782:
226:". Three years later, Hershey and DelbrĂĽck would independently discover that different strains of bacteriophage can both exchange genetic material when infecting the same bacterial cell. This process results in hybrid phages containing genetic material from both sources, which Hershey referred to as "genetic recombination".
33:
268:
Hershey died from congestive heart failure on May 22, 1997 at his home in Laurel Hollow, New York. He was 88 years old. At the time, he was survived by his wife
Harriet Davidson (1918–2000) and their only child, Peter Manning Hershey (1956–1999).
253:
Hershey's work with bacteriophage would earn him a share of the 1969 Nobel Prize in
Physiology or Medicine with DelbrĂĽck and Luria, "for their discoveries concerning the replication mechanism and the genetic structure of viruses."
249:
In 1962, Hershey was named director of the
Department of Genetics, a position he held until his retirement in 1970. He would live on the grounds of Cold Spring Harbor Laboratory (CSHL) for the rest of his life.
3277:
2620:
206:
viruses that infect and replicate inside bacteria. Hershey's work on the factors impacting the virus' ability to infect its targets brought him to the attention of fellow phage researchers
222:
In 1943, DelbrĂĽck invited
Hershey to Vanderbilt University to discuss his phage research. Together, with Luria, they would form the core of an informal network of researchers called "the
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manner made him the pope, of course, and Luria was the hard-working, socially sensitive priest-confessor. And Al (Hershey) was the saint."
2319:
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At
Washington University, Hershey worked closely with department head Jacques Bronfenbrenner to investigate bacteriophages, or phages
147:
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2354:
261:, an extensive volume on the subject, published by CSHL Press that same year. In 1981, Hershey became a founding member of the
543:
359:
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831:
684:
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111:
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Although officially retired from scientific research, Hershey would continue to pursue new projects. In 1971, he edited
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to Robert Day and Alma Wilbur
Hershey. He earned a B.S. in chemistry in 1930, and Ph.D. in bacteriology in 1934 from
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544:"The Hershey-Chase Experiments (1952), by Alfred Hershey and Martha Chase | The Embryo Project Encyclopedia"
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280:), was led by the Trinity of DelbrĂĽck, Luria, and Hershey. DelbrĂĽck's status as founder and his
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276:, a member of The Phage Group, wrote: "The Phage Church, as we were sometimes called (see
8:
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467:
376:
337:
329:
76:
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1791:
1759:
1500:
575:"[The Nobel Prize in Physiology or Medicine to 3 bacteriophage researchers]"
438:"Independent functions of viral protein and nucleic acid in growth of bacteriophage"
3150:
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229:
Hershey left
Washington University in 1950 for the Department of Genetics of the
2479:
1560:
241:. Their work confirmed that DNA, not protein, was the genetic material of life.
207:
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1456:
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1376:
1253:
943:
887:
743:
654:
592:
510:
333:
194:, serving as an instructor of bacteriology and immunology from 1934 to 1950.
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129:
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528:
341:
3107:
3027:
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2078:
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1987:
1963:
1871:
1536:
1332:
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879:
871:
855:
277:
223:
453:
3097:
3019:
2394:
2014:
1273:
1221:
1168:
171:
2674:
2215:
1767:
645:
627:
2010:
133:
2628:
734:
707:
32:
3278:
Recipients of the Albert Lasker Award for Basic
Medical Research
237:. Two years later, he and Martha Chase would conduct the famous
3293:
Members of the United States
National Academy of Sciences
792:
Linus
Pauling and the Race for DNA: A Documentary History
137:
368:
Biographical Memoirs of the National Academy of Sciences
190:. Shortly after, Hershey accepted a faculty position at
306:
Campbell, Allan; Stahl, Franklin W. (December 1998).
625:
628:"Alfred Hershey--Nobel Prize for work in virology"
162:(December 4, 1908 – May 22, 1997) was an American
3303:Washington University School of Medicine faculty
3209:
677:"Founding Members of the World Cultural Council"
3243:Founding members of the World Cultural Council
2369:
548:Embryo Project Encyclopediadate= June 23, 2019
107:Albert Lasker Award for Basic Medical Research
2614:
2355:
812:
305:
239:Hershey–Chase, or "Waring Blender" experiment
435:
3283:Washington University in St. Louis faculty
2621:
2607:
2362:
2348:
819:
805:
780:
626:Shampo, Marc A; Kyle Robert A (May 2004).
