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the
Animals Live. Due to large amounts of food left by people feeding them, colonies of cats are continuing to increase, with estimates putting the population within the city of Jerusalem at 2,000 cats per square kilometer. Efforts to trap, neuter, return the cats within Israel are not working, as the population is too large to feasibly catch enough cats to make a difference. Moreover, there is no national agreement on what to do regarding the cat population. As a result their population is increasing with no future plan of action. In January 2019, from a push by people who feed the cats, Jerusalem planned to instill "feeding stations" throughout the city. The goal was to facilitate specific areas for feeding to help the populations of stray cats and to improve their welfare. This plan was criticized by ecologists and conservationists, stating that it does nothing to help the welfare of the cats, with Amir Balaban of the Society for the Protection of Nature stating that "If someone cares about animals, they should take them home."
329:, particularly of intact cats, has been noted to be a barrier to decreasing cat populations over time through TNR efforts. It has become apparent that while the TNR process can reduce or limit the growth rate of the colony through reproduction, it may not reduce the population numbers if it is the sole method of intervention. Population reduction occurs primarily through adoptions of non-feral cats, natural death or euthanasia of sick animals, and disappearance or emigration of cats. TNR works together with these factors to reduce reproduction and thus to minimize replacement of animals lost from the colony. Other factors such as immigration of cats from surrounding areas can counteract its effect. Thus, the impact of TNR interventions on unowned cat populations can be complex, and ongoing management of colonies becomes an important component in optimizing reductions in the cat population.
208:: "As a long term strategy, in most of Australia, is difficult to recommend. Since resources for cat control are limited, resources in and around towns and cities would be better spent on education, increased community awareness about responsible cat ownership, targeted desexing programs particularly for low-income earners, and better laws and regulations. These strategies should reduce the number of owned animals and their offspring entering the unowned cat population. For remote Australia where feral cats are completely unsocialised and therefore not candidates for rehoming, the most cost-effective and humane option is likely to be targeted and ongoing lethal control in priority areas where adverse environmental impacts are highest."
238:(HSUS): "we support Trap-Neuter-Return (TNR) and similar sterilization programs, legislation that allows for and supports non-lethal population control, and coalition-based approaches that involve community leaders, citizens, and stakeholders to implement effective community cat management programs." They further clarify this view by stating: "The HSUS believes that the humane reduction and eventual elimination of unowned cat populations should be the end goal for all TNR participants and supporters. TNR should be considered a humane means to an end, not a method of permanently maintaining outdoor cat populations. The HSUS estimates that the population of feral cats in the US ranges from 50 to 70
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permanently removed by a combination of rehoming and euthanasia on an ongoing basis, as well as by an unusually high rate of death and disappearance. TNR colonies often increase in population because cats breed quickly and the trapping and sterilization rates are frequently too low to stop this population growth, because food is usually being provided to the cats, and because public awareness of a TNR colony tends to encourage people in the surrounding community to dump their own unwanted pet cats there. The growing popularity of TNR, even near areas of particular ecological sensitivity, has been attributed in part to a lack of public interest regarding
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states require animal shelters to keep records about animals being euthanized, and there is no agency responsible for collecting or verifying this data. The
American Society for the Prevention of Cruelty to Animals (ASPCA) has noted a marked decrease in euthanasia rates since 2011 and the Humane Society of the United States reported that euthanasia of animals in shelters has been declining sharply since 1970. In addition, the reasons for euthanasia vary. Animals may be euthanized because of shelter over-crowding, for medical reasons (illness or injury), for court-mandated reasons, or because of financial/staff limitations.
31:
292:, but the methodology, analysis and conclusions of some of these studies have been called into question. Reviews of these studies, as well as mathematical models of population growth, have shown that TNR colonies only decrease in population where cats experience very high rates of permanent removal (of at least 50% of the colony cats per year, such as by a combination of adoption and euthanasia), combined with high sterilization rates, and low rates of immigration of new cats into the colony. Unless all of these conditions are met, TNR colonies will not decrease in size over time.
446:. Some are owned, but have been allowed to roam outdoors; some have escaped their homes or owners and are strays; some may have been abandoned or "dumped." The assessment, after trapping, of "social" (friendly and adoptable), "social but timid or scared" (may adjust and be adoptable), "not social" (not feral, prefers to not be handled, hard to adopt out) or "feral" is crucial if TNR is intended to be in the best interest of the animal. When programs provide for feral kittens to be socialized and adopted, and for friendly cats to be adopted, the welfare of those cats is improved.
61:, then releasing them back into the outdoors. If the location is deemed unsafe or otherwise inappropriate, the cats may be relocated to other appropriate areas (barn/farmyard homes are often considered best). Often, friendly adults and kittens young enough to be easily socialized are retained and placed for adoption. Feral cats cannot be socialized, shun most human interaction and do not fare well in confinement, so they are not retained. Cats suffering from severe medical problems such as terminal, contagious, or untreatable illnesses or injuries are often euthanized.
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742:, since 1988. A study in 2006 found almost 8,000 were neutered and reintroduced to their original colony from 1991 to 2000. It concluded that spay/neuter campaigns brought about a general decrease in cat numbers among registered colonies and censused cats, but the percentage of cat immigration (due to abandonment and spontaneous arrival) was around 21 percent. It suggested that TNR efforts without an effective education of people to control the reproduction of house cats (as a prevention for abandonment) are a waste of money, time and energy.
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for a couple of days" where they could be observed in recuperation. WFLA said they had possession of images of TNR cats bleeding from opened surgical wounds, images of TNR cat with maggots in their incisions, and an image of a bloodied cat trap whose occupant had bled out; the TV station's managers declined to allow these images to be broadcast. Officials overseeing that TNR program said that their greatly reduced shelter euthanasia rates were worth it even if their TNR outcomes "weren't perfect".
190:: "The IWRC supports the humane removal of feral cat and dog populations, including feral cat colonies, through the rehabilitation and adoption of suitable animals into domestic environments and humane euthanasia of animals that cannot be rehabilitated and re-homed." They point out that as domestic animals "subsidized" by people, they exceed the ability for the environment to support them without resulting in damage to wildlife.
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towards providing a better quality of life for feral cats, stemming the population expansion that is a direct result of breeding, improving the communities in which these cats are found, reducing "kill" rates at shelters that accept captured free-roaming cats, in turn improving public perceptions and possibly reducing costs, and eliminating or reducing nuisance behaviors to decrease public complaints about free-roaming cats.
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of a successful prosecution under the MBTA was United States v. Moon Lake
Electric Association Incorporated; in that case a power company was found guilty and sentenced to fines and probation because birds were being electrocuted on power poles that it owned.) Because TNR colonies are supported by people, it is possible that predation of birds by TNR cats might be ruled an illegal intentional take by the courts.
202:(ABC): "Cats have been introduced into new habitats across the globe with terrible results. Outdoors, cats are a non-native and invasive species that threaten birds and other wildlife, disrupt ecosystems, and spread diseases." They advocate responsible pet ownership and "oppose Trap, Neuter, Release (TNR) for feral cats because of the persistent and severe threats posed by feral cat colonies."
156:('trap, neuter, shelter return') are alternative approaches that simply focus on the trap and desex portion and do not include a colony management aspect. In some instances, a receiving shelter will return a cat to where it was found; in other cases shelters are completely bypassed – a person takes a free-roaming live-trapped cat in for desexing, then returns it to where it was found.
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control measures, has been a contentious point. In a widely cited example of cat control by relocation (cats from
Bidwell Park, California, were trapped and moved to a private sanctuary), the high visibility of the project encouraged more abandonment. In another study, to explain the ingression of cats it was found that "the high quality and visibility of the program
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supported TNR programs, but these programs are often scorned by the public and poorly managed. Negative attitudes and fear towards cats in general have been slow to change and free-roaming cats may be subjected to abuse or violence. In recent years, however, South Korea's attitude toward feral cats has improved.
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of TNR program implemented, the extent of animal control involvement, the volunteer base available, and the community's overall support of TNR. Over time, through attrition and sterilization efforts, if the free-roaming population declines, savings may be realized by total decreased expenditures on them.
