1935:
108:
1357:
404:
Marivaux, L.; Welcomme, J.-L.; Antoine, P.-O.; Metais, G.; Baloch, I. M.; Benammi, M.; Chaimanee, Y.; Ducrocq, S.; Jaeger, J.-J. (19 October 2001). "A fossil lemur from the
Oligocene of Pakistan".
768:
297:
Described from a few teeth, the specimen possesses a lower canine that, according to
Marivaux et al., confirms the presence of the strepsirrhine-specific
325:
was confirmed in a 2016 cladistic analysis that recovered it in the family
Ekgmowechashalidae, which is more consistent with the lemur fossil record.
1588:
1503:
1415:
1048:
791:
1013:
1558:
1521:
1493:
1234:
870:
321:
has been questioned, as well as many other dental features, suggesting that it is most likely an adapiform. The adapiform nature of
2018:
602:
317:
Malagasy genus and its toothcomb was shorter and broader. More recently, the structure and general presence of the toothcomb in
2087:
468:
2082:
2072:
460:
2092:
354:
301:. Furthermore, on the basis of cheektooth morphology, the molars share strong affinities with those of the genus
1934:
294:, complicating the picture of the early evolution of lemurs by suggesting that lemurs originated in Asia.
595:
107:
2077:
376:"Earliest Evidence Of Lemurs Discovered In Pakistan, Far From Their Current Home, Science Reports"
1728:
375:
2049:
1973:
1435:
588:
2044:
2036:
529:"Oligocene primates from China reveal divergence between African and Asian primate evolution"
229:
271:
belonging to the adapiform family
Ekgmowechashalidae.It is represented by only one species,
2005:
1483:
1449:
540:
413:
276:
1356:
8:
544:
417:
509:
437:
102:
2031:
1915:
1790:
1756:
1749:
566:
558:
501:
464:
429:
513:
457:
Lemurs: Ecology and
Adaptation (Developments in Primatology: Progress and Prospects)
441:
1602:
548:
493:
421:
1956:
1908:
1511:
1442:
1155:
798:
784:
1719:
1697:
1677:
1613:
1476:
1459:
1286:
1265:
1227:
986:
686:
403:
386:. American Association For The Advancement Of Science: 587–591. 22 October 2001
291:
182:
2066:
1996:
1894:
1837:
1578:
1185:
1100:
1059:
1000:
891:
863:
819:
805:
669:
656:
611:
562:
169:
61:
553:
528:
425:
1857:
1830:
1821:
1807:
1783:
1374:
1328:
1027:
1020:
856:
733:
710:
570:
505:
484:
Godinot, M. (2006). "Lemuriform origins as viewed from the fossil record".
433:
314:
265:
1951:
1885:
1742:
1735:
1687:
1636:
1548:
1199:
1137:
1123:
1068:
1041:
970:
956:
949:
928:
898:
884:
754:
696:
306:
36:
2023:
1800:
1776:
1622:
1595:
1387:
1317:
1295:
1220:
1178:
1130:
1109:
942:
935:
835:
740:
81:
46:
527:
Ni, Xijun; Li, Qiang; Li, Lüzhou; Beard, K. Christopher (2016-05-06).
497:
1901:
1848:
1767:
1532:
1404:
1302:
1256:
1241:
1213:
1206:
1192:
1171:
1164:
1116:
1082:
979:
747:
726:
334:
298:
119:
86:
1967:
1990:
1708:
1469:
1310:
1089:
912:
877:
842:
828:
629:
280:
139:
76:
71:
56:
51:
41:
1670:
1272:
1075:
1034:
849:
719:
641:
268:
159:
91:
66:
2010:
580:
1871:
1426:
921:
635:
623:
454:
149:
129:
1864:
1656:
1629:
1569:
1882:
1818:
1796:
1704:
1693:
1683:
1584:
1554:
1544:
1517:
1499:
1489:
1465:
1455:
1411:
1400:
1283:
1253:
1152:
997:
967:
909:
816:
765:
707:
692:
682:
2064:
455:Gould, Lisa; Sauther, Michelle L., eds. (2006).
