Hadrosauridae - Knowledge

5546: 944: 860:, was published in 1942, and looked back at the whole of understanding about the family. It was designed as a definitive work, covering all aspects of their biology and evolution, and as part of it every known species was re-evaluated and many of them redescribed. They agreed with prior authors on the semi-aquatic nature of hadrosaurs, but re-evaluated Cope's idea of weak jaws and found quite the opposite. The teeth were rooted in strong batteries and would be continuously replaced to prevent them getting worn down. Such a system seemed incredibly overbuilt for the job of eating soft Mesozoic plants, and this fact confused the authors. Though they still proposed a diet of water plants, they considered it likely this would be supplemented by occasional forrays into browsing on land plants. 9353: 4024: 1553: 4212: 1630: 4647: 4623: 1909: 9348: 4549: 4244: 4079: 1734: 1657: 1580: 4137: 3952: 1512: 1451: 4486: 4424: 4275: 1837: 1803: 8438: 4671: 4367: 3822: 2682: 1997: 1936: 3993: 4455: 4702: 4168: 1485: 4574: 4299: 1700: 8452: 4726: 1970: 4055: 9335: 884:. It challenged the idea that hadrosaurs were semi-aquatic animals, which had been held since the work of Leidy back in the 1850s. This new approach was backed using evidence of the environment and climate they lived in, co-existing flora and fauna, physical anatomy, and preserved stomach contents from mummies. Based on evaluation of all this data, Ostrom found the idea that hadrosaurs were adapted for aquatic life incredibly lacking, and instead proposed they were capable terrestrial animals that browsed on plants such as 5431: 3116: 2185: 3351: 8445: 147: 3140: 9359: 5176: 3846: 2655: 3217: 2235: 3622: 2508: 3790: 3591: 3304: 2621: 2587: 2474: 2269: 3375: 3023: 2364: 2131: 775: 3750: 3726: 3439: 3054: 2303: 2158: 5327: 3280: 1183: 120: 4909: 1136: 666: 1166: 3647: 3085: 2535: 3496: 2982: 5109: 5071: 4977: 9342: 864: 7369: 650:), identified as hadrosaurines (Hadrosaurinae) in most pre-2010 works, which lacked hollow cranial crests (solid crests were present in some forms). Saurolophines tended to be bulkier than lambeosaurines. Lambeosaurines included the aralosaurins, tsintaosaurins, lambeosaurins and parasaurolophins, while saurolophines included the brachylophosaurins, kritosaurins, saurolophins and edmontosaurins. 5091:

As a result, the study determined that the hadrosaur diet was probably made of leaves and lacked the bulkier items, such as twigs or stems, that might have required a different chewing method and created different wear patterns. However, Purnell said these conclusions were less secure than the more conclusive evidence regarding the motion of teeth while chewing.

5127:, during the Late Cretaceous. It was concluded that hadrosaurids could reach low-growing trees and shrubs that were out of the reach of ceratopsids, ankylosaurs, and other small herbivores. Hadrosaurids were capable of feeding up to a height of 2 m (6 ft 7 in) when standing quadrupedally, and up to a height of 5 m (16 ft) bipedally. 888:. He remained uncertain, however, as to the purpose of the paddle-like hand Osborn had described, as well as their long and somewhat paddle-like tails. Thus he agreed with the idea that hadrosaurs would have taken refuge from predators in water. Numerous important studies would follow this; Ostrom's student Peter Dodson published a paper about 935:

catalyst for this boom. Hadrosaur research experienced high levels of diversity within the decade, with previously uncommon subjects such as growth, phylogeny, and biogeography experiencing more attention, though the functional morphology of hadrosaurids was found to have declined in study since the Dinosaur Renaissance.

5105:. As a result of that finding, Tweet concluded in September 2008 that the animal was likely a browser, not a grazer. In response to such findings, Purnell said that preserved stomach contents are questionable because they do not necessarily represent the usual diet of the animal. The issue remains a subject of debate. 5323:

like modern reptiles, with the ventral and lateral regions correlating fairly closely. Also unlike modern reptiles, the brains of the juveniles did not seem to correlate any closer to the brain wall than those of adults. It was cautioned, however, that very young individuals were not included in the study.

771:, giving a slightly better picture of the form of a hadrosaur. Leidy provided additional description in a 1865 paper. Among his 1858 work Leidy briefly suggested that the animal was likely amphibious in nature; this school of thought about hadrosaurs would come to be dominant for over a century to come. 5090:

found that hadrosaurs likely grazed on horsetails and vegetation close to the ground, rather than browsing higher-growing leaves and twigs. This conclusion was based on the evenness of scratches on hadrosaur teeth, which suggested the hadrosaur used the same series of jaw motions over and over again.

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level), related taxa, and previous predictions, the first such large-scale look into the neurology of the subfamily. Contra the early works, Evans' studies indicate that only some regions of the hadrosaur brain (the dorsal portion and much of the hindbrain) were loosely correlated to the brain wall,

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that would have resulted from transport, meaning the fossils were buried near their point of origin. Bonebeds 23, 28, 47, and 50 are productive sources of young hadrosaur remains in the formation, especially bonebed 50. The bones of juvenile hadrosaurs and fossil eggshell fragments are not known to

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The hypothesis that hadrosaurs were likely grazers rather than browsers appears to contradict previous findings from preserved stomach contents found in the fossilized guts in previous hadrosaur studies. The most recent such finding before the publication of the Purnell study was conducted in 2008,

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Alarcón-Muñoz, Jhonatan; Vargas, Alexander O.; Püschel, Hans P.; Soto-Acuña, Sergio; Manríquez, Leslie; Leppe, Marcelo; Kaluza, Jonatan; Milla, Verónica; Gutstein, Carolina S.; Palma-Liberona, José; Stinnesbeck, Wolfgang; Frey, Eberhard; Pino, Juan Pablo; Bajor, Dániel; Núñez, Elaine (2023-06-16).

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2016. This cladogram is a recent modification of the original 2010 analysis, including more characters and taxa. The resulting cladistic tree of their analysis was resolved using Maximum-Parsimony. 61 hadrosauroid species were included, characterized for 273 morphological features: 189 for cranial

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Tanke, D.H. and Brett-Surman, M.K. 2001. Evidence of Hatchling and Nestling-Size Hadrosaurs (Reptilia:Ornithischia) from Dinosaur Provincial Park (Dinosaur Park Formation: Campanian), Alberta, Canada. pp. 206-218. In: Mesozoic Vertebrate Life—New Research Inspired by the Paleontology of Philip J.

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The limbs of the juvenile hadrosaurs are anatomically and proportionally similar to those of adult animals. However, the joints often show "predepositional erosion or concave articular surfaces", which was probably due to the cartilaginous cap covering the ends of the bones. The pelvis of a young

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Evidence suggests that young hadrosaurs would have walked on only their two hind legs, while adults would have walked on all four. As the animal aged, the front limbs became more robust in order to take on weight, while the back legs became less robust as they transitioned to walking on all four

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zones, or areas where the teeth contact one another in chewing, whereas in adulthood there is a "dual function" arrangement with two distinct areas of different tooth wear. This change during growth may have helped transition from a diet of softer plants when young to more tough and fibrous ones

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as the source of data on the amount of works published in each decade. Various periods of high and low activity were found, but the twenty-first century was found to overwhelmingly be the most prolific time, with over two-hundred papers published. The advent of the internet was cited as a likely

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collaborated to arrange the International Hadrosaur Symposium, a professional meeting about ongoing hadrosaur research that was held at the latter institution on September 22 and 23 in 2011. Over fifty presentations were made at the event, thirty-six of which were later incorporated into a book,

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This information comes from the aforementioned Alan Boyle source from June 29, 2009. However, this specific information is not included in the body of the article, but rather a response by Boyle to comments in the article. Since the comments were written by Boyle himself, and since they cite

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In contrast with eggshell fossils, the remains of very young hadrosaurs are somewhat common. Tanke has observed that an experienced collector could discover multiple juvenile hadrosaur specimens in a single day. The most common remains of young hadrosaurs in the Dinosaur Park Formation are

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The most recognizable aspect of hadrosaur anatomy is the flattened and laterally stretched rostral bones, which gives the distinct duck-bill look. Some members of the hadrosaurs also had massive crests on their heads, probably for display and/or to make noises. In some genera, including

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clams and snails. This association is not a coincidence, as the invertebrate shells would have slowly dissolved and released enough basic calcium carbonate to protect the eggshells from naturally occurring acids that otherwise would have dissolved them and prevented fossilization.

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was informed of numerous large bones accidentally uncovered by farmer John E. Hopkins some twenty years earlier. Foulke obtained permission to investigate the now scattered fossils in 1858, and these specimens as well were given to Leidy. They were described in the same year as

5492:. However, the authors were unable to determine what specific factors shaped nesting ground choice in the formation's hadrosaurs. They suggested that behavior, diet, soil condition, and competition between dinosaur species all potentially influenced where hadrosaurs nested. 5972:

2014, Fuentes-Buxó, R. & Dalla Vecchia, F.M. 2014. Les Serretes, a new Late Maastrichtian dinosaur site in the eastern Tremp Syncline (Southern Pyrenees, Spain). In: Paleontologia i Evolució. Reconstructing the Terrestrial end-cretaceous paleoenvironments in Europe. pp

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for use in palaeontology has allowed for more widespread application of this without the need for specimen destruction. Modern research using these methods has focused largely on hadrosaurs. In a 2009 study by palaeontologist David C. Evans and colleagues, the brains of

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was questioned in a comprehensive study of hadrosaurid relationships by Albert Prieto-Márquez in 2010. Prieto-Márquez noted that, though the name Hadrosaurinae had been used for the clade of mostly crestless hadrosaurids by nearly all previous studies, its type species,

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of Asia, Europe and North America. However, the back of the mouth contained thousands of teeth suitable for grinding food before it was swallowed. This has been hypothesized to have been a crucial factor in the success of this group in the Cretaceous compared to the

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in 1912, who dubbed it the "Dinosaur mummy". This specimen's skin was almost completely preserved in the form of impressions. The skin around its hands, thought to represent webbing, was seen as further bolstering the idea that hadrosaurs were very aquatic animals.

819:. The Eastern states, too, would never yield particularly informative specimens. Instead, other sites in the American West would come to provide many very complete specimens that would form the backbone of hadrosaur research. One such specimen was the very complete 5236:). Though still obviously small, Ostrom recognized that the brains may be more significantly developed than expected, but supported the view that dinosaur brains would have only filled some of the endocranial cavity, limiting possibility of analysis. In 1977 5960:

Fuentes Buxó, R. 2012. Anàlisi de restes paleontològiques del Maastrichtià superior (Cretaci superior) procedents d’un nou jaciment situat en Les Serretes (Vilamitjana, Conca de Tremp, Espanya). Treball de fi de Master de Paleontologia (UAB-UB-ICP). 129

806:. Cope had studied the jaws of hadrosaurs and come to the conclusion that the teeth were fragile and could have been dislodged incredibly easily. As such, he supposed the animals must have fed largely on soft water plants; he presented this idea to the 626:

dinosaurs and had a similar body layout. Hadrosaurs were among the most dominant herbivores during the Late Cretaceous in Asia and North America, and during the close of the Cretaceous several lineages dispersed into Europe, Africa, and South America.

5382:, a close relative of the taxa from the 2009 study, was the subject of a 2013 paper once again looking into a cranial endocast. A nearly identical EQ range of 2.3 to 3.8 was found, and it was again noted this was higher than that of living reptiles, 5962: 810:

in 1883, and this idea would come to be very influential on future study. Research would continue in the Judith River area for years to come, but the formation never yielded much more than fragmentary remains, and Cope's species as well as

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and Nelda Wright to work on a similar project decades later. Eventually they realized the whole of Ornithopoda was too broad of a scope, until eventually it was narrowed down to specifically North American hadrosaurs. Their monograph,

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analysis has increased the resolution of hadrosaurid relationships considerably, leading to the widespread usage of tribes (a taxonomic unit below subfamily) to describe the finer relationships within each group of hadrosaurids.

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Case, Judd A.; Martin, James E.; Chaney, Dan S.; Regurero, Marcelo; Marenssi, Sergio A.; Santillana, Sergio M.; Woodburne, Michael O. (25 September 2000). "The first duck-billed dinosaur (family Hadrosauridae) from Antarctica".

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features and 84 for postcranial features. When characters had multiple states that formed an evolutionary scheme, they were ordered to account for the evolution of one state into the next. The final tree was run through

5145:, all of which would have been nutritious. Examination of hadrosaur coprolites from the Grand Staircase-Escalante indicates that shellfish such as crustaceans were also an important component of the hadrosaur diet. 4963:) subfamily have solid crests or none at all. Lambeosaurine crests had air chambers that may have produced a distinct sound and meant that their crests could have been used for both an audio and visual display. 5507:

in Montana, United States. While present, dinosaur eggshell is very rare in the Dinosaur Park Formation and is only found in two different microfossil sites. These sites are distinguished by large numbers of

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Pérez-Pueyo, Manuel; Cruzado-Caballero, Penélope; Moreno-Azanza, Miguel; Vila, Bernat; Castanera, Diego; Gasca, José Manuel; Puértolas-Pascual, Eduardo; Bádenas, Beatriz; Canudo, José Ignacio (2021-04-02).

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concluded that hadrosaurs nested in both the ancient upland and lowlands of the formation's depositional environment. The upland nesting grounds may have been preferred by the less common hadrosaurs, like

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during adulthood. It's thought the transition between states, characterized by a more gradual transition from one wear state to another, occurred at different times in the growth of different species; in

6104:"The Tetrapod Fossil Record from the Uppermost Maastrichtian of the Ibero-Armorican Island: An Integrative Review Based on the Outcrops of the Western Tremp Syncline (Aragón, Huesca Province, NE Spain)" 892:

skull anatomy that made enormous changes to hadrosaur taxonomy in 1975, and Michael K. Brett-Surman conducted a full revision of the group as part of his Graduate studies through the 1970s and 1980s.

5374:, an early bird, had very similar proportions. This lends further support to the idea of complex behaviours and relatively high intelligence, for non-avian dinosaurs, in hadrosaurids. 5137:

hadrosaurs show that the animals sometimes deliberately ate rotting wood. Wood itself is not nutritious, but decomposing wood would have contained fungi, decomposed wood material and

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Skin impressions of multiple hadrosaurs have been found. From these impressions, the hadrosaurs were determined to be scaled, and not feathered like some dinosaurs of other groups.

6911:"Computed tomographic investigation of a hatchling skull reveals ontogenetic changes in the dentition and occlusal surface morphology of Hadrosauridae (Dinosauria: Ornithischia)" 5608:

has been documented in the spine of a 78-million year old hadrosaurid. Other examples of pathologies in hadrosaurs include healed wounds from predators, such as those found in

967:. During the late early Maastrichtian, several lineages of Lambeosaurinae from Asia migrated into the European Ibero-Armorican Island (what is now France and Spain), including 5974: 6010:

Weishampel, David B. (2015). "A history of the study of ornithopods: Where have we been? Where are we now? Where are we going?". In Eberth, David A.; Evans, David C. (eds.).

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bone and a pubic bone which was positioned backwards in the pelvis. Unlike more primitive iguanodonts, the teeth of hadrosaurids are stacked into complex structures known as

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Chin, K. (September 2007). "The Paleobiological Implications of Herbivorous Dinosaur Coprolites from the Upper Cretaceous Two Medicine Formation of Montana: Why Eat Wood?".

