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Asilidae

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860:, but doubts remain about the real nature of the trophic regime and its mechanisms. The entomophagy of some species had indeed been already hypothesized by some authors of the 19th century, based on the findings of larvae of asilids associated with larvae of other insects, but Melin (1923) asserted that, in reality, predation was occasional and secondary to the plant-based diet. More recent studies have confirmed the entomophagy of some asilids without extending this species' feeding behavior for the whole family. Less certain, however, is the mechanism of entomophagy: in general, the behavior is cited as predation, but for some species may be ectoparasitoids. Musso (1983) described the feeding behavior of the larvae of 646: 757:, and Therevidae; the radial R is always four-branched, with R2+3 unbranched. Details of wing venation determine subfamilies and lower taxa. The wings are most often hyaline, but sometimes smoky or dark colored, or partly infuscated in many genera or completely darkened. The abdomen consists of six to eight visible segments preceding the genitalia in males, but the eighth segment is sometimes entirely or partially concealed, and terminal forming the ovipositor. It is long and narrow conical in most species, but wide, dorsoventrally flattened and short in bee mimics. In the Leptogastrinae, the abdomen is extremely long and slender. In some tribes, the male undergoes axial torsion of 180°. 102: 958: 1043: 826: 1072: 346: 694: 728: 126: 359: 635: 818:, in both the juvenile and the adult stages, and feed on small arthropods, mainly insects. Although predatory forms in the adult stage are present in other taxonomic groups of Diptera, the Asilidae are the most representative for the number of species and for uniformity of feeding behavior (>7000 species, all of which are predatory). The combination of high biodiversity and high predatory activity leads to this family playing an important role in the 1020: 963: 962: 959: 964: 761: 961: 541: 872: 601:. Larvae are also predacious, feeding on eggs, larvae, or other soft-bodied insects. Robber flies overwinter as larvae and pupate in the soil. Pupae migrate to the soil surface and emerge as adults, often leaving behind their pupal casing. Complete development ranges from one to three years, depending on species and environmental conditions. 713:. The proboscis is rounded in cross section or laterally or dorsoventrally compressed and is usually stout, and straight and sometimes able to penetrate through the hard integument of Coleoptera. The maxillary palpi are at the base beside the labium, two-segmented in all Dasypogoninae or single segmented in Asilinae and Leptogastrinae. 864:: the larvae of the first instar does not feed on insects, those of the second instar feed on secretions by larvae of beetles (and may cause death), while the larvae of the third and fourth instars actually behave like predators. In short, the feeding behavior of larval asilids can be intermediate between predation and ectoparasitism. 1012:. Their biodiversity is lower in forested ecosystems, and where asilids do occur in such environments, they tend to concentrate in the glades and margins. In those conditions, the interrupted canopy leaves space for various species of shrubs and herbaceous plants suited to asilid styles of predation. 1015:
In general, the biology of the Asilidae is still poorly known, but various authors have studied the population distribution in particular regions and ecosystems. They have classified the behavioral patterns in terms of microenvironments, ecological, and trophic factors, showing how different species
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With regards to interspecific trophic relationships, a large number of reports exists on the prey captured by the Asilidae. Lavigne (2003) has developed a database comprising over 13,000 reports. The prey of Asilidae are predominantly represented by other insects, mostly winged, but several cases in
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Much better known and described in detail is the behavior of adults. In general, predation in adults is concentrated in the hottest hours in open, sunny spaces, while at night, they take refuge in dense vegetation. The Asilidae are excellent flyers, and in most of the family, capture prey in flight.
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is apodous, cylindrical, and elongated, more or less flattened dorsoventrally and tapered at the cephalic and caudal ends. The colour is white or yellowish. The head is small, rugged, dark-pigmented and hypognathous, the abdomen is composed of eight apparent urites, with the last two often fused and
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are generally relatively short and hairy; the third segment (or first flagellomere) has an oval or oblong shape, is generally longer than the two basal segments, and bears a stylus generally composed of two segments, of which the basal is very short. In some asilids, the stylus can be monoarticolate
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The prey is caught with the tarsi and injected with a paralyzing saliva. The asilid pierces the integument of the prey with the prepharyx (hyopharynx) in preferential points of least resistance such as the eyes, the membranous area of transition between the head and thorax (neck) or between thorax
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Egg-laying takes place, according to the species, with three different behaviors that relate to the structure and the morphology of the abdomen. Females with an undifferentiated ovipositor release eggs randomly and independently from the substrate. In other cases, however, the abdomen bears a
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With regards to the specificity of the trophic relationship, Wood (1981) mentions some studies in the literature on the subject. Some genera have been found to be monophagic, but more generally Asilidae are polyphagic, with behaviors that vary from narrow specialization to broad prey choice.
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Adults are generally medium to large in size, with an average body width of 1 to 1.5 cm (0.39 to 0.59 in), but with a range of 3 mm (0.12 in) to more than 5 cm (2.0 in) in length. The shape is generally elongated, due to the conformation of the long tapering
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Female robber flies deposit whitish-colored eggs on low-lying plants and grasses, or in crevices within soil, bark, or wood. Egg-laying habits depend on the species and their specific habitat; most species lay their eggs in masses, which are then covered with a chalky protective coating.
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species occur at altitudes exceeding 4000 meters/13,000 feet. However, the highest levels of biodiversity are in warm climates; tropical or subtropical and arid or semi-arid regions tend to have the greatest variety of species, followed by areas where rainfall is highly seasonal.
544: 543: 549: 547: 542: 876: 875: 879: 878: 874: 548: 853:) and one pupa. The larvae of the first instar differ from other stages in both ethology and trophic regime. The larvae of most known asilids live in the soil or in the case of some taxonomic groups, in rotting organic material, usually wood and the bark of dead trees. 880: 960: 546: 877: 673:
has a convex profile with a characteristic dense bundle of bristles, called a "mystax". The mystax helps protect the head and face when the fly encounters prey bent on defense. Other bristles are arranged on the ocellar tubercle.
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cause paralysis of the victim, while proteolytic enzymes lead to the breakup and liquefaction of internal tissues. In a short time, the predator is able to feed by sucking the internal fluids through the alimentary canal.
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Other studies have shown that the ratio between the size of the prey and the asilid varies from 1.8:1 to 3.7:1, with an average of 2.6:1. The ratio tends to increase with decreasing size of the predator.
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of Asilidae favour particular habitats suited to particular patterns of reproduction and predation. Specific studies show correlations between the floristic composition and predatory behaviour.
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Dennis, D. Steve; Barnes, Jeffrey K.; Knutson, Lloyd (17 June 2013). "Review and analysis of information on the biology and morphology of immature stages of robber flies (Diptera: Asilidae)".
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Asilidae generally occur in habitats that are open, sunny, and dry, even arid. They favour open or scattered vegetation, and some species even frequent bare ground. Typical habitats include
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which they have attacked spiders have also been reported. Within the insects, orders that include the most frequent prey of asilids include a wide range of families within the
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They are often seen stationed to ambush prey at strategic points. This behavior signifies that sight plays an essential role in the detection of prey and their capture.
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and even other Asilidae, in fact practically anything of a suitable size. Some Asilidae do, however, specialize in smaller prey, and this is reflected in their more
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and liveries are often showy, with colors ranging from brown to black to grey, sometimes in contrast with other colors such as red and yellow. Frequently they are
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Though they are a very characteristic group for such a large family, the Asilidae may easily be confused with the related and less widely known family
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meaning "moustache" or "upper lip". The mystax has been suggested to afford some protection for the head and face when the flies deal with struggling
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After hatching, robber fly larvae generally seem to live in soil, rotting wood, leaf mold, and similar materials, some being predatory and others
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The egg is hyaline or pigmented and of variable shape from spherical to oval and up to 2 mm in length. The surface is smooth or bears
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and ocular fringe typical of the Asilidae, with short, stout proboscis and spiny, powerful legs, adapted to the capture of prey in flight.
