2182:
3034:
2189:
145:
121:
3028:
3937:
3943:
102:
905:. Implying support for the Australosphenida-Boreosphenida hypothesis. It may also be that both studies are right, consequently Tribosphenida would be a monophyletic group that contains the crown group Mammalia.. Australosphenida possible
1508:
886:
An alternative hypothesis is offered by
Flannery and colleagues, who view part of australosphenidans as members of Tribosphenida (excluding monotremes and others australosphenidans with
862:
cuspules but lacking a continuous mesial cingulid. Boreosphenidans also lack the triangulated trigonid on the last premolar found in Early
Cretaceous mammals. They differ from
868:(a monotreme-relative) in having the talonid placed posterior to the trigonid (like in modern tribosphenic mammals) in the lower molars, but upper molars similar to those of
892:, which they consider to be an unrelated group), with Barremian-Aptian Australian Bisphopidae as the sister taxon to all Northern Hemisphere Theria (Boreosphenida).
1061:
Rich, T. H.; Hopson, J. A.; Musser, A. M.; Flannery, T. F.; Vickers-Rich, P. (2005). "Independent origins of middle ear bones in monotremes and therians".
2969:
3663:
3563:
3246:
2962:
2948:
1775:
2899:
2779:
2688:
3703:
3502:
3356:
3160:
2906:
2764:
2625:
963:
3823:
3783:
3363:
3349:
3253:
3210:
2941:
2920:
2913:
2855:
2512:
2491:
2379:
3723:
3390:
3327:
3305:
3232:
3217:
3196:
2639:
2449:
321:
4004:
3813:
3634:
3445:
3438:
3284:
2862:
2841:
2520:
2409:
2399:
2329:
1577:
1500:
3203:
2934:
2588:
2359:
1852:
270:
3984:
3773:
3753:
3459:
3298:
2927:
2505:
2419:
2369:
1516:
311:
3803:
3522:
3167:
2834:
2786:
2595:
3573:
410:
1176:
Flannery, Timothy F.; Rich, Thomas H.; Vickers-Rich, Patricia; Ziegler, Tim; Veatch, E. Grace; Helgen, Kristofer M. (2022-01-02).
2181:
1153:
991:
1484:
1337:"A diverse assemblage of monotremes (Monotremata) from the Cenomanian Lightning Ridge fauna of New South Wales, Australia"
3033:
1556:
1468:
1217:
Flannery, Timothy F.; Rich, Thomas H.; Vickers-Rich, Patricia; Veatch, E. Grace; Helgen, Kristofer M. (2022-11-01).
1269:
Mao, Fangyuan; Li, Zhiyu; Wang, Zhili; Zhang, Chi; Rich, Thomas; Vickers-Rich, Patricia; Meng, Jin (2024-04-03).
2188:
382:) molars. Since then, other unrelated mammal groups have been found to have tribosphenic molars, such as the
1426:
1376:"Conflict Resolution for Mesozoic Mammals: Reconciling Phylogenetic Incongruence Among Anatomical Regions"
1018:
Luo, Zhe-Xi; Cifelli, Richard L.; Kielan-Jaworowska, Zofia (2001). "Dual origin of tribosphenic mammals".
144:
1218:
4191:
858:.) They share the tribosphenic molars with the Australosphenida but differ from them by having
4186:
4163:
4126:
1549:
3027:
1524:
4158:
2653:
1332:
957:
839:
1143:
4030:
3936:
3312:
2097:
1341:
1282:
1442:
379:
8:
3901:
2138:
2104:
1683:
831:
1469:"MESOZOIC MAMMALS; Stem zatherians, zatherians & Peramuridae, an internet directory"
1286:
3475:
1978:
1828:
1656:
1402:
1375:
1248:
1086:
1043:
843:
835:
261:
139:
3942:
3872:
2825:
2005:
1542:
1446:
1407:
1356:
1306:
1298:
1270:
1252:
1240:
1199:
1149:
1121:
1078:
1035:
987:
927:
895:
On the other hand, the study that cites
Flannery's article, given in 2024, maintains
455:
378:
Tribosphenid mammals were originally grouped on the basis of triangular or V-shaped (
3291:
3098:
2883:
2482:
2144:
1844:
1837:
1748:
1438:
1397:
1387:
1346:
1290:
1230:
1189:
1117:
1113:
1090:
1070:
1047:
1027:
943:
880:
859:
855:
847:
515:
383:
132:
1351:
1336:
1235:
1194:
1177:
3374:
2869:
2741:
2710:
2695:
2609:
2429:
1919:
1880:
1765:
1492:
1139:
925:
897:
430:
252:
409:. They are also united by postcranial features such as the presence of a modern
401:, is still identifiable, united by characteristics such as the lack of a mesial
397:
However, a clade between the aforementioned groups, the "true
Tribosphenida" or
4096:
4061:
3971:
3532:
3404:
3383:
3319:
3271:
3188:
3112:
3083:
2976:
2955:
2726:
2719:
2548:
2439:
2339:
2056:
1625:
1294:
440:
4180:
4149:
4091:
4086:
4081:
4066:
3994:
3743:
3593:
2797:
2733:
2277:
2090:
2080:
1950:
1450:
1392:
1360:
1302:
1271:"Jurassic shuotheriids show earliest dental diversification of mammaliaforms"
1244:
1203:
1125:
851:
665:
611:
583:
569:
557:
398:
347:
292:
59:
1074:
879:(~145-140 mya). They were restricted to the Northern Hemisphere during the
4071:
3793:
3649:
3512:
3429:
3411:
3239:
3153:
3137:
2755:
2646:
2632:
2318:
2220:
2207:
2156:
2028:
1807:
1798:
1476:
1411:
1324:
1310:
1082:
1039:
948:
531:
497:
391:
235:
883:, but spread to South America and India during the end of the Cretaceous.
