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Punctuated equilibrium

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celebrated for the color and energy of his prose, as well as his interdisciplinary knowledge, critics such as Scott, Richard Dawkins, and Daniel Dennett have concerns that the theory has gained undeserved credence among non-scientists because of Gould's rhetorical skills. Philosopher John Lyne and biologist Henry Howe believed punctuated equilibrium's success has much more to do with the nature of the geological record than the nature of Gould's rhetoric. They state, a "re-analysis of existing fossil data has shown, to the increasing satisfaction of the paleontological community, that Eldredge and Gould were correct in identifying periods of evolutionary stasis which are interrupted by much shorter periods of evolutionary change."
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these species become modified and improved, others will have to be improved in a corresponding degree, or they will be exterminated. Each new form, also, as soon as it has been improved, will be able to spread over the open and continuous area, and will thus come into competition with many other forms ... the new forms produced on large areas, which have already been victorious over many competitors, will be those that will spread most widely, and will give rise to the greatest number of new varieties and species. They will thus play a more important role in the changing history of the organic world.
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speedists", on the other hand, advance that "evolutionary rates fluctuate continuously from very fast to very slow and stop, with all intermediates. They see no particular reason to emphasize certain speeds more than others. In particular, stasis, to them, is just an extreme case of ultra-slow evolution. To a punctuationist, there is something very special about stasis."
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English professor Heidi Scott argues that Gould's talent for writing vivid prose, his use of metaphor, and his success in building a popular audience of nonspecialist readers altered the "climate of specialized scientific discourse" favorably in his promotion of punctuated equilibrium. While Gould is
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studies could revitalize various aspects of paleontology and macroevolution. Tom Schopf, who organized that year's meeting, assigned Gould the topic of speciation. Gould recalls that "Eldredge's 1971 publication had presented the only new and interesting ideas on the paleontological implications of
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Much confusion has arisen over what proponents of punctuated equilibrium actually argued, what mechanisms they advocated, how fast the punctuations were, what taxonomic scale their theory applied to, how revolutionary their claims were intended to be, and how punctuated equilibrium related to other
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in clarifying the meaning of stasis adds, "In claiming that species typically undergo no further evolutionary change once speciation is complete, they are not claiming that there is no change at all between one generation and the next. Lineages do change. But the change between generations does not
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Eldredge recalls that after much discussion the pair "each wrote roughly half. Some of the parts that would seem obviously the work of one of us were actually first penned by the other—I remember for example, writing the section on Gould's snails. Other parts are harder to reconstruct. Gould edited
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Dawkins also emphasizes that punctuated equilibrium has been "oversold by some journalists", but partly due to Eldredge and Gould's "later writings". Dawkins contends that the hypothesis "does not deserve a particularly large measure of publicity". It is a "minor gloss," an "interesting but minor
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Throughout a great and open area, not only will there be a greater chance of favourable variations, arising from the large number of individuals of the same species there supported, but the conditions of life are much more complex from the large number of already existing species; and if some of
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The fossil record includes well documented examples of both phyletic gradualism and punctuational evolution. As such, much debate persists over the prominence of stasis in the fossil record. Before punctuated equilibrium, most evolution biologists considered stasis to be rare or unimportant. The
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Darwin wrote that "the periods during which species have undergone modification, though long as measured in years, have probably been short in comparison with the periods during which they retain the same form." Thus punctuationism in general is consistent with Darwin's conception of evolution.
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speedism". Eldredge and Gould, proposing that evolution jumps between stability and relative rapidity, are described as "discrete variable speedists", and "in this respect they are genuinely radical." They assert that evolution generally proceeds in bursts, or not at all. "Continuously variable
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Simpson stated, "Evolutionary change is so nearly the universal rule that a state of motion is, figuratively, normal in evolving populations. The state of rest, as in bradytely, is the exception and it seems that some restraint or force must be required to maintain it." Despite such differences
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discontinuity (or "sudden jumps") found in the fossil record. Mayr later complimented Eldredge and Gould's paper, stating that evolutionary stasis had been "unexpected by most evolutionary biologists" and that punctuated equilibrium "had a major impact on paleontology and evolutionary biology."