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The TNR process itself can result in inhumane outcomes. In 2016, investigative reporting by WFLA found that in
Hillsborough County, Florida, TNR cats were being returned to the outdoors the day after their surgical sterilizations, because the Tampa Bay Humane Society did not have "a place to put them
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may have encouraged abandonment of cats if owners believed that the cats would be well taken care of after abandonment. Abandonment may also have occurred if owners believed that cats would be better off under the care of the program rather than surrendered to a shelter where they would face the risk
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achieved a population decrease of 66%, from 68 cats in 1996 (when the census was first completed after some trapping) to 23 cats in 2002. No new kittens were born after 1995, and newly arrived stray or abandoned cats were neutered or adopted to homes. However, as many proponents fail to note, TNR was
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any migratory bird, any part, nest, or egg of any such bird," including unintentional taking, except if otherwise regulated or permitted by the government, and provides for criminal penalties of up to $ 15,000 in fines and up to six months of incarceration for each bird unlawfully taken. (An example
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Across Canada, municipalities have replaced old animal control bylaws with "responsible pet ownership" rules intended to direct the obligations of pet behavior to their owners. A common feature of the accelerating trend is a requirement that owners get a license for their cats and ensure they do not
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The success of specific focused studies to advocate TNR as a solution for controlling and reducing free-roaming cat populations worldwide is problematic. More broad-based approaches include using matrix population models to estimate the efficacy of euthanasia versus trap-neuter-return for management
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Like Turkey, Israel struggles with a continually increasing population of stray cats. Cats exist in every location with people, from the southernmost city of Eilat to communities in the Golan
Heights. Moreover, it is illegal in Israel to remove cats from the streets as a result of pressure from Let
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A 2011 survey of U.S. pet owners found that 71% agreed that "Animal shelters should only be allowed to euthanize animals when they are too sick to be treated or too aggressive to be adopted," while only 25% agreed that "Sometimes animal shelters should be allowed to euthanize animals as a necessary
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them costs more. However, the cost savings associated with TNR are location-specific and accurate estimates involve taking into account numerous variables including volunteer support, donations, grants, and local spay-neuter agreements for low-cost services. Cost savings fluctuate based on the type
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A TNR program begun in 1992 by the
Merrimack River Feline Rescue Society (MRFRS) on the central waterfront of Newburyport, Massachusetts, has been widely cited as an example of TNR success on a community level; however, only superficial reports about what took place have been available and there is
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In the past, the main goal of most TNR programs was the reduction or eventual elimination of free-roaming cat populations. It is still the most widely implemented non-lethal method of managing them. While that is still a primary goal of many efforts, other programs and initiatives may be aimed more
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Stray animals in general may have significant impacts on public health due to factors such as a lack of preventive measures (e.g. vaccines, deworming), easy access to intermediate hosts (e.g. rats and birds), and unrestricted entry to public areas such as parks and playgrounds. Their presence is a
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TNR of cats is illegal in Alaska, owing to a law against the release of cats into the wild, even if they were originally captured there. This has left trap-and-kill the only legal method of controlling the feral cat population there; however, the law against TNR is not well enforced and there are
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It has been claimed that euthanasia in shelters is the leading cause of death of cats. Proponents of TNR use this statistic to promote methods of cat population control that do not include euthanasia. However, in the USA there are no exact numbers of animals being euthanized each year. Only a few
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gained publicity for her TNR work "at a time when euthanasia of feral cats was considered the only option". Hammond "fought many battles with local authorities, hospitals, environmental health departments" but stated that she succeeded over the years in showing that control "could be achieved by
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Turkey has a significant problem with free-roaming dogs and cats and the country is struggling with ways to manage the problem. Its Animal
Protection Law prohibits killing "ownerless animals" except where permitted by the Animal Health Police Law. They are required to be taken to animal shelters
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TNR may help reduce public complaints pertaining to free-roaming cats. Female cats will 'call' (come into season and be receptive to the male cat) regularly, about every three weeks during sexually active times of the year if they do not get pregnant. Having un-spayed female cats in an area will
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that need to be eliminated. Many international, national, and regional organizations and association, both professional and advocacy-based, have publicly aligned themselves into three basic groups: those that stridently oppose managing, maintaining, or tolerating free-roaming cats and hence TNR;
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The potential problem of TNR advocacy and increased public awareness of non-lethal intervention actually contributing to the increasing numbers of free-roaming pets, by enabling pet owners to make conscious decisions to illegally dump or abandon their animal without having to worry about lethal
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Scientific research has not found TNR to be an effective means of controlling the feral cat population. Literature reviews have found that when studies documented TNR colonies that declined in population, those declines were being driven primarily by substantial percentages of colony cats being
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There have been recorded instances of species extinction caused by them on islands. In 2004, targeted eradication programs had successfully removed free-roaming domestic cats from at least 49 islands. Citing eradication of invasive mammals from islands as a proven conservation tool, with clear
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The overall effect of TNR on the health and welfare of free-roaming cats as a whole is not possible to measure. In numerous studies, many of the cats simply disappeared, so follow-up was impossible. They are subject to injury, illness, or death from numerous things: trauma from humans or human
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Fighting may decline, thus reducing injuries: A study between four colonies, two of neutered males and two of intact males, found that the frequency of agonistic behavior was lower in the neutered groups. However, the agonistic behavior that was noted in the neutered groups was attributable to
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When the number of animals coming into a shelter exceeds its ability to care for, hold, or find foster placements, the facility may end up euthanizing animals. This could include even adoptable kittens or cats simply because they cannot be taken care of. A common outcome for a cat judged to be
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TNR programs may have a side effect of reducing the stress and strain volunteers and staff have related to euthanasia in shelters. A 2019 study concluded that euthanasia-related strain is prevalent among shelter employees. Such strain is associated with increased levels of general job stress,
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Alexandra Newman, David Smith, Ria R Ghai, Ryan M Wallace, Mia Kim
Torchetti, Christina Loiacono, Laura S Murrell, Ann Carpenter, Scott Moroff, Jane A Rooney, Casey Barton Behravesh. "First reported cases of SARS-CoV-2 infection in companion animals—New York, March–April 2020." Morbidity and
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The
Department of Conservation (DOC) is legislatively mandated to control feral cats on public conservation land. It has eradicated feral cats from several offshore islands. Control techniques include poisoning, trapping and shooting. Lethal controls follow efficient and humane best-practice
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Negative attitudes towards cats in general and free-roaming cats in particular are culturally entrenched in South Korea. The 2011 South Korean amendment of its Animal Protection Law required humane methods to be used in the transportation and euthanasia of animals. Some areas have government
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Improved overall heath: The American Veterinary Medical Association (AVMA) notes that properly managed programs can improve quality of life through better nutrition, vaccination to prevent disease, and euthanasia of sick and debilitated cats. However, management of colonies is not a mandated
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In Port Orange, Florida, a TNR program started in 2013 in the city's business areas resulted in fewer stray cats and money saved. In the first year, 214 cats were sterilized for $ 13,000, which was much less than over $ 50,000 spent in 2010, when most of the impounded cats were euthanized. A
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Facilities have reported notable decreases in intakes and euthanasia since implementing TNR programs. It is not clear, however, if these decreases can be directly attributed to TNR, or to concurrent efforts to increase and implement owned pet spay-neuter programs, new initiatives of adoption
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Efforts to assess the effectiveness have been hampered by the lack of sufficient monitoring data. Having some professional assistance, adapting the population monitoring framework developed over decades by wildlife biologists, and systematic monitoring can evolve into a relatively low-cost,
253:: "In some situations, safe cat colonies can be maintained by caretakers. American Humane Association supports trap, neuter and release programs for colony cats – especially for feral cats. Whenever possible, homes should be found for colony cats that might be successfully socialized."
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There are legal theories that TNR programs may have liability under the Migratory Bird Treaty Act of 1918, a strict liability statute that states that "it shall be unlawful at any time, by any means or in any manner, to pursue, hunt, take, capture, kill, attempt to take, capture, or
232:: "The ASPCA endorses Trap-Neuter-Return (TNR) as the only proven humane and effective method to manage community cat colonies." It clarifies its position by stating that managing "involves a colony caretaker who provides food and adequate shelter and monitors the cats' health."
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Ali Taghipour, Sahar Ghodsian, Mina Shajarizadeh, Mitra Sharbatkhori, Sasan Khazaei, Hamed Mirjalali. "Global prevalence of microsporidia infection in cats: a systematic review and meta-analysis of an emerging zoonotic pathogen." Preventive Veterinary Medicine 188, 105278,
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established or permitted by the local authorities. In keeping with the tenents of its main religion, most Turks are very much against euthanasia of animals for "population control"; recent efforts to curb an ever-increasing population include TNR for roaming cats and dogs.
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when a no-kill policy was introduced for both cats and dogs. Feral cats have the right to live free and cannot be permanently removed from their colony; cat caretakers can be formally registered; and TNR methods are outlined in the national law on the management of pets.
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Spayed female cats will no longer be burdened by pregnancy or nursing litters: Females have been found to be pregnant throughout the year. A study of the reproductive capacity of free-roaming cats showed they may have an average of 1.4 litters a year, with a median of 3
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In 2019, a study was published that concluded "for most Brisbane City (Australia) residents, when awareness is raised about the problem of urban stray cats and management strategies, the majority are supportive of a TNR community program with little or no persuasion
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Ali Rostami, Mahdi Sepidarkish, Guangxu Ma, Tao Wang, Maryam Ebrahimi, Yadolah Fakhri, Hamed Mirjalali, Andreas Hofmann, Calum NL Macpherson, Peter J Hotez, Robin B Gasser. "Global prevalence of Toxocara infection in cats". Advances in Parasitology 109, 615-639,
214:: "As typically applied, trap-neuter-release (TNR) alone is ineffective at reducing cat population sizes over reasonable time frames (e.g., less 10 years), with removal strategies such as high-volume adoptions, relocations and euthanasia being more efficacious."
497:. They cause considerable wildlife destruction and ecosystem disruption, including the deaths of hundreds of millions of birds, small mammals, reptiles, amphibians, and fish. They have been linked to the extinction of 63 species and pose a threat to 360 more.
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million feral cats – about a third of the population – by saying that they are "the single biggest threat to our native animals, and have already directly driven into extinction 20 out of 30 mammals lost." This cull was planned to continue until 2020.
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million of them are feral. Control of free-roaming dogs and cats is a worldwide problem. Beyond pragmatic and scientific considerations, cultural heritage, ethical beliefs, and social and economic impacts play critical roles in efforts to address it.
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in the mid-1970s, as reported at the 1980 Universities Federation for Animal Welfare (UFAW) symposium in London. Denmark's Society for the Protection of Cats practiced both tattooing and tipping the ear of the neutered cats to identify them.
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On January 29, 2019, the Hawaii Invasive Species Council adopted a resolution supporting the keeping of pet cats indoors and the use of peer-reviewed science in pursuing humane mitigation of the impacts of feral cats on wildlife and people.
695:, Ontario, includes TNR in its animal services and has a bylaw specifically addressing TNR and managed colonies. The Toronto Animal Services offers spay and neuter for colonies that are registered and have an assigned trained caretaker.
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In a 10-year study in Orange County, Florida, after a feral cat sterilization program was instituted in which 7,903 feral cats neutered, the cost was an estimated $ 442,568, as compared to $ 1,098,517 if they had been impounded and
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850:, a threatened species, was known to nest. Under a settlement and judicial order, the State of New York agreed to remove all of the cats to a shelter, and to trap and permanently remove any cats found in the park in the future.
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The Department of Conservation does not support TNR : "We don’t support the "trap, desex, release" approach to managing stray cats. This is because cats are predatory animals that continue to pose a threat to wildlife."
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In 1978, the city of Paris issued a Declaration of Rights of the Free-living Cat. In that year, Cambazard founded École du Chat and TNR'd its first cat, continuing to help thousands of cats in the following years.
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Natoli, Eugenia; Maragliano, Laura; Cariola, Giuseppe; Faini, Anna; Bonanni, Roberto; Cafazzo, Simona; Fantini, Claudio (2006-12-18). "Management of feral domestic cats in the urban environment of Rome (Italy)".