394:
this is fundamentally useless as it evolved multiple times among mammals.
366:. The current definition of Tribosphenida is more or less synonymous with
4101:
4076:
4051:
3878:
3713:
3642:
3625:
3603:
3543:
3397:
3342:
3174:
3070:
3051:
2994:
2983:
2892:
2681:
2618:
2578:
2564:
2534:
2527:
2498:
2389:
2349:
2307:
2267:
2257:
2017:
1967:
864:
715:
696:
639:
596:
485:
481:
34:
429:
showing one hypothesis of mammal relationships based on Rowe (1988) and
4056:
4024:
4014:
3890:
3850:
3693:
3583:
3181:
3105:
3091:
2848:
2674:
2660:
2557:
2459:
2297:
2237:
2126:
1936:
1928:
1897:
1890:
1814:
1104:
Rowe, Timothy (1988). "Definition, diagnosis, and origin of
Mammalia".
932:
from
Scotland, and posterior dentary morphology in early cladotherians"
888:
876:
692:
543:
511:
79:
44:
413:(though this too has evolved independently in many other groups, like
120:
3907:
3610:
3335:
3225:
3146:
3126:
2810:
2667:
2467:
2039:
1959:
1868:
1783:
1641:
1607:
1598:
1031:
906:
902:
426:
414:
387:
359:
355:
156:
84:
4120:
984:
Mammals from the Age of
Dinosaurs: Origins, Evolution, and Structure
4143:
3861:
3763:
3553:
3492:
3119:
2247:
2046:
1905:
1668:
1616:
1589:
1328:
711:
624:
467:
406:
402:
363:
245:
199:
176:
128:
74:
69:
54:
49:
39:
982:
Kielan-Jaworowska, Zofia; Cifelli, Richard L; Luo, Zhe-Xi (2004).
342:
is a group (infralegion) of mammals that includes the ancestor of
4035:
3656:
3452:
2772:
2287:
2068:
1994:
827:
417:), modern shoulder blades, and several features of the hindlimb.
89:
64:
3834:
3733:
1736:
1695:
1583:
1565:
1323:
1175:
678:
351:
327:
301:
223:
186:
166:
110:
1216:
1427:"Developmental Patterns in Mesozoic Evolution of Mammal Ears"
1534:
1017:
981:
1335:; Smith, Elizabeth T.; Helgen, Kristofer M. (2024-05-26).
926:
Panciroli E; Roger B.J. Benson; Richard J. Butler (2018).
101:
1060:
977:
975:
973:
1374:
Celik, Mรฉlina A.; Phillips, Matthew J. (8 July 2020).
362:
plus all of its descendants). It belongs to the group
970:
4020:
4010:
4000:
3990:
3980:
3846:
3819:
3809:
3799:
3789:
3779:
3769:
3759:
3749:
3739:
3729:
3719:
3709:
3699:
3689:
3622:
3599:
3589:
3579:
3569:
3559:
3549:
3539:
3528:
3518:
3508:
3498:
3488:
3426:
3268:
3067:
2880:
2822:
2752:
2707:
2606:
2575:
2545:
2479:
2455:
2445:
2435:
2425:
2415:
2405:
2395:
2385:
2375:
2365:
2355:
2345:
2335:
2325:
2314:
2303:
2293:
2283:
2273:
2263:
2253:
2243:
2233:
2123:
2086:
2076:
2052:
2035:
2024:
2013:
1990:
1947:
1916:
1877:
1825:
1795:
1771:
1761:
1431:
Annual Review of
Ecology, Evolution, and Systematics
1342:
1223:
1182:
Alcheringa: An Australasian Journal of Palaeontology
838:(three-cusped cheek teeth). In boreosphenidans, the
317:
307:
297:
289:
267:
258:
241:
232:
1219:"The Gondwanan Origin of Tribosphenida (Mammalia)"
1145:Classification of Mammals Above the Species Level
928:"New partial dentaries of amphitheriid mammalian
846:(hole in the mandible) is absent (in contrast to
4178:
1747:
1501:"Tribosphenida (after McKenna & Bell, 1997)"
1178:"A review of monotreme (Monotremata) evolution"
1268:
1132:
1550:
1373:
1317:
1148:. Columbia University Press. pp. 9โ10.
962:: CS1 maint: multiple names: authors list (
2218:
1138:
405:and of a triangulated trigonid on the last
1557:
1543:
119:
100:
1401:
1391:
1350:
1234:
1193:
947:
1640:
919:
875:The oldest boreosphenidans are from the
842:is placed posteriorly and the primitive
386:(a group that includes the still extant
1097:
986:. New York: Columbia University Press.
4179:
4125:
4124:
3958:
3049:
2205:
1734:
1576:
1538:
1511:from the original on August 22, 2023.
1443:10.1146/annurev-ecolsys-032511-142302
1013:
1011:
1009:
1007:
1005:
1003:
1264:
1262:
1103:
1424:
13:
1418:
1106:Journal of Vertebrate Paleontology
1000:
373:
14:
4203:
1461:
1259:
1210:
3941:
3935:
3032:
3026:
2206:
2187:
2180:
810:
143:
32:
3050:
1367:
3959:
1169:
1118:10.1080/02724634.1988.10011708
1054:
1:
1352:10.1080/03115518.2024.2348753
1236:10.1080/03115518.2022.2132288
1195:10.1080/03115518.2022.2025900
936:Acta Palaeontologica Polonica
912:
354:(the last common ancestor of
1735:
1564:
1491:(in Russian). Archived from
420:
7:
1517:"Infralegion Tribosphenida"
10:
4208:
1425:Luo, Zhe-Xi (2011-12-01).