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are diluted by the population's large size and are unable to reach fixation, due to such factors as constantly changing environments. If this is the case, then the transformation of whole lineages should be rare, as the fossil record indicates. Smaller populations on the other hand, which are
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regards the apparent gaps represented in the fossil record as documenting migratory events rather than evolutionary events. According to Dawkins, evolution certainly occurred but "probably gradually" elsewhere. However, the punctuational equilibrium model may still be inferred from both the
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is especially critical of Gould's presentation of punctuated equilibrium. Dennett argues that Gould alternated between revolutionary and conservative claims, and that each time Gould made a revolutionary statement—or appeared to do so—he was criticized, and thus retreated to a traditional
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accused Gould of being "something of an intellectual fraud" for using claims that were "well known from the time of Darwin" (that evolution displayed "periods of long stasis interspersed with periods of rapid change") to support distinct but more "grandiose" claims regarding
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Rapidly evolving peripherally isolated populations may be the place of origin of many evolutionary novelties. Their isolation and comparatively small size may explain phenomena of rapid evolution and lack of documentation in the fossil record, hitherto puzzling to the
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is especially intense, as peripheral isolated populations exist at the outer edges of ecological tolerance. If most evolution happens in these rare instances of allopatric speciation then evidence of gradual evolution in the fossil record should be rare. This
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states that "Species of different genera and classes have not changed at the same rate, or in the same degree. In the oldest tertiary beds a few living shells may still be found in the midst of a multitude of extinct forms... The Silurian
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that may underlie abrupt morphological transitions during evolution. Consequently, consideration of mechanisms of phylogenetic change that have been found in reality to be non-gradual is increasingly common in the field of
970:'s theories of developmental and genetic homeostasis. However this hypothesis was rejected over time, as evidence accumulated against it. Other plausible mechanisms which have been suggested include: habitat tracking, 846:
the entire manuscript for better consistency. We sent it in, and Schopf reacted strongly against it—thus signaling the tenor of the reaction it has engendered, though for shifting reasons, down to the present day."
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the subject—so I asked Schopf if we could present the paper jointly." According to Gould "the ideas came mostly from Niles, with yours truly acting as a sounding board and eventual scribe. I coined the term
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examining 58 published studies on speciation patterns in the fossil record showed that 71% of species exhibited stasis, and 63% were associated with punctuated patterns of evolutionary change. According to
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of organisms. This hypothesis differs from punctuated equilibrium in several respects. First, punctuated equilibrium was more modest in scope, in that it was addressing evolution specifically at the
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Thus punctuated equilibrium is incongruous with some of Darwin's ideas regarding the specific mechanisms of evolution, but generally accords with Darwin's theory of evolution by natural selection.
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research program. Punctuated equilibrium differs from Mayr's ideas mainly in that Eldredge and Gould placed considerably greater emphasis on stasis, whereas Mayr was concerned with explaining the
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level. Simpson's idea was principally concerned with evolution at higher taxonomic groups. Second, Eldredge and Gould relied upon a different mechanism. Where Simpson relied upon a
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was considered the "standard" model of speciation. This model was popularized by Ernst Mayr in his 1954 paper "Change of genetic environment and evolution," and his classic volume
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Although there exist some debate over how long the punctuations last, supporters of punctuated equilibrium generally place the figure between 50,000 and 100,000 years.
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and seemingly "rapid" episodes of gradual evolution could give the appearance of instantaneous change, since 10,000 years seldom registers in the geological record.
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stability followed by rare bursts of evolutionary change via rapid cladogenesis – vertical equilibrium states separated by horizontal "jump" phases. In contrast,
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According to early versions of punctuated equilibrium, "peripheral isolates" are considered to be of critical importance for speciation. However, Darwin wrote, "
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The punctuational nature of punctuated equilibrium has engendered perhaps the most confusion over Eldredge and Gould's theory. Gould's sympathetic treatment of
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as applied to the fossil record. Although the sudden appearance of species and its relationship to speciation was proposed and identified by Mayr in 1954,
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Although punctuated equilibrium generally applies to sexually reproducing organisms, some biologists have applied the model to non-sexual species like
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Some critics jokingly referred to the theory of punctuated equilibrium as "evolution by jerks", which reportedly prompted punctuationists to describe
5939: 3371: 4176: 974:, the Stenseth-Maynard Smith stability hypothesis, constraints imposed by the nature of subdivided populations, normalizing clade selection, and 1030: 713:. Cladogenesis is the process by which a species splits into two distinct species, rather than one species gradually transforming into another. 660: 4169: 2294: 1424:, particularly in studies of the origin of morphological novelty. A description of such mechanisms can be found in the multi-authored volume 5374: 1321:, who appealed to the imperfection of the record as the favored explanation. When presenting his ideas against the prevailing influences of 4674: 4145:
Abstract Profiles of Structural Stability Point to Universal Tendencies, Family-Specific Factors, and Ancient Connections between Languages
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Atkinson, Quentin D.; Meade, Andrew; Venditti, Chris; Greenhill, Simon J.; Pagel, Mark (2008). "Languages Evolve in Punctuational Bursts".
