454:
352:
390:(the stalk of the leaf, below the blade) is a shiny pale to medium brown, chestnut brown, or reddish-brown in color, measuring 2 to 7 centimeters (0.8 to 3 in) long and 0.50 to 0.75 millimeters (0.020 to 0.030 in) in diameter. It is rounded (rather than flattened or grooved) and lacks extensive hairs or scales; a few rhizome scales may be present at its base. The shiny brown color of the stipe continues into the rachis and the axis of the leaf segments, and blends with the leaf tissue at the base of each segment, without an abrupt end point. The
33:
83:
60:
769:
Kunze suggested the species was suitable for cultivation, noting that it was grown in Berlin in a well-lit part of the greenhouses. George
Schneider suggested it required "cool treatment", perhaps in contrast to the desert-dwelling members of the genus. It has been described as preferring medium
429:, and the dark color of the axes passes into their base, without a distinct joint. The blade tissue is bluish green and somewhat leafy in texture (rather than densely leathery); veins are often visible on the upper surface of the leaf. and a whitish (or rarely, yellowish)
798:
were formerly distinguished by the morphology of their false indusia: broken into multiple segments in the first genus and largely continuous in the second. This has since proven to be an artificial distinction, of little value in identifying the relationships between
326:. Its leaves are highly divided, with leaf segments joined by shiny, chestnut-brown axes, and their undersides are coated with white powder, giving the fern its name. First described as a species in 1814, it was transferred to the new genus
366:
is compact, more or less upright, and usually unbranching, and bears brown, pale brown or yellowish-brown scales that are uniform in color. These are about 3 millimeters (0.1 in) long, thin and delicate, narrowly
732:
is endemic to the United States. It is found from the western edge of
Illinois south and west through Arkansas and Missouri to southeastern Nebraska and thence south through Kansas and Oklahoma to Texas, with a
760:
is ranked as apparently secure (G4), but it is considered imperiled in
Arkansas and Oklahoma, and the single occurrences in Nebraska and Kentucky are considered extirpated and historical, respectively.
421:(pointed) at the base and acute at the tip. 3 to 5 pairs of pinnae are present. They are roughly deltate in shape and attached to the rachis by a long stalk. The final divisions of the blade are
433:(powder) covers the underside, while the upper surface is free of hairs, powder, etc., although it may bear a few glands. The edges of leaf segments curl back under and partially protect the
507:, he noted that the Missouri material was smaller and more highly divided, and commented on the white powder on the underside of the leaves, lacking in Pursh's description. The epithet
413:(cut into pinnae, pinnules, and pinnulets) to pentapinnate (five times divided). They are 2 to 6 centimeters (0.8 to 2 in) long and 1.5 to 6 centimeters (0.6 to 2 in) wide,
1474:; Huiet, Layne; Yatskievych, George; Pryer, Kathleen M. (2011). "Species Relationships and Farina Evolution in the Cheilanthoid Fern Genus Argyrochosma (Pteridaceae)".
1237:
601:
279:
1471:
644:
201:
532:
Delineating natural genera in the cheilanthoids has proven to be extremely difficult, and other placements of the species were subsequently put forward.
683:
was preoccupied), as part of a program to consolidate the cheilanthoid ferns into that genus. The epithet refers to the horse of Atreyu in the novel
1920:
1736:
1987:
1775:
538:
2007:
1814:
1697:
643:
suggested in the 1940s that this group of ferns might represent a distinct genus of its own. This was finally addressed in 1987 by
1933:
1749:
1460:
1227:
1938:
1754:
2002:
1390:
1251:
533:
231:
1997:
1280:
1780:
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were borne along the nerves and not densely clustered at the end of the nerves. He transferred the species there as
406:
368:
1907:
1723:
1649:
753:
402:
67:
422:
414:
376:
372:
1360:
426:
418:
1832:
1899:
1689:
1540:
441:
contain 64 spores, released in summer and fall. Plants are sexual diploids, with a chromosome number of 2
82:
546:
in 1852, breaking with most other authorities in recognizing and re-circumscribing that genus. In 1859,
1827:
1207:
666:
253:
1596:
485:, meaning "covered in white powder", refers to the coating of farina on the underside of the leaves.