5730:"Small hadrosaur manus and pes tracks from the Lower Cantwell Formation (Upper Cretaceous) Denali National Park, Alaska: implications for locomotion in juvenile hadrosaurs" 5495:

Sub-centimeter fragments of pebbly-textured hadrosaur eggshell have been reported from the Dinosaur Park Formation. This eggshell is similar to the hadrosaur eggshell of

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As with previous studies, EQ values were investigated, although a wider number range was given to account for uncertainty in brain and body mass. The range for the adult

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and other ornithischians, but different EQ estimates for theropods were cited, placing the hadrosaur numbers significantly below even more basal theropods like

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graduate student Justin S. Tweet found a homogeneous accumulation of millimeter-scale leaf fragments in the gut region of a well-preserved partially grown

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Dilkes, David W. (2001). "An ontogenetic perspective on locomotion in the Late Cretaceous dinosaur Maiasaura peeblesorum (Ornithischia: Hadrosauridae)".

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theropods. Reasonings suggested for their comparably high intelligence were the need for acute senses in the lack of defensive weapons, and more complex

916: 7397: 943: 5612:, and tumors such as Langerhans cell histiocytosis, hemangiomas, desmoplastic fibroma, metastatic cancer, and osteoblastomas, found in genera such as 920:, published in 2015. The volume was brought together primarily by palaeontologists David A. Eberth and David C. Evans, and featured an afterword from 5346:

dinosaurs (nearly all having EQs below two), but fell well short of maniraptorans themselves, which had quotients higher than four. The size of the

6303:"Diversity, relationships, and biogeography of the Lambeosaurine dinosaurs from the European archipelago, with description of the new aralosaurin 9857: 7387: 7445: 9870: 1071: 6635: 778:

From the mid 19th century through much of the 20th century, hadrosaurs were considered aquatic animals which subsisted on soft water plants

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about the rate of ornithopod research over history, and the interest in different aspects of it over that history, using the 2004 volume

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Hadrosaur research experienced a surge in the decade of the 2000s, similar to the research of other dinosaurs. In response to this, the

9939: 5496: 6544:"Quantitative analysis of dental microwear in hadrosaurid dinosaurs, and the implications for hypotheses of jaw mechanics and feeding" 6981:"Endocranial Anatomy of Lambeosaurine Hadrosaurids (Dinosauria: Ornithischia): A Sensorineural Perspective on Cranial Crest Function" 6758:"Feeding height stratification among the herbivorous dinosaurs from the Dinosaur Park Formation (upper Campanian) of Alberta, Canada" 6163:"The first duckbill dinosaur (Hadrosauridae: Lambeosaurinae) from Africa and the role of oceanic dispersal in dinosaur biogeography" 9831: 7306: 5148:

It's thought that chewing behaviour may have changed throughout life in hadrosaurids. Very young specimens show simple cup shapes

1070:, has almost always been excluded from the clade that bears its name, in violation of the rules for naming animals set out by the 4929:, the whole front of the skull was flat and broadened out to form a beak, which was ideal for clipping leaves and twigs from the 1090:, by Forster, in a 1997 abstract, as simply "Lambeosaurinae plus Hadrosaurinae and their most recent common ancestor". In 1998, 9944: 1132:

by default. Prieto-Marquez reviewed the phylogeny of Hadrosauridae in 2010, including many taxa potentially within the family.

924:, all of whom also contributed to one or more of the studies published therein. The first chapter of the volume was a study by 9352: 8437: 7424: 7348: 6019: 5994: 7392: 5038: 9929: 642:, which acted as effective grinding surfaces. Hadrosauridae is divided into two principal subfamilies: the lambeosaurines ( 7339:

Rothschild, Bruce; Tanke, Darren H. (2007). "Osteochondrosis is Late Cretaceous Hadrosauria". In Carpenter Kenneth (ed.).

5010: 851:, but never made much progress towards it before his death. This unrealized endeavor would come to be the inspiration for 9347: 8451: 6746:

information he received specifically from Purnell, they are as legitimate a source of information as the article itself.

5398:(with a range of 2.4 to 5.24, compared to only 1.6 in the 2009 study); more bird-like coelurosaurians theropods such as 9934: 6366:"A primitive hadrosaurid from southeastern North America and the origin and early evolution of 'duck-billed' dinosaurs" 5198:

dinosaurs as a whole. The brains of hadrosaurid dinosaurs have been studied as far back at the late 19th century, when

588:. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The 6666:

Tweet, Justin S.; Chin, Karen; Braman, Dennis R.; Murphy, Nate L. (2008). "Probable gut contents within a specimen of

6285: 5422:(both at 19%). This demonstrated a previously unrecognized level of variation in neuro-anatomy within Hadrosauridae. 5096: 5017: 5350:

was, for the first time, remarked upon. It was found to taking up around 43% of endocranial volume (not considering

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was found. Upon the fragments discovered he named seven new species in two genera, as well as assigning material to

9875: 6852:"Consumption of crustaceans by megaherbivorous dinosaurs: dietary flexibility and dinosaur life history strategies" 5905:

A revision of the Hadrosauridae (Reptilia: Ornithischia) and their evolution during the Campanian and Maastrichtian

731: 9334: 6718:"Plant-eating dinosaur spills his guts: Fossil suggests hadrosaur's last meal included lots of well-chewed leaves" 723: 716: 9713: 6717: 5414:(with 33%), though still distinctly larger than previously estimated numbers for more primitive iguanodonts like 5654: 4951:

The two major divisions of hadrosaurids are differentiated by their cranial ornamentation. While members of the

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Schmitz, L.; Motani, R. (2011). "Nocturnality in Dinosaurs Inferred from Scleral Ring and Orbit Morphology".

6460:"Standardized Terminology and Potential Taxonomic Utility for Hadrosaurid Skin Impressions: A Case Study for 996:. Since its creation, a major division has been recognized in the group between the hollow-crested subfamily 820: 560: 17: 5918:

Horner, J.R.; Makela, R. (1979). "Nest of juveniles provides evidence of family structure among dinosaurs".

1004:, historically known as Hadrosaurinae. Both of these have been robustly supported in all recent literature. 5083: 6412: 5006: 9358: 5078:

of hadrosaurs as semi-aquatic animals that could only chew soft water plants, a popular idea at the time.

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Longrich, Nicholas R.; Suberbiola, Xabier Pereda; Pyron, R. Alexander; Jalil, Nour-Eddine (April 2021).

146: 828: 679: 7035:"Cranial Endocast of the Lambeosaurine Hadrosaurid Amurosaurus riabinini from the Amur Region, Russia" 959:, a saurolophine hadrosaurid migrated into South America from North America, giving rise to the clade 951:

Hadrosaurids likely originated in North America, before shortly dispersing into Asia. During the late

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Currie. Edited by D.H. Tanke and K. Carpenter. Indiana University Press: Bloomington. xviii + 577 pp.

5305: 4023: 3415: 1552: 8444: 6211: 7524: 5241: 5155: 4640: 4517: 4211: 1629: 832: 657:, with the young of some species walking mostly on two legs and the adults walking mostly on four. 134: 6645: 6242:"Global phylogeny of Hadrosauridae (Dinosauria: Ornithopoda) using parsimony and Bayesian methods" 5985:

Eberth, David A.; Evans, David C. (2015). "Preface". In Eberth, David A.; Evans, David C. (eds.).

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have been preserved in association with each other, despite both being present in the formation.

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hadrosaurs, but far larger than in any ornithischians outside of Hadrosauriformes, and all large

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would come to be recognized as the first recognized hadrosaur specimens. Around the same time in

706: 129: 6038:"Relict duck-billed dinosaurs survived into the last age of the dinosaurs in subantarctic Chile" 5342:

was 2.3 to 3.7; the lowest end of this range was still higher than modern reptiles and most non-

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Brett-Surman, Michael K. (1979). "Phylogeny and paleobiogeography of hadrosaurian dinosaurs".

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posed with a nest; the naming of this genus was one of numerous important developments in the

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Ostrom, John H. (1961). "Cranial morphology of the hadrosaurian dinosaurs of North America".

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had stated EQs higher than seven. Additionally, the relative cerebral volume was only 30% in

5218: 5212:; only basic remarks were possible but it was noted that the organ was proportionally small. 5199: 5149: 4725: 4573: 4298: 3432: 1969: 1699: 990:

The family Hadrosauridae was first used by Edward Drinker Cope in 1869, then containing only

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The following cladogram is from Ramírez-Velasco (2022), including most recently named taxa.

9818: 9113: 7410: 7260: 7202: 7105: 7046: 6863: 6816: 6679: 6555: 6475: 6424: 6377: 6318: 6174: 6115: 6049: 5927: 5869: 5817: 5741: 5347: 5295: 5208: 5031: 3472: 3133: 910: 906: 874: 807: 9341: 6670:(Dinosauria: Hadrosauridae) from the Upper Cretaceous Judith River Formation of Montana". 5248:

to have an EQ of 1.5, above that of other ornithischians including earlier relatives like

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Twenty years later, in 1964, another very important work would be published, this time by

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Hopson, James A. (1977). "Relative brain size and behavior in archosaurian reptiles".

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have been noted as having the most complex brains among ornithopods, and indeed among

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This section relies far too heavily on the Tanke & Brett-Surman (2001) reference.

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Mallon, Jordan C.; Evans, David C.; Ryan, Michael J.; Anderson, Jason S. (2013).

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of hadrosaurids in 2009, the paleontologists Vincent Williams, Paul Barrett, and

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in 1854 through 1856, discovered the very first dinosaur fossils recognized from

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legs. Furthermore, the animals' front limbs were shorter than their back limbs.

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defined the clade Hadrosauridae as the most inclusive possible group containing

896:

would also begin to leave his impact on the field, including with the naming of

9708: 9703: 9641: 9578: 9451: 9300: 9263: 9245: 9190: 8822: 8638: 8631: 8560: 8482: 8469: 8397: 8385: 8354: 8329: 8193: 8115: 8076: 8062: 7939: 7850: 7772: 7750: 7743: 7484: 6909:

Warnock-Juteau, Trystan; Ryan, Michael; Patterson, Tim; Mallon, Jordan (2024).

6876: 6828: 6691: 6273: 5583: 5572: 5269: 5065: 4952: 3972: 3931: 3743: 3678: 3584: 3350: 3115: 2866: 2467: 2184: 1426: 1176: 1048: 997: 972: 930: 921: 893: 881: 643: 455: 223: 7272: 4955:

subfamily have hollow crests that differ depending on species, members of the

3139: 694:, who described and named them in 1856; two of the several species named were 9918: 9794: 9648: 9592: 9571: 9542: 9458: 9395: 9293: 9286: 9272: 9182: 9162: 9155: 9052: 9002: 8995: 8979: 8954: 8927: 8920: 8913: 8884: 8835: 8797: 8743: 8715: 8659: 8322: 8315: 8270: 8200: 8069: 7928: 7913: 7875: 7786: 7679: 7642: 7635: 7117: 6147: 6100: 6069: 5577: 5370: 5300: 5290: 5285: 5273: 5216:

would give a more informed analysis and review in 1961, pulling on data from

5195: 5112: 4960: 4956: 4913: 4541: 4339: 4205: 4181: 4130: 4072: 4017: 3986: 3814: 3783: 3368: 3273: 2961: 2951: 2675: 2614: 2580: 2357: 2100: 1902: 1760: 1727: 1623: 1573: 1546: 1505: 1411: 1215: 1193: 1118: 1017: 1001: 956: 687: 647: 616: 594: 463: 371: 77: 7343:. Bloomington and Indianapolis: Indiana University Press. pp. 171–183. 7214: 6774: 6568: 5175: 4948:, are noted for having their manus united in a fleshy, often nail-less pad. 3845: 3216: 2654: 2234: 732: 9556: 9532: 9520: 9486: 9479: 9430: 9423: 9229: 9173: 9148: 9120: 9009: 8811: 8776: 8722: 8652: 8624: 8510: 8503: 8343: 8308: 8235: 8228: 8150: 8091: 8046: 7948: 7822: 7815: 7764: 7728: 7686: 7536: 7475: 7280: 7222: 7078: 7006: 6895: 6793: 6587: 6507: 6350: 6087: 6061: 5472: 5388: 5364: 5355: 5343: 5315: 5310: 5265: 5261: 5250: 5237: 5160: 5142: 5087: 4566: 3570: 3196: 2975: 2938: 2914: 2330: 1693: 1332: 1269: 1171: 1124: 1052: 1022: 1005: 968: 964: 816: 754: 724: 717: 691: 683: 631: 623: 581: 434: 424: 392: 319: 299: 282: 207: 6301:

Prieto-Márquez, A.; Dalla Vecchia, F. M.; Gaete, R.; Galobart, À. (2013).

5830: 5805: 5753: 5314:(juvenile specimen ROM 758) were scanned and compared to each other (on a 3621: 2507: 714:". The former was based on a collection of teeth whilst the latter on two 9844: 9803: 9745: 9634: 9599: 9437: 9279: 9252: 9204: 9197: 9030: 9016: 8988: 8968: 8947: 8891: 8870: 8863: 8856: 8790: 8783: 8736: 8701: 8673: 8609: 8602: 8371: 8221: 8207: 8179: 8157: 8099: 7962: 7793: 7757: 7721: 7617: 7604: 7563: 7433: 6607: 5801: 5589: 5378: 5359: 5319: 5280: 5276:

behaviours as indicated by their acoustic and visual display structures.

5232: 5213: 4719: 4695: 4479: 4384: 4360: 4292: 4268: 3945: 3789: 3719: 3639: 3590: 3374: 3303: 3241: 3022: 2890: 2842: 2620: 2586: 2528: 2473: 2363: 2296: 2268: 2262: 2130: 1990: 1963: 1857: 1796: 1444: 1394: 1367: 1249: 1188: 1141: 1108: 1096: 1091: 1065: 992: 848: 361: 329: 239: 52: 6300: 6138: 5663: 3749: 3725: 3438: 3279: 3053: 2302: 2157: 1078:, and used the name Saurolophinae instead for the traditional grouping. 774: 9836: 9687: 9662: 9655: 9627: 9620: 9500: 9386: 9085: 9065: 9023: 8898: 8750: 8617: 8567: 8532: 8525: 8496: 8336: 8214: 8186: 8055: 8028: 8003: 7843: 7836: 7829: 7707: 6927: 6631: 6212:"Synopsis of the extinct batrachia, reptilia and aves of North America" 5672: 5594: 5394: 5326: 5191: 5134: 4664: 4448: 3016: 2124: 1929: 1305: 1182: 1106:(a well-known lambeosaurine), later emending the definition to include 889: 742: 635: 589: 381: 351: 119: 97: 62: 9849: 4908: 3646: 3084: 2534: 1165: 1135: 1074:. Prieto-Márquez defined Hadrosaurinae as just the lineage containing 665: 9669: 9585: 9564: 9402: 9099: 9037: 8546: 8518: 8378: 8242: 8172: 8143: 7969: 7955: 7698: 7244:

Rothschild, B.M.; Tanke, D.H.; Helbling II, M.; Martin, L.D. (2003).

