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Cultural group selection

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beliefs persist over time. While individuals will rely on copying high frequency behaviors under various conditions, this reliance increases when an individual is exposed to ambiguous environmental or social information. Conformist transmission can maintain between-group variation by reducing within-group variation, but it also facilitates the rapid spread of novel ideas, which increases between-group variation. Taken together, reduced within-group variation and increased between-group variation lead to the cultural divergence between groups that is the driving force of cultural group selection.
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environment. In a study comparing human children and young chimpanzees, it was shown that, when given a demonstration on how to retrieve a reward from a box, chimps copy relevant behaviour, while ignoring irrelevant behaviour, to solve the task. Meanwhile, human children will faithfully imitate both relevant and irrelevant behaviour to solve the same task. While this may seem like a negative quality, it is what allows for reliable, high-fidelity transmission of cultural information, and produces stable behavioural equilibria within cultural groups.
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models. Individual can determine the rank of potential models by how much deference they are shown by the rest of the group. Deference is shown to high-prestige individuals to gain the opportunity to copy their successful models. We can see evidence for this bias in how new technologies, or economic practices spread to different groups according to how quick "opinion leaders" adopt them.
128:. This involves following the gaze of an adult or using them as social reference points. Put simply, they become aware of the adult's attention and behaviour towards objects in the environment. In this sense, the child is beginning to understand people as goal-oriented intentional agents. This is vitally important for learning through imitation and, eventually, language acquisition. 155:
allows one to communicate various perspectives and shift attention to one aspect of the world over another. In learning a language, a child is inheriting a vast set of linguistic symbols that have been passed down many generations. What is inherited then is the methods of shifting attention and perspective that were historically of importance to the people of that culture.
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there must exist, between groups, cultural differences that when transmitted across time affect the persistence or proliferation of the groups. Cultural norms that provide these advantages will, in turn, lead to the displacement, absorption or even extinction of other, less successful cultural groups. However,
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Without between-group variation, cultural group selection could not occur, as there would be no group differentiation to select for. While processes such as cultural drift, epidemics, and natural disasters increase between-group variation, migration and genetic mixing decrease between-group variation
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While cultural norms are often beneficial to the individuals who hold them, they need not be. Norms can spread by cultural group selection when they are practiced within successful groups, and norms are more likely to spread from groups that are successful. But, for cultural group selection to occur,
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Demographic swamping occurs when one or more cultural groups reproduces individuals faster than other groups in the region because of stable, culturally transmitted ideas or practices. This is the slowest kind of cultural groups selection as it depends on natural selection of between-group cultural
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are a perfect example of how robust this effect is and its replication across many cultures shows that this behaviour is very common. Henrich suggests that normative conformity may have evolved to respond to the spread of punishing behaviour toward non-conformists. By appearing similar to the group,
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transmission is the tendency to copy those individuals who are similar in language, appearance, social standing and other behavioral and cultural traits. In the context of prestige-biased transmission, self-similarity means that individuals will preferentially imitate those high-prestige individuals
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Direct intergroup competition is the process by which cultural groups compete with each other over resources by engaging in warfare and raiding. The cultural practices and behaviour that gives an advantage to one group over another will proliferate at the expense of those who cannot compete. There
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Prestige-biased transmission is the tendency the copy those members of the group that are more successful. Preferentially copying successful members of the group allows individuals to avoid costly trial-and-error learning by imitating the better-than-average skills of the more prestigious cultural
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games, it was found that standards of fairness and inclination to punish were correlated with both participation in world religions and market integration. This indicates how many of the behaviours necessary for complex societies are the result of cultural exposure rather than any evolution of our
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In prestige-biased group selection, when individuals have opportunities to copy people from nearby groups, they will preferentially imitate the members of groups that are more cooperative than their own. Since cooperative groups have a higher average payoff than non-cooperative groups, members of
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Through imitatively learning, the child comprehends that linguistic symbols are intended to focus attention to some specific aspect of the shared experience. In doing this, the child must be able to take the perspective of the speaker. Due to the intersubjectivity of linguistic symbols, language
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uses real-world historical, archaeological and anthropological data to test hypotheses from cultural group selection theory and other competing explanations. The Collaborative for Information and Analysis; the International Institute of Social History; and the Database of Religious History also
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Normative conformity is the act of changing one's visible behaviour, simply to appear to match the majority, and without actually internalizing the groups opinions. This differs from conformist transmission since normative conformity does not consider frequency of a behaviour as an indicator of
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Conformist transmission refers to the psychological bias to preferentially imitate high frequency behaviors in the cultural group. This homogenizes the social group and reinforces widely held cultural norms. This explains why individuals within a social group hold the same beliefs and why these
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increase within-group variation. Variation is only maintained when cultural groups have mechanisms that prevent the norms of outside groups from invading the cultural group. These ‘mechanisms’ are those uniquely human psychological traits and behaviours that encourage imitation, conformity, and
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Cultural group selection theory can provide insight into human cooperation and is therefore a useful framework for generating hypothesis related to cultural evolution. These theories, however, must be tested using empirical data: a task addressed by several large-scale projects in the field of
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For cultural knowledge and behaviour to persist across multiple generations, humans need to have the capacity to acquire, retain, and transmit cultural information. While many species engage in social learning, humans consistently rely upon it for behavioural cues and information about the
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in acquiring behaviours. These instincts allow for the acquisition and persistence of culture. Through cultural group selection, culturally specific cooperative behaviour can evolve to support large societies. For example, in a study that spanned a variety of cultures, testing behaviour in
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Non-conformists threaten to increase within-group variation by introducing deviant behaviours to the group and must receive costly punishment to maintain a homogenous social group. As a consequence of being punished, non-conformists will be less successful than other members of the group.
