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Tsuga canadensis

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individuals reach 45 metres (148 ft) tall, and one tree has been measured in the Great Smoky Mountains National Park to 52.8 m (173 ft 3 in) tall, though this tree is now dead from hemlock woolly adelgids; the tallest now surviving, the "Noland Mountain tree", is 51.8 m (169 ft 11 in) tall. Altogether, ENTS has confirmed four trees to heights of 51 m (167 ft) or more by climb and tape drop. In the Northeast, the tallest accurately measured tree is 44 m (144 ft). This tree, named the Seneca hemlock, grows in Cook Forest State Park, PA. Above 43°N latitude, the maximum height of the species is less, under 39 m (128 ft). In New England, ENTS has measured hemlocks to 42 m (138 ft), although trees above 39 m (128 ft) are extremely rare in New England. By 44°N, the maximum height is probably not more than 35 m (115 ft). Diameters of mature hemlocks range from 0.75–1.8 m (2 ft 6 in – 5 ft 11 in), with trees over 1.6 m (5 ft 3 in) in diameter being very rare. In New England, the maximum diameter is 1.4 m (4 ft 7 in).
530: 791:. In cultivation, it prefers sites that are slightly acidic to neutral with nutrient-rich and moist but well-drained soil. It is most often used as a specimen, for a screen, or in small group plantings, though it can also be trained as a dense formal hedge. It should not be used on roadsides where salt is used in winter, as its foliage is sensitive to salt spray. It is also poorly adapted as a windbreak tree, as wind exposure causes dieback in winter. It has several drawbacks, such as a fairly low tolerance of urban stress, intolerance for very wet or very dry soils, and susceptibility to attack by the hemlock woolly adelgid, though this is treatable. Its tendency to shed needles rapidly after being cut down renders it unsuitable as a 598:
heavy snowfall; the higher number is common in southerly areas with high summer rainfall. Near the Atlantic coast and in the southern Appalachians where the trees often reach their greatest heights, annual rainfall often exceeds 1,520 mm (60 in). In the north of its range, the temperatures in January average −12 °C (10 °F), while in July they average only 16 °C (61 °F). In these areas, the frost-free season can last fewer than 80 days. In contrast, the southern end of the range experiences up to 200 days without frost and January temperatures as high as 6 °C (43 °F).
829: 274: 602: 260: 246: 105: 382: 615: 60: 690: 33: 82: 869:'Gentsch White' – a dwarf shrub growing to 1.3 m (4 ft 3 in) tall with an equal spread and new spring growth that turns creamy-white in autumn through winter, creating a dramatic contrast with the dark green old growth, it is easily scorched by the sun and requires partial shade. It is recommend to feather shear annually to keep it compact and create more tip growth. 396:, being at least 554 years old. The tree generally reaches heights of about 31 m (102 ft), but exceptional trees have been recorded up to 53 m (174 ft). The diameter of the trunk at breast height is often 1.5 m (4 ft 11 in), but again, outstanding trees have been recorded up to 1.75 m (5 ft 9 in). The trunk is usually straight and 686:, the pest could kill most of the region's hemlock trees within the next decade. According to the study, researchers found "hemlock woolly adelgid infestation is rapidly impacting the carbon cycle in tree stands," and "adelgid-infested hemlock trees in the South are declining much faster than the reported 9-year decline of some infested hemlock trees in the Northeast." 723:, which is native to the Pacific Northwest, naturally preys on the hemlock wooly adelgid. The particular site near Lansing was chosen because its hemlocks are only lightly infested with the woolly adelgid, and enough trees are found to sustain a long-term study. The site will be left untreated with pesticides for 10 years to study how well the 761:
analyses show the hemlock population experienced a pronounced decline approximately 5,500 BP that lasted for about 1,000 years. Continued research points to other, though less dramatic, dips in Holocene hemlock populations. Pathogens, insects, and climatic change, and a combination of these, have all
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to the United States in 1924, and first found in the native range of eastern hemlock in the late 1960s. The adelgid has spread very rapidly in southern parts of the range once becoming established, while its expansion northward is much slower. Virtually all the hemlocks in the southern Appalachian
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Eastern hemlock is generally confined to areas with highland climates, with cool and humid conditions. Precipitation in the areas where it grows is typically 740 mm (29 in) to more than 1,270 mm (50 in) per year. The lower number is more typical of northern forests that receive
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The leaves are typically 15 to 20 mm (0.59 to 0.79 in) in length, but may be as short as 5 mm (0.20 in) or as long as 25 mm (1 in).The leaves are arranged on little stalks, a characteristic that does not appear in the other evergreen trees. They are flattened and are
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has long been a popular tree in cultivation. The tree's preference for partial shade and tolerance of full shade allows it to be planted in areas where other conifers would not easily grow. In addition, its very fine-textured foliage that droops to the ground, its pyramidal growth habit, and its
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Mountains have seen infestations of the insect within the last five to seven years, with thousands of hectares of stands dying within the last two to three years. Attempts to save representative examples on both public and private lands are on-going. A project named "Tsuga Search", funded by the
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Trunk volume is the third dimension to receive attention by ENTS. Many eastern hemlocks have been modeled to over 30 m (1,100 cu ft) trunk volume, and the largest has been calculated to be 44.8 m (1,580 cu ft), making it the largest natural evergreen conifer in the
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The wood is soft, coarse-grained, and light buff in color. Air-dried, a cubic foot (0.028 m) weighs 28 pounds (13 kg). The lumber is used for general construction and crates. Because of its unusual power of holding spikes, it is also used for railroad ties. Untreated, the wood is not
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Due to it being a long-lived tree, several very large or otherwise impressive trees exist along the east coast of North America. One organization, the Eastern Native Tree Society (ENTS), has been particularly active in discovering and measuring these trees. In the southern Appalachians, many
646:, is being conducted to save the largest and tallest remaining eastern hemlocks in the Park. Through Tsuga Search, hemlocks have been found with trunk volumes up to 44.8 m³ within the park, making it the largest eastern evergreen conifer, eclipsing in volume both eastern white pine ( 892:'Sargentii' – a popular large weeping shrub that grows to 3 m (9.8 ft) tall with a wide spread up to 6 m (20 ft), it features numerous pendulous branches and is most effectively employed near water, in rock gardens, or on embankments. 360:
Eastern hemlocks are long lived trees, with many examples living for more than 500 years. They can grow to heights of more than 30 metres (100 ft), and are tolerant of shade, moist soil, and slopes. Hemlock wood is used in construction, and for
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American pioneers made tea from the tree's leafy twigs and used its branches as brooms. Tea can be made from the needles. The inner bark, which is best in winter and coming into spring, can be eaten raw or boiled; it can also be used to make flour.
