406:
33:
83:
60:
367:(leaf axis) is rounded or slightly flattened on the upper side, sometimes even shallowly grooved, and dark in color, as are the axes of the leaf segments. The color continues into the base of the leaf segments, without a distinct joint. The axes are straight, rather than zig-zagging. Each blade has 4 to 6 pairs of pinnae, ovate or deltoid-ovate in shape. These are in turn divided into 2 to 3 pairs of
379:(abruptly terminating) at the base and borne on a short stalk. The ones closest to the blade base may be subdivided into another set of segments, while the pinnules closer to the blade tip are lobed or entire. The leaf tissue is dark blue-green and of a leathery to fleshy texture, obscuring the veins from the upper surface, and does not bear hairs or scales on either surface. Unlike many species in the genus,
343:(the stalk of the leaf, below the blade) is chestnut brown or reddish-brown to dark brown, rounded and hairless, and 0.75 to 1.5 millimeters (0.030 to 0.059 in) in diameter. It is typically 2 to 6 centimeters (0.79 to 2.4 in) long, making up about one-half to two-thirds of the total length of the leaf. The leaf blade is
355:(cut into pinnae and pinnules) to tripinnate (with pinnules cut into pinnulets) at the base, where it is most divided. It is not reduced in width at the base. The blade measures from 3 to 7 centimeters (1 to 3 in) long and 0.8 to 3 centimeters (0.3 to 1 in) wide. The base is
607:. All four species lack farina, and their common ancestor is hypothesized to have diverged from the ancestor of the rest of the genus before farina production developed in the latter.
1317:; Huiet, Layne; Yatskievych, George; Pryer, Kathleen M. (2011). "Species Relationships and Farina Evolution in the Cheilanthoid Fern Genus Argyrochosma (Pteridaceae)".
427:
was thought to exist, but this is believed to be the result of plants occasionally producing 32 diploid (rather than 64 haploid) spores and giving rise to diploid
1314:
539:
201:
1942:
1927:
1932:
1804:
1620:
1897:
1947:
1659:
295:, it does not secrete white powder on the underside of its leaves. First described as a species in 1917, it was transferred to the new genus
1471:
485:
might represent a distinct genus of its own. Weatherby thought that, until that genus was described, the group might better be placed in
336:(toothless) margins and a thin texture. They are 5 to 7 millimeters (0.2 to 0.3 in) long and 0.3 millimeters (0.01 in) wide.
289:
rocks, and has small, finely-divided leaves with a leathery texture and dark axes connecting the leaf segments. Unlike many members of
1698:
398:. The leaf segments, at most, are slightly curled under, not concealing the sporangia, and their tissue is not modified into a false
1778:
1581:
1817:
1633:
1952:
1225:
1073:
1638:
390:
are close to the margin, borne along the further half of the secondary veins branching from the midrib of the segment. Each
1822:
1917:
646:. It grows on calcareous or igneous cliffs and ledges, at an altitude from 600 to 1,900 meters (2,000 to 6,200 ft).
1301:
1957:
1937:
1092:
348:
1664:
368:
376:
329:
1791:
1607:
655:
531:. John T. Mickel, carrying out a similar program of lumping in 1979, accidentally duplicated Munz's combination.
372:
67:
412:
leaflet, showing slightly curled leaf margins, exposed sporangia, and dark color entering base of leaf segments.
356:
333:
325:
1523:
1135:
360:
344:
1848:
1716:
1907:
1573:
1534:
82:
1843:
1711:
1401:
1053:
561:
278:
234:
1220:. Memoirs of the New York Botanical Garden. Vol. 88. Bronx, New York: New York Botanical Garden.
1922:
1912:
602:
476:
1651:
1457:
1703:
1463:
587:
436:
1809:
1625:
1568:
1485:
339:
The leaves range from 4 to 15 centimeters (1.6 to 5.9 in) long, and arise in clumps. The
1902:
1358:
180:
1677:
1103:
1555:
1412:. Vol. 2: Pteridophytes and Gymnosperms. New York and Oxford: Oxford University Press.