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Finkler, Hilit; Gunther, Idit; Terkel, Joseph (2011-05-01). "Behavioral differences between urban feeding groups of neutered and sexually intact free-roaming cats following a trap-neuter-return procedure".
578:. Contaminated soil is an important source of infection for humans, herbivores, rodents, and birds and several studies suggest that pet feces contribute to bacterial loading of streams and coastal waters.
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until an environmental review was completed under the California Environmental Quality Act. The judge did not rule on any environmental issues, or prohibit other organizations from doing TNR in the city.
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Managed TNR programs that involve continuous active intervention on detection along with treatment and prevention of some of the more common diseases and parasites may help improve their overall health.
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Alleviating unnecessary suffering of kittens: 75% of the kittens born to free-roaming cats being studied died or disappeared before 6 months of age. Trauma was found to be the most common cause of death.
846:(2014), conservationists sued Rose Harvey, the Commissioner of New York State Parks, under the Endangered Species Act for failing to prohibit activities supporting feral cats in a state park where the
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to consider criminal charges against those feeding the stray cats. No charges were laid, but the rescue group's business license was revoked and it was forced to move from its storefront location.
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practice, nor is ongoing monitoring and follow-up care once released after the neuter-spay. Vaccinations, medical examination, care, or treatments may or may not be a part of any given TNR effort.
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not the sole reason for success. The population reduction was primarily from adoption (47%) and euthanasia (11%), or due to the cats no longer living on site with their whereabouts unknown (15%).
138:–release') refers to a different method of cat population management, despite its similar name. TVHR differs in the type of sterilization surgery performed on the cats. Unlike traditional spays (
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The legal status of free-roaming and stray cats varies from location to location, as do the histories and efforts of TNR programs. There are numerous governments supporting trap–neuter–return.
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A study of a TNR program at Texas A&M University in 1998–2000 reported that the number of cat complaints received by the university's pest control service decreased from year 1 to year 2.
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Xuying Zhang, Kokila Jamwal, Ottmar Distl. "Tracking footprints of artificial and natural selection signatures in breeding and non-breeding cats." Scientific reports 12 (1), 18061, 2022.
260:(RSPCA): the RSPCA "supports Trap, Neuter and Release (TNR) programmes with veterinary support. Healthy cats should be neutered, ear-tipped and returned or, where appropriate, re-sited."
183:: PETA does not endorse TNR, citing research finding it is not an effective intervention along with data finding short life expectancies and inhumane living conditions for feral cats.
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Nogales, Manuel; Martín, Aurelio; Tershy, Bernie R.; Donlan, C. Josh; Veitch, Dick; Puerta, Néstor; Wood, Bill; Alonso, Jesús (2004). "A Review of Feral Cat Eradication on Islands".
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Butchart, Stuart H. M.; Micol, Thierry; Cranwell, Steve; Pasachnik, Stesha A.; Wanless, Ross; Fisher, Robert N.; Griffiths, Richard; Pierce, Ray; Borroto-Páez, Rafael (2019-03-27).
4567:"American Bird Conservancy, David A. Krauss, and Susan Scioli, Plaintiffs, v. Rose Harvey, Commissioner, New York Office of Parks, Recreation, And Historic Preservation, Defendant"
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Reeve, Charlie L.; Rogelberg, Steven G.; Spitzmüller, Christiane; Digiacomo, Natalie (2005). "The Caring-Killing Paradox: Euthanasia-Related Strain Among Animal-Shelter Workers1".
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below 2 kittens per mother, which can be extremely difficult given the high reproductive capacity of feral cats (with a queen being capable of weaning about 9.4 kittens/year).
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TNR as a method of managing free-roaming cat populations is controversial. Global attitudes towards these cats vary from those who see them as pets to those who target them as
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machines or other animals, predation by wildlife, toxins and poisons, contagious diseases, exposure to harsh weather, malnutrition, infections, and parasitic debilitation.
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campaigns where animals at risk of euthanasia are transported to areas where animals for adoptions are needed, or public awareness campaigns to enhance adoption rates.
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Judith L Stella, Candace C Croney. "Environmental aspects of domestic cat care and management: implications for cat welfare." The Scientific World Journal 2016, 2016.
120:('trap–neuter–release–maintain/manage') where 'maintain' generally means caregivers feed and monitor the feral cats after they are returned to their territories, and
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Montoya, A.; García, M.; Gálvez, R.; Checa, R.; Marino, V.; Sarquis, J.; Barrera, J. P.; Rupérez, C.; Caballero, L.; Chicharro, C.; Cruz, I.; Miró, G. (2018-02-15).
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After starting a TNR program in 1995, animal control in Orange County, Florida, received fewer complaints about cats, even after broadening the definition of a
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do not tolerate being caged or handled and many shelters are unable to manage them without putting the animal or the staff at risk. TNR could alleviate this.
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Governments have been sued to try to block their TNR efforts. In December, 2010, an injunction was granted to prevent a planned TNR program of the City of
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Control, World Health Organization Epidemic Disease; Tikhomirov, Evgueni; Poland, Jack D.; Gratz, Norman G.; Gage, Kenneth L.; Dennis, David T. (1999).
2312:"Decrease in Population and Increase in Welfare of Community Cats in a Twenty-Three Year Trap-Neuter-Return Program in Key Largo, FL: The ORCAT Program"
1411:"A renewed call for conservation leadership 10 years further in the feral cat Trap-Neuter-Return debate and new opportunities for constructive dialogue"
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Hughes, Kathy L.; Slater, Margaret R. (2002). "The Effects of Implementing a Feral Cat Spay/Neuter Program in a Florida County Animal Control Service".
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Thomas, J. E.; Staubus, L.; Goolsby, J. L.; Reichard, M. V. (2016-09-15). "Ectoparasites of free-roaming domestic cats in the central United States".
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The earliest documented practice of trap–neuter–return was in the 1950s, led by animal activist Ruth Plant in the UK. In the mid-1960s, former model
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4936:.Includes a wide range of solutions, including TNR, to surgical and non-surgical sterilization, and lethal methods; outlines a five-stage program.
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Schubach, Armando; Wanke, Bodo; Gremião, Isabella Dib; Coll, Jesana Ornellas; Schubach, Tania Pacheco; Barros, Monica Bastos de Lima (June 2010).
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evidence of subsequent native species recovery, it is gaining recognition as a recommended method of sustaining native biodiversity on islands.
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Feral or very fearful community cats have historically been euthanized either immediately or at the end of a holding period at most shelters.
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The first scientific conference on "the ecology and control of feral cats" was held in London in 1980 and its proceedings published by the
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3636:"Outdoor fecal deposition by free-roaming cats and attitudes of cat owners and nonowners toward stray pets, wildlife, and water pollution"
801:(UFAW). Subsequent UFAW publications in 1982, 1990, and 1995 were the primary scientific references for feral-cat control for many years.
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theoretical savings of $ 123,000 was projected based on not having to impound the offspring that the cats may have produced if not spayed.
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of euthanasia." Some of the cats that came in to the TNR colony had already been sterilized; some of these had ear-tips and some did not.
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1063:. Susan Richmond, Meredith Weiss, Anitra Frazier, joE. Needham, Lois McClurg & Laura Gay Senk (2nd ed.). Neighborhood Cats, Inc.
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663:, Threatened Species Commissioner Gregory Andrews reportedly summarized the reason for the federal government's intention to wipe out 2
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interactions involving intact males who had moved into them. As noted above, introgression of cats is a common factor noted in studies.
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2166:"Use of matrix population models to estimate the efficacy of euthanasia versus trap-neuter-return for management of free-roaming cats"
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Free-roaming domestic cats are considered an "alien" species and are listed as invasive in a multitude of countries around the world.
3699:
2728:"Public Opinions on Strategies for Managing Stray Cats and Predictors of Opposition to Trap-Neuter and Return in Brisbane, Australia"
1472:
526:
that can impact humans as well as other animals, domestic and wild. Transmissions can occur within the species and to other species.
4925:
Tens of millions of free-roaming felines take a huge toll on wildlife; what to do about them has spawned battles from coast to coast
4847:
834:
There is no federal law in the US that explicitly sets policy on feral cats, and state and local laws vary in their approaches. The
4024:
173:
management program (which includes community cat oversight and monitoring); and those who unconditionally support and endorse TNR.
4386:
1993:
4038:
2542:
235:
211:
1492:
3817:
277:
In general, trap-neuter-release programs are not effective at reducing the population of feral cats, as they do not affect the
4538:"Toolkit to Address Free-ranging Domestic Cats (Felis catus) on Agency Lands Managed for Native Wildlife and Ecosystem Health"
4140:
3787:
3011:
4940:
Guide to Trap-Neuter-Return (TNR) and Colony Care from Alley Cat Allies, the ASPCA and the Mayor's Alliance for NYC's Animals
4693:
3955:
2371:"Application of a Protocol Based on Trap-Neuter-Return (TNR) to Manage Unowned Urban Cats on an Australian University Campus"
4437:
3041:
146:(castration), which are done in TNR, the vasectomies and hysterectomies in TVHR result in sterile but sexually active cats.
2222:
1083:
493:
Numerous studies have shown that free-roaming cats can have a significant negative impact on native wildlife through their
1323:
4922:
3062:
Hughes, Kathy L.; Slater, Margaret R. (2002). "Implementation of a feral cat management program on a university campus".
337:
TNR is often presented to public officials and policymakers as a viable alternative to euthanasia with several benefits.
4353:
3927:
2952:
Elsheikha, Hany M.; Schnyder, Manuela; Traversa, Donato; Di Cesare, Angela; Wright, Ian; Lacher, David W. (2016-07-07).
2697:
2484:
2447:
1971:
1802:
4908:
4866:
4537:
1906:
1227:"Free-ranging domestic cat abundance and sterilization percentage following five years of a trap–neuter–return program"
884:
17:
3209:
4965:
Criticism of TNR, and advocacy of "a movement to overcome local and national challenges caused by free-roaming cats".
