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Asplenium arcanum

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37: 241:, and have no hairs or scales except at the very base. They range from 1 to 12 centimetres (0.39 to 4.72 in) long and 0.3 to 0.6 millimetres (0.01 to 0.02 in) wide, and typically make up one-half to two-thirds of the total length of the frond. 256:(leaf axis) is green beneath, sometimes with streaks of purple at the base, smooth, and has small wings on the upper side, from 0.1 to 0.2 millimetres (0.004 to 0.008 in) wide. 229:(underground stem). The rhizome bears narrow, blackish scales, 1 to 2 millimetres (0.04 to 0.08 in) long and 0.1 to 0.2 millimetres (0.004 to 0.008 in) wide, which are 237:(the stalk of the leaf, below the blade), 7 to 20 centimetres (2.8 to 7.9 in) long. The stipes are dark purple in color at the base, becoming green towards the blade, lack 329: 185: 300: 282: 294: 288: 272:
2 to 5 millimetres (0.08 to 0.2 in) long and 0.5 to 0.8 millimetres (0.02 to 0.03 in) wide, with jagged margins. The spores are kidney-shaped.
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in reference to the unclear and mysterious relationship between this species and other spleenworts; it is roughly intermediate in form between
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Each leaf bears from one to five pairs of pinnae (the first subdivisions of the leaf). The pair closest to the base is the largest.
624: 536: 650: 694: 384: 238: 233:(bear a lattice-like pattern) and have untoothed margins. The fronds grow in clumps and are, including the 230: 753: 602: 689: 531:. Memoirs of the New York Botanical Garden. Vol. 88. Bronx, New York: New York Botanical Garden. 36: 552: 748: 564: 668: 144: 383:, at altitudes from 200 to 700 metres (700 to 2,000 ft). It has also been collected at 8: 681: 743: 715: 168: 31: 663: 611: 532: 409: 720: 616: 320:
by the deeply cut and toothed pinnae and pinnules (rather than rounded and lobed).
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by the enlargement of the side of the lowest pinna towards the leaf base, and from
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per pinnule on both sides of the costule (pinnule midvein). They are covered by
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by the dark purple (rather than green) color at the base of its stipe, from
707: 111: 101: 91: 637: 596: 629: 642: 364:, but has well-formed spores, so it is not a first-generation hybrid. 312:
by the lack of long whitish hairs both above and below the leaf, from
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Specimens have in the past been identified as, or thought similar to,
206: 121: 655: 558: 702: 581: 459: 269: 503:"Specimen of Asplenium arcanum A. R. Sm. recorded on Sep 20, 1981" 380: 376: 341: 226: 375:
grows on stream banks and on wooded hillsides in the states of
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in 2004 based on material from Nayarit. He gave it the epithet
253: 202: 265: 48: 449: 447: 445: 443: 441: 439: 437: 435: 81: 71: 483: 471: 432: 420:. Saint Louis, Missouri: Missouri Botanical Garden 735: 264:Fertile fronds bear from one to three pairs of 526: 489: 477: 465: 453: 367: 328:Material of this species was described by 527:Mickel, John T.; Smith, Alan R. (2004). 390: 736: 563: 562: 402: 348:. It was described as a species by 332:in 1894 as a deeply cut variety of 217: 13: 695:urn:lsid:ipni.org:names:77094922-1 275: 259: 14: 765: 546: 340:, based on material collected in 225:has fibrous roots and an upright 35: 520: 495: 212: 201:is a fern known from western 1: 395: 385:Apoyo Lagoon Natural Reserve 7: 529:The Pteridophytes of Mexico 323: 10: 770: 571: 555:from JSTOR Global Plants. 468:, pp. 74, 76, 82–83. 