889:'Pendula' – an upright weeping form whose height is dependent on how long it is staked, but is typically seen 0.6–1.5 m (2 ft 0 in – 4 ft 11 in) tall with a 1.5-metre (4.9 ft) spread. It has also won the AGM. 924:. Author Edith Wharton described "hemlock boughs bent inward to their trunks by the weight of the snow," "intensely blue shadows of hemlocks on sunlit snow," and darkness "dropping down like a black veil from the heavy hemlock boughs." 851:'Bennett' – a dwarf shrub reaching 1 m (3 ft 3 in) high and 1.5 m (4 ft 11 in) wide, with upper branchlets that first ascend and then arch upper, this selection prefers partial shade. 580:, as well as western Wisconsin and eastern Minnesota. In Canada, it is present in Ontario and all provinces to the east except Newfoundland and Labrador. Its range completely overlaps that of the closely related 872:'Jeddeloh' – a dwarf shrub reaching to 1 m (3 ft 3 in) high and 1.5 m (4 ft 11 in) wide, showing a small concavity in the centre, it is an alternative to the bird's-nest spruce ( 1602:
Zhao, Yan; Yu, Zicheng; Zhao, Cheng (23 April 2010). "Hemlock (Tsuga canadensis) declines at 9800 and 5300 cal. yr BP caused by Holocene climatic shifts in northeastern North America".
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industry. Eastern hemlocks are popular as ornamental trees, thanks to their tolerance of a wide variety of soil and light conditions, as well as their characteristic drooping branches.
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beetles become established; if the experiment proves successful, researchers expect the population will take two to three years to build to levels where they can be readily detected.
866:, it slowly grows to only 30 cm (12 in) tall with a 1.3-metre (4.3 ft) spread, with the central stems eventually becoming visible. It also prefers partial shade. 802:, it is encountered frequently in gardens both large and small, as well as some parks, and is most common in the eastern areas of the country. It is sometimes employed as a 762:
been proposed to explain these anomalies. The eastern hemlock increased again after the major decline, but did not recover its former place as a dominant species.
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occurs at sea level in the north of its distribution, but is found primarily at elevations of 600–1,800 m (2,000–5,900 ft). It ranges from northeastern
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decline of hemlock populations is a much-studied phenomenon. From its foundation in the early Holocene (around 16,000 BP) in what is now the southeastern US,
2577: 1523: 2269: 848:'Beehive' – a very small dwarf shrub typically growing to 1 m (3 ft 3 in) high and 1.5 m wide, resembling a spreading beehive in form 812:(western hemlock); it is not well adapted to the UK climate and as a consequence often has a poorly developed, forked and sinuous trunk there. In 775:
eastern United States. The center of maximum size development for the species is the southern Appalachians, especially the Great Smoky Mountains.
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bands, while the top is a shiny green to yellow-green in color. The leaf margins are very slightly toothed, especially near the apex. The seed
2208: 666:, but this is based largely on its wide distribution and because the adelgid populations have not reached the northern areas of its range. 2572: 2562: 2552: 676:
suggests the hemlock woolly adelgid is killing hemlock trees faster than expected in the southern Appalachians, and rapidly altering the
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durable if exposed to the elements. As a fuel, it is low in value. The wood is also a source of pulp for paper manufacturing.
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have been selected for use, many of them being dwarf forms and shrubs. A partial list of popular cultivars includes:
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Eastern hemlock populations in North America are threatened in much of their range by the spread of the invasive
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Atlas of Relations Between Climatic Parameters and Distributions of Important Trees and Shrubs in North America
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is scaly and deeply fissured, especially with age. The twigs are a yellow-brown in color with darker red-brown
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Gove, J.H.; Fairweather, S.E. (1988). "Tree-ring analysis of a 500-year old hemlock in central Pennsylvania".
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The eastern hemlock grows well in shade and is very long lived, with the oldest recorded specimen, found in
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typically distichous, or two-ranked. The bottom of the leaf is glaucous with two broad and clearly visible
1267:. American Forestry Association. Washington, District of Columbia. 374 pp. ("EASTERN HEMLOCK", pp. 88-89.) 1250: 2316: 1120: 879: 504: in) in width. The apex is more or less rounded and is often projected outward. Twenty-four diploid 1777: 1755: 2481: 2434: 2303: 2086: 1951: 589:
It is found primarily on rocky ridges, ravines, and hillsides with relatively high levels of moisture.