1295:10.1038/s41586-024-07258-7
4133:
4044:
3969:
3965:
3954:
3932:
3889:
3860:
3842:
3833:
3685:
3674:
3621:
3484:
3473:
3425:
3373:
3279:
3267:
3136:
3078:
3066:
3062:
3058:
3045:
3023:
2993:
2879:
2821:
2805:
2796:
2751:
2706:
2605:
2574:
2544:
2478:
2229:
2214:
2201:
2177:
2155:
2122:
2067:
2004:
1986:
1977:
1946:
1915:
1876:
1867:
1824:
1794:
1757:
1743:
1730:
1694:
1676:
1667:
1649:
1636:
1572:
1505:Mikko's Phylogeny Archive
1142:; Bell, Susan K. (1997).
709:
690:
683:
662:
655:
636:
629:
608:
601:
580:
573:
554:
547:
528:
508:
501:
479:
472:
452:
445:
229:
222:
140:Scientific classification
138:
127:
118:
108:
99:
23:
1393:10.3389/fgene.2020.00651
390:), suggesting that as a
1327:; McCurry, Matthew R.;
1075:10.1126/science.1105717
930:Palaeoxonodon ooliticus
830:that originated in the
30:Early CretaceousโRecent
1485:"THERIA ะ MARSUPIONTA"
1333:Vickers-Rich, Patricia
949:10.4202/app.00434.2017
826:, "wedge") were early
822:, "northern wind" and
16:Infralegion of mammals
4159:Paleobiology Database
1380:Frontiers in Genetics
4031:Saint Bathans Mammal
1495:on October 31, 2013.
1325:Flannery, Timothy F.
1287:2024Natur.628..569M
1140:McKenna, Malcolm C.
836:tribosphenic molars
832:Northern Hemisphere
3476:Prototribosphenida
1979:Ornithorhynchoidea
1829:Ausktribosphenidae
844:postdentary trough
384:australosphenidans
262:Ausktribosphenidae
4174:
4173:
4127:Taxon identifiers
4118:
4117:
4114:
4113:
4110:
4109:
3950:
3949:
3930:
3929:
3926:
3925:
3922:
3921:
3918:
3917:
3469:
3468:
3421:
3420:
3263:
3262:
3041:
3040:
3021:
3020:
3017:
3016:
3013:
3012:
2970:Symmetrodontoides
2826:Zhangheotheriidae
2197:
2196:
2175:
2174:
2145:multituberculates
2118:
2117:
2114:
2113:
2006:Ornithorhynchidae
1863:
1862:
1726:
1725:
1722:
1721:
1718:
1717:
1714:
1713:
1527:on March 3, 2016.
1479:on June 18, 2013.
1281:(8008): 569โ575.
1155:978-0-231-11012-9
1069:(5711): 910โ914.
993:978-0-231-11918-4
901:as a relative of
807:
806:
798:
797:
789:
788:
780:
779:
771:
770:
762:
761:
753:
752:
744:
743:
735:
734:
726:
725:
456:Morganucodontidae
433:and Bell (1997):
337:
336:
286:
218:
4199:
4167:
4166:
4154:
4153:
4152:
4122:
4121:
4022:
4012:
4002:
3992:
3982:
3967:
3966:
3956:
3955:
3945:
3940:
3939:
3848:
3840:
3839:
3821:
3811:
3801:
3791:
3781:
3771:
3761:
3751:
3741:
3731:
3721:
3711:
3701:
3691:
3683:
3682:
3664:Tendagurutherium
3624:
3601:
3591:
3581:
3571:
3561:
3551:
3541:
3530:
3520:
3510:
3500:
3490:
3482:
3481:
3428:
3292:Austrotriconodon
3277:
3276:
3270:
3247:Portopinheirodon
3099:Drescheratherium
3076:
3075:
3069:
3064:
3063:
3060:
3059:
3047:
3046:
3036:
3031:
3030:
2963:Spalacotheroides
2949:Spalacotheridium
2884:Spalacotheriidae
2882:
2824:
2803:
2802:
2754:
2709:
2608:
2577:
2547:
2483:Gobiconodontidae
2481:
2457:
2447:
2437:
2427:
2417:
2407:
2397:
2387:
2377:
2367:
2357:
2347:
2337:
2327:
2316:
2305:
2295:
2285:
2275:
2265:
2255:
2245:
2235:
2227:
2226:
2216:
2215:
2203:
2202:
2192:
2191:
2185:
2184:
2125:
2088:
2078:
2054:
2037:
2026:
2015:
1992:
1984:
1983:
1949:
1918:
1879:
1874:
1873:
1838:Ausktribosphenos
1827:
1797:
1776:Tendagurutherium
1773:
1763:
1755:
1754:
1749:Australosphenida
1745:
1744:
1732:
1731:
1674:
1673:
1647:
1646:
1638:
1637:
1631:
1630:
1574:
1573:
1559:
1552:
1545:
1536:
1535:
1528:
1523:. Archived from
1512:
1496:
1480:
1475:. Archived from
1455:
1454:
1422:
1416:
1415:
1405:
1395:
1371:
1365:
1364:
1354:
1321:
1315:
1314:
1266:
1257:
1256:
1238:
1229:(3โ4): 277โ290.