244: 174: 167: 150: 143: 32:Scientific classification 30: 23: 368:Distribution and habitat 490:Mickel & Smith 2004 478:Mickel & Smith 2004 466:Mickel & Smith 2004 454:Mickel & Smith 2004 391:Notes and references 330:George E. Davenport 754:Flora of Nicaragua 358:A. tenerrimum 314:A. tenerrimum 304:. It differs from 301:A. tenerrimum 731: 730: 664:Open Tree of Life 603:Asplenium arcanum 573:Asplenium arcanum 565:Taxon identifiers 538:978-0-89327-488-7 412:Asplenium arcanum 373:Asplenium arcanum 223:Asplenium arcanum 198:Asplenium arcanum 194: 193: 188: 178:Asplenium pumilum 154:Asplenium arcanum 25:Asplenium arcanum 761: 724: 723: 711: 710: 698: 697: 685: 684: 672: 671: 659: 658: 646: 645: 633: 632: 620: 619: 607: 606: 605: 592: 591: 590: 560: 559: 542: 514: 513: 511: 509: 499: 493: 487: 481: 475: 469: 463: 457: 451: 430: 429: 427: 425: 406: 306:A. fragrans 283:A. fragrans 218:Roots and stipes 184: 156: 40: 39: 21: 20: 769: 768: 764: 763: 762: 760: 759: 758: 749:Ferns of Mexico 734: 733: 732: 727: 719: 714: 706: 701: 693: 688: 680: 675: 667: 662: 654: 649: 641: 636: 628: 623: 615: 610: 601: 600: 595: 586: 585: 580: 567: 549: 539: 523: 518: 517: 507: 505: 501: 500: 496: 488: 484: 476: 472: 464: 460: 452: 433: 423: 421: 414:Baker synonyms" 408: 407: 403: 398: 393: 370: 362:A. pumilum 334:A. pumilum 326: 318:A. minimum 310:A. pumilum 295:A. pumilum 289:A. minimum 278: 276:Similar species 262: 260:Sori and spores 247: 220: 215: 163: 158: 152: 139: 136:A. arcanum 34: 17: 16:Species of fern 12: 11: 5: 767: 757: 756: 751: 746: 729: 728: 726: 725: 721:wfo-0001240213 712: 699: 686: 673: 660: 647: 634: 621: 608: 593: 577: 575: 569: 568: 557: 556: 548: 547:External links 545: 544: 543: 537: 522: 519: 516: 515: 494: 482: 470: 458: 431: 400: 399: 397: 394: 392: 389: 387:in Nicaragua. 369: 366: 325: 322: 277: 274: 261: 258: 246: 243: 219: 216: 214: 211: 192: 191: 190: 189: 172: 171: 165: 164: 159: 148: 147: 141: 140: 133: 131: 127: 126: 119: 115: 114: 109: 105: 104: 99: 95: 94: 89: 85: 84: 82:Polypodiopsida 79: 75: 74: 72:Polypodiophyta 69: 65: 64: 59: 52: 51: 46: 42: 41: 28: 27: 15: 9: 6: 4: 3: 2: 766: 755: 752: 750: 747: 745: 742: 741: 739: 722: 717: 713: 709: 704: 700: 696: 691: 687: 683: 678: 674: 670: 665: 661: 657: 652: 648: 644: 639: 635: 631: 626: 622: 618: 613: 609: 604: 598: 594: 589: 583: 579: 578: 576: 574: 570: 566: 561: 554: 551: 550: 540: 534: 530: 525: 524: 504: 498: 492:, p. 81. 491: 486: 480:, p. 74. 479: 474: 467: 462: 456:, p. 82. 455: 450: 448: 446: 444: 442: 440: 438: 436: 419: 415: 413: 405: 401: 388: 386: 382: 378: 374: 365: 363: 359: 355: 351: 350:Alan R. Smith 347: 343: 339: 335: 331: 321: 319: 315: 311: 307: 303: 302: 297: 296: 291: 290: 285: 284: 273: 271: 267: 257: 255: 250: 242: 240: 236: 232: 228: 224: 210: 208: 204: 200: 199: 187: 183: 179: 176: 175: 173: 170: 166: 162: 157: 155: 149: 146: 145:Binomial name 142: 138: 137: 132: 129: 128: 125: 124: 120: 117: 116: 113: 110: 107: 106: 103: 100: 97: 96: 93: 90: 87: 86: 83: 80: 77: 76: 73: 70: 67: 66: 63: 62:Tracheophytes 60: 57: 54: 53: 50: 47: 44: 43: 38: 33: 29: 26: 22: 19: 682:tro-50106525 572: 528: 506:. Retrieved 497: 485: 473: 461: 422:. Retrieved 417: 411: 404: 372: 371: 361: 357: 353: 337: 336:, var.  333: 327: 317: 313: 309: 305: 299: 293: 287: 281: 279: 263: 251: 248: 222: 221: 197: 196: 195: 181: 177: 153: 151: 135: 134: 122: 112:Aspleniaceae 102:Aspleniineae 92:Polypodiales 55: 24: 18: 638:iNaturalist 597:Wikispecies 521:Works cited 213:Description 738:Categories 677:Plant List 656:77094922-1 396:References 338:laciniatum 182:laciniatum 180:var.  98:Suborder: 68:Division: 744:Asplenium 588:Q15609551 231:clathrate 207:Nicaragua 130:Species: 123:Asplenium 45:Kingdom: 708:50106525 703:Tropicos 582:Wikidata 553:Holotype 418:Tropicos 410:"Name - 324:Taxonomy 169:Synonyms 108:Family: 669:6008561 630:4257058 508:July 1, 424:July 1, 381:Sinaloa 377:Nayarit 354:arcanum 346:Pringle 342:Jalisco 270:indusia 227:rhizome 186:Davenp. 161:A.R.Sm. 118:Genus: 88:Order: 78:Class: 49:Plantae 643:282619 535:  298:, and 254:rachis 245:Leaves 203:Mexico 617:5VV6S 239:wings 235:stipe 56:Clade 690:POWO 651:IPNI 625:GBIF 533:ISBN 510:2016 426:2016 379:and 360:and 266:sori 252:The 205:and 716:WFO 612:CoL 344:by 740:: 718:: 705:: 692:: 679:: 666:: 653:: 640:: 627:: 614:: 599:: 584:: 434:^ 416:. 292:, 286:, 209:. 58:: 541:. 512:. 428:.

Index

Scientific classification
Edit this classification
Plantae
Tracheophytes
Polypodiophyta
Polypodiopsida
Polypodiales
Aspleniineae
Aspleniaceae
Asplenium
Binomial name
A.R.Sm.
Synonyms
Davenp.
Mexico
Nicaragua
rhizome
clathrate
stipe
wings
rachis
sori
indusia
A. fragrans
A. minimum
A. pumilum
A. tenerrimum
George E. Davenport
Jalisco
Pringle

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