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Pseudanthium

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438: 608: 644: 572: 626: 255: 590: 560: 350: 82: 402: 215:) can be used as an exact synonym for pseudanthium and flower head; however, this use is generally but not always restricted to the family Asteraceae. At least one source defines it as a small flower head. In addition to its botanical use as a term meaning flower head it is also used to mean the top of the 160:
Pseudanthia may be grouped into types. The first type has units of individual flowers that are recognizable as single flowers even if fused. In the second type, the flowers do not appear as individual units and certain organs like stamens and carpels can not be associated with any individual flowers.
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In all cases, a pseudanthium is superficially indistinguishable from a flower, but closer inspection of its anatomy will reveal that it is composed of multiple flowers. Thus, the pseudanthium represents an evolutionary convergence of the inflorescence to a reduced reproductive unit that may function
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The term pseudanthium was originally applied to flowers with stamens in two whorls with the outer whorl opposite the petals (obdiplostemonate) or polyandric flowers; by the early 1900s the term was repurposed by the advocates of the 'pseudanthium theory' which assumed flower evolution originated
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In some families, it is not yet clear whether the "flower" represents a pseudanthium because the anatomical work has not been done (or is still ambiguous due to considerable evolutionary reduction). Possible pseudanthia of this type may occur in the following families:
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are grouped together to form a single flower-like structure. Pseudanthia take various forms. The real flowers (the florets) are generally small and often greatly reduced, but the pseudanthium itself can sometimes be quite large (as in the heads of some varieties of
607: 643: 589: 625: 559: 393:, composed of a single carpal flower with few to many single stamen staminate flowers contained within a cup-shaped structure or bracts; the bracts are often rimmed with 125:
and the corolla has one large lobe (the so-called "petals" of a daisy are individual ray flowers, for example). Either ray or disk flowers may be absent in some plants:
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that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include
1248:"Floral ontogeny and gene protein localization rules out euanthial interpretation of reproductive units in Lepironia (Cyperaceae, Mapanioideae, Chrysitricheae)" 185: 113:), whose flowers are differentiated into ray flowers and disk flowers, unique to this family. The disk flowers in the center of the pseudanthium are 1505:
Claßen-Bockhoff, R.; Arndt, M. (2018). "Flower-like heads from flower-like meristems: pseudanthium development in Davidia involucrata (Nyssaceae)".
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Petra Hoffmann, Hashendra S. Kathriarachchi, and Kenneth J. Wurdack. 2006. "A Phylogenetic Classification of Phyllanthaceae (Malpighiales)."
340:— Where individual male and female flowers are grouped together and wrapped in bracts forming a pseudanthium appearing as a bisexual flower. 242:, as a specific term for a flower head of a plant in the family Asteraceae. However, on-line botanical glossaries do not define it, and 571: 1393:"The unique pseudanthium of Actinodium (Myrtaceae) - morphological reinvestigation and possible regulation by CYCLOIDEA -like genes" 458:— the pseudanthia is a head-like structure with fertile flowers in the center and showy ray-like structures along the outside. 325:. Commonly the capitulum has ray flowers specialized to attract pollinators arranged surrounding disc flowers responsible for 1375: 1348: 1321: 1159: 1132: 1021: 995: 948: 923: 897: 690: 321:, which are collections of different types of flowers, is a pseudanthium. The individual flowers of a capitulum are called 137:
lacks disk flowers. The individual flowers of a pseudanthium in the family Asteraceae (or Compositae) are commonly called
69:, which are special types of inflorescences in which anything from a small cluster to hundreds or sometimes thousands of 376:
sp the pseudanthium is greatly condensed with staminate flowers surrounding a central terminal pistillate female flower.
17: 93:. Because the collection has the overall appearance of a single flower, the collection of flowers in the head of this 717: 1201:"Phylogenetic relationships in Cyperaceae subfamily Mapanioideae inferred from pollen and plastid DNA sequence data" 619:; every "petal" is actually a separate five-petaled flower complete with its own stamens and making its own fruit. 1391:
Claßen-Bockhoff, Regine; Ruonala, Raili; Bull-Hereñu, Kester; Marchant, Neville; Albert, Victor A. (2013-03-01).