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one can gain the advantages of in-group membership, while also avoiding punishment. A curious byproduct of normative conformity is that it can contribute to the conformity transmission of norms that the transmitter does not hold, because they were mistakenly attributed by the imitator.
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can explain the behaviour of small social groups common in many species, it is unable to explain the large complex societies of unrelated, anonymous individuals that we see in the human species. However, one of the major distinctions between humans and other species is our reliance on
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study found that 14-month-old children will, after seeing an adult do it, bend at the waist and press a panel with their head to turn on a light, instead of using their hands. According to Tomasello, imitative learning is necessary for learning the symbolic conventions of language.
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By about 1 year of age, children begin to learn by imitation. At this point, children are capable of discriminating intentional actions from unintentional ones, and will attempt to accurately copy those intentional actions to accomplish tasks they've seen adults do. Because of
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are many possible traits that could contribute to a group's success, such as technological development, social and political organization, economic development, nationalism, etc. According to Joseph Soltis, it would take 500–1000 years for group selection to happen this way.
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who are similar to them. From the perspective of an imitator, this trait is adaptive. By only imitating those high-prestige individuals who are similar, the imitator avoids adopting traits or behaviors that are not compatible with his or her knowledge or social environment.
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models suggest that if individuals are able to migrate between groups (which is common in small-scale societies), differences between groups should be difficult to maintain. Research in psychology reveals that humans have a particular set of traits, which include
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Richerson, Peter; Baldini, Ryan; Bell, Adrian V.; Demps, Kathryn; Frost, Karl; hillis, Vickens; Mathew, Sarah; Newton, Emily K.; Naar, Nicole; Newson, Lesley; Ross, Cody; Smaldino, Paul E.; Waring, Timothy M.; Zefferman, Matthew (2016).
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Prestige-biased transmission would suggest that non-conformist behaviors would, therefore, not spread through the population. Papers on the topic suggest that this kind of punishment is prevalent across many different societies.
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Baron, R., Vandello, J., Brunsman, B., 1996. The forgotten variable in conformity research: impact of task importance on social influence. Journal of Personality and Social Psychology 71 (5), 915–927.
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Henrich, J., Boyd, R., Bowles, S., Gintis, H., Camerer, C., Fehr, E., McElreath, R., 2001. In search of Homo economicus: experiments in 15 small-scale societies. American Economic Review 91, 73–78.
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These two social biases act together in reducing within-group variation. Additionally, prestige-biased transmission increases between-group variation by contributing to the spread of novel ideas.
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of how cultural traits evolve according to the competitive advantage they bestow upon a group. This multidisciplinary approach to the question of human culture engages research from the fields of
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Campbell, D.T. Variation and selective retention in sociocultural evolution. Social change in developing areas: A reinterpretation of evolutionary theory. Cambridge: Schenkman. 1965. 19-49.
141:, children will copy those intentional acts which have no perceivable effect on the outcome, as well as strange or unnatural actions when easier methods are available. For example, an 548:
Asch, S.E., 1951. Effects of group pressure upon the modification and distortion of judgments. In: Guetzkow, H. (Ed.), Groups, Leadership and Men. Carnegie, Pittsburgh, pp. 177–190.
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Henrich, J. Cultural group selection, coevolutionary processes and large-scale cooperation. Journal of Economic Behavior & Organization. Volume 53, Issue 1, January 2004, 3–35.
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says, for cultural group selection to occur, there must be cultural differences between groups which affect their persistence or proliferation. This means groups are selected
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Horner, V. Whiten, A. Causal knowledge and imitation/emulation switching in chimpanzees (Pan troglodytes) and children (Homo sapiens). Animal Cognition (2005) 8: 164–181
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Soltis, J., Boyd, R., Richerson, P.J., 1995. Can group-functional behaviours evolve by cultural group selection? An empirical test. Current Anthropology 36 (3), 473–494.
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Turchin, Peter; Currie, Thomas E. (2016). "Cultural Group selection is plausible but the predictions of its hypotheses should be tested with real-world data".
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Zefferman, Matthew; Waring, Timothy M.; Smaldino, Paul E.; Ross, Cody; Newson, Lesley; Naar, Nicole; Newton, Emily K.; Mathew, Sarah; Hillis, Vicken (2016).
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variation operating on a scale of millennia. It has been suggested that this is how early agriculturalist displaced hunter-gatherer societies.
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Cultural group selection gives a compelling explanation for how large-scale complex societies have formed. While altruistic behaviour such as
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Cavalli-Sforza, L.L., Menozzi, P., Piazza, A., 1994. The History and Geography of Human Genes. Princeton University Press, Princeton, NJ.
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Henrich, J., et al. Markets, Religion, Community Size, and the Evolution of Fairness and Punishment. Science 327, 1480 (2010)
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provide datasets and analytical tools for assessing the validity of competing hypotheses about human cultural evolution.
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Joseph Henrich describes the three mechanisms through which this process occurs:
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cooperative groups will be considered more prestigious and worthy of imitation.