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Cells: d: discal; br: 1st basal; bm: 2nd basal; r1: marginal; r3: 1st submarginal; r5: 1st posterior; m3: 4th posterior; cup: cell cup
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which very rapidly paralyze the victim and soon digest the insides; the fly then sucks the liquefied material through the proboscis.
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The fly attacks its prey by stabbing it with its short, strong proboscis, injecting the victim with saliva containing neurotoxic and
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chemicals the prey may have at their disposal. Many Asilidae when attacked in turn do not hesitate to defend themselves with their
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is generally well developed, with the exception of the Leptogastrinae and part of Dasypogoninae. The venation is much as in the
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The mouthparts are short and modified for piercing-sucking. They consist of a strongly sclerotized proboscis which includes the
1823: 1838: 1529: 1482: 1439: 1091:. Their taxonomy is still under study in the light of new specimens and cladistic analysis. The 14 accepted subfamilies are: 4190: 1501: 641:: a powerful predator with body adapted to fast flight. Note the mystax, the proboscis and the depression between the eyes. 619:, however there are also compact species with broad abdomens. The integument is covered with thick hair, especially on the 1522:
Imms' General Textbook of Entomology: Volume 1: Structure, Physiology and Development Volume 2: Classification and Biology
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in an early work of 1803 erected four genera, three of which now represent subfamilies. He also described many species in
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type, composed typically of five segments but sometimes from three to four, depending on the structure of the stylus. The
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differentiated, specialized ovipositor to lay eggs in the soil or sand, or lay them in cavities within plant tissues.
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Musso, Joseph-Jean (1983). "Nutritive and Ecological Requirements of Robber Flies (Diptera: Brachycera: Asilidae)".
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The thorax is robust and compact. Unlike in other lower Brachycera, it bears long bristles (macrochaeta) useful as
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Section of Asilidae mouthparts a: labrum; b: hypopharynx; c: maxillary palp; d: maxillae; e: food canal; f: labium
4138: 1968: 4198: 1892: 1545: 1357: 706: 645: 1430:. In McAlpine, J. F.; Peterson, B. V.; Shewell, G. E.; Teskey, H. J.; Vockeroth, J. R.; Wood, D. M. (eds.). 4306: 1365: 1353: 791: 786: 746: 721: 710: 702: 686: 682: 670: 666: 662: 658: 624: 620: 616: 101: 4055: 4032: 1704:
D. Grimaldi. 1990. Insects from the Santana formation, Lower Cretaceous, of Brazil - Chapter 9. Diptera.
1445: 1042: 665:. This feature is clearly visible in the front view and is a morphological peculiarity of Asilidae. The 1631:"Fritz Geller-Grimm. Information on Robber Flies: Ecological Classification in Robber flies (Asilidae)" 4265: 4060: 1906: 4278: 4226: 1961: 1329: 522: 382: 125: 825: 390: 222: 3957: 1382: 897:. Puncture is followed by the injection of saliva, whose active components perform two functions: 1352:
in particular. Other prominent authors dealing with the Asilidae during the 19th century included
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Recherches sur le développement, la nutrition et l'écologie des Asilidae (Diptera - Brachycera)
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Head-on view showing the characteristic depression formed by the elevation of the compound eyes
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Photographic atlas and identification key to the robber flies of Germany (Diptera: Asilidae)
1348:(1800 to 1838). During the rest of the 19th century, significant contributions were made by 492: 4094: 4019: 1325: 819: 533:
comprising the mystax of the Asilidae. Furthermore, in the Asilidae the depression on the
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except Antarctica. In the Northern Hemisphere, some species are even adapted to tundra.
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takes place in 1–3 years. The postembryonic development consists of four larval stages (
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Hull, Frank M. (1962). "Robber Flies of the World: The Genera of the Family Asilidae".
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function. The wings are well developed, often relatively narrow for speedy flight; the
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Longitudinal veins: C: costa; Sc: subcosta; R: radius; M: media; Cu: cubitus; A: anal.
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With regards to feeding behavior, most of the literature describes Asilidae larvae as
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are short, have three segments, and sometimes bear a bristle-like structure called an
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has one or more rows of bristles aligned behind the posterior margin of the eye. The
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Robber flies of North America – reference photographs, descriptions, natural history
1793:. Handbooks for the identification of British insects. Royal Entomological Society. 3857: 3763: 3746: 3624: 3066: 2669: 2426: 2337: 1942: 1754: 1746: 1679: 1568: 1001: 525:
adapted to the sucking of liquid foods. Again, the Therevidae commonly have fluffy
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was the authority for establishing the family in 1802. The Asilidae, together with
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Comprehensive overview by Fritz Geller-Grimm, Torsten Dikow and Robert J. Lavigne
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In general, the family attacks a very wide range of prey, including other flies,
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more or less reduced. The respiratory system is amphineustic, with two pairs of
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Development research, nutrition and ecology of Asilidae (Diptera - Brachycera)
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Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten
1305: 1110: 1047: 1028: 989: 777:, which are generally polygonal and visible only in the electron microscope. 741:
The legs are relatively long and strong, with many macrochaetes performing a
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in the Therevidae is not a piercing, predatory organ, but ends in two fleshy
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Crossveins: h: humeral; r-m: radio-medial; m-m: medial; m-cu: medio-cubital
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and may deliver intensely painful bites to humans if handled incautiously.
417: 1953: 1812: 1406: 1318:, including 11 species, and added four others in the 12th edition (1767). 4159: 4047: 4004: 3844: 3823: 3554: 3537: 3513: 3500: 3424: 3403: 3316: 3308: 3298: 3241: 3212: 3172: 3146: 3037: 3011: 2956: 2840: 2807: 2750: 2687: 2644: 2498: 2457: 2449: 2431: 2324: 2306: 2119: 1274: 1236: 1024: 923: 911: 857: 750: 628: 598: 534: 484: 394: 363: 358: 197: 34: 634: 3836: 3779: 3713: 3689: 3663: 3608: 3440: 3411: 3378: 3355: 3277: 3164: 3131: 3118: 3092: 3027: 2915: 2742: 2729: 2674: 2624: 2581: 2573: 2290: 2256: 2248: 2095: 2068: 2060: 2044: 1759: 1644: 1216: 1120: 931: 898: 834: 717: 661:
arranged in a characteristic depression formed by the elevation of the
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Comparative behavior of Wyoming robber flies II (Diptera: Asilidae)
1324:(1758) is the type of the genus. The rank of family is credited to 1095: 1019: 985: 565: 433: 350: 238: 74: 69: 54: 49: 39: 537:
between the eyes tends to be more obvious than in the Therevidae.
