:
2310:
2305:
2293:
2285:
2280:
2278:Lophocateridae
2275:
2267:
2262:
2254:
2246:
2233:
2231:
2225:
2224:
2222:
2221:
2216:
2211:
2206:
2201:
2196:
2193:(leaf beetles)
2191:Chrysomelidae
2188:
2179:
2177:
2175:Chrysomeloidea
2168:
2162:
2161:
2158:
2157:
2155:
2154:
2145:
2143:
2137:
2136:
2134:
2133:
2125:
2120:Nosodendridae
2117:
2112:Jacobsoniidae
2109:
2104:
2101:(skin beetles)
2096:
2087:
2085:
2076:
2066:
2065:
2052:
2051:
2048:
2047:
2045:
2044:
2039:
2034:
2029:
2020:
2017:
2016:
2003:
2002:
1999:
1998:
1995:
1994:
1992:
1991:
1983:
1975:
1970:Meru phyllisae
1963:
1958:
1950:
1942:
1934:
1926:
1918:
1913:
1904:
1902:
1891:
1890:
1877:
1876:
1873:
1872:
1870:
1869:
1864:
1862:Micromalthidae
1859:
1847:
1839:
1826:
1823:
1822:
1809:
1808:
1806:
1805:
1799:
1793:
1787:
1781:
1775:
1768:
1765:
1764:
1755:
1754:
1747:
1740:
1732:
1725:
1724:
1719:
1714:
1709:
1704:
1699:
1691:
1685:
1680:
1679:at Wikispecies
1667:
1665:
1664:External links
1662:
1660:
1659:
1652:
1645:
1638:
1626:
1624:
1621:
1618:
1617:
1604:
1580:
1556:
1536:
1523:
1493:
1474:(1–3): 24–32.
1458:
1449:
1440:
1427:
1414:
1363:
1304:
1281:
1272:
1248:
1238:
1229:
1220:
1184:
1125:
1124:
1122:
1119:
1118:
1117:
1110:
1107:
1103:
1102:
1097:
1090:
1088:
1083:
1076:
1074:
1069:
1062:
1060:
1055:
1048:
1046:
1041:
1034:
1032:
1027:
1020:
1018:
1013:
1006:
1004:
999:
992:
989:
988:
987:
981:
978:
976:in their way.
919:
916:
891:
888:
871:Microhisterids
827:
826:
809:September 2018
785:
783:
776:
770:
767:
718:
715:
705:
702:
690:sclerotization
684:
681:
660:
657:
628:
625:
594:
593:Feeding habits
591:
577:sister species
558:
555:
527:
524:
477:
474:
461:
458:
402:
401:
399:
396:
348:hister beetles
327:
326:
320:
319:
313:
312:
311:
310:
302:
294:
286:
278:
270:
262:
254:
246:
241:Dendrophilinae
238:
233:Chlamydopsinae
230:
219:
218:
214:
213:
201:
197:
196:
191:
187:
186:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
141:
137:
136:
131:
127:
126:
113:
112:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
26:
15:
9:
6:
4:
3:
2:
4018:
4007:
4004:
4002:
3999:
3997:
3994:
3992:
3989:
3988:
3986:
3969:
3964:
3960:
3956:
3950:
3946:
3941:
3937:
3933:
3928:
3924:
3920:
3915:
3911:
3907:
3901:
3897:
3892:
3888:
3884:
3879:
3875:
3871:
3866:
3862:
3858:
3853:
3849:
3845:
3840:
3836:
3832:
3827:
3823:
3819:
3814:
3810:
3806:
3801:
3797:
3793:
3788:
3784:
3780:
3775:
3771:
3767:
3762:
3758:
3754:
3749:
3745:
3741:
3736:
3732:
3728:
3722:
3718:
3713:
3709:
3705:
3700:
3696:
3691:
3685:
3681:
3676:
3670:
3666:
3665:
3663:
3661:
3657:
3653:
3648:
3638:
3633:
3615:
3609:
3607:
3601:
3599:
3593:
3591:
3585:
3583:
3580:
3578:
3572:
3571:
3569:
3567:
3563:
3557:
3554:
3552:
3546:
3544:
3541:
3539:
3536:
3534:
3528:
3526:
3523:
3522:
3520:
3518:
3514:
3508:
3505:
3503:
3499:Sphaeritidae
3497:
3495:
3489:
3488:
3486:
3484:
3480:
3477:
3475:
3471:
3461:
3455:
3453:
3449:Scarabaeidae
3447:
3445:
3439:
3437:
3431:
3429:
3423:
3421:
3415:
3413:
3407:
3405:
3399:
3397:
3391:
3389:
3387:(dor beetles)
3383:
3381:
3375:
3373:
3370:
3368:
3365:
3358:
3357:
3355:
3353:
3352:Scarabaeoidea
3349:
3346:
3344:
3340:
3330:
3327:
3325:
3319:
3317:
3314:
3307:
3305:
3302:
3301:
3299:
3297:
3293:
3287:
3284:
3277:
3276:
3274:
3272:
3268:
3262:
3256:
3254:
3251:
3249:
3246:
3244:
3238:
3236:
3230:
3228:
3222:
3220:
3214:
3212:
3209:
3207:
3201:
3199:
3193:
3191:
3187:Cerophytidae
3185:
3183:
3177:
3175:
3169:
3167:
3161:
3160:
3158:
3156:
3152:
3146:
3142:Rhipiceridae
3140:
3138:
3132:
3131:
3129:
3127:
3123:
3117:
3114:
3112:
3106:
3104:
3098:
3096:
3090:
3088:
3082:
3080:
3076:Eulichadidae
3074:
3072:
3066:
3064:
3058:
3056:
3055:Cneoglossidae
3053:
3051:
3045:
3043:
3037:
3035:
3029:
3028:
3026:
3024:
3020:
3014:
3013:Schizopodidae
3011:
3009:
3003:
3002:
3000:
2998:
2994:
2991:
2989:
2988:Elateriformia
2985:
2975:
2969:
2967:
2964:
2962:
2959:
2957:
2953:Tetratomidae
2951:
2949:
2943:
2941:
2935:
2933:
2927:
2925:
2919:
2917:
2911:
2909:
2905:Ripiphoridae
2903:
2901:
2895:
2893:
2889:Pyrochroidae
2887:
2885:
2884:Pterogeniidae
2882:
2880:
2874:
2872:
2869:
2867:
2861:
2859:
2853:
2851:
2845:
2843:
2837:
2835:
2829:
2827:
2823:Melandryidae
2821:
2819:
2813:
2811:
2810:Chalcodryidae
2808:
2806:
2800:
2798:
2792:
2790:
2784:
2782:
2776:
2775:
2773:
2771:
2767:
2761:
2755:
2754:
2752:
2750:
2746:
2740:
2736:Nemonychidae
2734:
2732:
2726:
2724:
2721:
2719:
2713:
2711:
2705:
2703:
2697:
2695:
2689:
2688:
2686:
2676:
2672:
2666:
2663:
2661:
2655:
2653:
2647:
2645:
2639:
2637:
2634:
2632:
2631:Propalticidae
2629:
2627:
2626:Priasilphidae
2624:
2622:
2619:
2617:
2611:
2609:
2605:Passandridae
2603:
2601:
2599:(sap beetles)
2595:
2593:
2590:
2588:
2582:
2580:
2577:
2570:
2568:
2562:
2560:
2554:
2552:
2549:
2547:
2544:
2542:
2536:
2534:
2531:
2529:
2526:
2524:
2518:
2516:
2510:
2508:
2507:Cavognathidae
2505:
2503:
2500:
2498:
2495:
2488:
2487:
2485:
2483:
2479:
2473:
2470:
2468:
2465:
2463:
2460:
2458:
2452:
2450:
2444:
2442:
2441:Eupsilobiidae
2439:
2437:
2433:Endomychidae
2431:
2429:
2426:
2424:
2420:Corylophidae
2418:
2416:
2410:
2408:
2402:
2400:
2394:
2392:
2389:
2387:
2384:
2382:
2379:
2378:
2376:
2374:
2370:
2364:
2358:
2356:
2353:
2351:
2348:
2346:
2343:
2341:
2338:
2336:
2335:Protopeltidae
2333:
2331:
2330:Prionoceridae
2328:
2326:
2323:
2321:
2318:
2311:
2309:
2306:
2304:
2301:
2294:
2292:
2286:
2284:
2283:Mauroniscidae
2281:
2279:
2276:
2274:
2268:
2266:
2263:
2261:
2255:
2253:
2247:
2245:
2242:
2235:
2234:
2232:
2230:
2226:
2220:
2217:
2215:
2212:
2210:
2207:
2205:
2204:Megalopodidae
2202:
2200:
2197:
2195:
2189:
2187:
2183:Cerambycidae
2181:
2180:
2178:
2176:
2172:
2169:
2167:
2163:
2153:
2149:Derodontidae
2147:
2146:
2144:
2142:
2141:Derodontoidea
2138:
2132:
2126:
2124:
2118:
2116:
2110:
2108:
2107:Endecatomidae
2105:
2103:
2097:
2095:
2091:Bostrichidae
2089:
2088:
2086:
2084:
2083:Bostrichoidea
2080:
2077:
2075:
2071:
2067:
2063:
2057:
2053:
2043:
2040:
2038:
2037:Sphaeriusidae
2035:
2033:
2030:
2028:
2022:
2021:
2018:
2014:
2008:
2004:
1990:
1984:
1982:
1976:
1974:
1971:
1964:
1962:
1959:
1957:
1951:
1949:
1943:
1941:
1935:
1933:
1929:Cicindelidae
1927:
1925:
1919:
1917:
1914:
1912:
1906:
1905:
1903:
1900:
1896:
1892:
1888:
1882:
1878:
1868:
1865:
1863:
1860:
1858:
1855:
1848:
1846:
1840:
1838:
1835:
1828:
1827:
1824:
1820:
1814:
1810:
1804:
1800:
1798:
1794:
1792:
1788:
1786:
1782:
1780:
1776:
1774:
1770:
1769:
1766:
1761:
1753:
1748:
1746:
1741:
1739:
1734:
1733:
1730:
1723:
1720:
1718:
1715:
1713:
1710:
1708:
1705:
1703:
1700:
1695:
1692:
1689:
1688:Histeridae de
1686:
1684:
1681:
1678:
1673:
1669:
1668:
1657:
1653:
1650:
1646:
1643:
1639:
1636:
1632:
1628:
1627:
1614:
1608:
1594:
1590:
1584:
1570:
1566:
1560:
1553:
1549:
1546:
1540:
1533:
1527:
1520:
1516:
1513:
1510:
1506:
1503:
1497:
1489:
1485:
1481:
1477:
1473:
1469:
1462:
1453:
1444:
1437:
1431:
1424:
1418:
1410:
1406:
1402:
1398:
1394:
1390:
1386:
1382:
1378:
1376:
1367:
1359:
1355:
1350:
1345:
1341:
1337:
1332:
1327:
1323:
1319:
1315:
1308:
1301:
1297:
1294:
1288:
1286:
1276:
1269:
1265:
1261:
1258:
1252:
1242:
1233:
1224:
1216:
1215:
1210:
1206:
1199:
1197:
1195:
1193:
1191:
1189:
1180:
1176:
1171:
1166:
1161:
1156:
1152:
1148:
1145:(88): 1–972.
1144:
1140:
1133:
1131:
1126:
1116:
1113:
1112:
1100:
1094:
1089:
1086:
1080:
1075:
1072:
1066:
1061:
1058:
1052:
1047:
1044:
1038:
1033:
1030:
1024:
1019:
1016:
1010:
1005:
1002:
996:
991:
990:
986:
977:
973:
970:
965:
963:
959:
955:
951:
945:
943:
942:decomposition
939:
935:
931:
928:Flesh eating
926:
924:
915:
913:
909:
905:
900:
898:
887:
883:
882:
877:
873:
872:
867:
865:
861:
855:
854:
849:
847:
842:
841:
836:
834:
823:
820:
812:
802:
798:
792:
791:
786:This section
784:
780:
775:
774:
766:
764:
760:
759:fertilization
756:
752:
748:
744:
740:
736:
732:
728:
724:
714:
711:
710:proteinaceous
701:
699:
695:
691:
680:
678:
674:
670:
666:
656:
654:
650:
646:
642:
638:
634:
624:
620:
618:
617:
610:
607:
605:
601:
590:
589:
587:
582:
581:Burmese amber
578:
574:
570:
566:
565:
554:
552:
547:
545:
541:
536:
534:
523:
521:
517:
513:
509:
504:
502:
498:
494:
491:
487:
483:
473:
471:
467:
457:
455:
451:
447:
443:
439:
435:
431:
427:
423:
419:
410:
406:
395:
393:
389:
385:
381:
377:
373:
369:
365:
361:
357:
353:
349:
345:
344:clown beetles
341:
337:
333:
325:
321:
318:
314:
306:
303:
300:Marseul, 1857
298:
295:
290:
287:
282:
279:
276:MacLeay, 1819
274:
273:Onthophilinae
271:
266:
263:
258:
255:
252:Marseul, 1857
250:
247:
244:Reitter, 1909
242:
239:
234:
231:
228:MacLeay, 1819
226:
223:
222:
220:
215:
210:
205:
202:
199:
198:
195:
192:
190:Superfamily:
189:
188:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
145:
142:
139:
138:
135:
132:
129:
128:
123:
118:
114:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
3659:
3538:Helophoridae
3530:Georissidae
3525:Epimetopidae
3490:
3441:Pleocomidae
3425:Ochodaeidae
3409:Hybosoridae
3393:Glaphyridae
3385:Geotrupidae
3363:
3360:Belohinidae
3321:Eucinetidae
3312:
3309:Decliniidae