301:
299:
31:
3263:Nobel laureates in Physiology or Medicine
733:
644:
518:
461:
416:The American Association of Immunologists
323:
197:
3268:People from Cold Spring Harbor, New York
572:
535:
244:
148:Washington University School of Medicine
98:Proof of DNA as genetic material of life
566:
478:
353:
351:
296:
3210:
705:
699:
687:from the original on February 27, 2023
607:from the original on February 27, 2023
554:from the original on February 27, 2023
16:American bacteriologist and geneticist
2602:
2343:
832:Nobel Prize in Physiology or Medicine
800:
619:
541:
484:
357:
112:Nobel Prize in Physiology or Medicine
407:
405:
390:from the original on January 7, 2021
348:
788:Key Participants: Alfred D. Hershey
13:
436:Hershey, AD; Chase, M (May 1952).
231:Carnegie Institution of Washington
217:
192:Washington University in St. Louis
14:
3314:
2443:International Labour Organization
770:
485:Stahl, Franklin W (May 1, 1998).
442:The Journal of General Physiology
402:
3298:Scientists from New York (state)
3258:Michigan State University alumni
3228:20th-century American chemists
669:
429:
177:
1:
3288:People from Syosset, New York
289:
235:Cold Spring Harbor Laboratory
3273:People from Owosso, Michigan
573:Malmgren, B (October 1969).
325:10.1146/annurev.genet.32.1.1
7:
1924:Christiane NĂĽsslein-Volhard
10:
3319:
708:"Alfred Hershey (1908-97)"
413:"Alfred D. Hershey, Ph.D."
2637:
2526:
2501:
2470:
2451:
2428:
2404:
2379:
1997:
1662:
1323:
1121:Charles Scott Sherrington
1097:Frederick Gowland Hopkins
1059:
838:
639:(5). United States: 590.
375:. United States: 142–59.
312:Annual Review of Genetics
188:Michigan State University
153:
143:
125:
118:
102:
94:
89:Michigan State University
84:
65:
39:
30:
23:
3238:American Nobel laureates
2629:Founding members of the
2111:Françoise Barré-Sinoussi
503:10.1093/genetics/149.1.1
2690:André Frédéric Cournand
706:Cairns, J (July 1997).
587:(44). Sweden: 1369–75.
2631:World Cultural Council
2532:Nobel Prize recipients
2472:Physiology or Medicine
892:Santiago RamĂłn y Cajal
720:(6638). England: 130.
681:World Cultural Council
263:World Cultural Council
198:Bacteriophage research
2946:Marshall W. Nirenberg
2941:William A. Nierenberg
2901:Vernon B. Mountcastle
2710:Jean-François Denisse
2642:Christian B. Anfinsen
1393:Dickinson W. Richards
1077:Julius Wagner-Jauregg
542:Hernandez, Victoria.
272:Following his death,
245:Later years and death
3233:American geneticists
3041:William G. Schneider
3036:Menahem Max Schiffer
3020:Hakim Muhammad Saeed
2994:Chintamani N. R. Rao
2732:Jacques-Émile Dubois
1816:Rita Levi-Montalcini
1177:Albert Szent-Györgyi
360:"Alfred Day Hershey"
182:Hershey was born in
3248:History of genetics
2974:Comlan A. A. Quenum
2936:Bernhard H. Neumann
2833:Vassos Karageorghis
2770:José Rafael Estrada
2698:Eduardo de Robertis
2208:William C. Campbell
2119:Elizabeth Blackburn
1956:Robert F. Furchgott
1948:Stanley B. Prusiner
1940:Rolf M. Zinkernagel
1840:George H. Hitchings
1804:Joseph L. Goldstein
1764:Bengt I. Samuelsson
1593:Earl Sutherland Jr.
726:1997Natur.388..130.
454:10.1085/jgp.36.1.39
358:Stahl, F W (2001).