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and they form one of the most characteristic groups of the lower
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Key to the North American genera and a catalog of species - 2007
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in five publications dated from 1775 to 1805, erected the genus
760: 517:. Some points of contrast between the families include that the 3637: 2695: 2010: 1998: 1173: 1005: 997: 993: 939: 907: 894: 850: 488: 452: 167: 147: 709:
which form a food canal, the labrum and a piercing organ, the
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characters. Bristles of this type are always present on the
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Robber flies have stout, spiny legs and three simple eyes (
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The pupa is naked, as in the majority of the Orthorrhapha,
530: 526: 468: 460: 1865: 1985: 1600: 915: 464: 378: 177: 432:; various Asilidae prey on formidable species including 1546:"Backyard Gardener - Robber Flies - September 3, 2014" 1669: 1515: 1513: 1499:
http://www.bt-images.net/beautiful-eyes/robber-fly/
1083:The Asilidae currently include over 7500 described 1706:Bulletin of the American Museum of Natural History 1470: 2382: 2196: 4293: 2481: 1637: 1510: 1462: 1378:List of soldierflies and allies of Great Britain 2897:(scuttle flies, coffin flies, humpbacked flies) 1418: 1416: 1414: 2434:(minute black scavenger flies, or dung midges) 1519: 1336:and described 76 exotic and European species. 1046:Asilidae Robber fly from the Anaimalai hills, 3887: 3754: 2408: 1969: 1743:Bulletin of the United States National Museum 1606: 1023:Robber flies (Asilidae) mating in a blade of 3918: 2238: 2059: 1813:Database service for Asilidae – Robber flies 1411: 491:. They do so apparently irrespective of any 2518: 2085: 1983: 1607:Dennis, D Steve; Lavigne, Robert J (1975). 1590:"Database Asilidae: predator-prey database" 2643: 1976: 1962: 1925:Courting-dance of the robber fly (gallery) 1663: 926:; prey belonging to various other orders ( 100: 1824:West Palaearctic species including Russia 1758: 1552: 1166:The oldest known member of the family is 1297:Clade showing relationship of Asiloidea 1070: 1041: 1018: 956: 870: 824: 759: 726: 692: 644: 633: 539: 357: 344: 3619:(blow-flies: bluebottles, greenbottles) 2622: 2042: 1788: 1520:Richards, O. W.; Davies, R. G. (1977). 1468: 1407:http://www.geller-grimm.de/genera15.htm 631:, imitating the livery of Hymenoptera. 529:above the mouthparts, unlike the stiff 4294: 2852: 1730:, CD-ROM, Amphx-Verlag Halle (Saale). 1308:(LinnĂ©; 1758), in the 10th edition of 1300: 1004:, and related shrubland types such as 979: 832:, a robber fly that closely resembles 362:A member of the Asilidae feeding on a 3971: 3970: 2621: 2041: 1957: 1767: 1558: 325:. They are powerfully built, bristly 4312:Taxa named by Pierre AndrĂ© Latreille 4266:7708F17A-B83F-E502-63F1-FA1D66F1FCD3 4227:ea9011a8-9132-4beb-95a5-f85257bea94f 4139:e3f594ab-2f9a-4a07-9e15-ac2b5fbc1c98 1740: 1584: 1582: 1422: 13: 3082:(cactus flies, banana stalk flies) 1950:Images. Flowers visited by adults. 1718: 571:Many Asilidae have long, tapering 14: 4323: 1806: 1579: 893:and abdomen, or between the last 374:The Asilidae are a family in the 1776:] (PhD thesis) (in French). 731:Basal plan of the wing venation. 653:The head is free and mobile and 424:, a term derived from the Greek 124: 32: 1854:Family Asilidae at Bugguide.net 1698: 1623: 1037: 1939:Excellent. Includes structure. 1538: 1491: 1400: 1066: 575:, sometimes with a sword-like 1: 2766:(dagger flies, balloon flies) 1888:Family description and images 1829:Australasian/Oceanian species 1393: 604: 333:enclosing the sharp, sucking 1366:Jacques-Marie-Frangile Bigot 1354:Pierre-Justin-Marie Macquart 1183: 971:lays eggs on an ear of grass 946:, etc.) are also mentioned. 806:and therefore able to move. 657:in both sexes and has three 7: 3958:List of families of Diptera 3590:(house flies, stable flies) 1921:Featured Creatures Web site 1903:Featured Creatures Web site 1768:Musso, Joseph-Jean (1978). 1573:10.1127/entom.gen/9/1983/35 1371: 1172:from the Early Cretaceous ( 838:paper wasp species such as 385:, with over 7000 described 340: 10: 4328: 2361:(dark-winged fungus gnats) 2353:(long-winged fungus gnats) 2340:(long-beaked fungus gnats) 1432:Manual of Nearctic Diptera 809: 3979: 3955: 3909: 3878: 3835: 3804: 3795: 3745: 3741: 3680: 3607: 3562: 3553: 3525: 3460: 3435:(vinegar and fruit flies) 3402: 3345:(lekking, or druid flies) 3307: 3268: 3232: 3155: 3091: 3057: 2994: 2973: 2964: 2955: 2951: 2914: 2861: 2848: 2839: 2798: 2741: 2652: 2639: 2635: 2631: 2617: 2568:(long-bodied crane flies) 2548: 2527: 2514: 2477: 2448: 2417: 2404: 2378: 2289: 2268: 2247: 2234: 2192: 2139: 2094: 2081: 2055: 2051: 2037: 1993: 1871:Images (text in Japanese) 1684:10.11646/zootaxa.3673.1.1 1651:. Smithsonian Institution 1645:"Asiloid Flies: Asilidae" 1477:. London: Edward Arnold. 1469:Jeffrey, Charles (1973). 1330:Johan Christian Fabricius 1271: 1249: 1229: 1209: 1202: 1194: 235: 230: 121:Scientific classification 119: 109:Fan-bristled robber fly ( 108: 99: 23: 2332:(predatory fungus gnats) 1866:Atlas of German Asilidae 1791:Tabanoidea and Asiloidea 1789:Oldroyd, Harold (1969). 1383:List of Asilidae species 780: 677:The antennae are of the 609: 366:. This asilid shows the 3223:(black scavenger flies) 2584:(hairy-eyed craneflies) 2501:(primitive crane flies) 2259:(march flies, lovebugs) 1751:10.5479/si.03629236.224 1724:Geller-Grimm F (2003): 1473:Biological nomenclature 1423:Wood, Grace C. (1981). 