596:
667:
448:
526:
368:
603:
589:
477:
552:
483:
397:
347:
2065:
1972:
1971:
1372:
654:
584:
313:appears to be much smaller than the
290:was classified in the lemur family
13:
14:
2104:
655:
610:
1933:
1373:
1355:
106:
34:
520:
1:
2088:Fossil taxa described in 2001
340:
7:
357:. The Paleobiology Database
328:
10:
2109:
2083:Prehistoric primate genera
2073:Prehistoric strepsirrhines
1980:
1946:
1931:
1881:
1847:
1817:
1766:
1718:
1665:
1654:
1612:
1568:
1540:
1531:
1425:
1396:
1385:
1381:
1368:
1353:
1327:
1282:
1252:
1151:
1099:
1058:
1008:
996:
966:
908:
815:
764:
706:
678:
663:
650:
618:
275:, which was found in the
235:
228:
103:Scientific classification
101:
23:
16:Extinct genus of primates
554:10.1126/science.aaf2107
426:10.1126/science.1065257
264:is an extinct genus of
2093:Eocene mammals of Asia
286:When first described,
2045:Paleobiology Database
1982:Bugtilemur mathesoni
277:Chitarwata Formation
242:Bugtilemur mathesoni
1886:Palaeopropithecidae
545:2016Sci...352..673N
486:Folia Primatologica
418:2001Sci...294..587M
769:Ekgmowechashalidae
32:Early Oligocene
2060:
2059:
2032:Open Tree of Life
1974:Taxon identifiers
1965:
1964:
1942:
1941:
1929:
1928:
1925:
1924:
1916:Palaeopropithecus
1650:
1649:
1646:
1645:
1364:
1363:
1351:
1350:
1347:
1346:
1147:
1146:
539:(6286): 673–677.
498:10.1159/000095391
470:978-0-387-34585-7
412:(5542): 587–591.
257:
256:
221:B. mathesoni
209:
2100:
2053:
2052:
2040:
2039:
2027:
2026:
2014:
2013:
2001:
2000:
1999:
1969:
1968:
1937:
1884:
1822:Archaeolemuridae
1820:
1798:
1706:
1695:
1685:
1663:
1662:
1603:Xanthonycticebus
1586:
1556:
1546:
1538:
1537:
1519:
1501:
1491:
1467:
1457:
1413:
1402:
1394:
1393:
1383:
1382:
1370:
1369:
1359:
1285:
1255:
1154:
1006:
1005:
999:
969:
911:
818:
767:
709:
694:
684:
676:
675:
665:
664:
652:
651:
605:
598:
591:
582:
581:
575:
574:
556:
524:
518:
517:
481:
475:
474:
463:. pp. 8–9.
452:
446:
445:
401:
395:
394:
392:
391:
372:
366:
365:
363:
362:
351:
244:
240:
203:
196:
181:
111:
110:
96:
33:
29:Temporal range:
21:
20:
2108:
2107:
2103:
2102:
2101:
2099:
2098:
2097:
2078:Eocene primates
2063:
2062:
2061:
2056:
2048:
2043:
2035:
2030:
2022:
2017:
2009:
2004:
1995:
1994:
1989:
1976:
1966:
1961:
1957:Subfossil lemur
1938:
1921:
1909:Mesopropithecus
1877:
1843:
1813:
1762:
1714:
1659:
1642:
1608:
1564:
1527:
1421:
1390:
1377:
1360:
1343:
1323:
1278:
1248:
1156:Caenopithecidae
1143:
1095:
1054:
992:
962:
904:
811:
799:Muangthanhinius
785:Ekgmowechashala
760:
702:
672:
659:
646:
614:
609:
579:
578:
525:
521:
482:
478:
471:
453:
449:
402:
398:
389:
387:
374:
373:
369:
360:
358:
353:
352:
348:
343:
331:
253:
246:
238:
237:
224:
202:
194:
179:
105:
97:
95:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
31:
30:
27:
17:
12:
11:
5:
2106:
2096:
2095:
2090:
2085:
2080:
2075:
2058:
2057:
2055:
2054:
2041:
2028:
2015:
2002:
1986:
1984:
1978:
1977:
1963:
1962:
1960:
1959:
1954:
1947:
1944:
1943:
1940:
1939:
1932:
1930:
1927:
1926:
1923:
1922:
1920:
1919:
1912:
1905:
1898:
1890:
1888:
1879:
1878:
1876:
1875:
1868:
1861:
1853:
1851:
1845:
1844:
1842:
1841:
1834:
1826:
1824:
1815:
1814:
1812:
1811:
1804:
1794:
1787:
1780:
1772:
1770:
1764:
1763:
1761:
1760:
1753:
1746:
1739:
1732:
1724:
1722:
1720:Cheirogaleidae
1716:
1715:
1713:
1712:
1702:
1698:Plesiopithecus
1691:
1681:
1674:
1666:
1660:
1655:
1652:
1651:
1648:
1647:
1644:
1643:
1641:
1640:
1633:
1626:
1618:
1616:
1614:Perodicticinae
1610:
1609:
1607:
1606:
1599:
1592:
1582:
1574:
1572:
1566:
1565:
1563:
1562:
1552:
1541:
1535:
1529:
1528:
1526:
1525:
1515:
1508:
1497:
1487:
1480:
1473:
1463:
1453:
1446:
1439:
1431:
1429:
1423:
1422:
1420:
1419:
1409:
1397:
1391:
1386:
1379:
1378:
1366:
1365:
1362:
1361:
1354:
1352:
1349:
1348:
1345:
1344:
1342:
1341:
1340:
1339:
1333:
1331:
1325:
1324:
1322:
1321:
1314:
1307:
1299:
1291:
1289:
1287:Djebelemuridae
1280:
1279:
1277:
1276:
1269:
1266:Algeripithecus
1261:
1259:
1250:
1249:
1247:
1246:
1238:
1231:
1228:Mescalerolemur
1224:
1217:
1210:
1203:
1196:
1189:
1182:
1175:
1168:
1160:
1158:
1149:
1148:
1145:
1144:
1142:
1141:
1134:
1127:
1120:
1113:
1105:
1103:
1097:
1096:
1094:
1093:
1086:
1079:
1072:
1064:
1062:
1056:
1055:
1053:
1052:
1045:
1038:
1031:
1024:
1017:
1009:
1003:
994:
993:
991:
990:
987:Marcgodinotius
983:
975:
973:
964:
963:
961:
960:
953:
946:
939:
932:
925:
917:
915:
906:
905:
903:
902:
895:
888:
881:
874:
867:
860:
853:
846:
839:
832:
824:
822:
813:
812:
810:
809:
802:
795:
788:
781:
773:
771:
762:
761:
759:
758:
751:
744:
737:
730:
723:
715:
713:
704:
703:
701:
700:
690:
687:Plesiopithecus
679:
673:
668:
661:
660:
648:
647:
645:
644:
638:
632:
626:
619:
616:
615:
608:
607:
600:
593:
585:
577:
576:
519:
492:(6): 446–464.
476:
469:
459:(1 ed.).