3282:
3240:Phengodidae
3179:Cantharidae
3134:Dascillidae
3126:Dascilloidea
3108:Psephenidae
3100:Lutrochidae
3092:Limnichidae
3005:Buprestidae
2997:Buprestoidea
2937:Synchroidae
2921:Scraptiidae
2913:Salpingidae
2876:Prostomidae
2863:Oedemeridae
2839:Mordellidae
2757:Lymexylidae
2749:Lymexyloidea
2699:Attelabidae
2691:Anthribidae
2649:Smicripidae
2613:Phalacridae
2597:Nitidulidae
2592:Myraboliidae
2584:Monotomidae
2575:
2556:Kateretidae
2533:Cyclaxyridae
2493:
2490:Agapythidae
2454:Latridiidae
2404:Cerylonidae
2391:Anamorphidae
2325:Phycosecidae
2316:
2299:
2296:Metaxinidae
2249:Biphyllidae
2240:
2209:Orsodacnidae
2166:Cucujiformia
2099:Dermestidae
1969:
1908:Amphizoidae
1853:
1833:
1819:Archostemata
1803:Holometabola
1801:Superorder:
1795:Infraclass:
1683:Tree of life
1656:Hister nomas
1655:
1630:
1607:
1596:. Retrieved
1592:
1589:"Histeridae"
1583:
1572:. Retrieved
1568:
1559:
1539:
1526:
1496:
1471:
1467:
1461:
1452:
1443:
1430:
1417:
1384:
1380:
1374:
1366:
1321:
1317:
1307:
1275:
1251:
1241:
1232:
1223:
1212:
1142:
1098:
1084:
1070:
1056:
1042:
1028:
1014:
1000:
983:
974:
966:
961:
957:
953:
946:
927:
921:
901:
897:colonization
893:
884:
880:
878:
874:
870:
868:
856:
852:
850:
843:
839:
837:
830:
815:
806:
795:Please help
790:verification
787:
720:
707:
694:pigmentation
686:
677:Platylomalus
676:
672:
662:
630:
621:
614:
611:
608:
596:
584:
564:Antigracilus
562:
560:
548:
537:
529:
505:
501:antennomeres
479:
463:
415:
408:
347:
343:
331:
330:
305:Trypeticinae
297:Trypanaeinae
268:Fowler, 1912
217:Subfamilies
203:
180:Infraorder:
18:
3996:Histeroidea
3839:iNaturalist
3684:Wikispecies
3582:Hydraenidae
3556:Spercheidae
3543:Hydrochidae
3507:Synteliidae
3491:Histeridae
3483:Histeroidea
3433:Passalidae
3401:Glaresidae
3258:Throscidae
3218:(fireflies)
3216:Lampyridae
3203:Eucnemidae
3195:Elateridae
3155:Elateroidea
2971:Zopheridae
2855:Mycteridae
2786:Anthicidae
2657:Sphindidae
2641:Silvanidae
2551:Hobartiidae
2538:Erotylidae
2467:Mycetaeidae
2462:Murmidiidae
2446:Euxestidae
2340:Rentoniidae
2214:Oxypeltidae
2199:Disteniidae
2032:Lepiceridae
1961:Hygrobiidae
1953:Haliplidae
1937:Dytiscidae
1916:Aspidytidae
1697:(in German)
1690:German site
1375:Onthophilus
1270:2009-10-31.
840:Dendrobites
763:spermatheca
727:mesosternum
667:larva. The
627:Development
616:Psiloscelis
586:Onthophilus
510:within its
249:Haeteriinae
194:Histeroidea
3991:Histeridae
3985:Categories
3717:HISTERIDAE
3704:Histeridae
3690:Histeridae
3660:Histeridae
3603:Silphidae
3595:Ptiliidae
3587:Leiodidae
3574:Agyrtidae
3417:Lucanidae
3296:Scirtoidea
3232:Omethidae
3211:Jurasaidae
3060:Dryopidae
3031:Byrrhidae
3023:Byrrhoidea
2715:Brentidae
2520:Cucujidae
2502:Boganiidae
2482:Cucujoidea
2355:Thymalidae
2345:Rhadalidae
2288:Melyridae
2257:Byturidae
2219:Vesperidae
2128:Ptiniidae
1978:Noteridae
1945:Gyrinidae
1921:Carabidae
1850:Jurodidae
1842:Cupedidae
1789:Subclass:
1779:Arthropoda
1760:Coleoptera
1677:Histeridae
1598:2019-06-21
1574:2019-06-21
1387:: 104443.