7152: 6997: 6980: 6725: 5939: 5907:. Ph.D. dissertation. Washington, D.C.: George Washington University. 5881: 5806:"A reconsideration of the paleoecology of the hadrosaurian dinosaurs" 5551: 5256: 5130: 5124: 4945: 4417: 3495: 3047: 2981: 2151: 1830: 1128:) outside the family, which led them to define the family to include 952: 898: 869: 844: 758: 605: 566: 549: 158: 102: 46: 9765: 7398:"Mummified" Dinosaur Discovered In Montana, National Geographic 2002 7093: 6034: 5989:. Bloomington, Indiana: Indina University Press. pp. xiii–xiv. 5714: 5108: 5070: 4987: 4976: 646:), which had hollow cranial crests or tubes; and the saurolophines ( 9788: 9549: 9218: 9106: 8906: 8877: 8804: 8729: 8694: 8595: 8581: 8539: 8363: 8164: 8129: 8035: 8021: 7983: 7493: 7466: 7457: 5509: 5460: 5383: 5203: 5138: 4935: 4161: 3489: 3078: 1478: 885: 746: 577: 402: 191: 178: 92: 87: 72: 67: 57: 8842: 8277: 6908: 6542:

Williams, Vincent S.; Barrett, Paul M.; Purnell, Mark A. (2009).

6413:"Phylogenetic and biogeography analysis of Mexican hadrosauroids" 5525: 5516: 5464: 5441: 5123:(2013) examined herbivore coexistence on the island continent of 4048: 977: 863: 107: 82: 9823: 6410: 6014:. Bloomington, Indiana: Indiana University Press. pp. 2–7. 5593:) and modern birds and reptiles suggest that they may have been 8708: 8301: 7451: 4930: 168: 7402: 6850:

Chin, Karen; Feldmann, Rodney M.; Tashman, Jessica N. (2017).

6363: 8553: 8083: 6640: 6280:. Jefferson, North Carolina: McFarland & Co. p. 69. 5512: 5334:

brain in right lateral (A), dorsal (B), and ventral (C) views

1087: 1028: 340: 6160: 7921: 6364:

Prieto-Marquez, A.; Erickson, G.M.; Ebersole, J.A. (2016).

5655:

10.1671/0272-4634(2000)020[0612:tfdbdf]2.0.co;2

5453: 7338: 5159:

it did not occur until a nearly adult stage, whereas in a

5639: 5448:. Tiny hadrosaur footprints have been discovered in the 6755: 1012:

Lambeosaurines have also been traditionally split into

7307:"Evidence of predatory behavior by theropod dinosaurs" 7091: 6541: 5559:

hadrosaur was similar to that of an older individual.

798:

led an expedition to the Judith River Formation where

7341:

Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs

6665: 7364: 6009: 5444:

sized hadrosaur fossils have been documented in the

2672: 2645: 2611: 2577: 2525: 2498: 2464: 2433: 2354: 2327: 2293: 2259: 2225: 2175: 2148: 2121: 1987: 1960: 1926: 1899: 1854: 1827: 1793: 1757: 1724: 1690: 1647: 1620: 1570: 1543: 1502: 1475: 1441: 1423: 1408: 1391: 1364: 1347: 1329: 1302: 1266: 1246: 1112:, the type genus of the family. According to Horner 460: 452: 440: 430: 419: 408: 398: 387: 377: 367: 357: 346: 335: 325: 315: 289: 6849: 1039:, and Parasaurolophini as all those taxa closer to 7032: 6404: 6216:Transactions of the American Philosophical Society 5849:. M.A. thesis. Berkeley: University of California. 5847:The appendicular anatomy of hadrosaurian dinosaurs 5783:Lull, Richard Swann; and Wright, Nelda E. (1942). 5528:, bones from limbs and feet, as well as vertebral 5515:and other less common shelled invertebrates, like 5354:) in ROM 702. This is comparable to their size in 947:Map of various hadrosaur taxa across North America 6902: 5304:(juvenile specimen ROM 759 and subadult specimen 9916: 6944: 5597:, active throughout the day at short intervals. 7138: 6978: 6548:Proceedings of the National Academy of Sciences 6209: 5727: 5244:to the topic of dinosaur intelligence, finding 7508: 7237: 7028: 7026: 7024: 6739: 6239: 5163:specimen likely less than a year old from the 8480: 7418: 7295: 7192: 6940: 6938: 6711: 6709: 5721: 827:), recovered in 1908 by the fossil collector 7246:"Epidemiologic study of tumors in dinosaurs" 7178: 7176: 7174: 7172: 7170: 7168: 7166: 7164: 7162: 6974: 6972: 6970: 6968: 6626: 6624: 5917: 5902: 5859: 5844: 5532:. The material showed little or none of the 5179:A 1905 diagram showing the small size of an 5115:dentary with teeth, typical of hadrosauridae 4996:introducing citations to additional sources 7021: 6915:Vertebrate Anatomy Morphology Palaeontology 6603:"Study hints at what and how dinosaurs ate" 6458:Bell, P. R. (2012). Farke, Andrew A (ed.). 6003: 5984: 5978: 5499:in southern Alberta as well as that of the 4919:Oxford University Museum of Natural History 7615: 7425: 7411: 7132: 6935: 6706: 6659: 5566: 118: 7301: 7159: 7085: 7068: 7058: 6996: 6965: 6926: 6885: 6875: 6783: 6773: 6621: 6577: 6567: 6535: 6497: 6487: 6411:Ramírez-Velasco, Angel Alejandro (2022). 6340: 6330: 6257: 6246:Zoological Journal of the Linnean Society 6235: 6233: 6231: 6229: 6186: 6137: 6127: 6077: 5829: 5662: 5540: 1153:Below is a cladogram from Prieto-Marquez 7033:Lauters, Pascaline; et al. (2013). 6947:Annual Review of Ecology and Systematics 6203: 5544: 5429: 5325: 5174: 5107: 5069: 4986:Relevant discussion may be found on the 4907: 1181: 1164: 1134: 963:, which is closely related to the tribe 942: 862: 773: 664: 5796: 5794: 5792: 5785:Hadrosaurian Dinosaurs of North America 5392:(with an EQ range of 3.31 to 5.07) and 858:Hadrosaurian Dinosaurs of North America 660: 14: 9917: 7332: 6600: 6266: 6226: 5800: 5779: 5777: 5775: 5773: 5771: 5728:Fiorillo, A.R.; Tykoski, R.S. (2016). 5700: 5624:is also commonly found in hadrosaurs. 757:, on the other side of the continent, 615:. Hadrosaurids are descendants of the 592:family, which includes genera such as 9770: 9769: 9374: 8467: 7602: 7444: 7406: 6979:Evans, David C.; et al. (2009). 6715: 6630: 5896: 5853: 5838: 1086:Hadrosauridae was first defined as a 9375: 6806: 6457: 6357: 6272: 5789: 5696: 5694: 5692: 5690: 5187:, top) commented on in early sources 5133:(fossilized droppings) of some Late 4970: 1145:, the taxon named by Prieto-Marquez 7094:"The skull and brain of Claosaurus" 6959:10.1146/annurev.es.08.110177.002241 5768: 5410:, closer to that of theropods like 815:itself would in time be seen as of 690:. These specimens were obtained by 24: 8468: 6451: 6370:Journal of Vertebrate Paleontology 5703:Canadian Journal of Earth Sciences 5643:Journal of Vertebrate Paleontology 5167:the transition had already begun. 1027:. Lambeosaurini is defined as all 25: 9956: 9940:Taxa named by Edward Drinker Cope 7360: 5903:Brett-Surman, Michael K. (1989). 5845:Brett-Surman, Michael K. (1975). 5687: 5183:brain (bottom; alongside that of 5097:University of Colorado at Boulder 985: 556: 'stout, thick' and 9357: 9351: 9346: 9340: 9333: 8450: 8443: 8436: 7367: 6259:10.1111/j.1096-3642.2009.00617.x 5240:introduced the use of estimated 4975: 4724: 4700: 4669: 4645: 4621: 4572: 4547: 4484: 4453: 4422: 4365: 4297: 4273: 4242: 4210: 4166: 4135: 4077: 4053: 4022: 3991: 3950: 3844: 3820: 3788: 3748: 3724: 3645: 3620: 3589: 3494: 3437: 3373: 3349: 3302: 3278: 3215: 3138: 3114: 3083: 3052: 3021: 2980: 2680: 2653: 2619: 2585: 2533: 2506: 2472: 2362: 2301: 2267: 2233: 2183: 2156: 2129: 1995: 1968: 1934: 1907: 1835: 1801: 1732: 1698: 1655: 1628: 1578: 1551: 1510: 1483: 1449: 1100:(a well-known saurolophine) and 794:" would come from the East, and 145: 50: 9714:List of informal dinosaur names 7603: 7432: 7186: 7092:Marsh, Othniel Charles (1893). 6843: 6800: 6749: 6594: 6514: 6294: 6154: 6094: 6028: 5966: 5954: 5549:Juvenile specimen of the genus 5425: 5206:of a specimen then referred to 4966: 938: 9724:Timeline of hadrosaur research 7383:Timeline of hadrosaur research 5911: 5633: 5459:In a 2001 review of hadrosaur 5406:, significantly lower than in 5230:(then considered a synonym of 741:teeth turned out to belong to 682:, during expeditions near the 13: 1: 9945:Prehistoric dinosaur families 6601:Bryner, Jeanna (2009-06-29). 6437:10.1016/j.cretres.2022.105267 6390:10.1080/02724634.2015.1054495 6188:10.1016/j.cretres.2020.104678 5627: 5575:of several hadrosaur genera ( 782:Further discoveries such as " 7060:10.1371/journal.pone.0078899 6668:Brachylophosaurus canadensis 6489:10.1371/journal.pone.0031295 6332:10.1371/journal.pone.0069835 5600: 5463:and hatchling material from 5170: 5084:studying the chewing methods 1081: 843:Cope had planned to write a 7: 9930:Santonian first appearances 7098:American Journal of Science 6716:Lloyd, Robin (2008-09-25). 6278:Dinosaurs: The Encyclopedia 6129:10.3390/geosciences11040162 5810:American Journal of Science 27:Extinct family of dinosaurs 10: 9961: 6877:10.1038/s41598-017-11538-w 6829:10.2110/palo.2006.p06-087r 6692:10.2110/palo.2007.p07-044r 6522:"Hadrosaur Forelimb Study" 6240:Prieto-Márquez, A (2010). 5063: 4903: 982:a member of Arenysaurini. 829:Charles Hazelius Sternberg 680:Ferdinand Vandeveer Hayden 559: 542: 9935:Maastrichtian extinctions 9778: 9679: 9530: 9510: 9385: 9381: 9370: 9331: 9262: 9228: 9172: 9131: 9075: 9060: 9051: 8978: 8937: 8830: 8821: 8771: 8760: 8491: 8476: 8463: 8434: 8396: 8353: 8296: 8287: 8124: 8113: 8045: 8016: 8012: 8001: 7938: 7902: 7870: 7861: 7697: 7660: 7630: 7626: 7611: 7598: 7562: 7544: 7535: 7517: 7504: 7440: 7317:: 135–144. Archived from 7273:10.1007/s00114-003-0473-9 6464:from Canada and Mongolia" 5242:encephalization quotients 4716: 4692: 4685: 4661: 4637: 4613: 4606: 4563: 4538: 4531: 4514: 4507: 4500: 4476: 4469: 4445: 4438: 4414: 4407: 4381: 4357: 4350: 4343: 4289: 4265: 4258: 4233: 4226: 4202: 4195: 4185: 4158: 4151: 4127: 4120: 4069: 4045: 4038: 4014: 4007: 3983: 3976: 3966: 3942: 3935: 3836: 3811: 3804: 3780: 3773: 3740: 3716: 3709: 3702: 3689: 3682: 3636: 3612: 3605: 3581: 3574: 3564: 3486: 3469: 3462: 3429: 3416:Gryposaurus monumentensis 3412: 3405: 3398: 3365: 3341: 3334: 3327: 3294: 3270: 3263: 3256: 3238: 3231: 3207: 3200: 3190: 3130: 3106: 3099: 3075: 3068: 3044: 3037: 3013: 3006: 2996: 2972: 2965: 2955: 2935: 2928: 2911: 2904: 2887: 2880: 2863: 2856: 2839: 2832: 2669: 2642: 2635: 2608: 2601: 2574: 2567: 2522: 2495: 2488: 2461: 2454: 2447: 2430: 2423: 2351: 2344: 2324: 2317: 2290: 2283: 2256: 2249: 2222: 2215: 2208: 2172: 2145: 2118: 2111: 2104: 1984: 1957: 1950: 1923: 1896: 1889: 1851: 1824: 1817: 1790: 1783: 1776: 1755: 1748: 1721: 1714: 1687: 1680: 1644: 1617: 1610: 1603: 1567: 1540: 1533: 1526: 1499: 1472: 1465: 1438: 1431: 1416: 1388: 1381: 1361: 1354: 1326: 1319: 1299: 1292: 1263: 1243: 1236: 1229: 1211: 1204: 515:(Lull & Wright, 1942) 480: 473: 312: 307: 288: 281: 142:Scientific classification 140: 126: 117: 34: 7118:10.2475/ajs.s3-45.265.83 6524:. Palaeo-electronica.org 6210:Cope, Edward D. (1969). 5571:Comparisons between the 5436:Hypacrosaurus stebingeri 5156:Hypacrosaurus stebingeri 4641:Hypacrosaurus altispinus 4518:Hypacrosaurus stebingeri 833:Converse County, Wyoming 604:, was a common group of 135:Oxford University Museum 7215:10.1126/science.1200043 6775:10.1186/1472-6785-13-14 6569:10.1073/pnas.0812631106 5610:Edmontosaurus annectens 5567:Daily activity patterns 5505:Judith River Formations 5469:Dinosaur Park Formation 5362:dinosaurs; maniraptors 5260:and similar to that of 5181:Edmontosaurus annectens 5165:Dinosaur Park Formation 5059: 1037:Parasaurolophus walkeri 825:Edmontosaurus annectens 737:. Although most of the 712:Great Lignite Formation 707:Thespesius occidentalis 130:Edmontosaurus annectens 6636:"How dinosaurs chewed" 6062:10.1126/sciadv.adg2456 5555: 5541:Growth and development 5438: 5335: 5188: 5116: 5079: 4944:Hadrosaurs, much like 4921: 3298:Kritosaurus navajovius 1196: 1179: 1150: 948: 877: 835:. It was described by 831:and his three sons in 779: 702:Judith River Formation 676: 9897:Paleobiology Database 9694:Basturs Poble bonebed 6985:The Anatomical Record 5831:10.2475/ajs.262.8.975 5754:10.2110/palo.2016.049 5548: 5446:scientific literature 5433: 5329: 5264:theropods and modern 5228:Gryposaurus notabilis 5219:Edmontosaurus regalis 5200:Othniel Charles Marsh 5178: 5111: 5074:Early restoration by 5073: 4911: 3691:"Sabinosaur" PASAC-1 3433:Gryposaurus notabilis 1185: 1168: 1138: 1031:more closely related 946: 866: 791:Ornithotarsus immanis 777: 762:William Parker Foulke 668: 634:, hadrosaurids had a 580:, are members of the 9716:(contains hadrosaur 9114:Probrachylophosaurus 7141:Bulletin of the AMNH 6305:Canardia garonnensis 5434:Eggs of the species 5348:cerebral hemispheres 5209:Claosaurus annectens 5185:Triceratops horridus 4992:improve this article 3473:Gryposaurus latidens 3134:Probrachylophosaurus 2436:Sabinosaur" PASAC-1 1192:, the type taxon of 1175:, the type taxon of 1058:The use of the term 911:Royal Tyrrell Museum 907:Royal Ontario Museum 875:Dinosaur Renaissance 808:Philadelphia Academy 730: 722: 715: 661:History of discovery 573: 'lizard'), or 127:Mounted skeleton of 7265:2003NW.....90..495R 7253:Naturwissenschaften 7207:2011Sci...332..705S 7110:1893AmJS...45...83M 7051:2013PLoSO...878899L 6868:2017NatSR...711163C 6821:2007Palai..22..554C 6728:on February 4, 2016 6684:2008Palai..23..624T 6560:2009PNAS..10611194W 6554:(27): 11194–11199. 6480:2012PLoSO...731295B 6429:2022CrRes.13805267R 6417:Cretaceous Research 6382:2016JVPal..36E4495P 6323:2013PLoSO...869835P 6179:2021CrRes.12004678L 6167:Cretaceous Research 6120:2021Geosc..11..162P 6054:2023SciA....9G2456A 5932:1979Natur.282..296H 5874:1979Natur.277..560B 5822:1964AmJS..262..975O 5746:2016Palai..31..479F 5606:Spondyloarthropathy 5450:Blackhawk Formation 5095:when a team led by 3274:Kritosaurus horneri 1055:have also emerged. 1033:Lambeosaurus lambei 926:David B. Weishampel 796:Edward Drinker Cope 768:Hadrosaurus foulkii 697:Trachodon mirabilis 672:Trachodon mirabilis 294:Hadrosaurus foulkii 9466:Ornithopodichnites 9077:Brachylophosaurini 7662:Hypsilophodontidae 7303:Carpenter, Kenneth 6928:10.18435/vamp29395 6856:Scientific Reports 5556: 5477:M. K. Brett-Surman 5439: 5336: 5268:but below that of 5189: 5117: 5080: 4922: 3003:Brachylophosaurini 1197: 1180: 1151: 1047:. In recent years 1000:and the subfamily 949: 878: 853:Richard Swann Lull 780: 677: 655:facultative bipeds 653:Hadrosaurids were 9912: 9911: 9884:Open Tree of Life 9772:Taxon identifiers 9763: 9762: 9759: 9758: 9755: 9754: 9473:Shiraminesauropus 9366: 9365: 9329: 9328: 9325: 9324: 9321: 9320: 9317: 9316: 9212:Naashoibitosaurus 9133:Austrokritosauria 9093:Brachylophosaurus 9047: 9046: 8459: 8458: 8432: 8431: 8428: 8427: 8424: 8423: 8420: 8419: 8416: 8415: 8109: 8108: 7997: 7996: 7993: 7992: 7977:Transylvanosaurus 7863:Rhabdodontomorpha 7809:Notohypsilophodon 7780:Macrogryphosaurus 7594: 7593: 7590: 7589: 7586: 7585: 7582: 7581: 7388:UCMP Tree of Life 7350:978-0-253-34817-3 7201:(6030): 705–708. 