146:
102:
123:
696:
Timothy P. Topper; Christian B. Skovsted; Glenn A. Brock & John R. Paterson (2011). "The oldest bivalved arthropods from the early
Cambrian of East Gondwana: systematics, biostratigraphy and biogeography".
610:, eggs have been found preserved attached to the posteriormost three pairs of biramous limbs, suggesting it engaged in brood care. Around 50–80 eggs, each around 150–180 μm across were attached in total.
778:
Zhai, Dayou; Williams, Mark; Siveter, David J.; Harvey, Thomas H. P.; Sansom, Robert S.; Gabbott, Sarah E.; Siveter, Derek J.; Ma, Xiaoya; Zhou, Runqing; Liu, Yu; Hou, Xianguang (2019-09-03).
642:
fossils. Their taxonomic composition broadly reflects two geographical provinces ("European" and "4A", i.e. America, Asia, Australia, Antarctica) which approximately mirror
658:) and was relatively low along the Laurentian coast. Bradoriids severely declined in abundance from the late Middle Cambrian onwards, with only a handful of
565:
224:
259:
280:
885:
399:
311:
287:
976:
453:
439:
425:
364:
304:
357:
319:
268:
208:
446:
432:
325:
232:
385:
333:
294:
214:
646:
provinces, with the 4A area representing warmer waters closer to the palaeo equator. Bradoriid diversity was highest along the coasts of
371:
241:
606:
has 4 pairs of appendages on the head each with a distinct morphology, with 9 pairs of trunk appendages, the last being uniramous. In
248:
837:"Reproductive strategy of the bradoriid arthropod Kunmingella douvillei from the Lower Cambrian Chengjiang Lagerstätte, South China"
392:
544:
Most bradoriids are only known from their bivalved carapaces, which are small in size, typically up to around 5 millimetres (
835:
Duan, Yanhong; Han, Jian; Fu, Dongjing; Zhang, Xingliang; Yang, Xiaoguang; Komiya, Tsuyoshi; Shu, Degan (April 2014).
1017:
780:"Variation in appendages in early Cambrian bradoriids reveals a wide range of body plans in stem-euarthropods"
880:
881:"Biogeography and affinities of the bradoriid arthropods: Cosmopolitan microbenthos of the Cambrian seas"
602:, and a posterior 5 with only a single row of endites, as well as two terminal pairs of uniramous limbs.
517:
does not support such a relationship; neither are they related to the
Cambrian bivalved arthropod group
509:
Whilst the
Bradoriida were traditionally considered as relatives of the modern bivalved arthropod group
145:
1022:
695:
836:
994:
929:
989:
981:
967:
894:
848:
745:
706:
571:
8:
622:) or swimming close to the seafloor (nektobenthic). Bradoriids are proposed to have been
898:
879:
Williams, Mark; Siveter, David J.; Popov, Leonid E.; Vannier, Jean M. C. (14 May 2007).
852:
749:
710:
812:
779:
140:
114:
817:
799:
579:
902:
856:
807:
791:
753:
714:
498:
906:
518:
482:
758:
733:
952:
413:
795:
1011:
860:
803:
718:
635:
554: in) in length. Preserved soft tissues known from some members, such as
342:
59:
821:
594:
limbs, including four anterior pairs of appendages bearing double rows of
961:
647:
623:
560:
556:
529:
522:
406:
132:
34:
667:
525:
378:
79:
44:
618:
Bradoriids are thought to have lived either crawling on the seafloor (
655:
643:
639:
583:
533:
514:
486:
353:
Other genera (may fall into above orders or others; please confirm):
177:
157:
101:
84:
923:
946:
663:
651:
591:
510:
494:
493:, which globally distributed, forming a significant portion of the
490:
74:
69:
54:
49:
39:
634:
Bradoriida are geographically widespread, and first occur in the
619:
599:
590:
had 12 appendages, including a pair of antennae as well pairs of
479:
89:
64:
595:
167:
122:
659:
878:
734:"Bradoriids (Arthropoda) and the Cambrian Diversification"
777:
626:, scavengers, or micro predators of soft-bodied prey.
731:
886:Palaeogeography, Palaeoclimatology, Palaeoecology
586:appendages which were all similar to each other.