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suggests the following three adaptations are necessary for human culture:
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New York: Penguin Group, 1-11. 313: â€“ Theory of human behavior 3449: 2705:Computer-mediated communication 1865:Extended evolutionary synthesis 1054:Gene-centered view of evolution 765: 752: 711: 687: 658: 609: 596: 587: 578: 569: 560: 551: 542: 533: 524: 514: 505: 496: 487: 478: 469: 297:Seshat: Global History Databank 281:Prestige-biased group selection 22:is an explanatory model within 2982:Empathising–systemising theory 2285:female intrasexual competition 2222:Evolutionarily stable strategy 1993:Hologenome theory of evolution 1860:History of molecular evolution 1086:Evolutionarily stable strategy 975:Last universal common ancestor 446: 437: 398: 343: 248:according to their respective 1: 3463:Behavioral and Brain Sciences 3342:Standard social science model 2395:Cognitive tradeoff hypothesis 1787:Renaissance and Enlightenment 667:Behavioral and Brain Sciences 622:Behavioral and Brain Sciences 336: 272:Direct intergroup competition 231: 209:Punishment of non-conformists 105:Human adaptations for culture 3190:Missing heritability problem 2782:Social aspects of television 2405:Evolution of nervous systems 2373:Computational theory of mind 1998:Missing heritability problem 1625:Gamete differentiation/sexes 7: 3436:Evolutionary biology portal 303: 225:Asch conformity experiments 16:Model of cultural evolution 10: 3535: 3397:Evolutionary psychologists 3270:Trivers–Willard hypothesis 3185:Human–animal communication 2897:Ovulatory shift hypothesis 2747:Imprinted brain hypothesis 2715:Human–computer interaction 1630:Life cycles/nuclear phases 1182:Trivers–Willard hypothesis 256:relative to other groups. 3476:10.1017/S0140525X1400106X 3391: 3317:Environmental determinism 3288:Cultural selection theory 3280: 3175:Evolutionary epistemology 3162: 3089:evolutionary neuroscience 3051: 3044: 2942: 2817: 2762:Rank theory of depression 2685: 2609: 2511: 2317: 2310: 2264:Parent–offspring conflict 2173: 2116: 2048: 1948: 1873: 1777: 1704: 1660: 1515: 1419: 1236: 1195: 1128:Parent–offspring conflict 1064: 933:Earliest known life forms 854: 821: 635:10.1017/S0140525X1400106X 3509:Evolutionary game theory 3210:Cultural group selection 3094:Biocultural anthropology 2787:Societal impacts of cars 2720:Media naturalness theory 2410:Fight-or-flight response 1981:Cultural group selection 1845:The eclipse of Darwinism 1817:On the Origin of Species 1792:Transmutation of species 317:Evolutionary game theory 40:evolutionary game theory 20:Cultural group selection 3410:Evolutionary psychology 3374:Sociocultural evolution 3215:Dual inheritance theory 2672:Personality development 2133:Theoretical foundations 2110:Evolutionary psychology 1986:Dual inheritance theory 1825:History of paleontology 311:Dual inheritance theory 180:Conformist transmission 3332:Social constructionism 3327:Psychological nativism 3302:Biological determinism 3250:Recent human evolution 3245:Punctuated equilibrium 3068:Behavioral epigenetics 3063:evolutionary economics 3032:Variability hypothesis 2977:Emotional intelligence 2710:Engineering psychology 2400:Evolution of the brain 1674:Punctuated equilibrium 995:Non-adaptive radiation 943:Evolutionary arms race 602:Turchin, Peter. 2007. 404:Turchin, Peter. 2018. 