1512:
269:
Various studies and arguments have been presented both in support of and in opposition to free-roaming cats and TNR.
217:
4778:
4469:
2879:
1532:
196:: "TNR undermines the work of wildlife professionals and severely jeopardizes the integrity of native biodiversity."
3956:"Merritt, B.C., overrun by stray cats: Bylaw officers ask RCMP to consider mischief charges against animal feeders"
1746:
2048:"Evaluation of the effect of a long-term trap–neuter–return and adoption program on a free-roaming cat population"
1349:
1770:
1143:
2252:
1479:, Corporation of Delta, accessed August 6, 2014. Quote: "...our Trap–Neuter/Spay–Return program for feral cats".
4093:
1113:
915:"Better trap–neuter–return for free-roaming cats: Using models and monitoring to improve population management"
3634:
Dabritz, Haydee A.; Atwill, E. Robert; Gardner, Ian A.; Miller, Melissa A.; Conrad, Patricia A. (2006-07-01).
2509:
2047:
1005:
4969:
The Outdoor Cat: Science and Policy from a Global Perspective, December 3-4, 2012, Marina del Rey, California
4650:"Domestic cats and their impacts on biodiversity: A blind spot in the application of nature conservation law"
4407:
685:
531:
297:
288:
Some long-term studies have claimed or been cited to show that TNR is effective in stopping reproduction and
30:
4323:
4277:
1733:"What is PETA's stance on programs that advocate trapping, spaying and neutering, and releasing feral cats?"
4594:"New York State Parks, American Bird Conservancy Announce Feral Cats Relocated from Jones Beach State Park"
4522:
3978:
3188:
1580:"Estimation of effectiveness of three methods of feral cat population control by use of a simulation model"
805:
4939:
3334:"Globally important islands where eradicating invasive mammals will benefit highly threatened vertebrates"
1747:"IWRC Education and Resources for Wildlife Conservation Worldwide: Feral Cats and Dogs Position Statement"
1058:
413:
work-to-family conflict, somatic complaints, and substance use, and with lower levels of job satisfaction.
4123:
250:
2609:"The Effects of Implementing a Feral Cat Spay/Neuter Program in a Florida County Animal Control Service"
4900:
4001:
3113:
Dickman, Chris R.; Ritchie, Euan G.; Nimmo, Dale G.; Glen, Alistair S.; Doherty, Tim S. (2016-10-04).
1268:"Rabies Prevention and Management of Cats in the Context of Trap, Neuter, Vaccinate, Release Programs"
503:
Cats are now thought to be the single largest cause of anthropogenic bird mortality in North America.
4953:
4890:
Free Ranging and Feral Cats. US Department of Agriculture Wildlife Damage Management Technical Series
4566:
3536:
2586:
839:
681:
494:
488:
289:
199:
70:
2677:
1350:"Do trap-neuter-return (TNR) practices contribute to human–coyote conflicts in southern California?"
1181:
4622:
4593:
2107:"An Examination of an Iconic Trap-Neuter-Return Program: The Newburyport, Massachusetts Case Study"
220:: "These programs cannot be supported as a generalised and key strategy in the management of cats."
108:
are returned to their original locations under such a program. Variant acronyms and terms include:
4988:
4918:
An Examination of an Iconic Trap-Neuter-Return Program: The Newburyport, Massachusetts Case Study
1913:, ASPCA, second paragraph, accessed June 20, 2014. Resources provided here on TNR and feral cats.
4719:
4298:
1660:
914:
176:
Organizations that oppose TNR on the basis of animal welfare and wildlife preservation include:
1176:
1106:"Trap-Neuter-Return (TNR) Program For Community Cats | Humane Society of the Pikes Peak Region"
894:
835:
2543:"County of Santa Clara Animal Care & Control Places 100% of Healthy Dogs and Cats in 2011"
2008:
1689:
467:
attract un-neutered males with the attendant problems of spraying, fighting and caterwauling.
4973:
3750:
889:
527:
523:
422:
It has been claimed that TNR programs improve the welfare of free-roaming cats in many ways:
310:
of free-roaming cats, such as the one researchers established for use in urban environments.
282:
1923:
281:
of the environment. In order to be successful, a trap-neuter-release program must drive the
4945:
3471:"Implications of zoonotic and vector-borne parasites to free-roaming cats in central Spain"
3412:
3345:
3126:
2223:"A Generalized Population Monitoring Program to Inform the Management of Free-Roaming Cats"
1718:
1060:
Neighborhood Cats TNR Handbook: The Guide to Trap-Neuter-Return for the Feral Cat Caretaker
542:
193:
3691:
2552:. San Jose, California: Office of the County Executive, County of Santa Clara. p. 2.
1469:
8:
4993:
4842:
4145:
1550:"Trap-Vasectomy-Hysterectomy-Return (TVHR) Is No Substitute for Trap-Neuter-Return (TNR)"
4917:
3537:"Critical Assessment of Claims Regarding Management of Feral Cats by Trap–Neuter–Return"
3416:
3349:
3130:
1629:"Tufts researchers say TNR is not most effective method to control feral cat population"
939:
4671:
4408:"Managing Street Dogs and Cats in Turkey | Dog and Cat Behavior | Dog and Cat Behavior"
4378:
3673:
3567:
3443:
3400:
3376:
3333:
3314:
3157:
3114:
3095:
2988:
2953:
2798:
2762:
2727:
2644:
2405:
2370:
2346:
2311:
2203:
2141:
2106:
1989:
1609:
1432:
1292:
1248:
1202:
952:
4035:
979:"Stray Cats Boosting Rabies Risk in Brevard. 16 Rabid Cats Reported Statewide in 2018"
4904:
4675:
4244:
4226:
4218:
4089:
3665:
3657:
3616:
3559:
3555:
3492:
3448:
3430:
3399:
McGregor, Hugh; Legge, Sarah; Jones, Menna E.; Johnson, Christopher N. (2015-08-19).
3381:
3363:
3306:
3292:
3261:
3162:
3144:
3087:
3079:
2993:
2975:
2934:
2926:
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2846:
2802:
2767:
2749:
2636:
2628:
2410:
2392:
2351:
2333:
2277:
2195:
2187:
2146:
2128:
2072:
2028:
1640:
1601:
1489:
1436:
1297:
1252:
1194:
1025:
944:
654:
382:
356:
278:
85:
A cat displaying an ear tip, a common practice to identify cats that have been TNRed.
4214:
3809:
3775:
3677:
3318:
3099:
2648:
2207:
1613:
1206:
956:
81:
73:, and the unwillingness of both scientific communities and TNR advocates to engage.
4661:
4210:
3725:
3647:
3608:
3551:
3482:
3438:
3420:
3371:
3353:
3296:
3288:
3251:
3152:
3134:
3071:
2983:
2965:
2918:
2836:
2794:
2757:
2739:
2620:
2608:
2400:
2382:
2341:
2323:
2269:
2177:
2136:
2118:
2064:
2020:
1591:
1422:
1361:
1287:
1279:
1238:
1186:
1017:
1006:"Review and synthesis of the global literature on domestic cat impacts on wildlife"
934:
926:
809:
510:
Free-roaming cats have been documented hunting and killing prey without eating it.
165:
2954:"Updates on feline aelurostrongylosis and research priorities for the next decade"
4870:
4851:
4127:
4042:
4031:
4008:
3985:
3962:
3744:"QUESTIONS AND ANSWERS ON AVIAN INFLUENZA In relation to animals, food and water"
3612:
3571:
3487:
3470:
3425:
3358:
3033:
2684:
2624:
1910:
1536:
1516:
1496:
1476:
1457:
1190:
534:, ectoparasites (fleas, mites, lice, ticks), intestinal and protozoan parasites,
352:
205:
3401:"Feral Cats Are Better Killers in Open Habitats, Revealed by Animal-Borne Video"
3075:
2841:
2824:
2273:
2182:
2165:
1075:
2024:
2970:
2825:"Reproductive capacity of free-roaming domestic cats and kitten survival rate"
2068:
1383:
4982:
4299:"South Korea's Animal Protection Laws | International Aid for Korean Animals"
4222:
3661:
3434:
3367:
3310:
3148:
3083:
2979:
2930:
2922:
2850:
2806:
2753:
2744:
2632:
2476:
2396:
2337:
2328:
2191:
2132:
2032:
1794:
1578:
McCarthy, Robert J.; Levine, Stephen H.; Reed, J. Michael (August 15, 2013).
930:
847:
790:
594:
326:
170:
4863:
3535:
Longcore, Travis; Rich, Catherine; Sullivan, Lauren M.; et al. (2009).
3139:
1903:
1596:
1579:
1021:
4230:
4086:
TNR: Past, Present and Future: A History of the Trap-Neuter-Return Movement
3669:
3652:
3635:
3620:
3563:
3496:
3452:
3385:
3265:
3240:"The impact of free-ranging domestic cats on wildlife of the United States"
3166:
3091:
2997:
2938:
2858:
2771:
2640:
2414:
2355:
2281:
2199:
2150:
2076:
1605:
1301:
1198:
1029:
948:
735:
633:
Domestic cats can be found on every continent except Antarctica. Of the 700
135:
2541:
Mitchell, Gwendolyn; Anderson, Laurel; Wasson, Brigid (January 25, 2012).
1509:
581:
Free-roaming cat populations have been identified as a source for several
242:
million. In contrast, the number of pet cats in the US is approximately 76
4746:"Resolution 19-2: Keeping Cats Indoors & Using Peer-Reviewed Science"
4720:"Electric Utility Sentenced for Killing Eagles and Hawks (Press Release)"
4461:
1824:"A review of trap-neuter-return (TNR) for the management of unowned cats"
1529:
871:
1267:
4666:
4649:
4245:"Stray dog and cat laws and enforcement in Czech Republic and in Italy"
3729:
3301:
3256:
3239:
2823:
Nutter, Felicia B.; Levine, Jay F.; Stoskopf, Michael K. (2004-11-01).
2387:
2123:
1451:"Denver Animal Shelter partners up to help reduce feral cat population"
1427:
1410:
1384:"How Effective and Humane Is Trap-Neuter-Release (TNR) for Feral Cats?"