6021:978-0-253-01385-9 5996:978-0-253-01385-9 5926:(5736): 296–298. 5868:(5697): 560–562. 5614:Brachylophosaurus 5483:Brachylophosaurus 5288:hadrosaur genera 5102:Brachylophosaurus 5076:Charles R. Knight 5057: 5056: 5042: 4900: 4899: 4891: 4890: 4882: 4881: 4873: 4872: 4864: 4863: 4855: 4854: 4846: 4845: 4837: 4836: 4828: 4827: 4819: 4818: 4810: 4809: 4801: 4800: 4792: 4791: 4783: 4782: 4774: 4773: 4765: 4764: 4756: 4755: 4747: 4746: 4738: 4737: 4595: 4594: 4586: 4585: 4396: 4395: 4329: 4328: 4320: 4319: 4311: 4310: 4109: 4108: 4100: 4099: 4091: 4090: 3921: 3920: 3912: 3911: 3903: 3902: 3894: 3893: 3885: 3884: 3876: 3875: 3867: 3866: 3858: 3857: 3762: 3761: 3668: 3667: 3659: 3658: 3553: 3552: 3544: 3543: 3535: 3534: 3526: 3525: 3517: 3516: 3508: 3507: 3451: 3450: 3387: 3386: 3316: 3315: 3211:Naashoibitosaurus 3179: 3178: 3170: 3169: 3161: 3160: 3152: 3151: 3110:Brachylophosaurus 2820: 2819: 2811: 2810: 2802: 2801: 2793: 2792: 2784: 2783: 2775: 2774: 2766: 2765: 2757: 2756: 2748: 2747: 2739: 2738: 2730: 2729: 2721: 2720: 2712: 2711: 2703: 2702: 2694: 2693: 2556: 2555: 2547: 2546: 2412: 2411: 2403: 2402: 2394: 2393: 2385: 2384: 2376: 2375: 2229:Naashoibitosaurus 2197: 2196: 2179:Brachylophosaurus 2090: 2089: 2081: 2080: 2072: 2071: 2063: 2062: 2054: 2053: 2045: 2044: 2036: 2035: 2027: 2026: 2018: 2017: 2009: 2008: 1878: 1877: 1869: 1868: 1669: 1668: 1592: 1591: 1281: 1280: 961:Austrokritosauria 817:doubtful validity 785:Hadrosaurus minor 530: 529: 277: 16:(Redirected from 9952: 9905: 9904: 9892: 9891: 9879: 9878: 9866: 9865: 9853: 9852: 9840: 9839: 9827: 9826: 9814: 9813: 9812: 9799: 9798: 9797: 9767: 9766: 9740:mummy S.M. R4036 9614:Microhadrosaurus 9607:Mandschurosaurus 9417:Camptosaurichnus 9410:Bonaparteichnium 9383: 9382: 9376:Related articles 9372: 9371: 9361: 9355: 9350: 9345: 9344: 9337: 9058: 9057: 8939:Parasaurolophini 8850:Angulomastacator 8828: 8827: 8769: 8768: 8688:Shuangmiaosaurus 8589:Huehuecanauhtlus 8575:Gongpoquansaurus 8489: 8488: 8478: 8477: 8465: 8464: 8454: 8448: 8447: 8440: 8294: 8293: 8289:Hadrosauriformes 8257:Ratchasimasaurus 8122: 8121: 8014: 8013: 8010: 8009: 7892:Muttaburrasaurus 7868: 7867: 7801:Muttaburrasaurus 7715:Atlascopcosaurus 7650:Thescelosauridae 7628: 7627: 7624: 7623: 7613: 7612: 7600: 7599: 7542: 7541: 7515: 7514: 7506: 7505: 7499: 7498: 7442: 7441: 7427: 7420: 7413: 7404: 7403: 7377: 7375:Dinosaurs portal 7372: 7371: 7370: 7355: 7354: 7336: 7330: 7329: 7327: 7326: 7299: 7293: 7292: 7250: 7241: 7235: 7234: 7190: 7184: 7180: 7157: 7156: 7136: 7130: 7129: 7089: 7083: 7082: 7072: 7062: 7030: 7019: 7018: 7000: 6998:10.1002/ar.20984 6991:(9): 1315–1337. 6976: 6963: 6962: 6942: 6933: 6932: 6930: 6906: 6900: 6899: 6889: 6879: 6847: 6841: 6840: 6804: 6798: 6797: 6787: 6777: 6753: 6747: 6743: 6737: 6736: 6734: 6733: 6724:. Archived from 6713: 6704: 6703: 6663: 6657: 6656: 6654: 6653: 6644:. Archived from 6628: 6619: 6618: 6616: 6615: 6598: 6592: 6591: 6581: 6571: 6539: 6533: 6532: 6530: 6529: 6518: 6512: 6511: 6501: 6491: 6455: 6449: 6448: 6408: 6402: 6401: 6361: 6355: 6354: 6344: 6334: 6298: 6292: 6291: 6270: 6264: 6263: 6261: 6237: 6224: 6223: 6207: 6201: 6200: 6190: 6158: 6152: 6151: 6141: 6131: 6098: 6092: 6091: 6081: 6048:(24): eadg2456. 6042:Science Advances 6032: 6026: 6025: 6007: 6001: 6000: 5982: 5976: 5970: 5964: 5958: 5952: 5951: 5940:10.1038/282296a0 5915: 5909: 5908: 5900: 5894: 5893: 5882:10.1038/277560a0 5857: 5851: 5850: 5842: 5836: 5835: 5833: 5798: 5787: 5781: 5766: 5765: 5725: 5719: 5718: 5709:(8): 1205–1227. 5698: 5685: 5684: 5666: 5637: 5294:(adult specimen 5052: 5049: 5043: 5041: 5000: 4979: 4971: 4729: 4728: 4705: 4704: 4688: 4687: 4674: 4673: 4650: 4649: 4626: 4625: 4617:Angulomastacator 4609: 4608: 4577: 4576: 4552: 4551: 4534: 4533: 4510: 4509: 4503: 4502: 4489: 4488: 4472: 4471: 4458: 4457: 4441: 4440: 4427: 4426: 4410: 4409: 4370: 4369: 4353: 4352: 4346: 4345: 4302: 4301: 4278: 4277: 4261: 4260: 4247: 4246: 4229: 4228: 4215: 4214: 4198: 4197: 4192:Parasaurolophini 4188: 4187: 4171: 4170: 4154: 4153: 4140: 4139: 4123: 4122: 4082: 4081: 4058: 4057: 4041: 4040: 4027: 4026: 4010: 4009: 3996: 3995: 3979: 3978: 3969: 3968: 3955: 3954: 3938: 3937: 3849: 3848: 3825: 3824: 3807: 3806: 3793: 3792: 3776: 3775: 3753: 3752: 3729: 3728: 3712: 3711: 3705: 3704: 3685: 3684: 3650: 3649: 3625: 3624: 3608: 3607: 3594: 3593: 3577: 3576: 3567: 3566: 3499: 3498: 3465: 3464: 3442: 3441: 3408: 3407: 3401: 3400: 3378: 3377: 3354: 3353: 3337: 3336: 3330: 3329: 3307: 3306: 3283: 3282: 3266: 3265: 3259: 3258: 3234: 3233: 3220: 3219: 3203: 3202: 3193: 3192: 3143: 3142: 3119: 3118: 3102: 3101: 3088: 3087: 3071: 3070: 3057: 3056: 3040: 3039: 3026: 3025: 3009: 3008: 2999: 2998: 2985: 2984: 2968: 2967: 2958: 2957: 2931: 2930: 2907: 2906: 2883: 2882: 2859: 2858: 2835: 2834: 2827: 2826: 2685: 2684: 2674: 2658: 2657: 2647: 2638: 2637: 2624: 2623: 2613: 2604: 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1014:Parasaurolophini 847:about the group 734: 726: 719: 669:Illustration of 640:dental batteries 621:Early Cretaceous 570: 563: 553: 546: 524: 516: 508: 500: 492: 462: 454: 442: 432: 421: 410: 400: 389: 379: 369: 359: 348: 337: 327: 317: 291: 272: 267: 256:Hadrosauromorpha 254: 238: 222: 206: 150: 149: 122: 112: 49: 38:Temporal range: 32: 31: 21: 9960: 9959: 9955: 9954: 9953: 9951: 9950: 9949: 9915: 9914: 9913: 9908: 9900: 9895: 9887: 9882: 9874: 9869: 9861: 9856: 9848: 9843: 9835: 9830: 9822: 9817: 9808: 9807: 9802: 9793: 9792: 9787: 9774: 9764: 9751: 9732:mummy AMNH 5060 9699:Dakota (fossil) 9675: 9526: 9506: 9494:Tetrapodosaurus 9445:Hadrosaurichnus 9377: 9362: 9339: 9313: 9308:Shantungosaurus 9258: 9239:Augustynolophus 9224: 9168: 9142:Bonapartesaurus 9127: 9071: 9043: 8974: 8962:Parasaurolophus 8933: 8817: 8765: 8756: 8681:Qianjiangsaurus 8667:Probactrosaurus 8646:Penelopognathus 8485: 8472: 8455: 8442: 8412: 8392: 8349: 8283: 8264:Riabininohadros 8137:Bayannurosaurus 8118: 8105: 8041: 8006: 7989: 7934: 7904:Tenontosauridae 7898: 7857: 7693: 7672:Gideonmantellia 7656: 7620: 7607: 7578: 7558: 7531: 7525:Avemetatarsalia 7510:Avemetatarsalia 7500: 7447: 7446: 7436: 7431: 7373: 7368: 7366: 7363: 7358: 7351: 7337: 7333: 7324: 7322: 7300: 7296: 7259:(11): 495–500. 7248: 7242: 7238: 7191: 7187: 7181: 7160: 7137: 7133: 7090: 7086: 7031: 7022: 6977: 6966: 6943: 6936: 6907: 6903: 6848: 6844: 6805: 6801: 6754: 6750: 6744: 6740: 6731: 6729: 6714: 6707: 6664: 6660: 6651: 6649: 6629: 6622: 6613: 6611: 6599: 6595: 6540: 6536: 6527: 6525: 6520: 6519: 6515: 6456: 6452: 6409: 6405: 6376:(2): e1054495. 6362: 6358: 6299: 6295: 6288: 6274:Glut, Donald F. 6271: 6267: 6238: 6227: 6208: 6204: 6159: 6155: 6099: 6095: 6033: 6029: 6022: 6008: 6004: 5997: 5983: 5979: 5971: 5967: 5959: 5955: 5916: 5912: 5901: 5897: 5858: 5854: 5843: 5839: 5802:Ostrom, John H. 5799: 5790: 5782: 5769: 5740:(10): 479–482. 5726: 5722: 5715:10.1139/e01-016 5699: 5688: 5638: 5634: 5630: 5622:Osteochondrosis 5603: 5569: 5543: 5489:Parasaurolophus 5428: 5352:olfactory bulbs 5330:Endocast of an 5173: 5068: 5062: 5053: 5047: 5044: 5007:"Hadrosauridae" 5001: 4999: 4985: 4980: 4969: 4906: 4901: 4892: 4883: 4874: 4865: 4856: 4847: 4838: 4829: 4820: 4811: 4802: 4793: 4784: 4775: 4766: 4757: 4748: 4739: 4723: 4699: 4668: 4644: 4620: 4596: 4587: 4571: 4546: 4483: 4452: 4421: 4397: 4364: 4330: 4321: 4312: 4296: 4272: 4241: 4237:Parasaurolophus 4209: 4165: 4134: 4110: 4101: 4092: 4076: 4052: 4021: 3990: 3949: 3922: 3913: 3904: 3895: 3886: 3877: 3868: 3859: 3843: 3840:Shantungosaurus 3819: 3787: 3763: 3747: 3723: 3669: 3660: 3644: 3619: 3616:Augustynolophus 3588: 3554: 3545: 3536: 3527: 3518: 3509: 3493: 3452: 3436: 3388: 3372: 3348: 3345:Bonapartesaurus 3317: 3301: 3277: 3214: 3180: 3171: 3162: 3153: 3137: 3113: 3082: 3051: 3020: 2979: 2821: 2812: 2803: 2794: 2785: 2776: 2767: 2758: 2749: 2740: 2731: 2722: 2713: 2704: 2695: 2679: 2652: 2649:Shantungosaurus 2618: 2584: 2557: 2548: 2532: 2505: 2502:Augustynolophus 2471: 2413: 2404: 2395: 2386: 2377: 2361: 2300: 2266: 2232: 2198: 2182: 2155: 2128: 2091: 2082: 2073: 2064: 2055: 2046: 2037: 2028: 2019: 2010: 1994: 1967: 1933: 1906: 1879: 1870: 1834: 1800: 1731: 1697: 1670: 1654: 1651:Parasaurolophus 1627: 1593: 1577: 1550: 1509: 1482: 1448: 1429: 1414: 1352: 1282: 1160:TNT version 1.0 1103:Parasaurolophus 1084: 988: 941: 751:T. occidentalis 749:and remains of 736: 728: 721: 663: 610:Late Cretaceous 601:Parasaurolophus 522: 520:Ornithotarsidae 514: 506: 498: 486: 414:Penelopognathus 303: 297: 271: 265: 252: 236: 220: 204: 144: 113: 111: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 44: 43: 40:Late Cretaceous 36: 28: 23: 22: 15: 12: 11: 5: 9958: 9948: 9947: 9942: 9937: 9932: 9927: 9910: 9909: 9907: 9906: 9893: 9880: 9867: 9854: 9841: 9828: 9815: 9800: 9784: 9782: 9776: 9775: 9761: 9760: 9757: 9756: 9753: 9752: 9750: 9749: 9742: 9734: 9726: 9721: 9711: 9709:Hypsilophodont 9706: 9704:Hadrosaur diet 9701: 9696: 9691: 9683: 9681: 9680:Other articles 9677: 9676: 9674: 9673: 9666: 9659: 9652: 9645: 9642:Stephanosaurus 9638: 9631: 9624: 9617: 9610: 9603: 9596: 9589: 9582: 9579:Fulgurotherium 9575: 9568: 9561: 9553: 9546: 9538: 9536: 9528: 9527: 9525: 9524: 9516: 9514: 9508: 9507: 9505: 9504: 9497: 9490: 9483: 9476: 9469: 9462: 9455: 9452:Iguanodontipus 9448: 9441: 9434: 9427: 9420: 9413: 9406: 9399: 9391: 9389: 9379: 9378: 9368: 9367: 9364: 9363: 9332: 9330: 9327: 9326: 9323: 9322: 9319: 9318: 9315: 9314: 9312: 9311: 9304: 9301:Laiyangosaurus 9297: 9290: 9283: 9276: 9268: 9266: 9264:Edmontosaurini 9260: 9259: 9257: 9256: 9249: 9246:Prosaurolophus 9242: 9234: 9232: 9226: 9225: 9223: 9222: 9215: 9208: 9201: 9194: 9191:Coahuilasaurus 9187: 9178: 9176: 9170: 9169: 9167: 9166: 9159: 9152: 9145: 9137: 9135: 9129: 9128: 9126: 9125: 9117: 9110: 9103: 9096: 9089: 9081: 9079: 9073: 9072: 9070: 9069: 9061: 9055: 9049: 9048: 9045: 9044: 9042: 9041: 9034: 9027: 9020: 9013: 9006: 8999: 8992: 8984: 8982: 8976: 8975: 8973: 8972: 8965: 8958: 8951: 8943: 8941: 8935: 8934: 8932: 8931: 8924: 8917: 8910: 8903: 8895: 8888: 8881: 8874: 8867: 8860: 8853: 8846: 8839: 8831: 8825: 8823:Lambeosaurinae 8819: 8818: 8816: 8815: 8808: 8801: 8794: 8787: 8780: 8772: 8766: 8761: 8758: 8757: 8755: 8754: 8747: 8740: 8733: 8726: 8719: 8712: 8705: 8698: 8691: 8684: 8677: 8670: 8663: 8656: 8649: 8642: 8639:Nanyangosaurus 8635: 8632:Nanningosaurus 8628: 8621: 8614: 8606: 8599: 8592: 8585: 8578: 8571: 8564: 8561:Gilmoreosaurus 8557: 8550: 8543: 8536: 8529: 8522: 8515: 8507: 8500: 8492: 8486: 8483:Hadrosauroidea 8481: 8474: 8473: 8470:Hadrosauroidea 8461: 8460: 8457: 8456: 8435: 8433: 8430: 8429: 8426: 8425: 8422: 8421: 8418: 8417: 8414: 8413: 8411: 8410: 8409: 8408: 8402: 8400: 8398:Hadrosauroidea 8394: 8393: 8391: 8390: 8386:Mantellisaurus 8382: 8375: 8368: 8359: 8357: 8355:Iguanodontidae 8351: 8350: 8348: 8347: 8340: 8333: 8330:Mantellisaurus 8326: 8319: 8312: 8305: 8297: 8291: 8285: 8284: 8282: 8281: 8274: 8267: 8260: 8253: 8246: 8239: 8232: 8225: 8218: 8211: 8204: 8197: 8194:Iguanacolossus 8190: 8183: 8176: 8169: 8161: 8154: 8147: 8140: 8133: 8125: 8119: 8116:Ankylopollexia 8114: 8111: 8110: 8107: 8106: 8104: 8103: 8096: 8088: 8080: 8077:Eousdryosaurus 8073: 8066: 8063:Dysalotosaurus 8059: 8051: 8049: 8043: 8042: 8040: 8039: 8032: 8025: 8017: 8007: 8002: 7999: 7998: 7995: 7994: 7991: 7990: 7988: 7987: 7980: 7973: 7966: 7959: 7952: 7944: 7942: 7940:Rhabdodontidae 7936: 7935: 7933: 7932: 7925: 7918: 7909: 7907: 7900: 7899: 7897: 7896: 7888: 7880: 7871: 7865: 7859: 7858: 7856: 7855: 7851:Weewarrasaurus 7847: 7840: 7833: 7826: 7819: 7812: 7805: 7797: 7790: 7783: 7776: 7773:Leaellynasaura 7769: 7761: 7754: 7751:Gasparinisaura 7747: 7744:Galleonosaurus 7740: 7732: 7725: 7718: 7711: 7703: 7701: 7695: 7694: 7692: 7691: 7683: 7676: 7667: 7665: 7658: 7657: 7655: 7654: 7646: 7639: 7631: 7621: 7616: 7609: 7608: 7596: 7595: 7592: 7591: 7588: 7587: 7584: 7583: 7580: 7579: 7577: 7576: 7575: 7574: 7568: 7566: 7560: 7559: 7557: 7556: 7555: 7554: 7545: 7539: 7533: 7532: 7530: 7529: 7528: 7527: 7518: 7512: 7502: 7501: 7497: 7496: 7487: 7485:Neornithischia 7478: 7469: 7460: 7454: 7438: 7437: 7430: 7429: 7422: 7415: 7407: 7401: 7400: 7395: 7390: 7385: 7379: 7378: 7362: 7361:External links 7359: 7357: 7356: 7349: 7331: 7294: 7236: 7185: 7158: 7131: 7104:(265): 83–86. 