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Thompson, Robert S.; Anderson, Katherine H.; Bartlein, Patrick J. (1999). "Tsuga canadensis".
1524:"Middle-Holocene dynamics of Tsuga canadensis (eastern hemlock) in northern New England, USA" 1291: 883: 561: 327: 215: 2249: 2151: 2006: 1611: 565: 464: in) in width. The scales are ovate to cuneate in shape and measure 8 to 12 mm ( 8: 1835: 1648: 711: 670: 49: 2282: 1615: 1460:(November 1985). "Notes on Clavarioid Fungi. XX. New Taxa and Distributional Records in 2362: 1627: 1546: 1493: 808: 702: 534: 323: 99: 1293: 1263:
Collingwood, C.H. and Warren D. Brush (Revised and Edited by Devereux Butcher). 1974.
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The tree can be found living in association with many forest mushrooms, such as
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are ovoid in shape and are very small, measuring only 1.5 to 2.5 mm (0 to
339: 259: 245: 129: 2259: 1962: 1505: 2526: 2146: 2037: 1853: 1623: 1542: 1489: 1278: 954: 816:, it is the most frequently seen hemlock in cultivation, and is also used in 682: 663: 651: 422: in) in length. These are usually not resinous, but may be slightly so. 401: 354: 232: 69: 400:, but very rarely is forked. The crown is broadly conic, while the brownish 2499: 2334: 1457: 677: 614: 431: 381: 362: 315: 1831:
Edible Wild Plants: A North American Field Guide to Over 200 Natural Foods
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Numerous place names in North America are named Hemlock. For a list, see
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of Pennsylvania. Eastern hemlocks are widespread throughout much of the
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The Audubon Society Field Guide to North American Trees: Eastern Region
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expanded rapidly and successfully into its potential range. However,
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A line drawing of the leaves and cones from Britton and Brown's 1913
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and mainland Europe, where they are used as ornamental trees.
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93. Auflage, p. 424. Quelle & Meyer Verlag, Wiebelsheim.
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are ovoid in shape and typically measure 1.5 to 2.5 cm (
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Bulletin of the Connecticut Agricultural Experiment Station
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Hemlock Trees Dying Rapidly, Affecting Forest Carbon Cycle
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A Field Guide to the Trees of Britain and Northern Europe
1570: 509: 409: 806:, but is considered inferior for this usage compared to 1109:. In Flora of North America Editorial Committee (ed.). 799: 715:
beetles into a stand of adelgid-infested hemlocks near
541:, Pennsylvania; note the hemlocks' deeply fissured bark 1030:. Greensboro, North Carolina: National Plant Data Team 787:
ability to withstand hard pruning make it a desirable
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Schmeil, O., Fitschen, J., & Seybold, S. (2006).
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It was introduced to British gardens in 1736. In the
1333:. In Burns, Russell M.; Honkala, Barbara H. (eds.). 605:
Hemlock boughs in the autumn, shedding older foliage
1577:(1st ed.). London: Chapman and Hall. pp.  1521: 1161:
U.S. Forest Service General Technical Report NC-120
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Illustrated flora of the northern states and Canada
