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Elpistostegalia

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Being shallow-water fishes, the elpistostegalians evolved many of the basic adaptions that later allowed the tetrapods to become terrestrial animals. The most important ones were the shift of main propulsion apparatus from the tail fin to the pectoral and pelvic fins, and a shift to reliance on lungs
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A rise in global oxygen content allowed for the evolution of large, predatory fish that were able to exploit the shallow tidal areas and swamplands as top predators. Several groups evolved to fill these niches, the most successful were the elpistiostegalians. In such environments, they would have
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Lebedev, O. A.; Clément, G. (2018). "New tetrapodomorph vertebrates from the Yam-Tesovo locality (Amata Regional Stage, Middle–Upper Devonian) of Leningrad Region, northwestern Russia".
453:(around 380–375 million years old). Thus, the oldest bone remains of elpistostegalians appear in the late Middle Devonian or in the early Late Devonian. However, 1603: 2623: 3215: 283: 2133: 436:
rather than gills as the main means of obtaining oxygen. Both of these appear to be a direct result of moving to an inland freshwater mode of living.
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Chang, M.-M.; Yu, X. (1997). "Reexamination of the relationship of Middle Devonian osteolepids–fossil characters and their interpretations".
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from the early Middle Devonian of Poland suggests that tetrapod-limbed animals may have existed as early as in the
1831:"Devonian rise in atmospheric oxygen correlated to the radiations of terrestrial plants and large predatory fish" 4503: 4510: 4637: 2279: 2174: 146: 1547:
Niedźwiedzki, Grzegorz; Szrek, Piotr; Narkiewicz, Katarzyna; Narkiewicz, Marek; Ahlberg, Per E. (2010).
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been challenged by periodic oxygen deficiency. In comparable modern aquatic environments like shallow
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Stewart, Thomas A.; Lemberg, Justin B.; Daly, Ailis; Daeschler, Edward B.; Shubin, Neil H. (2022).
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Cloutier, R.; Clement, A. M.; Lee, M. S. Y.; Noël, R.; Béchard, I.; Roy, V.; Long, J. A. (2020). "
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The anal and posterior dorsal fin supported by a basal plate and three unjointed radials.
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Earth and Environmental Science Transactions of the Royal Society of Edinburgh
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Boisvert, Catherine A.; Mark-Kurik, Elga; Ahlberg, Per E. (December 2008).
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The pectoral fin skeleton is composed of bones homologous to the tetrapod
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Long, J.A. (1990). "Heterochrony and the origin of tetrapods".
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Dahl TW, Hammarlund EU, Anbar AD, et al. (October 2010).
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as the sister taxon of all unequivocally digited vertebrates.
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taxa. The earliest elpistostegalians, combining fishlike and
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also rely on the more stable, atmospheric source of oxygen.
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Ahlberg, P. E.; Luksevics, E.; Mark-Kurik, E. (2000).
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foramen. The cranial kinesis is also visible in the
347: 309: 299: 289: 279: 269: 259: 248: 2294: 1828: 1172:. The analysis carried out in this study recovered 2197: 2013: 2011: 1627:"A near-tetrapod from the Baltic Middle Devonian" 4785: 2008: 1472: 1470: 2444: 2134:"Osteolepiforms and the ancestry of tetrapods" 1957: 1660: 2280: 2067: 2368: 1608:Proceedings of the Royal Society of Victoria 1467: 1601: 1542: 1540: 2287: 2273: 2127: 2125: 1963: 1936: 477:for Elpistostegalia (and thus Tetrapoda): 123: 2087: 2075:Zoological Journal of the Linnean Society 1864: 1854: 1805: 1795: 1733: 1715: 1656: 1654: 1642: 2217:and the origin of the vertebrate hand". 