1214:
1208:
1207:
1197:
1173:
1167:
1166:
1164:
1162:
1136:
1130:
1129:
1101:
1095:
1094:
1058:
1052:
1051:
1032:10.1038/35051023
1015:
998:
997:
979:
968:
967:
961:
953:
951:
923:
881:Early Cretaceous
856:Australosphenida
848:Kuehneotheriidae
840:mandibular angle
686:
685:
658:
657:
632:
631:
604:
603:
576:
575:
550:
549:
516:Multituberculata
504:
503:
475:
474:
448:
447:
437:
436:
319:
309:
299:
291:
280:
269:
260:
243:
234:
216:
148:
147:
123:
104:
94:
31:
27:Temporal range:
21:
20:
4207:
4206:
4202:
4201:
4200:
4198:
4197:
4196:
4192:Mammal taxonomy
4177:
4176:
4175:
4170:
4162:
4157:
4148:
4147:
4142:
4129:
4119:
4106:
4040:
3961:
3946:
3934:
3914:
3885:
3856:
3829:
3677:
3676:Tribosphenida /
3670:
3617:
3478:
3465:
3417:
3375:Mesungulatoidea
3369:
3259:
3132:
3054:
3037:
3025:
3009:
2989:
2875:
2870:Zhangheotherium
2817:
2792:
2780:Trishulotherium
2747:
2742:Volaticotherium
2702:
2689:Victoriaconodon
2610:Triconodontidae
2601:
2570:
2540:
2474:
2430:Phascolotherium
2223:
2210:
2193:
2186:
2179:
2171:
2151:
2110:
2063:
2047:Ornithorhynchus
2000:
1973:
1942:
1920:Steropodontidae
1911:
1881:Kollikodontidae
1859:
1820:
1790:
1739:
1710:
1690:
1663:
1632:
1579:
1578:
1568:
1563:
1532:
1515:
1499:
1483:
1467:
1464:
1459:
1458:
1423:
1419:
1372:
1368:
1329:Rich, Thomas H.
1322:
1318:
1267:
1260:
1215:
1211:
1174:
1170:
1160:
1158:
1156:
1137:
1133:
1102:
1098:
1059:
1055:
1026:(6816): 53โ57.
1016:
1001:
994:
980:
971:
955:
954:
924:
920:
915:
813:
808:
799:
790:
781:
772:
763:
754:
745:
736:
727:
423:
376:
374:Characteristics
253:Monotremaformes
215:
142:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
24:Tribosphenidans
17:
12:
11:
5:
4205:
4195:
4194:
4189:
4172:
4171:
4169:
4168:
4155:
4139:
4137:
4131:
4130:
4116:
4115:
4112:
4111:
4108:
4107:
4105:
4104:
4099:
4097:Amphitheriidae
4094:
4089:
4084:
4079:
4074:
4069:
4064:
4062:Eutriconodonta
4059:
4054:
4048:
4046:
4042:
4041:
4039:
4038:
4033:
4028:
4018:
4008:
3998:
3988:
3977:
3975:
3972:Incertae sedis
3963:
3962:
3952:
3951:
3948:
3947:
3933:
3931:
3928:
3927:
3924:
3923:
3920:
3919:
3916:
3915:
3913:
3912:
3911:
3910:
3904:
3895:
3893:
3887:
3886:
3884:
3883:
3882:
3881:
3875:
3866:
3864:
3858:
3857:
3855:
3854:
3843:
3837:
3831:
3830:
3828:
3827:
3817:
3807:
3797:
3787:
3777:
3767:
3757:
3747:
3737:
3727:
3717:
3707:
3704:Argaliatherium
3697:
3686:
3680:
3672:
3671:
3669:
3668:
3660:
3653:
3646:
3639:
3630:
3628:
3619:
3618:
3616:
3615:
3607:
3597:
3587:
3577:
3567:
3557:
3547:
3537:
3533:Brancatherulum
3526:
3516:
3506:
3503:Amphibetulimus
3496:
3485:
3479:
3474:
3471:
3470:
3467:
3466:
3464:
3463:
3456:
3449:
3442:
3434:
3432:
3423:
3422:
3419:
3418:
3416:
3415:
3408:
3405:Peligrotherium
3401:
3394:
3387:
3384:Coloniatherium
3379:
3377:
3371:
3370:
3368:
3367:
3360:
3357:Quirogatherium
3353:
3346:
3339:
3332:
3324:
3320:Groebertherium
3316:
3309:
3302:
3295:
3288:
3280:
3274:
3272:Meridiolestida
3265:
3264:
3261:
3260:
3258:
3257:
3250:
3243:
3236:
3229:
3222:
3214:
3207:
3200:
3193:
3189:Groebertherium
3185:
3178:
3171:
3164:
3161:Anthracolestes
3157:
3150:
3142:
3140:
3134:
3133:
3131:
3130:
3123:
3116:
3113:Henkelotherium
3109:
3102:
3095:
3088:
3084:Brancatherulum
3079:
3073:
3056:
3055:
3043:
3042:
3039:
3038:
3024:
3022:
3019:
3018:
3015:
3014:
3011:
3010:
3008:
3007:
3006:
3005:
2999:
2997:
2991:
2990:
2988:
2987:
2980:
2977:Symmetrolestes
2973:
2966:
2959:
2956:Spalacotherium
2952:
2945:
2938:
2931:
2924:
2917:
2910:
2907:Cifellitherium
2903:
2896:
2888:
2886:
2877:
2876:
2874:
2873:
2866:
2859:
2852:
2845:
2838:
2830:
2828:
2819:
2818:
2816:
2815:
2806:
2800:
2794:
2793:
2791:
2790:
2783:
2776:
2769:
2765:Gobiotheriodon
2760:
2758:
2749:
2748:
2746:
2745:
2738:
2730:
2727:Ichthyoconodon
2723:
2720:Argentoconodon
2715:
2713:
2711:Volaticotheria
2704:
2703:
2701:
2700:
2696:Volaticotheria
2692:
2685:
2678:
2671:
2664:
2657:
2650:
2643:
2636:
2629:
2626:Arundelconodon
2622:
2614:
2612:
2603:
2602:
2600:
2599:
2592:
2584:
2582:
2572:
2571:
2569:
2568:
2561:
2553:
2551:
2549:Jeholodentidae
2542:
2541:
2539:
2538:
2531:
2524:
2517:
2509:
2502:
2495:
2487:
2485:
2476:
2475:
2473:
2472:
2464:
2453:
2443:
2440:Sangarotherium
2433:
2423:
2413:
2403:
2393:
2383:
2373:
2363:
2353:
2343:
2340:Indotriconodon
2333:
2323:
2312:
2301:
2291:
2281:
2271:
2261:
2251:
2241:
2230:
2224:
2219:
2212:
2211:
2199:
2198:
2195:
2194:
2178:
2176:
2173:
2172:
2170:
2169:
2168:
2167:
2161:
2159:
2153:
2152:
2150:
2149:
2148:
2147:
2141:
2132:
2130:
2120:
2119:
2116:
2115:
2112:
2111:
2109:
2108:
2101:
2094:
2084:
2073:
2071:
2069:Tachyglossidae
2065:
2064:
2062:
2061:
2057:Patagorhynchus
2050:
2043:
2033:
2022:
2010:
2008:
2002:
2001:
1999:
1998:
1987:
1981:
1975:
1974:
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1626:Mammaliaformes
1619:
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1570:
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1554:
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1462:External links
1460:
1457:
1456:
1437:(1): 355โ380.