839:"Flower-like terminal structures in racemose inflorescences: a tool in morphogenetic and evolutionary research" 260: 1199:
Simpson, David A.; Furness, Carol A.; Hodkinson, Trevor R.; Muasya, A. Muthama; Chase, Mark W. (July 2003).
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Diagram of a flower head. Note bracts surrounding the flowers, which would be absent on a capitulum.
437: 1039:"Floral development and evolution of capitulum structure in Anacyclus (Anthemideae, Asteraceae)" 397:
and less commonly petal-like structures. The central cyathia maybe composed of all male flowers.
42: 1550:"Monocot Pseudanthia Revisited: Floral Structure of the Mycoheterotrophic Family Triuridaceae" 1365: 1311: 1149: 1122: 1011: 707: 985: 913: 817: 680: 513: 615: 133: 85:
What appear to be "petals" of an individual flower, are actually each individual complete
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is an equivalent term for flower head and pseudanthium when used in the botanical sense.
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Wild Crop Relatives: Genomic and Breeding Resources: Plantation and Ornamental Crops
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Bello, M. Angélica; Álvarez, Ines; Torices, Rubén; Fuertes-Aguilar, Javier (2013).
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Progress in Botany: Structural Botany Physiology Genetics Taxonomy Geobotany
1526: 1452:"Ontogeny and Diversity in Staminate Flowers of Nothofagus (Nothofagaceae)" 1436: 1297: 1232: 1088: 872: 540: 90: 1409: 1216: 1390: 1263: 1054: 855: 535: 508: 379: 246:
does not link to any significant usage of the term in a botanical sense.
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Floral Diagrams: An Aid to Understanding Flower Morphology and Evolution
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The collection of independent organs into a complex structure is called
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Sokoloff, Dmitry; Rudall, Paula J.; Remizowa, Margarita (2006-10-01).
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like a single flower, at least in plants that are animal pollinated.
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Developmental Genetics of the Flower: Advances in Botanical Research
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The Molecular Genetics of Floral Transition and Flower Development
655:) with petaloid ray florets and tubular disc florets in the middle 482: 295: 749: 394: 238:) is a very rarely used term. It was defined in the 1966 book, 70: 1340:
Photographic Atlas of Botany and Guide to Plant Identification
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Pseudanthia are characteristic of the daisy and sunflower
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Plant Identification Terminology An illustrated Glossary
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from a polyaxial instead of a monoaxial configuration.
89:, and at the center is a dense pack of individual tiny 1370:. Springer Science & Business Media. p. 125. 1304: 836: 329:, perianth symmetry can be variable within the family. 990:. Springer Science & Business Media. p. 83. 889:
Second International Rubiaceae Conference Proceedings
366:— In subfamily Mapanioideae, pseudanthia are termed 1120: 892:. National Botanic Garden of Belgium. p. 330. 631:Discoid (having only disk flowers) flower heads of 595:
Flowers open in succession in head of a sunflower (
121:is fused into a tube. Flowers on the periphery are 812: 810: 737: 268:Pseudanthia occur in 40 plant families including: 1450:Rozefelds, Andrew C.; Drinnan, Andrew N. (1998). 1363: 1617: 1449: 1336: 1147: 761: 759: 30:"Flower head" redirects here. For the band, see 1116: 1114: 1112: 1110: 1108: 1106: 1104: 1102: 1100: 1098: 1016:. Elsevier Science. 16 June 2014. p. 299. 885: 807: 918:. Elsevier. 29 September 2006. pp. 35–. 756: 740:The genera of flowering plants (Angiospermae) 240:The genera of flowering plants (Angiospermae) 1316:. OUP Oxford. 14 January 2016. p. 176. 1245: 1095: 879: 731: 729: 679:Louis P. Ronse De Craene (4 February 2010). 674: 672: 670: 1330: 1176:"Cyperaceae - Evolution and classification" 709:The Arctic Guide: Wildlife of the Far North 699: 1141: 983: 785: 735: 685:. Cambridge University Press. p. 23. 464:in subgenus Lophozonia - a three‐flowered 27:Type of inflorescence, clusters of flowers 1426: 1408: 1287: 1078: 977: 854: 726: 667: 1357: 601:), with ray florets forming the 'petals' 436: 400: 348: 253: 80: 1554:International Journal of Plant Sciences 1456:International Journal of Plant Sciences 705: 14: 1618: 1547: 1364:Chittaranjan Kole (1 September 2011). 938: 1246:Prychid, C. J.; Bruhl, J. J. (2013). 1121:Rolf H. J. Schlegel (22 July 2020). 649:Flower head of creeping groundsel ( 46: 24: 178: 141:. The pseudanthium has a whorl of 25: 1642: 249: 1313:Plant Ecology in the Middle East 642: 624: 606: 588: 570: 558: 173: 1596: 1541: 1498: 1443: 1384: 1239: 1192: 1168: 1030: 1004: 957: 932: 769:Missouri Plants. Archived from 613:Close up of the ray corolla of 577:Flower head of a common daisy ( 984:Karl Esser (6 December 2012). 906: 843:Journal of Experimental Botany 830: 712:, Princeton University Press, 261:Spathiphyllum cochlearispathum 145:below the flowers, forming an 13: 1: 1343:. Feline Press. p. 145. 660: 1124:Dictionary of Plant Breeding 203: 45:for 'false flower'; 7: 1154:. UNSW Press. p. 407. 744:. Oxford: Clarendon Press. 222: 10: 1647: 1205:American Journal of Botany 1127:. CRC Press. p. 459. 818:"calflora Botanical Terms" 551: 405:Euphorbia caput-medusae 01 353:compressed pseudanthia of 164: 29: 1548:Rudall, Paula J. (2003). 1519:10.1007/s10265-018-1029-6 1337:James L. Castner (2004). 1148:Gwen Jean Harden (1990). 736:Hutchinson, John (1964). 387:— pseudanthia are called 298:— pseudanthia are called 287:— pseudanthia are called 1151:Flora of New South Wales 886:Elmar Robbrecht (1996). 706:Chester, Sharon (2016), 1180:Encyclopedia Britannica 965:"Australian bryophytes" 442:Actinodium cunninghamii 192: 939:Harris, James (2001). 