1537: 2122: 547:The name, originally coined around the 473:has identified the following traits as 4786: 1769: 1765: 1763: 1761: 1651: 572:. Elpistostegalia was re-defined as a 449:(around 385 million years old) or the 4694: 4693: 4533: 3339: 2431: 2313: 2268: 2102: 2027:Physiological and Biochemical Zoology 1888:Lewis, William M. Jr. (1 June 1970). 1887: 560:, the last "fishes" of the tetrapod 2432: 2132:Ahlberg, P.E; Johanson, Z. (1998). 1822: 1758: 13: 4534: 2089:10.1111/j.1096-3642.1998.tb02526.x 1951:10.1111/j.1502-3931.1990.tb01357.x 14: 4810: 4799:Middle Devonian first appearances 2018:Gordon, M.S.; Long, J.A. (2004). 1602:Long, J. A.; Holland, T. (2008). 1156:revealed that the paired fins of 485:is hinged, the hinge forming the 4674: 4673: 4516: 4509: 4502: 4495: 4488: 4481: 3322: 3316: 3311: 3304: 1964:Armbruster, Jonathan W. (1998). 1644:10.1111/j.0031-0239.2000.00138.x 1380: 1356: 1326: 1296: 1239: 1202: 1018: 996: 968: 938: 909: 880: 850: 820: 790: 732: 695: 613: 461:, around 395 million years ago. 411:redefined to include tetrapods. 350: 145: 48: 2206: 2191: 2096: 2061: 1930: 414: 1881: 1681: 1618: 1595: 1: 4638:Polydactyly in stem-tetrapods 1460: 469:Paleontologist and professor 1835:Proc. Natl. Acad. Sci. U.S.A 1797:10.1371/journal.pone.0033683 542: 424:lakes and swampland, modern 7: 1152:The 2020 study by Cloutier 10: 4815: 2105:"Getting a Leg Up on Land" 1717:10.1038/s41586-022-04990-w 385:-like characters, such as 4702: 4669: 4625: 4544: 4540: 4529: 4479: 4433: 4398: 4385: 4321: 4284: 4139: 4094: 4071: 4059: 4031: 3986: 3971: 3959: 3915: 3877: 3842: 3827: 3818: 3791: 3762: 3750: 3602: 3587: 3546: 3376: 3364: 3360: 3335: 3301: 3255: 3176: 3167: 3009: 2990: 2981: 2869: 2816: 2693: 2455: 2440: 2427: 2395: 2377: 2364: 2309: 2239:10.1038/s41586-020-2100-8 2200:American Museum Novitates 1675:10.1017/S1755691018000907 1483:and the origin of digits" 1372: 1348: 1341: 1319: 1312: 1288: 1281: 1264: 1257: 1232: 1225: 1218: 1194: 1187: 1012: 990: 983: 961: 954: 932: 925: 903: 896: 872: 865: 843: 836: 812: 805: 783: 776: 747: 725: 718: 711: 687: 680: 660: 653: 636: 629: 607: 600: 464: 455:a series of trace fossils 334: 327: 245: 240: 142:Scientific classification 140: 131: 122: 23: 1168:(digit bones) of modern 235:Camp & Allison, 1961 16:Clade of tetrapodomorphs 1856:10.1073/pnas.1011287107 391:, are sometimes called 4633:Evolution of tetrapods 2068:Ahlberg, P.E. (1998). 4766:Paleobiology Database 2103:Clack, J. A. (2005). 1479:"The pectoral fin of 377:and all more derived 399:) was considered an 4653:Zachelmie trackways 3833:Pseudophlegethontia 2231:2020Natur.579..549C 2155:1998Natur.395..792A 2109:Scientific American 1847:2010PNAS..10717911D 1788:2012PLoSO...733683S 1770:Swartz, B. (2012). 1708:2022Natur.608..563S 1573:10.1038/nature08623 1565:2010Natur.463...43N 1507:10.1038/nature07339 1499:2008Natur.456..636B 589:from Swartz, 2012. 428:and some genera of 358:Paleontology portal 4575:Megalichthyiformes 3137:Platycephalichthys 664:Platycephalichthys 558:evolutionary grade 405:lobe-finned fishes 4794:Elpistostegalians 4781: 4780: 4696:Taxon identifiers 4687: 4686: 4665: 4664: 4661: 4660: 4581:Tristichopteridae 4525: 4524: 4477: 4476: 4473: 4472: 4469: 4468: 4135: 4134: 4027: 4026: 3873: 3872: 3869: 3868: 3820:Phlegethontioidea 3583: 3582: 3551:Elginerpetontidae 3331: 3330: 3299: 3298: 3295: 3294: 3291: 3290: 3287: 3286: 3216:Parapanderichthys 3014:Tristichopteridae 2983:Eotetrapodiformes 2423: 2422: 2419: 2418: 2415: 2414: 2225:(7800): 549–554. 1702:(7923): 563–568. 1493:(7222): 636–638. 1457: 1456: 1448: 1447: 1439: 1438: 1430: 1429: 1421: 1420: 1412: 1411: 1403: 1402: 1394: 1393: 1149: 1148: 1140: 1139: 1131: 1130: 1122: 1121: 1113: 1112: 1104: 1103: 1095: 1094: 1086: 1085: 1077: 1076: 1068: 1067: 1059: 1058: 1050: 1049: 1041: 1040: 1032: 1031: 762: 761: 610:Tristichopteridae 597:Eotetrapodiformes 445:are dated to the 346: 345: 340: 284:Parapanderichthys 236: 218:Eotetrapodiformes 117: 4806: 4774: 4773: 4761: 4760: 4748: 4747: 4738: 4737: 4736: 4723: 4722: 4721: 4691: 4690: 4677: 4676: 4587:Labyrinthodontia 4542: 4541: 4531: 4530: 4520: 4514: 4513: 4507: 4506: 4500: 4499: 4493: 4492: 4485: 4442: 4438: 4404: 4396: 4395: 4388: 4327: 4324: 4287: 4142: 4069: 4068: 4062: 4034: 3969: 3968: 3962: 3918: 3880: 3844:Phlegethontiidae 3825: 3824: 3760: 3759: 3753: 3738: 3728: 3718: 3708: 3698: 3688: 3678: 3668: 3658: 3648: 3638: 3628: 3618: 3608: 3600: 3599: 3596: 3590: 3549: 3534: 3524: 3514: 3504: 3494: 3483: 3473: 3463: 3453: 3443: 3433: 3423: 3412: 3402: 3392: 3382: 3374: 