1365:
808:
began utilizing TNR of feral cats to protect the regionally endangered
612:
547:
are examples of inter and intra-species shared diseases and parasites.
536:
4600:. New York State Office of Parks, Recreation and Historic Preservation
1283:
1243:
1226:
1135:
4438:"Stray Dog and Cat Feral Populations in Europe: from Culling to Care"
825:
Establishing buffer zones to prevent feral cats returning to the area
443:
378:
360:
143:
139:
131:
105:
54:
50:
1508:
Society for the Prevention of Cruelty to Animals, Montreal, Quebec,
1105:
550:
There are numerous zoonotic pathogens shed in feline feces, such as
3722:
Plague manual: epidemiology, distribution, surveillance and control
2164:
Andersen, Mark C.; Martin, Brent J.; Roemer, Gary W. (2004-12-15).
2046:
Levy, Julie K.; Gale, David W.; Gale, Leslie A. (January 1, 2003).
582:
472:
49:, is a controversial method that attempts to manage populations of
4933:
3012:"Quick release after surgery results in gruesome feral cat deaths"
2951:
2784:
2432:. Michigan State University Animal Legal & Historical Center.
2310:
Levy, Julie K.; Cornell, Heather N.; Kreisler, Rachael E. (2019).
442:
Not all free-roaming cats, or cats that end up in live traps, are
351:"feral" after being taken to a traditional shelter not practicing
4273:
2726:
Hayward, Andrea; Lamb, Kate; Fisher, Gina; Rand, Jacquie (2019).
2607:
Hughes, Kathy L.; Slater, Margaret R.; Haller, Linda (Oct 2002).
704:
692:
58:
4518:
3899:"Australian Feral Cats Eat More Than a Million Reptiles Per Day"
3180:
2009:"Reproductive performance in a cat colony over a 10-year period"
4864:"Get the Facts about the Los Angeles Trap–Neuter–Return Ruling"
1530:"NPA's TTVAR Program: Trap, Test, Vaccinate, Alter and Release"
589:
272:
4623:"United States v. Moon Lake Electric Association Incorporated"
4120:
3331:
3238:
Loss, Scott R.; Will, Tom; Marra, Peter P. (29 January 2013).
2698:"New Survey Reveals Widespread Support for Trap-Neuter-Return"
1690:"The Global Impact of Feral Cats | The Ecology Global Network"
1409:
Debrot, Adolphe O.; Ruijter, Martin N. M. (17 January 2022).
746:
745:
Since August 1991, feral cats have been protected throughout
4694:"Detailed Discussion of Feral Cat and Wild Bird Controversy"
4199:
4109:"Declaration of Rights of the Free-living Cat (translation)"
3719:
3598:
842:
may be relevant to the legality of TNR. In a federal case,
739:
224:
Some organizations that conditionally support TNR include:
180:
53:. The process involves live-trapping the cats, having them
3633:
2578:
2251:
Levy, Julie K.; Crawford, Patti Cynda (November 1, 2004).
1490:"Trap / Neuter / Vaccinate / Return (TNVR): What is TNVR?"
1348:
Bucklin, Danielle M.; Shedden, Jennifer M. (Spring 2023).
4897:
Cat Wars: The Devastating Consequences of a Cuddly Killer
3776:"Esporotricose: a evolução e os desafios de uma epidemia"
3398:
3278:
2687:, Ray Weiss, The Daytona Beach News-Journal, May 7, 2013.
2253:"Humane strategies for controlling feral cat populations"
417:
230:
American Society for the Prevention of Cruelty to Animals
4141:"Jerusalem Dilemma in Dealing with Hordes of Stray Cats"
3773:
3468:
4648:
Trouwborst, Arie; McCormack, Phillipa (February 2020).
4496:. Washington DC: Humane Society Press. pp. 43–53.
4462:"Celia Hammond Animal Trust - UK Animal Rescue Charity"
3112:
4818:. Alaska Department of Fish and Game. pp. 72–76.
4351:
3640:
Journal of the American Veterinary Medical Association
2911:
Journal of the American Veterinary Medical Association
2829:
Journal of the American Veterinary Medical Association
2673:
2671:
2669:
2540:
2261:
Journal of the American Veterinary Medical Association
2170:
Journal of the American Veterinary Medical Association
2056:
Journal of the American Veterinary Medical Association
1584:
Journal of the American Veterinary Medical Association
768:
techniques and adhere to the Animal Welfare Act 1999.
519:
major risk for the transmission of zoonotic diseases.
258:
Royal Society for the Prevention of Cruelty to Animals
3869:"War on feral cats: Australia aims to cull 2 million"
2725:
1225:
Coe, Seraiah T; Elmore, Jared A. (23 December 2021).
599:
Various nematode parasites including intestinal worms
398:
3534:
2907:
2822:
2503:
2501:
2163:
867:
proposals to exempt sterilized cats from the rules.
4934:
International Companion Animal Management Coalition
4647:
4172:"Jerusalem Approves Feeding Program for Stray Cats"
4056:"Trap, Neuter, Return Program (TNR) for Feral Cats"
2666:
2309:
513:
482:
305:
very little statistical data to support the claims.
169:those who conditionally support TNR as a part of a
57:, ear-tipped for identification, and, if possible,
4931:ICAM - Humane Cat Population Management Guidelines
2606:
1577:
314:high-value adjunct to ongoing management efforts.
4529:
4025:"The Trap, Neuter, Return Program for Feral Cats"
3115:"Invasive predators and global biodiversity loss"
2498:
1972:"Animal Welfare Policy & Position Statements"
1719:"Trap-Neuter-Release: Does TNR Really Save Cats?"
1082:. Silverdale, Washington: Kitsap Humane Society.
27:Strategy for controlling feral animal populations
4980:
4844:Urban Wildlands Group v. The City of Los Angeles
4687:
4685:
4034:, City of Toronto, accessed August 3, 2014; and
2105:Spehar, Daniel D.; Wolf, Peter J. (2017-10-31).
1266:Roebling, Allison D; Johnson, Dana (June 2014).
822:Using humane, neutering-based feral cat controls
3119:Proceedings of the National Academy of Sciences
2534:
1853:"Free-roaming owned, Abandoned, and Feral Cats"
1771:"Effects of an Invasive Species: Domestic Cats"
1265:
264:
4854:, Court of Appeals of California, No. B222696.
4490:Slater, Margaret R.; Shain, Stephanie (2005).
4036:"Toronto Municipal Code, Chapter 349, Animals"
2510:"Feline Shelter Intake Reduction Program FAQs"
2369:Swarbrick, Helen; Rand, Jacquie (2018-05-17).
1347:
585:that can and have affected humans, including:
405:way of controlling the population of animals."
340:
4974:A Beginners Guide to TNR (Trap-Neuter-Return)
4946:"Information on Managed Cat Colonies and TNR"
4894:
4682:
4641:
4617:
4615:
4547:. Association of Fish & WIldlife Agencies
4435:
4274:"Cats | International Aid for Korean Animals"
3979:"City's approach to feral cats not effective"
3528:
3237:
3034:"Neutering your cat | International Cat Care"
2368:
1408:
1324:"How the "No Kill" Movement Betrays its Name"
1162:
1160:
999:
997:
995:
296:An eleven-year study of a TNR program at the
188:International Wildlife Rehabilitation Council
4877:, Alley Cat Allies, accessed August 2, 2014.
4535:
3061:
2250:
2246:
2244:
2242:
2045:
1377:
1375:
1166:
1003:
816:Saving the genetically pure Scottish wildcat
273:Purported reductions in population over time
4489:
2220:
2006:
1961:Mortality Weekly Report 69 (23), 710, 2020.
1645:: CS1 maint: numeric names: authors list (
1402:
1381:
734:Killing feral cats has been illegal in the
71:the environmental harm caused by feral cats
4779:"Alaska has far too many impractical cats"
4700:. Michigan State University College of Law
4629:. Michigan State University College of Law
4612:
4436:Furler-Mihali, Andrea Irina (April 2017).
3692:"CAPC Parasite ProtocolsFeline Arthropods"
2508:Hurley, Kate; Levy, Julie (January 2013).
2507:
2427:
2104:
1220:
1218:
1216:
1157:
992:
799:Universities Federation for Animal Welfare
159:
4895:Marra, Peter P.; Santella, Chris (2016).
4665:
3651:
3486:
3442:
3424:
3375:
3357:
3300:
3255:
3231:
3156:
3138:
3064:Journal of Applied Animal Welfare Science
2987:
2969:
2840:
2761:
2743:
2678:"Port Orange expanding feral cat program"
2613:Journal of Applied Animal Welfare Science
2404:
2386:
2345:
2327:
2239:
2181:
2140:
2122:
2039:
1661:"Million Cat Challenge | Return to Field"
1595:
1426:
1372:
1291:
1242:
1224:
1180:
1169:Journal of Applied Animal Welfare Science
1004:Loss, Scott R.; Boughton, Brooke (2022).
938:
624:Fungal diseases, including sporotrichosis
4130:, École du Chat, accessed Oct. 30, 2014.
4121:"History of École du chat (translation)"
4111:, École du Chat, accessed Oct. 30, 2014.
4083:
4002:"Merritt storefront clear of stray cats"
2430:"Animal Euthanasia: Detailed Discussion"
2007:Robinson, Roy; Cox, H. W. (1970-04-01).
359:(humanely putting the animal to death).
80:
29:
4776:
4591:
4536:Schweitzer, Sarah H.; Gillin, Colin M.
3464:
3462:
3210:"issg Database: Ecology of Felis catus"
2880:"Free-roaming Abandoned and Feral Cats"
2481:The Humane Society of the United States
1928:The Humane Society of the United States
1857:Canadian Veterinary Medical Association
1321:
1213:
819:Removing all feral cats from the region
628:
236:The Humane Society of the United States
212:Canadian Veterinary Medical Association
124:('trap–test–vaccinate–alter–release').
14:
4981:
4805:
4803:
4772:
4770:
2221:Boone, J.D.; Slater, M.A. (Nov 2014).