1388:List of Asilidae genera 1056:zoogeographical regions 888:. Includes slow motion. 830:Blepharepium sonorensis 797: 579:. Others, for instance 3865:(horse and deer flies) 3137:Strongylophthalmyiidae 3048:(picture-winged flies) 2576:(limoniid crane flies) 1948:Family Asilidae at EOL 1937:Robberflies of Germany 1860:Family Asilidae at EOL 1192:Asiloidea   1080: 1054:Asilidae occur in all 1051: 1034: 972: 889: 842: 768: 765: 738: 698: 650: 642: 561: 371: 355: 4248:Paleobiology Database 3451:(frightful hairy fly) 3280:(beetle-backed flies) 2818:(tangle-veined flies) 2703:(hilarimorphid flies) 2664:(flower-loving flies) 2493:(phantom crane flies) 1561:Entomologia Generalis 1342:Johann Wilhelm Meigen 1328:in Samouelle (1819). 1321:Asilus crabroniformis 1079:feeding on a honeybee 1074: 1045: 1022: 967: 883: 828: 763: 730: 696: 648: 637: 552: 361: 348: 115:) with honeybee prey 4134:Fauna Europaea (new) 3782:(wood soldier flies) 3632:(New Zealand batfly) 3582:(little house flies) 3074:(stilt-legged flies) 2985:(thick-headed flies) 2881:(spear-winged flies) 2810:(small-headed flies) 2539:(winter crane flies) 2114:(frog-biting midges) 1524:. Berlin: Springer. 1314:, erected the genus 884:A robber fly with a 820:ecological stability 559:Rhynocoris annulatus 434:stinging Hymenoptera 329:with a short, stout 321:family, also called 4307:Brachycera families 3847:(water snipe flies) 3368:(upside-down flies) 2905:(flat-footed flies) 2889:(flat-footed flies) 2758:(long-legged flies) 2592:(large crane flies) 2208:(net-winged midges) 2175:(non-biting midges) 1649:asiloidflies.si.edu 1594:www.geller-grimm.de 1301:Notable researchers 980:Habitat and ecology 969:Didysmachus picipes 639:Choerades fimbriata 566:proteolytic enzymes 3516:(freeloader flies) 3319:(leaf miner flies) 3259:(small dung flies) 3103:(stalk-eyed flies) 2926:(big-headed flies) 2384:Perissommatomorpha 2214:Deuterophlebiidae 2198:Blephariceromorpha 1930:2019-12-27 at the 1876:2020-01-30 at the 1844:World list species 1081: 1077:Stenopogon martini 1052: 1035: 973: 895:abdominal tergites 890: 843: 766: 739: 699: 651: 643: 562: 372: 356: 112:Dysmachus trigonus 4289: 4288: 4235:Open Tree of Life 3973:Taxon identifiers 3964: 3963: 3951: 3950: 3947: 3946: 3943: 3942: 3939: 3938: 3931: 3905: 3904: 3874: 3873: 3866: 3848: 3827: 3791: 3790: 3783: 3775: 3767: 3764:Pantophthalmidae 3737: 3736: 3733: 3732: 3729: 3728: 3725: 3724: 3717: 3709: 3701: 3693: 3667: 3659: 3641: 3633: 3620: 3599: 3591: 3583: 3575: 3549: 3548: 3541: 3517: 3504: 3491: 3483: 3470:Acartophthalmidae 3452: 3444: 3436: 3428: 3420: 3419:(quasimodo flies) 3369: 3346: 3338: 3320: 3289: 3281: 3260: 3224: 3216: 3168: 3127: 3104: 3083: 3075: 3049: 3041: 3023: 3015: 3007: 2986: 2947: 2946: 2943: 2942: 2935: 2927: 2906: 2898: 2890: 2882: 2874: 2835: 2834: 2831: 2830: 2827: 2826: 2819: 2811: 2780: 2767: 2759: 2733: 2725: 2712: 2704: 2691: 2683: 2665: 2613: 2612: 2609: 2608: 2605: 2604: 2601: 2600: 2593: 2585: 2577: 2569: 2559: 2540: 2510: 2509: 2502: 2494: 2483:Ptychopteromorpha 2473: 2472: 2469: 2468: 2461: 2435: 2400: 2399: 2374: 2373: 2370: 2369: 2362: 2354: 2341: 2338:Lygistorrhinidae 2333: 2315: 2300: 2281: 2260: 2230: 2229: 2217: 2216:(mountain midges) 2209: 2188: 2187: 2184: 2183: 2176: 2168: 2160: 2152: 2151:(solitary midges) 2131: 2123: 2115: 2107: 2106:(meniscus midges) 2077: 2076: 1531:978-0-412-61390-6 1497:Robber fly eyes: 1484:978-0-7131-2431-6 1441:978-0-660-10731-8 1294: 1293: 1285: 1284: 1260: 1259: 1251: ? Asilidae 1200: N.N.  965: 881: 862:Machimus rusticus 814:The Asilidae are 550: 311: 310: 226: 4319: 4282: 4281: 4269: 4268: 4256: 4255: 4243: 4242: 4230: 4229: 4220: 4219: 4207: 4206: 4204:NBNSYS0000159448 4194: 4193: 4181: 4180: 4168: 4167: 4155: 4154: 4142: 4141: 4129: 4128: 4116: 4115: 4103: 4102: 4090: 4089: 4077: 4076: 4064: 4063: 4051: 4050: 4041: 4040: 4028: 4027: 4015: 4014: 4013: 4000: 3999: 3998: 3968: 3967: 3929: 3916: 3915: 3885: 3884: 3880:Vermileonomorpha 3864: 3858:Pelecorhynchidae 3846: 3825: 3802: 3801: 3781: 3773: 3765: 3752: 3751: 3747:Stratiomyomorpha 3743: 3742: 3715: 3707: 3699: 3691: 3665: 3657: 3639: 3631: 3630:Mystacinobiidae 3625:Mesembrinellidae 3618: 3597: 3589: 3581: 3573: 3560: 3559: 3539: 3515: 3502: 3489: 3481: 3450: 3442: 3434: 3426: 3418: 3367: 3344: 3336: 3335:Aulacigastridae 3318: 3287: 3279: 3258: 3222: 3214: 3166: 3142:Syringogastridae 3125: 3102: 3081: 3073: 3067:Cypselosomatidae 3047: 3039: 3021: 3020:Platystomatidae 3013: 3005: 2984: 2971: 2970: 2962: 2961: 2953: 2952: 2933: 2925: 2904: 2896: 2888: 2880: 2872: 2859: 2858: 2850: 2849: 2846: 2845: 2817: 2809: 2778: 2765: 2757: 2732:(stiletto flies) 2731: 2723: 2710: 2702: 2701:Hilarimorphidae 2689: 2681: 2670:Apsilocephalidae 2663: 2650: 2649: 2641: 2640: 2637: 2636: 2633: 2632: 2619: 2618: 2591: 2583: 2575: 2567: 2566:Cylindrotomidae 2558: 2557: 2553: 2538: 2525: 2524: 2516: 2515: 2500: 2492: 2479: 2478: 2459: 2433: 2427:Canthyloscelidae 2415: 2414: 2406: 2405: 2380: 2379: 2360: 2352: 2339: 2331: 2313: 2299: 2298: 2294: 2279: 2258: 2245: 2244: 2236: 2235: 2215: 2207: 2206:Blephariceridae 2194: 2193: 2174: 2166: 2165:Ceratopogonidae 2158: 2150: 2129: 2122:(phantom midges) 2121: 2113: 2105: 2092: 2091: 2083: 2082: 2057: 2056: 2053: 2052: 2039: 2038: 1978: 1971: 1964: 1955: 1954: 1834:Nearctic species 1802: 1785: 1764: 1762: 1712: 1702: 1696: 1695: 1667: 1661: 1660: 1658: 1656: 1641: 1635: 1634: 1627: 1621: 1620: 1604: 1598: 1597: 1586: 1577: 1576: 1556: 1550: 1549: 1542: 1536: 1535: 1517: 1508: 1495: 1489: 1488: 1476: 1466: 1460: 1459: 1457: 1456: 1450: 1444:. Archived from 1429: 1420: 1409: 1404: 1205: 1204: 1197: 1196: 1189: 1188: 1106:Brachyrhopalinae 1002:maquis shrubland 966: 882: 822:of entomofauna. 