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desert, and coast to caves, mammalian burrows, and vegetation. They live in accordance with where their prey lives. One division of the
Geobiote feeds on maggots and eggs that are found in forest vegetation or in carrion. Naturally, these are found in heavily forested areas. The second and third divisions hunt for arthropods that feed on dead plant matter. Thus, these types of Geobiotes are found in the sand and burrowed in soil. The fourth type of Geobiotes feed on fly eggs that grow on fresh
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fly breeding resources, and they can be distinguished from fly larvae by the following: Beetle larvae possess a hard, head capsule, often brown in colour. Fly larvae lack a head capsule, instead having distinct, internal, black mouth hooks (cephalopharyngeal skeleton of mouthparts) at the anterior end of their body." Understanding how long ago the eggs were laid and the time period of the developmental stages is important for determining the time of death.
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910:. If the Histeridae beetles are present, the investigator can assume that some of the other insects have been eaten by the hister beetles. Due to their importance in forensic entomology, research is continually being conducted on this family's life cycle and development, their prevalence in specific locations, and their geographical distribution.
546:, dead vegetation, sandy areas, under tree bark, mammal burrows, and ant/termite colonies. Charactertistics of the hister beetle are dependent upon its habitat. For example, the flat hister beetles are found under bark, while the cylindrical beetles are not. The shape of the beetle will vary from species to species.
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cement. This makes their outer shell harder and protects them during this vulnerable stage. While they pupate, they breathe through spiracles on the abdomen. The beetle is non-feeding and immobile in this stage, as their internal structure is breaking down and rebuilding to its adult form. Under good
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The hister beetles will hide under the dead body in the soil during the day and come out at night to feed. For this reason, the dead body must be examined at different times of the day. After collecting the hister beetles from a body, isolate them because they are predacious beetles and eat anything
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are destructive and when they hatch they eat the maggots in the dead body. The stage of the beetle and other insects in the body at the time of investigation helps to determine the time of death. "Beetle larvae are ... helpful when determining post-mortem time interval. Beetle larvae often reside in
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The family
Histeridae is very diverse, so different species can be found on the body at different times because of their various feeding habits. This needs to be noted when examining a dead body. They are predacious beetles and arrive when there is material to feed on, like other beetles or maggots.
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Because members of the family
Histeridae can be found on carrion, they have proven to be important in certain forensic investigations. The predacious hister beetles will feed on the various insects on the body, primarily Diptera. To estimate a person's time of death, forensic investigators must look
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This subclade is the most diverse and the largest of the hister predator subclasses. The
Geobites' body structures are generally circular, and they are known for their digging tendencies. This subclass is separated into five more divisions. The members of this subclass live anywhere from the soil,
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This division consists of those
Histeridae that live in close proximity with social colonies of arthropods such as ants and termites. Histeridae that live near ants can live in a harmonious or hostile relationship. The hostile hister beetles feed on the ants. The harmonious hister beetles eat the
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This subclade is the smallest of the four. They live on plant litter and feed on the tiny arthropods found there. Microhisterids, like all other
Histeridae, become specialized to hunt their prey and live in their habitats. Like the fifth division of Geobiotes, some Microhisterids are known to be
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The larval stage of the beetle typically goes through two instars, and the second instar is the longest stage of its entire development, taking up 39% of overall development time. It takes 5.1±0.1 days on average for the first instar to develop into the second. The larval form of the insect will
564:. As new food sources are introduced into an environment, the predators of that food source will soon follow. Histeridae live in areas where their prey feed. Some examples are mammalian nests where other arthropods scavenge for food or carrion where maggots will arrive.
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Because the
Histeridae are predacious, they can be utilized as control agents, but must be stored in isolation. The hister beetles have proven useful in both the control of pest flies in poultry houses and pastures, and against pest beetles of stored food products.
925:. Forty species of Coleoptera, including the hister beetles, were observed on twelve pig carcasses over a one-year period. The beetles’ distribution and their time of colonization in the various decomposition stages over the year were examined and recorded.
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Histeridae is a large and diverse family. There are more than 410 genera and 4,800 described species in
Histeridae worldwide, with more than 500 species in North America. The beetles range in size, shape and color. Many of the adults are predacious.
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Some
Histeridae species live in an integrated nest with ants and termites. Some species have been found to be fed by the ants, while others simply feed on leftover insect larvae that the ants do not want. In contrast, in their adult stage
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Hister beetles are found throughout the world in various habitats. Histeridae have been located in North
America, Central America, Europe, Asia, and Australia, but each hister beetle occupies certain niches. The beetles live in dung,
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479:. Histeridae has two common names, the “clown” beetle and the “hister” beetle. There have been several theories explaining the origin of these common names. One theory for the “hister” nickname comes from the work of
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The larvae and adult forms of Histeridae have been known to feed on dung, carrion, decomposing vegetation, other insects, larvae, and pupae. Hister beetles are able to locate both dung and carrion through
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Dictionary of word roots and combining forms compiled from the Greek, Latin, and other languages, with special reference to biological terms and scientific names. Mountain View, Calif: Mayfield Pub. Co.,
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The average time of development from egg to adult at 30 °C is 20.5 days. The eggs of most species are off-white and oval in shape. The egg takes on average 3.8±0.02 days to hatch into the first
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eggs, are found near tree bark. The latter also feed on insects and prefer to live in forested areas. Species of the cylindrical Dendrobites usually hunt prey that is unique for that species.
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and their mandibular mouth parts can cause excess damage to the body. Insects can also help determine the season the body died in. The hister beetle is more prevalent in spring and summer.
483:, a Roman poet. Juvenal used the word “hister” to mean a dirty, lowly being. Another theory for the origin of this beetle's name stems from the fact that in Latin, “hister” means actor.
613:. When found on dung, carrion and vegetation, the hister beetles will feed on the fly larvae found there. The predacious hister beetle will feed on soft-bodied insect eggs and larvae,
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They have two common body shapes. One is flattened while the other has a more cylindrical appearance. The former usually lives near tree barks. This is because the prey they feed on,
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same food as the ants, however they may not be in direct competition for the food. These beetles have an excretory organ that produces an odor telling the ants they mean no harm.
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are located on the abdomen and are the tracheal system of the beetle. Oxygen is brought into the body by spiracles and tiny sacs exchange the oxygen like a mammal's lung.
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and forewings are normally well developed in beetles, but in Histeridae the elytra are shortened and rectangular. The shortened elytra expose the final two of the seven
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Arnett, R. H. 1968. Histeridae. In: The Beetles of the United States (A manual for identification). The American Entomological Institute, Ann Arbor, xii + 1112 pp.
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takes place, the male leaves enough sperm in the female to fertilize all the eggs in the female's ovaries. The excess sperm is kept in a special structure called
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Davies, A. "Family Histeridae." Checklist of beetles of Canada and Alaska. Canadian National Collection of Insects, Arachnids and Nematodes. 11 Mar. 2009 <
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is the study of insects related to crime scenes. Insects can be very helpful at a crime scene and give people an idea of what happened and when it happened.