1519:, July 5, 2013, accessed June 15, 2014
1382:Hostetler, Mark; Wisely, Samantha M..
1322:Franzen, Jonathan (25 December 2023).
1056:
976:
919:Journal of Feline Medicine and Surgery
418:Improving the cats' health and welfare
3928:"The problem with cats - Macleans.ca"
3866:
2874:
2872:
2870:
2868:
2818:
2816:
2583:SPCA of Martinsville and Henry County
2579:"FAQs About Trap/Neuter/Return (TNR)"
1626:
1510:"Do you know about our TNRM program?"
912:
4698:Animal Legal & Historical Center
4627:Animal Legal & Historical Center
4571:Animal Legal & Historical Center
3511:"Gastrointestinal Parasites of Cats"
3459:
2787:Journal of Applied Social Psychology
2291:from the original on August 19, 2014
1052:
1050:
1048:
1046:
680:In January 2012, a bylaw officer in
332:
4800:
4767:
4691:
4400:
3807:
1528:for example, National Pet Alliance
844:American Bird Conservancy v. Harvey
461:
377:Proponents of TNR claim that while
24:
4883:
4379:"Turkey | World Animal Protection"
4088:. Alley Cat Allies. pp. 2–3.
2865:
2813:
2799:10.1111/j.1559-1816.2005.tb02096.x
2559:from the original on July 23, 2014
2428:Kleinfeldt, Alexandra (Sep 2017).
2086:from the original on July 18, 2014
1990:"RSPCA policies on animal welfare"
1470:"How to Help: Financial Donations"
885:Overpopulation in domestic animals
566:spp); and the protozoan parasites
399:Improved morale and public support
116:('trap–neuter–vaccinate–return'),
25:
5010:
4045:, City of Toronto, June 13, 2013.
3896:
1883:Australian Veterinary Association
1879:"Management of cats in Australia"
1415:Conservation Science and Practice
1086:from the original on July 9, 2014
1043:
913:Boone, John D. (25 August 2015).
784:
684:, removed cat food and asked the
290:reducing the population over time
218:Australian Veterinary Association
112:(for 'trap–neuter/spay–return'),
34:Cat caught in a live-trap for TNR
4999:Trap–neuter–return organizations
4857:
4836:
4812:Permits for Possessing Live Game
4738:
4712:
4585:
4559:
4525:from the original on 2017-01-10.
4511:
4483:
4454:
4450:from the original on 2019-03-21.
4429:
4371:
4345:
4316:
4291:
4266:
3556:10.1111/j.1523-1739.2009.01174.x
3293:10.1111/j.1523-1739.2004.00442.x
2436:from the original on 2020-09-01.
2235:from the original on 2016-03-14.
1996:from the original on 2018-08-18.
1978:from the original on 2017-10-03.
1783:from the original on 2018-05-17.
1759:from the original on 2015-02-22.
1460:, The Denver Post, June 12, 2014
829:
514:Risks to human and animal health
483:Effects on wildlife from hunting
4825:from the original on 2018-04-15
4789:from the original on 2020-06-26
4756:from the original on 2019-03-21
4750:Hawaii Invasive Species Council
4500:from the original on 2020-09-01
4472:from the original on 2019-03-26
4418:from the original on 2019-03-21
4389:from the original on 2019-10-30
4334:from the original on 2019-03-21
4305:from the original on 2019-04-04
4280:from the original on 2019-04-04
4255:from the original on 2020-09-01
4237:
4215:10.1016/j.prevetmed.2006.06.005
4193:
4182:from the original on 2019-12-05
4164:
4153:from the original on 2019-11-20
4133:
4114:
4102:
4077:
4066:from the original on 2019-04-02
4048:
4018:
3995:
3972:
3949:
3938:from the original on 2019-04-02
3920:
3909:from the original on 2019-03-21
3890:
3879:from the original on 2019-03-21
3860:
3851:
3841:
3831:
3820:from the original on 2019-03-31
3801:
3790:from the original on 2019-03-30
3767:
3736:
3713:
3702:from the original on 2019-03-30
3684:
3627:
3592:
3588:– via UrbanWildlands.org.
3581:from the original on 2013-09-08
3517:from the original on 2019-03-30
3503:
3392:
3325:
3272:
3220:from the original on 2019-03-30
3202:
3191:from the original on 2019-03-29
3173:
3106:
3055:
3044:from the original on 2019-03-23
3026:
3004:
2945:
2901:
2890:from the original on 2019-03-25
2778:
2732:Frontiers in Veterinary Science
2719:
2708:from the original on 2019-03-23
2690:
2655:from the original on 2020-09-01
2600:
2589:from the original on 2019-03-23
2571:
2520:from the original on 2018-07-24
2487:from the original on 2019-03-23
2469:
2458:from the original on 2019-03-23
2440:
2421:
2362:
2316:Frontiers in Veterinary Science
2303:
2214:
2157:
2098:
2000:
1982:
1964:
1954:
1945:
1934:from the original on 2019-11-21
1916:
1897:
1871:
1845:
1816:
1805:from the original on 2019-03-22
1787:
1763:
1739:
1725:
1711:
1700:from the original on 2019-06-11
1682:
1671:from the original on 2019-03-30
1653:
1620:
1571:
1560:from the original on 2020-09-01
1542:
1522:
1502:
1482:
1463:
1443:
1341:
1315:
1259:
1146:from the original on 2019-03-23
1133:
1116:from the original on 2019-03-23
977:Waymer, Jim (8 February 2019).
371:
96:. It is sometimes described as
4203:Preventive Veterinary Medicine
4084:Berkeley, Ellen Perry (2004).
1627:Staff, DVM360.com (Sep 2013).
1127:
1098:
1067:
970:
906:
762:
753:
659:In a 2017 news release in the
637:million cats in the world, 480
76:
13:
1:
4383:api.worldanimalprotection.org
3808:Toenjes, Nicolle Birch Anna.
1488:Humane Society of Tampa Bay,
1449:for example, Joe Vaccarelli,
900:
686:Royal Canadian Mounted Police
532:feline immunodeficiency virus
522:Free-roaming cats can act as
298:University of Central Florida
4777:Sinnott, Rick (2017-11-06).
4726:. U.S. Department of Justice
4412:animalbehaviorassociates.com
4352:Department of Conservation.
3810:"Felis catus (domestic cat)"
3613:10.1016/j.vetpar.2016.07.034
3488:10.1016/j.vetpar.2018.01.009
3426:10.1371/journal.pone.0133915
3359:10.1371/journal.pone.0212128
2625:10.1207/s15327604jaws0504_03
1191:10.1207/S15327604JAWS0504_03
1140:Noah's Ark Animal Foundation
806:Scottish Wildcat Association
794:neutering and not killing".
648:
621:Avian Influenza A/H5N1 virus
265:Advantages and disadvantages
7:
4598:parks.ny.gov Press Releases
4592:Simons, Randy; Keefe, Dan.
4573:. Michigan State University
3867:Power, Julie (2017-02-17).
3696:Today's Veterinary Practice
3076:10.1207/S15327604JAWS0501_2
2842:10.2460/javma.2004.225.1399
2274:10.2460/javma.2004.225.1354
2183:10.2460/javma.2004.225.1871
1665:www.millioncatchallenge.org
1354:Human–Wildlife Interactions
878:
611:Bacterial diseases such as
341:Reducing euthanasia numbers
251:American Humane Association
10:
5015:
4901:Princeton University Press
4358:Department of Conservation
2025:10.1258/002367770781036616
698:
652:
486:
4954:American Bird Conservancy
3988:, Joni Hughes, letter to
3873:The Sydney Morning Herald
2971:10.1186/s13071-016-1671-6
2069:10.2460/javma.2003.222.42
1799:American Bird Conservancy
1539:, accessed June 15, 2014.
1499:, accessed June 15, 2014.
1010:Journal of Animal Ecology
840:Migratory Bird Treaty Act
775:
720:
711:
682:Merritt, British Columbia
671:
489:Cat predation on wildlife
200:American Bird Conservancy
2923:10.2460/javma.238.9.1141
2745:10.3389/fvets.2018.00290
2329:10.3389/fvets.2019.00007
931:10.1177/1098612X15594995
729:
4493:Feral Cats: An Overview
3780:Rev Panam Salud Publica
3601:Veterinary Parasitology
3475:Veterinary Parasitology
3140:10.1073/pnas.1602480113
2958:Parasites & Vectors
1597:10.2460/javma.243.4.502
1390:. University of Florida
1022:10.1111/1365-2656.13745
812:. Their goals include:
160:Advocacy and opposition
152:('return to field') or
3653:10.2460/javma.229.1.74
1924:"Our position on cats"
1694:Ecology Global Network
1272:Zoonoses Public Health
1057:Kortis, Bryan (2013).
895:Wildlife contraceptive
836:Endangered Species Act
86:
35:
4519:"Wildcat Haven C.I.C"
3244:Nature Communications
3014:. WFLA News Channel 8
890:Spaying and neutering
703:TNR was practiced in
661:Sydney Morning Herald
653:Further information:
558:spp; ascarids (e.g.,
528:Feline leukemia virus
283:net reproduction rate
104:emphasizes that most
84:
33:
4783:Anchorage Daily News
4466:www.celiahammond.org
3814:Animal Diversity Web
3544:Conservation Biology
3281:Conservation Biology
629:Treatment by country
524:vectors for diseases
381:cats may be costly,
194:The Wildlife Society
4923:The Cat Conundrum.
4146:The Times of Israel
4011:, Phillip Woolgar,
3992:, January 12, 2012.
3969:, January 11, 2012.
3417:2015PLoSO..1033915M
3350:2019PLoSO..1412128H
3131:2016PNAS..11311261D
3125:(40): 11261–11265.
1721:. 10 February 2021.
1136:"Feral Cat Program"
613:cat-scratch fever (
98:trap–neuter–release
92:usually stands for
47:trap–neuter–release
4869:2014-08-12 at the
4850:2014-08-10 at the
4692:Sanchez, Ariahna.