98:Third-party punishment 3504:Cultural anthropology 3359:Multilineal evolution 3322:Nature versus nurture 3281:Theoretical positions 3129:Functional psychology 3124:Evolutionary medicine 3099:Biological psychiatry 2807:Texting while driving 2797:Lead–crime hypothesis 2657:Cognitive development 2642:Caregiver deprivation 2153:Gene selection theory 1966:Evolutionary medicine 1840:Mendelian inheritance 1548:Biological complexity 1536:Programmed cell death 1228:Phenotypic plasticity 948:Evolutionary pressure 938:Evidence of evolution 836:Timeline of evolution 32:behavioural economics 3312:Cultural determinism 3119:Evolutionary biology 3104:Cognitive psychology 3052:Academic disciplines 2700:Cognitive ergonomics 2667:Language acquisition 2647:Childhood attachment 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2363:modularity of mind 1976:Cultural evolution 1091:Fisher's principle 1020:Handicap principle 1010:Parallel evolution 874:Adaptive radiation 760:Annual Report 2016 290:Testing the theory 238:Donald T. Campbell 139:imitative learning 132:Imitative learning 24:cultural evolution 3445: 3444: 3423:Psychology portal 3387: 3386: 3230:Hologenome theory 3200:Unit of selection 3195:Primate cognition 3109:Cognitive science 3040: 3039: 2911:Sexual attraction 2887:Mating strategies 2652:Cinderella effect 2582:Moral foundations 2486:Visual perception 2378:Domain generality 2347:Facial expression 2295:Sexual dimorphism 2254:Natural selection 2200:Hamiltonian spite 2076: 2075: 1692:Uniformitarianism 1645:Sex-determination 1150:Sexual dimorphism 1145:Natural selection 1049:Unit of selection 1015:Signalling theory 169:in-group biases. 114:Michael Tomasello 3526: 3489: 3488: 3478: 3453: 3432: 3419: 3406: 3405: 3049: 3048: 3045:Related subjects 2832:Adult attachment 2359:Cognitive module 2315: 2314: 2302:Social selection 2276:Costly signaling 2271:Sexual selection 2158:Modern synthesis 2103: 2096: 2089: 2080: 2079: 2066: 2056: 2055: 1855:Modern synthesis 1615:Multicellularity 1610:Mosaic evolution 1495:auditory ossicle 1177:Social selection 1160:Flowering plants 1155:Sexual selection 808: 801: 794: 785: 784: 776: 769: 763: 756: 750: 749: 739: 715: 709: 708: 706: 705: 691: 685: 684: 682: 662: 656: 655: 637: 613: 607: 600: 594: 591: 585: 582: 576: 573: 567: 564: 558: 555: 549: 546: 540: 537: 531: 528: 522: 518: 512: 509: 503: 500: 494: 491: 485: 482: 476: 473: 467: 464: 453: 450: 444: 441: 435: 432: 423: 420: 409: 402: 396: 393: 374: 371: 362: 359: 350: 347: 3534: 3533: 3529: 3528: 3527: 3525: 3524: 3523: 3494: 3493: 3492: 3454: 3450: 3446: 3441: 3383: 3369:Neoevolutionism 3276: 3260:Species complex 3225:Group selection 3163:Research topics 3158: 3134:Neuropsychology 3036: 3022:Substance abuse 2944:Sex differences 2938: 2852:Coolidge effect 2813: 2725:Neuroergonomics 2690: 2681: 2605: 2507: 2441:Folk psychology 2322: 2306: 2176: 2169: 2112: 2107: 2077: 2072: 2044: 1971:Group selection 1944: 1869: 1773: 1700: 1662:Tempo and modes 1656: 1511: 1415: 1232: 1191: 1067: 1060: 1037:Species complex 850: 841:History of life 817: 812: 780: 779: 775:85 (2): 312-47. 770: 766: 757: 753: 716: 712: 703: 701: 693: 692: 688: 663: 659: 614: 610: 601: 597: 592: 588: 583: 579: 574: 570: 565: 561: 556: 552: 547: 543: 538: 534: 529: 525: 519: 515: 510: 506: 501: 497: 492: 488: 483: 479: 474: 470: 465: 456: 451: 447: 442: 438: 433: 426: 421: 412: 403: 399: 394: 377: 372: 365: 360: 353: 348: 344: 339: 334: 323:Group selection 306: 292: 283: 274: 265: 234: 220: 211: 199:self-similarity 191: 182: 161: 152: 143:Andrew Meltzoff 134: 126:joint attention 122: 120:Joint attention 107: 85:social learning 17: 12: 11: 5: 3532: 3522: 3521: 3516: 3511: 3506: 3491: 3490: 3447: 3443: 3442: 3440: 3439: 3426: 3413: 3400: 3392: 3389: 3388: 3385: 3384: 3382: 3381: 3376: 3371: 3366: 3361: 3356: 3351: 3346: 3345: 3344: 3339: 3334: 3329: 3324: 3319: 3314: 3309: 3304: 3290: 3284: 3282: 3278: 3277: 3275: 3274: 3273: 3272: 3267: 3262: 3257: 3252: 3247: 3242: 3237: 3232: 3227: 3222: 3217: 3212: 3207: 3197: 3192: 3187: 3182: 3177: 3172: 3166: 3164: 3160: 3159: 3157: 3156: 3151: 3146: 3141: 3136: 3131: 3126: 3121: 3116: 3111: 3106: 3101: 3096: 3091: 3074: 3065: 3055: 3053: 3046: 3042: 3041: 3038: 3037: 3035: 3034: 3029: 3024: 3019: 3014: 3009: 3004: 2999: 2994: 2989: 2984: 2979: 2974: 2969: 2964: 2959: 2954: 2948: 2946: 2940: 2939: 2937: 2936: 2931: 2926: 2913: 2904: 2899: 2894: 2889: 2884: 2879: 2874: 2869: 2864: 2859: 2854: 2849: 2844: 2839: 2834: 2829: 2823: 2821: 2815: 2814: 2812: 2811: 2810: 2809: 2804: 2799: 2794: 2784: 2779: 2774: 2769: 2764: 