447:
396:
367:
345:
344:
342:
339:
338:
337:
330:
327:
292:Cheirogaleidae
255:
254:
247:
233:
232:
226:
225:
217:
215:
211:
210:
192:
188:
187:
184:incertae sedis
177:
173:
172:
167:
163:
162:
157:
153:
152:
147:
143:
142:
137:
133:
132:
127:
123:
122:
117:
113:
112:
99:
98:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
35:
28:
15:
9:
6:
4:
3:
2:
2105:
2094:
2091:
2089:
2086:
2084:
2081:
2079:
2076:
2074:
2071:
2070:
2068:
2051:
2046:
2042:
2038:
2033:
2029:
2025:
2020:
2016:
2012:
2007:
2003:
1998:
1992:
1988:
1987:
1985:
1983:
1979:
1975:
1970:
1958:
1955:
1953:
1949:
1948:
1945:
1936:
1918:
1917:
1913:
1911:
1910:
1906:
1904:
1903:
1899:
1897:
1896:
1895:Archaeoindris
1892:
1891:
1889:
1887:
1880:
1874:
1873:
1869:
1867:
1866:
1862:
1860:
1859:
1855:
1854:
1852:
1850:
1846:
1840:
1839:
1838:Hadropithecus
1835:
1833:
1832:
1828:
1827:
1825:
1823:
1816:
1810:
1809:
1805:
1803:
1802:
1795:
1793:
1792:
1788:
1786:
1785:
1781:
1779:
1778:
1774:
1773:
1771:
1769:
1765:
1759:
1758:
1754:
1752:
1751:
1747:
1745:
1744:
1740:
1738:
1737:
1733:
1731:
1730:
1726:
1725:
1723:
1721:
1717:
1711:
1710:
1703:
1700:
1699:
1692:
1690:
1689:
1682:
1680:
1679:
1675:
1673:
1672:
1668:
1667:
1664:
1661:
1658:
1653:
1639:
1638:
1634:
1632:
1631:
1627:
1625:
1624:
1620:
1619:
1617:
1615:
1611:
1605:
1604:
1600:
1598:
1597:
1593:
1591:
1590:
1589:Nycticeboides
1583:
1581:
1580:
1576:
1575:
1573:
1571:
1567:
1561:
1560:
1553:
1551:
1550:
1543:
1542:
1539:
1536:
1534:
1530:
1524:
1523:
1516:
1514:
1513:
1512:Sciurocheirus
1509:
1506:
1505:
1498:
1496:
1495:
1488:
1486:
1485:
1481:
1479:
1478:
1474:
1472:
1471:
1464:
1462:
1461:
1454:
1452:
1451:
1447:
1445:
1444:
1440:
1438:
1437:
1433:
1432:
1430:
1428:
1424:
1418:
1417:
1410:
1407:
1406:
1399:
1398:
1395:
1392:
1389:
1384:
1380:
1376:
1371:
1367:
1358:
1337:
1336:
1335:
1334:
1332:
1330:
1326:
1320:
1319:
1315:
1313:
1312:
1308:
1305:
1304:
1300:
1298:
1297:
1293:
1292:
1290:
1288:
1281:
1275:
1274:
1270:
1268:
1267:
1263:
1262:
1260:
1258:
1251:
1244:
1243:
1239:
1237:
1236:
1232:
1230:
1229:
1225:
1223:
1222:
1218:
1216:
1215:
1211:
1209:
1208:
1204:
1202:
1201:
1197:
1195:
1194:
1190:
1188:
1187:
1186:Caenopithecus
1183:
1181:
1180:
1176:
1174:
1173:
1169:
1167:
1166:
1162:
1161:
1159:
1157:
1150:
1140:
1139:
1135:
1133:
1132:
1128:
1126:
1125:
1121:
1119:
1118:
1114:
1112:
1111:
1107:
1106:
1104:
1102:
1101:Sivaladapinae
1098:
1092:
1091:
1087:
1085:
1084:
1080:
1078:
1077:
1073:
1071:
1070:
1066:
1065:
1063:
1061:
1060:Hoanghoniinae
1057:
1051:
1050:
1046:
1044:
1043:
1039:
1037:
1036:
1032:
1030:
1029:
1025:
1023:
1022:
1018:
1016:
1015:
1011:
1010:
1007:
1004:
1002:
1001:Sivaladapidae
995:
989:
988:
984:
982:
981:
977:
976:
974:
972:
965:
959:
958:
954:
952:
951:
947:
945:
944:
940:
938:
937:
933:
931:
930:
926:
924:
923:
919:
918:
916:
914:
907:
901:
900:
896:
894:
893:
892:Pronycticebus
889:
887:
886:
882:
880:
879:
875:
873:
872:
868:
866:
865:
864:Mazateronodon
861:
859:
858:
854:
852:
851:
847:
845:
844:
840:
838:
837:
833:
831:
830:
826:
825:
823:
821:
820:Cercamoniidae
814:
808:
807:
806:Palaeohodites
803:
801:
800:
796:
794:
793:
789:
787:
786:
782:
780:
779:
775:
774:
772:
770:
763:
757:
756:
752:
750:
749:
745:
743:
742:
738:
736:
735:
731:
729:
728:
724:
722:
721:
717:
716:
714:
712:
705:
699:
698:
691:
689:
688:
681:
680:
677:
674:
671:
670:Strepsirrhini
666:
662:
658:
657:Strepsirrhini
653:
649:
643:
639:
637:
633:
631:
627:
625:
621:
620:
617:
613:
612:Strepsirrhini
606:
601:
599:
594:
592:
587:
586:
583:
572:
568:
564:
560:
555:
550:
546:
542:
538:
534:
530:
523:
515:
511:
507:
503:
499:
495:
491:
487:
480:
472:
466:
462:
458:
451:
443:
439:
435:
431:
427:
423:
419:
415:
411:
407:
400:
385:
381:
377:
371:
356:
350:
346:
336:
333:
332:
326:
324:
320:
316:
312:
308:
304:
300:
295:
293:
289:
284:
282:
278:
274:
270:
267:
263:
262:
251:
245:
243:
234:
231:
230:Binomial name
227:
223:
222:
216:
213:
212:
207:
201:
200:
193:
190:
189:
186:
185:
178:
175:
174:
171:
170:Strepsirrhini
168:
165:
164:
161:
158:
155:
154:
151:
148:
145:
144:
141:
138:
135:
134:
131:
128:
125:
124:
121:
118:
115:
114:
109:
104:
100:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
1981:
1914:
1907:
1900:
1893:
1870:
1863:
1856:
1836:
1831:Archaeolemur
1829:
1806:
1799:
1789:
1782:
1775:
1755:
1748:
1741:
1736:Cheirogaleus
1734:
1727:
1707:
1696:
1686:
1676:
1669:
1635:
1630:Perodicticus
1628:
1621:
1601:
1594:
1587:
1577:
1557:
1547:
1520:
1510:
1504:Saharagalago
1502:
1492:
1482:
1475:
1468:
1458:
1448:
1441:
1434:
1416:Saharagalago
1414:
1403:
1375:Lemuriformes
1329:Lemuriformes
1316:
1309:
1301:
1294:
1271:
1264:
1240:
1233:
1226:
1219:
1212:
1205:
1198:
1191:
1184:
1177:
1170:
1163:
1136:
1129:
1122:
1115:
1108:
1088:
1081:
1074:
1067:
1049:Yunnanadapis
1047:
1040:
1033:
1028:Kyitchaungia
1026:
1021:Guangxilemur
1019:
1012:
985:
978:
955:
948:
941:
934:
927:
920:
897:
890:
883:
876:
869:
862:
857:Donrussellia
855:
848:
841:
834:
827:
804:
797:
792:Gatanthropus
790:
783:
777:
776:
753:
746:
739:
734:Hesperolemur
732:
725:
718:
711:Notharctidae
695:
685:
536:
532:
522:
489:
485:
479:
456:
450:
409:
405:
399:
388:. Retrieved
383:
380:ScienceDaily
379:
370:
359:. Retrieved
355:"Bugtilemur"
349:
322:
318:
310:
309:). However,
307:dwarf lemurs
303:Cheirogaleus
302:
296:
287:
285:
273:B. mathesoni
272:
266:Strepsirhine
260:
259:
258:
249:
241:
236:
220:
219:
205:
198:
197:
183:
24:
18:
1952:Adapiformes
1872:Propithecus
1688:Megaladapis
1671:Daubentonia
1637:Pseudopotto
1549:Mioeuoticus
1200:Europolemur
1138:Sivaladapis
1124:Siamoadapis
1069:Hoanghonius