3373: 3370: 3362: 3361: 3355: 3346: 3337: 3336: 3326: 3320: 3315: 3309: 3308: 3270: 3261: 3243: 3233: 3223: 3213: 3203: 3193: 3182: 3174: 3173: 3012: 2996: 2988: 2987: 2872: 2819: 2696: 2681: 2671: 2661: 2651: 2641: 2631: 2621: 2611: 2601: 2591: 2581: 2571: 2561: 2551: 2541: 2531: 2521: 2511: 2501: 2491: 2481: 2471: 2461: 2453: 2452: 2442: 2441: 2429: 2428: 2375: 2374: 2366: 2365: 2359: 2358: 2311: 2310: 2303: 2299: 2296:Tetrapodomorpha 2289: 2282: 2275: 2266: 2265: 2259: 2258: 2210: 2204: 2203: 2195: 2189: 2188: 2186: 2185: 2179: 2173:. Archived from 2138: 2129: 2120: 2119: 2117: 2115: 2100: 2094: 2093: 2091: 2065: 2059: 2058: 2024: 2015: 2006: 2005: 2003: 2001: 1970: 1961: 1955: 1954: 1934: 1928: 1927: 1925: 1919:. Archived from 1894: 1885: 1879: 1878: 1868: 1858: 1826: 1820: 1819: 1809: 1799: 1767: 1756: 1755: 1737: 1719: 1685: 1679: 1678: 1658: 1649: 1648: 1646: 1622: 1616: 1615: 1599: 1593: 1592: 1544: 1535: 1534: 1474: 1385: 1384: 1361: 1360: 1344: 1343: 1330: 1315: 1314: 1301: 1300: 1284: 1283: 1260: 1259: 1243: 1228: 1227: 1221: 1220: 1207: 1206: 1190: 1189: 1180: 1179: 1160:contained bones 1023: 1022: 1001: 1000: 986: 985: 972: 957: 956: 943: 942: 928: 927: 914: 913: 899: 898: 885: 884: 868: 867: 854: 839: 838: 825: 824: 808: 807: 794: 779: 778: 736: 721: 720: 714: 713: 700: 699: 683: 682: 656: 655: 632: 631: 618: 617: 603: 602: 593: 592: 360: 355: 354: 353: 338: 337:Panderichthyida 311: 301: 291: 281: 271: 261: 250: 234: 150: 149: 127: 111: 110: 47: 27:Temporal range: 21: 20: 4814: 4813: 4809: 4808: 4807: 4805: 4804: 4803: 4784: 4783: 4782: 4777: 4769: 4764: 4756: 4751: 4743: 4741: 4734:Elpistostegalia 4732: 4731: 4726: 4717: 4716: 4711: 4704:Elpistostegalia 4698: 4688: 4683: 4657: 4621: 4597:Ichthyostegalia 4565:Osteolepiformes 4549: 4536: 4521: 4508: 4501: 4494: 4487: 4465: 4440: 4436: 4435:Reptiliomorpha 4429: 4402: 4389: 4386: 4381: 4325: 4317: 4289:Gephyrostegidae 4280: 4181:Carbonoherpeton 4131: 4090: 4055: 4023: 4018:Palaeomolgophis 3982: 3955: 3911: 3865: 3838: 3814: 3793:Oestocephalidae 3787: 3768:Andersonerpeton 3746: 3741:Tantallognathus 3594: 3588: 3579: 3567:Obruchevichthys 3542: 3368: 3356: 3350: 3344: 3327: 3303: 3283: 3265: 3259: 3251: 3169:Elpistostegalia 3163: 3158:Tristichopterus 3005: 2999:Palaeospondylus 2977: 2971:Sterropterygion 2905:Ectosteorhachis 2890:Cladarosymblema 2874:Megalichthyidae 2865: 2812: 2749:Propycnoctenion 2689: 2449: 2447:Tetrapodomorpha 2436: 2434:Tetrapodomorpha 2411: 2397:Tetrapodomorpha 2391: 2360: 2316: 2315: 2305: 2301: 2297: 2293: 2263: 2262: 2211: 2207: 2196: 2192: 2183: 2181: 2177: 2149:(6704): 792–4. 2136: 2130: 2123: 2113: 2111: 2101: 2097: 2082:(1–2): 99–141. 2066: 2062: 2022: 2016: 2009: 1999: 1997: 1987:10.2307/1447796 1968: 1962: 1958: 1935: 1931: 1923: 1909:10.2307/1441653 1892: 1886: 1882: 1841:(42): 17911–5. 1827: 1823: 1768: 1759: 1686: 1682: 1659: 1652: 1623: 1619: 1600: 1596: 1559:(7277): 43–48. 1545: 1538: 1475: 1468: 1463: 1458: 1449: 1440: 1431: 1422: 1413: 1404: 1395: 1379: 1355: 1295: 1201: 1184:Elpistostegalia 1150: 1141: 1132: 1123: 1114: 1105: 1096: 1087: 1078: 1069: 1060: 1051: 1042: 1033: 1017: 995: 937: 908: 879: 819: 763: 694: 677:Elpistostegalia 612: 582:and tetrapods. 545: 506:shoulder girdle 504:A rather small 487:profundus nerve 467: 417: 403:of prehistoric 397:Panderichthyida 365:Elpistostegalia 356: 351: 349: 339:Vorobyeva, 1989 233: 231:Elpistostegalia 205:Tetrapodomorpha 144: 118: 116:record, 395 Ma) 109: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 42: 41: 30:Middle Devonian 28: 25: 24:Elpistostegalia 17: 12: 11: 5: 4812: 4802: 4801: 4796: 4779: 4778: 4776: 4775: 4762: 4749: 4739: 4724: 4708: 4706: 4700: 4699: 4685: 4684: 4682: 4681: 4670: 4667: 4666: 4663: 4662: 4659: 4658: 4656: 4655: 4650: 4645: 4640: 4635: 4629: 4627: 4623: 4622: 4620: 4619: 4617:Anthracosauria 4614: 4612:Phyllospondyli 4609: 4604: 4599: 4594: 4589: 4584: 4578: 4572: 4570:Osteolepididae 4567: 4562: 4560:Stem tetrapoda 4556: 4554: 4538: 4537: 4535:Related topics 4527: 4526: 4523: 4522: 4480: 4478: 4475: 4474: 4471: 4470: 4467: 4466: 4464: 4463: 4462: 4461: 4455: 4453:Reptiliomorpha 4446: 4444: 4431: 4430: 4428: 4427: 4426: 4425: 4419: 4410: 4408: 4406:Batrachomorpha 4393: 4383: 4382: 4380: 4379: 4372: 4365: 4358: 4351: 4343: 4335: 4333: 4319: 4318: 4316: 4315: 4308: 4301: 4298:Bruktererpeton 4293: 4291: 4282: 4281: 4279: 4278: 4275:Spondylerpeton 4271: 4264: 4257: 4254:Proterogyrinus 4250: 4243: 4236: 4233:Palaeoherpeton 