2759: 2754: 2752:Mind-blindness 2749: 2744: 2739: 2734: 2729: 2728: 2727: 2722: 2717: 2712: 2707: 2696: 2694: 2683: 2682: 2680: 2679: 2674: 2669: 2664: 2659: 2654: 2649: 2644: 2639: 2626: 2621: 2615: 2613: 2607: 2606: 2604: 2603: 2598: 2597: 2596: 2586: 2585: 2584: 2574: 2573: 2572: 2567: 2562: 2552: 2547: 2546: 2545: 2535: 2534: 2533: 2528: 2517: 2515: 2509: 2508: 2506: 2505: 2504: 2503: 2498: 2493: 2483: 2478: 2473: 2464: 2463: 2462: 2457: 2447: 2445:theory of mind 2438: 2429: 2428: 2427: 2422: 2417: 2407: 2402: 2397: 2392: 2391: 2390: 2385: 2380: 2375: 2370: 2356: 2351: 2350: 2349: 2344: 2339: 2328: 2326: 2312: 2308: 2307: 2305: 2304: 2299: 2298: 2297: 2292: 2287: 2278: 2268: 2267: 2266: 2256: 2251: 2246: 2241: 2240: 2239: 2229: 2224: 2219: 2214: 2212:Baldwin effect 2209: 2208: 2207: 2202: 2197: 2187: 2181: 2179: 2171: 2170: 2168: 2167: 2162: 2161: 2160: 2155: 2150: 2145: 2140: 2130: 2129: 2128: 2117: 2114: 2113: 2106: 2105: 2098: 2091: 2083: 2074: 2073: 2071: 2070: 2060: 2049: 2046: 2045: 2043: 2042: 2037: 2032: 2027: 2022: 2021: 2020: 2010: 2005: 2000: 1995: 1990: 1989: 1988: 1983: 1978: 1968: 1963: 1958: 1952: 1950: 1946: 1945: 1943: 1942: 1937: 1936: 1935: 1930: 1925: 1924: 1923: 1913: 1908: 1903: 1898: 1893: 1883: 1877: 1875: 1871: 1870: 1868: 1867: 1862: 1857: 1852: 1847: 1842: 1837: 1832: 1827: 1822: 1821: 1820: 1811:Charles Darwin 1808: 1807: 1806: 1794: 1789: 1783: 1781: 1775: 1774: 1772: 1771: 1766: 1761: 1756: 1751: 1749:Non-ecological 1746: 1741: 1736: 1731: 1726: 1721: 1716: 1710: 1708: 1702: 1701: 1699: 1698: 1689: 1680: 1666: 1664: 1658: 1657: 1655: 1654: 1649: 1648: 1647: 1642: 1637: 1632: 1627: 1617: 1612: 1607: 1602: 1597: 1592: 1587: 1582: 1577: 1572: 1567: 1566: 1565: 1555: 1550: 1545: 1540: 1539: 1538: 1533: 1522: 1520: 1513: 1512: 1510: 1509: 1508: 1507: 1502: 1500:nervous system 1497: 1492: 1487: 1479: 1478: 1477: 1472: 1467: 1462: 1457: 1452: 1442: 1437: 1432: 1426: 1424: 1417: 1416: 1414: 1413: 1408: 1403: 1398: 1393: 1392: 1391: 1381: 1380: 1379: 1374: 1373: 1372: 1367: 1357: 1352: 1347: 1342: 1337: 1336: 1335: 1330: 1320: 1310: 1305: 1304: 1303: 1293: 1288: 1283: 1278: 1277: 1276: 1266: 1261: 1260: 1259: 1249: 1243: 1241: 1234: 1233: 1231: 1230: 1225: 1220: 1215: 1210: 1205: 1199: 1197: 1193: 1192: 1190: 1189: 1184: 1179: 1174: 1173: 1172: 1167: 1162: 1152: 1147: 1142: 1137: 1132: 1131: 1130: 1125: 1115: 1110: 1105: 1104: 1103: 1093: 1088: 1083: 1078: 1072: 1070: 1062: 1061: 1059: 1058: 1057: 1056: 1046: 1041: 1040: 1039: 1034: 1024: 1023: 1022: 1012: 1007: 1002: 1000:Origin of life 997: 992: 987: 985:Microevolution 982: 980:Macroevolution 977: 972: 967: 966: 965: 955: 950: 945: 940: 935: 930: 925: 920: 918:Common descent 915: 914: 913: 903: 898: 896:Baldwin effect 893: 892: 891: 886: 876: 871: 866: 860: 858: 852: 851: 849: 848: 843: 838: 833: 828: 822: 819: 818: 811: 810: 803: 796: 788: 778: 777: 764: 751: 710: 686: 657: 608: 595: 586: 577: 568: 559: 550: 541: 532: 523: 513: 504: 495: 486: 477: 468: 454: 445: 436: 424: 410: 397: 375: 363: 351: 341: 340: 338: 335: 333: 332: 326: 320: 314: 307: 305: 302: 291: 288: 282: 279: 273: 270: 264: 261: 233: 230: 219: 216: 210: 207: 190: 187: 181: 178: 174:Joseph Henrich 160: 157: 151: 148: 133: 130: 121: 118: 106: 103: 56:game theoretic 15: 9: 6: 4: 3: 2: 3531: 3520: 3517: 3515: 3512: 3510: 3507: 3505: 3502: 3501: 3499: 3486: 3482: 3477: 3472: 3468: 3464: 3460: 3452: 3448: 3438: 3437: 3431: 3427: 3425: 3424: 3418: 3414: 3412: 3411: 3401: 3399: 3398: 3394: 3393: 3390: 3380: 3377: 3375: 3372: 3370: 3367: 3365: 3364:Neo-Darwinism 3362: 3360: 3357: 3355: 3352: 3350: 3349:Functionalism 3347: 3343: 3340: 3338: 3335: 3333: 3330: 3328: 3325: 3323: 3320: 3318: 3315: 3313: 3310: 3308: 3307:Connectionism 3305: 3303: 3300: 3299: 3298: 3297:indeterminism 3294: 3291: 3289: 3286: 3285: 3283: 3279: 3271: 3268: 3266: 3263: 3261: 3258: 3256: 3253: 3251: 3248: 3246: 3243: 3241: 3238: 3236: 3233: 3231: 3228: 3226: 3223: 3221: 3218: 3216: 3213: 3211: 3208: 