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The pupal form of the beetle is similar in appearance to the adult form. They have outer cells produced in the larval stages that are reinforced with
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Myers, P., R. Espinosa, and C. S. Parr. "Family Histeridae." Animal Diversity Web. 1998. University of Michigan Museum of Zoology. 22 Mar. 2009 <
873:. This division of Geobiotes is found near animal homes such as nests and burrows. The last division of Geobiotes lives in caves. They feed on the
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Stephens, Stephanie A. "An Overview of the Coleopteran Family Histeridae and its Significance to Forensic Entomology." 2003. 13 Mar. 2009 <
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1669:(Coleoptera: Histeridae)." Journal of Entomological Science. CABI. Vet. Toxicology & Entomology Res. Lab., College station. 20 Mar. 2009.
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Brundage, Adrienne. "Importance of Beetles Part 2." Ento 431 Class Period. Texas A&M University: Kleberg, College Station. 11 Mar. 2009.
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and other arthropods that occupy the vegetation and fungi found there. This type of Geobiote is known to include species that become blind.
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which is typically shiny black or metallic green. The two main shapes for this family are oval and flat. The elytra is shorter than the
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pests that infest dung or to control house flies. Histeridae are most active at night and they will play dead if they feel threatened.
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Brundage, Adrienne. "Development and Orders." Ento 431 Class Period. Texas A&M University: Kleberg, College Station. 11 Mar. 2009.
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feeders are most active at night and will fake death if they feel threatened. This family of beetles will occupy almost any kind of
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Zhou, Yu-Lingzi; Ślipiński, Adam; Ren, Dong; Parker, Joseph (2019-04-16). Long, John A; Tautz, Diethard; Thayer, Margaret (eds.).
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The hister beetles lay their eggs in dead bodies and over a short period of time they develop into various stages. The Histeridae
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https://web.archive.org/web/20110714075700/http://resources.metapress.com/pdf-preview.axd?code=362v3k1361r2x385&size=largest
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Byrd, Dr. J. H. "Definition and Information." Forensic Entomology: Insects in Legal Investigations. 1998. 21 Mar. 2009 <
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Dillon, Elizabeth S., and Lawrence R. Dillon. A Manual of Common Beetles of Eastern North America. Peterson and Co., 1961.
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Fuller, John. "What do bugs have to do with forensic science?" How Stuff Works? University of Georgia. 21 Mar. 2009 <
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arrive at the dead body within a few hours. Certain species of the hister beetles follow shortly behind and prey on the
1477:Ă–zdemir, Senem & Sert, Osman (2009). "Determination of Coleoptera fauna on carcasses in Ankara province, Turkey".
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557:(termitophiles). The hister beetles can live in harmony with the ants or prey on the ants, depending on the species.
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range in length from three millimeters to several centimeters. They have a membranous body with a limited amount of
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594:, around 99 million years old, including those belonging to the extant subfamily Haeteriinae, and the living genus
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1266:"Beetle - MSN Encarta." MSN Encarta : Online Encyclopedia, Dictionary, Atlas, and Homework. 07 Apr. 2009 <
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1651:"Hister beetles - Family Histeridae." Decomposition: Hister Beetles. 2003. Australian Museum. 11 Mar. 2009 <
1445:"Hister beetles - Family Histeridae." Decomposition: Hister Beetles. 2003. Australian Museum. 11 Mar. 2009 <
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Gillott, C. 1995. The Remaining Endopterygote Orders. In: Entomology. Plenum Press, New York, xviii + 798 pp.
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arrive at the body first, when it is fresh to late (beginning of decomposition to advanced decomposition).
662:. The larval form does not resemble the adult form and the pupa, non-feeding and has internally developing
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525:, also known as a body cavity. They have a tube shaped heart that spans the length of the body, and use
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Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011).
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Dadour, Dr. Ian, and Dr. David Cook. "Collecting From a Corpse." Web Ring. 2001. 23 Mar. 2009 <
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to all living members of the family. The oldest crown group representatives are known from the
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arrive to feed when the carrion is bloat to dry (full of maggots to no maggots). In contrast,
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A remarkable ability of some hister beetles is their capacity to live in close proximity with
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around the body and it is horizontally segmented. The legs are short and do not help much in
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as blood. This blood does not contain any oxygen, but carries nutrients throughout the body.
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temperature conditions, the hister beetle will stay in the pupal stage for about a week.
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1644:(Erichson) (Coleoptera:Histeridae)." BioControl (2005). 14 Mar. 2005. 20 Mar. 2009 <
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Summerlin, J. W., and G. T. Fincher. "Laboratory observations on the life cycle of
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http://animaldiversity.ummz.umich.edu/site/accounts/classification/Histeridae.html
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510:. A distinctive characteristic is that the antennae are elbowed and contain three
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https://web.archive.org/web/20090330011619/http://www.beetlelady.com/?page_id=7
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http://encarta.msn.com/encnet/refpages/RefArticle.aspx?refid=761555773&pn=2
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in particular. Some species of Histeridae will even feed on other Histeridae.
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and will even eat other hister beetles, they must be isolated when collected.
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Jiang, Rixin; Song, Weicai; Yang, Huiyu; Shi, Chao; Wang, Shuo (July 2020).
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The male and female reproductive organs are hidden underneath the last few
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exposed. Hister beetles have specialized heads that can retract into their
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http://www.deathonline.net/decomposition/corpse_fauna/beetles/hister.htm
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http://www.deathonline.net/decomposition/corpse_fauna/beetles/hister.htm
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Mazur, Slawomir. "Ecology and Habitats of Histeridae." 13 Mar. 2009 <
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Achiano, K. A., and J. H. Giliomee. "Biology of the house fly predator
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while the male's is adapted as a copulatory structure. The female has
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http://www.museum.hokudai.ac.jp/exhibition/kikaku03/6sympoMazur.html
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Peter W. Kovarik & Michael S. Caterino (2001). "Histeridae". In
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The majority of Histeridae species prefer dried, decaying habitats.
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species from mid-Cretaceous Burmese amber (Coleoptera: Histeridae)"
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http://www.canacoll.org/Coleo/Checklist/PDF%20files/HISTERIDAE.pdf
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Histeridae live in widespread habitats because they feed on other
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stages of other insects. Certain species are also used to control
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investigations to help in time of death estimation. Also, certain
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found worldwide. They can be easily identified by their shortened
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1325:"A Mesozoic clown beetle myrmecophile (Coleoptera: Histeridae)"
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present. Insects that feed on dead bodies increase the rate of
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throughout the world. Hister beetles have proved useful during
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which holds the sperm until the eggs are fully developed.
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http://science.howstuffworks.com/forensic-entomology2.htm
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to the copulatory structure, which stays concealed until
740:) side. The female's structure is modified to work as an
2141:(furniture beetles, death watch beetles, spider beetles)
1241:
427:
The hister beetles are easily identified by their shiny
393:
are used in the control of livestock pests that infest
1322:
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361:. This very diverse group of beetles contains 3,900
1293:
690:it can look pale brown and be leathery in texture.
1556:http://agspsrv34.agric.wa.gov.au/ento/forensic.htm
906:at the insects on the body and determine time of
711:. They move mostly through muscular contraction.