1235:
596:
506:
494:
253:
1066:
Plant
Gateway's the Global Flora: A practical flora to vascular plant species of the world
380:
332:, uniformly colored, brown or reddish-brown scales, sometimes nearly black in color, with
301:(the "false cloak ferns") in 1987, recognizing their distinctness from the "cloak ferns" (
8:
1835:
1690:
1145:
1057:
461:
453:
197:
49:
662:
is ranked as apparently secure (G4), but it is considered critically imperiled in Utah.
383:(powder) is not present on either surface of the leaf. The leaves do not curl when dry.
1869:
1737:
1438:
1388:
1380:
1354:
1342:
1334:
1259:
1173:
1119:
444:, but the latter has a darker leaf stalk and axes and the leaf is less highly divided.
209:
77:
1275:
493:, but died in 1949 before he could circumscribe and publish it. Accordingly, in 1950,
1672:
1542:
1392:
1221:
1131:
1124:
1088:
1069:
1362:
1346:
1239:
1186:
1149:
670:
It prefers high light, and moist-dry to dry, well-drained soil, perhaps of high pH.
1874:
1742:
1430:
1370:
1326:
1287:
1251:
1202:
1165:
481:
1547:
1769:
465:
1418:
1830:
1685:
1508:
1271:
1126:
A Field Manual of the Ferns & Fern-Allies of the United States & Canada
643:
582:
514:
472:
340:
223:
107:
1292:
574:, as part of a program to consolidate the cheilanthoid ferns into that genus.
542:, who was carrying out phylogenetic studies of the cheilanthoids. He elevated
1891:
1330:
1061:
639:
308:
1861:
1729:
1207:
994:
982:
552:
297:
291:
157:
137:
850:
1646:
1594:
1517:
523:
428:
405:
147:
32:
1384:
1338:
517:, preparing a flora of California and following Copeland's opinion that
1783:
1586:
1442:
1263:
1177:
619:
566:
391:
387:
303:
286:
1599:
1375:
352:
1479:
1434:
1255:
1169:
1856:
1763:
1724:
1502:
424:
399:
1796:
1612:
970:
623:
468:, who collected the material on which the description was based.
321:
1560:
635:
627:
441:
371:(circular) or deltoid to ovate pinnules, obtuse at the tip and
364:
282:
840:
838:
836:
834:
832:
784:
782:
780:
778:
776:
774:
324:
is short and horizontal, with leaves closely spaced. It bears
1030:
592:
395:
94:
1312:
1000:
988:
862:
856:
728:
726:
724:
722:
1367:
Contributions from the Gray
Herbarium of Harvard University
829:
771:
761:
720:
718:
716:
714:
712:
710:
708:
706:
704:
702:
631:
274:
127:
117:
759:
757:
755:
753:
751:
749:
747:
745:
743:
741:
910:
886:
689:
687:
1018:
699:
479:
suggested in the 1940s that a group of ferns related to
434:
It is most similar in appearance to the closely related
1408:. In Flora of North America Editorial Committee (ed.).
738:
819:
817:
684:
958:
946:
934:
898:
815:
813:
811:
809:
807:
805:
803:
801:
799:
797:
1052:
1006:
976:
874:
591:, a rare Sonoran species; these two species form a
1363:"A revision of the American species of Notholaena"
1123:
922:
794:
538:group as a genus was finally addressed in 1987 by
1419:"Argyrochosma, a new genus of cheilanthoid ferns"
1240:"Notes on the ferns of the eastern United States"
1130:. Washington, DC: Smithsonian Institution Press.
1083:Hoshizaki, Barbara Joe; Moran, Robbin C. (2001).
419:is a sexual diploid, with a chromosome count of 2
1889:
556:, and transferred this species to that genus as
1353:
1082:
1036:
844:
788:
1300:
1215:
868:
732:
693:
610:
1943:Flora of the Sierra Nevada (United States)
58:
31:
1464:Jepson Manual eFlora (TJM2) treatment of
1374:
1291:
1206:
1118:
1024:
765:
16:Species of fern in the family Pteridaceae
1216:Mickel, John T.; Smith, Alan R. (2004).
404:
1928:Flora of the Southwestern United States
1416:
1399:
1101:
1012:
964:
952:
904:
823:
673:
521:was best lumped into a broadly defined
1933:Flora of the California desert regions
1890:
1410:Flora of North America North of Mexico
1234:
1184:
940:
916:
892:
511:Flowering Plants and ferns of Arizona.