4229: 4222: 4215: 4207: 4200: 4197:Diplovertebron 4193: 4184: 4177: 4174:Calligenethlon 4170: 4163: 4156: 4153:Anthracosaurus 4148: 4146: 4137: 4136: 4133: 4132: 4130: 4129: 4126:Megalocephalus 4122: 4115: 4108: 4100: 4098: 4092: 4091: 4089: 4088: 4085:Spathicephalus 4081: 4072: 4066: 4057: 4056: 4054: 4053: 4052: 4051: 4042: 4040: 4029: 4028: 4025: 4024: 4022: 4021: 4014: 4011:Dolichopareias 4007: 4004:Adelospondylus 4000: 3992: 3990: 3988:Adelogyrinidae 3984: 3983: 3981: 3980: 3972: 3966: 3957: 3956: 3954: 3953: 3946: 3939: 3932: 3924: 3922: 3913: 3912: 3910: 3909: 3902: 3895: 3886: 3884: 3882:Whatcheeriidae 3875: 3874: 3871: 3870: 3867: 3866: 3864: 3863: 3856: 3848: 3846: 3840: 3839: 3837: 3836: 3828: 3822: 3816: 3815: 3813: 3812: 3805: 3802:Coloraderpeton 3797: 3795: 3789: 3788: 3786: 3785: 3778: 3771: 3763: 3757: 3748: 3747: 3745: 3744: 3734: 3724: 3714: 3704: 3694: 3684: 3674: 3664: 3654: 3644: 3634: 3624: 3614: 3603: 3597: 3585: 3584: 3581: 3580: 3578: 3577: 3570: 3563: 3555: 3553: 3544: 3543: 3541: 3540: 3530: 3520: 3510: 3500: 3490: 3479: 3469: 3459: 3449: 3439: 3429: 3419: 3408: 3398: 3388: 3377: 3371: 3358: 3357: 3333: 3332: 3329: 3328: 3302: 3300: 3297: 3296: 3293: 3292: 3289: 3288: 3285: 3284: 3282: 3281: 3280: 3279: 3273: 3271: 3253: 3252: 3250: 3249: 3239: 3229: 3219: 3209: 3199: 3189: 3177: 3171: 3165: 3164: 3162: 3161: 3154: 3147: 3140: 3133: 3126: 3122:Marsdenichthys 3118: 3111: 3103: 3096: 3089: 3082: 3075: 3072:Eusthenopteron 3068: 3061: 3054: 3047: 3040: 3037:Callistipterus 3033: 3026: 3018: 3016: 3007: 3006: 3004: 3003: 2991: 2985: 2979: 2978: 2976: 2975: 2967: 2964:Sengoerichthys 2960: 2953: 2950:Palatinichthys 2946: 2938: 2930: 2923: 2916: 2908: 2901: 2893: 2886: 2878: 2876: 2867: 2866: 2864: 2863: 2855: 2851:Marsdenichthys 2847: 2840: 2833: 2825: 2823: 2814: 2813: 2811: 2810: 2802: 2794: 2787: 2780: 2773: 2766: 2759: 2752: 2745: 2738: 2731: 2724: 2717: 2710: 2702: 2700: 2691: 2690: 2688: 2687: 2677: 2667: 2657: 2647: 2637: 2634:Platyethmoidia 2627: 2617: 2607: 2597: 2594:Megadonichthys 2587: 2577: 2567: 2557: 2547: 2537: 2534:Harajicadectes 2527: 2517: 2507: 2497: 2487: 2477: 2467: 2456: 2450: 2445: 2438: 2437: 2425: 2424: 2421: 2420: 2417: 2416: 2413: 2412: 2410: 2409: 2408: 2407: 2401: 2399: 2393: 2392: 2390: 2389: 2388: 2387: 2378: 2372: 2362: 2361: 2357: 2356: 2347: 2338: 2329: 2323: 2307: 2306: 2292: 2291: 2284: 2277: 2269: 2261: 2260: 2205: 2190: 2121: 2095: 2060: 2039:10.1086/425183 2033:(5): 700–719. 2007: 1981:(3): 663–675. 1956: 1945:(2): 157–166. 1929: 1926:on 2011-08-13. 1903:(2): 319–326. 1880: 1821: 1757: 1680: 1669:(1–2): 61–73. 1650: 1637:(3): 533–548. 1617: 1594: 1536: 1465: 1464: 1462: 1459: 1455: 1454: 1451: 1450: 1446: 1445: 1442: 1441: 1437: 1436: 1433: 1432: 1428: 1427: 1424: 1423: 1419: 1418: 1415: 1414: 1410: 1409: 1406: 1405: 1401: 1400: 1397: 1396: 1392: 1391: 1388: 1387: 1371: 1368: 1367: 1364: 1363: 1347: 1342: 1340: 1337: 1336: 1333: 1332: 1318: 1313: 1311: 1308: 1307: 1304: 1303: 1287: 1282: 1280: 1277: 1276: 1273: 1272: 1263: 1258: 1256: 1250: 1249: 1246: 1245: 1231: 1226: 1224: 1219: 1217: 1214: 1213: 1210: 1209: 1193: 1188: 1186: 1178: 1147: 1146: 1143: 1142: 1138: 1137: 1134: 1133: 1129: 1128: 1125: 1124: 1120: 1119: 1116: 1115: 1111: 1110: 1107: 1106: 1102: 1101: 1098: 1097: 1093: 1092: 1089: 1088: 1084: 1083: 1080: 1079: 1075: 1074: 1071: 1070: 1066: 1065: 1062: 1061: 1057: 1056: 1053: 1052: 1048: 1047: 1044: 1043: 1039: 1038: 1035: 1034: 1030: 1029: 1026: 1025: 1011: 1008: 1007: 1004: 1003: 989: 984: 982: 979: 978: 975: 974: 960: 955: 953: 950: 949: 946: 945: 931: 926: 924: 921: 920: 917: 916: 906:Whatcheeriidae 902: 897: 895: 892: 891: 888: 887: 871: 866: 864: 861: 860: 857: 856: 842: 837: 835: 832: 831: 828: 827: 811: 806: 804: 801: 800: 797: 796: 782: 777: 775: 769: 768: 765: 764: 760: 759: 756: 755: 746: 743: 742: 739: 738: 724: 719: 717: 712: 710: 707: 706: 703: 702: 686: 681: 679: 673: 672: 669: 668: 659: 654: 652: 649: 648: 645: 644: 635: 630: 628: 625: 624: 621: 620: 606: 601: 599: 591: 544: 541: 540: 539: 512: 509: 502: 495:parietal bones 493:, between the 471:Per E. Ahlberg 466: 463: 416: 413: 379:tetrapodomorph 362: 361: 344: 343: 342: 341: 332: 331: 325: 324: 323: 322: 317: 307: 297: 287: 277: 267: 257: 243: 242: 238: 237: 228: 221: 220: 215: 208: 207: 202: 195: 194: 189: 182: 181: 176: 172: 171: 166: 162: 161: 156: 152: 151: 138: 137: 129: 128: 120: 119: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 