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It is often incorrectly assumed that he insisted that the rate of change must be constant, or nearly so, but even the first edition of
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language family there. Although his model has raised considerable interest, it does not command majority support within linguistics.
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is among the few brachiopods surviving today but also known from fossils over 500 million years old. In the fourth edition (1866) of
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Allopatric speciation suggests that species with large central populations are stabilized by their large volume and the process of
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which suggested that gradual evolution was seldom seen in the fossil record and argued that Ernst Mayr's standard mechanism of
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Koeslag, J. H. (1990). "Koinophilia groups sexual creatures into species, promotes stasis, and stabilizes social behaviour".
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The sudden appearance of most species in the geologic record and the lack of evidence of substantial gradual change in most
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and wrote most of our 1972 paper, but Niles is the proper first author in our pairing of Eldredge and Gould." In his book
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has proposed a punctuated equilibrium model for language histories, with reference particularly to the prehistory of the
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The importance of isolation in forming species had played a significant part in Darwin's early thinking, as shown in his
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According to Gould, "stasis may emerge as the theory's most important contribution to evolutionary science." Philosopher
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as well as Simpson himself—have argued that punctuated equilibrium is little more than quantum evolution relabeled.
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Lyne, John; Howe, Henry F. (1986). ""Punctuated equilibria": Rhetorical dynamics of a scientific controversy".
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observation of stasis and examples of rapid and episodic speciation events documented in the fossil record.
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wrinkle on the surface of neo-Darwinian theory," and "lies firmly within the neo-Darwinian synthesis".
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also failed to grasp the implication(s), primarily because they did not think at geological scales".
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Alternative explanations for the punctuated pattern of evolution observed in the fossil record. Both
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missed this insight because they had not studied evolutionary theory and either did not know about
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Wake, David B.; Roth, G.; Wake, M. H. (1983). "On the problem of stasis in organismal evolution".
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Punctuational evolution has been argued to explain changes in folktales and mythology over time.
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This article is about the theory in evolutionary biology. For the theory of social change, see
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A Cosmic Hunt in the Berber sky : a phylogenetic reconstruction of Palaeolithic mythology
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Punctuated equilibrium has also been cited as contributing to the hypothesis that species are
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for most of its geological history. This state of little or no morphological change is called
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is virtually nonexistent in the fossil record, and that stasis dominates the history of most
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he had downplayed its importance. He explained the reasons for his revised view as follows:
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Gemmell, Michael R.; Trewick, Steven A.; Hills, Simon F. K.; Morgan-Richards, Mary (2019).
2771: 2703:; Maynard Smith, John (1984). "Coevolution in ecosystems: Red Queen evolution or stasis?". 2672: 2424: 2229: 1906: 1500: 1201: 1178: 804: 674: 451: 223: 108: 65: 46: 5413: 5283: 2340:"Speciation in real time and historical-archaeological and its absence in geological time" 853:
proposed an "anticipation of the theory of punctuated equilibrium of Gould and Eldredge."
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record, and to portray contemporary evolutionary biology as advancing neo-saltationism.
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sp.) also confirm morphological stasis in multiple lineages over three million years.
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p. 450–451, 487–490, 499–501. Redwood City CA: Benjamin Cummings Publishing Company.
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de Brito Neto, S. G.; Fernando Alves, E.; Cavalcante e Almeida Sá, Mariana (2017).
2237: 2049:. Cambridge, Massachusetts: The Belknap Press of Harvard University Press. p.  1914: 1897: 1820: 1812: 1681: 1671: 1631: 1621: 1598: 1239: 1133: 763:. Eldredge and Gould proposed that the degree of gradualism commonly attributed to 320: 6339: 5293: 3737:
Scott, Heidi (2007). "Stephen Jay Gould and the Rhetoric of Evolutionary Theory".
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in his terminology) comprised 90% of evolution. More modern studies, including a
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methods claims to show that punctuational bursts play an important factor when
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Polyphemus (Aa. Th. 1137) A phylogenetic reconstruction of a prehistoric tale.
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generally recognize the 1972 Eldredge and Gould paper as the basis of the new
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Tempo and mode in evolution: genetics and paleontology 50 years after Simpson
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between the two models, earlier critiques—from such eminent commentators as
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Michaux, B. (1989). "Morphological variation of species through time".