7084: 7045:(11): e78899. 7020: 6964: 6953:(1): 429–448. 6934: 6901: 6842: 6799: 6748: 6738: 6705: 6678:(9): 624–635. 6658: 6634:(2009-06-29). 6620: 6593: 6534: 6513: 6450: 6403: 6356: 6293: 6286: 6265: 6252:(2): 435–502. 6225: 6202: 6153: 6093: 6027: 6020: 6002: 5995: 5977: 5965: 5953: 5910: 5895: 5852: 5837: 5816:(8): 975–997. 5788: 5767: 5720: 5686: 5649:(3): 612–614. 5631: 5629: 5626: 5602: 5599: 5584:Prosaurolophus 5568: 5565: 5542: 5539: 5497:Devil's Coulee 5427: 5424: 5279:The advent of 5270:coelurosaurian 5172: 5169: 5066:Hadrosaur diet 5064:Main article: 5061: 5058: 5055: 5054: 4990:. Please help 4983: 4981: 4974: 4968: 4965: 4953:Lambeosaurinae 4905: 4902: 4898: 4897: 4894: 4893: 4889: 4888: 4885: 4884: 4880: 4879: 4876: 4875: 4871: 4870: 4867: 4866: 4862: 4861: 4858: 4857: 4853: 4852: 4849: 4848: 4844: 4843: 4840: 4839: 4835: 4834: 4831: 4830: 4826: 4825: 4822: 4821: 4817: 4816: 4813: 4812: 4808: 4807: 4804: 4803: 4799: 4798: 4795: 4794: 4790: 4789: 4786: 4785: 4781: 4780: 4777: 4776: 4772: 4771: 4768: 4767: 4763: 4762: 4759: 4758: 4754: 4753: 4750: 4749: 4745: 4744: 4741: 4740: 4736: 4735: 4732: 4731: 4715: 4712: 4711: 4708: 4707: 4691: 4686: 4684: 4681: 4680: 4677: 4676: 4660: 4657: 4656: 4653: 4652: 4636: 4633: 4632: 4629: 4628: 4612: 4607: 4605: 4602: 4601: 4598: 4597: 4593: 4592: 4589: 4588: 4584: 4583: 4580: 4579: 4562: 4559: 4558: 4555: 4554: 4537: 4532: 4530: 4527: 4526: 4523: 4522: 4513: 4508: 4506: 4501: 4499: 4496: 4495: 4492: 4491: 4475: 4470: 4468: 4465: 4464: 4461: 4460: 4444: 4439: 4437: 4434: 4433: 4430: 4429: 4413: 4408: 4406: 4403: 4402: 4399: 4398: 4394: 4393: 4390: 4389: 4380: 4377: 4376: 4373: 4372: 4356: 4351: 4349: 4344: 4342: 4336: 4335: 4332: 4331: 4327: 4326: 4323: 4322: 4318: 4317: 4314: 4313: 4309: 4308: 4305: 4304: 4288: 4285: 4284: 4281: 4280: 4264: 4259: 4257: 4254: 4253: 4250: 4249: 4232: 4227: 4225: 4222: 4221: 4218: 4217: 4201: 4196: 4194: 4186: 4184: 4178: 4177: 4174: 4173: 4157: 4152: 4150: 4147: 4146: 4143: 4142: 4126: 4121: 4119: 4116: 4115: 4112: 4111: 4107: 4106: 4103: 4102: 4098: 4097: 4094: 4093: 4089: 4088: 4085: 4084: 4068: 4065: 4064: 4061: 4060: 4044: 4039: 4037: 4034: 4033: 4030: 4029: 4013: 4008: 4006: 4003: 4002: 3999: 3998: 3982: 3977: 3975: 3973:Tsintaosaurini 3967: 3965: 3962: 3961: 3958: 3957: 3941: 3936: 3934: 3932:Lambeosaurinae 3928: 3927: 3924: 3923: 3919: 3918: 3915: 3914: 3910: 3909: 3906: 3905: 3901: 3900: 3897: 3896: 3892: 3891: 3888: 3887: 3883: 3882: 3879: 3878: 3874: 3873: 3870: 3869: 3865: 3864: 3861: 3860: 3856: 3855: 3852: 3851: 3835: 3832: 3831: 3828: 3827: 3810: 3805: 3803: 3800: 3799: 3796: 3795: 3779: 3774: 3772: 3769: 3768: 3765: 3764: 3760: 3759: 3756: 3755: 3744:Laiyangosaurus 3739: 3736: 3735: 3732: 3731: 3715: 3710: 3708: 3703: 3701: 3698: 3697: 3694: 3693: 3688: 3683: 3681: 3679:Edmontosaurini 3675: 3674: 3671: 3670: 3666: 3665: 3662: 3661: 3657: 3656: 3653: 3652: 3635: 3632: 3631: 3628: 3627: 3611: 3606: 3604: 3601: 3600: 3597: 3596: 3585:Prosaurolophus 3580: 3575: 3573: 3565: 3563: 3560: 3559: 3556: 3555: 3551: 3550: 3547: 3546: 3542: 3541: 3538: 3537: 3533: 3532: 3529: 3528: 3524: 3523: 3520: 3519: 3515: 3514: 3511: 3510: 3506: 3505: 3502: 3501: 3485: 3482: 3481: 3478: 3477: 3468: 3463: 3461: 3458: 3457: 3454: 3453: 3449: 3448: 3445: 3444: 3428: 3425: 3424: 3421: 3420: 3411: 3406: 3404: 3399: 3397: 3394: 3393: 3390: 3389: 3385: 3384: 3381: 3380: 3364: 3361: 3360: 3357: 3356: 3340: 3335: 3333: 3328: 3326: 3323: 3322: 3319: 3318: 3314: 3313: 3310: 3309: 3293: 3290: 3289: 3286: 3285: 3269: 3264: 3262: 3257: 3255: 3252: 3251: 3248: 3247: 3237: 3232: 3230: 3227: 3226: 3223: 3222: 3206: 3201: 3199: 3191: 3189: 3186: 3185: 3182: 3181: 3177: 3176: 3173: 3172: 3168: 3167: 3164: 3163: 3159: 3158: 3155: 3154: 3150: 3149: 3146: 3145: 3129: 3126: 3125: 3122: 3121: 3105: 3100: 3098: 3095: 3094: 3091: 3090: 3074: 3069: 3067: 3064: 3063: 3060: 3059: 3043: 3038: 3036: 3033: 3032: 3029: 3028: 3012: 3007: 3005: 2997: 2995: 2992: 2991: 2988: 2987: 2971: 2966: 2964: 2956: 2954: 2948: 2947: 2944: 2943: 2934: 2929: 2927: 2924: 2923: 2920: 2919: 2910: 2905: 2903: 2900: 2899: 2896: 2895: 2886: 2881: 2879: 2876: 2875: 2872: 2871: 2867:Nanyangosaurus 2862: 2857: 2855: 2852: 2851: 2848: 2847: 2838: 2833: 2831: 2830:Hadrosauridae 2825: 2818: 2817: 2814: 2813: 2809: 2808: 2805: 2804: 2800: 2799: 2796: 2795: 2791: 2790: 2787: 2786: 2782: 2781: 2778: 2777: 2773: 2772: 2769: 2768: 2764: 2763: 2760: 2759: 2755: 2754: 2751: 2750: 2746: 2745: 2742: 2741: 2737: 2736: 2733: 2732: 2728: 2727: 2724: 2723: 2719: 2718: 2715: 2714: 2710: 2709: 2706: 2705: 2701: 2700: 2697: 2696: 2692: 2691: 2688: 2687: 2668: 2665: 2664: 2661: 2660: 2641: 2636: 2634: 2631: 2630: 2627: 2626: 2607: 2602: 2600: 2597: 2596: 2593: 2592: 2573: 2568: 2566: 2563: 2562: 2559: 2558: 2554: 2553: 2550: 2549: 2545: 2544: 2541: 2540: 2521: 2518: 2517: 2514: 2513: 2494: 2489: 2487: 2484: 2483: 2480: 2479: 2468:Prosaurolophus 2460: 2455: 2453: 2448: 2446: 2443: 2442: 2439: 2438: 2429: 2424: 2422: 2419: 2418: 2415: 2414: 2410: 2409: 2406: 2405: 2401: 2400: 2397: 2396: 2392: 2391: 2388: 2387: 2383: 2382: 2379: 2378: 2374: 2373: 2370: 2369: 2350: 2345: 2343: 2340: 2339: 2336: 2335: 2323: 2318: 2316: 2313: 2312: 2309: 2308: 2289: 2284: 2282: 2279: 2278: 2275: 2274: 2255: 2250: 2248: 2245: 2244: 2241: 2240: 2221: 2216: 2214: 2209: 2207: 2204: 2203: 2200: 2199: 2195: 2194: 2191: 2190: 2171: 2168: 2167: 2164: 2163: 2144: 2141: 2140: 2137: 2136: 2117: 2112: 2110: 2105: 2103: 2097: 2096: 2093: 2092: 2088: 2087: 2084: 2083: 2079: 2078: 2075: 2074: 2070: 2069: 2066: 2065: 2061: 2060: 2057: 2056: 2052: 2051: 2048: 2047: 2043: 2042: 2039: 2038: 2034: 2033: 2030: 2029: 2025: 2024: 2021: 2020: 2016: 2015: 2012: 2011: 2007: 2006: 2003: 2002: 1983: 1980: 1979: 1976: 1975: 1956: 1951: 1949: 1946: 1945: 1942: 1941: 1922: 1919: 1918: 1915: 1914: 1895: 1890: 1888: 1885: 1884: 1881: 1880: 1876: 1875: 1872: 1871: 1867: 1866: 1863: 1862: 1850: 1847: 1846: 1843: 1842: 1823: 1818: 1816: 1813: 1812: 1809: 1808: 1789: 1784: 1782: 1777: 1775: 1772: 1771: 1768: 1767: 1754: 1749: 1747: 1744: 1743: 1740: 1739: 1720: 1715: 1713: 1710: 1709: 1706: 1705: 1686: 1681: 1679: 1676: 1675: 1672: 1671: 1667: 1666: 1663: 1662: 1643: 1640: 1639: 1636: 1635: 1616: 1611: 1609: 1604: 1602: 1599: 1598: 1595: 1594: 1590: 1589: 1586: 1585: 1566: 1563: 1562: 1559: 1558: 1539: 1534: 1532: 1527: 1525: 1522: 1521: 1518: 1517: 1498: 1495: 1494: 1491: 1490: 1471: 1466: 1464: 1461: 1460: 1457: 1456: 1437: 1432: 1430: 1427:Lambeosaurinae 1422: 1417: 1415: 1407: 1404: 1403: 1400: 1399: 1387: 1382: 1380: 1377: 1376: 1373: 1372: 1360: 1355: 1353: 1345: 1342: 1341: 1338: 1337: 1325: 1320: 1318: 1315: 1314: 1311: 1310: 1298: 1293: 1291: 1288: 1287: 1284: 1283: 1279: 1278: 1275: 1274: 1262: 1259: 1258: 1255: 1254: 1242: 1237: 1235: 1230: 1228: 1225: 1224: 1221: 1220: 1210: 1205: 1203: 1198: 1177:Lambeosaurinae 1139:Premaxilla of 1083: 1080: 1049:Tsintaosaurini 1025:magnicristatus 998:Lambeosaurinae 987: 986:Classification 984: 973:Tsintaosaurini 940: 937: 931:The Dinosauria 922:John R. Horner 894:John R. Horner 882:John H. Ostrom 823:(belonging to 662: 659: 644:Lambeosaurinae 632:ornithischians 528: 527: 526: 525: 517: 512:Cheneosauridae 509: 504:Lambeosauridae 501: 493: 484:Trachodontidae 478: 477: 471: 470: 469: 468: 467: 466: 458: 456:Lambeosaurinae 438: 428: 417: 406: 396: 385: 375: 365: 355: 344: 333: 323: 310: 309: 305: 304: 298: 286: 285: 279: 278: 263: 259: 258: 250: 243: 242: 234: 227: 226: 224:Neornithischia 218: 211: 210: 202: 195: 194: 189: 182: 181: 176: 172: 171: 166: 162: 161: 156: 152: 151: 138: 137: 124: 123: 115: 114: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 37: 26: 9: 6: 4: 3: 2: 9957: 9946: 9943: 9941: 9938: 9936: 9933: 9931: 9928: 9926: 9923: 9922: 9920: 9903: 9898: 9894: 9890: 9885: 9881: 9877: 9872: 9868: 9864: 9859: 9855: 9851: 9846: 9842: 9838: 9833: 9829: 9825: 9820: 9816: 9811: 9810:Hadrosauridae 9805: 9801: 9796: 9790: 9786: 9785: 9783: 9781: 9780:Hadrosauridae 9777: 9773: 9768: 9748: 9747: 9743: 9741: 9739: 9738:Edmontosaurus 9735: 9733: 9731: 9730:Edmontosaurus 9727: 9725: 9722: 9719: 9715: 9712: 9710: 9707: 9705: 9702: 9700: 9697: 9695: 9692: 9690: 9689: 9685: 9684: 9682: 9678: 9672: 9671: 9667: 9665: 9664: 9660: 9658: 9657: 9653: 9651: 9650: 9649:Syngonosaurus 9646: 9644: 9643: 9639: 9637: 9636: 9632: 9630: 9629: 9625: 9623: 9622: 9618: 9616: 9615: 9611: 9609: 9608: 9604: 9602: 9601: 9597: 9595: 9594: 9593:Koutalisaurus 9590: 9588: 9587: 9583: 9581: 9580: 9576: 9574: 9573: 9572:Eucercosaurus 9569: 9567: 9566: 9562: 9559: 9558: 9554: 9552: 9551: 9547: 9545: 9544: 9543:Arstanosaurus 9540: 9539: 9537: 9535: 9534: 9529: 9523: 9522: 9518: 9517: 9515: 9513: 9509: 9503: 9502: 9498: 9496: 9495: 9491: 9489: 9488: 9484: 9482: 9481: 9477: 9475: 9474: 9470: 9468: 9467: 9463: 9461: 9460: 9459:Orcauichnites 9456: 9454: 9453: 9449: 9447: 9446: 9442: 9440: 9439: 9435: 9433: 9432: 9428: 9426: 9425: 9421: 9419: 9418: 9414: 9412: 9411: 9407: 9405: 9404: 9400: 9398: 9397: 9396:Amblydactylus 9393: 9392: 9390: 9388: 9384: 9380: 9373: 9369: 9360: 9356: 9354: 9349: 9343: 9336: 9310: 9309: 9305: 9303: 9302: 9298: 9296: 9295: 9294:Kundurosaurus 9291: 9289: 9288: 9287:Kerberosaurus 9284: 9282: 9281: 9277: 9275: 9274: 9273:Edmontosaurus 9270: 9269: 9267: 9265: 9261: 9255: 9254: 9250: 9248: 9247: 9243: 9241: 9240: 9236: 9235: 9233: 9231: 9227: 9221: 9220: 9216: 9214: 9213: 9209: 9207: 9206: 9202: 9200: 9199: 9195: 9193: 9192: 9188: 9185: 9184: 9183:Anasazisaurus 9180: 9179: 9177: 9175: 9171: 9165: 9164: 9163:Secernosaurus 9160: 9158: 9157: 9156:Kelumapusaura 9153: 9151: 9150: 9146: 9144: 9143: 9139: 9138: 9136: 9134: 9130: 9123: 9122: 9118: 9116: 9115: 9111: 9109: 9108: 9104: 9102: 9101: 9097: 9095: 9094: 9090: 9088: 9087: 9083: 9082: 9080: 9078: 9074: 9068: 9067: 9063: 9062: 9059: 9056: 9054: 9053:Saurolophinae 9050: 9040: 9039: 9035: 9033: 9032: 9028: 9026: 9025: 9021: 9019: 9018: 9014: 9012: 9011: 9007: 9005: 9004: 9003:Hypacrosaurus 9000: 8998: 8997: 8996:Corythosaurus 8993: 8991: 8990: 8986: 8985: 8983: 8981: 8980:Lambeosaurini 8977: 8971: 8970: 8966: 8964: 8963: 8959: 8957: 8956: 8955:Charonosaurus 8952: 8950: 8949: 8945: 8944: 8942: 8940: 8936: 8930: 8929: 8928:Tsintaosaurus 8925: 8923: 8922: 8921:Pararhabdodon 8918: 8916: 8915: 8914:Nipponosaurus 8911: 8909: 8908: 8904: 8901: 8900: 8896: 8894: 8893: 8889: 8887: 8886: 8885:Jaxartosaurus 8882: 8880: 8879: 8875: 8873: 8872: 8868: 8866: 8865: 8861: 8859: 8858: 8854: 8852: 8851: 8847: 8845: 8844: 8840: 8838: 8837: 8836:Adynomosaurus 8833: 8832: 8829: 8826: 8824: 8820: 8814: 8813: 8809: 8807: 8806: 8802: 8800: 8799: 8798:Lapampasaurus 8795: 8793: 8792: 8788: 8786: 8785: 8781: 8779: 8778: 8774: 8773: 8770: 8767: 8764: 8763:Hadrosauridae 8759: 8753: 8752: 8748: 8746: 8745: 