365:. Historically its bark was an important source of 1903:Eastern Native Tree Society's Tsuga Search Project 302:, and in the French-speaking regions of Canada as 1574:Quaternary Ecology: a Paleoecological Perspective 1230:Pinaceae. Drawings and Descriptions of the Genera 2524: 1279:"Eastern hemlock found in Macon County, Alabama" 444:to 1 in) in length and 1.0 to 1.5 cm ( 1375:"Biology and control of hemlock woolly adelgid" 1324: 1158: 568:and Alabama. Disjunct populations occur in the 1571:Delcourt, Hazel R.; Delcourt, Paul A. (1991). 1522:Oswald, W. W.; Foster, D. R. (8 August 2011). 1439: 669:A 2009 study conducted by scientists with the 1828:Elias, Thomas S.; Dykeman, Peter A. (2009) . 969:10.2305/IUCN.UK.2013-1.RLTS.T42431A2979676.en 2578:Natural history of the Great Smoky Mountains 1827: 1112:Flora of North America North of Mexico (FNA) 918:Hemlocks are a recurring image in the novel 878:'Nidiformis'). This cultivar has gained the 625:The species is currently threatened by the 524: 1325:Godman, R. M.; Lancaster, Kenneth (1990). 1014: 943: 484: in) in length by 7.0 to 10 mm ( 258: 244: 80: 58: 31: 1517: 1515: 1224: 1222: 1220: 1218: 1216: 967: 609: 1643: 1641: 1564: 1456: 1450: 1285: 1096: 1094: 1092: 1090: 1088: 1086: 1084: 1082: 1059:Botanical Society of Britain and Ireland 827: 688: 613: 600: 528: 380: 272: 1703: 1428: 1372: 1353:United States Department of Agriculture 1320: 1318: 1152: 353:being listed as Near Threatened on the 2525: 1802: 1672: 1670: 1512: 1446:Predator Beetle to Battle Hemlock Pest 1362:– via Southern Research Station. 1213: 1196: 1173: 2538:NatureServe apparently secure species 2533:IUCN Red List near threatened species 1912: 1911: 1638: 1411: 1407: 1405: 1403: 1276: 1190: 1079: 1042: 1028:The PLANTS Database (plants.usda.gov) 905: 658:). The tree is currently listed as a 2224:ed2625e1-8b1b-4666-80ed-3d0480fd85b8 1733:Welch, H., & Haddow, G. (1993). 1690: 1595: 1315: 1048: 765: 318:to eastern North America. It is the 1866: 1748: 1667: 1412:Earle, Christopher J., ed. (2018). 1167: 955:IUCN Red List of Threatened Species 862:as an alternative to the prostrate 707:University of Massachusetts-Amherst 644:Great Smoky Mountains National Park 338:. They have been introduced in the 13: 2573:Trees of humid continental climate 2563:Flora of the Appalachian Mountains 2553:Trees of the Eastern United States 1898:images at bioimages.vanderbilt.edu 1400: 1197:Blozan, Will (December 18, 2006). 1174:Blozan, Will (February 16, 2007). 1133: 1100: 14: 2604: 1888: 1700:. Forestry Commission Booklet 33. 1200:The Laurel Branch Leviathan Climb 408:, and are densely pubescent. The 279:Kortright Centre for Conservation 277:An Eastern Hemlock branch at the 2487:urn:lsid:ipni.org:names:325665-2 2309:urn:lsid:ipni.org:names:264005-1 1807:. New York: Knopf. p. 300. 1163:. Vol. 1. pp. 483–489. 103: 1860: 1821: 1796: 1770: 1735:The World Checklist of Conifers 1727: 1366: 1270: 1257: 1243: 744: 680:of these forests. According to 2583:Garden plants of North America 1127: 1008: 983: 858:form that can also be used in 778: 701:from the U.S. Forest Service, 508:are present within the trees' 376: 1: 1758:. Royal Horticultural Society 1756:"Tsuga canadensis 'Jeddeloh'" 1698:Conifers in the British Isles 928: 533:Stand of eastern hemlock and 1349:United States Forest Service 823: 572:, northern Alabama, western 7: 1655:. University of Connecticut 1121:Harvard University Herbaria 880:Royal Horticultural Society 10: 2609: 1803:Little, Elbert L. (1980). 1232:. Koeltz Scientific Books 991:"NatureServe Explorer 2.0" 730: 592: 552:eastward through southern 332:Northeastern United States 2558:Trees of Northern America 2378: 1920: 1117:Missouri Botanical Garden 674:Southern Research Station 266: 257: 252: 243: 221: 214: 100:Scientific classification 98: 78: 56: 47: 39: 30: 23: 2593:Plants described in 1763 1696:Mitchell, A. F. (1972). 1676:Mitchell, A. F. (1974). 1624:10.1177/0959683610365932 1543:10.1177/0959683611409774 1341:Silvics of North America 1277:South, David B. (2016). 995:explorer.natureserve.org 525:Distribution and habitat 2568:Symbols of Pennsylvania 1422:The Gymnosperm Database 1373:McClure, M. S. (1987). 