3993:
3018:(jewel beetles, or metallic wood-boring beetles)
1634:Further references (not necessarily used inline)
1513:http://www.forensicentomology.com/definition.htm
1381:
846:worth mentioning. These subclades are known as:
842:There are four subclades of Histeridae that are
2741:(true weevils, bark beetles, ambrosia beetles)
1908:
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1209:
1207:
1205:
1203:
1201:
1199:
1143:
1141:
756:. The males also have a duct that carries the
1754:
727:
682:is shiny and smooth. In certain species like
2984:(ironclad beetles, cylindrical bark beetles)
1150:"Family-group names in Coleoptera (Insecta)"
1476:
1196:
1138:
1761:
1747:
1733:Lectures on Forensically Important Insects
1523:http://www.forensicentomology.com/info.htm
928:
111:
3648:List of subgroups of the order Coleoptera
2728:(straight snout weevils, New York weevil)
1827:
1358:
1340:
1179:
1169:
830:Learn how and when to remove this message
646:, meaning it goes through four stages of
3155:(cicada beetle, cicada parasite beetles)
414:
373:exposed, and their geniculate (elbowed)
1580:Integrated Taxonomic Information System
748:that carry the developed eggs from the
567:
14:
3994:
3245:(false fireflies, long-lipped beetles)
2070:
2021:
451:, hister beetles commonly feed on the
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3660:
2069:
2020:
1895:
1894:
1826:
1742:
990:
3956:B718293D-FFCE-F20F-FF0D-FC9EBD8DF65D
3917:c9c5cba9-392d-4db4-818b-e6d3d8b48861
3829:6a08aaed-bb27-42ac-8a42-576a697dd7bf
3271:(false metallic wood-boring beetles)
1592:
1568:
812:adding citations to reliable sources
783:
586:of China, which was resolved as the
900:
572:The oldest record of the family is
122:with larvae and anatomical details
24:
630:will actually feed on adult ants.
408:
25:
4028:
1674:
959:They do not actually feed on the
603:
4017:Taxa named by Leonard Gyllenhaal
2882:Perimylopidae, or Promecheilidae
2467:(minute brown scavenger beetles)
1681:
1102:
1088:
1074:
1060:
1046:
1032:
1018:
1004:
788:
499:. The head has compound eyes, a
435:with typically two of the seven
131:
43:
3097:(variegated mud-loving beetles)
1616:
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1535:
1505:
1491:10.1016/j.forsciint.2008.09.018
1470:
1461:
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799:needs additional citations for
703:around the head. There is some
1479:Forensic Science International
1316:
1284:
1260:
1250:
1232:
637:
120:Central European clown beetles
13:
1:
2926:(narrow-waisted bark beetles)
2618:(parasitic flat bark beetles)
2569:(short-winged flower beetles)
2425:(lady beetles, or God's cows)
2162:(tooth-necked fungus beetles)
1412:10.1016/j.cretres.2020.104443
1131:
514:that form a club at the end.
443:and two geniculate (elbowed)
369:that leaves two of the seven
3438:(sand-loving scarab beetles)
2828:(minute tree-fungus beetles)
2654:(silvanid flat bark beetles)
779:
470:
7:
3587:(primitive carrion beetles)
3543:(minute mud-loving beetles)
3406:(bumble bee scarab beetles)
3176:(soft-bodied plant beetles)
3147:(soft bodied plant beetles)
1950:(predaceous diving beetles)
1119:
517:Clown beetles have an open
10:
4033:
3422:(scavenger scarab beetles)
3414:(enigmatic scarab beetles)
2587:Lamingtonium binnaberrense
2301:(soft-wing flower beetles)
1110:Paromalus parallelepipedus
728:Male and female structures
644:holometabolous development
536:
486:
3669:
3645:
3575:
3561:(water scavenger beetles)
3526:
3492:
3483:
3361:
3352:
3305:
3294:Rhinorhipus tamborinensis
3280:
3164:
3135:
3073:(long-toed water beetles)
3032:
3006:
2997:
2966:(polypore fungus beetles)
2868:(palm and flower beetles)
2852:(tumbling flower beetles)
2799:(ant-like flower beetles)
2779:
2758:
2684:
2577:(lined flat bark beetles)
2551:(pleasing fungus beetles)
2491:
2446:(handsome fungus beetles)
2382:
2238:
2184:
2175:
2150:
2092:
2083:
2079:
2065:
2030:
2016:
1991:(burrowing water beetles)
1904:
1890:
1836:
1822:
1778:
1228:. Vol. 1. CRC Press.
1126:List of Histeridae genera
913:A study was conducted at
693:
377:with clubbed ends. These
333:
326:
232:
227:
128:Scientific classification
126:
119:
110:
34:
3608:(feather-winged beetles)
2942:(false longhorn beetles)
2889:(jugular-horned beetles)
2836:(false darkling beetles)
2807:(cryptic fungus beetles)
2626:(shining flower beetles)
1966:(crawling water beetles)
1865:Sikhotealinia zhiltzovae
1384:"Discovery of the first
714:
642:Histeridae goes through
475:Histeridae was named by
41:Early Cretaceous–Present
3089:(forest stream beetles)
2950:(synchroa bark beetles)
2910:(dead log bark beetles)
2902:(fire-coloured beetles)
2791:(ant-like leaf beetles)
2525:(silken fungus beetles)
2459:(well polished beetles)
2433:(minute fungus beetles)
2252:Acanthocnemus nigricans
973:Saprinus pennsylvanicus
929:Importance in forensics
3600:(round fungus beetles)
2934:(false flower beetles)
2918:(wedge-shaped beetles)
2876:(false blister beetle)
2860:(hairy fungus beetles)
2815:(conifer bark beetles)
2712:(leaf-rolling weevils)
2373:(bark-gnawing beetles)
2133:(wounded-tree beetles)
1999:(false ground beetles)
1921:(trout-stream beetles)
1713:Overview of Histeridae
1171:10.3897/zookeys.88.807
1096:Pachylister inaequalis
1068:Hister quadrimaculatus
669:
447:with clubbed ends. As
422:
3938:Paleobiology Database
3512:(false clown beetles)
3334:(plate-thigh beetles)
3216:(false click beetles)
3121:(water-penny beetles)
2770:(ship-timber beetles)
2749:(pine flower weevils)
2597:(root-eating beetles)
2417:(minute bark beetles)
2328:Phloiophilus edwardsi
1855:(reticulated beetles)
1054:Haeterius ferrugineus
418:
3824:Fauna Europaea (new)
3390:(false stag beetles)
3375:Belohina inexpectata
3237:(net-winged beetles)
3200:(rare click beetles)
3113:(travertine beetles)
3105:(minute mud beetles)
2670:(dry-fungus beetles)
2505:Agapytho foveicollis
2262:(false skin beetles)
2125:(Jacobson's beetles)
1845:Crowsoniella relicta
1705:Histeridae of Europe
1082:Onthophilus striatus
965:Hister quadrinotatus
915:Hacettepe University
808:improve this article
736:on the mesosternal (
568:Evolutionary history
553:(myrmecophiles) and
3388:Diphyllostomatidae
2720:(primitive weevils)
2533:(flat bark beetles)
2283:(checkered beetles)
2270:(fruitworm beetles)
1958:(whirligig beetles)
1723:Forensic Entomology
1576:"Histeridae Report"
1404:2020CrRes.11104443J
1392:Cretaceous Research
1342:10.7554/eLife.44985
1216:Arnett, Ross H. Jr.