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Evidence for stasis has also been corroborated from the genetics of
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Punctuated equilibrium is often portrayed to oppose the concept of
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was alluded to by Mayr in the closing paragraph of his 1954 paper:
559: 158: 3570:(1994). "Quantitative genetics of bryozoan phenotypic evolution". 2840: 2661:"An evaluation of stabilizing selection as a mechanism for stasis" 7093: 5156: 4870: 4477: 4312: 2544:
Futuyma, Douglas (1987). "On the role of species in anagenesis".
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John Wilkins and Gareth Nelson have argued that French architect
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published a landmark paper developing their theory and called it
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Brett, Carlton E.; Ivany, Linda C.; Schopf, Kenneth M. (1996).
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neo-Darwinian position. Gould responded to Dennett's claims in
987: 768: 615: 4096: 1571:"Punctuated equilibria: An alternative to phyletic gradualism" 1189:
or had not considered its translation to geological time. Our
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accumulate. Instead, over time, the species wobbles about its
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Punctuated equilibrium originated as a logical consequence of
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work on the importance of reproductive isolating mechanisms.
947: 3484:. New York: W. W. Norton & Co., Chapter 9. (p. 224-252) 2847:
Journal of Zoological Systematics and Evolutionary Research
1359:
differs but little from the living species of this genus".
832:
in 1971. The symposium focused its attention on how modern
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All you need to know about Punctuated Equilibrium (almost)
3538:
Dawkins, Richard (1996). The Blind Watchmaker, p. 245-246.
1740:. Vol. 94. pp. 137–139 – via Google Books. 1561: 1313:—from their initial appearance until their extinction—has 2589:. Cambridge, Massachusetts: Harvard University Press, p. 2279: 2277: 1205:
to correcting, in his view, the wide confusion regarding
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isolated from the parental stock, are decoupled from the
4173:
Nouvelle Mythologie Comparée / New Comparative Mythology
1400: 2410: 2411:
Bomfleur, B.; McLoughlin, S.; Vajda, V. (March 2014).
2274: 2015:. New York: Columbia University Press, pp. 78–85. 3847:
The Richness of Life: The Essential Stephen Jay Gould
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Nichol, S.T, Joan Rowe, and Walter M. Fitch (1993).
1732:"Punctuated equilibrium — a different way of seeing" 1415:
has identified dynamical and physical mechanisms of
904:
When Eldredge and Gould published their 1972 paper,
697:, the population will become stable, showing little 3408:"Character change, speciation, and the higher taxa" 2804:Koeslag, J.H. (1995). On the engine of speciation. 2100:
Time Frames: The evolution of punctuated equilibria
1722: 1720: 966:causes of stasis. Gould was initially attracted to 830:
Annual Meeting of the Geological Society of America