1898:NatureServe apparently secure species
1484:
1483:
1144:
880:
577:Phylogenetic studies have shown that
1948:Natural history of the Mojave Desert
1270:
928:
618:is known in the United States from
595:sister to another clade containing
13:
1087:. Portland, Oregon: Timber Press.
527:, transferred it to that genus as
14:
1969:
1451:
1191:in the continental United States"
977:Christenhusz, Fay & Byng 2018
1849:urn:lsid:ipni.org:names:170429-2
1717:urn:lsid:ipni.org:names:272606-2
81:
1308:. NatureServe. October 3, 2020.
656:NatureServe conservation status
649:
1112:Biota of North America Program
1045:
665:
505:, to provide a name for it in
386:In fertile leaf segments, the
315:
1:
678:
1953:Flora of the Sonoran Deserts
1417:Windham, Michael D. (1987).
1400:Windham, Michael D. (1993).
7:
1218:The Pteridophytes of Mexico
1054:Christenhusz, Maarten J. M.
564:transferred the species to
497:transferred the species to
447:
10:
1974:
1918:Ferns of the United States
1276:"California Miscellany IV"
1037:Hoshizaki & Moran 2001
845:Tryon & Weatherby 1956
789:Tryon & Weatherby 1956
638:, particularly within the
562:Maarten J. M. Christenhusz
440:, and both are present in
359:(blunt), while the tip is
279:southwestern United States
1753:
1492:
1293:10.5642/aliso.19580401.04
1102:Kartesz, John T. (2014).
215:
208:
186:
179:
78:Scientific classification
76:
56:
47:
39:
30:
23:
1958:Plants described in 1917
1938:Flora of the Great Basin
1331:10.1600/036364411X583547
1185:Mickel, John T. (1979).
611:Distribution and habitat
477:Charles Alfred Weatherby
1460:at JSTOR Plant Science.
869:Mickel & Smith 2004
733:Mickel & Smith 2004
534:The recognition of the
271:Jones' false cloak fern
1458:Isotype of the species
1208:10.5962/bhl.part.20791
452:The species was first
413:
285:, Mexico. It grows on
1474:Argyrochosma jonesii
1466:Argyrochosma jonesii
1423:American Fern Journal
1244:American Fern Journal
1158:American Fern Journal
408:
1524:Argyrochosma jonesii
1494:Argyrochosma jonesii
1472:UC Photos gallery —
1404:Argyrochosma jonesii
1304:Argyrochosma jonesii
1085:Fern Grower's Manual
674:Notes and references
616:Argyrochosma jonesii
550:to become the genus
507:Thomas Henry Kearney
495:Conrad Vernon Morton
410:Argyrochosma jonesii
266:Argyrochosma jonesii
190:Argyrochosma jonesii
41:Argyrochosma jonesii
25:Argyrochosma jonesii
1908:Ferns of California
1315:Windham, Michael D.
1154:from the Southwest"
1120:Lellinger, David B.
1064:(8 February 2018).
919:, pp. 249–251.
895:, pp. 249–250.
634:and in Mexico from
604:A. microphylla
464:. It was named for
462:William Ralph Maxon
423:= 54. A tetraploid
219:Cheilanthes jonesii
50:Conservation status
43:growing on a cliff
1755:Notholaena jonesii
871:, pp. 67, 69.
588:A. lumholtzii
540:Michael D. Windham
458:Notholaena jonesii
437:A. lumholtzii
414:
375:(heart-shaped) or
351:, and ranges from
273:, is a species of
241:Notholaena jonesii
230:Hemionitis jonesii
64:Apparently Secure
1885:
1884:
1673:Open Tree of Life
1486:Taxon identifiers
1359:Weatherby, Una F.
1319:Systematic Botany
1227:978-0-89327-488-7
1146:Maxon, William R.