1162:2011ZooK...88....1B
1040:Atholus bimaculatus
1026:Asolenus julietteae
934:Forensic entomology
401:. Because they are
335:at least 410 genera
3253:(glowworm beetles)
2958:(darkling beetles)
2805:Archeocrypticidae
2662:(palmetto beetles)
2409:(dry bark beetles)
2196:(longhorn beetles)
1561:2006-08-19 at the
1528:2018-02-15 at the
1518:2018-02-14 at the
1309:2008-02-03 at the
1273:2003-08-21 at the
1220:Thomas, Michael C.
991:Genera and species
519:circulatory system
477:Leonard Gyllenhaal
423:
353:commonly known as
3989:
3988:
3925:Open Tree of Life
3663:Taxon identifiers
3654:
3653:
3641:
3640:
3637:
3636:
3633:
3632:
3625:
3617:
3616:(carrion beetles)
3609:
3601:
3588:
3562:
3544:
3513:
3505:
3479:
3478:
3471:
3463:
3455:
3447:
3439:
3431:
3423:
3415:
3407:
3399:
3391:
3378:
3348:
3347:
3335:
3327:
3297:
3272:
3254:
3246:
3238:
3230:
3217:
3209:
3201:
3193:
3192:(soldier beetles)
3185:
3182:Brachypsectridae
3177:
3156:
3148:
3122:
3114:
3106:
3098:
3090:
3082:
3074:
3061:
3053:
3045:
3019:
2993:
2992:
2985:
2967:
2959:
2951:
2943:
2940:Stenotrachelidae
2935:
2927:
2919:
2911:
2903:
2890:
2877:
2869:
2861:
2853:
2845:
2844:(blister beetles)
2837:
2829:
2816:
2808:
2800:
2792:
2771:
2750:
2742:
2729:
2721:
2713:
2705:
2695:
2671:
2663:
2655:
2627:
2619:
2611:
2598:
2590:
2578:
2570:
2552:
2534:
2526:
2508:
2468:
2460:
2447:
2434:
2426:
2418:
2410:
2392:Akalyptoischiidae
2374:
2331:
2314:
2302:
2284:
2271:
2263:
2255:
2205:
2197:
2171:
2170:
2163:
2142:
2134:
2126:
2113:
2105:
2061:
2060:
2038:
2012:
2011:
2008:
2007:
2000:
1992:
1984:
1967:
1959:
1951:
1943:
1935:
1922:
1886:
1885:
1868:
1856:
1848:
1642:Carcinops pumilio
840:
839:
832:
340:
339:
320:
312:
304:
296:
288:
280:
272:
264:
256:
248:
240:
223:
27:Family of beetles
16:(Redirected from
4024:
3982:
3981:
3969:
3968:
3959:
3958:
3946:
3945:
3933:
3932:
3920:
3919:
3910:
3909:
3897:
3896:
3894:NHMSYS0001718459
3884:
3883:
3871:
3870:
3858:
3857:
3845:
3844:
3832:
3831:
3819:
3818:
3806:
3805:
3793:
3792:
3780:
3779:
3767:
3766:
3754:
3753:
3741:
3740:
3731:
3730:
3718:
3717:
3705:
3704:
3703:
3690:
3689:
3688:
3658:
3657:
3623:
3615:
3607:
3599:
3586:
3560:
3542:
3511:
3503:
3490:
3489:
3485:Staphyliniformia
3469:
3461:
3453:
3445:
3437:
3429:
3421:
3413:
3405:
3397:
3389:
3372:
3359:
3358:
3333:
3324:Declinia relicta
3321:
3291:
3270:
3264:Sinopyrophoridae
3259:Rhagophthalmidae
3252:
3244:
3236:
3228:
3215:
3207:
3199:
3191:
3183:
3175:
3174:Artematopodidae
3154:
3146:
3120:
3112:
3104:
3096:
3088:
3081:(riffle beetles)
3080:
3072:
3060:(turtle beetles)
3059:
3051:
3043:
3017:
3004:
3003:
2983:
2965:
2957:
2949:
2941:
2933:
2925:
2917:
2909:
2901:
2888:
2875:
2867:
2859:
2851:
2843:
2835:
2827:
2814:
2806:
2798:
2790:
2769:
2748:
2740:
2727:
2719:
2711:
2704:(fungus weevils)
2703:
2694:
2693:
2689:
2676:Tasmosalpingidae
2669:
2661:
2653:
2625:
2617:
2609:
2596:
2584:
2576:
2568:
2550:
2532:
2524:
2502:
2466:
2458:
2445:
2432:
2424:
2416:
2408:
2372:
2325:
2308:
2300:
2282:
2269:
2261:
2249:
2248:Acanthocnemidae
2203:
2195:
2182:
2181:
2161:
2140:
2132:
2124:
2111:
2103:
2090:
2089:
2081:
2080:
2067:
2066:
2036:
2018:
2017:
1998:
1990:
1978:
1965:
1957:
1949:
1941:
1934:(ground beetles)
1933:
1920:
1906:
1905:
1892:
1891:
1862:
1854:
1842:
1841:Crowsoniellidae
1824:
1823:
1763:
1756:
1749:
1740:
1739:
1709:
1686:Data related to
1685:
1627:
1620:
1614:
1613:
1611:
1610:
1596:
1590:
1589:
1587:
1586:
1572:
1566:
1552:
1546:
1539:
1533:
1509:
1503:
1502:
1474:
1468:
1465:
1459:
1456:
1450:
1443:
1437:
1430:
1424:
1423:
1379:
1373:
1372:
1362:
1344:
1320:
1314:
1300:
1291:
1288:
1282:
1264:
1258:
1254:
1248:
1245:
1239:
1236:
1230:
1229:
1225:American Beetles
1211:
1194:
1193:
1183:
1173:
1145:
1106:
1092:
1078:
1064:
1050:
1036:
1022:
1008:
901:Current research
835:
828:
824:
821:
815:
792:
784:
584:Yixian Formation
506:, and shortened
318:
310:
302:
294:
286:
278:
270:
262:
254:
246:
238:
218:
195:Staphyliniformia
136:
135:
115:
105:
42:
38:Temporal range:
32:
31:
21:
4032:
4031:
4027:
4026:
4025:
4023:
4022:
4021:
4012:Beetle families
3992:
3991:
3990:
3985:
3977:
3972:
3964:
3962:
3954:
3949:
3941:
3936:
3928:
3923:
3915:
3913:
3905:
3900:
3892:
3887:
3879:
3874:
3866:
3861:
3853:
3848:
3840:
3835:
3827:
3822:
3814:
3809:
3801:
3796:
3788:
3783:
3775:
3770:
3762:
3757:
3749:
3744:
3736:
3734:
3726:
3721:
3713:
3708:
3699:
3698:
3693:
3684:
3683:
3678:
3665:
3655:
3650:
3629:
3571:
3522:
3504:(clown beetles)
3475:
3446:(betsy beetles)
3354:Scarabaeiformia
3344:
3301:
3276:
3208:(click beetles)
3184:(Texas beetles)
3160:
3131:
3127:Ptilodactylidae
3052:(cedar beetles)
3028:
2989:
2972:Trictenotomidae
2858:Mycetophagidae
2775:
2754:
2691:
2690:
2688:
2680:
2632:Phloeostichidae
2583:Lamingtoniidae
2575:Laemophloeidae
2523:Cryptophagidae
2487:
2378:
2361:Thanerocleridae
2324:Phloiophilidae
2311:Metaxina ornata
2276:Chaetosomatidae
2234:
2167:
2146:
2104:(auger beetles)
2085:Bostrichiformia
2075:
2057:
2053:Torridincolidae
2037:(skiff beetles)
2035:Hydroscaphidae
2026:
2004:
1997:Trachypachidae
1942:(tiger beetles)
1900:
1882:
1832:
1818:
1774:
1767:
1737:
1707:
1677:
1672:
1636:
1631:
1630:
1621:
1617:
1608:
1606:
1598:
1597:
1593:
1584:
1582:
1574:
1573:
1569:
1563:Wayback Machine
1553:
1549:
1540:
1536:
1530:Wayback Machine
1520:Wayback Machine
1510:
1506:
1475:
1471:
1466:
1462:
1457:
1453:
1444:
1440:
1431:
1427:
1380:
1376:
1321:
1317:
1311:Wayback Machine
1301:
1294:
1289:
1285:
1275:Wayback Machine
1265:
1261:
1255:
1251:
1246:
1242:
1237:
1233:
1212:
1197:
1146:
1139:
1134:
1122:
1117:
1116:
1115:
1112:
1107:
1098:
1093:
1084:
1079:
1070:
1065:
1056:
1051:
1042:
1037:
1028:
1023:
1014:
1012:Acritus minutus
1009:
993:
969:Hister sedakovi
963:. For example,
931:
903:
887:blind as well.