717:
are commonly referred to as "false cloak ferns"; the name apparently refers to their resemblance to
1422:
1370:
640:
547:
470:
242:
197:
1767:
1819:
1402:
571:
494:
298:
1925:
1894:
1741:
1684:
1611:
1448:
702:
1992:
1515:
1241:
734:
685:
180:
1793:
1291:
1671:
1551:. Vol. 2: Pteridophytes and Gymnosperms. New York and Oxford: Oxford University Press.
386:
spring from the rhizome in clusters and are 3 to 15 centimeters (1 to 6 in) long. The
1220:
Plant
Gateway's the Global Flora: A practical flora to vascular plant species of the world
430:
332:(the "false cloak ferns") in 1987, recognizing their distinctness from the "cloak ferns" (
8:
1806:
1211:
721:
proper, known as "cloak ferns" for the reflexed leaf margins, which cloak the sporangia.
466:
457:
Underside of a leaf, showing the coating of white farina that gives the species its name.
49:
1959:
1853:
1577:
1527:
1511:
1499:
1491:
1344:
567:
209:
77:
453:
351:
1788:
1658:
1456:
1356:
1349:
1276:
1223:
1519:
1503:
1437:
465:
ferns, this species has been placed in many genera at different times. It was first
1964:
1858:
1569:
1483:
503:
1663:
770:
light, and relatively dry, well-drained soil mixed with sand, perhaps of high pH.
706:, of the southwestern United States and northern Mexico; these two species form a
1885:
1634:
576:
318:, is a small fern endemic to the central and southern United States. It grows on
1557:
1374:
1328:
1311:
1255:
1148:
398:(pinna axes) are more or less straight. The axes tend to curl upwards when dry.
1801:
1411:
Botanische JahrbĂĽcher fĂĽr
Systematik, Pflanzengeschichte und Pflanzengeographie
1351:
A Field Manual of the Ferns & Fern-Allies of the United States & Canada
697:
636:
478:
395:
387:
107:
1426:
1406:
511:
means "small and beautiful". He subsequently recognized it as synonymous with
32:
1981:
1487:
1215:
339:
580:
to include several genera in which he perceived close affinities, including
1951:
1845:
1307:
657:
486:
462:
328:
264:
157:
137:
1762:
1710:
1643:
647:, who was carrying out phylogenetic studies of these genera. He elevated
147:
1531:
1495:
409:, broadest at the base, and are most highly divided there, ranging from
1702:
1581:
741:
671:
559:
517:
438:
410:
394:(leaf axis) is rounded or slightly flattened on the upper surface. The
334:
319:
1715:
323:
1605:
1573:
1946:
1879:
1840:
1628:
1260:
Mémoires de la Société du muséum d'histoire naturelle de
Strasbourg
498:
1912:
1728:
1136:
363:
1379:
619:
By the late 20th century, most authorities favored placement of
1676:
1439:
The book of choice ferns for the garden, conservatory and stove
910:
842:
840:
838:
836:
834:
832:
830:
391:
908:
906:
904:
902:
900:
898:
896:
894:
892:
890:
707:
434:
358:
leaf blade, showing a few glands dotted on the upper surface.
94:
1469:
1154:
827:
570:
for a different species in 1825, rendering it illegitimate.
525:
in 1848, noting that he "would not have looked for it under
1524:
Contributions from the Gray
Herbarium of Harvard University
968:
887:
710:
sister to another clade of several South
American species.
127:
117:
1243:
Catalogue of the
Davenport Herbarium Supplement—March 1883
1052:
1016:
958:
956:
954:
952:
939:
937:
935:
933:
931:
929:
927:
925:
923:
1312:"Nothochlaena pulchella, ein neues Farrnkraut der Gärten"
980:
817:
815:
1184:
1004:
1547:. In Flora of North America Editorial Committee (ed.).
1088:
1064:
949:
920:
1028:
877:
875:
812:
1124:
1112:
1100:
1076:
873:
871:
869:
867:
865:
863:
861:
859:
857:
855:
437:, which are borne along veins close to the edge. The
1172:
1206:
1160:
1142:
992:
1520:"A revision of the American species of Notholaena"
1455:. Cambridge, England: Cambridge University Press.