26: 15: 9: 6: 4: 3: 2: 4811: 4800: 4797: 4795: 4792: 4791: 4789: 4772: 4767: 4763: 4759: 4754: 4750: 4746: 4740: 4735: 4729: 4725: 4720: 4714: 4710: 4709: 4707: 4705: 4701: 4697: 4692: 4680: 4672: 4671: 4668: 4654: 4651: 4649: 4646: 4644: 4641: 4639: 4636: 4634: 4631: 4630: 4628: 4624: 4618: 4615: 4613: 4610: 4608: 4605: 4603: 4600: 4598: 4595: 4593: 4590: 4588: 4585: 4582: 4579: 4576: 4573: 4571: 4568: 4566: 4563: 4561: 4558: 4557: 4555: 4552: 4547: 4543: 4539: 4532: 4528: 4519: 4515: 4512: 4505: 4498: 4491: 4484: 4460: 4456: 4454: 4450: 4449: 4448: 4447: 4445: 4443: 4432: 4424: 4420: 4418: 4417:Temnospondyli 4414: 4413: 4412: 4411: 4409: 4407: 4401: 4400:Temnospondyli 4397: 4394: 4392: 4384: 4378: 4377: 4376:Termonerpeton 4373: 4371: 4370: 4369:Silvanerpeton 4366: 4364: 4363: 4359: 4357: 4356: 4352: 4349: 4348: 4344: 4342: 4341: 4337: 4336: 4334: 4331: 4330:anthracosaurs 4320: 4314: 4313: 4312:Gephyrostegus 4309: 4307: 4306: 4305:Eusauropleura 4302: 4300: 4299: 4295: 4294: 4292: 4290: 4283: 4277: 4276: 4272: 4270: 4269: 4265: 4263: 4262: 4258: 4256: 4255: 4251: 4249: 4248: 4247:Pholiderpeton 4244: 4242: 4241: 4237: 4235: 4234: 4230: 4228: 4227: 4223: 4221: 4220: 4219:Leptophractus 4216: 4213: 4212: 4208: 4206: 4205: 4201: 4199: 4198: 4194: 4191: 4190: 4185: 4183: 4182: 4178: 4176: 4175: 4171: 4169: 4168: 4164: 4162: 4161: 4157: 4155: 4154: 4150: 4149: 4147: 4145: 4138: 4128: 4127: 4123: 4121: 4120: 4116: 4114: 4113: 4109: 4107: 4106: 4102: 4101: 4099: 4097: 4093: 4087: 4086: 4082: 4079: 4078: 4074: 4073: 4070: 4067: 4065: 4058: 4050: 4046: 4045: 4044: 4043: 4041: 4038: 4030: 4020: 4019: 4015: 4013: 4012: 4008: 4006: 4005: 4001: 3999: 3998: 3994: 3993: 3991: 3989: 3985: 3979: 3978: 3977:Acherontiscus 3974: 3973: 3970: 3967: 3965: 3964:Adelospondyli 3958: 3952: 3951: 3950:Pholidogaster 3947: 3945: 3944: 3940: 3938: 3937: 3936:Deltaherpeton 3933: 3931: 3930: 3926: 3925: 3923: 3921: 3914: 3908: 3907: 3903: 3901: 3900: 3896: 3893: 3892: 3888: 3887: 3885: 3883: 3876: 3862: 3861: 3857: 3855: 3854: 3853:Phlegethontia 3850: 3849: 3847: 3845: 3841: 3835: 3834: 3830: 3829: 3826: 3823: 3821: 3817: 3811: 3810: 3809:Oestocephalus 3806: 3804: 3803: 3799: 3798: 3796: 3794: 3790: 3784: 3783: 3779: 3777: 3776: 3772: 3770: 3769: 3765: 3764: 3761: 3758: 3756: 3749: 3743: 3742: 3735: 3733: 3732: 3725: 3723: 3722: 3715: 3713: 3712: 3705: 3703: 3702: 3695: 3693: 3692: 3685: 3683: 3682: 3675: 3673: 3672: 3671:Ichthyerpeton 3665: 3663: 3662: 3655: 3653: 3652: 3645: 3643: 3642: 3635: 3633: 3632: 3631:Crassigyrinus 3625: 3623: 3622: 3615: 3613: 3612: 3605: 3604: 3601: 3598: 3593: 3586: 3576: 3575: 3571: 3569: 3568: 3564: 3562: 3561: 3557: 3556: 3554: 3552: 3545: 3539: 3538: 3531: 3529: 3528: 3521: 3519: 3518: 3511: 3509: 3508: 3501: 3499: 3498: 3491: 3488: 3487: 3480: 3478: 3477: 3470: 3468: 3467: 3460: 3458: 3457: 3456:Metaxygnathus 3450: 3448: 3447: 3440: 3438: 3437: 3430: 3428: 3427: 3420: 3417: 3416: 3409: 3407: 3406: 3399: 3397: 3396: 3395:Brittagnathus 3389: 3387: 3386: 3379: 3378: 3375: 3372: 3367: 3363: 3359: 3353: 3349: 3343: 3338: 3334: 3325: 3321: 3319: 3314: 3307: 3277: 3276: 3275: 3274: 3272: 3268: 3264: 3258: 3254: 3248: 3247: 3240: 3238: 3237: 3230: 3228: 3227: 3220: 3218: 3217: 3210: 3208: 3207: 3206:Panderichthys 3200: 3198: 3197: 3190: 3187: 3186: 3185:Howittichthys 3179: 3178: 3175: 3172: 3170: 3166: 3160: 3159: 3155: 3153: 3152: 3148: 3146: 3145: 3141: 3139: 3138: 3134: 3132: 3131: 3127: 3124: 3123: 3119: 3117: 3116: 3112: 3109: 3108: 3104: 3102: 3101: 3097: 3095: 3094: 3090: 3088: 3087: 3083: 3081: 3080: 3079:Heddleichthys 3076: 3074: 3073: 3069: 3067: 3066: 3062: 3060: 3059: 3055: 3053: 3052: 3048: 3046: 3045: 3041: 3039: 3038: 3034: 3032: 3031: 3030:Cabonnichthys 3027: 3025: 3024: 3023:Bruehnopteron 3020: 3019: 3017: 3015: 3008: 3001: 3000: 2993: 2992: 2989: 2986: 2984: 2980: 2973: 2972: 2968: 2966: 2965: 2961: 2959: 2958: 2954: 2952: 2951: 2947: 2944: 2943: 2939: 2936: 2935: 2931: 2929: 2928: 2924: 2922: 2921: 2917: 2914: 2913: 2909: 2907: 2906: 2902: 2899: 2898: 2894: 2892: 2891: 2887: 2885: 2884: 2880: 2879: 2877: 2875: 2868: 2861: 2860: 2856: 2853: 2852: 2848: 2846: 2845: 2841: 2839: 2838: 2834: 2832: 2831: 2827: 2826: 2824: 2822: 2821:Canowindridae 2815: 2808: 2807: 2803: 2800: 2799: 2795: 2793: 2792: 2788: 2786: 2785: 2781: 2779: 2778: 2774: 2772: 2771: 2767: 2765: 2764: 2760: 2758: 2757: 2753: 2751: 2750: 2746: 2744: 2743: 2739: 2737: 2736: 2732: 2730: 2729: 2725: 