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(eds). 2125:History and Philosophy of the Life Sciences 1788: 1757: 1726: 1124:was a controversial hypothesis advanced by 5940: 5926: 4645: 4631: 4262: 4248: 4063: 3983: 2932:Dawkins vs. Gould: Survival of the Fittest 2830:. New York: Chapman and Hall. p. 126. 2646:. Princeton University Press, pp. 139-141. 2072: 2070: 1950: 1948: 1946: 1944: 1753: 1751: 1749: 1747: 661: 647: 4120: 3378:. New York: Columbia Univ. Press, p. 390. 3356:Washington, D.C.: National Academy Press. 3073:"The genetics of stasis and punctuations" 2970: 2884:Biological Journal of the Linnean Society 2858: 2684: 2649: 2162: 1978:. New York: Oxford University Press, pp. 1685: 1675: 1635: 1625: 1097:In an often quoted remark, Gould stated, 1016: 894: 7104:Transgenerational epigenetic inheritance 5879:Transgenerational epigenetic inheritance 4652: 4177:Les mythes Ă©volueraient par ponctuations 3771: 3732: 3730: 3340:. New York: Columbia Univ. 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But by the time he wrote the 709:events of branching speciation called 6596:Psychological effects of Internet use 5921: 5642:Dialogues Concerning Natural Religion 4626: 4243: 3999:"Punctuated equilibrium comes of age" 3736: 3727: 3259: 3235: 3128: 3012: 2935:. Cambridge, U.K.: Icon Books, p. 96. 2184:Erwin, D. H. and R. L. Anstey (1995) 2037: 1877: 1853: 1548: 1401:Supplemental modes of rapid evolution 825:might suggest a possible resolution. 4581: 3977:. London: John Murray. 5th edition, 3961:. London: John Murray. 5th edition, 3944:. London: John Murray. 5th edition, 3849:. New York: W. W. Norton & Co., 3349:Fitch, W. J. and F. J. Ayala (1995) 3046:. 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Cape Town: Struik Publishers. 1465:languages split from one another 628: 627: 614: 72: 6544:Computer-mediated communication 5704:Extended evolutionary synthesis 4893:Gene-centered view of evolution 4154: 4090: 4057: 4044: 3967: 3951: 3934: 3917: 3870: 3855: 3836: 3817: 3792: 3765: 3702: 3676: 3661: 3646: 3631: 3613: 3598: 3556: 3541: 3532: 3514: 3502: 3487: 3456: 3399: 3368:The Major Features of Evolution 3359: 3343: 3309: 3253: 3229: 3195: 3175:. Simon and Schuster. pp.  3116: 3097: 3061: 3030: 3026:. 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Vol. 2. pp. 34–37. 3241:"Evolution as fact and theory" 2764:Journal of Theoretical Biology 2222:Journal of Theoretical Biology 1968: 1847: 1770: 1668:"3rd draft (final, published)" 1459:Separately, recent work using 1427:Origination of Organismal Form 1272:, and in his technical volume 1107:abundant between larger groups 1013:describes this very process." 580:Creation–evolution controversy 334:History of evolutionary theory 36:Punctuated Equilibrium (album) 27:Theory in evolutionary biology 13: 1: 7181:Standard social science model 6234:Cognitive tradeoff hypothesis 5626:Renaissance and Enlightenment 3911:The Virginia Quarterly Review 3900:"Confessions of a Darwinist." 3845:and Steven Rose, ed. 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Michael Lerner 952:Douglas Futuyma's 930:natural selection 863:Rate of evolution 834:microevolutionary 733:Stephen Jay Gould 671: 670: 362:Origin of Species 164:Natural selection 16:(Redirected from 7336: 7304:Biology theories 7271: 7258: 7245: 7244: 6888: 6887: 6884:Related subjects 6671:Adult attachment 6198:Cognitive module 6154: 6153: 6141:Social selection 6115:Costly signaling 6110:Sexual selection 5997:Modern synthesis 5942: 5935: 5928: 5919: 5918: 5905: 5895: 5894: 5694:Modern synthesis 5454:Multicellularity 5449:Mosaic evolution 