667:occipital region 551: 249:Brachyrhopalinae 221: 129: 128: 104: 94: 31: 27:Temporal range: 21: 20: 4327: 4326: 4322: 4321: 4320: 4318: 4317: 4316: 4292: 4291: 4290: 4285: 4277: 4272: 4264: 4259: 4251: 4246: 4238: 4233: 4225: 4223: 4215: 4210: 4202: 4197: 4189: 4184: 4176: 4171: 4163: 4158: 4150: 4145: 4137: 4132: 4124: 4119: 4111: 4106: 4098: 4093: 4085: 4080: 4072: 4067: 4059: 4054: 4046: 4044: 4036: 4031: 4023: 4018: 4009: 4008: 4003: 3994: 3993: 3988: 3975: 3965: 3960: 3935: 3911:Xylophagomorpha 3901: 3870: 3831: 3787: 3774:(soldier flies) 3721: 3676: 3666:(tachina flies) 3603: 3596:Scathophagidae 3574:(cabbage flies) 3545: 3521: 3475:Australimyzidae 3456: 3449:Mormotomyiidae 3398: 3366:Neurochaetidae 3303: 3264: 3257:Sphaeroceridae 3234:Sphaeroceroidea 3228: 3188:Heterocheilidae 3183:Helosciomyzidae 3151: 3087: 3053: 3040:(peacock flies) 3006:(flutter flies) 2990: 2939: 2910: 2879:Lonchopteridae 2823: 2794: 2772:Homalocnemiidae 2756:Dolichopodidae 2737: 2627: 2597: 2555: 2554: 2552: 2544: 2506: 2491:Ptychopteridae 2465: 2444: 2396: 2392:Perissommatidae 2366: 2351:Rangomaramidae 2296: 2295: 2293: 2285: 2264: 2226: 2180: 2167:(biting midges) 2135: 2112:Corethrellidae 2073: 2047: 2033: 1989: 1982: 1932:Wayback Machine 1878:Wayback Machine 1809: 1721: 1719:Further reading 1716: 1715: 1703: 1699: 1668: 1664: 1654: 1652: 1643: 1642: 1638: 1629: 1628: 1624: 1605: 1601: 1588: 1587: 1580: 1557: 1553: 1544: 1543: 1539: 1532: 1518: 1511: 1496: 1492: 1485: 1467: 1463: 1454: 1452: 1448: 1442: 1427: 1421: 1412: 1405: 1401: 1396: 1374: 1362:Camillo Rondani 1311:Systema Naturae 1303: 1295: 1286: 1261: 1186: 1178:Crato Formation 1161:Willistonininae 1151:Tillobromatinae 1146:Stichopogoninae 1069: 1040: 982: 957: 871: 812: 800: 783: 775:microsculptures 771: 736: 734: 732: 612: 607: 586:are fat-bodied 540: 481:ichneumon wasps 343: 304:Willistonininae 294:Tillobromatinae 289:Stichopogoninae 220: 123: 95: 93: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 42: 37: 29: 28: 25: 17: 16:Family of flies 12: 11: 5: 4325: 4315: 4314: 4309: 4304: 4287: 4286: 4284: 4283: 4270: 4257: 4244: 4231: 4221: 4208: 4195: 4182: 4169: 4156: 4143: 4130: 4121:Fauna Europaea 4117: 4104: 4091: 4078: 4065: 4052: 4042: 4029: 4016: 4001: 3985: 3983: 3977: 3976: 3962: 3961: 3956: 3953: 3952: 3949: 3948: 3945: 3944: 3941: 3940: 3937: 3936: 3934: 3933: 3924: 3922: 3913: 3907: 3906: 3903: 3902: 3900: 3899: 3893: 3891: 3889:Vermileonoidea 3882: 3876: 3875: 3872: 3871: 3869: 3868: 3860: 3855: 3850: 3841: 3839: 3833: 3832: 3830: 3829: 3821: 3816: 3814:Austroleptidae 3810: 3808: 3799: 3793: 3792: 3789: 3788: 3786: 3785: 3777: 3772:Stratiomyidae 3769: 3766:(timber flies) 3760: 3758: 3756:Stratiomyoidea 3749: 3739: 3738: 3735: 3734: 3731: 3730: 3727: 3726: 3723: 3722: 3720: 3719: 3711: 3706:Nycteribiidae 3703: 3698:Hippoboscidae 3695: 3692:(tsetse flies) 3686: 3684: 3682:Hippoboscoidea 3678: 3677: 3675: 3674: 3669: 3661: 3656:Sarcophagidae 3653: 3648: 3643: 3635: 3627: 3622: 3617:Calliphoridae 3613: 3611: 3605: 3604: 3602: 3601: 3593: 3585: 3577: 3568: 3566: 3557: 3551: 3550: 3547: 3546: 3544: 3543: 3535: 3533:Cryptochetidae 3529: 3527: 3523: 3522: 3520: 3519: 3511: 3506: 3498: 3493: 3485: 3477: 3472: 3466: 3464: 3458: 3457: 3455: 3454: 3446: 3438: 3433:Drosophilidae 3430: 3422: 3414: 3408: 3406: 3400: 3399: 3397: 3396: 3391: 3386: 3384:Periscelididae 3381: 3376: 3371: 3363: 3358: 3353: 3351:Fergusoninidae 3348: 3340: 3332: 3327: 3322: 3313: 3311: 3305: 3304: 3302: 3301: 3296: 3291: 3286:Chamaemyiidae 3283: 3274: 3272: 3266: 3265: 3263: 3262: 3254: 3252:Nannodastiidae 3249: 3244: 3238: 3236: 3230: 3229: 3227: 3226: 3218: 3210: 3205: 3200: 3195: 3190: 3185: 3180: 3175: 3170: 3161: 3159: 3153: 3152: 3150: 3149: 3144: 3139: 3134: 3129: 3121: 3116: 3111: 3106: 3097: 3095: 3089: 3088: 3086: 3085: 3077: 3069: 3063: 3061: 3055: 3054: 3052: 3051: 3043: 3035: 3030: 3025: 3022:(signal flies) 3017: 3014:(cheese flies) 3009: 3004:Pallopteridae 3000: 2998: 2992: 2991: 2989: 2988: 2979: 2977: 2968: 2959: 2949: 2948: 2945: 2944: 2941: 2940: 2938: 2937: 2929: 2920: 2918: 2912: 2911: 2909: 2908: 2900: 2892: 2884: 2876: 2873:(ironic flies) 2867: 2865: 2856: 2843: 2837: 2836: 2833: 2832: 2829: 2828: 2825: 2824: 2822: 2821: 2816:Nemestrinidae 2813: 2804: 2802: 2800:Nemestrinoidea 2796: 2795: 2793: 2792: 2787: 2782: 2774: 2769: 2761: 2753: 2747: 2745: 2739: 2738: 2736: 2735: 2727: 2724:(window flies) 2719: 2717:Mythicomyiidae 2714: 2706: 2698: 2693: 2685: 2682:(robber flies) 2677: 2672: 2667: 2658: 2656: 2647: 2629: 2628: 2615: 2614: 2611: 2610: 2607: 2606: 2603: 2602: 2599: 2598: 2596: 2595: 2587: 2579: 2571: 2562: 2560: 2546: 2545: 2543: 2542: 2537:Trichoceridae 2533: 2531: 2529:Trichoceroidea 2522: 2512: 2511: 2508: 2507: 2505: 2504: 2496: 2487: 2485: 2475: 2474: 2471: 2470: 2467: 2466: 2464: 2463: 2454: 2452: 2446: 2445: 2443: 2442: 2437: 2429: 2423: 2421: 2412: 2410:Psychodomorpha 2402: 2401: 2398: 2397: 2395: 2394: 2388: 2386: 2376: 2375: 2372: 2371: 2368: 2367: 2365: 2364: 2356: 2348: 2346:Mycetophilidae 2343: 2335: 2327: 2322: 2317: 2312:Cecidomyiidae 2309: 2307:Bolitophilidae 2303: 2301: 2297:(fungus gnats) 2287: 2286: 2284: 2283: 2274: 2272: 2266: 2265: 2263: 2262: 2253: 2251: 2242: 2232: 2231: 2228: 2227: 2225: 2224: 2219: 2211: 2202: 2200: 2190: 2189: 2186: 2185: 2182: 2181: 2179: 2178: 2170: 2162: 2154: 2145: 2143: 2137: 2136: 2134: 2133: 2125: 2117: 2109: 2100: 2098: 2089: 2079: 2078: 2075: 2074: 2072: 2071: 2065: 2063: 2049: 2048: 2035: 2034: 2032: 2031: 2025: 2019: 2013: 2007: 2001: 1994: 1991: 1990: 1981: 1980: 1973: 1966: 1958: 1952: 1951: 1945: 1940: 1934: 1922: 1904: 1890: 1885: 1880: 1868: 1863: 1857: 1851: 1846: 1841: 1836: 1831: 1826: 1821: 1816: 1808: 1807:External links 1805: 1804: 1803: 1786: 1765: 1745:(224): 1–907. 