1042:Paukkaungia
1014:Anthradapis
971:Asiadapidae
957:Palaeolemur
950:Microadapis
929:Cryptadapis
899:Protoadapis
885:Periconodon
755:Smilodectes
697:Sulaimanius
2067:Categories
1950:See also:
1801:Pachylemur
1743:Microcebus
1657:Lemuroidea
1623:Arctocebus
1596:Nycticebus
1484:Paragalago
1450:Galagoides
1388:Lorisoidea
1338:see below↓
1318:Shizarodon
1296:Djebelemur
1221:Masradapis
1179:Aframonius
1131:Sinoadapis
1110:Indraloris
943:Magnadapis
936:Leptadapis
836:Anchomomys
778:Bugtilemur
741:Notharctus
390:2009-07-08
361:2009-07-08
341:References
323:Bugtilemur
319:Bugtilemur
311:Bugtilemur
288:Bugtilemur
261:Bugtilemur
199:Bugtilemur
166:Suborder:
25:Bugtilemur
1902:Babakotia
1849:Indriidae
1784:Hapalemur
1768:Lemuridae
1729:Allocebus
1678:Lepilemur
1570:Lorisinae
1559:Namaloris
1533:Lorisidae
1522:Wadilemur
1494:Progalago
1427:Galagidae
1405:Karanisia
1303:Notnamaia
1257:Azibiidae
1242:Notnamaia
1235:Namadapis
1214:Mahgarita
1207:Godinotia
1193:Darwinius
1172:Afradapis
1165:Adapoides
1117:Ramadapis
1083:Rencunius
980:Asiadapis
748:Pelycodus
727:Copelemur
622:Kingdom:
563:0036-8075
335:Karanisia
299:toothcomb
248:Marivaux
214:Species:
204:Marivaux
126:Kingdom:
120:Eukaryota
1991:Wikidata
1709:Propotto
1477:Otolemur
1470:Laetolia
1436:Euoticus
1311:Omanodon
1090:Wailekia
913:Adapidae
878:Panobius
871:Nievesia
843:Barnesia
829:Agerinia
642:Primates
636:Mammalia
630:Chordata
628:Phylum:
624:Animalia
571:27151861
514:24163044
506:17053330
461:Springer
442:10585152
434:11641497
329:See also
281:Pakistan
176:Family:
160:Primates
150:Mammalia
140:Chordata
136:Phylum:
130:Animalia
116:Domain:
2037:4117714
2024:4974648
2011:4454000
1997:Q926781
1808:Varecia
1777:Eulemur
1273:Azibius
1076:Lushius
1035:Laomaki
850:Buxella
720:Cantius
640:Order:
634:Class:
541:Bibcode
533:Science
414:Bibcode
406:Science
269:primate
252:., 2001
239:†
218:†
208:., 2001
191:Genus:
156:Order:
146:Class:
2050:131786
1757:Phaner
1443:Galago
922:Adapis
569:
561:
512:
504:
467:
440:
432:
315:extant
1865:Indri
1858:Avahi
1791:Lemur
1750:Mirza
1579:Loris
1460:Komba
510:S2CID
438:S2CID
250:et al
206:et al
2019:GBIF
567:PMID
559:ISSN
502:PMID
465:ISBN
430:PMID
37:PreꞒ
2006:EoL
549:doi
537:352
494:doi
422:doi
410:294
384:294
279:of
2069::
2047::
2034::
2021::
2008::
1993::
565:.
557:.
547:.
535:.
531:.
508:.
500:.
490:77
488:.
436:.
428:.
420:.
408:.
382:.
378:.
283:.
87:Pg
1883:†
1819:†
1797:†
1705:†
1701:?
1694:†
1684:†
1585:†
1555:†
1545:†
1518:†
1507:?
1500:†
1490:†
1466:†
1456:†
1412:†
1408:?
1401:†
1306:?
1284:†
1254:†
1245:?
1153:†
998:†
968:†
910:†
817:†
766:†
708:†
693:†
683:†
604:e
597:t
590:v
573:.
551::
543::
516:.
496::
473:.
444:.
424::
416::
393:.
364:.
305:(
195:†
180:†
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:Ꞓ
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