308:"Alfred D. Hershey"
259:The Bacteriophage λ
233:, a predecessor of
3167:Eugene van Tamelen
3156:Jean L. F. Tricart
3146:Jan Peter Toennies
3132:Peter C. Thonemann
3094:Heikki Suomalainen
3032:Stuart J. Saunders
3010:Stanley K. Runcorn
3006:Abraham J. A. Roux
2984:G. N. Ramachandran
2921:Jayant V. Narlikar
2916:Ilie G. Murgulescu
2906:Robert S. Mulliken
2865:William D. McElroy
2707:Guy Blaudin de Thé
2665:Christiaan Barnard
2268:William Kaelin Jr.
2264:Peter J. Ratcliffe
2007:Leland H. Hartwell
1896:Richard J. Roberts
1776:Barbara McClintock
1720:Godfrey Hounsfield
1621:Nikolaas Tinbergen
1553:Marshall Nirenberg
1521:Charles B. Huggins
1310:Tadeusz Reichstein
1298:AntĂłnio Egas Moniz
1093:Christiaan Eijkman
779:on Nobelprize.org
160:Alfred Day Hershey
44:Alfred Day Hershey
3205:
3204:
3182:William J. Whelan
3141:Alexander R. Todd
3136:Phillip V. Tobias
3090:Frederick Stewart
3060:Leonard T. Skeggs
2886:Desmond J. Morris
2853:Georges B. Koelle
2820:Adrian Kantrowitz
2815:Reginald V. Jones
2797:Alfred D. Hershey
2767:Kenneth O. Emergy
2713:Venancio Deulofeu
2703:Cornelis de Jager
2596:
2595:
2503:Economic Sciences
2337:
2336:
2302:Ardem Patapoutian
2151:Ralph M. Steinman
2147:Jules A. Hoffmann
2135:Robert G. Edwards
2103:Harald zur Hausen
2027:H. Robert Horvitz
1928:Eric F. Wieschaus
1864:E. Donnall Thomas
1848:J. Michael Bishop
1836:Gertrude B. Elion
1708:Hamilton O. Smith
1633:Christian de Duve
1369:Frederick Robbins
1246:Alexander Fleming
1185:Corneille Heymans
1031:Frederick Banting
936:Allvar Gullstrand
777:Alfred D. Hershey
157:
156:
120:Scientific career
77:Syosset, New York
3310:
3192:Rosalyn S. Yalow
3187:Karel F. Wiesner
3172:Ulf S. von Euler
3151:Andrzej Trautman
3076:Leonard Sosnowky
3046:Glenn T. Seaborg
3015:Donald H. Sadler
2970:Jens J. Pindborg
2956:Arthur B. Pardee
2951:George E. Palade
2926:Louis E. F. NĂ©el
2891:Giuseppe Moruzzi
2876:Jerrold Meinwald
2856:Walther Manshard
2825:Nathan O. Kaplan
2802:Gerhard Herzberg
2793:Carl G. Gahmberg
2759:Mohammed El Fasi
2694:William J. Darby
2685:Harold G. Callan
2623:
2616:
2609:
2600:
2599:
2534:
2461:Murray Gell-Mann
2364:
2357:
2350:
2341:
2340:
2330:
2315:
2304:
2289:
2283:Michael Houghton
2270:
2260:Gregg L. Semenza
2254:
2248:James P. Allison
2242:
2240:Michael W. Young
2226:
2224:Yoshinori Ohsumi
2218:
2202:
2186:
2184:Thomas C. SĂĽdhof
2170:
2158:
2137:
2129:
2123:Carol W. Greider
2113:
2097:
2081:
2069:
2057:
2045:
2033:
2017:
1990:
1974:
1966:
1950:
1942:
1936:Peter C. Doherty
1930:
1914:
1908:Alfred G. Gilman
1902:
1890:
1878:
1866:
1854:
1852:Harold E. Varmus
1842:
1826:
1818:
1806:
1794:
1778:
1770:
1754:
1738:
1728:Baruj Benacerraf
1722:
1710:
1694:
1678:
1655:
1639:
1623:
1607:
1595:
1587:
1571:
1555:
1545:Robert W. Holley
1539:
1523:
1511:
1495:
1483:
1467:
1451:
1449:Georg von Békésy
1443:
1431:
1419:
1417:Joshua Lederberg
1403:
1395:
1389:Werner Forssmann
1379:
1371:
1355:
1343:
1335:
1316:
1300:
1288:
1280:
1278:Bernardo Houssay
1264:
1256:
1240:
1228:
1216:
1215:
1209:
1208:
1202:
1201:
1195:
1187:
1179:
1171:
1159:
1151:
1135:
1127:
1115:
1107:
1105:Karl Landsteiner
1099:
1087:
1079:
1071:
1069:Johannes Fibiger
1052:
1051:
1045:
1043:Willem Einthoven
1037:
1025:
1013:
1012:
1006:
998:
990:
989:
983:
982:
976:
975:
969:
968:
962:
954:
946:
938:
930:
922:
914:
908:Élie Metchnikoff
902:
900:Alphonse Laveran
894:
882:
874:
866:
858:
850:
821:
814:
807:
798:
797:
784:
764:
763:
737:
703:
697:
696:
694:
692:
673:
667:
666:
648:
646:10.4065/79.5.590
633:Mayo Clin. Proc.