1738: 1720: 1717: 1714: 1713: 1697: 1662: 1636: 1622: 1599: 1578: 1567:(1–2): 35–50. 1551: 1537: 1530: 1509: 1504:2014-04-23 at 1490: 1483: 1461: 1440: 1410: 1398: 1397: 1395: 1392: 1391: 1390: 1385: 1380: 1373: 1370: 1358:Francis Walker 1302: 1299: 1292: 1291: 1288: 1287: 1283: 1282: 1279: 1278: 1270: 1267: 1266: 1263: 1262: 1258: 1257: 1254: 1253: 1248: 1245: 1244: 1241: 1240: 1228: 1225: 1224: 1221: 1220: 1208: 1203: 1201: 1195: 1193: 1187: 1185: 1182: 1164: 1163: 1158: 1156:Trigonomiminae 1153: 1148: 1143: 1141:Stenopogoninae 1138: 1133: 1128: 1126:Leptogastrinae 1123: 1118: 1113: 1108: 1103: 1101:Bathypogoninae 1098: 1068: 1065: 1039: 1036: 981: 978: 811: 808: 799: 796: 782: 779: 770: 767: 611: 608: 606: 603: 342: 339: 323:assassin flies 309: 308: 307: 306: 301: 299:Trigonomiminae 296: 291: 286: 284:Stenopogoninae 281: 276: 271: 269:Leptogastrinae 266: 261: 256: 251: 246: 244:Bathypogoninae 241: 233: 232: 228: 227: 215: 211: 210: 205: 201: 200: 195: 191: 190: 185: 181: 180: 175: 171: 170: 165: 161: 160: 155: 151: 150: 145: 141: 140: 135: 131: 130: 117: 116: 106: 105: 97: 96: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 38: 33: 30:Aptian–Present 26: 15: 9: 6: 4: 3: 2: 4324: 4313: 4310: 4308: 4305: 4303: 4300: 4299: 4297: 4280: 4275: 4271: 4267: 4262: 4258: 4254: 4249: 4245: 4241: 4236: 4232: 4228: 4222: 4218: 4213: 4209: 4205: 4200: 4196: 4192: 4187: 4183: 4179: 4174: 4170: 4166: 4161: 4157: 4153: 4148: 4144: 4140: 4135: 4131: 4127: 4122: 4118: 4114: 4109: 4105: 4101: 4096: 4092: 4088: 4083: 4079: 4075: 4070: 4066: 4062: 4057: 4053: 4049: 4043: 4039: 4034: 4030: 4026: 4021: 4017: 4012: 4006: 4002: 3997: 3991: 3987: 3986: 3984: 3982: 3978: 3974: 3969: 3959: 3954: 3932: 3928:Xylophagidae 3926: 3925: 3923: 3921: 3920:Xylophagoidea 3917: 3914: 3912: 3908: 3898: 3897:Vermileonidae 3895: 3894: 3892: 3890: 3886: 3883: 3881: 3877: 3867: 3861: 3859: 3856: 3854: 3851: 3849: 3843: 3842: 3840: 3838: 3834: 3828: 3826:(snipe flies) 3822: 3820: 3817: 3815: 3812: 3811: 3809: 3807: 3803: 3800: 3798: 3794: 3784: 3778: 3776: 3770: 3768: 3762: 3761: 3759: 3757: 3753: 3750: 3748: 3744: 3740: 3718: 3712: 3710: 3704: 3702: 3700:(louse flies) 3696: 3694: 3688: 3687: 3685: 3683: 3679: 3673: 3670: 3668: 3662: 3660: 3658:(flesh flies) 3654: 3652: 3651:Rhinophoridae 3649: 3647: 3644: 3642: 3636: 3634: 3628: 3626: 3623: 3621: 3615: 3614: 3612: 3610: 3606: 3600: 3594: 3592: 3586: 3584: 3578: 3576: 3572:Anthomyiidae 3570: 3569: 3567: 3565: 3561: 3558: 3556: 3552: 3542: 3540:(lance flies) 3536: 3534: 3531: 3530: 3528: 3524: 3518: 3512: 3510: 3507: 3505: 3499: 3497: 3494: 3492: 3490:(beach flies) 3486: 3484: 3478: 3476: 3473: 3471: 3468: 3467: 3465: 3463: 3459: 3453: 3447: 3445: 3443:(shore flies) 3439: 3437: 3431: 3429: 3423: 3421: 3417:Curtonotidae 3415: 3413: 3410: 3409: 3407: 3405: 3401: 3395: 3392: 3390: 3389:Teratomyzidae 3387: 3385: 3382: 3380: 3377: 3375: 3372: 3370: 3364: 3362: 3359: 3357: 3354: 3352: 3349: 3347: 3341: 3339: 3333: 3331: 3328: 3326: 3323: 3321: 3315: 3314: 3312: 3310: 3306: 3300: 3297: 3295: 3294:Cremifaniidae 3292: 3290: 3288:(aphid flies) 3284: 3282: 3276: 3275: 3273: 3271: 3267: 3261: 3255: 3253: 3250: 3248: 3245: 3243: 3240: 3239: 3237: 3235: 3231: 3225: 3219: 3217: 3215:(marsh flies) 3211: 3209: 3208:Ropalomeridae 3206: 3204: 3201: 3199: 3198:Natalimyzidae 3196: 3194: 3191: 3189: 3186: 3184: 3181: 3179: 3176: 3174: 3171: 3169: 3163: 3162: 3160: 3158: 3154: 3148: 3145: 3143: 3140: 3138: 3135: 3133: 3130: 3128: 3122: 3120: 3117: 3115: 3114:Megamerinidae 3112: 3110: 3107: 3105: 3099: 3098: 3096: 3094: 3090: 3084: 3078: 3076: 3072:Micropezidae 3070: 3068: 3065: 3064: 3062: 3060: 3056: 3050: 3044: 3042: 3036: 3034: 3031: 3029: 3026: 3024: 3018: 3016: 3010: 3008: 3002: 3001: 2999: 2997: 2993: 2987: 2981: 2980: 2978: 2976: 2972: 2969: 2967: 2963: 2960: 2958: 2954: 2950: 2936: 2930: 2928: 2924:Pipunculidae 2922: 2921: 2919: 2917: 2913: 2907: 2903:Platypezidae 2901: 2899: 2893: 2891: 2885: 2883: 2877: 2875: 2871:Ironomyiidae 2869: 2868: 2866: 2864: 2863:Platypezoidea 2860: 2857: 2855: 2851: 2847: 2844: 2842: 2838: 2820: 2814: 2812: 2806: 2805: 2803: 2801: 2797: 2791: 2788: 2786: 2785:Oreogetonidae 2783: 2781: 2779:(dance flies) 2775: 2773: 2770: 2768: 2762: 2760: 2754: 2752: 2749: 2748: 2746: 2744: 2740: 2734: 2728: 2726: 2722:Scenopinidae 2720: 2718: 2715: 2713: 2711:(mydas flies) 2707: 2705: 2699: 2697: 2694: 2692: 2686: 2684: 2678: 2676: 2675:Apystomyiidae 2673: 2671: 2668: 2666: 2660: 2659: 2657: 2655: 2651: 2648: 2646: 2642: 2638: 2634: 2630: 2626: 2620: 2616: 2594: 2588: 2586: 2580: 2578: 2572: 2570: 2564: 2563: 2561: 2556:(crane flies) 2551: 2547: 2541: 2535: 2534: 2532: 2530: 2526: 2523: 2521: 2517: 2513: 2503: 2497: 2495: 2489: 2488: 2486: 2484: 2480: 2476: 2462: 2456: 2455: 2453: 2451: 2447: 2441: 2440:Valeseguyidae 2438: 2436: 2430: 2428: 2425: 2424: 2422: 2420: 2416: 2413: 2411: 2407: 2403: 2393: 2390: 2389: 2387: 2385: 2381: 2377: 2363: 2357: 2355: 2349: 2347: 2344: 2342: 2336: 2334: 2330:Keroplatidae 2328: 2326: 2323: 2321: 