3206: 3203: 3202: 3201: 3198: 3196: 3193: 3191: 3188: 3186: 3183: 3181: 3178: 3176: 3173: 3171: 3168: 3167: 3165: 3161: 3155: 3152: 3150: 3147: 3145: 3142: 3140: 3137: 3135: 3132: 3130: 3127: 3125: 3122: 3120: 3117: 3115: 3112: 3110: 3107: 3105: 3102: 3100: 3097: 3095: 3092: 3090: 3086: 3082: 3078: 3075: 3073: 3069: 3066: 3064: 3060: 3057: 3056: 3054: 3050: 3047: 3043: 3033: 3030: 3028: 3025: 3023: 3020: 3018: 3017:Schizophrenia 3015: 3013: 3010: 3008: 3005: 3003: 3002:Mental health 3000: 2998: 2995: 2993: 2990: 2988: 2985: 2983: 2980: 2978: 2975: 2973: 2970: 2968: 2965: 2963: 2960: 2958: 2955: 2953: 2950: 2949: 2947: 2945: 2941: 2935: 2932: 2930: 2927: 2925: 2921: 2917: 2914: 2912: 2908: 2905: 2903: 2900: 2898: 2895: 2893: 2890: 2888: 2885: 2883: 2880: 2878: 2877:Mate guarding 2875: 2873: 2870: 2868: 2865: 2863: 2860: 2858: 2855: 2853: 2850: 2848: 2845: 2843: 2840: 2838: 2837:Age disparity 2835: 2833: 2830: 2828: 2825: 2824: 2822: 2820: 2816: 2808: 2805: 2803: 2800: 2798: 2795: 2793: 2790: 2789: 2788: 2785: 2783: 2780: 2778: 2775: 2773: 2770: 2768: 2767:Schizophrenia 2765: 2763: 2760: 2758: 2755: 2753: 2750: 2748: 2745: 2743: 2740: 2738: 2735: 2733: 2730: 2726: 2723: 2721: 2718: 2716: 2713: 2711: 2708: 2706: 2703: 2702: 2701: 2698: 2697: 2695: 2693: 2692:Mental health 2688: 2687:Human factors 2684: 2678: 2677:Socialization 2675: 2673: 2670: 2668: 2665: 2663: 2660: 2658: 2655: 2653: 2650: 2648: 2645: 2643: 2640: 2638: 2637:paternal bond 2634: 2630: 2627: 2625: 2622: 2620: 2617: 2616: 2614: 2612: 2608: 2602: 2599: 2595: 2592: 2591: 2590: 2587: 2583: 2580: 2579: 2578: 2575: 2571: 2568: 2566: 2563: 2561: 2558: 2557: 2556: 2553: 2551: 2548: 2544: 2541: 2540: 2539: 2536: 2532: 2529: 2527: 2524: 2523: 2522: 2519: 2518: 2516: 2514: 2510: 2502: 2501:NaĂŻve physics 2499: 2497: 2494: 2492: 2489: 2488: 2487: 2484: 2482: 2479: 2477: 2474: 2472: 2468: 2467:Motor control 2465: 2461: 2458: 2456: 2453: 2452: 2451: 2448: 2446: 2442: 2439: 2437: 2433: 2430: 2426: 2425:Ophidiophobia 2423: 2421: 2418: 2416: 2415:Arachnophobia 2413: 2412: 2411: 2408: 2406: 2403: 2401: 2398: 2396: 2393: 2389: 2386: 2384: 2381: 2379: 2376: 2374: 2371: 2369: 2366: 2365: 2364: 2360: 2357: 2355: 2352: 2348: 2345: 2343: 2342:Display rules 2340: 2338: 2335: 2334: 2333: 2330: 2329: 2327: 2325: 2320: 2316: 2313: 2309: 2303: 2300: 2296: 2293: 2291: 2288: 2286: 2282: 2279: 2277: 2274: 2273: 2272: 2269: 2265: 2262: 2261: 2260: 2257: 2255: 2252: 2250: 2247: 2245: 2244:Kin selection 2242: 2238: 2235: 2234: 2233: 2230: 2228: 2225: 2223: 2220: 2218: 2215: 2213: 2210: 2206: 2203: 2201: 2198: 2196: 2193: 2192: 2191: 2188: 2186: 2183: 2182: 2180: 2178: 2172: 2166: 2163: 2159: 2156: 2154: 2151: 2149: 2146: 2144: 2141: 2139: 2138:Adaptationism 2136: 2135: 2134: 2131: 2127: 2124: 2123: 2122: 2119: 2118: 2115: 2111: 2104: 2099: 2097: 2092: 2090: 2085: 2084: 2081: 2069: 2065: 2061: 2059: 2051: 2050: 2047: 2041: 2038: 2036: 2033: 2031: 2028: 2026: 2023: 2019: 2016: 2015: 2014: 2013:Phylogenetics 2011: 2009: 2006: 2004: 2001: 1999: 1996: 1994: 1991: 1987: 1984: 1982: 1979: 1977: 1974: 1973: 1972: 1969: 1967: 1964: 1962: 1959: 1957: 1954: 1953: 1951: 1947: 1941: 1938: 1934: 1931: 1929: 1926: 1922: 1919: 1918: 1917: 1916:Structuralism 1914: 1912: 1909: 1907: 1904: 1902: 1899: 1897: 1894: 1892: 1891:Catastrophism 1889: 1888: 1887: 1884: 1882: 1879: 1878: 1876: 1872: 1866: 1863: 1861: 1858: 1856: 1853: 1851: 1850:Neo-Darwinism 1848: 1846: 1843: 1841: 1838: 1836: 1833: 1831: 1828: 1826: 1823: 1819: 1818: 1814: 1813: 1812: 1809: 1805: 1804: 1800: 1799: 1798: 1795: 1793: 1790: 1788: 1785: 1784: 1782: 1780: 1776: 1770: 1767: 1765: 1764:Reinforcement 1762: 1760: 1757: 1755: 1752: 1750: 1747: 1745: 1742: 1740: 1737: 1735: 1732: 1730: 1727: 1725: 1722: 1720: 1717: 1715: 1712: 1711: 1709: 1707: 1703: 1697: 1696:Catastrophism 1693: 1690: 1688: 1687:Macromutation 1684: 1683:Micromutation 1681: 1679: 1675: 1671: 1668: 1667: 1665: 1663: 1659: 1653: 1650: 1646: 1643: 1641: 1638: 1636: 1633: 