623:
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346:
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327:
303:
184:Owosso, Michigan
72:
58:Owosso, Michigan
54:December 4, 1908
53:
51:
35:
21:
20:
3318:
3317:
3313:
3312:
3311:
3309:
3308:
3307:
3208:
3207:
3206:
3201:
3163:Constantin Vago
3128:Harold Thompson
3103:Charles Tanford
3080:Roger W. Sperry
3072:George D. Snell
3068:George J. Smets
3056:Frederick Seitz
3051:Ernest R. Sears
2966:Jacques Piccard
2846:Seymour S. Kety
2841:Donald W. Kerst
2784:David J. Finney
2774:Hans J. Eysenck
2763:Ernest L. Eliel
2741:Francisco J. Dy
2727:Audouin Dollfus
2722:Richard S. Doll
2670:Colin Blakemore
2633:
2627:
2597:
2592:
2530:
2522:
2497:
2494:(United States)
2488:(United States)
2482:(United States)
2466:
2463:(United States)
2447:
2424:
2400:
2391:(Great Britain)
2375:
2368:
2338:
2333:
2318:
2307:
2292:
2287:Charles M. Rice
2279:Harvey J. Alter
2273:
2257:
2245:
2236:Michael Rosbash
2232:Jeffrey C. Hall
2229:
2221:
2205:
2196:May-Britt Moser
2189:
2173:
2168:Shinya Yamanaka
2161:
2140:
2132:
2127:Jack W. Szostak
2116:
2100:
2095:Oliver Smithies
2084:
2072:
2060:
2048:
2043:Peter Mansfield
2036:
2031:John E. Sulston
2020:
2004:
1993:
1977:
1969:
1953:
1945:
1933:
1920:Edward B. Lewis
1917:
1905:
1893:
1881:
1869:
1857:
1845:
1829:
1824:Susumu Tonegawa
1821:
1809:
1797:
1781:
1773:
1757:
1741:
1725:
1713:
1697:
1684:Roger Guillemin
1681:
1676:Daniel Gajdusek
1672:Baruch Blumberg
1669:
1658:
1649:Renato Dulbecco
1645:David Baltimore
1642:
1626:
1613:Karl von Frisch
1610:
1598:
1590:
1574:
1558:
1542:
1533:Haldan Hartline
1526:
1514:
1498:
1486:
1470:
1465:Maurice Wilkins
1454:
1446:
1434:
1429:Arthur Kornberg
1422:
1406:
1398:
1382:
1374:
1358:
1346:
1338:
1330:
1319:
1303:
1291:
1283:
1267:
1259:
1243:
1234:Joseph Erlanger
1231:
1219:
1213:
1212:
1206:
1205:
1199:
1198:
1190:
1182:
1174:
1162:
1154:
1138:
1130:
1118:
1110:
1102:
1090:
1085:Charles Nicolle
1082:
1074:
1066:
1055:
1049:
1048:
1040:
1028:
1016:
1010:
1009:
1001:
993:
987:
986:
980:
979:
973:
972:
966:
965:
957:
949:
941:
933:
928:Albrecht Kossel
925:
917:
905:
897:
885:
877:
869:
861:
853:
845:
834:
825:
773:
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704:
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688:
675:
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670:
624:
620:
610:
608:
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2789:Val L. Fitch
2750:Paul Ehrlich
2675:Aage N. Bohr
2647:Werner Arber
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2485:
2480:Max DelbrĂĽck
2389:Derek Barton
2313:Svante Pääbo
2298:David Julius
2252:Tasuku Honjo
2200:Edvard Moser
2192:John O'Keefe
2155:posthumously
2154:
2091:Martin Evans
2067:Robin Warren
2051:Richard Axel
1998:2001–present
1876:Bert Sakmann
1744:Roger Sperry
1736:George Snell
1732:Jean Dausset
1700:Werner Arber
1653:Howard Temin
1577:Bernard Katz
1564:
1561:Max DelbrĂĽck
1517:Francis Rous
1493:Feodor Lynen
1489:Konrad Bloch
1477:Alan Hodgkin
1461:James Watson
1437:Frank Burnet
1425:Severo Ochoa
1413:Edward Tatum
1401:Daniel Bovet
1314:Philip Hench
1226:Edward Doisy
1157:Hans Spemann
1145:George Minot
1125:Edgar Adrian
1113:Otto Warburg
1035:John Macleod
1004:August Krogh
996:Jules Bordet
912:Paul Ehrlich
864:Niels Finsen
848:Emil Behring
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691:February 27,
689:. Retrieved
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609:. Retrieved
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558:February 27,
556:. Retrieved
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448:(1): 39–56.