2320:Diadocidiidae 2318: 2316: 2314:(gall midges) 2310: 2308: 2305: 2304: 2302: 2292: 2288: 2282: 2278:Anisopodidae 2276: 2275: 2273: 2271: 2270:Anisopodoidea 2267: 2261: 2255: 2254: 2252: 2250: 2246: 2243: 2241: 2240:Bibionomorpha 2237: 2233: 2223: 2222:Nymphomyiidae 2220: 2218: 2212: 2210: 2204: 2203: 2201: 2199: 2195: 2191: 2177: 2173:Chironomidae 2171: 2169: 2163: 2161: 2159:(black flies) 2155: 2153: 2149:Thaumaleidae 2147: 2146: 2144: 2142: 2141:Chironomoidea 2138: 2132: 2126: 2124: 2118: 2116: 2110: 2108: 2102: 2101: 2099: 2097: 2093: 2090: 2088: 2084: 2080: 2070: 2067: 2066: 2064: 2062: 2061:Axymyiomorpha 2058: 2054: 2050: 2046: 2040: 2036: 2030: 2026: 2024: 2020: 2018: 2014: 2012: 2008: 2006: 2002: 2000: 1996: 1995: 1992: 1987: 1979: 1974: 1972: 1967: 1965: 1960: 1959: 1956: 1949: 1946: 1944: 1943:Wing venation 1941: 1938: 1935: 1933: 1929: 1926: 1923: 1920: 1916: 1912: 1910: 1907:bee killers, 1905: 1902: 1898: 1894: 1891: 1889: 1886: 1884: 1881: 1879: 1875: 1872: 1869: 1867: 1864: 1861: 1858: 1855: 1852: 1850: 1847: 1845: 1842: 1840: 1839:Japan species 1837: 1835: 1832: 1830: 1827: 1825: 1822: 1820: 1817: 1814: 1811: 1810: 1800: 1796: 1792: 1787: 1783: 1779: 1775: 1771: 1766: 1761: 1756: 1752: 1748: 1744: 1739: 1737: 1736:3-932795-18-0 1733: 1729: 1728: 1723: 1722: 1710: 1707: 1701: 1693: 1689: 1685: 1681: 1677: 1673: 1666: 1650: 1646: 1640: 1632: 1626: 1618: 1614: 1610: 1603: 1595: 1591: 1585: 1583: 1574: 1570: 1566: 1562: 1555: 1547: 1541: 1533: 1527: 1523: 1516: 1514: 1507: 1506:archive.today 1503: 1500: 1494: 1486: 1480: 1475: 1474: 1465: 1451:on 2016-11-04 1447: 1443: 1437: 1433: 1426: 1419: 1417: 1415: 1408: 1403: 1399: 1389: 1386: 1384: 1381: 1379: 1376: 1375: 1369: 1367: 1363: 1359: 1355: 1351: 1347: 1343: 1339: 1335: 1331: 1327: 1323: 1322: 1317: 1313: 1312: 1307: 1306:Carl Linnaeus 1298: 1290: 1289: 1281: 1280: 1277: 1276: 1269: 1268: 1265: 1264: 1256: 1255: 1252: 1247: 1246: 1243: 1242: 1239: 1238: 1234: 1227: 1226: 1223: 1222: 1219: 1218: 1214: 1207: 1206: 1199: 1198: 1191: 1190: 1181: 1179: 1175: 1171: 1170: 1162: 1159: 1157: 1154: 1152: 1149: 1147: 1144: 1142: 1139: 1137: 1134: 1132: 1129: 1127: 1124: 1122: 1119: 1117: 1114: 1112: 1111:Dasypogoninae 1109: 1107: 1104: 1102: 1099: 1097: 1094: 1093: 1092: 1090: 1087:in about 556 1086: 1078: 1073: 1064: 1061: 1057: 1049: 1048:Western Ghats 1044: 1033: 1030: 1029:Western ghats 1026: 1021: 1017: 1013: 1011: 1007: 1003: 999: 995: 991: 990:forest steppe 987: 977: 970: 955: 951: 947: 945: 941: 937: 933: 929: 925: 921: 917: 913: 909: 903: 900: 896: 887: 869: 865: 863: 859: 858:entomophagous 854: 852: 848: 841: 837: 836: 831: 827: 823: 821: 817: 807: 805: 795: 793: 788: 778: 776: 762: 758: 756: 752: 748: 744: 729: 725: 723: 719: 714: 712: 708: 704: 695: 691: 688: 684: 680: 675: 672: 671:facial region 668: 664: 663:compound eyes 660: 656: 647: 640: 636: 632: 630: 626: 622: 618: 602: 600: 595: 591: 589: 585: 583: 578: 574: 569: 567: 560: 557:feeding on a 556: 555:Laphria flava 538: 536: 532: 528: 524: 520: 516: 511: 509: 505: 500: 498: 494: 493:repugnatorial 490: 486: 482: 478: 474: 470: 466: 462: 458: 454: 449: 447: 443: 439: 435: 431: 427: 423: 419: 418:compound eyes 415: 410: 408: 404: 400: 396: 392: 388: 384: 380: 377: 369: 365: 360: 353: 352: 347: 338: 336: 332: 328: 324: 320: 316: 305: 302: 300: 297: 295: 292: 290: 287: 285: 282: 280: 277: 275: 272: 270: 267: 265: 262: 260: 257: 255: 254:Dasypogoninae 252: 250: 247: 245: 242: 240: 237: 236: 234: 229: 224: 219: 216: 213: 212: 209: 206: 204:Superfamily: 203: 202: 199: 196: 193: 192: 189: 186: 183: 182: 179: 176: 173: 172: 169: 166: 163: 162: 159: 156: 153: 152: 149: 146: 143: 142: 139: 136: 133: 132: 127: 122: 118: 114: 113: 107: 103: 98: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 41: 36: 22: 19: 3980: 3853:Oreoleptidae 3845:Athericidae 3824:Rhagionidae 3819:Bolbomyiidae 3806:Rhagionoidea 3797:Tabanomorpha 3690:Glossinidae 3672:Ulurumyiidae 3598:(dung flies) 3538:Lonchaeidae 3526:Lonchaeoidea 3514:Milichiidae 3509:Inbiomyiidae 3503:(frit flies) 3501:Chloropidae 3425:Diastatidae 3394:Xenasteiidae 3325:Anthomyzidae 3317:Agromyzidae 3270:Lauxanioidea 3247:Heleomyzidae 3213:Sciomyzidae 3203:Phaeomyiidae 3193:Huttoninidae 3178:Helcomyzidae 3167:(kelp flies) 3157:Sciomyzoidea 3126:(rust flies) 3038:Tephritidae 3033:Richardiidae 3012:Piophilidae 2996:Tephritoidea 2966:Acalyptratae 2934:(hoverflies) 2808:Acroceridae 2688:Bombyliidae 2679: 2662:Apioceridae 2520:Tipulomorpha 2499:Tanyderidae 2460:(moth flies) 2458:Psychodidae 2450:Psychodoidea 2432:Scatopsidae 2419:Scatopsoidea 2280:(wood gnats) 2130:(mosquitoes) 2120:Chaoboridae 2087:Culicomorpha 2029:Holometabola 2027:Superorder: 2021:Infraclass: 1908: 1893:robber flies 1883:Diptera.info 1790: 1773: 1769: 1742: 1725: 1708: 1705: 1700: 1675: 1671: 1665: 1653:. 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Retrieved 1446:the original 1431: 1402: 1350:Hermann Loew 1345: 1333: 1319: 1315: 1309: 1304: 1296: 1272: 1250: 1230: 1213:Scenopinidae 1210: 1167: 1165: 1082: 1076: 1053: 1038:Distribution 1014: 983: 974: 968: 952: 948: 940:Thysanoptera 904: 891: 866: 861: 855: 844: 839: 833: 829: 813: 801: 784: 772: 740: 715: 700: 676: 652: 638: 613: 596: 592: 580: 570: 563: 558: 554: 512: 501: 485:grasshoppers 450: 438:grasshoppers 425: 421: 411: 383:cosmopolitan 373: 367: 349: 322: 318: 314: 312: 231:Subfamilies 217: 194:Infraorder: 110: 18: 4160:iNaturalist 4005:Wikispecies 3930:(awl flies) 3780:Xylomyidae 3716:(bat flies) 3714:Streblidae 3708:(bat flies) 3664:Tachinidae 3555:Calyptratae 3441:Ephydridae 3427:(bog flies) 3404:Ephydroidea 3337:(sap flies) 3309:Opomyzoidea 3299:Lauxaniidae 3278:Celyphidae 3242:Chyromyidae 3173:Dryomyzidae 3165:Coelopidae 3147:Tanypezidae 2957:Schizophora 2841:Muscomorpha 2751:Atelestidae 2730:Therevidae 2690:(bee flies) 2645:Asilomorpha 2582:Pediciidae 2574:Limoniidae 2325:Ditomyiidae 2257:Bibionidae 2249:Bibionoidea 2157:Simuliidae 1760:10088/10126 1678:(1): 1–64. 