8744:Zhanghenglong 8741: 8739: 8738: 8734: 8732: 8731: 8727: 8725: 8724: 8720: 8718: 8717: 8716:Telmatosaurus 8713: 8711: 8710: 8706: 8704: 8703: 8699: 8697: 8696: 8692: 8690: 8689: 8685: 8683: 8682: 8678: 8676: 8675: 8671: 8669: 8668: 8664: 8662: 8661: 8660:Portellsaurus 8657: 8655: 8654: 8650: 8648: 8647: 8643: 8641: 8640: 8636: 8634: 8633: 8629: 8627: 8626: 8622: 8620: 8619: 8615: 8612: 8611: 8607: 8605: 8604: 8600: 8598: 8597: 8593: 8591: 8590: 8586: 8584: 8583: 8579: 8577: 8576: 8572: 8570: 8569: 8565: 8563: 8562: 8558: 8556: 8555: 8551: 8549: 8548: 8544: 8542: 8541: 8537: 8535: 8534: 8530: 8528: 8527: 8523: 8521: 8520: 8516: 8513: 8512: 8508: 8506: 8505: 8501: 8499: 8498: 8494: 8493: 8490: 8487: 8484: 8479: 8475: 8471: 8466: 8462: 8453: 8449: 8446: 8439: 8406: 8405: 8404: 8403: 8401: 8399: 8395: 8388: 8387: 8383: 8381: 8380: 8376: 8374: 8373: 8369: 8366: 8365: 8361: 8360: 8358: 8356: 8352: 8346: 8345: 8341: 8339: 8338: 8334: 8332: 8331: 8327: 8325: 8324: 8323:Jinzhousaurus 8320: 8318: 8317: 8316:Hypselospinus 8313: 8311: 8310: 8306: 8304: 8303: 8299: 8298: 8295: 8292: 8290: 8286: 8280: 8279: 8275: 8273: 8272: 8271:Theiophytalia 8268: 8266: 8265: 8261: 8259: 8258: 8254: 8252: 8251: 8247: 8245: 8244: 8240: 8238: 8237: 8233: 8231: 8230: 8226: 8224: 8223: 8219: 8217: 8216: 8212: 8210: 8209: 8205: 8203: 8202: 8201:Lanzhousaurus 8198: 8196: 8195: 8191: 8189: 8188: 8184: 8182: 8181: 8177: 8175: 8174: 8170: 8167: 8166: 8162: 8160: 8159: 8155: 8153: 8152: 8148: 8146: 8145: 8141: 8139: 8138: 8134: 8132: 8131: 8127: 8126: 8123: 8120: 8117: 8112: 8102: 8101: 8097: 8094: 8093: 8089: 8086: 8085: 8081: 8079: 8078: 8074: 8072: 8071: 8070:Elrhazosaurus 8067: 8065: 8064: 8060: 8058: 8057: 8053: 8052: 8050: 8048: 8044: 8038: 8037: 8033: 8031: 8030: 8026: 8024: 8023: 8019: 8018: 8015: 8011: 8008: 8005: 8000: 7986: 7985: 7981: 7979: 7978: 7974: 7972: 7971: 7967: 7965: 7964: 7960: 7958: 7957: 7953: 7951: 7950: 7946: 7945: 7943: 7941: 7937: 7931: 7930: 7929:Tenontosaurus 7926: 7924: 7923: 7919: 7916: 7915: 7914:Convolosaurus 7911: 7910: 7908: 7905: 7901: 7894: 7893: 7889: 7886: 7885: 7881: 7878: 7877: 7876:Ampelognathus 7873: 7872: 7869: 7866: 7864: 7860: 7853: 7852: 7848: 7846: 7845: 7841: 7839: 7838: 7834: 7832: 7831: 7827: 7825: 7824: 7820: 7818: 7817: 7813: 7811: 7810: 7806: 7803: 7802: 7798: 7796: 7795: 7791: 7789: 7788: 7787:Mahuidacursor 7784: 7782: 7781: 7777: 7775: 7774: 7770: 7767: 7766: 7762: 7760: 7759: 7755: 7753: 7752: 7748: 7746: 7745: 7741: 7738: 7737: 7733: 7731: 7730: 7726: 7724: 7723: 7719: 7717: 7716: 7712: 7710: 7709: 7705: 7704: 7702: 7700: 7696: 7689: 7688: 7684: 7682: 7681: 7680:Hypsilophodon 7677: 7674: 7673: 7669: 7668: 7666: 7663: 7659: 7652: 7651: 7647: 7645: 7644: 7643:Callovosaurus 7640: 7638: 7637: 7636:Burianosaurus 7633: 7632: 7629: 7625: 7622: 7619: 7614: 7610: 7606: 7601: 7597: 7572: 7571: 7570: 7569: 7567: 7565: 7561: 7553: 7549: 7548: 7547: 7546: 7543: 7540: 7538: 7534: 7526: 7522: 7521: 7520: 7519: 7516: 7513: 7511: 7507: 7503: 7495: 7491: 7488: 7486: 7482: 7479: 7477: 7473: 7470: 7468: 7464: 7461: 7459: 7455: 7453: 7449: 7448: 7443: 7439: 7435: 7428: 7423: 7421: 7416: 7414: 7409: 7408: 7405: 7399: 7396: 7394: 7391: 7389: 7386: 7384: 7381: 7380: 7376: 7365: 7352: 7346: 7342: 7335: 7321:on 2007-11-17 7320: 7316: 7312: 7308: 7304: 7298: 7290: 7286: 7282: 7278: 7274: 7270: 7266: 7262: 7258: 7254: 7247: 7240: 7232: 7228: 7224: 7220: 7216: 7212: 7208: 7204: 7200: 7196: 7189: 7179: 7177: 7175: 7173: 7171: 7169: 7167: 7165: 7163: 7154: 7150: 7146: 7142: 7135: 7127: 7123: 7119: 7115: 7111: 7107: 7103: 7099: 7095: 7088: 7080: 7076: 7071: 7066: 7061: 7056: 7052: 7048: 7044: 7040: 7036: 7029: 7027: 7025: 7016: 7012: 7008: 7004: 6999: 6994: 6990: 6986: 6982: 6975: 6973: 6971: 6969: 6960: 6956: 6952: 6948: 6941: 6939: 6929: 6924: 6920: 6916: 6912: 6905: 6897: 6893: 6888: 6883: 6878: 6873: 6869: 6865: 6861: 6857: 6853: 6846: 6838: 6834: 6830: 6826: 6822: 6818: 6814: 6810: 6803: 6795: 6791: 6786: 6781: 6776: 6771: 6767: 6763: 6759: 6752: 6742: 6727: 6723: 6719: 6712: 6710: 6701: 6697: 6693: 6689: 6685: 6681: 6677: 6673: 6669: 6662: 6648:on 2009-07-02 6647: 6643: 6642: 6637: 6633: 6627: 6625: 6610: 6609: 6604: 6597: 6589: 6585: 6580: 6575: 6570: 6565: 6561: 6557: 6553: 6549: 6545: 6538: 6523: 6517: 6509: 6505: 6500: 6495: 6490: 6485: 6481: 6477: 6474:(2): e31295. 6473: 6469: 6465: 6463: 6454: 6446: 6442: 6438: 6434: 6430: 6426: 6422: 6418: 6414: 6407: 6399: 6395: 6391: 6387: 6383: 6379: 6375: 6371: 6367: 6360: 6352: 6348: 6343: 6338: 6333: 6328: 6324: 6320: 6317:(7): e69835. 6316: 6312: 6308: 6306: 6297: 6289: 6287:0-89950-917-7 6283: 6279: 6275: 6269: 6260: 6255: 6251: 6247: 6243: 6236: 6234: 6232: 6230: 6221: 6217: 6213: 6206: 6198: 6194: 6189: 6184: 6180: 6176: 6172: 6168: 6164: 6157: 6149: 6145: 6140: 6135: 6130: 6125: 6121: 6117: 6113: 6109: 6105: 6097: 6089: 6085: 6080: 6075: 6071: 6067: 6063: 6059: 6055: 6051: 6047: 6043: 6039: 6031: 6023: 6017: 6013: 6006: 5998: 5992: 5988: 5981: 5975: 5969: 5963: 5957: 5949: 5945: 5941: 5937: 5933: 5929: 5925: 5921: 5914: 5906: 5899: 5891: 5887: 5883: 5879: 5875: 5871: 5867: 5863: 5856: 5848: 5841: 5832: 5827: 5823: 5819: 5815: 5811: 5807: 5803: 5797: 5795: 5793: 5786: 5780: 5778: 5776: 5774: 5772: 5763: 5759: 5755: 5751: 5747: 5743: 5739: 5735: 5731: 5724: 5716: 5712: 5708: 5704: 5697: 5695: 5693: 5691: 5682: 5678: 5674: 5670: 5665: 5660: 5656: 5652: 5648: 5644: 5636: 5632: 5625: 5623: 5619: 5618:Edmontosaurus 5615: 5611: 5607: 5598: 5596: 5592: 5591: 5586: 5585: 5580: 5579: 5578:Corythosaurus 5574: 5573:scleral rings 5564: 5560: 5554: 5553: 5547: 5538: 5535: 5531: 5527: 5521: 5518: 5514: 5511: 5506: 5502: 5498: 5493: 5491: 5490: 5485: 5484: 5478: 5474: 5470: 5466: 5462: 5457: 5455: 5451: 5447: 5443: 5437: 5432: 5423: 5421: 5417: 5413: 5412:Tyrannosaurus 5409: 5408:Hypacrosaurus 5405: 5401: 5397: 5396: 5391: 5390: 5385: 5381: 5380: 5375: 5373: 5372: 5371:Archaeopteryx 5367: 5366: 5361: 5357: 5353: 5349: 5345: 5341: 5340:Hypacrosaurus 5333: 5328: 5324: 5321: 5317: 5313: 5312: 5307: 5303: 5302: 5301:Corythosaurus 5297: 5293: 5292: 5291:Hypacrosaurus 5287: 5286:lambeosaurine 5282: 5277: 5275: 5274:intraspecific 5271: 5267: 5263: 5259: 5258: 5253: 5252: 5247: 5246:Edmontosaurus 5243: 5239: 5235: 5234: 5229: 5225: 5221: 5220: 5215: 5211: 5210: 5205: 5201: 5197: 5196:ornithischian 5193: 5186: 5182: 5177: 5168: 5166: 5162: 5158: 5157: 5151: 5146: 5144: 5143:invertebrates 5140: 5136: 5132: 5128: 5126: 5122: 5114: 5113:Edmontosaurus 5110: 5106: 5104: 5103: 5098: 5092: 5089: 5085: 5077: 5072: 5067: 5051: 5040: 5037: 5033: 5030: 5026: 5023: 5019: 5016: 5012: 5009: – 5008: 5004: 5003:Find sources: 4997: 4993: 4989: 4982: 4978: 4973: 4972: 4964: 4962: 4961:Hadrosaurinae 4958: 4957:Saurolophinae 4954: 4949: 4947: 4942: 4939: 4937: 4932: 4928: 4927:Edmontosaurus 4920: 4916: 4915: 4914:Edmontosaurus 4910: 4896: 4895: 4887: 4886: 4878: 4877: 4869: 4868: 4860: 4859: 4851: 4850: 4842: 4841: 4833: 4832: 4824: 4823: 4815: 4814: 4806: 4805: 4797: 4796: 4788: 4787: 4779: 4778: 4770: 4769: 4761: 4760: 4752: 4751: 4743: 4742: 4734: 4733: 4730: 4727: 4722: 4721: 4714: 4713: 4710: 4709: 4706: 4703: 4698: 4697: 4690: 4689: 4683: 4682: 4679: 4678: 4675: 4672: 4667: 4666: 4659: 4658: 4655: 4654: 4651: 4648: 4643: 4642: 4635: 4634: 4631: 4630: 4627: 4624: 4619: 4618: 4611: 4610: 4604: 4603: 4600: 4599: 4591: 4590: 4582: 4581: 4578: 4575: 4569: 4568: 4561: 4560: 4557: 4556: 4553: 4550: 4544: 4543: 4542:Corythosaurus 4536: 4535: 4529: 4528: 4525: 4524: 4521: 4520: 4519: 4512: 4511: 4505: 4504: 4498: 4497: 4494: 4493: 4490: 4487: 4482: 4481: 4474: 4473: 4467: 4466: 4463: 4462: 4459: 4456: 4451: 4450: 4443: 4442: 4436: 4435: 4432: 4431: 4428: 4425: 4420: 4419: 4412: 4411: 4405: 4404: 4401: 4400: 4392: 4391: 4388: 4387: 4386: 4379: 4378: 4375: 4374: 4371: 4368: 4363: 4362: 4355: 4354: 4348: 4347: 4341: 4340:Lambeosaurini 4338: 4337: 4334: 4333: 4325: 4324: 4316: 4315: 4307: 4306: 4303: 4300: 4295: 4294: 4287: 4286: 4283: 4282: 4279: 4276: 4271: 4270: 4263: 4262: 4256: 4255: 4252: 4251: 4248: 4245: 4239: 4238: 4231: 4230: 4224: 4223: 4220: 4219: 4216: 4213: 4208: 4207: 4206:Charonosaurus 4200: 4199: 4193: 4190: 4189: 4183: 4182:Corythosauria 4180: 4179: 4176: 4175: 4172: 4169: 4164: 4163: 4156: 4155: 4149: 4148: 4145: 4144: 4141: 4138: 4133: 4132: 4131:Jaxartosaurus 4125: 4124: 4118: 4117: 4114: 4113: 4105: 4104: 4096: 4095: 4087: 4086: 4083: 4080: 4075: 4074: 4073:Pararhabdodon 4067: 4066: 4063: 4062: 4059: 4056: 4051: 4050: 4043: 4042: 4036: 4035: 4032: 4031: 4028: 4025: 4020: 4019: 4018:Tsintaosaurus 4012: 4011: 4005: 4004: 4001: 4000: 3997: 3994: 3989: 3988: 3987:Adynomosaurus 3981: 3980: 3974: 3971: 3970: 3964: 3963: 3960: 3959: 3956: 3953: 3948: 3947: 3940: 3939: 3933: 3930: 3929: 3926: 3925: 3917: 3916: 3908: 3907: 3899: 3898: 3890: 3889: 3881: 3880: 3872: 3871: 3863: 3862: 3854: 3853: 3850: 3847: 3842: 3841: 3834: 3833: 3830: 3829: 3826: 3823: 3817: 3816: 3815:Edmontosaurus 3809: 3808: 3802: 3801: 3798: 3797: 3794: 3791: 3786: 3785: 3784:Kerberosaurus 3778: 3777: 3771: 3770: 3767: 3766: 3758: 3757: 3754: 3751: 3746: 3745: 3738: 3737: 3734: 3733: 3730: 3727: 3722: 3721: 3714: 3713: 3707: 3706: 3700: 3699: 3696: 3695: 3692: 3687: 3686: 3680: 3677: 3676: 3673: 3672: 3664: 3663: 3655: 3654: 3651: 3648: 3642: 3641: 3634: 3633: 3630: 3629: 3626: 3623: 3618: 3617: 3610: 3609: 3603: 3602: 3599: 3598: 3595: 3592: 3587: 3586: 3579: 3578: 3572: 3569: 3568: 3562: 3561: 3558: 3557: 3549: 3548: 3540: 3539: 3531: 3530: 3522: 3521: 3513: 3512: 3504: 3503: 3500: 3497: 3492: 3491: 3484: 3483: 3480: 3479: 3476: 3475: 3474: 3467: 3466: 3460: 3459: 3456: 3455: 3447: 3446: 3443: 3440: 3435: 3434: 3427: 3426: 3423: 3422: 3419: 3418: 3417: 3410: 3409: 3403: 3402: 3396: 3395: 3392: 3391: 3383: 3382: 3379: 3376: 3371: 3370: 3369:Secernosaurus 3363: 3362: 3359: 3358: 3355: 3352: 3347: 3346: 3339: 3338: 3332: 3331: 3325: 3324: 3321: 3320: 3312: 3311: 3308: 3305: 3300: 3299: 3292: 3291: 3288: 3287: 3284: 3281: 3276: 3275: 3268: 3267: 3261: 3260: 3254: 3253: 3250: 3249: 3246: 3244: 3243: 3236: 3235: 3229: 3228: 3225: 3224: 3221: 3218: 3213: 3212: 3205: 3204: 3198: 3195: 3194: 3188: 3187: 3184: 3183: 3175: 3174: 3166: 3165: 3157: 3156: 3148: 3147: 3144: 3141: 3136: 3135: 3128: 3127: 3124: 3123: 3120: 3117: 3112: 3111: 3104: 3103: 3097: 3096: 3093: 3092: 3089: 3086: 3081: 3080: 3073: 3072: 3066: 3065: 3062: 3061: 3058: 3055: 3050: 3049: 3042: 3041: 3035: 3034: 3031: 3030: 3027: 3024: 3019: 3018: 3011: 3010: 3004: 3001: 3000: 2994: 2993: 2990: 2989: 2986: 2983: 2978: 2977: 2970: 2969: 2963: 2962:Saurolophinae 2960: 2959: 2953: 2952:Euhadrosauria 2950: 2949: 2946: 2945: 2942: 2941: 2940: 2933: 2932: 2926: 2925: 2922: 2921: 2918: 2917: 2916: 2909: 2908: 2902: 2901: 2898: 2897: 2894: 2893: 2892: 2885: 2884: 2878: 2877: 2874: 2873: 2870: 2869: 2868: 2861: 2860: 2854: 2853: 2850: 2849: 2846: 2845: 2844: 2837: 2836: 2829: 2828: 2824: 2816: 2815: 2807: 2806: 2798: 2797: 2789: 2788: 2780: 2779: 2771: 2770: 2762: 2761: 2753: 2752: 2744: 2743: 2735: 2734: 2726: 2725: 2717: 2716: 2708: 2707: 2699: 2698: 2690: 2689: 2686: 2683: 2678: 2677: 2676:Edmontosaurus 2667: 2666: 2663: 2662: 2659: 2656: 2651: 2650: 2640: 2639: 2633: 2632: 2629: 2628: 2625: 2622: 2617: 2616: 2615:Kundurosaurus 2606: 2605: 2599: 2598: 2595: 2594: 2591: 2588: 2583: 2582: 2581:Kerberosaurus 2572: 2571: 2565: 2564: 2561: 2560: 2552: 2551: 2543: 2542: 2539: 2536: 2531: 2530: 2520: 2519: 2516: 2515: 2512: 2509: 2504: 2503: 2493: 2492: 2486: 2485: 2482: 2481: 2478: 2475: 2470: 2469: 2459: 2458: 2452: 2451: 2445: 2444: 2441: 2440: 2437: 2428: 2427: 2421: 2420: 2417: 2416: 2408: 2407: 2399: 2398: 2390: 2389: 2381: 2380: 2372: 2371: 2368: 2365: 2360: 2359: 