793:"Taraxacum Officinale" 445: 406: 359: 265: 131:lacks ray flowers and 102: 1410:10.1186/2041-9139-4-8 1217:10.3732/ajb.90.7.1071 767:"Senecio vulgaris L." 440: 404: 352: 257: 84: 637:(rubber rabbitbrush) 616:Hieracium lachenalii 356:Lepironia articulata 134:Taraxacum officinale 634:Ericameria nauseosa 327:sexual reproduction 1264:10.1093/aob/mct111 1055:10.1093/aob/mcs301 856:10.1093/jxb/erl126 446: 407: 360: 266: 103: 18:Capitulum (flower) 1560:(S5): S307–S320. 1377:978-3-642-21201-7 1350:978-0-9625150-0-2 1323:978-0-19-107873-6 1161:978-0-86840-188-1 1134:978-1-00-006698-2 1023:978-0-12-417181-7 997:978-3-642-78020-2 950:978-0-9640221-6-4 925:978-0-08-046463-3 899:978-90-72619-29-7 849:(13): 3517–3530. 692:978-1-139-48455-8 652:Senecio angulatus 598:Helianthus annuus 468:without branches. 63:composite flowers 16:(Redirected from 1638: 1626:Plant morphology 1610: 1600: 1594: 1593: 1545: 1539: 1538: 1502: 1496: 1495: 1447: 1441: 1440: 1430: 1412: 1388: 1382: 1381: 1361: 1355: 1354: 1334: 1328: 1327: 1308: 1302: 1301: 1291: 1252:Annals of Botany 1243: 1237: 1236: 1211:(7): 1071–1086. 1196: 1190: 1189: 1187: 1186: 1172: 1166: 1165: 1145: 1139: 1138: 1118: 1093: 1092: 1082: 1049:(8): 1597–1612. 1043:Annals of Botany 1034: 1028: 1027: 1008: 1002: 1001: 981: 975: 974: 972: 971: 961: 955: 954: 936: 930: 929: 910: 904: 903: 883: 877: 876: 858: 834: 828: 827: 825: 824: 814: 805: 804: 802: 800: 789: 783: 782: 780: 778: 763: 754: 753: 743: 733: 724: 723: 703: 697: 696: 676: 646: 628: 610: 592: 574: 562: 338:Centrolepidaceae 128:Senecio vulgaris 48: 21: 1646: 1645: 1641: 1640: 1639: 1637: 1636: 1635: 1616: 1615: 1614: 1613: 1601: 1597: 1546: 1542: 1503: 1499: 1448: 1444: 1389: 1385: 1378: 1362: 1358: 1351: 1335: 1331: 1324: 1310: 1309: 1305: 1244: 1240: 1197: 1193: 1184: 1182: 1174: 1173: 1169: 1162: 1146: 1142: 1135: 1119: 1096: 1035: 1031: 1024: 1010: 1009: 1005: 998: 982: 978: 969: 967: 963: 962: 958: 951: 943:. Spring Lake. 937: 933: 926: 912: 911: 907: 900: 884: 880: 835: 831: 822: 820: 816: 815: 808: 798: 796: 791: 790: 786: 776: 774: 773:on 27 June 2012 765: 764: 757: 734: 727: 720: 704: 700: 693: 677: 668: 663: 656: 647: 638: 629: 620: 611: 602: 593: 584: 580:Bellis perennis 575: 566: 563: 554: 252: 225: 206: 195: 186:synorganization 181: 179:Synorganization 176: 167: 101:or a composite. 35: 28: 23: 22: 15: 12: 11: 5: 1644: 1634: 1633: 1628: 1612: 1611: 1595: 1582:10.1086/376879 1566:10.1086/376879 1540: 1513:(3): 443–458. 1497: 1484:10.1086/314090 1468:10.1086/314090 1462:(6): 906–922. 1442: 1383: 1376: 1356: 1349: 1329: 1322: 1303: 1258:(1): 161–177. 1238: 1191: 1167: 1160: 1140: 1133: 1094: 1029: 1022: 1003: 996: 976: 956: 949: 931: 924: 905: 898: 878: 829: 806: 795:. Florida Data 784: 755: 725: 718: 698: 691: 665: 664: 662: 659: 658: 657: 648: 641: 639: 630: 623: 621: 612: 605: 603: 594: 587: 585: 576: 569: 567: 564: 557: 553: 550: 549: 548: 546:Phyllanthaceae 543: 538: 533: 528: 518: 517: 506: 501: 496: 488:Pontederiaceae 485: 480: 469: 459: 435: 434: 429: 427:Marcgraviaceae 424: 416:Hamamelidaceae 413: 399: 398: 382: 377: 347: 346: 341: 335: 330: 304: 293: 282: 264:) pseudanthium 251: 250:Plant families 248: 244:Google Scholar 224: 221: 205: 202: 194: 191: 180: 177: 175: 172: 166: 163: 26: 9: 6: 4: 3: 2: 1643: 1632: 1629: 1627: 1624: 1623: 1621: 1608: 1605: 1604:Kew Bulletin. 