1631: 1628: 1626: 1623: 1622: 1621: 1618: 1616: 1613: 1611: 1608: 1606: 1603: 1601: 1598: 1596: 1593: 1591: 1590:Immune system 1588: 1586: 1583: 1581: 1578: 1576: 1573: 1571: 1568: 1564: 1561: 1560: 1559: 1556: 1554: 1551: 1549: 1546: 1544: 1541: 1537: 1534: 1532: 1529: 1528: 1527: 1524: 1523: 1521: 1519: 1514: 1506: 1503: 1501: 1498: 1496: 1493: 1491: 1488: 1486: 1483: 1482: 1480: 1476: 1473: 1471: 1468: 1466: 1463: 1461: 1458: 1456: 1453: 1451: 1450:symbiogenesis 1448: 1447: 1446: 1443: 1441: 1438: 1436: 1433: 1431: 1428: 1427: 1425: 1423: 1418: 1412: 1409: 1407: 1404: 1402: 1399: 1397: 1394: 1390: 1387: 1386: 1385: 1382: 1378: 1375: 1371: 1368: 1366: 1363: 1362: 1361: 1358: 1356: 1353: 1351: 1348: 1346: 1343: 1341: 1338: 1334: 1331: 1329: 1326: 1325: 1324: 1321: 1319: 1316: 1315: 1314: 1311: 1309: 1306: 1302: 1299: 1298: 1297: 1294: 1292: 1289: 1287: 1284: 1282: 1279: 1275: 1272: 1271: 1270: 1267: 1265: 1262: 1258: 1255: 1254: 1253: 1250: 1248: 1245: 1244: 1242: 1240: 1235: 1229: 1226: 1224: 1221: 1219: 1216: 1214: 1211: 1209: 1206: 1204: 1201: 1200: 1198: 1194: 1188: 1185: 1183: 1180: 1178: 1175: 1171: 1168: 1166: 1163: 1161: 1158: 1157: 1156: 1153: 1151: 1148: 1146: 1143: 1141: 1138: 1136: 1133: 1129: 1126: 1124: 1121: 1120: 1119: 1118:Kin selection 1116: 1114: 1113:Genetic drift 1111: 1109: 1106: 1102: 1099: 1098: 1097: 1094: 1092: 1089: 1087: 1084: 1082: 1079: 1077: 1074: 1073: 1071: 1069: 1063: 1055: 1052: 1051: 1050: 1047: 1045: 1042: 1038: 1035: 1033: 1030: 1029: 1028: 1025: 1021: 1018: 1017: 1016: 1013: 1011: 1008: 1006: 1003: 1001: 998: 996: 993: 991: 988: 986: 983: 981: 978: 976: 973: 971: 968: 964: 961: 960: 959: 956: 954: 951: 949: 946: 944: 941: 939: 936: 934: 931: 929: 926: 924: 921: 919: 916: 912: 909: 908: 907: 904: 902: 899: 897: 894: 890: 887: 885: 882: 881: 880: 877: 875: 872: 870: 867: 865: 862: 861: 859: 857: 853: 847: 844: 842: 839: 837: 834: 832: 829: 827: 824: 823: 820: 816: 809: 804: 802: 797: 795: 790: 789: 786: 782: 774: 768: 761: 755: 747: 743: 738: 733: 729: 725: 721: 714: 700: 696: 690: 681: 676: 672: 668: 661: 653: 649: 645: 641: 636: 631: 627: 623: 619: 612: 605: 599: 590: 581: 572: 563: 554: 545: 536: 527: 517: 508: 499: 490: 481: 472: 463: 461: 459: 449: 440: 431: 429: 419: 417: 415: 407: 401: 392: 390: 388: 386: 384: 382: 380: 370: 368: 358: 356: 346: 342: 330: 327: 324: 321: 318: 315: 312: 309: 308: 301: 298: 287: 278: 269: 260: 257: 255: 251: 247: 243: 239: 229: 226: 215: 206: 203: 200: 195: 186: 177: 175: 172:According to 170: 167: 156: 147: 144: 140: 129: 127: 117: 115: 111: 102: 99: 95: 91: 86: 81: 77: 76:kin selection 72: 70: 69:in-group bias 66: 62: 57: 51: 49: 45: 41: 37: 33: 29: 25: 21: 3466: 3462: 3451: 3434: 3421: 3408: 3395: 3209: 3154:Sociobiology 3012:Neuroscience 2992:Intelligence 2538:Anthropology 2491:Color vision 2476:Multitasking 2455:Flynn effect 2450:Intelligence 2432:Folk biology 2175:Evolutionary 2025:Polymorphism 2008:Astrobiology 1980: 1956:Biogeography 1911:Saltationism 1901:Orthogenesis 1886:Alternatives 1815: 1801: 1734:Cospeciation 1729:Cladogenesis 1678:Saltationism 1635:Mating types 1558:Color vision 1543:Avian flight 1465:mitochondria 1203:Canalisation 1081:Biodiversity 826:Introduction 781: 772: 767: 759: 754: 727: 723: 713: 702:. Retrieved 698: 689: 670: 666: 660: 625: 621: 611: 603: 598: 589: 580: 571: 562: 553: 544: 535: 526: 516: 507: 498: 489: 480: 471: 448: 439: 405: 400: 345: 293: 284: 275: 266: 258: 253: 249: 245: 241: 235: 221: 212: 204: 196: 192: 183: 171: 165: 162: 153: 135: 123: 112: 108: 101:psychology. 73: 52: 28:anthropology 19: 18: 3469:(39): e30. 3293:Determinism 3205:Coevolution 3149:Primatology 2987:Gender role 2892:Orientation 2772:Screen time 2629:Affectional 2611:Development 2290:Mate choice 2217:By-products 2185:Adaptations 2148:Cognitivism 2035:Systematics 1906:Mutationism 1724:Catagenesis 1652:Snake venom 1585:Eusociality 1563:in primates 1553:Cooperation 1481:In animals 1301:butterflies 1274:Cephalopods 1264:Brachiopods 1196:Development 1170:Mate choice 923:Convergence 906:Coevolution 864:Abiogenesis 730:(1): 1–14. 