836:
825:
819:
816:
805:
793:
782:
730:
717:
696:
672:
640:
606:
570:
539:
489:
473:
425:
424:
420:Hister unicolor
411:
409:Characteristics
397:and to control
319:Bickhardt, 1913
303:Bickhardt, 1914
295:Blanchard, 1845
271:Gyllenhal, 1808
247:Bickhardt, 1914
217:
130:
121:
106:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
40:
39:
36:
28:
23:
22:
15:
12:
11:
5:
4030:
4020:
4019:
4014:
4009:
4004:
3987:
3986:
3984:
3983:
3970:
3960:
3947:
3934:
3921:
3911:
3898:
3885:
3872:
3859:
3846:
3833:
3820:
3811:Fauna Europaea
3807:
3794:
3781:
3768:
3755:
3742:
3732:
3719:
3706:
3691:
3675:
3673:
3667:
3666:
3652:
3651:
3646:
3643:
3642:
3639:
3638:
3635:
3634:
3631:
3630:
3628:
3627:
3624:(rove beetles)
3622:Staphylinidae
3619:
3611:
3603:
3595:
3590:
3581:
3579:
3577:Staphylinoidea
3573:
3572:
3570:
3569:
3564:
3559:Hydrophilidae
3556:
3551:
3546:
3538:
3532:
3530:
3528:Hydrophiloidea
3524:
3523:
3521:
3520:
3515:
3507:
3498:
3496:
3487:
3481:
3480:
3477:
3476:
3474:
3473:
3470:(hide beetles)
3465:
3457:
3454:(rain beetles)
3449:
3441:
3433:
3430:(stag beetles)
3425:
3417:
3409:
3401:
3393:
3385:
3383:Bolboceratidae
3380:
3367:
3365:
3356:
3350:
3349:
3346:
3345:
3343:
3342:
3337:
3329:
3317:
3311:
3309:
3303:
3302:
3300:
3299:
3290:Rhinorhipidae
3286:
3284:
3282:Rhinorhipoidea
3278:
3277:
3275:
3274:
3266:
3261:
3256:
3248:
3240:
3232:
3224:
3219:
3211:
3203:
3195:
3187:
3179:
3170:
3168:
3162:
3161:
3159:
3158:
3150:
3141:
3139:
3133:
3132:
3130:
3129:
3124:
3116:
3108:
3100:
3095:Heteroceridae
3092:
3084:
3076:
3068:
3063:
3058:Chelonariidae
3055:
3050:Callirhipidae
3047:
3044:(pill beetles)
3038:
3036:
3030:
3029:
3027:
3026:
3021:
3012:
3010:
3001:
2995:
2994:
2991:
2990:
2988:
2987:
2979:
2974:
2969:
2961:
2956:Tenebrionidae
2953:
2945:
2937:
2929:
2921:
2913:
2905:
2897:
2892:
2884:
2879:
2871:
2863:
2855:
2847:
2839:
2831:
2823:
2818:
2810:
2802:
2794:
2785:
2783:
2781:Tenebrionoidea
2777:
2776:
2774:
2773:
2764:
2762:
2756:
2755:
2753:
2752:
2744:
2739:Curculionidae
2736:
2731:
2723:
2715:
2707:
2698:
2696:
2686:Curculionoidea
2682:
2681:
2679:
2678:
2673:
2665:
2657:
2649:
2647:Protocucujidae
2644:
2639:
2634:
2629:
2621:
2613:
2605:
2600:
2592:
2580:
2572:
2564:
2559:
2554:
2546:
2541:
2539:Cybocephalidae
2536:
2528:
2520:
2515:
2510:
2497:
2495:
2489:
2488:
2486:
2485:
2480:
2475:
2470:
2462:
2454:
2449:
2441:
2439:Discolomatidae
2436:
2428:
2423:Coccinellidae
2420:
2412:
2407:Bothrideridae
2404:
2399:
2394:
2388:
2386:
2384:Coccinelloidea
2380:
2379:
2377:
2376:
2371:Trogossitidae
2368:
2363:
2358:
2353:
2348:
2343:
2338:
2333:
2321:
2316:
2304:
2296:
2291:
2289:Lophocateridae
2286:
2278:
2273:
2265:
2257:
2244:
2242:
2236:
2235:
2233:
2232:
2227:
2222:
2217:
2212:
2207:
2204:(leaf beetles)
2202:Chrysomelidae
2199:
2190:
2188:
2186:Chrysomeloidea
2179:
2173:
2172:
2169:
2168:
2166:
2165:
2156:
2154:
2148:
2147:
2145:
2144:
2136:
2131:Nosodendridae
2128:
2123:Jacobsoniidae
2120:
2115:
2112:(skin beetles)
2107:
2098:
2096:
2087:
2077:
2076:
2063:
2062:
2059:
2058:
2056:
2055:
2050:
2045:
2040:
2031:
2028:
2027:
2014:
2013:
2010:
2009:
2006:
2005:
2003:
2002:
1994:
1986:
1981:Meru phyllisae
1974:
1969:
1961:
1953:
1945:
1937:
1929:
1924:
1915:
1913:
1902:
1901:
1888:
1887:
1884:
1883:
1881:
1880:
1875:
1873:Micromalthidae
1870:
1858:
1850:
1837:
1834:
1833:
1820:
1819:
1817:
1816:
1810:
1804:
1798:
1792:
1786:
1779:
1776:
1775:
1766:
1765:
1758:
1751:
1743:
1736:
1735:
1730:
1725:
1720:
1715:
1710:
1702:
1696:
1691:
1690:at Wikispecies
1678:
1676:
1675:External links
1673:
1671:
1670:
1663:
1656:
1649:
1637:
1635:
1632:
1629:
1628:
1615:
1591:
1567:
1547:
1534:
1504:
1485:(1–3): 24–32.