5334:auditory ossicle 5016:Social selection 4999:Flowering plants 4994:Sexual selection 4647: 4640: 4633: 4624: 4623: 4608: 4607: 4596: 4584: 4583: 4572: 4571: 4423:Sexual selection 4352:Geographic modes 4264: 4257: 4250: 4241: 4240: 4184: 4158: 4152: 4142: 4124: 4094: 4088: 4087: 4074:10.1075/cilt.249 4061: 4055: 4048: 4042: 4041: 4023:10.1038/366223a0 3987: 3981: 3971: 3965: 3955: 3949: 3938: 3932: 3921: 3915: 3896: 3883: 3874: 3868: 3859: 3853: 3840: 3834: 3833: 3821: 3815: 3814: 3812: 3811: 3805:Psychology Today 3796: 3790: 3789: 3769: 3763: 3762: 3734: 3725: 3724: 3722: 3721: 3706: 3700: 3680: 3674: 3665: 3659: 3650: 3644: 3635: 3629: 3620:Dawkins, Richard 3617: 3611: 3602: 3596: 3595: 3560: 3554: 3545: 3539: 3536: 3530: 3521:Dawkins, Richard 3518: 3512: 3509:Dawkins, Richard 3506: 3500: 3491: 3485: 3476:Dawkins, Richard 3473: 3467: 3460: 3454: 3453: 3451: 3445:. 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Archived from 3045: 3034: 3028: 3027: 3010: 3004: 2983: 2977: 2976: 2974: 2942: 2936: 2927: 2921: 2906: 2900: 2899: 2879: 2873: 2872: 2862: 2838: 2832: 2831: 2820: 2814: 2802: 2796: 2795: 2759: 2753: 2743: 2737: 2736: 2697: 2691: 2690: 2688: 2671:(1–4): 229–238. 2656: 2647: 2640: 2634: 2633: 2600: 2594: 2578: 2572: 2571: 2541: 2535: 2525: 2519: 2509: 2503: 2486: 2480: 2466: 2457: 2456: 2408: 2402: 2384: 2378: 2377: 2371: 2363: 2335: 2329: 2314: 2308: 2281: 2272: 2265: 2259: 2252: 2246: 2245: 2217: 2211: 2195: 2189: 2182: 2176: 2169: 2160: 2150: 2141: 2140: 2122: 2113: 2107: 2093: 2087: 2074: 2065: 2064: 2048: 2035: 2029: 2022: 2016: 2005: 1999: 1988: 1982: 1972: 1966: 1952: 1939: 1938: 1919:10.1038/305269a0 1881: 1875: 1874: 1851: 1845: 1844: 1795: 1786: 1785: 1774: 1768: 1767: 1755: 1742: 1741: 1724: 1715: 1712: 1706: 1701: 1699: 1691: 1689: 1679: 1662: 1656: 1651: 1649: 1641: 1639: 1629: 1612: 1610: 1609: 1603: 1596: 1578: 1559: 1444:In linguistics, 1240:Push of the past 1084:hopeful monsters 942:palaeontologist. 759:, and their own 663: 656: 649: 636: 631: 630: 623: 619: 618: 595:Level of support 388:Current research 373:Modern synthesis 368:Before synthesis 321:Extinction event 79:Darwin's finches 76: 57: 56: 21: 18:Stasis (biology) 7344: 7343: 7339: 7338: 7337: 7335: 7334: 7333: 7289: 7288: 7285: 7280: 7222: 7208:Neoevolutionism 7115: 7099:Species complex 7064:Group selection 7002:Research topics 6997: 6973:Neuropsychology 6875: 6861:Substance abuse 6783:Sex differences 6777: 6691:Coolidge effect 6652: 6564:Neuroergonomics 6529: 6520: 6444: 6346: 6280:Folk psychology 6161: 6145: 6015: 6008: 5951: 5946: 5916: 5911: 5883: 5810:Group selection 5783: 5708: 5612: 5539: 5501:Tempo and modes 5495: 5350: 5254: 5071: 5030: 4906: 4899: 4876:Species complex 4689: 4680:History of life 4656: 4651: 4621: 4616: 4560: 4543:Paleopolyploidy 4507: 4462:Hybrid concepts 4456: 4399: 4347: 4317:Species complex 4301: 4273: 4268: 4192: 4187: 4159: 4155: 4095: 4091: 4084: 4062: 4058: 4049: 4045: 3995:Eldredge, Niles 3988: 3984: 3972: 3968: 3956: 3952: 3939: 3935: 3922: 3918: 3907:Wayback Machine 3897: 3886: 3875: 3871: 3860: 3856: 3841: 3837: 3822: 3818: 3809: 3807: 3797: 3793: 3780:(2): 132–147 . 