1348:
852:
477:, based on material collected on the banks of the
1558:"Argyrochosma, a new genus of cheilanthoid ferns"
1355:. Washington, DC: Smithsonian Institution Press.
1271:Hoshizaki, Barbara Joe; Moran, Robbin C. (2001).
1979:
1431:. Vol. 2. London: White, Cochrane & Co.
740:It grows in the crevices of cliffs or ledges of
661:, and transferred this species to that genus as
1040:
1510:
1270:
914:
846:
554:as a separate genus, but recognized the genus
1407:"Die Farngattungen Cryptogramme und Pellaea"
493:in 1843. His specimen was collected for the
1453:A Primer of Botanical Latin with Vocabulary
1446:
1389:
1022:
986:
821:
724:
1329:"Notes on some ferns of the United States"
58:
31:
1435:
1369:
1343:
1236:
1190:
1094:
1070:
1058:
974:
962:
943:
566:, but this name had already been used by
16:Species of fern in the family Pteridaceae
1375:"Uber einige Farngattungen: Cheilanthes"
452:
350:
1555:
1538:
1289:
1166:
1130:
1118:
1106:
881:
773:
489:independently described the species as
1980:
1549:Flora of North America North of Mexico
1442:. Vol. 2. London: L. Upcott Gill.
1401:
1082:
1988:NatureServe apparently secure species
1610:
1609:
1421:
1326:
1306:
1178:
1034:
1010:
998:
692:Phylogenetic studies have shown that
623:and closely related ferns, including
1333:American Journal of Science and Arts
550:rejected FĂ©e's attempt to delineate
1250:
1046:
13:
2008:Endemic flora of the United States
1275:. Portland, Oregon: Timber Press.
1246:. Boston: George Edward Davenport.
515:and transferred the older name to
14:
2019:
1590:
1143:Christenhusz, Fay & Byng 2018
1833:urn:lsid:ipni.org:names:272601-2
81:
1597:Pictures and description of an
1397:. NatureServe. October 3, 2020.
1256:"Genera filicum. Polypodiacées"
785:
754:NatureServe conservation status
747:
1428:Flora Americae Septentrionalis
1300:Biota of North America Program
1199:
764:
584:. Accordingly, he transferred
346:
1:
806:
501:. Initially misidentified as
1556:Windham, Michael D. (1987).
1539:Windham, Michael D. (1993).
7:
1208:Christenhusz, Maarten J. M.
737:in south-central Kentucky.
669:transferred the species to
448:
43:growing in a rock crevice.
10:
2024:
2003:Ferns of the United States
1436:Schneider, George (1892).
915:Tryon & Weatherby 1956
847:Hoshizaki & Moran 2001
667:Maarten J. M. Christenhusz
604:lumped it as a variety of
379:(toothless) at the edges.
1869:
1618:
1290:Kartesz, John T. (2014).
215:
208:
186:
179:
78:Scientific classification
76:
56:
47:
39:
30:
23:
1998:Plants described in 1814
1488:10.1600/036364411X583547
1238:Davenport, George Edward
778:
725:Distribution and habitat
316:powdery false cloak fern
1023:Short & George 2013
987:Short & George 2013
602:George Edward Davenport
495:Berlin Botanical Garden
1327:Kunze, Gustav (1848).
458:
359:
1650:Argyrochosma dealbata
1620:Argyrochosma dealbata
1562:American Fern Journal
1543:Argyrochosma dealbata
1393:Argyrochosma dealbata
1310:(15 September 1843).
730:Argyrochosma dealbata
686:The Neverending Story
456:
356:Argyrochosma dealbata
354:
311:Argyrochosma dealbata
245:, nom. illeg. hom.
190:Argyrochosma dealbata
41:Argyrochosma dealbata
25:Argyrochosma dealbata
1871:Cheilanthes dealbata
1273:Fern Grower's Manual
1157:, pp. 558, 561.