1075:978-0-9929993-9-1
1001:Sigel et al. 2011
989:Sigel et al. 2011
857:Sigel et al. 2011
489:, rather than in
262:
261:
256:
245:
237:
226:
71:
1965:
1878:
1877:
1865:
1864:
1852:
1851:
1839:
1838:
1826:
1825:
1813:
1812:
1800:
1799:
1787:
1786:
1774:
1773:
1772:
1746:
1745:
1733:
1732:
1720:
1719:
1707:
1706:
1694:
1693:
1681:
1680:
1668:
1667:
1655:
1654:
1642:
1641:
1629:
1628:
1616:
1615:
1603:
1602:
1590:
1589:
1577:
1576:
1564:
1563:
1551:
1550:
1538:
1537:
1528:
1527:
1526:
1513:
1512:
1511:
1481:
1480:
1446:
1413:
1396:
1378:
1376:10.5962/p.336378
1350:
1313:Sigel, Erin M.;
1309:
1297:
1295:
1267:
1231:
1212:
1210:
1187:"The fern genus
1181:
1141:
1129:
1115:
1098:
1079:
1040:
1034:
1028:
1022:
1016:
1010:
1004:
998:
992:
986:
980:
974:
968:
962:
956:
950:
944:
938:
932:
926:
920:
914:
908:
902:
896:
890:
884:
878:
872:
866:
860:
854:
848:
842:
827:
821:
792:
786:
769:
763:
736:
730:
697:
694:NatureServe 2020
691:
482:Notholaena nivea
251:
243:
232:
221:
192:
86:
85:
65:
62:
61:
35:
21:
20:
1973:
1972:
1968:
1967:
1966:
1964:
1963:
1962:
1923:Flora of Sonora
1913:Ferns of Mexico
1888:
1887:
1886:
1881:
1873:
1868:
1860:
1855:
1847:
1842:
1834:
1829:
1821:
1816:
1808:
1803:
1795:
1790:
1782:
1777:
1768:
1767:
1762:
1749:
1741:
1736:
1728:
1723:
1715:
1710:
1702:
1697:
1689:
1684:
1676:
1671:
1663:
1658:
1650:
1645:
1637:
1632:
1624:
1619:
1611:
1606:
1598:
1593:
1585:
1580:
1572:
1567:
1559:
1554:
1546:
1541:
1533:
1531:
1522:
1521:
1516:
1507:
1506:
1501:
1488:
1454:
1449:
1435:10.2307/1547438
1355:Tryon, Rolla M.
1272:Munz, Philip A.
1256:10.2307/1545864
1228:
1170:10.2307/1543948
1138:
1095:
1076:
1068:. Vol. 4.
1058:Fay, Michael F.
1048:
1043:
1035:
1031:
1023:
1019:
1011:
1007:
999:
995:
987:
983:
975:
971:
963:
959:
951:
947:
939:
935:
927:
923:
915:
911:
903:
899:
891:
887:
879:
875:
867:
863:
855:
851:
843:
830:
822:
795:
787:
772:
764:
739:
731:
700:
692:
685:
681:
676:
668:
660:A. jonesii
652:
644:Sonoran Deserts
630:, and southern
613:
598:A. formosa
579:A. jonesii
572:H. jonesii
558:A. jonesii
529:C. jonesii
503:P. jonesii
466:Marcus E. Jones
450:
417:A. jonesii
363:(pointed). The
318:
249:Pellaea jonesii
204:
194:
188:
175:
172:A. jonesii
80:
72:
63:
59:
52:
17:
12:
11:
5:
1971:
1961:
1960:
1955:
1950:
1945:
1940:
1935:
1930:
1925:
1920:
1915:
1910:
1905:
1900:
1883:
1882:
1880:
1879:
1875:wfo-0001109192
1866:
1853:
1840:
1827:
1814:
1801:
1788:
1775:
1759:
1757:
1751:
1750:
1748:
1747:
1743:wfo-0001114273
1734:
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1452:External links
1450:
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1397:
1369:(179): 1–106.
1351:
1325:(3): 554–564.
1310:
1298:
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1250:(4): 241–252.
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1201:(6): 431–437.
1182:
1164:(4): 106–109.
1142:
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1080:
1074:
1062:Byng, James W.
1049:
1047:
1044:
1042:
1041:
1039:, p. 189.
1029:
1027:, p. 154.
1025:Lellinger 1985
1017:
1005:
1003:, p. 559.
993:
991:, p. 558.
981:
969:
957:
945:
943:, p. 434.
933:
921:
909:
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885:
883:, p. 108.
873:
861:
859:, p. 560.