1009:
834:
691:
689:
687:
685:
683:
521:. They have been alternatively recovered as
121:
100:
811:
757:
680:
1010:
928:
927:
874:
872:
870:
773:
771:
769:
109:Stereographic pair of a 3D scan of
13:
14:
1034:
867:
766:
144:
32:
30:Cambrian stage 3–Mid Ordovician
828:
725:
539:
1:
732:Mark A. S. McMenamin (2020).
673:
629:
536:, depending on the analysis.
907:10.1016/j.palaeo.2006.12.004
638:shortly before the earliest
7:
759:10.3390/geosciences10040119
570:suggest that the group was
504:
16:Extinct order of arthropods
10:
1039:
613:
253:Jones & McKenzie, 1980
936:
796:10.1038/s42003-019-0573-5
662:continuing into the Late
501:soft-bodied communities.
205:
200:
141:Scientific classification
139:
129:
120:
108:
99:
23:
861:10.1016/j.gr.2013.03.011
719:10.1016/j.gr.2010.05.012
582:followed by 11 pairs of
513:, the anatomy of their
263:Sylvester-Bradley, 1961
1018:Prehistoric arthropods
784:Communications Biology
990:Paleobiology Database
201:Families and genera
130:Life restoration of
899:2007PPP...248..202W
853:2014GondR..25..983D
750:2020Geosc..10..119M
711:2011GondR..19..310T
225:Parahoulongdongella
115:Maotianshan Shales
1005:
1004:
930:Taxon identifiers
841:Gondwana Research
699:Gondwana Research
469:
468:
348:
338:
300:
264:
260:Hipponicharionida
254:
237:
220:
196:
1030:
1023:Arthropod orders
998:
997:
985:
984:
972:
971:
970:
957:
956:
955:
925:
924:
918:
917:
915:
913:
893:(1–2): 202–232.
876:
865:
864:
832:
826:
825:
815:
775:
764:
763:
761:
729:
723:
722:
693:
624:detritus feeders
553:
552:
548:
532:, or stem-group
528:, as stem-group
499:Early Ordovician
489:with a bivalved
485:of small marine
346:
336:
298:
281:Cambroarchilocus
262:
252:
235:
218:
194:
189:
149:
148:
125:
104:
94:
31:
27:Temporal range:
21:
20:
1038:
1037:
1033:
1032:
1031:
1029:
1028:
1027:
1008:
1007:
1006:
1001:
993:
988:
980:
975:
966:
965:
960:
951:
950:
945:
932:
922:
921:
911:
909:
877:
868:
833:
829:
776:
767:
730:
726:
694:
681:
676:
632:
616:
572:morphologically
550:
546:
545:
542:
519:Phosphatocopina
507:
193:
187:
143:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
1036:
1026:
1025:
1020:
1003:
1002:
1000:
999:
986:
973:
958:
942:
940:
934:
933:
920:
919:
866:
847:(3): 983–990.
827:
765:
724:
705:(1): 310–326.
678:
677:
675:
672:
631:
628:
615:
612:
578:had a pair of
541:
538:
506:
503:
474:, also called
467:
466:
465:
464:
460:
459:
458:
457:
450:
443:
436:
429:
422:
414:Mongolitubulus
410:
403:
396:
389:
382:
375:
368:
361:
351:
350:
349:
331:
330:
329:
317:
316:
315:
312:Neokunmingella
308:
301:
291:
288:Hipponicharion
284:
277:
257:
256:
255:
245:
230:
229:
228:
221:
203:
202:
198:
197:
185:
181:
180:
175:
171:
170:
165:
161:
160:
155:
151:
150:
137:
136:
127:
126:
118:
117:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
26:
15:
9:
6:
4:
3:
2:
1035:
1024:
1021:
1019:
1016:
1015:
1013:
996:
991:
987:
983:
978:
974:
969:
963:
959:
954:
948:
944:
943:
941:
939:
935:
931:
926:
908:
904:
900:
896:
892:
888:
887:
882:
875:
873:
871:
862:
858:
854:
850:
846:
842:
838:
831:
823:
819:
814:
809:
805:
801:
797:
793:
789:
785:
781:
774:
772:
770:
760:
755:
751:
747:
743:
739:
735:
728:
720:
716:
712:
708:
704:
700:
692:
690:
688:
686:
684:
679:
671:
669:
665:
661:
657:
653:
649:
645:
641:
637:
636:fossil record
627:
625:
621:
611:
609:
605:
601:
597:
593:
589:
585:
581:
577:
573:
569:
568:
563:
562:
558:
537:
535:
531:
527:
524:
520:
516:
512:
502:
500:
496:
492:
488:
484:
481:
477:
473:
463:and others...