896: 854:'Cole's Prostrate' – a 832:The weeping shrub form 539:Tiadaghton State Forest 515: 2548:Hardwood forest plants 1303:U.S. Geological Survey 1177:The Usis Hemlock Climb 1119:, St. Louis, MO & 837: 738:Ramaria flavosaponaria 697:In a 2009 case study, 694: 627:hemlock woolly adelgid 622: 620:hemlock woolly adelgid 610:Hemlock woolly adelgid 606: 542: 394:Tionesta, Pennsylvania 389: 347:Hemlock woolly adelgid 296:eastern hemlock-spruce 282: 267:Closeup view of range 1714:Flora von Deutschland 884:Award of Garden Merit 831: 692: 617: 604: 562:Appalachian Mountains 532: 384: 328:Appalachian Mountains 276: 1653:UConn Plant Database 962:: e.T42431A2979676. 618:Shoot infested with 1616:2010Holoc..20..877Z 1458:Petersen, Ronald H. 1228:Farjon, A. (1990). 1140:www.srs.fs.usda.gov 944:Farjon, A. (2013). 712:Laricobius nigrinus 671:U.S. Forest Service 560:, and south in the 50:Conservation status 40:Large specimens at 1649:"Tsuga canadensis" 1265:Knowing Your Trees 1251:"Studies of Trees" 1101:Taylor, Ronald J. 906:In popular culture 838: 809:Tsuga heterophylla 703:Cornell University 695: 623: 607: 543: 535:eastern white pine 390: 324:Great Lakes region 283: 207:T. canadensis 86:Apparently Secure 16:Species of conifer 2520: 2519: 2245:Open Tree of Life 1914:Taxon identifiers 1873:Project Gutenberg 1845:978-1-4027-6715-9 1588:978-0-412-29790-8 1136:"Eastern Hemlock" 766:Exceptional trees 717:Lansing, New York 633:), a sap-sucking 583:Tsuga caroliniana 271: 270: 93: 73: 2600: 2588:Ornamental trees 2513: 2512: 2503: 2502: 2490: 2489: 2477: 2476: 2464: 2463: 2451: 2450: 2438: 2437: 2425: 2424: 2412: 2411: 2399: 2398: 2397: 2380:Pinus canadensis 2371: 2370: 2358: 2357: 2348: 2347: 2338: 2337: 2325: 2324: 2312: 2311: 2299: 2298: 2286: 2285: 2273: 2272: 2270:tsuga-canadensis 2263: 2262: 2260:Tsuga_canadensis 2253: 2252: 2240: 2239: 2227: 2226: 2217: 2216: 2204: 2203: 2201:NHMSYS0000464649 2191: 2190: 2178: 2177: 2165: 2164: 2155: 2154: 2142: 2141: 2129: 2128: 2116: 2115: 2103: 2102: 2090: 2089: 2077: 2076: 2064: 2063: 2051: 2050: 2041: 2040: 2028: 2027: 2015: 2014: 2002: 2001: 1989: 1988: 1979: 1978: 1976:tsuga-canadensis 1966: 1965: 1956: 1955: 1954: 1952:Tsuga canadensis 1941: 1940: 1939: 1922:Tsuga canadensis 1909: 1908: 1896:Tsuga canadensis 1883: 1882: 1880: 1879: 1867:Wharton, Edith. 1864: 1858: 1857: 1825: 1819: 1818: 1800: 1794: 1793: 1791: 1789: 1780:Tsuga canadensis 1774: 1768: 1767: 1765: 1763: 1752: 1746: 1731: 1725: 1711: 1707: 1701: 1694: 1688: 1674: 1665: 1664: 1662: 1660: 1645: 1636: 1635: 1599: 1593: 1592: 1568: 1562: 1561: 1559: 1557: 1528: 1519: 1510: 1509: 1454: 1448: 1443: 1437: 1432: 1426: 1425: 1416:Tsuga canadensis 1409: 1398: 1397: 1395: 1393: 1379: 1370: 1364: 1363: 1361: 1360: 1345:Washington, D.C. 1329:Tsuga canadensis 1322: 1313: 1312: 1310: 1309: 1300: 1289: 1283: 1282: 1274: 1268: 1261: 1255: 1254: 1247: 1241: 1226: 1211: 1210: 1208: 1207: 1194: 1188: 1187: 1185: 1184: 1171: 1165: 1164: 1156: 1150: 1149: 1147: 1146: 1131: 1125: 1124: 1123:, Cambridge, MA. 1105:Tsuga canadensis 1098: 1077: 1076: 1074: 1073: 1067: 1061:. Archived from 1056: 1046: 1040: 1039: 1037: 1035: 1022:Tsuga canadensis 1012: 1006: 1005: 1003: 1001: 987: 981: 980: 978: 976: 971: 948:Tsuga canadensis 941: 784:Tsuga canadensis 637:introduced from 503: 502: 498: 493: 492: 488: 483: 482: 478: 473: 472: 468: 463: 462: 458: 453: 452: 448: 443: 442: 438: 421: 420: 416: 369:for the leather 351:Tsuga canadensis 305:pruche du Canada 300:Canadian hemlock 290:, also known as 287:Tsuga canadensis 262: 248: 227: 225:Tsuga canadensis 108: 107: 87: 84: 83: 67: 62: 61: 42:Morton Arboretum 35: 25:Tsuga canadensis 21: 20: 2608: 2607: 2603: 2602: 2601: 2599: 2598: 2597: 2523: 2522: 2521: 2516: 2508: 2506: 2498: 2493: 2485: 2480: 2472: 2467: 2459: 2454: 2446: 2441: 2433: 2428: 2420: 2415: 2407: 2402: 2393: 2392: 2387: 2374: 2366: 2361: 2353: 2351: 2343: 2341: 2333: 2328: 2320: 2315: 2307: 2302: 2294: 2289: 2281: 2276: 2268: 2266: 2258: 2256: 2248: 2243: 2235: 2232:Observation.org 2230: 2222: 2220: 2212: 2207: 2199: 2194: 2186: 2181: 2173: 2168: 2160: 2159:MichiganFlora: 2158: 2150: 2145: 2137: 2132: 2124: 2119: 2111: 2106: 2098: 2093: 2085: 2080: 2072: 2067: 2059: 2054: 2046: 2044: 2036: 2031: 2023: 2018: 2010: 2005: 1997: 1992: 1984: 1982: 1974: 1969: 1961: 1959: 1950: 1949: 1944: 1935: 1934: 1929: 1916: 1891: 1886: 1877: 1875: 1865: 1861: 1846: 1838:. p. 122. 