774:Notes and references
655:to become the genus
491:Notholaena pulchella
475:Cheilanthes dealbata
401:The leaf blades are
286:Notholaena pulchella
238:Gymnogramma dealbata
219:Cheilanthes dealbata
1472:Windham, Michael D.
1345:Lellinger, David B.
1013:, pp. 633–634.
977:, pp. 160–161.
544:Cincinalis dealbata
260:Notholaena dealbata
227:Cincinalis dealbata
50:Conservation status
1316:Botanische Zeitung
645:Michael D. Windham
558:for species where
536:transferred it to
459:
360:
64:Apparently Secure
1975:
1974:
1789:Open Tree of Life
1612:Taxon identifiers
1516:Weatherby, Una F.
1476:Systematic Botany
1462:978-1-107-69375-3
1229:978-0-9929993-9-1
1155:Sigel et al. 2011
1061:, pp. 49–51.
1037:, pp. 82–83.
703:A. limitanea
307:
306:
301:
290:
282:
271:Notholaena nivea
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1599:A. dealbata
1585:
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1535:
1507:
1470:Sigel, Erin M.;
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1423:Pursh, Frederick
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822:NatureServe 2020
819:
800:
789:
758:A. dealbata
735:disjunct station
694:A. dealbata
663:A. dealbata
625:N. dealbata
621:Notholaena nivea
598:P. dealbata
586:C. dealbata
564:G. dealbata
523:N. dealbata
513:C. dealbata
504:Notholaena nivea
405:(triangular) or
384:A. dealbata
296:
294:Pellaea dealbata
288:
277:
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252:
250:Hemionitis artax
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172:A. dealbata
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1574:10.2307/1547438
1512:Tryon, Rolla M.
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1363:
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1222:. Vol. 4.
1212:Fay, Michael F.
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322:rocks, such as
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1591:External links
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1526:(179): 1–106.
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1482:(3): 554–564.
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1322:(37): 633–635.
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1228:
1216:Byng, James W.
1203:
1201:
1198:
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1195:
1193:, p. 606.
1191:Schneider 1892
1183:
1181:, p. 635.
1171:
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1111:
1099:
1095:Davenport 1883
1087:
1085:, p. 417.
1075:
1071:Mettenius 1859
1063:
1059:Mettenius 1859
1051:
1039:
1027:
1025:, p. 236.
1015:
1003:
1001:, p. 671.
991:
989:, p. 153.
979:
975:Lellinger 1985
967:
965:, p. 161.
963:Lellinger 1985
948:
946:, p. 160.
944:Lellinger 1985
919:
886:
851:
849:, p. 189.
826:
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799:cheilanthoids.
783:
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766:
763:
749:
746:
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698:sister species
637:Edwin Copeland
497:from rocks in
481:. The epithet
479:Missouri River
450:
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382:The fronds of
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1447:Short, Emma;
1445:
1441:
1440:
1434:
1430:
1429:
1424:
1420:
1417:(5): 403–430.
1416:
1412:
1408:
1404:
1400:
1396:
1394:
1388:
1384:
1380:
1376:
1372:
1371:Mettenius, G.
1368:
1364:
1358:
1353:
1352:
1346:
1342:
1338:
1334:
1330:
1325:
1321:
1317:
1313:
1309:
1308:Kunze, Gustav
1305:
1301:
1297:
1295:
1288:
1284:
1282:9780881924954
1278:
1274:
1269:
1265:
1261:
1257:
1253:
1252:FĂ©e, A. L. A.
1249:
1245:
1244:
1239:
1235:
1231:
1225:
1221:
1217:
1213:
1209:
1205:
1204:
1192:
1187:
1180:
1175:
1168:
1163:
1156:
1151:
1145:, p. 10.
1144:
1139:
1133:, p. 40.
1132:
1127:
1121:, p. 38.
1120:
1115:
1109:, p. 37.
1108:
1103:
1097:, p. 44.
1096:
1091:
1084:
1079:
1073:, p. 50.
1072:
1067:
1060:
1055:
1048:
1043:
1036:
1031:
1024:
1019:
1012:
1007:
1000:
995:
988:
983:
976:
971:
964:
959:
957:
955:
953:
945:
940:
938:
936:
934:
932:
930:
928:
926:
924:
917:, p. 89.