4667:10.1002/pan3.10073
4126:2014-10-13 at the
4041:2014-07-15 at the
4030:2014-08-08 at the
4007:2014-08-08 at the
3984:2014-08-08 at the
3961:2014-08-08 at the
3257:10.1038/ncomms3961
2683:2014-10-06 at the
2388:10.3390/ani8050077
2124:10.3390/ani7110081
2013:Laboratory Animals
1909:2014-07-02 at the
1535:2014-09-02 at the
1515:2014-10-06 at the
1495:2014-10-05 at the
1475:2014-10-06 at the
1456:2015-05-04 at the
1428:10.1111/csp2.12641
1366:10.26077/b86e-600f
1076:"Barn Cat Program"
353:no-kill sheltering
140:ovariohysterectomy
94:trap–neuter–return
87:
39:Trap–neuter–return
36:
18:Trap-Neuter-Return
4654:People and Nature
4015:, August 7, 2012.
2516:. Maddie's Fund.
2176:(12): 1871–1876.
1833:. RSPCA Australia
1284:10.1111/zph.12070
1244:10.2981/wlb.00799
1080:Kitsap-Humane.org
738:, which includes
655:Cats in Australia
583:zoonotic diseases
333:Euthanasia debate
279:carrying capacity
45:), also known as
16:(Redirected from
5006:
4964:
4962:
4960:
4914:
4878:
4861:
4855:
4840:
4834:
4833:
4831:
4830:
4824:
4817:
4807:
4798:
4797:
4795:
4794:
4774:
4765:
4764:
4762:
4761:
4742:
4736:
4735:
4733:
4731:
4716:
4710:
4709:
4707:
4705:
4689:
4680:
4679:
4669:
4645:
4639:
4638:
4636:
4634:
4619:
4610:
4609:
4607:
4605:
4589:
4583:
4582:
4580:
4578:
4563:
4557:
4556:
4554:
4552:
4545:FishWildlife.Org
4542:
4533:
4527:
4526:
4515:
4509:
4508:
4506:
4505:
4487:
4481:
4480:
4478:
4477:
4458:
4452:
4451:
4449:
4442:
4433:
4427:
4426:
4424:
4423:
4404:
4398:
4397:
4395:
4394:
4375:
4369:
4368:
4366:
4364:
4349:
4343:
4342:
4340:
4339:
4320:
4314:
4313:
4311:
4310:
4295:
4289:
4288:
4286:
4285:
4270:
4264:
4263:
4261:
4260:
4241:
4235:
4234:
4209:(3–4): 180–185.
4197:
4191:
4190:
4188:
4187:
4168:
4162:
4161:
4159:
4158:
4137:
4131:
4118:
4112:
4106:
4100:
4099:
4081:
4075:
4074:
4072:
4071:
4052:
4046:
4022:
4016:
3999:
3993:
3976:
3970:
3953:
3947:
3946:
3944:
3943:
3924:
3918:
3917:
3915:
3914:
3894:
3888:
3887:
3885:
3884:
3864:
3858:
3855:
3849:
3845:
3839:
3835:
3829:
3828:
3826:
3825:
3805:
3799:
3798:
3796:
3795:
3771:
3765:
3764:
3762:
3761:
3755:
3749:. Archived from
3748:
3740:
3734:
3733:
3717:
3711:
3710:
3708:
3707:
3688:
3682:
3681:
3655:
3631:
3625:
3624:
3596:
3590:
3589:
3587:
3586:
3580:
3541:
3532:
3526:
3525:
3523:
3522:
3507:
3501:
3500:
3490:
3466:
3457:
3456:
3446:
3428:
3396:
3390:
3389:
3379:
3361:
3329:
3323:
3322:
3304:
3276:
3270:
3269:
3259:
3235:
3229:
3228:
3226:
3225:
3206:
3200:
3199:
3197:
3196:
3185:www.iucngisd.org
3177:
3171:
3170:
3160:
3142:
3110:
3104:
3103:
3059:
3053:
3052:
3050:
3049:
3030:
3024:
3023:
3021:
3019:
3008:
3002:
3001:
2991:
2973:
2949:
2943:
2942:
2917:(9): 1141–1149.
2905:
2899:
2898:
2896:
2895:
2876:
2863:
2862:
2844:
2835:(9): 1399–1402.
2820:
2811:
2810:
2782:
2776:
2775:
2765:
2747:
2723:
2717:
2716:
2714:
2713:
2694:
2688:
2675:
2664:
2663:
2661:
2660:
2604:
2598:
2597:
2595:
2594:
2575:
2569:
2568:
2566:
2564:
2558:
2547:
2538:
2532:
2531:
2526:
2525:
2505:
2496:
2495:
2493:
2492:
2473:
2467:
2466:
2464:
2463:
2448:"Pet Statistics"
2444:
2438:
2437:
2425:
2419:
2418:
2408:
2390:
2366:
2360:
2359:
2349:
2331:
2307:
2301:
2300:
2298:
2296:
2290:
2257:
2248:
2237:
2236:
2234:
2227:
2218:
2212:
2211:
2185:
2161:
2155:
2154:
2144:
2126:
2102:
2096:
2095:
2093:
2091:
2085:
2052:
2043:
2037:
2036:
2004:
1998:
1997:
1986:
1980:
1979:
1968:
1962:
1958:
1952:
1949:
1943:
1942:
1940:
1939:
1920:
1914:
1904:"Feral Cats FAQ"
1901:
1895:
1894:
1892:
1890:
1875:
1869:
1868:
1866:
1864:
1849:
1843:
1842:
1840:
1838:
1828:
1820:
1814:
1813:
1811:
1810:
1791:
1785:
1784:
1782:
1775:
1767:
1761:
1760:
1758:
1751:
1743:
1737:
1736:
1729:
1723:
1722:
1715:
1709:
1708:
1706:
1705:
1686:
1680:
1679:
1677:
1676:
1657:
1651:
1650:
1644:
1636:
1631:. Archived from
1624:
1618:
1617:
1599:
1575:
1569:
1568:
1566:
1565:
1554:Alley Cat Allies
1546:
1540:
1526:
1520:
1506:
1500:
1486:
1480:
1467:
1461:
1447:
1441:
1440:
1430:
1406:
1400:
1399:
1397:
1395:
1379:
1370:
1369:
1345:
1339:
1338:
1336:
1334:
1319:
1313:
1312:
1310:
1308:
1295:
1263:
1257:
1256:
1246:
1231:Wildlife Biology
1222:
1211:
1210:
1184:
1164:
1155:
1154:
1152:
1151:
1131:
1125:
1124:
1122:
1121:
1102:
1096:
1095:
1093:
1091:
1071:
1065:
1064:
1054:
1041:
1040:
1038:
1036:
1016:(7): 1361–1372.
1001:
990:
989:
987:
985:
974:
968:
967:
965:
963:
942:
910:
857:
810:Scottish wildcat
666:
640:
636:
462:Fewer complaints
321:
245:
241:
166:invasive species
21:
5014:
5013:
5009:
5008:
5007:
5005:
5004:
5003:
4979:
4978:
4958:
4956:
4944:
4911:
4886:
4884:Further reading
4881:
4871:Wayback Machine
4862:
4858:
4852:Wayback Machine
4841:
4837:
4828:
4826:
4822:
4815:
4809:
4808:
4801:
4792:
4790:
4775:
4768:
4759:
4757:
4744:
4743:
4739:
4729:
4727:
4718:
4717:
4713:
4703:
4701:
4690:
4683:
4646:
4642:
4632:
4630:
4621:
4620:
4613:
4603:
4601:
4590:
4586:
4576:
4574:
4565:
4564:
4560:
4550:
4548:
4540:
4534:
4530:
4517:
4516:
4512:
4503:
4501:
4488:
4484:
4475:
4473:
4460:
4459:
4455:
4447:
4440:
4434:
4430:
4421:
4419:
4406:
4405:
4401:
4392:
4390:
4377:
4376:
4372:
4362:
4360:
4350:
4346:
4337:
4335:
4328:www.doc.govt.nz
4322:
4321:
4317:
4308:
4306:
4297:
4296:
4292:
4283:
4281:
4272:
4271:
4267:
4258:
4256:
4243:
4242:
4238:
4198:
4194:
4185:
4183:
4170:
4169:
4165:
4156:
4154:
4139:
4138:
4134:
4128:Wayback Machine
4119:
4115:
4107:
4103:
4096:
4082:
4078:
4069:
4067:
4060:City of Toronto
4054:
4053:
4049:
4043:Wayback Machine
4032:Wayback Machine
4023:
4019:
4009:Wayback Machine
4000:
3996:
3986:Wayback Machine
3977:
3973:
3963:Wayback Machine
3954:
3950:
3941:
3939:
3932:www.macleans.ca
3926:
3925:
3921:
3912:
3910:
3895:
3891:
3882:
3880:
3865:
3861:
3856:
3852:
3846:
3842:
3836:
3832:
3823:
3821:
3806:
3802:
3793:
3791:
3772:
3768:
3759:
3757:
3753:
3746:
3742:
3741:
3737:
3718:
3714:
3705:
3703:
3690:
3689:
3685:
3632:
3628:
3597:
3593:
3584:
3582:
3578:
3539:
3533:
3529:
3520:
3518:
3509:
3508:
3504:
3467:
3460:
3411:(8): e0133915.
3397:
3393:
3344:(3): e0212128.