1826: 1822: 1815: 1801: 1797: 1787: 1785: 1776: 1775: 1771: 1761: 1759: 1754: 1753: 1749: 1732: 1728: 1709: 1708: 1704: 1695: 1691: 1675: 1668: 1658: 1656: 1647: 1646: 1639: 1600: 1596: 1589: 1569: 1565: 1555: 1553: 1526: 1520: 1513: 1482:10.2307/3793302 1455: 1451: 1444: 1440: 1433: 1429: 1410: 1401: 1391: 1389: 1377: 1371: 1367: 1358: 1356: 1343:. Vol. 1. 1323: 1316: 1307: 1305: 1298: 1290: 1286: 1275: 1271: 1262: 1258: 1249: 1248: 1244: 1227: 1214: 1205: 1203: 1195: 1191: 1182: 1180: 1172: 1168: 1157: 1153: 1144: 1142: 1132: 1128: 1099: 1080: 1071: 1069: 1065: 1054: 1047: 1043: 1033: 1031: 1013: 1009: 999: 997: 989: 988: 984: 974: 972: 942: 935: 931: 908: 899: 826: 789:ornamental tree 781: 768: 747: 733: 693:Closeup of bark 662:species in the 660:near threatened 612: 595: 570:Piedmont region 527: 518: 500: 496: 495: 490: 486: 485: 480: 476: 475: 470: 466: 465: 460: 456: 455: 450: 446: 445: 440: 436: 435: 418: 414: 413: 379: 336:Maritime Canada 292:eastern hemlock 239: 229: 223: 210: 102: 94: 85: 81: 74: 65:Near Threatened 63: 59: 52: 17: 12: 11: 5: 2606: 2596: 2595: 2590: 2585: 2580: 2575: 2570: 2565: 2560: 2555: 2550: 2545: 2540: 2535: 2518: 2517: 2515: 2514: 2504: 2491: 2478: 2465: 2452: 2439: 2426: 2413: 2400: 2384: 2382: 2376: 2375: 2373: 2372: 2368:wfo-0000456302 2359: 2349: 2339: 2326: 2313: 2300: 2287: 2274: 2264: 2254: 2241: 2228: 2218: 2205: 2192: 2179: 2166: 2156: 2143: 2130: 2117: 2104: 2091: 2078: 2065: 2052: 2042: 2029: 2016: 2003: 1990: 1980: 1967: 1957: 1942: 1926: 1924: 1918: 1917: 1906: 1905: 1900: 1890: 1889:External links 1887: 1885: 1884: 1859: 1844: 1820: 1813: 1795: 1769: 1747: 1737:. Landsman's. 1726: 1702: 1689: 1666: 1637: 1610:(6): 877–886. 1594: 1587: 1563: 1511: 1476:(6): 903–919. 1449: 1438: 1427: 1399: 1365: 1314: 1284: 1269: 1256: 1242: 1212: 1189: 1166: 1151: 1134:Godman, R. M. 1126: 1078: 1051:BSBI List 2007 1041: 1007: 982: 932: 930: 927: 926: 925: 916: 907: 904: 898: 895: 894: 893: 890: 887: 870: 867: 852: 849: 825: 822: 793:Christmas tree 780: 777: 767: 764: 746: 743: 732: 729: 631:Adelges tsugae 611: 608: 594: 591: 526: 523: 517: 514: 378: 375: 340:United Kingdom 269: 268: 264: 263: 255: 254: 253:Natural range 250: 249: 241: 240: 230: 219: 218: 212: 211: 204: 202: 198: 197: 190: 186: 185: 180: 176: 175: 170: 166: 165: 160: 156: 155: 150: 146: 145: 140: 133: 132: 127: 120: 119: 114: 110: 109: 96: 95: 79: 76: 75: 57: 54: 53: 48: 45: 44: 37: 36: 28: 27: 15: 9: 6: 4: 3: 2: 2605: 2594: 2591: 2589: 2586: 2584: 2581: 2579: 2576: 2574: 2571: 2569: 2566: 2564: 2561: 2559: 2556: 2554: 2551: 2549: 2546: 2544: 2541: 2539: 2536: 2534: 2531: 2530: 2528: 2511: 2505: 2501: 2496: 2492: 2488: 2483: 2479: 2475: 2470: 2466: 2462: 2457: 2453: 2449: 2444: 2440: 2436: 2431: 2427: 2423: 2418: 2414: 2410: 2405: 2401: 2396: 2390: 2386: 2385: 2383: 2381: 2377: 2369: 2364: 2360: 2356: 2350: 2346: 2340: 2336: 2331: 2327: 2323: 2318: 2314: 2310: 2305: 2301: 2297: 2292: 2288: 2284: 2279: 2275: 2271: 2265: 2261: 2255: 2251: 2246: 2242: 2238: 2233: 2229: 2225: 2219: 2215: 2210: 2206: 2202: 2197: 2193: 2189: 2184: 2180: 2176: 2171: 2167: 2163: 2157: 2153: 2148: 2144: 2140: 2135: 2131: 2127: 2122: 2118: 2114: 2109: 2105: 2101: 2096: 2092: 2088: 2083: 2079: 2075: 2070: 2066: 2062: 2057: 2053: 2049: 2043: 2039: 2034: 2030: 2026: 2021: 2017: 2013: 2008: 2004: 2000: 1995: 1991: 1987: 1981: 1977: 1972: 1968: 1964: 1958: 1953: 1947: 1943: 1938: 1932: 1928: 1927: 1925: 1923: 1919: 1915: 1910: 1904: 1901: 1899: 1897: 1893: 1892: 1874: 