1599: 1591: 1587: 1583: 1579: 1575: 1571: 1567: 1563: 1559: 1555: 1551: 1544: 1536: 1532: 1528: 1524: 1520: 1516: 1512: 1508: 1501: 1493: 1489: 1485: 1481: 1477: 1473: 1469: 1465: 1461: 1457: 1453: 1446: 1438: 1434: 1429: 1424: 1420: 1416: 1411: 1406: 1402: 1398: 1394: 1387: 1379: 1373: 1369: 1368: 1360: 1352: 1346: 1342: 1341: 1333: 1325: 1319: 1315: 1314: 1307: 1299: 1295: 1290: 1285: 1281: 1277: 1273: 1269: 1265: 1261: 1257: 1253: 1249: 1242: 1234: 1230: 1226: 1222: 1218: 1214: 1210: 1206: 1202: 1195: 1181: 1177: 1171: 1163: 1157: 1153: 1152: 1144: 1136: 1130: 1126: 1125: 1117: 1115: 1113: 1111: 1109: 1107: 1105: 1103: 1101: 1099: 1090: 1086: 1081: 1076: 1072: 1068: 1064: 1060: 1056: 1052: 1048: 1044: 1040: 1033: 1025: 1019: 1015: 1014: 1007: 999: 993: 989: 988: 980: 966: 960: 952: 946: 942: 935: 927: 921: 917: 916: 909: 901: 895: 891: 890: 882: 874: 870: 866: 862: 857: 852: 848: 844: 840: 833: 819: 813: 811: 794: 788: 772: 768: 762: 760: 751: 747: 742: 741: 732: 730: 721: 719:9781400865963 715: 711: 710: 702: 694: 688: 684: 683: 675: 673: 671: 666: 654: 653: 645: 640: 636: 635: 627: 622: 618: 617: 609: 604: 600: 599: 591: 586: 582: 581: 573: 568: 561: 556: 555: 547: 544: 542: 539: 537: 534: 532: 529: 527: 526:Hydatellaceae 524: 523: 522: 516: 515: 510: 507: 505: 502: 500: 497: 495: 494: 489: 486: 484: 481: 479: 478: 473: 470: 467: 463: 462:Nothofagaceae 460: 457: 456: 451: 448: 447: 443: 439: 433: 430: 428: 425: 423: 422: 417: 414: 412: 411:Eriocaulaceae 409: 408: 403: 396: 392: 391: 386: 385:Euphorbiaceae 383: 381: 378: 375: 374: 369: 365: 362: 361: 358: 357: 351: 345: 342: 339: 336: 334: 333:Campanulaceae 331: 328: 324: 320: 316: 312: 308: 305: 303: 302: 297: 294: 292: 291: 286: 283: 280: 279: 274: 271: 270: 269: 263: 262: 256: 247: 245: 241: 237: 233: 229: 220: 218: 214: 210: 201: 199: 190: 188: 187: 174:Related terms 171: 162: 158: 156: 150: 148: 144: 140: 136: 135: 130: 129: 124: 120: 116: 115:actinomorphic 112: 108: 100: 96: 92: 88: 83: 79: 77: 72: 68: 64: 60: 56: 55:inflorescence 52: 44: 43:Ancient Greek 40: 33: 19: 1606: 1603: 1598: 1557: 1553: 1543: 1510: 1506: 1500: 1459: 1455: 1445: 1400: 1396: 1386: 1366: 1359: 1339: 1332: 1312: 1306: 1255: 1251: 1241: 1208: 1204: 1194: 1183:. Retrieved 1179: 1170: 1150: 1143: 1123: 1046: 1042: 1032: 1012: 1006: 986: 979: 968:. Retrieved 959: 940: 934: 914: 908: 888: 881: 846: 842: 832: 821:. Retrieved 797:. Retrieved 787: 775:. Retrieved 771:the original 739: 708: 701: 681: 650: 632: 614: 596: 578: 541:Triuridaceae 519: 512: 491: 475: 453: 441: 419: 388: 371: 367: 354: 322: 319:flower heads 318: 314: 310: 299: 288: 276: 267: 259: 258:Peace Lily ( 239: 235: 231: 227: 226: 212: 208: 207: 197: 196: 184: 182: 168: 159: 151: 138: 132: 126: 104: 99:pseudanthium 98: 97:is called a 91:disc flowers 66: 62: 59:flower heads 58: 50: 39:pseudanthium 38: 36: 1507:J Plant Res 536:Pandanaceae 509:Saururaceae 444:pseudanthia 380:Dipsacaceae 155:pollination 123:zygomorphic 87:ray flowers 51:pseudanthia 