849:
828:
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770:
768:, p. 153.
766:Lellinger 1985
737:
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667:
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612:
609:
583:sister species
515:Philip A. Munz
473:Edwin Copeland
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277:native to the
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118:Polypodiophyta
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1286:(1): 87–100.
1285:
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1269:
1265:
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1257:
1253:
1249:
1245:
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1237:
1236:Morton, C. V.
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1143:
1139:
1133:
1128:
1127:
1121:
1117:
1113:
1109:
1107:
1100:
1096:
1094:9780881924954
1090:
1086:
1081:
1077:
1071:
1067:
1063:
1059:
1055:
1051:
1050:
1038:
1033:
1026:
1021:
1014:
1009:
1002:
997:
990:
985:
979:, p. 16.
978:
973:
967:, p. 40.
966:
961:
955:, p. 38.
954:
949:
942:
937:
931:, p. 87.
930:
925:
918:
913:
907:, p. 37.
906:
901:
894:
889:
882:
877:
870:
865:
858:
853:
847:, p. 84.
846:
841:
839:
837:
835:
833:
825:
820:
818:
816:
814:
812:
810:
808:
806:
804:
802:
800:
798:
791:, p. 83.
790:
785:
783:
781:
779:
777:
775:
767:
762:
760:
758:
756:
754:
752:
750:
748:
746:
744:
742:
735:, p. 67.
734:
729:
727:
725:
723:
721:
719:
717:
715:
713:
711:
709:
707:
705:
703:
695:
690:
688:
683:
671:
663:
661:
657:
647:
645:
641:
637:
633:
629:
625:
621:
617:
608:
606:
605:
600:
599:
594:
590:
589:
584:
580:
575:
573:
569:
568:
563:
559:
555:
554:
549:
545:
541:
537:
536:N. nivea
532:
530:
526:
525:
520:
516:
512:
508:
504:
500:
496:
492:
488:
484:
483:
478:
474:
469:
467:
463:
459:
455:
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432:
430:
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397:
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389:
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366:
362:
358:
354:
350:
346:
342:
337:
335:
331:
327:
323:
313:
311:
310:
309:sensu stricto
306:
305:
300:
299:
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293:
288:
284:
280:
276:
272:
268:
267:
255:
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231:
228:
225:
220:
217:
216:
214:
211:
207:
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199:
193:
191:
185:
182:
181:Binomial name
178:
174:
173:
168:
165:
164:
161:
160:
156:
153:
152:
149:
146:
143:
142:
139:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
109:
108:Tracheophytes
106:
103:
100:
99:
96:
93:
90:
89:
84:
79:
75:
69:
55:
51:
46:
42:
38:
34:
29:
26:
22:
19:
1903:Argyrochosma
1836:tro-26602346
1754:
1691:tro-26608667
1493:
1473:
1465:
1429:(2): 37–41.
1426:
1422:
1409:
1403:
1366:
1322:
1318:
1303:
1283:
1279:
1247:
1243:
1217:
1198:
1194:
1188:
1161:
1157:
1151:
1125:
1111:
1106:Argyrochosma
1105:
1084:
1065:
1032:
1020:
1013:Kartesz 2014
1008:
996:
984:
972:
965:Windham 1987
960:
953:Windham 1987
948:
936:
924:
912:
905:Windham 1987
900:
888:
876:
864:
852:
824:Windham 1993
669:
659:
653:
650:Conservation
615:
614:
603:
597:
586:
578:
576:
571:
565:
557:
553:Argyrochosma
551:
548:Argyrochosma
547:
543:
535:
533:
528:
522:
518:
510:
502:
498:
490:
486:
480:
470:
457:
451:
435:
433:
429:gametophytes
420:
416:
415:
409:
394:contains 64
385:
338:
319:
307:
302:
298:Argyrochosma
296:
292:Argyrochosma
290:
270:
265:
264:
263:
248:
240:
229:
218:
189:
187:
171:
170:
159:Argyrochosma
158:
138:Polypodiales
101:
40:
24:
18:
1647:NatureServe
1595:iNaturalist
1518:Wikispecies
1189:Cheilanthes
1046:Works cited
941:Mickel 1979
917:Morton 1950
893:Morton 1950
666:Cultivation
560:. In 2018,
524:Cheilanthes
456:in 1917 as
316:Description
269:, known as
148:Pteridaceae
68:NatureServe
1892:Categories
1831:Plant List
1686:Plant List
1532:Calflora:
1195:Phytologia
1152:Notholaena
1137:0874746035
881:Maxon 1917
679:References
654:Under the
620:California
567:Hemionitis
544:Notholaena
519:Notholaena
491:Notholaena
392:sporangium
349:lanceolate
328:or linear-
304:Notholaena
287:calcareous
254:C.V.Morton
235:Christenh.