2358:Secernosaurus 2349: 2348: 2342: 2341: 2338: 2337: 2334: 2333: 2332: 2322: 2321: 2315: 2314: 2311: 2310: 2307: 2304: 2299: 2298: 2288: 2287: 2281: 2280: 2277: 2276: 2273: 2270: 2265: 2264: 2254: 2253: 2247: 2246: 2243: 2242: 2239: 2236: 2231: 2230: 2220: 2219: 2213: 2212: 2206: 2205: 2202: 2201: 2193: 2192: 2189: 2186: 2181: 2180: 2170: 2169: 2166: 2165: 2162: 2159: 2154: 2153: 2143: 2142: 2139: 2138: 2135: 2132: 2127: 2126: 2116: 2115: 2109: 2108: 2102: 2101:Saurolophinae 2099: 2098: 2095: 2094: 2086: 2085: 2077: 2076: 2068: 2067: 2059: 2058: 2050: 2049: 2041: 2040: 2032: 2031: 2023: 2022: 2014: 2013: 2005: 2004: 2001: 1998: 1993: 1992: 1982: 1981: 1978: 1977: 1974: 1971: 1966: 1965: 1955: 1954: 1948: 1947: 1944: 1943: 1940: 1937: 1932: 1931: 1921: 1920: 1917: 1916: 1913: 1910: 1905: 1904: 1903:Hypacrosaurus 1894: 1893: 1887: 1886: 1883: 1882: 1874: 1873: 1865: 1864: 1861: 1860: 1859: 1849: 1848: 1845: 1844: 1841: 1838: 1833: 1832: 1822: 1821: 1815: 1814: 1811: 1810: 1807: 1804: 1799: 1798: 1788: 1787: 1781: 1780: 1774: 1773: 1770: 1769: 1766: 1765: 1762: 1761:Hypacrosaurus 1753: 1752: 1746: 1745: 1742: 1741: 1738: 1735: 1730: 1729: 1728:Corythosaurus 1719: 1718: 1712: 1711: 1708: 1707: 1704: 1701: 1696: 1695: 1685: 1684: 1678: 1677: 1674: 1673: 1665: 1664: 1661: 1658: 1653: 1652: 1642: 1641: 1638: 1637: 1634: 1631: 1626: 1625: 1624:Charonosaurus 1615: 1614: 1608: 1607: 1601: 1600: 1597: 1596: 1588: 1587: 1584: 1581: 1576: 1575: 1574:Pararhabdodon 1565: 1564: 1561: 1560: 1557: 1554: 1549: 1548: 1547:Tsintaosaurus 1538: 1537: 1531: 1530: 1524: 1523: 1520: 1519: 1516: 1513: 1508: 1507: 1506:Jaxartosaurus 1497: 1496: 1493: 1492: 1489: 1486: 1481: 1480: 1470: 1469: 1463: 1462: 1459: 1458: 1455: 1452: 1447: 1446: 1436: 1435: 1428: 1421: 1420: 1413: 1412:Saurolophidae 1406: 1405: 1402: 1401: 1398: 1397: 1396: 1386: 1385: 1379: 1378: 1375: 1374: 1371: 1370: 1369: 1359: 1358: 1351: 1350:Hadrosauridae 1344: 1343: 1340: 1339: 1336: 1335: 1334: 1324: 1323: 1317: 1316: 1313: 1312: 1309: 1308: 1307: 1297: 1296: 1290: 1289: 1286: 1285: 1277: 1276: 1273: 1272: 1271: 1261: 1260: 1257: 1256: 1253: 1252: 1251: 1241: 1240: 1234: 1233: 1227: 1226: 1223: 1222: 1219: 1218: 1217: 1216:Telmatosaurus 1209: 1208: 1202: 1201: 1195: 1194:Saurolophinae 1191: 1190: 1184: 1178: 1174: 1173: 1167: 1163: 1161: 1156: 1148: 1144: 1143: 1137: 1133: 1131: 1130:Telmatosaurus 1127: 1126: 1121: 1120: 1119:Telmatosaurus 1115: 1111: 1110: 1105: 1104: 1099: 1098: 1093: 1089: 1079: 1077: 1073: 1069: 1067: 1061: 1060:Hadrosaurinae 1056: 1054: 1050: 1046: 1042: 1038: 1034: 1030: 1026: 1024: 1019: 1018:Lambeosaurini 1015: 1010: 1007: 1003: 1002:Saurolophinae 999: 995: 994: 983: 981: 979: 974: 970: 966: 962: 958: 957:Maastrichtian 954: 945: 936: 933: 932: 927: 923: 919: 918: 912: 908: 903: 901: 900: 895: 891: 887: 883: 876: 872: 871: 865: 861: 859: 854: 850: 846: 841: 838: 834: 830: 826: 822: 818: 814: 809: 805: 801: 797: 793: 792: 787: 786: 776: 772: 770: 769: 763: 760: 756: 752: 748: 744: 740: 735: 727: 720: 713: 709: 708: 703: 699: 698: 693: 689: 688:North America 685: 681: 674: 673: 667: 658: 656: 651: 649: 648:Saurolophinae 645: 641: 637: 633: 628: 625: 624:iguanodontian 622: 618: 617:Late Jurassic 614: 611: 607: 603: 602: 597: 596: 595:Edmontosaurus 591: 587: 586:Hadrosauridae 583: 582:ornithischian 579: 576: 572: 569: 562: 558: 555: 552: 545: 541: 538: 537:Ancient Greek 534: 521: 518: 513: 510: 507:(Parks, 1923) 505: 502: 499:(Brown, 1914) 497: 496:Saurolophidae 494: 490: 485: 482: 481: 479: 476: 472: 465: 464:Saurolophinae 459: 457: 451: 450: 449: 448:Saurolophidae 445: 444:Euhadrosauria 439: 437: 436: 429: 427: 426: 418: 416: 415: 407: 405: 404: 397: 395: 394: 386: 384: 383: 376: 374: 373: 372:Lapampasaurus 366: 364: 363: 356: 354: 353: 345: 343: 342: 334: 332: 331: 324: 322: 321: 314: 313: 311: 306: 301: 296: 295: 287: 284: 280: 275: 270: 269:Hadrosauridae 264: 261: 260: 257: 251: 248: 245: 244: 241: 235: 232: 229: 228: 225: 219: 216: 213: 212: 209: 203: 200: 197: 196: 193: 190: 187: 184: 183: 180: 177: 174: 173: 170: 167: 164: 163: 160: 157: 154: 153: 148: 143: 139: 136: 132: 131: 125: 121: 116: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 48: 41: 33: 30: 19: 18:Euhadrosauria 9779: 9744: 9737: 9729: 9717: 9686: 9668: 9661: 9654: 9647: 9640: 9633: 9626: 9619: 9612: 9605: 9598: 9591: 9584: 9577: 9570: 9563: 9557:Claorhynchus 9555: 9548: 9541: 9533:Nomina dubia 9531: 9521:Guegoolithus 9519: 9499: 9492: 9487:Staurichnium 9485: 9480:Sousaichnium 9478: 9471: 9464: 9457: 9450: 9443: 9436: 9431:Dakotasaurus 9429: 9424:Caririchnium 9422: 9415: 9408: 9401: 9394: 9338: 9306: 9299: 9292: 9285: 9278: 9271: 9251: 9244: 9237: 9230:Saurolophini 9217: 9210: 9203: 9196: 9189: 9181: 9174:Kritosaurini 9161: 9154: 9149:Huallasaurus 9147: 9140: 9121:Wulagasaurus 9119: 9112: 9105: 9098: 9091: 9084: 9064: 9036: 9029: 9022: 9015: 9010:Lambeosaurus 9008: 9001: 8994: 8987: 8967: 8960: 8953: 8946: 8926: 8919: 8912: 8905: 8897: 8890: 8883: 8876: 8869: 8862: 8855: 8848: 8841: 8834: 8812:Yamatosaurus 8810: 8803: 8796: 8789: 8782: 8777:Aquilarhinus 8775: 8762: 8749: 8742: 8735: 8728: 8723:Tethyshadros 8721: 8714: 8707: 8700: 8693: 8686: 8679: 8672: 8665: 8658: 8653:Plesiohadros 8651: 8644: 8637: 8630: 8625:Lophorhothon 8623: 8616: 8608: 8601: 8594: 8587: 8580: 8573: 8566: 8559: 8552: 8545: 8538: 8531: 8524: 8517: 8511:Batyrosaurus 8509: 8504:Bactrosaurus 8502: 8495: 8441: 8384: 8377: 8370: 8362: 8344:Ouranosaurus 8342: 8335: 8328: 8321: 8314: 8309:Brighstoneus 8307: 8300: 8276: 8269: 8262: 8255: 8248: 8241: 8236:Osmakasaurus 8234: 8229:Oblitosaurus 8227: 8220: 8213: 8206: 8199: 8192: 8185: 8178: 8171: 8163: 8156: 8151:Camptosaurus 8149: 8142: 8135: 8128: 8098: 8092:Kangnasaurus 8090: 8082: 8075: 8068: 8061: 8054: 8047:Dryosauridae 8034: 8027: 8020: 7982: 7975: 7968: 7961: 7954: 7949:Matheronodon 7947: 7927: 7920: 7912: 7890: 7882: 7874: 7849: 7842: 7835: 7828: 7823:Sektensaurus 7821: 7816:Qantassaurus 7814: 7807: 7799: 7792: 7785: 7778: 7771: 7765:Kangnasaurus 7763: 7756: 7749: 7742: 7734: 7729:Diluvicursor 7727: 7720: 7713: 7706: 7687:Vectidromeus 7685: 7678: 7670: 7648: 7641: 7634: 7552:Ornithischia 7537:Ornithischia 7489: 7480: 7476:Ornithischia 7471: 7462: 7340: 7334: 7323:. Retrieved 7319:the original 7314: 7310: 7297: 7256: 7252: 7239: 7198: 7194: 7188: 7144: 7140: 7134: 7101: 7097: 7087: 7042: 7038: 6988: 6984: 6950: 6946: 6918: 6914: 6904: 6862:(1): 11163. 6859: 6855: 6845: 6812: 6808: 6802: 6765: 6761: 6751: 6741: 6730:. Retrieved 6726:the original 6721: 6675: 6671: 6667: 6661: 6650:. Retrieved 6646:the original 6639: 6612:. Retrieved 6606: 6596: 6551: 6547: 6537: 6526:. Retrieved 6516: 6471: 6467: 6461: 6453: 6420: 6416: 6406: 6373: 6369: 6359: 6314: 6310: 6304: 6296: 6277: 6268: 6249: 6245: 6219: 6215: 6205: 6170: 6166: 6156: 6139:10362/145947 6111: 6107: 6096: 6045: 6041: 6030: 6011: 6005: 5986: 5980: 5968: 5956: 5923: 5919: 5913: 5904: 5898: 5865: 5861: 5855: 5846: 5840: 5813: 5809: 5784: 5737: 5733: 5723: 5706: 5702: 5664:11336/105444 5646: 5642: 5635: 5617: 5613: 5609: 5604: 5588: 5582: 5576: 5570: 5561: 5557: 5550: 5522: 5501:Two Medicine 5494: 5487: 5481: 5473:Darren Tanke 5458: 5440: 5435: 5426:Reproduction 5419: 5415: 5411: 5407: 5403: 5399: 5393: 5389:Ceratosaurus 5387: 5377: 5376: 5369: 5365:Conchoraptor 5363: 5356:saurolophine 5344:maniraptoran 5339: 5337: 5331: 5316:phylogenetic 5311:Lambeosaurus 5309: 5308:34825), and 5299: 5289: 5278: 5266:crocodilians 5262:carnosaurian 5255: 5251:Camptosaurus 5249: 5245: 5238:James Hopson 5231: 5227: 5224:E. annectens 5223: 5217: 5207: 5190: 5184: 5180: 5161:saurolophine 5154: 5147: 5129: 5120: 5118: 5100: 5093: 5088:Mark Purnell 5081: 5045: 5035: 5028: 5021: 5014: 5002: 4967:Paleobiology 4950: 4943: 4940: 4926: 4923: 4912: 4718: 4717: 4694: 4693: 4663: 4662: 4639: 4638: 4615: 4614: 4567:Lambeosaurus 4565: 4564: 4540: 4539: 4516: 4515: 4478: 4477: 4447: 4446: 4416: 4415: 4383: 4382: 4359: 4358: 4291: 4290: 4267: 4266: 4235: 4234: 4204: 4203: 4160: 4159: 4129: 4128: 4071: 4070: 4047: 4046: 4016: 4015: 3985: 3984: 3944: 3943: 3838: 3837: 3813: 3812: 3782: 3781: 3742: 3741: 3718: 3717: 3690: 3638: 3637: 3614: 3613: 3583: 3582: 3571:Saurolophini 3488: 3487: 3471: 3470: 3431: 3430: 3414: 3413: 3367: 3366: 3343: 3342: 3296: 3295: 3272: 3271: 3240: 3239: 3209: 3208: 3197:Kritosaurini 3132: 3131: 3108: 3107: 3077: 3076: 3046: 3045: 3015: 3014: 2976:Wulagasaurus 2974: 2973: 2939:Yamatosaurus 2937: 2936: 2915:Aquilarhinus 2913: 2912: 2889: 2888: 2865: 2864: 2841: 2840: 2822: 2671: 2670: 2644: 2643: 2610: 2609: 2576: 2575: 2524: 2523: 2497: 2496: 2463: 2462: 2431: 2353: 2352: 2331:Aquilarhinus 2326: 2325: 2292: 2291: 2258: 2257: 2224: 2223: 2174: 2173: 2147: 2146: 2120: 2119: 1986: 1985: 1959: 1958: 1925: 1924: 1898: 1897: 1853: 1852: 1826: 1825: 1792: 1791: 1763: 1756: 1723: 1722: 1694:Lambeosaurus 1689: 1688: 1646: 1645: 1619: 1618: 1569: 1568: 1542: 1541: 1501: 1500: 1474: 1473: 1440: 1439: 1390: 1389: 1363: 1362: 1346: 1333:Tethyshadros 1328: 1327: 1301: 1300: 1270:Lophorhothon 1265: 1264: 1245: 1244: 1213: 1212: 1187: 1172:Lambeosaurus 1170: 1154: 1152: 1146: 1140: 1129: 1125:Bactrosaurus 1123: 1117: 1113: 1107: 1101: 1095: 1085: 1075: 1064: 1059: 1057: 1053:Aralosaurini 1044: 1040: 1036: 1032: 1023:Lambeosaurus 1021: 1011: 1006:Phylogenetic 991: 989: 976: 969:Arenysaurini 965:Kritosaurini 950: 939:Distribution 929: 915: 904: 897: 879: 868: 867:Skeleton of 857: 842: 837:Henry Osborn 824: 812: 803: 799: 789: 783: 781: 766: 755:Philadelphia 750: 738: 705: 695: 692:Joseph Leidy 684:Judith River 678: 670: 652: 629: 599: 593: 585: 574: 567: 564: 557: 550: 547: 540: 533:Hadrosaurids 532: 531: 523:(Cope, 1871) 519: 511: 503: 495: 483: 447: 443: 435:Yamatosaurus 433: 425:Plesiohadros 423: 412: 401: 393:Lophorhothon 391: 380: 370: 360: 350: 339: 328: 320:Aquilarhinus 318: 292: 283:Type species 268: 246: 230: 214: 208:Ornithischia 198: 185: 128: 35:Hadrosaurids 29: 9845:iNaturalist 9804:Wikispecies 9746:Willinakaqe 9718:nomina nuda 9635:Siluosaurus 9600:Loncosaurus 9438:Dinehichnus 9280:Kamuysaurus 9253:Saurolophus 9205:Kritosaurus 9198:Gryposaurus 9031:Sahaliyania 9017:Magnapaulia 8989:Amurosaurus 8969:Tlatolophus 8948:Adelolophus 8892:Kazaklambia 8871:Blasisaurus 8864:Arenysaurus 8857:Aralosaurus 8791:Hadrosaurus 8784:Eotrachodon 8737:Yunganglong 8702:Sirindhorna 8674:Protohadros 8610:Koshisaurus 8603:Jintasaurus 8372:Comptonatus 8222:Napaisaurus 8208:Lurdusaurus 8180:Fukuisaurus 8158:Cedrorestes 8100:Valdosaurus 7963:Pareisactus 7794:Morrosaurus 7758:Isasicursor 7722:Chakisaurus 7618:Ornithopoda 7605:Ornithopoda 7564:Ornithopoda 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9663:Tichosteus 9656:Thespesius 9628:Pteropelyx 9621:Orthomerus 9501:Wintonopus 9086:Acristavus 9066:Barsboldia 9024:Olorotitan 8899:Latirhinus 8751:Zuoyunlong 8618:Levnesovia 8568:Gobihadros 8533:Datonglong 8526:Claosaurus 8497:Altirhinus 8407:see below↓ 8337:Morelladon 8215:Magnamanus 8187:Hippodraco 8056:Dryosaurus 8029:Hesperonyx 8004:Dryomorpha 7844:Trinisaura 7837:Tietasaura 7830:Talenkauen 7708:Anabisetia 7573:see below↓ 7467:Dinosauria 7325:2009-03-08 6815:(5): 554. 6732:2009-06-03 6652:2009-06-03 6614:2009-06-03 6528:2013-07-23 6423:: 105267. 6173:: 104678. 6114:(4): 162. 6012:Hadrosaurs 5987:Hadrosaurs 5628:References 5595:cathemeral 5395:Allosaurus 5192:Hadrosaurs 5135:Cretaceous 5131:Coprolites 5048:March 2014 5018:newspapers 4665:Olorotitan 4449:Latirhinus 3017:Acristavus 2125:Acristavus 1930:Olorotitan 1764:stebingeri 1306:Claosaurus 1076:H. foulkii 1041:P. walkeri 917:Hadrosaurs 890:lambeosaur 636:predentary 606:herbivores 590:ornithopod 535:(from 382:Latirhinus 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