1469:
1460:
1451:
1438:
1425:
1374:
1315:
1292:
1283:
1259:
1249:
1240:
1231:
1195:
1136:
1135:
1133:
1130:
1129:
1128:
1121:
1118:
1114:
1113:
1108:
1101:
1099:
1094:
1087:
1085:
1080:
1073:
1071:
1066:
1059:
1057:
1052:
1045:
1043:
1038:
1031:
1029:
1024:
1017:
1015:
1010:
1003:
1000:
999:
998:
992:
989:
987:in their way.
930:
927:
902:
899:
882:Microhisterids
838:
837:
820:September 2018
796:
794:
787:
781:
778:
729:
726:
716:
713:
701:sclerotization
695:
692:
671:
668:
639:
636:
605:
604:Feeding habits
602:
588:sister species
569:
566:
538:
535:
488:
485:
472:
469:
413:
412:
410:
407:
359:hister beetles
338:
337:
331:
330:
324:
323:
322:
321:
313:
305:
297:
289:
281:
273:
265:
257:
252:Dendrophilinae
249:
244:Chlamydopsinae
241:
230:
229:
225:
224:
212:
208:
207:
202:
198:
197:
192:
188:
187:
182:
178:
177:
172:
168:
167:
162:
158:
157:
152:
148:
147:
142:
138:
137:
124:
123:
117:
116:
108:
107:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
37:
26:
9:
6:
4:
3:
2:
4029:
4018:
4015:
4013:
4010:
4008:
4005:
4003:
4000:
3999:
3997:
3980:
3975:
3971:
3967:
3961:
3957:
3952:
3948:
3944:
3939:
3935:
3931:
3926:
3922:
3918:
3912:
3908:
3903:
3899:
3895:
3890:
3886:
3882:
3877:
3873:
3869:
3864:
3860:
3856:
3851:
3847:
3843:
3838:
3834:
3830:
3825:
3821:
3817:
3812:
3808:
3804:
3799:
3795:
3791:
3786:
3782:
3778:
3773:
3769:
3765:
3760:
3756:
3752:
3747:
3743:
3739:
3733:
3729:
3724:
3720:
3716:
3711:
3707:
3702:
3696:
3692:
3687:
3681:
3677:
3676:
3674:
3672:
3668:
3664:
3659:
3649:
3644:
3626:
3620:
3618:
3612:
3610:
3604:
3602:
3596:
3594:
3591:
3589:
3583:
3582:
3580:
3578:
3574:
3568:
3565:
3563:
3557:
3555:
3552:
3550:
3547:
3545:
3539:
3537:
3534:
3533:
3531:
3529:
3525:
3519:
3516:
3514:
3510:Sphaeritidae
3508:
3506:
3500:
3499:
3497:
3495:
3491:
3488:
3486:
3482:
3472:
3466:
3464:
3460:Scarabaeidae
3458:
3456:
3450:
3448:
3442:
3440:
3434:
3432:
3426:
3424:
3418:
3416:
3410:
3408:
3402:
3400:
3398:(dor beetles)
3394:
3392:
3386:
3384:
3381:
3379:
3376:
3369:
3368:
3366:
3364:
3363:Scarabaeoidea
3360:
3357:
3355:
3351:
3341:
3338:
3336:
3330:
3328:
3325:
3318:
3316:
3313:
3312:
3310:
3308:
3304:
3298:
3295:
3288:
3287:
3285:
3283:
3279:
3273:
3267:
3265:
3262:
3260:
3257:
3255:
3249:
3247:
3241:
3239:
3233:
3231:
3225:
3223:
3220:
3218:
3212:
3210:
3204:
3202:
3198:Cerophytidae
3196:
3194:
3188:
3186:
3180:
3178:
3172:
3171:
3169:
3167:
3163:
3157:
3153:Rhipiceridae
3151:
3149:
3143:
3142:
3140:
3138:
3134:
3128:
3125:
3123:
3117:
3115:
3109:
3107:
3101:
3099:
3093:
3091:
3087:Eulichadidae
3085:
3083:
3077:
3075:
3069:
3067:
3066:Cneoglossidae
3064:
3062:
3056:
3054:
3048:
3046:
3040:
3039:
3037:
3035:
3031:
3025:
3024:Schizopodidae
3022:
3020:
3014:
3013:
3011:
3009:
3005:
3002:
3000:
2999:Elateriformia
2996:
2986:
2980:
2978:
2975:
2973:
2970:
2968:
2964:Tetratomidae
2962:
2960:
2954:
2952:
2946:
2944:
2938:
2936:
2930:
2928:
2922:
2920:
2916:Ripiphoridae
2914:
2912:
2906:
2904:
2900:Pyrochroidae
2898:
2896:
2895:Pterogeniidae
2893:
2891:
2885:
2883:
2880:
2878:
2872:
2870:
2864:
2862:
2856:
2854:
2848:
2846:
2840:
2838:
2834:Melandryidae
2832:
2830:
2824:
2822:
2821:Chalcodryidae
2819:
2817:
2811:
2809:
2803:
2801:
2795:
2793:
2787:
2786:
2784:
2782:
2778:
2772:
2766:
2765:
2763:
2761:
2757:
2751:
2747:Nemonychidae
2745:
2743:
2737:
2735:
2732:
2730:
2724:
2722:
2716:
2714:
2708:
2706:
2700:
2699:
2697:
2687:
2683:
2677:
2674:
2672:
2666:
2664:
2658:
2656:
2650:
2648:
2645:
2643:
2642:Propalticidae
2640:
2638:
2637:Priasilphidae
2635:
2633:
2630:
2628:
2622:
2620:
2616:Passandridae
2614:
2612:
2610:(sap beetles)
2606:
2604:
2601:
2599:
2593:
2591:
2588:
2581:
2579:
2573:
2571:
2565:
2563:
2560:
2558:
2555:
2553:
2547:
2545:
2542:
2540:
2537:
2535:
2529:
2527:
2521:
2519:
2518:Cavognathidae
2516:
2514:
2511:
2509:
2506:
2499:
2498:
2496:
2494:
2490:
2484:
2481:
2479:
2476:
2474:
2471:
2469:
2463:
2461:
2455:
2453:
2452:Eupsilobiidae
2450:
2448:
2444:Endomychidae
2442:
2440:
2437:
2435:
2431:Corylophidae
2429:
2427:
2421:
2419:
2413:
2411:
2405:
2403:
2400:
2398:
2395:
2393:
2390:
2389:
2387:
2385:
2381:
2375:
2369:
2367:
2364:
2362:
2359:
2357:
2354:
2352:
2349:
2347:
2346:Protopeltidae
2344:
2342:
2341:Prionoceridae
2339:
2337:
2334:
2332:
2329:
2322:
2320:
2317:
2315:
2312:
2305:
2303:
2297:
2295:
2294:Mauroniscidae
2292:
2290:
2287:
2285:
2279:
2277:
2274:
2272:
2266:
2264:
2258:
2256:
2253:
2246:
2245:
2243:
2241:
2237:
2231:
2228:
2226:
2223:
2221:
2218:
2216:
2215:Megalopodidae
2213:
2211:
2208:
2206:
2200:
2198:
2194:Cerambycidae
2192:
2191:
2189:
2187:
2183:
2180:
2178:
2174:
2164:
2160:Derodontidae
2158:
2157:
2155:
2153:
2152:Derodontoidea
2149:
2143:
2137:
2135:
2129:
2127:
2121:
2119:
2118:Endecatomidae
2116:
2114:
2108:
2106:
2102:Bostrichidae
2100:
2099:
2097:
2095:
2094:Bostrichoidea
2091:
2088:
2086:
2082:
2078:
2074:
2068:
2064:
2054:
2051:
2049:
2048:Sphaeriusidae
2046:
2044:
2041:
2039:
2033:
2032:
2029:
2025:
2019:
2015:
2001:
1995:
1993:
1987:
1985:
1982:
1975:
1973:
1970:
1968:
1962:
1960:
1954:
1952:
1946:
1944:
1940:Cicindelidae
1938:
1936:
1930:
1928:
1925:
1923:
1917:
1916:
1914:
1911:
1907:
1903:
1899:
1893:
1889:
1879:
1876:
1874:
1871:
1869:
1866:
1859:
1857:
1851:
1849:
1846:
1839:
1838:
1835:
1831:
1825:
1821:
1815:
1811:
1809:
1805:
1803:
1799:
1797:
1793:
1791:
1787:
1785:
1781:
1780:
1777:
1772:
1764:
1759:
1757:
1752:
1750:
1745:
1744:
1741:
1734:
1731:
1729:
1726:
1724:
1721:
1719:
1716:
1714:
1711:
1706:
1703:
1700:
1699:Histeridae de
1697:
1695:
1692:
1689:
1684:
1680:
1679:
1668:
1664:
1661:
1657:
1654:
1650:
1647:
1643:
1639:
1638:
1625:
1619:
1605:
1601:
1595:
1581:
1577:
1571:
1564:
1560:
1557:
1551:
1544:
1538:
1531:
1527:
1524:
1521:
1517:
1514:
1508:
1500:
1496:
1492:
1488:
1484:
1480:
1473:
1464:
1455:
1448:
1442:
1435:
1429:
1421:
1417:
1413:
1409:
1405:
1401:
1397:
1393:
1389:
1387:
1378:
1370:
1366:
1361:
1356:
1352:
1348:
1343:
1338:
1334:
1330:
1326:
1319:
1312:
1308:
1305:
1299:
1297:
1287:
1280:
1276:
1272:
1269:
1263:
1253:
1244:
1235:
1227:
1226:
1221:
1217:
1210:
1208:
1206:
1204:
1202:
1200:
1191:
1187:
1182:
1177:
1172:
1167:
1163:
1159:
1156:(88): 1–972.
1155:
1151:
1144:
1142:
1137:
1127:
1124:
1123:
1111:
1105:
1100:
1097:
1091:
1086:
1083:
1077:
1072:
1069:
1063:
1058:
1055:
1049:
1044:
1041:
1035:
1030:
1027:
1021:
1016:
1013:
1007:
1002:
1001:
997:
988:
984:
981:
976:
974:
970:
966:
962:
956:
954:
953:decomposition
950:
946:
942:
939:Flesh eating
937:
935:
926:
924:
920:
916:
911:
909:
898:
894:
893:
888:
884:
883:
878:
876:
872:
866:
865:
860:
858:
853:
852:
847:
845:
834:
831:
823:
813:
809:
803:
802:
797:This section
795:
791:
786:
785:
777:
775:
771:
770:fertilization
767:
763:
759:
755:
751:
747:
743:
739:
735:
725:
722:
721:proteinaceous
712:
710:
706:
702:
691:
689:
685:
681:
677:
667:
665:
661:
657:
653:
649:
645:
635:
631:
629:
628:
621:
618:
616:
612:
601:
600:
598:
593:
592:Burmese amber
589:
585:
581:
577:
576:
565:
563:
558:
556:
552:
547:
545:
534:
532:
528:
524:
520:
515:
513:
509:
505:
502:
498:
494:
484:
482:
478:
468:
466:
462:
458:
454:
450:
446:
442:
438:
434:
430:
421:
417:
406:
404:
400:
396:
392:
388:
384:
380:
376:
372:
368:
364:
360:
356:
355:clown beetles
352:
348:
344:
336:
332:
329:
325:
317:
314:
311:Marseul, 1857
309:
306:
301:
298:
293:
290:
287:MacLeay, 1819
285:
284:Onthophilinae
282:
277:
274:
269:
266:
263:Marseul, 1857
261:
258:
255:Reitter, 1909
253:
250:
245:
242:
239:MacLeay, 1819
237:
234:
233:
231:
226:
221:
216:
213:
210:
209:
206:
203:
201:Superfamily:
200:
199:
196:
193:
190:
189:
186:
183:
180:
179:
176:
173:
170:
169:
166:
163:
160:
159:
156:
153:
150:
149:
146:
143:
140:
139:
134:
129:
125:
118:
114:
109:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
33:
30:
19:
3670:
3549:Helophoridae
3541:Georissidae
3536:Epimetopidae
3501:
3452:Pleocomidae
3436:Ochodaeidae
3420:Hybosoridae
3404:Glaphyridae
3396:Geotrupidae
3374:
3371:Belohinidae
3332:Eucinetidae
3323:
3320:Decliniidae
3293:
3251:Phengodidae
3190:Cantharidae
3145:Dascillidae
3137:Dascilloidea
3119:Psephenidae
3111:Lutrochidae
3103:Limnichidae
3016:Buprestidae
3008:Buprestoidea
2948:Synchroidae
2932:Scraptiidae
2924:Salpingidae
2887:Prostomidae
2874:Oedemeridae
2850:Mordellidae
2768:Lymexylidae
2760:Lymexyloidea
2710:Attelabidae
2702:Anthribidae
2660:Smicripidae
2624:Phalacridae
2608:Nitidulidae
2603:Myraboliidae
2595:Monotomidae
2586:
2567:Kateretidae
2544:Cyclaxyridae
2504:
2501:Agapythidae
2465:Latridiidae
2415:Cerylonidae
2402:Anamorphidae
2336:Phycosecidae
2327:
2310:
2307:Metaxinidae
2260:Biphyllidae
2251:
2220:Orsodacnidae
2177:Cucujiformia
2110:Dermestidae
1980:
1919:Amphizoidae
1864:
1844:
1830:Archostemata
1814:Holometabola
1812:Superorder:
1806:Infraclass:
1694:Tree of life
1667:Hister nomas
1666:
1641:
1618:
1607:. Retrieved
1603:
1600:"Histeridae"
1594:
1583:. Retrieved
1579:
1570:
1550:
1537:
1507:
1482:
1478:
1472:
1463:
1454:
1441:
1428:
1395:
1391:
1385:
1377:
1332:
1328:
1318:
1286:
1262:
1252:
1243:
1234:
1223:
1153:
1109:
1095:
1081:
1067:
1053:
1039:
1025:
1011:
994:
985:
977:
972:
968:
964:
957:
938:
932:
912:
908:colonization
904:
895:
891:
889:
885:
881:
879:
867:
863:
861:
854:
850:
848:
841:
826:
817:
806:Please help
801:verification
798:
731:
718:
705:pigmentation
697:
688:Platylomalus
687:
683:
673:
641:
632:
625:
622:
619:
607:
595:
575:Antigracilus
573:
571:
559:
548:
540:
516:
512:antennomeres
490:
474:
426:
419:
358:
354:
342:
341:
316:Trypeticinae
308:Trypanaeinae
279:Fowler, 1912
228:Subfamilies
214:
191:Infraorder:
29:
18:Clown beetle
4007:Histeroidea
3850:iNaturalist
3695:Wikispecies
3593:Hydraenidae
3567:Spercheidae
3554:Hydrochidae
3518:Synteliidae
3502:Histeridae
3494:Histeroidea
3444:Passalidae
3412:Glaresidae
3269:Throscidae
3229:(fireflies)
3227:Lampyridae
3214:Eucnemidae
3206:Elateridae
3166:Elateroidea
2982:Zopheridae
2866:Mycteridae
2797:Anthicidae
2668:Sphindidae
2652:Silvanidae
2562:Hobartiidae
2549:Erotylidae
2478:Mycetaeidae
2473:Murmidiidae
2457:Euxestidae
2351:Rentoniidae
2225:Oxypeltidae
2210:Disteniidae
2043:Lepiceridae
1972:Hygrobiidae
1964:Haliplidae
1948:Dytiscidae
1927:Aspidytidae
1708:(in German)
1701:German site
1386:Onthophilus
1281:2009-10-31.