3770: 3766: 3739:Rhetoric Review 3735: 3728: 3719: 3717: 3708: 3707: 3703: 3681: 3677: 3666: 3662: 3651: 3647: 3636: 3632: 3618: 3614: 3603: 3599: 3584:10.2307/2410098 3561: 3557: 3546: 3542: 3537: 3533: 3519: 3515: 3507: 3503: 3492: 3488: 3474: 3470: 3461: 3457: 3449: 3427:10.2307/2408092 3410: 3404: 3400: 3386: 3382: 3376:Wayback Machine 3364: 3360: 3348: 3344: 3333: 3326: 3314: 3310: 3301: 3299: 3295: 3264: 3258: 3254: 3234: 3230: 3223: 3200: 3196: 3187: 3162: 3121: 3117: 3102: 3098: 3090: 3075: 3066: 3062: 3054: 3043: 3035: 3031: 3023:Natural History 3011: 3007: 2997:Wayback Machine 2984: 2980: 2943: 2939: 2928: 2924: 2907: 2903: 2880: 2876: 2839: 2835: 2821: 2817: 2807:J. theor. Biol. 2803: 2799: 2760: 2756: 2746:Williams, G. C. 2744: 2740: 2717:10.2307/2408397 2701:Stenseth, N. C. 2698: 2694: 2657: 2650: 2641: 2637: 2630: 2604:Eldredge, Niles 2601: 2597: 2579: 2575: 2542: 2538: 2526: 2522: 2510: 2506: 2489:Eldredge, Niles 2487: 2483: 2467: 2460: 2409: 2405: 2395:Wayback Machine 2385: 2381: 2365: 2364: 2336: 2332: 2315: 2311: 2299:Wayback Machine 2288:Eldredge, Niles 2282: 2275: 2266: 2262: 2253: 2249: 2218: 2214: 2196: 2192: 2183: 2179: 2170: 2163: 2151: 2144: 2120: 2114: 2110: 2096:Eldredge, Niles 2094: 2090: 2084:Natural History 2075: 2068: 2061: 2036: 2032: 2023: 2019: 2006: 2002: 1989: 1985: 1973: 1969: 1953: 1942: 1882: 1878: 1865:) in Bermuda". 1852: 1848: 1817:10.2307/2406508 1799:Eldredge, Niles 1796: 1789: 1775: 1771: 1756: 1745: 1725: 1718: 1704: 1702: 1693: 1692: 1654: 1652: 1643: 1642: 1607: 1605: 1601: 1594: 1586: 1563:Eldredge, Niles 1560: 1549: 1545: 1540: 1481: 1473: 1442: 1440:Language change 1436: 1434:Language change 1411:Recent work in 1409: 1403: 1317:, including by 1315:long been noted 1307: 1305:Darwin's theory 1245:Richard Dawkins 1242: 1236:Species complex 1232:Species problem 1228: 1217:speedism" and " 1207:rates of change 1197:Richard Dawkins 1175: 1128:paleontologist 1119: 1092:paleontological 1060: 1052:mass extinction 1039: 1025:, and not just 1019: 1005:Jonathan Weiner 983:sibling species 960: 919:. New and even 902: 897: 868:paleontologist 865: 859: 801:paleobiological 791:and especially 777: 751:'s theories of 693:appears in the 667: 626: 613: 612: 605: 604: 555: 547: 546: 417: 409: 408: 407: 335: 327: 326: 325: 274:Human evolution 264:History of life 248: 247:Natural history 240: 239: 238: 138: 130: 85: 39: 28: 23: 22: 15: 12: 11: 5: 7342: 7332: 7331: 7326: 7321: 7316: 7311: 7306: 7301: 7282: 7281: 7279: 7278: 7265: 7252: 7239: 7231: 7228: 7227: 7224: 7223: 7221: 7220: 7215: 7210: 7205: 7200: 7195: 7190: 7185: 7184: 7183: 7178: 7173: 7168: 7163: 7158: 7153: 7148: 7143: 7129: 7123: 7121: 7117: 7116: 7114: 7113: 7112: 7111: 7106: 7101: 7096: 7091: 7086: 7081: 7076: 7071: 7066: 7061: 7056: 7051: 7046: 7036: 7031: 7026: 7021: 7016: 7011: 7005: 7003: 6999: 6998: 6996: 6995: 6990: 6985: 6980: 6975: 6970: 6965: 6960: 6955: 6950: 6945: 6940: 6935: 6930: 6913: 6904: 6894: 6892: 6885: 6881: 6880: 6877: 6876: 6874: 6873: 6868: 6863: 6858: 6853: 6848: 6843: 6838: 6833: 6828: 6823: 6818: 6813: 6808: 6803: 6798: 6793: 6787: 6785: 6779: 6778: 6776: 6775: 6770: 6765: 6752: 6743: 6738: 6733: 6728: 6723: 6718: 6713: 6708: 6703: 6698: 6693: 6688: 6683: 6678: 6673: 6668: 6662: 6660: 6654: 6653: 6651: 6650: 6649: 6648: 6643: 6638: 6633: 6623: 6618: 6613: 6608: 6603: 6598: 6593: 6591:Mind-blindness 6588: 6583: 6578: 6573: 6568: 6567: 6566: 6561: 6556: 6551: 6546: 6535: 6533: 6522: 6521: 6519: 6518: 6513: 6508: 6503: 6498: 6493: 6488: 6483: 6478: 6465: 6460: 6454: 6452: 6446: 6445: 6443: 6442: 6437: 6436: 6435: 6425: 6424: 6423: 6413: 6412: 6411: 6406: 6401: 6391: 6386: 6385: 6384: 6374: 6373: 6372: 6367: 6356: 6354: 6348: 6347: 6345: 6344: 6343: 