1417:
1366:
1316:
1258:
1209:
1168:
1154:
1131:
1112:(3): 241โ264.
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441:Mammaliaformes
435:
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38:
33:
26:
15:
9:
6:
4:
3:
2:
4204:
4193:
4190:
4188:
4187:Tribosphenida
4185:
4184:
4182:
4165:
4160:
4156:
4151:
4145:
4141:
4140:
4138:
4136:
4135:Tribosphenida
4132:
4128:
4123:
4103:
4100:
4098:
4095:
4093:
4092:Paurodontidae
4090:
4088:
4087:Eupantotheria
4085:
4083:
4082:Symmetrodonta
4080:
4078:
4075:
4073:
4070:
4068:
4067:Amphilestidae
4065:
4063:
4060:
4058:
4055:
4053:
4050:
4049:
4047:
4043:
4037:
4034:
4032:
4029:
4027:
4026:
4019:
4017:
4016:
4009:
4007:
4006:
3999:
3997:
3996:
3995:Chronoperates
3989:
3987:
3986:
3979:
3978:
3976:
3974:
3973:
3968:
3964:
3957:
3953:
3944:
3938:
3909:
3905:
3903:
3899:
3898:
3897:
3896:
3894:
3892:
3888:
3880:
3876:
3874:
3870:
3869:
3868:
3867:
3865:
3863:
3859:
3853:
3852:
3845:
3844:
3841:
3838:
3836:
3832:
3826:
3825:
3824:Tribactonodon
3818:
3816:
3815:
3808:
3806:
3805:
3798:
3796:
3795:
3788:
3786:
3785:
3784:Kielantherium
3778:
3776:
3775:
3768:
3766:
3765:
3758:
3756:
3755:
3748:
3746:
3745:
3744:Holoclemensia
3738:
3736:
3735:
3728:
3726:
3725:
3718:
3716:
3715:
3708:
3706:
3705:
3698:
3696:
3695:
3688:
3687:
3684:
3681:
3679:
3678:Boreosphenida
3673:
3666:
3665:
3661:
3659:
3658:
3654:
3652:
3651:
3647:
3645:
3644:
3640:
3637:
3636:
3632:
3631:
3629:
3627:
3620:
3613:
3612:
3608:
3606:
3605:
3598:
3596:
3595:
3594:Palaeoxonodon
3588:
3586:
3585:
3578:
3576:
3575:
3568:
3566:
3565:
3558:
3556:
3555:
3548:
3546:
3545:
3538:
3535:
3534:
3527:
3525:
3524:
3517:
3515:
3514:
3507:
3505:
3504:
3497:
3495:
3494:
3487:
3486:
3483:
3480:
3477:
3472:
3462:
3461:
3457:
3455:
3454:
3450:
3448:
3447:
3443:
3441:
3440:
3436:
3435:
3433:
3431:
3424:
3414:
3413:
3409:
3407:
3406:
3402:
3400:
3399:
3395:
3393:
3392:
3388:
3386:
3385:
3381:
3380:
3378:
3376:
3372:
3366:
3365:
3364:Solanutherium
3361:
3359:
3358:
3354:
3352:
3351:
3350:Paraungulatum
3347:
3345:
3344:
3340:
3338:
3337:
3333:
3330:
3329:
3325:
3322:
3321:
3317:
3315:
3314:
3310:
3308:
3307:
3303:
3301:
3300:
3296:
3294:
3293:
3289:
3287:
3286:
3282:
3281:
3278:
3275:
3273:
3266:
3256:
3255:
3254:Treslagosodon
3251:
3249:
3248:
3244:
3242:
3241:
3237:
3235:
3234:
3230:
3228:
3227:
3223:
3220:
3219:
3215:
3213:
3212:
3211:Krebsotherium
3208:
3206:
3205:
3201:
3199:
3198:
3194:
3191:
3190:
3186:
3184:
3183:
3179:
3177:
3176:
3172:
3170:
3169:
3165:
3163:
3162:
3158:
3156:
3155:
3151:
3149:
3148:
3144:
3143:
3141:
3139:
3135:
3129:
3128:
3124:
3122:
3121:
3117:
3115:
3114:
3110:
3108:
3107:
3103:
3101:
3100:
3096:
3094:
3093:
3089:
3086:
3085:
3081:
3080:
3077:
3074:
3072:
3065:
3061:
3057:
3053:
3048:
3044:
3035:
3029:
3003:
3002:
3001:
3000:
2998:
2996:
2992:
2986:
2985:
2981:
2979:
2978:
2974:
2972:
2971:
2967:
2965:
2964:
2960:
2958:
2957:
2953:
2951:
2950:
2946:
2944:
2943:
2942:Spalacolestes
2939:
2937:
2936:
2932:
2930:
2929:
2925:
2923:
2922:
2921:Infernolestes
2918:
2916:
2915:
2914:Heishanlestes
2911:
2909:
2908:
2904:
2902:
2901:
2897:
2895:
2894:
2890:
2889:
2887:
2885:
2878:
2872:
2871:
2867:
2865:
2864:
2860:
2858:
2857:
2856:Ningchengodon
2853:
2851:
2850:
2846:
2844:
2843:
2839:
2837:
2836:
2832:
2831:
2829:
2827:
2820:
2813:
2812:
2808:
2807:
2804:
2801:
2799:
2798:Trechnotheria
2795:
2789:
2788:
2784:
2782:
2781:
2777:
2775:
2774:
2770:
2767:
2766:
2762:
2761:
2759:
2757:
2750:
2744:
2743:
2739:
2736:
2735:
2734:Triconolestes
2731:
2729:
2728:
2724:
2722:
2721:
2717:
2716:
2714:
2712:
2705:
2698:
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2637:
2635:
2634:
2630:
2628:
2627:
2623:
2621:
2620:
2616:
2615:
2613:
2611:
2604:
2598:
2597:
2593:
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2590:
2586:
2585:
2583:
2580:
2573:
2567:
2566:
2562:
2560:
2559:
2555:
2554:
2552:
2550:
2543:
2537:
2536:
2532:
2530:
2529:
2525:
2523:
2522:
2518:
2515:
2514:
2513:Huasteconodon
2510:
2508:
2507:
2503:
2501:
2500:
2496:
2494:
2493:
2492:Fuxinoconodon
2489:
2488:
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2484:
2477:
2470:
2469:
2465:
2462:
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2454:
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2412:
2411:
2404:
2402:
2401:
2394:
2392:
2391:
2384:
2382:
2381:
2380:Kryptotherium
2374:
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2364:
2362:
2361:
2354:
2352:
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2344:
2342:
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2334:
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2320:
2313:
2310:
2309:
2302:
2300:
2299:
2292:
2290:
2289:
2282:
2280:
2279:
2278:Chaoyangodens
2272:
2270:
2269:
2262:
2260:
2259:
2252:
2250:
2249:
2242:
2240:
2239:
2232:
2231:
2228:
2225:
2222:
2217:
2213:
2209:
2204:
2200:
2190:
2183:
2165:
2164:
2163:
2162:
2160:
2158:
2154:
2146:
2142:
2140:
2136:
2135:
2134:
2133:
2131:
2128:
2121:
2107:
2106:
2102:
2100:
2099:
2095:
2093:
2092:
2091:Murrayglossus
2085:
2083:
2082:
2081:Megalibgwilia
2075:
2074:
2072:
2070:
2066:
2059:
2058:
2051:
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2041:
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2019:
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2011:
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1996:
1989:
1988:
1985:
1982:
1980:
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1969:
1965:
1962:
1961:
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1956:
1954:
1952:
1951:Teinolophidae
1945:
1939:
1938:
1934:
1931:
1930:
1926:
1925:
1923:
1921:
1914:
1908:
1907:
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1123:
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893:
891:
890:
884:
882:
878:
873:
871:
867:
866:
861:
857:
853:
852:Eupantotheria
849:
845:
841:
837:
833:
829:
825:
821:
817:
816:Boreosphenida
811:Boreosphenida
803:
802:
794:
793:
785:
784:
776:
775:
767:
766:
758:
757:
749:
748:
740:
739:
731:
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722:
721:
718:
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713:
707:
706:
703:
702:
699:
698:
694:
688:
687:
680:
676:
675:
672:
671:
668:
667:
666:Aegialodontia
660:
659:
653:
652:Tribosphenida
650:
649:
646:
645:
642:
641:
634:
633:
626:
622:
621:
618:
617:
614:
613:
612:Dryolestoidea
606:
605:
598:
594:
593:
590:
589:
586:
585:
584:Symmetrodonta
578:
577:
571:
570:Trechnotheria
568:
567:
564:
563:
560:
559:
558:Kuehneotheria
552:
551:
545:
542:
541:
538:
537:
534:
533:
526:
525:
522:
521:
518:
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513:
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477:
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450:
449:
442:
439:
438:
434:
432:
428:
418:
416:
412:
408:
404:
400:
399:Boreosphenida
395:
393:
389:
385:
381:
371:
369:
368:Boreosphenida
365:
361:
357:
353:
349:
348:Aegialodontia
345:
341:
340:Tribosphenida
329:
326:
324:
323:
316:
314:
313:
306:
304:
303:
296:
294:
293:Aegialodontia
288:
287:
284:
278:
277:Boreosphenida
275:
272:
266:
263:
257:
254:
251:
248:
247:
240:
237:
231:
230:
228:
225:
221:
217:McKenna, 1975
214:
213:Tribosphenida
211:
208:
205:
204:
201:
198:
195:
192:
191:
188:
185:
182:
181:
178:
175:
172:
171:
168:
165:
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146:
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137:
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126:
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117:
113:
112:
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
4134:
4072:Theriiformes
4023:
4013:
4003:
3993:
3983:
3970:
3849:
3822:
3812:
3802:
3794:Pappotherium
3792:
3782:
3772:
3762:
3752:
3742:
3732:
3724:Carinalestes
3722:
3712:
3702:
3692:
3675:
3662:
3655:
3650:Peramuroides
3648:
3641:
3633:
3609:
3602:
3592:
3582:
3572:
3562:
3552:
3542:
3531:
3521:
3513:Amphitherium
3511:
3501:
3491:
3458:
3451:
3444:
3437:
3430:Donodontidae
3412:Reigitherium
3410:
3403:
3396:
3391:Mesungulatum
3389:
3382:
3362:
3355:
3348:
3341:
3334:
3328:Lakotalestes
3326:
3318:
3311:
3306:Casamiquelia
3304:
3297:
3290:
3283:
3252:
3245:
3240:Phascolestes
3238:
3233:Minutolestes
3231:
3224:
3218:Lakotalestes
3216:
3209:
3202:
3197:Guimarotodus
3195:
3187:
3180:
3173:
3166:
3159:
3154:Amblotherium
3152:
3145:
3138:Dryolestidae
3125:
3118:
3111:
3104:
3097:
3090:
3082:
2982:
2975:
2968:
2961:
2954:
2947:
2940:
2933:
2926:
2919:
2912:
2905:
2898:
2891:
2868:
2861:
2854:
2847:
2840:
2833:
2809:
2785:
2778:
2771:
2763:
2756:Tinodontidae
2740:
2732:
2725:
2718:
2694:
2687:
2680:
2673:
2666:
2659:
2652:
2647:Eotriconodon
2645:
2640:Corviconodon
2638:
2633:Astroconodon
2631:
2624:
2617:
2594:
2587:
2563:
2556:
2533:
2526:
2519:
2511:
2504:
2497:
2490:
2466:
2458:
2450:Tendagurodon
2448:
2438:
2428:
2418:
2408:
2398:
2388:
2378:
2368:
2358:
2348:
2338:
2328:
2319:Fruitafossor
2317:
2306:
2296:
2286:
2276:
2266:
2256:
2246:
2236:
2221:Theriimorpha
2208:Theriimorpha
2157:Theriimorpha
2103:
2098:Tachyglossus
2096:
2089:
2079:
2055:
2045:
2038:
2029:Monotrematum
2027:
2016:
1993:
1966:
1958:
1935:
1927:
1904:
1896:
1889:
1851:
1843:
1836:
1813:
1808:Asfaltomylos
1806:
1799:Henosferidae
1782:
1774:
1764:
1621:
1612:
1603:
1594:
1531:
1525:the original
1520:
1504:
1493:the original
1488:
1477:the original
1472:
1434:
1430:
1420:
1383:
1379:
1369:
1340:
1319:
1278:
1274:
1226:
1222:
1212:
1185:
1181:
1171:
1159:. Retrieved
1144:
1134:
1109:
1105:
1099:
1066:
1062:
1056:
1023:
1019:
983:
958:cite journal
939:
935:
929:
921:
896:
894:
887:
885:
874:
869:
863:
823:
819:
815:
814:
710:
691:
663:
651:
637:
609:
581:
555:
532:Triconodonta
529:
509:
498:Theriiformes
480:
453:
424:
396:
392:synapomorphy
380:tribosphenic
377:
367:
343:
339:
338:
322:Tribotherium
320:
310:
300:
282:
276:
244:
236:Henosferidae
212:
206:
193:
109:
18:
4102:Marsupionta
4077:Pantotheria
4052:Prototheria
4005:Microderson
3814:Tirotherium
3714:Brasilestes
3643:Kouriogenys
3635:Afriquiamus
3626:Peramuridae
3611:Monotremata
3604:Vincelestes
3544:Chunnelodon
3446:Anoualestes
3439:Amazighodon
3398:Orretherium
3343:Necrolestes
3285:Amarillodon
3175:Crusafontia
3071:Dryolestida
3052:Cladotheria
2995:Cladotheria
2984:Yaverlestes
2893:Akidolestes
2863:Origolestes
2842:Kiyatherium
2811:Monotremata
2682:Trioracodon
2619:Alticonodon
2579:Klameliidae
2565:Yanoconodon
2535:Spinolestes
2528:Repenomamus
2521:Meemannodon
2499:Gobiconodon
2468:Monotremata
2410:Manchurodon
2400:Liaotherium
2390:Liaoconodon
2350:Juchilestes
2330:Hakusanodon
2308:Dyskritodon
2268:Aploconodon
2258:Amphilestes
2018:Dharragarra
1968:Teinolophos
1869:Monotremata
1784:Monotremata
1188:(1): 3โ20.
903:monotremata
870:Shuotherium
865:Shuotherium
716:Placentalia
697:Marsupialia
640:Peramuridae
597:Cladotheria
486:Monotremata
482:Prototheria
425:Below is a
4181:Categories
4057:Yinotheria
4045:Other taxa
4025:Tingamarra
4015:Thereuodon
3960:Other taxa
3908:marsupials
3902:Metatheria
3891:Metatheria
3879:placentals
3851:Patagomaia
3694:Aegialodon
3584:Nanolestes
3204:Hercynodon
3182:Dryolestes
3106:Euthlastus
3092:Dorsetodon
3004:see belowโ
2935:Shalbaatar
2849:Maotherium
2675:Triconodon
2661:Meiconodon
2589:Ferganodon
2558:Jeholodens
2460:Allotheria
2360:Jueconodon
2298:Condorodon
2238:Acinacodus
2166:see belowโ
2139:Allotheria
2127:Allotheria
1937:Steropodon
1929:Parvopalus
1898:Kryoryctes
1891:Kollikodon
1853:Kryoparvus
1815:Henosferus
1705:see belowโ
1684:Cynodontia
1669:Cynodontia
1617:Cynodontia
1608:Therapsida
1473:Home.arcor
913:References
889:Steropodon
877:Berriasian
693:Metatheria
544:Holotheria
512:Allotheria
415:monotremes
388:monotremes
360:placentals
356:marsupials
271:Bishopidae
3985:Atlasodon
3906:includes
3877:includes
3774:Kermackia
3754:Hypomylos
3460:Stylodens
3336:Leonardus
3299:Bondesius
3226:Laolestes
3147:Achyrodon
3127:Tathiodon
2928:Lactodens
2668:Priacodon
2654:Jugulator
2506:Hangjinia
2420:Nakunodon
2370:Kemchugia
2143:includes
2105:Zaglossus
2040:Obdurodon
1960:Stirtodon
1657:Synapsida
1642:Synapsida
1599:Synapsida
1582:Kingdom:
1451:1543-592X
1361:0311-5518
1303:0028-0836
1253:253323862
1245:0311-5518
1204:0311-5518
1126:0272-4634
907:paraphyly
427:cladogram
421:Phylogeny
344:Hypomylos
312:Hypomylos
163:Kingdom:
157:Eukaryota
131:with her
4144:Wikidata
3873:Eutheria
3862:Eutheria
3804:Picopsis
3764:Juramaia
3554:Magnimus
3523:Arguimus
3493:Abelodon
3313:Cronopio
3168:Beckumia
3120:Paurodon
2835:Anebodon
2787:Yermakia
2596:Klamelia
2248:Amphidon
1906:Sundrius
1766:Ambondro
1737:Mammalia
1696:Mammalia
1590:Chordata
1588:Phylum:
1584:Animalia
1566:Mammalia
1521:Taxonomy
1509:Archived
1489:Elementy
1412:32774343
1386:: 0651.