2723: 2722: 2718: 2716: 2715: 2711: 2709: 2708: 2704: 2703: 2701: 2699: 2692: 2686: 2685: 2678: 2676: 2675: 2668: 2666: 2665: 2658: 2656: 2655: 2654:Thaumatolepis 2648: 2646: 2645: 2638: 2636: 2635: 2628: 2626: 2625: 2618: 2616: 2615: 2608: 2606: 2605: 2598: 2596: 2595: 2588: 2586: 2585: 2578: 2576: 2575: 2568: 2566: 2565: 2564:Lamprotolepis 2558: 2556: 2555: 2548: 2546: 2545: 2538: 2536: 2535: 2528: 2526: 2525: 2518: 2516: 2515: 2514:Greiserolepis 2508: 2506: 2505: 2498: 2496: 2495: 2488: 2486: 2485: 2478: 2476: 2475: 2468: 2466: 2465: 2458: 2457: 2454: 2451: 2448: 2443: 2439: 2435: 2430: 2426: 2405: 2404: 2403: 2402: 2400: 2398: 2394: 2386: 2385:Gnathostomata 2382: 2381: 2380: 2379: 2376: 2373: 2371: 2370:Gnathostomata 2367: 2363: 2355: 2351: 2348: 2346: 2345:Sarcopterygii 2342: 2339: 2337: 2333: 2330: 2328: 2324: 2322: 2318: 2317: 2312: 2308: 2304: 2300:Pan-Tetrapoda 2290: 2285: 2283: 2278: 2276: 2271: 2270: 2267: 2256: 2252: 2248: 2244: 2240: 2236: 2232: 2228: 2224: 2220: 2216: 2209: 2202:(3189): 1–20. 2201: 2194: 2180:on 2014-11-24 2176: 2172: 2168: 2164: 2163:10.1038/27421 2160: 2156: 2152: 2148: 2144: 2143: 2135: 2128: 2126: 2110: 2106: 2099: 2090: 2085: 2081: 2077: 2076: 2071: 2064: 2056: 2052: 2048: 2044: 2040: 2036: 2032: 2028: 2021: 2014: 2012: 1996: 1992: 1988: 1984: 1980: 1976: 1975: 1967: 1960: 1952: 1948: 1944: 1940: 1933: 1922: 1918: 1914: 1910: 1906: 1902: 1898: 1891: 1884: 1876: 1872: 1867: 1862: 1857: 1852: 1848: 1844: 1840: 1836: 1832: 1825: 1817: 1813: 1808: 1803: 1798: 1793: 1789: 1785: 1782:(3): e33683. 1781: 1777: 1773: 1766: 1764: 1762: 1753: 1749: 1745: 1741: 1736: 1731: 1727: 1723: 1718: 1713: 1709: 1705: 1701: 1697: 1696: 1691: 1684: 1676: 1672: 1668: 1664: 1657: 1655: 1645: 1640: 1636: 1632: 1631:Palaeontology 1628: 1621: 1614:(1): 184–193. 1613: 1609: 1605: 1598: 1590: 1586: 1582: 1578: 1574: 1570: 1566: 1562: 1558: 1554: 1550: 1543: 1541: 1532: 1528: 1524: 1520: 1516: 1512: 1508: 1504: 1500: 1496: 1492: 1488: 1484: 1482: 1481:Panderichthys 1473: 1471: 1466: 1453: 1452: 1444: 1443: 1435: 1434: 1426: 1425: 1417: 1416: 1408: 1407: 1399: 1398: 1390: 1389: 1386: 1383: 1378: 1377: 1370: 1369: 1366: 1365: 1362: 1359: 1354: 1353: 1346: 1345: 1339: 1338: 1335: 1334: 1331: 1329: 1325: 1324: 1317: 1316: 1310: 1309: 1306: 1305: 1302: 1299: 1294: 1293: 1286: 1285: 1279: 1278: 1275: 1274: 1271: 1270: 1269: 1262: 1261: 1255: 1252: 1251: 1248: 1247: 1244: 1242: 1238: 1237: 1230: 1229: 1223: 1222: 1216: 1215: 1212: 1211: 1208: 1205: 1200: 1199: 1198:Panderichthys 1192: 1191: 1185: 1182: 1181: 1177: 1175: 1171: 1167: 1163: 1159: 1155: 1145: 1144: 1136: 1135: 1127: 1126: 1118: 1117: 1109: 1108: 1100: 1099: 1091: 1090: 1082: 1081: 1073: 1072: 1064: 1063: 1055: 1054: 1046: 1045: 1037: 1036: 1028: 1027: 1024: 1021: 1016: 1010: 1009: 1006: 1005: 1002: 999: 994: 988: 987: 981: 980: 977: 976: 973: 971: 967: 966: 965:Crassigyrinus 959: 958: 952: 951: 948: 947: 944: 941: 936: 930: 929: 923: 922: 919: 918: 915: 912: 907: 901: 900: 894: 893: 890: 889: 886: 883: 878: 877: 870: 869: 863: 862: 859: 858: 855: 853: 849: 848: 841: 840: 834: 833: 830: 829: 826: 823: 818: 817: 810: 809: 803: 802: 799: 798: 795: 793: 789: 788: 781: 780: 774: 771: 770: 767: 766: 758: 757: 754: 753: 752: 745: 744: 741: 740: 737: 735: 731: 730: 723: 722: 716: 715: 709: 708: 705: 704: 701: 698: 693: 692: 691:Panderichthys 685: 684: 678: 675: 674: 671: 670: 667: 666: 665: 658: 657: 651: 650: 647: 646: 643: 642: 641: 634: 633: 627: 626: 623: 622: 619: 616: 611: 605: 604: 598: 595: 594: 590: 588: 583: 581: 580: 579:Panderichthys 575: 571: 567: 563: 559: 555: 554: 550: 537: 533: 529: 525: 521: 517: 513: 510: 507: 503: 500: 496: 492: 488: 484: 480: 479: 478: 476: 475:synapomorphic 472: 462: 460: 456: 452: 448: 444: 443: 442:Panderichthys 437: 433: 431: 427: 423: 412: 410: 409:cladistically 406: 402: 398: 394: 390: 389: 384: 380: 376: 375: 374:Panderichthys 370: 366: 359: 348: 336: 335: 333: 330: 326: 321: 318: 316: 315: 308: 306: 305: 298: 296: 295: 288: 286: 285: 278: 276: 275: 274:Panderichthys 268: 266: 265: 258: 255: 254: 253:Howittichthys 247: 246: 244: 239: 232: 229: 226: 223: 222: 219: 216: 213: 210: 209: 206: 203: 200: 197: 196: 193: 192:Sarcopterygii 190: 187: 184: 183: 180: 177: 174: 173: 170: 167: 164: 163: 160: 157: 154: 153: 148: 143: 139: 136: 135: 134:Panderichthys 130: 126: 121: 115: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 46: 39: 35: 31: 22: 19: 4703: 4626:Other topics 4607:Holospondyli 