1275:Bombyliidae 1237:Apioceridae 1180:of Brazil. 1116:Dioctriinae 1067:Systematics 1025:lemon grass 924:Lepidoptera 912:Hymenoptera 899:neurotoxins 751:Rhagionidae 722:notopleuron 711:hypopharynx 690:or absent. 599:detrivorous 487:, and some 477:damselflies 473:dragonflies 457:butterflies 442:dragonflies 436:, powerful 395:Bombyliidae 364:grasshopper 335:hypopharynx 259:Dioctriinae 198:Asilomorpha 4296:Categories 3863:Tabanidae 3837:Tabanoidea 3640:(botflies) 3638:Oestridae 3609:Oestroidea 3580:Fanniidae 3488:Canacidae 3482:(bee lice) 3480:Braulidae 3412:Camillidae 3379:Opomyzidae 3356:Marginidae 3343:Clusiidae 3132:Somatiidae 3119:Nothybidae 3101:Diopsidae 3093:Diopsoidea 3046:Ulidiidae 3028:Pyrgotidae 2983:Conopidae 2975:Conopoidea 2932:Syrphidae 2916:Syrphoidea 2887:Opetiidae 2777:Hybotidae 2764:Empididae 2743:Empidoidea 2625:Brachycera 2590:Tipulidae 2550:Tipuloidea 2359:Sciaridae 2291:Sciaroidea 2128:Culicidae 2096:Culicoidea 2069:Axymyiidae 2045:Nematocera 2015:Subclass: 2005:Arthropoda 1909:Mallophora 1655:10 January 1455:2016-04-18 1425:"Asilidae" 1394:References 1217:Therevidae 1169:Araripogon 1131:Ommatiinae 1121:Laphriinae 932:Neuroptera 908:Coleoptera 847:life cycle 840:P. apachus 629:aposematic 605:Morphology 577:ovipositor 515:Therevidae 463:, various 407:Brachycera 399:Therevidae 319:robber fly 274:Ommatiinae 264:Laphriinae 188:Brachycera 184:Suborder: 158:Arthropoda 3646:Rhiniidae 3588:Muscidae 3564:Muscoidea 3462:Carnoidea 3374:Odiniidae 3361:Neminidae 3330:Asteiidae 3221:Sepsidae 3124:Psilidae 3109:Gobryidae 3080:Neriidae 3059:Nerioidea 2895:Phoridae 2790:Ragadidae 2696:Evocoidae 2680:Asilidae 2654:Asiloidea 2623:Suborder 2043:Suborder 2017:Pterygota 1997:Kingdom: 1799:256410648 1338:Wiedemann 1184:Phylogeny 1136:Phellinae 1010:chaparral 944:Blattodea 920:Hemiptera 886:honey bee 816:predators 792:spiracles 764:Egglaying 755:Tabanidae 743:raptorial 718:taxonomic 655:dichoptic 588:bumblebee 497:probosces 403:Asiloidea 391:Latreille 331:proboscis 279:Phellinae 223:Latreille 208:Asiloidea 144:Kingdom: 138:Eukaryota 4302:Asilidae 4069:BugGuide 4045:BioLib: 4038:ASILIDAE 4025:Asilidae 4011:Asilidae 3990:Wikidata 3981:Asilidae 3496:Carnidae 2709:Mydidae 2104:Dixidae 2023:Neoptera 2003:Phylum: 1999:Animalia 1988:families 1928:Archived 1874:Archived 1782:30534417 1711::164-183 1692:26146701 1617:10795887 1502:Archived 1372:See also 1231: ? 1211: ? 1096:Asilinae 986:savannah 936:Isoptera 914:, other 835:Polistes 707:maxillae 679:aristate 590:mimics. 573:abdomens 504:antennae 426:mystakos 351:Zosteria 341:Overview 317:are the 315:Asilidae 239:Asilinae 218:Asilidae 214:Family: 154:Phylum: 148:Animalia 134:Domain: 24:Asilidae 3996:Q837089 2854:Aschiza 2011:Insecta 2009:Class: 1986:Diptera 1984:Extant 1913:on the 1895:on the 1672:Zootaxa 1334:Damalis 1233:Mydidae 1085:species 1075:Female 1050:, India 992:, open 928:Odonata 916:Diptera 851:instars 810:Biology 804:exarate 687:pedicel 617:abdomen 582:Laphria 553:A male 531:chaetae 523:labella 489:spiders 453:beetles 448:build. 446:gracile 387:species 379:Diptera 178:Diptera 174:Order: 168:Insecta 164:Class: 4279:150931 4240:653876 4224:NZOR: 4191:132394 4178:101287 4113:1ASILF 1862:Images 1856:Images 1797:  1780:  1734:  1690:  1615:  1528:  1481:  1438:  1364:, and 1316:Asilus 1273:  1174:Aptian 1089:genera 1060:Alpine 1006:fynbos 998:desert 996:, semi 994:steppe 922:, and 703:labium 659:ocelli 625:thorax 535:vertex 519:labium 508:arista 422:mystax 414:ocelli 368:mystax 225:, 1802 4274:WoRMS 4261:Plazi 4253:76780 4217:50673 4173:IRMNG 4165:47982 4126:10884 4048:17396 1772:[ 1449:(PDF) 1428:(PDF) 1326:Leach 1032:India 1027:, in 787:larva 781:Larva 747:alula 683:scape 610:Adult 527:setae 461:moths 376:order 327:flies 4212:NCBI 4186:ITIS 4152:7275 4147:GBIF 4108:EPPO 4056:BOLD 1919:IFAS 1911:spp. 1901:IFAS 1795:OCLC 1778:OCLC 1732:ISBN 1688:PMID 1676:3673 1657:2017 1613:OCLC 1526:ISBN 1479:ISBN 1436:ISBN 1235:and 1215:and 1008:and 845:The 798:Pupa 785:The 705:and 685:and 623:and 621:head 502:The 475:and 469:ants 465:bees 459:and 430:prey 397:and 313:The 35:Preęž’ 4199:NBN 4100:508 4095:EoL 4087:6SY 4082:CoL 4074:151 4061:693 4033:AFD 4020:ADW 1755:hdl 1747:doi 1709:195 1680:doi 1569:doi 769:Egg 354:sp. 4298:: 4276:: 4263:: 4250:: 4237:: 4214:: 4201:: 4188:: 4175:: 4162:: 4149:: 4136:: 4123:: 4110:: 4097:: 4084:: 4071:: 4058:: 4035:: 4022:: 4007:: 3992:: 1917:/ 1915:UF 1899:/ 1897:UF 1753:. 1686:. 1674:. 1647:. 1592:. 1581:^ 1563:. 1512:^ 1413:^ 1368:. 1360:, 1356:, 1176:) 1000:, 988:, 942:, 938:, 934:, 930:, 918:, 910:, 753:, 510:. 483:, 479:, 471:, 467:, 455:, 440:, 409:. 389:. 85:Pg 1977:e 1970:t 1963:v 1801:. 1784:. 1763:. 1757:: 1749:: 1694:. 1682:: 1659:. 1633:. 1619:. 1596:. 1575:. 1571:: 1565:9 1548:. 1534:. 1487:. 1458:. 584:, 90:N 80:K 75:J 70:T 65:P 60:C 55:D 50:S 45:O 40:ęž’

Index

Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N

Dysmachus trigonus
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Diptera
Brachycera
Asilomorpha
Asiloidea
Asilidae
Latreille
Asilinae
Bathypogoninae
Brachyrhopalinae
Dasypogoninae

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