1870: 1869:"Ethan Frome" 1863: 1855: 1851: 1847: 1841: 1837: 1833: 1832: 1824: 1816: 1814:0-394-50760-6 1810: 1806: 1799: 1783: 1781: 1773: 1757: 1751: 1744: 1743:0-900513-09-8 1740: 1736: 1730: 1723: 1722:3-494-01413-2 1719: 1715: 1706: 1699: 1693: 1687: 1686:0-00-212035-6 1683: 1679: 1673: 1671: 1654: 1650: 1644: 1642: 1633: 1629: 1625: 1621: 1617: 1613: 1609: 1605: 1598: 1590: 1584: 1580: 1576: 1575: 1567: 1552: 1548: 1544: 1540: 1536: 1532: 1525: 1518: 1516: 1507: 1503: 1499: 1495: 1491: 1487: 1483: 1479: 1475: 1471: 1467: 1463: 1459: 1453: 1447: 1442: 1436: 1431: 1423: 1419: 1417: 1408: 1406: 1404: 1387: 1383: 1376: 1369: 1354: 1350: 1346: 1342: 1338: 1337: 1332: 1330: 1321: 1319: 1304: 1297: 1296: 1288: 1280: 1273: 1266: 1260: 1252: 1246: 1239: 1238:3-87429-298-3 1235: 1231: 1225: 1223: 1221: 1219: 1217: 1202: 1201: 1193: 1179: 1178: 1170: 1162: 1155: 1141: 1137: 1130: 1122: 1118: 1114: 1113: 1108: 1106: 1097: 1095: 1093: 1091: 1089: 1087: 1085: 1083: 1068:on 2015-06-26 1064: 1060: 1053: 1052: 1045: 1029: 1025: 1023: 1017: 1011: 996: 992: 986: 970: 965: 961: 957: 956: 951: 949: 940: 938: 933: 923: 922: 917: 914: 910: 909: 903: 891: 888: 885: 881: 877: 876: 871: 868: 865: 861: 857: 853: 850: 847: 846: 845: 843: 835: 834:T. canadensis 830: 821: 819: 815: 811: 810: 805: 801: 796: 794: 790: 785: 776: 772: 763: 760: 759:palynological 756: 755:T. canadensis 752: 742: 740: 739: 728: 726: 722: 718: 714: 713: 709:released 900 708: 704: 700: 699:entomologists 691: 687: 685: 684: 683:Science Daily 679: 675: 672: 667: 665: 664:IUCN Red List 661: 657: 653: 652:loblolly pine 649: 648:Pinus strobus 645: 640: 636: 632: 628: 621: 616: 603: 599: 590: 587: 585: 584: 579: 575: 571: 567: 563: 559: 555: 551: 547: 546:T. canadensis 540: 536: 531: 522: 513: 511: 507: 433: 429: 423: 411: 407: 403: 399: 395: 388: 383: 374: 372: 368: 364: 363:railroad ties 358: 356: 355:IUCN Red List 352: 348: 343: 341: 337: 333: 329: 325: 321: 317: 314: 311: 307: 306: 301: 297: 293: 289: 288: 280: 275: 265: 261: 256: 251: 247: 242: 238: 234: 228: 226: 220: 217: 216:Binomial name 213: 209: 208: 203: 200: 199: 196: 195: 191: 188: 187: 184: 181: 178: 177: 174: 171: 168: 167: 164: 161: 158: 157: 154: 151: 148: 147: 144: 141: 138: 135: 134: 131: 130:Tracheophytes 128: 125: 122: 121: 118: 115: 112: 111: 106: 101: 97: 91: 77: 71: 66: 55: 51: 46: 43: 38: 34: 29: 26: 22: 19: 2379: 1921: 1895: 1876:. Retrieved 1872: 1862: 1834:. New York: 1830: 1823: 1804: 1798: 1786:. Retrieved 1779: 1772: 1760:. Retrieved 1750: 1734: 1729: 1713: 1705: 1697: 1692: 1677: 1657:. Retrieved 1652: 1607: 1604:The Holocene 1603: 1597: 1573: 1566: 1554:. Retrieved 1537:(1): 71–78. 1534: 1531:The Holocene 1530: 1473: 1469: 1465: 1461: 1452: 1441: 1430: 1421: 1415: 1390:. Retrieved 1385: 1381: 1368: 1357:. Retrieved 1340: 1335: 1328: 1306:. Retrieved 1294: 1287: 1272: 1264: 1259: 1245: 1229: 1204:. Retrieved 1199: 1192: 1181:. Retrieved 1176: 1169: 1160: 1154: 1143:. Retrieved 1139: 1129: 1111: 1104: 1070:. Retrieved 1063:the original 1050: 1044: 1032:. Retrieved 1027: 1021: 1010: 998:. Retrieved 994: 985: 973:. Retrieved 959: 953: 947: 919: 900: 873: 839: 833: 807: 797: 783: 782: 773: 769: 754: 748: 745:Paleoecology 736: 734: 724: 720: 710: 696: 681: 678:carbon cycle 668: 655: 647: 630: 624: 596: 588: 581: 564:to northern 545: 544: 519: 424: 391: 386: 359: 350: 344: 304: 303: 299: 295: 291: 286: 285: 284: 224: 222: 206: 205: 193: 143:Gymnospermae 136: 123: 24: 18: 2283:kew-2516507 2183:NatureServe 2095:iNaturalist 1946:Wikispecies 1710:(in German) 1392:October 24, 1034:12 December 975:11 November 921:Ethan Frome 875:Picea abies 856:groundcover 836:'Sargentii' 779:Cultivation 725:L. nigrinus 721:L. nigrinus 656:Pinus taeda 558:Nova Scotia 506:chromosomes 377:Description 90:NatureServe 2527:Categories 2352:WisFlora: 2278:Plant List 1878:2024-09-18 1782:'Pendula'" 1680:. Collins 1506:7377077277 1359:2007-07-05 1308:2007-07-05 