680:10871/24291 223:worth. The 197:Meanwhile, 80:reciprocity 3498:Categories 3240:Population 3235:Lamarckism 3081:behavioral 3059:Behavioral 3007:Narcissism 2952:Aggression 2742:Hypophobia 2732:Depression 2619:Attachment 2601:Universals 2565:Psychology 2543:Biological 2531:Musicology 2521:Aesthetics 2420:Basophobia 2227:Exaptation 2205:Reciprocal 1896:Lamarckism 1874:Philosophy 1797:David Hume 1759:Peripatric 1754:Parapatric 1739:Ecological 1719:Anagenesis 1714:Allopatric 1706:Speciation 1670:Gradualism 1595:Metabolism 1455:chromosome 1445:Eukaryotes 1223:Modularity 1140:Population 1066:Population 1027:Speciation 1005:Panspermia 958:Extinction 953:Exaptation 928:Divergence 901:Cladistics 889:Reciprocal 869:Adaptation 704:2019-01-03 521:1068–1095. 337:References 232:Mechanisms 65:conformity 48:psychology 3514:Selection 3085:cognitive 3077:Affective 2962:Cognition 2916:Sexuality 2902:Pair bond 2662:Education 2319:Cognition 2237:Inclusive 2177:processes 2165:Criticism 2030:Protocell 1881:Darwinism 1769:Sympatric 1518:processes 1406:Tetrapods 1355:Kangaroos 1281:Dinosaurs 1218:Inversion 1187:Variation 1108:Gene flow 1101:Inclusive 911:Mutualism 856:Evolution 746:2169-0812 673:: 43–44. 644:1469-1825 90:Ultimatum 61:imitation 44:sociology 3519:Memetics 3485:25347943 3354:Memetics 3114:Ethology 3072:genetics 2907:Physical 2872:Jealousy 2827:Activity 2633:maternal 2589:Religion 2577:Morality 2555:Language 2436:taxonomy 2249:Mismatch 2195:Cheating 2190:Altruism 2058:Category 1933:Vitalism 1928:Theistic 1921:Spandrel 1605:Morality 1600:Monogamy 1475:plastids 1440:Flagella 1396:Reptiles 1377:sea cows 1360:primates 1269:Molluscs 1247:Bacteria 1135:Mutation 1068:genetics 1044:Taxonomy 990:Mismatch 970:Homology 884:Cheating 879:Altruism 652:25347943 329:Memetics 304:See also 94:Dictator 3255:Species 3027:Suicide 2862:Fantasy 2842:Arousal 2624:Bonding 2513:Culture 2337:Display 2324:Emotion 2232:Fitness 2121:History 1949:Related 1779:History 1640:Meiosis 1575:Empathy 1570:Emotion 1470:nucleus 1411:Viruses 1401:Spiders 1313:Mammals 1296:Insects 1096:Fitness 1032:Species 831:Outline 628:: e30. 246:against 3483:  3433:  3420:  3407:  2997:Memory 2957:Autism 2924:female 2857:Desire 2594:Origin 2570:Speech 2560:Origin 2332:Affect 2068:Portal 1744:Hybrid 1580:Ethics 1422:organs 1384:Plants 1370:lemurs 1365:humans 1350:horses 1340:hyenas 1328:wolves 1323:canids 1257:origin 744:  650:  642:  254:losses 96:, and 67:, and 46:, and 2967:Crime 2550:Crime 2481:Sleep 2471:skill 2311:Areas 1531:Death 1526:Aging 1505:brain 1291:Fungi 1252:Birds 1165:Fungi 963:Event 846:Index 250:gains 3481:PMID 2920:male 2281:Male 2018:Tree 1490:hair 1430:Cell 1333:dogs 1318:cats 1308:Life 1286:Fish 1239:taxa 742:ISSN 648:PMID 640:ISSN 78:and 3471:doi 2819:Sex 2496:Eye 1516:Of 1485:eye 1435:DNA 1420:Of 1237:Of 732:doi 675:hdl 671:e55 630:doi 252:or 244:or 242:for 236:As 166:and 3500:: 3479:. 3467:39 3465:. 3461:. 740:. 726:. 722:. 697:. 669:. 646:. 638:. 626:39 624:. 620:. 457:^ 427:^ 413:^ 378:^ 366:^ 354:^ 92:, 63:, 50:. 42:, 38:, 34:, 30:, 3487:. 3473:: 3295:/ 3087:/ 3083:/ 3079:/ 3070:/ 3061:/ 2922:/ 2918:/ 2909:/ 2689:/ 2635:/ 2631:/ 2469:/ 2443:/ 2434:/ 2361:/ 2321:/ 2283:/ 2102:e 2095:t 2088:v 1694:/ 1685:/ 1676:/ 1672:/ 807:e 800:t 793:v 748:. 734:: 728:1 707:. 683:. 677:: 654:. 632::

Index

cultural evolution
anthropology
behavioural economics
evolutionary biology
evolutionary game theory
sociology
psychology
game theoretic
imitation
conformity
in-group bias
kin selection
reciprocity
social learning
Ultimatum
Dictator
Third-party punishment
Michael Tomasello
joint attention
imitative learning
Andrew Meltzoff
Joseph Henrich
self-similarity
Asch conformity experiments
Donald T. Campbell
Seshat: Global History Databank
Dual inheritance theory
Evolutionary game theory
Group selection
Memetics

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