916:
911:
909:
907:
905:
903:
901:
899:
897:
895:
893:
891:
883:
878:
876:
874:
872:
870:
868:
866:
864:
862:
860:
858:
856:
848:
843:
841:
839:
837:
835:
833:
831:
823:
818:
816:
811:
797:
793:
788:
784:
771:
762:
759:
755:
745:
743:
738:
736:
731:
722:
720:
716:
711:
709:
705:
704:
699:
695:
690:
688:
687:
682:
679:(the epithet
678:
677:H. artax
674:
673:
668:
664:
660:
659:
654:
650:
646:
642:
638:
634:
630:
626:
622:
617:
615:
611:
610:N. nivea
607:
606:N. nivea
603:
599:
595:
591:
587:
583:
579:
578:
573:
569:
565:
561:
557:
553:
549:
545:
541:
540:
535:
530:
528:
524:
520:
519:
514:
510:
506:
505:
500:
496:
492:
488:
484:
480:
476:
472:
468:
464:
455:
446:
444:
440:
436:
432:
428:
424:
420:
416:
412:
408:
404:
399:
397:
393:
389:
385:
380:
378:
374:
370:
365:
357:
353:
344:
342:
341:
340:sensu stricto
337:
336:
331:
330:
325:
321:
317:
313:
312:
300:
295:
292:
287:
284:
281:
276:
269:
266:
261:
258:
255:
251:
248:
244:
239:
236:
233:
228:
225:
220:
217:
216:
214:
211:
207:
203:
199:
193:
191:
185:
182:
181:Binomial name
178:
174:
173:
168:
165:
164:
161:
160:
156:
153:
152:
149:
146:
143:
142:
139:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
109:
108:Tracheophytes
106:
103:
100:
99:
96:
93:
90:
89:
84:
79:
75:
69:
55:
51:
46:
42:
38:
34:
29:
26:
22:
19:
1993:Argyrochosma
1870:
1807:tro-26608662
1619:
1598:
1568:(2): 37–41.
1565:
1561:
1548:
1542:
1523:
1479:
1475:
1452:
1449:George, Alex
1438:
1427:
1414:
1410:
1392:
1382:
1378:
1350:
1339:(16): 80–88.
1336:
1335:. Series 2.
1332:
1319:
1315:
1299:
1294:Argyrochosma
1293:
1272:
1263:
1259:
1242:
1219:
1186:
1174:
1167:Kartesz 2014
1162:
1150:
1138:
1131:Windham 1987
1126:
1119:Windham 1987
1114:
1107:Windham 1987
1102:
1090:
1078:
1066:
1054:
1042:
1030:
1018:
1006:
994:
982:
970:
882:Windham 1993
795:
791:
787:
768:
757:
751:
748:Conservation
739:
729:
728:
718:
715:Argyrochosma
714:
712:
701:
693:
691:
684:
680:
676:
670:
662:
658:Argyrochosma
656:
653:Argyrochosma
652:
648:
632:
628:
627:, either in
624:
620:
618:
613:
609:
605:
597:
593:
589:
585:
581:
575:
563:
555:
551:
543:
537:
531:
526:
522:
516:
512:
508:
502:
490:
487:Gustav Kunze
482:
474:
463:cheilanthoid
460:
442:
400:
383:
381:
361:
355:
338:
333:
329:Argyrochosma
327:
315:
310:
309:
308:
293:
285:
270:
259:
249:
237:
226:
218:
189:
187:
171:
170:
159:Argyrochosma
158:
138:Polypodiales
101:
40:
24:
18:
1763:NatureServe
1711:iNaturalist
1644:Wikispecies
1385:(1): 47–99.
1200:Works cited
1083:Prantl 1882
792:Cheilanthes
765:Cultivation
713:Species of
665:. In 2018,
616:, in 1883.
556:Gymnogramma
527:Cheilanthes
473:in 1814 as
417:(blunt) to
347:Description
148:Pteridaceae
68:NatureServe
1982:Categories
1802:Plant List
1601:occurrence
1403:Prantl, K.