114:Division:
1770:Q38241728
1574:233500151
1393:249085059
929:Munz 1958
513:In 1958,
454:described
388:sporangia
369:orbicular
353:bipinnate
166:Species:
91:Kingdom:
1862:26602346
1857:Tropicos
1810:10257141
1797:170429-2
1764:Wikidata
1730:26608667
1725:Tropicos
1652:2.131280
1626:10804536
1613:272606-2
1509:Q4789955
1503:Wikidata
1385:41764632
1361:(1956).
1347:16214744
1339:23028975
1274:(1958).
1238:(1950).
1148:(1917).
1122:(1985).
658:system,
448:Taxonomy
425:cytotype
400:indusium
377:truncate
330:subulate
252:(Maxon)
244:Maxon
233:(Maxon)
222:(Maxon)
210:Synonyms
144:Family:
1784:2651666
1587:2651663
1443:1547438
1264:1545864
1178:1543948
1150:"A New
624:Arizona
499:Pellaea
487:Pellaea
373:cordate
322:rhizome
202:Windham
154:Genus:
134:Order:
124:Class:
95:Plantae
66: (
1699:PLANTS
1678:513465
1665:414590
1639:500967
1561:489985
1441:
1391:
1383:
1345:
1337:
1262:
1224:
1176:
1134:
1091:
1072:
640:Mojave
636:Sonora
628:Nevada
546:sect.
442:Sonora
396:spores
381:farina
365:rachis
357:obtuse
334:entire
326:linear
283:Sonora
1823:17624
1805:IRMNG
1704:ARJO2
1621:IRMNG
1600:75555
1439:JSTOR
1389:S2CID
1381:JSTOR
1343:S2CID
1335:JSTOR
1280:Aliso
1260:JSTOR
1174:JSTOR
593:clade
581:is a
471:Both
361:acute
345:ovate
341:stipe
198:Maxon
102:Clade
1844:POWO
1818:ITIS
1792:IPNI
1779:GBIF
1712:POWO
1660:NCBI
1634:ITIS
1608:IPNI
1582:GBIF
1548:GJMV
1222:ISBN
1132:ISBN
1089:ISBN
1070:ISBN
642:and
632:Utah
601:and
475:and
320:The
281:and
275:fern
224:Munz
1870:WFO
1738:WFO
1569:FNA
1556:EoL
1543:CoL
1535:659
1431:doi
1371:doi
1327:doi
1288:doi
1252:doi
1203:doi
1166:doi
585:to
570:as
509:'s
501:as
460:by
347:to
312:).
1894::
1872::
1859::
1846::
1833::
1820::
1807::
1794::
1781::
1766::
1740::
1727::
1714::
1701::
1688::
1675::
1662::
1649::
1636::
1623::
1610::
1597::
1584::
1571::
1558::
1545::
1520::
1505::
1437:.
1427:77
1425:.
1421:.
1387:.
1379:.
1365:.
1357:;
1341:.
1333:.
1323:36
1321:.
1282:.
1278:.
1258:.
1248:40
1246:.
1242:.
1199:41
1197:.
1193:.
1172:.
1160:.
1156:.
1110:.
1060:;
1056:;
831:^
796:^
773:^
740:^
701:^
686:^
626:,
622:,
431:.
402:.
200:)
104::
1445:.
1433::
1406:"
1402:"
1395:.
1373::
1349:.
1329::
1306:"
1302:"
1296:.
1290::
1284:4
1266:.
1254::
1230:.
1211:.
1205::
1180:.
1168::
1162:7
1140:.
1114:.
1108:"
1104:"
1097:.
1078:.
1015:.
826:.
696:.
421:n
196:(
70:)
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