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421:November 11,
419:. Retrieved
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394:February 27,
392:. Retrieved
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144:Institutions
130:Bacteriology
119:
71:(1997-05-22)
69:May 22, 1997
18:
3223:1997 deaths
3218:1908 births
3108:Henry Taube
3028:Abdus Salam
2372:Nobel Prize
2164:John Gurdon
2079:Craig Mello
2075:Andrew Fire
1988:Eric Kandel
1964:Ferid Murad
1872:Erwin Neher
1784:Niels Jerne
1549:Har Khorana
1537:George Wald
1505:André Lwoff
1473:John Eccles
1361:John Enders
1333:Max Theiler
1294:Walter Hess
1286:Paul MĂĽller
1250:Ernst Chain
920:Emil Kocher
880:Robert Koch
872:Ivan Pavlov
856:Ronald Ross
283:ex cathedra
278:Phage group
274:Frank Stahl
224:Phage group
178:Early years
164:Nobel Prize
3212:Categories
3177:Alan Walsh
3098:Pol Swings
3003:Jean Rösch
2406:Literature
2395:Odd Hassel
2015:Paul Nurse
1349:Hans Krebs
1274:Gerty Cori
1222:Henrik Dam
1169:Otto Loewi
1165:Henry Dale
497:(1): 1–6.
318:(1): 1–6.
290:References
172:geneticist
50:1908-12-04
3160:Ioan Ursu
2421:(Ireland)
2381:Chemistry
2374:laureates
2216:Tu Youyou
1768:John Vane
1663:1976–2000
1324:1951–1975
1270:Carl Cori
1060:1926–1950
839:1901–1925
828:Laureates
760:205027895
744:0028-0836
655:0025-6196
593:0029-1420
511:0016-6731
487:"Hershey"
334:0066-4197
166:–winning
2513:(Norway)
2397:(Norway)
2011:Tim Hunt
685:Archived
663:15132399
605:Archived
552:Archived
491:Genetics
472:12981234
385:Archived
381:15202470
134:genetics
2453:Physics
830:of the
752:9217149
722:Bibcode
601:4903832
529:9584081
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342:9928472
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126:Fields
114:(1969)
109:(1958)
103:Awards
2563:1969
2430:Peace
2370:1969
756:S2CID
388:(PDF)
363:(PDF)
2588:1974
2583:1973
2578:1972
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2568:1970
2558:1968
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2434:1969
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1214:1942
1207:1941
1200:1940
1050:1925
1011:1921
988:1918
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974:1916
967:1915
748:PMID
740:ISSN
693:2023
659:PMID
651:ISSN
613:2023
597:PMID
589:ISSN
560:2023
525:PMID
507:ISSN
468:PMID
423:2023
396:2023
377:PMID
338:PMID
330:ISSN
210:and
170:and
79:, US
66:Died
60:, US
40:Born
730:doi
718:388
641:doi
515:PMC
499:doi
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458:PMC
450:doi
320:doi
138:DNA
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204:—
52:)
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