1362:0874746035
1179:Kunze 1843
1035:Kunze 1848
1011:Kunze 1843
999:Pursh 1814
807:References
796:Notholaena
752:Under the
742:calcareous
719:Notholaena
672:Hemionitis
649:Notholaena
629:Notholaena
612:var.
594:Cincinalis
582:Cincinalis
552:Cincinalis
539:Cincinalis
518:Notholaena
461:Like many
411:tripinnate
407:lanceolate
335:Notholaena
320:calcareous
254:Christenh.
114:Division:
1900:242312228
1886:Q38241697
1690:233500148
1635:Q15597199
600:in 1882.
574:expanded
560:sporangia
548:Mettenius
509:pulchella
467:described
439:sporangia
324:limestone
166:Species:
91:Kingdom:
1952:26608669
1947:Tropicos
1926:10861173
1913:274451-2
1880:Wikidata
1846:26608662
1841:Tropicos
1768:2.158326
1742:10207911
1729:272601-2
1629:Wikidata
1532:41764632
1518:(1956).
1504:16214744
1496:23028975
1451:(2013).
1425:(1814).
1405:(1882).
1373:(1859).
1347:(1985).
1254:(1852).
1240:(1883).
1218:(2018).
1047:FĂ©e 1852
756:system,
681:dealbata
614:dealbata
592:section
499:Missouri
483:dealbata
449:Taxonomy
369:lanceate
297:(Pursh)
289:Kunze
278:(Pursh)
275:dealbata
263:(Pursh)
241:(Pursh)
230:(Pursh)
222:Pursh
210:Synonyms
144:Family:
1781:1055476
1703:2651656
1582:1547438
635:. Both
633:Pellaea
590:Pellaea
577:Pellaea
403:deltate
364:rhizome
280:Davenp.
202:Windham
154:Genus:
134:Order:
124:Class:
95:Plantae
66: (
1939:510777
1815:PLANTS
1794:420011
1755:500964
1716:158622
1677:488511
1580:
1530:
1502:
1494:
1459:
1359:
1279:
1226:
744:rock.
651:sect.
572:Prantl
445:= 54.
431:farina
423:oblong
415:obtuse
396:costae
392:rachis
377:entire
373:linear
314:, the
299:Prantl
1921:IRMNG
1820:ARDE5
1737:IRMNG
1578:JSTOR
1528:JSTOR
1500:S2CID
1492:JSTOR
779:Notes
708:clade
696:is a
568:Presl
471:Pursh
427:ovate
419:acute
388:stipe
265:Kunze
243:Mett.
198:Pursh
102:Clade
1934:ITIS
1908:IPNI
1828:POWO
1776:NCBI
1750:ITIS
1724:IPNI
1698:GBIF
1664:GJMQ
1457:ISBN
1357:ISBN
1277:ISBN
1224:ISBN
794:and
639:and
435:sori
375:and
362:The
273:var.
1960:WFO
1895:FNA
1854:WFO
1685:FNA
1672:EoL
1659:CoL
1570:doi
1484:doi
700:to
675:as
631:or
596:as
588:to
542:as
534:FĂ©e
529:".
521:as
469:by
425:to
371:or
343:).
232:FĂ©e
1984::
1962::
1949::
1936::
1923::
1910::
1897::
1882::
1856::
1843::
1830::
1817::
1804::
1791::
1778::
1765::
1752::
1739::
1726::
1713::
1700::
1687::
1674::
1661::
1646::
1631::
1576:.
1566:77
1564:.
1560:.
1522:.
1514:;
1498:.
1490:.
1480:36
1478:.
1413:.
1409:.
1381:.
1377:.
1331:.
1318:.
1314:.
1298:.
1262:.
1258:.
1214:;
1210:;
951:^
922:^
889:^
854:^
829:^
814:^
689:.
608:,
200:)
104::
1584:.
1572::
1545:"
1541:"
1534:.
1506:.
1486::
1465:.
1415:3
1395:"
1391:"
1383:3
1365:.
1337:6
1320:1
1302:.
1296:"
1292:"
1285:.
1266:.
1264:5
1232:.
1169:.
1049:.
884:.
824:.
443:n
196:(
70:)
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