3330:
3326:
3277:
3273:
3236:
3232:
3223:
3221:
3208:
3207:
3203:
3194:
3192:
3179:
3178:
3174:
3111:
3107:
3060:
3056:
3047:
3045:
3032:
3031:
3027:
3017:
3015:
3010:
3009:
3005:
2950:
2946:
2906:
2902:
2893:
2891:
2878:
2877:
2866:
2821:
2814:
2783:
2779:
2724:
2720:
2711:
2709:
2696:
2695:
2691:
2685:Wayback Machine
2676:
2667:
2658:
2656:
2605:
2601:
2592:
2590:
2577:
2576:
2572:
2562:
2560:
2556:
2545:
2539:
2535:
2523:
2521:
2514:MaddiesFund.org
2506:
2499:
2490:
2488:
2475:
2474:
2470:
2461:
2459:
2446:
2445:
2441:
2426:
2422:
2367:
2363:
2308:
2304:
2294:
2292:
2288:
2255:
2249:
2240:
2232:
2225:
2219:
2215:
2162:
2158:
2103:
2099:
2089:
2087:
2083:
2050:
2044:
2040:
2005:
2001:
1988:
1987:
1983:
1970:
1969:
1965:
1959:
1955:
1950:
1946:
1937:
1935:
1922:
1921:
1917:
1911:Wayback Machine
1902:
1898:
1888:
1886:
1877:
1876:
1872:
1862:
1860:
1851:
1850:
1846:
1836:
1834:
1831:RSPCA Australia
1826:
1822:
1821:
1817:
1808:
1806:
1793:
1792:
1788:
1780:
1773:
1769:
1768:
1764:
1756:
1749:
1745:
1744:
1740:
1731:
1730:
1726:
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1659:
1658:
1654:
1638:
1637:
1625:
1621:
1576:
1572:
1563:
1561:
1548:
1547:
1543:
1537:Wayback Machine
1527:
1523:
1517:Wayback Machine
1507:
1503:
1497:Wayback Machine
1487:
1483:
1477:Wayback Machine
1468:
1464:
1458:Wayback Machine
1448:
1444:
1407:
1403:
1393:
1391:
1380:
1373:
1346:
1342:
1332:
1330:
1320:
1316:
1306:
1304:
1264:
1260:
1223:
1214:
1182:10.1.1.566.9583
1165:
1158:
1149:
1147:
1132:
1128:
1119:
1117:
1104:
1103:
1099:
1089:
1087:
1074:
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1068:
1055:
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1002:
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981:
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765:
756:
732:
723:
714:
701:
674:
664:
657:
651:
638:
634:
631:
568:Cryptosporidium
516:
491:
485:
464:
427:kittens/litter.
420:
401:
374:
343:
335:
319:
275:
267:
243:
239:
206:RSPCA Australia
162:
79:
28:
23:
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15:
12:
11:
5:
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5002:
5001:
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4991:
4989:Animal welfare
4977:
4976:
4971:
4966:
4942:
4937:
4928:
4920:
4915:
4910:978-0691167411
4909:
4892:
4885:
4882:
4880:
4879:
4856:
4835:
4799:
4766:
4752:. 2019-02-01.
4737:
4711:
4681:
4660:(1): 235–250.
4640:
4611:
4584:
4558:
4528:
4510:
4482:
4453:
4428:
4399:
4370:
4344:
4315:
4290:
4265:
4236:
4192:
4163:
4132:
4113:
4101:
4094:
4076:
4062:. 2017-10-27.
4047:
4017:
4013:Merritt Herald
3994:
3990:Merritt Herald
3971:
3948:
3919:
3897:Daley, Jason.
3889:
3859:
3850:
3840:
3830:
3800:
3766:
3735:
3712:
3698:. 2015-01-01.
3683:
3626:
3591:
3550:(4): 887–894.
3527:
3513:. 2017-10-11.
3502:
3458:
3391:
3324:
3287:(2): 310–319.
3271:
3230:
3201:
3172:
3105:
3054:
3025:
3003:
2944:
2900:
2864:
2812:
2793:(1): 119–143.
2777:
2718:
2704:. 2015-03-17.
2689:
2665:
2619:(4): 285–298.
2599:
2570:
2533:
2497:
2477:"Our policies"
2468:
2439:
2420:
2361:
2302:
2268:(9): 1354–60.
2238:
2213:
2156:
2097:
2038:
1999:
1981:
1963:
1953:
1944:
1915:
1896:
1870:
1844:
1815:
1795:"Cats Indoors"
1786:
1762:
1738:
1735:. 7 July 2010.
1724:
1710:
1696:. 2013-08-27.
1681:
1652:
1635:on 2019-03-06.
1619:
1590:(4): 502–511.
1570:
1541:
1521:
1501:
1481:
1462:
1442:
1401:
1388:IFAS Extension
1371:
1340:
1328:The New Yorker
1314:
1278:(4): 290–296.
1258:
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1175:(4): 285–298.
1156:
1126:
1097:
1066:
1042:
991:
969:
925:(9): 800–807.
904:
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892:
887:
880:
877:
831:
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827:
826:
823:
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785:United Kingdom
783:
777:
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650:
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626:
625:
622:
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606:
603:
600:
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592:
562:); hookworms (
515:
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487:Main article:
484:
481:
480:
479:
476:
463:
460:
440:
439:
435:
431:
428:
419:
416:
415:
414:
410:
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400:
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368:
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342:
339:
334:
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307:
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302:
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271:
266:
263:
262:
261:
254:
247:
233:
222:
221:
215:
209:
203:
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191:
184:
161:
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3770:
3756:on 2020-07-14
3752:
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2101:
2090:September 24,
2082:
2078:
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2062:
2058:
2057:
2049:
2042:
2034:
2030:
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2022:
2019:(1): 99–112.
2018:
2014:
2010:
2003:
1995:
1991:
1985:
1977:
1973:
1967:
1957:
1948:
1933:
1929:
1925:
1919:
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1908:
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1900:
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1134:Ark, Noah's.
1130:
1115:
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1110:www.hsppr.org
1107:
1101:
1085:
1081:
1077:
1073:For example:
1070:
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1049:
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1019:
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848:Piping Plover
845:
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837:
830:United States
824:
821:
818:
815:
814:
813:
811:
807:
804:In 2008, the
802:
800:
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792:
791:Celia Hammond
782:
773:
769:
760:
751:
748:
743:
741:
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696:
694:
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687:
683:
678:
669:
662:
656:
646:
643:
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620:
618:
616:
610:
607:
604:
601:
598:
596:
595:Toxoplasmosis
593:
591:
588:
587:
586:
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577:
573:
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565:
561:
560:Toxocara cati
557:
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552:Campylobacter
548:
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520:
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508:
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490:
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384:
380:
376:
375:
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362:
358:
354:
349:
345:
344:
338:
330:
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327:Introgression
324:
315:
311:
303:
299:
295:
294:
293:
291:
286:
284:
280:
270:
259:
255:
252:
248:
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226:
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216:
213:
210:
207:
204:
201:
198:
195:
192:
189:
185:
182:
179:
178:
177:
174:
172:
171:community cat
167:
157:
155:
151:
147:
145:
141:
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133:
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125:
123:
119:
115:
111:
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4957:. Retrieved
4950:ABCBirds.org
4949:
4924:
4896:
4875:AlleyCat.org
4874:
4859:
4843:
4838:
4827:. Retrieved
4811:
4791:. Retrieved
4782:
4758:. Retrieved
4749:
4740:
4728:. Retrieved
4723:
4714:
4702:. Retrieved
4697:
4657:
4653:
4643:
4631:. Retrieved
4626:
4602:. Retrieved
4597:
4587:
4575:. Retrieved
4570:
4561:
4549:. Retrieved
4544:
4531:
4513:
4502:. Retrieved
4492:
4485:
4474:. Retrieved
4465:
4456:
4431:
4420:. Retrieved
4411:
4402:
4391:. Retrieved
4382:
4373:
4361:. Retrieved
4357:
4354:"Feral cats"
4347:
4336:. Retrieved
4327:
4324:"Feral cats"
4318:
4307:. Retrieved
4293:
4282:. Retrieved
4268:
4257:. Retrieved
4249:ResearchGate
4248:
4239:
4206:
4202:
4195:
4184:. Retrieved
4175:
4166:
4155:. Retrieved
4144:
4135:
4116:
4104:
4085:
4079:
4068:. Retrieved
4059:
4050:
4020:
4012:
3997:
3989:
3974:
3966:
3951:
3940:. Retrieved
3931:
3922:
3911:. Retrieved
3902:
3892:
3881:. Retrieved
3872:
3862:
3853:
3843:
3833:
3822:. Retrieved
3813:
3803:
3792:. Retrieved
3783:
3779:
3769:
3758:. Retrieved
3751:the original
3738:
3721:
3715:
3704:. Retrieved
3695:
3686:
3646:(1): 74–81.
3643:
3639:
3629:
3604:
3600:
3594:
3583:. Retrieved
3547:
3543:
3530:
3519:. Retrieved
3505:
3478:
3474:
3408:
3404:
3394:
3341:
3337:
3327:
3284:
3280:
3274:
3247:
3243:
3233:
3222:. Retrieved
3213:
3204:
3193:. Retrieved
3184:
3175:
3122:
3118:
3108:
3070:(1): 15–28.
3067:
3063:
3057:
3046:. Retrieved
3038:icatcare.org
3037:
3028:
3016:. Retrieved
3006:
2961:
2957:
2947:
2914:
2910:
2903:
2892:. Retrieved
2884:www.avma.org
2883:
2832:
2828:
2790:
2786:
2780:
2735:
2731:
2721:
2710:. Retrieved
2701:
2692:
2657:. Retrieved
2616:
2612:
2602:
2591:. Retrieved
2582:
2573:
2561:. Retrieved
2549:
2536:
2528:
2522:. Retrieved
2513:
2489:. Retrieved
2480:
2471:
2460:. Retrieved
2451:
2442:
2423:
2378:
2374:
2364:
2319:
2315:
2305:
2293:. Retrieved
2265:
2259:
2216:
2173:
2169:
2159:
2114:
2110:
2100:
2088:. Retrieved
2063:(1): 42–46.
2060:
2054:
2041:
2016:
2012:
2002:
1984:
1974:. Aug 2012.
1966:
1956:
1947:
1936:. Retrieved
1927:
1918:
1899:
1887:. Retrieved
1882:
1873:
1861:. Retrieved
1856:
1847:
1835:. Retrieved
1830:
1818:
1807:. Retrieved
1798:
1789:
1776:. Mar 2017.
1765:
1752:. Sep 2014.
1741:
1727:
1713:
1702:. Retrieved
1693:
1684:
1673:. Retrieved
1664:
1655:
1633:the original
1622:
1587:
1583:
1573:
1562:. Retrieved
1553:
1544:
1524:
1504:
1484:
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