851:Dendrobites
774:spermatheca
738:mesosternum
678:larva. The
638:Development
627:Psiloscelis
597:Onthophilus
521:within its
260:Haeteriinae
205:Histeroidea
4002:Histeridae
3996:Categories
3728:HISTERIDAE
3715:Histeridae
3701:Histeridae
3671:Histeridae
3614:Silphidae
3606:Ptiliidae
3598:Leiodidae
3585:Agyrtidae
3428:Lucanidae
3307:Scirtoidea
3243:Omethidae
3222:Jurasaidae
3071:Dryopidae
3042:Byrrhidae
3034:Byrrhoidea
2726:Brentidae
2531:Cucujidae
2513:Boganiidae
2493:Cucujoidea
2366:Thymalidae
2356:Rhadalidae
2299:Melyridae
2268:Byturidae
2230:Vesperidae
2139:Ptiniidae
1989:Noteridae
1956:Gyrinidae
1932:Carabidae
1861:Jurodidae
1853:Cupedidae
1800:Subclass:
1790:Arthropoda
1771:Coleoptera
1688:Histeridae
1609:2019-06-21
1585:2019-06-21
1398:: 104443.
1335:: e44985.
1132:References
949:arthropods
947:and other
892:Inquilines
766:copulation
754:ovipositor
742:ovipositor
709:locomotion
562:arthropods
403:predacious
399:houseflies
343:Histeridae
300:Tribalinae
292:Saprininae
276:Niponiinae
268:Histerinae
215:Histeridae
185:Coleoptera
165:Arthropoda
35:Histeridae
3468:Trogidae
3462:(scarabs)
3340:Scirtidae
3315:Clambidae
2908:Pythidae
2842:Meloidae
2789:Aderidae
2692:(weevils)
2557:Helotidae
2483:Teredidae
2397:Alexiidae
2281:Cleridae
2240:Cleroidea
2073:Polyphaga
2071:Suborder
2024:Myxophaga
2022:Suborder
1977:Meruidae
1896:Suborder
1878:Ommatidae
1828:Suborder
1802:Pterygota
1782:Kingdom:
1728:Bug Guide
1420:216238703
1351:2050-084X
864:Geobiotes
844:predators
780:Subclades
760:from the
734:sternites
611:olfaction
578:from the
531:Spiracles
527:hemolymph
504:mouthpart
471:Etymology
465:livestock
449:predators
441:prothorax
379:predatory
328:Diversity
236:Abraeinae
220:Gyllenhal
151:Kingdom:
145:Eukaryota
3759:BugGuide
3735:BioLib:
3680:Wikidata
3235:Lycidae
3079:Elmidae
2977:Ulodidae
2813:Boridae
2718:Belidae
2319:Peltidae
1912:families
1898:Adephaga
1808:Neoptera
1788:Phylum:
1784:Animalia
1773:families
1559:Archived
1526:Archived
1516:Archived
1499:19019588
1369:30990167
1307:Archived
1279:Archived
1271:Archived
1222:(eds.).
1190:21594053
1120:See also
746:oviducts
555:termites
523:hemocoel
508:antennae
501:mandible
497:tergites
445:antennae
437:tergites
387:forensic
375:antennae
371:tergites
211:Family:
161:Phylum:
155:Animalia
141:Domain:
3966:4779751
3686:Q596252
2826:Ciidae
2734:Caridae
1796:Insecta
1794:Class:
1769:Extant
1718:Summary
1400:Bibcode
1360:6467565
1181:3088472
1158:Bibcode
1154:ZooKeys
961:carrion
945:maggots
941:insects
768:. When
752:to the
750:ovaries
684:Epierus
680:chorion
615:Diptera
544:carrion
537:Habitat
487:Anatomy
481:Juvenal
433:abdomen
391:species
363:species
351:beetles
181:Order:
175:Insecta
171:Class:
3979:150732
3963:uBio:
3930:738371
3914:NZOR:
3907:110043
3881:114959
3868:114574
3803:1HISTF
1910:Extant
1497:
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1367:
1357:
1349:
1277:>.
1218:&
1188:
1178:
980:larvae
923:Turkey
919:Ankara
762:testes
694:Larval
676:instar
580:Aptian
493:elytra
459:, and
457:larval
429:elytra
367:elytra
347:family
222:, 1808
3974:WoRMS
3951:Plazi
3943:69273
3863:IRMNG
3855:68869
3816:11102
3751:80462
1662:>.
1655:>.
1648:>.
1626:>.
1565:>.
1545:>.
1532:>.
1449:>.
1436:>.
1416:S2CID
1329:eLife
1313:>.
1257:1988.
875:mites
758:sperm
715:Pupal
664:wings
660:adult
652:larva
582:aged
461:adult
383:niche
345:is a
3902:NCBI
3876:ITIS
3842:4733
3837:GBIF
3798:EPPO
3764:6577
3746:BOLD
3738:5034
1604:GBIF
1495:PMID
1365:PMID
1347:ISSN
1186:PMID
967:and
871:dung
658:and
656:pupa
551:ants
491:The
395:dung
46:Preęž’
3889:NBN
3790:364
3785:EoL
3777:B44
3772:CoL
3723:AFD
3710:ADW
1487:doi
1483:183
1408:doi
1396:111
1355:PMC
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1176:PMC
1166:doi
917:in
890:4.
880:3.
862:2.
857:fly
849:1.
810:by
686:or
670:Egg
648:egg
453:egg
357:or
349:of
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