4602:Lepospondyli 4551:Polyphyletic 4546:Paraphyletic 4486: 4423:Lissamphibia 4374: 4367: 4360: 4353: 4345: 4340:Caerorhachis 4338: 4310: 4303: 4296: 4273: 4268:Seroherpeton 4266: 4259: 4252: 4245: 4238: 4231: 4226:Neopteroplax 4224: 4217: 4209: 4202: 4195: 4187: 4179: 4172: 4165: 4158: 4151: 4124: 4117: 4110: 4103: 4083: 4075: 4016: 4009: 4002: 3997:Adelogyrinus 3995: 3975: 3948: 3943:Greererpeton 3941: 3934: 3927: 3904: 3897: 3889: 3858: 3851: 3831: 3807: 3800: 3782:Ophiderpeton 3780: 3773: 3766: 3739: 3729: 3719: 3709: 3699: 3691:Mesanerpeton 3689: 3679: 3669: 3659: 3649: 3639: 3629: 3621:Aytonerpeton 3619: 3609: 3574:Webererpeton 3572: 3565: 3560:Elginerpeton 3558: 3535: 3525: 3515: 3505: 3495: 3484: 3474: 3464: 3454: 3444: 3436:Ichthyostega 3434: 3424: 3415:Elpistostege 3413: 3405:Densignathus 3403: 3393: 3385:Acanthostega 3383: 3351: 3342:Stegocephali 3310: 3266: 3257:Stegocephali 3244: 3236:Rubrognathus 3234: 3224: 3214: 3204: 3194: 3183: 3168: 3156: 3149: 3142: 3135: 3130:Notorhizodon 3128: 3120: 3113: 3105: 3098: 3091: 3084: 3077: 3070: 3063: 3056: 3049: 3042: 3035: 3028: 3021: 2997: 2969: 2962: 2955: 2948: 2942:Megistolepis 2940: 2932: 2927:Megalichthys 2925: 2918: 2910: 2903: 2895: 2888: 2883:Askerichthys 2881: 2857: 2849: 2842: 2835: 2828: 2804: 2798:Thysanolepis 2796: 2789: 2782: 2775: 2768: 2761: 2756:Pycnoctenion 2754: 2747: 2740: 2733: 2728:Gooloogongia 2726: 2719: 2712: 2705: 2698:Rhizodontida 2682: 2672: 2662: 2652: 2642: 2632: 2622: 2612: 2604:Muranjilepis 2602: 2592: 2582: 2572: 2562: 2552: 2542: 2532: 2524:Gyroptychius 2522: 2512: 2502: 2492: 2482: 2472: 2462: 2349: 2340: 2336:Osteichthyes 2331: 2222: 2218: 2215:Elpistostege 2214: 2208: 2199: 2193: 2182:. Retrieved 2175:the original 2146: 2140: 2112:. Retrieved 2108: 2098: 2079: 2073: 2063: 2030: 2026: 1998:. Retrieved 1978: 1972: 1959: 1942: 1938: 1932: 1921:the original 1900: 1896: 1883: 1838: 1834: 1824: 1779: 1775: 1699: 1693: 1683: 1666: 1662: 1634: 1630: 1620: 1611: 1607: 1597: 1556: 1552: 1490: 1486: 1480: 1374: 1373: 1352:Ichthyostega 1350: 1349: 1323:Acanthostega 1321: 1320: 1290: 1289: 1268:Elpistostege 1266: 1265: 1254:Stegocephali 1234: 1233: 1196: 1195: 1183: 1174:Elpistostege 1173: 1158:Elpistostege 1157: 1153: 1151: 1013: 991: 963: 962: 933: 904: 876:Ichthyostega 874: 873: 847:Acanthostega 845: 844: 814: 813: 787:Elginerpeton 785: 784: 773:Stegocephali 751:Elpistostege 749: 748: 727: 726: 689: 688: 676: 662: 661: 638: 637: 608: 584: 577: 553:Elpistostege 551: 546: 499:postparietal 468: 440: 438: 434: 418: 415:Paleobiology 396: 392: 386: 372: 364: 363: 320:Stegocephali 312: 304:Rubrognathus 302: 292: 282: 272: 262: 251: 230: 224: 211: 198: 185: 132: 18: 4728:Wikispecies 4643:Romer's gap 4439:Pan-Amniota 4387:Crown group 4240:Papposaurus 4064:Baphetoidea 3920:Colosteidae 3906:Whatcheeria 3860:Sillerpeton 3651:Doragnathus 3611:Antlerpeton 3144:Spodichthys 3107:Litoptychus 3065:Eusthenodon 3058:Edenopteron 3051:Devonosteus 2957:Rhizodopsis 2920:Mahalalepis 2897:Cryptolepis 2844:Koharalepis 2830:Beelarongia 2806:Vorobjevaia 2784:Screbinodus 2777:Sauripterus 2770:Rossichthys 2742:Letognathus 2707:Archichthys 2684:Viluichthys 2554:Litoptychus 2494:Glyptopomus 2474:Chrysolepis 2354:Rhipidistia 935:Colosteidae 585:Below is a 576:containing 508:is present. 483:endocranium 439:Fossils of 371:containing 43:385–0  4788:Categories 4362:Eoherpeton 4211:Eoherpeton 4204:Eobaphetes 4144:Embolomeri 4096:Baphetidae 3721:Perittodus 3641:Diploradus 3527:Ventastega 3497:Tulerpeton 3466:Parmastega 3446:Jakubsonia 3426:Hynerpeton 3352:sensu lato 3278:see below↓ 3267:sensu lato 3226:Qikiqtania 3115:Mandageria 3100:Langlieria 3044:Canningius 2837:Canowindra 2791:Strepsodus 2644:Shirolepis 2614:Osteolepis 2544:Kenichthys 2484:Geptolepis 2464:Bogdanovia 2406:see below↓ 2184:2011-03-12 1461:References 1376:Tulerpeton 1292:Ventastega 1162:homologous 993:Baphetidae 816:Ventastega 491:skull roof 294:Qikiqtania 241:Subgroups 112:(Possible 4592:Rachitomi 4457:includes 4421:includes 4391:Tetrapoda 4355:Eldeceeon 4347:Casineria 4261:Pteroplax 4049:Nectridea 4037:Nectridea 3929:Colosteus 3891:Ossinodus 3775:Lethiscus 3755:Aistopoda 3731:Sigournea 3711:Ossirarus 3517:Umzantsia 3486:Tiktaalik 3476:Sinostega 3348:Tetrapoda 3263:Tetrapoda 3246:Tiktaalik 3196:Livoniana 3093:Jarvikina 2934:Megapomus 2674:Tungsenia 2624:Peregrina 2504:Gogonasus 2319:Kingdom: 2255:213171029 1752:250730904 1726:0028-0836 1515:1476-4687 1236:Tiktaalik 