1620:Categories 1185:2021-02-14 970:2012-02-26 823:2012-02-26 799:2 December 777:2 December 661:References 499:Proteaceae 493:Hydrothrix 455:Actinodium 364:Cyperaceae 313:(singular 307:Asteraceae 275:— in some 236:calathidia 111:Asteraceae 32:Flowerhead 1574:1058-5893 1476:1058-5893 1419:2041-9139 1272:0305-7364 1225:0002-9122 1063:0305-7364 865:0022-0957 531:Lemnaceae 514:Anemopsis 504:Rubiaceae 472:Nyssaceae 466:dichasium 450:Myrtaceae 421:Rhodoleia 395:nectaries 373:Lepironia 344:Cornaceae 315:capitulum 273:Adoxaceae 232:calathids 209:Capitulum 204:Capitulum 147:involucre 95:sunflower 76:sunflower 1590:85115689 1527:29569169 1492:83956542 1437:23448118 1403:(1): 8. 1298:23723258 1280:42801396 1233:21659207 1089:23287557 1071:42801622 873:17005921 750:65000676 432:Moraceae 368:spicoids 311:capitula 301:spadices 285:Apiaceae 278:Viburnum 230:(plural 228:Calathid 223:Calathid 217:sphagnum 213:capitula 211:(plural 117:and the 67:capitula 53:) is an 1631:Flowers 1609:(1):40. 1535:4202581 1428:3610234 1397:EvoDevo 1289:3690996 1080:3828941 552:Gallery 483:Poaceae 477:Davidia 390:cyathia 323:florets 296:Araceae 219:plant. 165:History 139:florets 119:corolla 71:flowers 1588:  1580:  1572:  1533:  1525:  1490:  1482:  1474:  1435:  1425:  1417:  1374:  1347:  1320:  1296:  1286:  1278:  1270:  1231:  1223:  1158:  1131:  1087:  1077:  1069:  1061:  1020:  994:  947:  922:  896:  871:  863:  748:  716:  689:  309:— The 290:umbels 143:bracts 107:family 1586:S2CID 1578:JSTOR 1531:S2CID 1488:S2CID 1480:JSTOR 1276:JSTOR 1067:JSTOR 511:— in 490:— in 474:— in 452:— in 418:— in 370:. In 317:) or 65:, or 1570:ISSN 1523:PMID 1472:ISSN 1433:PMID 1415:ISSN 1372:ISBN 1345:ISBN 1318:ISBN 1294:PMID 1268:ISSN 1229:PMID 1221:ISSN 1156:ISBN 1129:ISBN 1085:PMID 1059:ISSN 1018:ISBN 992:ISBN 945:ISBN 920:ISBN 894:ISBN 869:PMID 861:ISSN 801:2012 779:2012 746:LCCN 714:ISBN 687:ISBN 281:spp. 198:Head 193:Head 78:). 1562:doi 1558:164 1515:doi 1511:131 1464:doi 1460:159 1423:PMC 1405:doi 1284:PMC 1260:doi 1256:112 1213:doi 1075:PMC 1051:doi 1047:112 851:doi 234:or 153:in 47:pl. 1622:: 1607:61 1584:. 1576:. 1568:. 1556:. 1552:. 1529:. 1521:. 1509:. 1486:. 1478:. 1470:. 1458:. 1454:. 1431:. 1421:. 1413:. 1399:. 1395:. 1292:. 1282:. 1274:. 1266:. 1254:. 1250:. 1227:. 1219:. 1209:90 1207:. 1203:. 1178:. 1097:^ 1083:. 1073:. 1065:. 1057:. 1045:. 1041:. 867:. 859:. 847:57 845:. 841:. 809:^ 758:^ 728:^ 669:^ 189:. 149:. 61:, 49:: 37:A 1592:. 1564:: 1537:. 1517:: 1494:. 1466:: 1439:. 1407:: 1401:4 1380:. 1353:. 1326:. 1300:. 1262:: 1235:. 1215:: 1188:. 1164:. 1137:. 1091:. 1053:: 1026:. 1000:. 973:. 953:. 928:. 902:. 875:. 853:: 826:. 803:. 781:. 752:. 722:. 695:. 583:) 109:( 41:( 34:. 20:)

Index

Capitulum (flower)
Flowerhead
Ancient Greek
inflorescence
flowers
sunflower

ray flowers
disc flowers
sunflower
family
Asteraceae
actinomorphic
corolla
zygomorphic
Senecio vulgaris
Taraxacum officinale
bracts
involucre
pollination
synorganization
sphagnum
Google Scholar

Spathiphyllum cochlearispathum
Adoxaceae
Viburnum
Apiaceae
umbels
Araceae

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