462:
461:
456:
455:
454:Wimanicharion
451:
449:
448:
444:
442:
441:
440:Spinospitella
437:
435:
434:
430:
428:
427:
426:Onagrocharion
423:
420:
416:
415:
411:
409:
408:
404:
402:
401:
397:
395:
394:
390:
388:
387:
383:
381:
380:
376:
374:
373:
369:
367:
366:
365:Amphikeropsis
362:
360:
359:
355:
354:
352:
345:
344:
343:Liangshanella
340:
339:
335:
332:
328:
327:
323:
322:
321:
318:
314:
313:
309:
307:
306:
305:Meishucunella
302:
297:
296:
292:
290:
289:
285:
283:
282:
278:
275:
271:
270:
266:
265:
261:
258:
251:
250:
246:
244:
243:
239:
238:
234:
231:
227:
226:
222:
219:Matthew, 1886
217:
216:
212:
211:
210:
207:
206:
204:
199:
195:Raymond, 1935
192:
186:
183:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
147:
142:
138:
135:
134:
128:
124:
119:
116:
113:sp. from the
112:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
937:
910:. Retrieved
890:
884:
844:
840:
830:
787:
783:
741:
737:
727:
702:
698:
650:and eastern
633:
617:
607:
603:
587:
575:
566:
555:
543:
530:mandibulates
508:
475:
471:
470:
452:
445:
438:
431:
424:
418:
412:
405:
398:
391:
384:
377:
370:
363:
358:Albrunnicola
356:
341:
324:
320:Monasteriida
310:
303:
299:Topper, 2010
293:
286:
279:
273:
269:Albrunnicola
267:
247:
240:
223:
213:
209:Beryichonida
190:
131:
110:
18:
962:Wikispecies
912:14 November
738:Geosciences
648:South China
608:Kunmingella
604:Kunyangella
588:Kunmingella
561:Kunyangella
557:Kunmingella
540:Description
526:crustaceans
447:Walcottella
433:Shangsiella
407:Kunmingella
326:Epactridion
233:Compatulida
133:Kunmingella
1012:Categories
968:Bradoriida
938:Bradoriida
790:(1): 329.
744:(4): 119.
674:References
668:Ordovician
666:and Early
630:Occurrence
620:epibenthic
534:arthropods
523:stem-group
515:appendages
487:arthropods
476:bradoriids
472:Bradoriida
386:Dielymella
379:Comptaluta
337:Öpik, 1968
334:Svealutida
295:Quadricona
274:Beyrichona
236:Öpik, 1968
215:Beyrichona
191:Bradoriida
178:Arthropoda
24:Bradoriida
804:2399-3642
656:Australia
644:trilobite
640:trilobite
598:on their
584:uniramous
574:diverse.
511:Ostracoda
478:, are an
347:Huo, 1956
164:Kingdom:
158:Eukaryota
953:Q4955096
947:Wikidata
822:31508504
664:Cambrian
652:Gondwana
600:endopods
592:biramous
580:antennae
505:Affinity
495:Cambrian
491:carapace
419:Spinella
372:Bradoria
242:Altuella
174:Phylum:
168:Animalia
154:Domain:
895:Bibcode
849:Bibcode
813:6722085
746:Bibcode
707:Bibcode
614:Ecology
596:endites
576:Indiana
567:Indiana
549:⁄
480:extinct
400:Indiana
249:Zepaera
184:Order:
111:Indiana
820:
810:
802:
660:genera
995:92005
982:10239
977:IRMNG
483:order
393:Haoia
914:2022
818:PMID
800:ISSN
564:and
497:and
35:PreꞒ
903:doi
891:248
857:doi
808:PMC
792:doi
754:doi
715:doi
1014::
992::
979::
964::
949::
901:.
889:.
883:.
869:^
855:.
845:25
843:.
839:.
816:.
806:.
798:.
786:.
782:.
768:^
752:.
742:10
740:.
736:.
713:.
703:19
701:.
682:^
670:.
559:,
551:16
85:Pg
916:.
905::
897::
863:.
859::
851::
824:.
794::
788:2
762:.
756::
748::
721:.
717::
709::
654:(
547:3
421:)
417:(
276:)
272:(
188:†
90:N
80:K
75:J
70:T
65:P
60:C
55:D
50:S
45:O
40:Ꞓ
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