1170:tetrapods 1166:phalanges 1015:Tetrapoda 729:Tiktaalik 587:cladogram 570:Tetrapoda 562:stem line 543:Phylogeny 422:eutrophic 407:, it was 393:fishapods 388:Tiktaalik 314:Tiktaalik 264:Livoniana 165:Kingdom: 159:Eukaryota 4742:BioLib: 4713:Wikidata 4679:Category 4648:Timeline 4189:Cricotus 4160:Archeria 4112:Kyrinion 4105:Baphetes 4077:Eucritta 3899:Pederpes 3701:Occidens 3592:Devonian 3507:Tutusius 3366:Devonian 2912:Lohsania 2763:Rhizodus 2721:Barameda 2664:Thursius 2584:Medoevia 2327:Chordata 2325:Phylum: 2321:Animalia 2247:32214248 2114:28 March 2047:15547790 1875:20884852 1816:22448265 1776:PLOS ONE 1744:35859171 1581:20054388 1523:18806778 497:and the 459:Eifelian 451:Frasnian 447:Givetian 426:lungfish 383:tetrapod 329:Synonyms 179:Chordata 175:Phylum: 169:Animalia 155:Domain: 114:Eifelian 34:Givetian 4719:Q150821 4459:Amniota 4167:Aversor 4119:Loxomma 3681:Koilops 3661:Gaiasia 3151:Tinirau 3086:Hyneria 2859:Owensia 2714:Aztecia 2574:Latvius 2227:Bibcode 2171:4430783 2151:Bibcode 2055:1260442 2000:25 June 1995:1447796 1939:Lethaia 1917:1441653 1866:2964239 1843:Bibcode 1807:3308997 1784:Bibcode 1735:9385497 1704:Bibcode 1589:4428903 1561:Bibcode 1531:2588617 1495:Bibcode 1164:to the 640:Tinirau 516:humerus 430:catfish 38:Present 4771:219423 4553:groups 3537:Ymeria 2735:Hongyu 2253:  2245:  2219:Nature 2169:  2142:Nature 2053:  2045:  1993:  1974:Copeia 1915:  1897:Copeia 1873:  1863:  1814:  1804:  1750:  1742:  1732:  1724:  1695:Nature 1587:  1579:  1553:Nature 1529:  1521:  1513:  1487:Nature 1154:et al. 536:fibula 534:, and 524:radius 522:, and 501:bones. 465:Traits 4758:11907 4753:IRMNG 4745:16201 4326:Other 3589:Post- 2350:Clade 2341:Clade 2332:Clade 2251:S2CID 2178:(PDF) 2167:S2CID 2137:(PDF) 2051:S2CID 2023:(PDF) 1991:JSTOR 1969:(PDF) 1924:(PDF) 1913:JSTOR 1893:(PDF) 1748:S2CID 1585:S2CID 1527:S2CID 574:clade 566:clade 549:genus 532:tibia 528:femur 401:order 369:clade 367:is a 225:Clade 212:Clade 199:Clade 186:Clade 4451:see 4415:see 4047:see 3595:taxa 3369:taxa 2383:see 2243:PMID 2116:2024 2043:PMID 2002:2009 1979:1998 1871:PMID 1812:PMID 1740:PMID 1722:ISSN 1577:PMID 1519:PMID 1511:ISSN 520:ulna 481:The 51:PreꞒ 36:) – 2235:doi 2223:579 2159:doi 2147:395 2084:doi 2080:122 2035:doi 1983:doi 1947:doi 1905:doi 1861:PMC 1851:doi 1839:107 1802:PMC 1792:doi 1730:PMC 1712:doi 1700:608 1671:doi 1667:109 1639:doi 1612:120 1569:doi 1557:463 1503:doi 1491:456 568:to 4790:: 4768:: 4755:: 4730:: 4715:: 2352:: 2343:: 2334:: 2249:. 2241:. 2233:. 2221:. 2165:. 2157:. 2145:. 2139:. 2124:^ 2107:. 2078:. 2072:. 2049:. 2041:. 2031:77 2029:. 2025:. 2010:^ 1989:. 1977:. 1971:. 1943:23 1941:. 1911:. 1899:. 1895:. 1869:. 1859:. 1849:. 1837:. 1833:. 1810:. 1800:. 1790:. 1778:. 1774:. 1760:^ 1746:. 1738:. 1728:. 1720:. 1710:. 1698:. 1692:. 1665:. 1653:^ 1635:43 1633:. 1629:. 1610:. 1606:. 1583:. 1575:. 1567:. 1555:. 1551:. 1539:^ 1525:. 1517:. 1509:. 1501:. 1489:. 1485:. 1469:^ 530:, 518:, 227:: 214:: 201:: 188:: 101:Pg 45:Ma 40:, 4583:? 4577:? 4548:/ 4441:) 4437:( 4403:( 4350:? 4332:" 4328:" 4323:† 4286:† 4214:? 4192:" 4186:" 4141:† 4080:? 4061:† 4039:" 4035:" 4033:† 3961:† 3917:† 3894:? 3879:† 3752:† 3737:† 3727:† 3717:† 3707:† 3697:† 3687:† 3677:† 3667:† 3657:† 3647:† 3637:† 3627:† 3617:† 3607:† 3548:† 3533:† 3523:† 3513:† 3503:† 3493:† 3489:? 3482:† 3472:† 3462:† 3452:† 3442:† 3432:† 3422:† 3418:? 3411:† 3401:† 3391:† 3381:† 3354:) 3345:( 3269:) 3260:( 3242:† 3232:† 3222:† 3212:† 3202:† 3192:† 3188:? 3181:† 3125:? 3110:? 3011:† 3002:? 2995:† 2974:? 2945:? 2937:? 2915:? 2900:? 2871:† 2862:? 2854:? 2818:† 2809:? 2801:? 2695:† 2680:† 2670:† 2660:† 2650:† 2640:† 2630:† 2620:† 2610:† 2600:† 2590:† 2580:† 2570:† 2560:† 2550:† 2540:† 2530:† 2520:† 2510:† 2500:† 2490:† 2480:† 2470:† 2460:† 2302:) 2298:( 2288:e 2281:t 2274:v 2257:. 2237:: 2229:: 2187:. 2161:: 2153:: 2118:. 2092:. 2086:: 2057:. 2037:: 2004:. 1985:: 1953:. 1949:: 1907:: 1901:2 1877:. 1853:: 1845:: 1818:. 1794:: 1786:: 1780:7 1754:. 1714:: 1706:: 1677:. 1673:: 1647:. 1641:: 1591:. 1571:: 1563:: 1533:. 1505:: 1497:: 538:. 310:† 300:† 290:† 280:† 270:† 260:† 256:? 249:† 106:N 96:K 91:J 86:T 81:P 76:C 71:D 66:S 61:O 56:Ꞓ 32:(

Index

Middle Devonian
Givetian
Present
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N
Eifelian

Panderichthys
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Sarcopterygii
Tetrapodomorpha
Eotetrapodiformes
Elpistostegalia
Howittichthys
Livoniana

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