1324:: e44985.
1121:References
938:arthropods
936:and other
881:Inquilines
755:copulation
743:ovipositor
731:ovipositor
698:locomotion
551:arthropods
392:predacious
388:houseflies
332:Histeridae
289:Tribalinae
281:Saprininae
265:Niponiinae
257:Histerinae
204:Histeridae
174:Coleoptera
154:Arthropoda
24:Histeridae
3457:Trogidae
3451:(scarabs)
3329:Scirtidae
3304:Clambidae
2897:Pythidae
2831:Meloidae
2778:Aderidae
2681:(weevils)
2546:Helotidae
2472:Teredidae
2386:Alexiidae
2270:Cleridae
2229:Cleroidea
2062:Polyphaga
2060:Suborder
2013:Myxophaga
2011:Suborder
1966:Meruidae
1885:Suborder
1867:Ommatidae
1817:Suborder
1791:Pterygota
1771:Kingdom:
1717:Bug Guide
1409:216238703
1340:2050-084X
853:Geobiotes
833:predators
769:Subclades
749:from the
723:sternites
600:olfaction
567:from the
520:Spiracles
516:hemolymph
493:mouthpart
460:Etymology
454:livestock
438:predators
430:prothorax
368:predatory
317:Diversity
225:Abraeinae
209:Gyllenhal
140:Kingdom:
134:Eukaryota
3748:BugGuide
3724:BioLib:
3669:Wikidata
3224:Lycidae
3068:Elmidae
2966:Ulodidae
2802:Boridae
2707:Belidae
2308:Peltidae
1901:families
1887:Adephaga
1797:Neoptera
1777:Phylum:
1773:Animalia
1762:families
1548:Archived
1515:Archived
1505:Archived
1488:19019588
1358:30990167
1296:Archived
1268:Archived
1260:Archived
1211:(eds.).
1179:21594053
1109:See also
735:oviducts
544:termites
512:hemocoel
497:antennae
490:mandible
486:tergites
434:antennae
426:tergites
376:forensic
364:antennae
360:tergites
200:Family:
150:Phylum:
144:Animalia
130:Domain:
3955:4779751
3675:Q596252
2815:Ciidae
2723:Caridae
1785:Insecta
1783:Class:
1758:Extant
1707:Summary
1389:Bibcode
1349:6467565
1170:3088472
1147:Bibcode
1143:ZooKeys
950:carrion
934:maggots
930:insects
757:. When
741:to the
739:ovaries
673:Epierus
669:chorion
604:Diptera
533:carrion
526:Habitat
476:Anatomy
470:Juvenal
422:abdomen
380:species
352:species
340:beetles
170:Order:
164:Insecta
160:Class:
3968:150732
3952:uBio:
3919:738371
3903:NZOR:
3896:110043
3870:114959
3857:114574
3792:1HISTF
1899:Extant
1486:
1407:
1356:
1346:
1338:
1266:>.
1207:&
1177:
1167:
969:larvae
912:Turkey
908:Ankara
751:testes
683:Larval
665:instar
569:Aptian
482:elytra
448:, and
446:larval
418:elytra
356:elytra
336:family
211:, 1808
3963:WoRMS
3940:Plazi
3932:69273
3852:IRMNG
3844:68869
3805:11102
3740:80462
1651:>.
1644:>.
1637:>.
1615:>.
1554:>.
1534:>.
1521:>.
1438:>.
1425:>.
1405:S2CID
1318:eLife
1302:>.
1246:1988.
864:mites
747:sperm
704:Pupal
653:wings
649:adult
641:larva
571:aged
450:adult
372:niche
334:is a
3891:NCBI
3865:ITIS
3831:4733
3826:GBIF
3787:EPPO
3753:6577
3735:BOLD
3727:5034
1593:GBIF
1484:PMID
1354:PMID
1336:ISSN
1175:PMID
956:and
860:dung
647:and
645:pupa
540:ants
480:The
384:dung
35:Preęž’
3878:NBN
3779:364
3774:EoL
3766:B44
3761:CoL
3712:AFD
3699:ADW
1476:doi
1472:183
1397:doi
1385:111
1344:PMC
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1165:PMC
1155:doi
906:in
879:4.
869:3.
851:2.
846:fly
838:1.
799:by
675:or
659:Egg
637:egg
442:egg
346:or
338:of
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90:N
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