1345:: 1โ19.
1311:38570681
1161:16 March
1083:15705848
1040:11343108
898:Ambondro
860:cingulid
834:and had
712:Eutheria
625:Zatheria
468:Mammalia
407:premolar
403:cingulid
364:Zatheria
246:Ambondro
200:Zatheria
187:Mammalia
177:Chordata
173:Phylum:
167:Animalia
153:Domain:
129:Kangaroo
4150:Q941624
4036:UA 8699
3657:Peramus
3574:Mozomus
3564:Minimus
3453:Donodon
2773:Tinodon
2288:Comodon
1995:Opalios
1845:Bishops
1403:7381353
1283:Bibcode
1091:3048437
1063:Science
1048:4342585
828:mammals
681:
627:
599:
470:
443:
431:McKenna
285:., 2001
183:Class:
114:fossil
3835:Theria
3734:Eomaia
2900:Aliaga
1449:
1410:
1400:
1359:
1309:
1301:
1275:Nature
1251:
1243:
1202:
1152:
1124:
1089:
1081:
1046:
1038:
1020:Nature
990:
854:, and
820:boreas
818:(from
679:Theria
677:
623:
595:
352:Theria
328:Theria
302:Eomaia
224:Clades
111:Eomaia
4164:97116
1622:Clade
1613:Clade
1604:Clade
1595:Clade
1249:S2CID
1087:S2CID
1044:S2CID
942:(2).
824:sphen
283:et al
207:Clade
194:Clade
3900:see
3871:see
2137:see
1682:see
1655:see
1447:ISSN
1408:PMID
1357:ISSN
1307:PMID
1299:ISSN
1241:ISSN
1200:ISSN
1163:2015
1150:ISBN
1122:ISSN
1079:PMID
1036:PMID
988:ISBN
964:link
358:and
350:and
281:Luo
133:joey
35:Pre๊
1439:doi
1398:PMC
1388:doi
1347:doi
1291:doi
1279:628
1231:doi
1190:doi
1114:doi
1071:doi
1067:307
1028:doi
1024:409
944:doi
411:ear
4183::
4161::
4146::
1624::
1615::
1606::
1597::
1519:.
1507:.
1503:.
1487:.
1471:.
1445:.
1435:42
1433:.
1429:.
1406:.
1396:.
1384:11
1382:.
1378:.
1355:.
1339:.
1331:;
1305:.
1297:.
1289:.
1277:.
1273:.
1261:^
1247:.
1239:.
1227:46
1225:.
1221:.
1198:.
1186:46
1184:.
1180:.
1120:.
1108:.
1085:.
1077:.
1065:.
1042:.
1034:.
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1002:^
972:^
960:}}
956:{{
940:63
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934:.
909:.
872:.
850:,
514:โโ
370:.
346:,
279:?
209::
196::
85:Pg
4021:โ
4011:โ
4001:โ
3991:โ
3981:โ
3847:โ
3820:โ
3810:โ
3800:โ
3790:โ
3780:โ
3770:โ
3760:โ
3750:โ
3740:โ
3730:โ
3720:โ
3710:โ
3700:โ
3690:โ
3667:?
3638:?
3623:โ
3614:?
3600:โ
3590:โ
3580:โ
3570:โ
3560:โ
3550:โ
3540:โ
3536:?
3529:โ
3519:โ
3509:โ
3499:โ
3489:โ
3427:โ
3331:?
3323:?
3269:โ
3221:?
3192:?
3087:?
3068:โ
2881:โ
2823:โ
2814:?
2768:?
2753:โ
2737:?
2708:โ
2699:?
2607:โ
2581:?
2576:โ
2546:โ
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2480:โ
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2463:?
2456:โ
2446:โ
2436:โ
2426:โ
2416:โ
2406:โ
2396:โ
2386:โ
2376:โ
2366:โ
2356:โ
2346:โ
2336:โ
2326:โ
2322:?
2315:โ
2311:?
2304:โ
2294:โ
2284:โ
2274:โ
2264:โ
2254:โ
2244:โ
2234:โ
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2124:โ
2087:โ
2077:โ
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2053:โ
2036:โ
2032:?
2025:โ
2021:?
2014:โ
1991:โ
1963:?
1948:โ
1932:?
1917:โ
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1762:โ
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1255:.
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1073::
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952:.
946::
714:โ
695:โ
664:โ
638:โ
610:โ
582:โ
556:โ
530:โ
510:โ
484:โ
454:โ
318:โ
308:โ
298:โ
290:โ
273:?
268:โ
264:?
259:โ
255:?
249:?
242:โ
238:?
233:โ
90:N
80:K
75:J
70:T
65:P
60:C
55:D
50:S
45:O
40:๊
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