1206:2007-06-08 1183:2007-06-08 1145:2021-12-13 1072:2014-10-17 1016:USDA, NRCS 929:References 705:, and the 398:monopodial 320:state tree 310:coniferous 149:Division: 2395:Q17359215 2061:233501320 1854:244766414 1659:19 August 1632:220988296 1490:0027-5514 1470:Mycologia 1462:Clavulina 842:cultivars 840:Over 300 824:Cultivars 639:East Asia 576:and into 556:and into 550:Minnesota 201:Species: 163:Pinopsida 153:Pinophyta 113:Kingdom: 2507:VASCAN: 2500:24900561 2495:Tropicos 2461:10872510 2448:325665-2 2389:Wikidata 2342:VASCAN: 2335:24900236 2330:Tropicos 2257:PalDat: 2188:2.131718 2126:10586135 2113:264005-1 1983:BioLib: 1937:Q1137143 1931:Wikidata 1836:Sterling 1762:13 March 1551:42106379 1351:(USFS), 1336:Conifers 1018:(n.d.). 1000:12 April 864:junipers 818:forestry 751:Holocene 749:The mid- 428:stomatal 237:Carrière 183:Pinaceae 179:Family: 70:IUCN 3.1 2422:5285847 2170:MoBotPF 2074:2687182 2012:1033654 1788:5 March 1612:Bibcode 1556:6 March 1498:3793302 1466:Ramaria 913:Hemlock 814:Germany 731:Ecology 593:Climate 578:Indiana 566:Georgia 499:⁄ 489:⁄ 479:⁄ 469:⁄ 459:⁄ 449:⁄ 439:⁄ 417:⁄ 406:pulvini 371:tanning 308:, is a 189:Genus: 173:Pinales 169:Order: 159:Class: 117:Plantae 88: ( 68: ( 2474:183398 2291:PLANTS 2250:117276 2221:NZOR: 2175:285010 2139:183397 2048:tsucan 2045:FEIS: 2038:150566 1971:ARKive 1852:  1842:  1811:  1741:  1720:  1684:  1630:  1585:  1549:  1504:  1496:  1488:  1355:(USDA) 1236:  860:bonsai 650:) and 554:Quebec 367:tannin 334:, and 330:, the 326:, the 316:native 2543:Tsuga 2510:22101 2456:IRMNG 2435:28424 2409:4J242 2322:18463 2267:PPE: 2214:66173 2152:42431 2121:IRMNG 2100:48734 2087:40698 2033:EUNIS 2025:TSUCA 1999:59HM6 1960:APA: 1784:. RHS 1628:S2CID 1579:43–44 1547:S2CID 1527:(PDF) 1494:JSTOR 1388:: 1–9 1378:(PDF) 1299:(PDF) 1066:(xls) 1055:(xls) 804:hedge 432:cones 298:, or 194:Tsuga 137:Clade 124:Clade 2482:POWO 2469:ITIS 2443:IPNI 2430:GRIN 2417:GBIF 2355:5291 2345:7202 2304:POWO 2296:TSCA 2237:7587 2209:NCBI 2162:1927 2147:IUCN 2134:ITIS 2108:IPNI 2082:GRIN 2069:GBIF 2020:EPPO 1986:2391 1850:OCLC 1840:ISBN 1809:ISBN 1790:2021 1764:2021 1739:ISBN 1718:ISBN 1682:ISBN 1661:2013 1583:ISBN 1558:2013 1502:OCLC 1486:ISSN 1464:and 1394:2011 1234:ISBN 1036:2015 1002:2022 977:2021 960:2013 897:Uses 574:Ohio 516:Wood 410:buds 402:bark 313:tree 2404:CoL 2363:WFO 2317:RHS 2196:NBN 2056:FNA 2007:EoL 1994:CoL 1963:219 1620:doi 1539:doi 1478:doi 1468:". 1386:851 964:doi 882:'s 635:bug 537:in 510:DNA 494:to 474:to 454:to 2529:: 2497:: 2484:: 2471:: 2458:: 2445:: 2432:: 2419:: 2406:: 2391:: 2365:: 2332:: 2319:: 2306:: 2293:: 2280:: 2247:: 2234:: 2211:: 2198:: 2185:: 2172:: 2149:: 2136:: 2123:: 2110:: 2097:: 2084:: 2071:: 2058:: 2035:: 2022:: 2009:: 1996:: 1973:: 1948:: 1933:: 1871:. 1848:. 1669:^ 1651:. 1640:^ 1626:. 1618:. 1608:20 1606:. 1581:. 1545:. 1535:22 1533:. 1529:. 1514:^ 1500:. 1492:. 1484:. 1474:77 1472:. 1420:. 1402:^ 1384:. 1380:. 1347:: 1339:. 1317:^ 1301:. 1215:^ 1138:. 1081:^ 1057:. 1026:. 993:. 958:. 952:. 936:^ 820:. 800:UK 795:. 741:. 719:. 586:. 512:. 357:. 294:, 235:) 233:L. 139:: 126:: 1881:. 1856:. 1817:. 1792:. 1778:" 1766:. 1745:. 1724:. 1663:. 1634:. 1622:: 1614:: 1591:. 1560:. 1541:: 1508:. 1480:: 1424:. 1418:" 1414:" 1396:. 1331:" 1327:" 1311:. 1281:. 1253:. 1240:. 1209:. 1186:. 1148:. 1107:" 1103:" 1075:. 1038:. 1024:" 1020:" 1004:. 979:. 966:: 950:" 946:" 915:. 886:. 654:( 629:( 501:8 497:3 491:4 487:1 481:2 477:1 471:8 467:3 461:8 457:5 451:8 447:3 441:8 437:5 419:8 415:1 281:. 231:( 92:) 72:)

Index


Morton Arboretum
Conservation status
Near Threatened
IUCN 3.1
NatureServe
Scientific classification
Edit this classification
Plantae
Tracheophytes
Gymnospermae
Pinophyta
Pinopsida
Pinales
Pinaceae
Tsuga
Binomial name
L.
Carrière



Kortright Centre for Conservation
coniferous
tree
native
state tree
Great Lakes region
Appalachian Mountains
Northeastern United States

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