6342: 6337: 6332: 6322: 6317: 6312: 6303: 6302: 6301: 6296: 6286: 6284:theory of mind 6277: 6268: 6267: 6266: 6261: 6256: 6246: 6241: 6236: 6231: 6230: 6229: 6224: 6219: 6214: 6209: 6195: 6190: 6189: 6188: 6183: 6178: 6167: 6165: 6151: 6147: 6146: 6144: 6143: 6138: 6137: 6136: 6131: 6126: 6117: 6107: 6106: 6105: 6095: 6090: 6085: 6080: 6079: 6078: 6068: 6063: 6058: 6053: 6051:Baldwin effect 6048: 6047: 6046: 6041: 6036: 6026: 6020: 6018: 6010: 6009: 6007: 6006: 6001: 6000: 5999: 5994: 5989: 5984: 5979: 5969: 5968: 5967: 5956: 5953: 5952: 5945: 5944: 5937: 5930: 5922: 5913: 5912: 5910: 5909: 5899: 5888: 5885: 5884: 5882: 5881: 5876: 5871: 5866: 5861: 5860: 5859: 5849: 5844: 5839: 5834: 5829: 5828: 5827: 5822: 5817: 5807: 5802: 5797: 5791: 5789: 5785: 5784: 5782: 5781: 5776: 5775: 5774: 5769: 5764: 5763: 5762: 5752: 5747: 5742: 5737: 5732: 5722: 5716: 5714: 5710: 5709: 5707: 5706: 5701: 5696: 5691: 5686: 5681: 5676: 5671: 5666: 5661: 5660: 5659: 5650:Charles Darwin 5647: 5646: 5645: 5633: 5628: 5622: 5620: 5614: 5613: 5611: 5610: 5605: 5600: 5595: 5590: 5588:Non-ecological 5585: 5580: 5575: 5570: 5565: 5560: 5555: 5549: 5547: 5541: 5540: 5538: 5537: 5528: 5519: 5505: 5503: 5497: 5496: 5494: 5493: 5488: 5487: 5486: 5481: 5476: 5471: 5466: 5456: 5451: 5446: 5441: 5436: 5431: 5426: 5421: 5416: 5411: 5406: 5405: 5404: 5394: 5389: 5384: 5379: 5378: 5377: 5372: 5361: 5359: 5352: 5351: 5349: 5348: 5347: 5346: 5341: 5339:nervous system 5336: 5331: 5326: 5318: 5317: 5316: 5311: 5306: 5301: 5296: 5291: 5281: 5276: 5271: 5265: 5263: 5256: 5255: 5253: 5252: 5247: 5242: 5237: 5232: 5231: 5230: 5220: 5219: 5218: 5213: 5212: 5211: 5206: 5196: 5191: 5186: 5181: 5176: 5175: 5174: 5169: 5159: 5149: 5144: 5143: 5142: 5132: 5127: 5122: 5117: 5116: 5115: 5105: 5100: 5099: 5098: 5088: 5082: 5080: 5073: 5072: 5070: 5069: 5064: 5059: 5054: 5049: 5044: 5038: 5036: 5032: 5031: 5029: 5028: 5023: 5018: 5013: 5012: 5011: 5006: 5001: 4991: 4986: 4981: 4976: 4971: 4970: 4969: 4964: 4954: 4949: 4944: 4943: 4942: 4932: 4927: 4922: 4917: 4911: 4909: 4901: 4900: 4898: 4897: 4896: 4895: 4885: 4880: 4879: 4878: 4873: 4863: 4862: 4861: 4851: 4846: 4841: 4839:Origin of life 4836: 4831: 4826: 4824:Microevolution 4821: 4819:Macroevolution 4816: 4811: 4806: 4805: 4804: 4794: 4789: 4784: 4779: 4774: 4769: 4764: 4759: 4757:Common descent 4754: 4753: 4752: 4742: 4737: 4735:Baldwin effect 4732: 4731: 4730: 4725: 4715: 4710: 4705: 4699: 4697: 4691: 4690: 4688: 4687: 4682: 4677: 4672: 4667: 4661: 4658: 4657: 4650: 4649: 4642: 4635: 4627: 4618: 4617: 4615: 4614: 4602: 4590: 4578: 4565: 4562: 4561: 4559: 4558: 4551:Macroevolution 4536: 4531: 4526: 4521: 4515: 4513: 4509: 4508: 4506: 4505: 4500: 4495: 4485: 4466: 4464: 4458: 4457: 4455: 4454: 4452:Haldane's rule 4449: 4444: 4439: 4425: 4420: 4415: 4409: 4407: 4401: 4400: 4398: 4397: 4392: 4378: 4375:Founder effect 4355: 4353: 4349: 4348: 4346: 4345: 4340: 4335: 4330: 4325: 4320: 4309: 4307: 4306:Basic concepts 4303: 4302: 4300: 4299: 4294: 4289: 4284: 4278: 4275: 4274: 4267: 4266: 4259: 4252: 4244: 4238: 4237: 4228: 4222: 4216: 4207: 4201: 4191: 4190:External links 4188: 4186: 4185: 4160:Julien d'Huy, 4153: 4089: 4082: 4056: 4043: 3982: 3966: 3950: 3933: 3916: 3884: 3869: 3854: 3835: 3816: 3791: 3764: 3745:(2): 120–141. 3726: 3701: 3675: 3660: 3645: 3630: 3612: 3597: 3578:(2): 360–375. 3568:Hayek, Lee-Ann 3564:Cheetham, Alan 3555: 3540: 3531: 3513: 3501: 3486: 3468: 3455: 3452:on 2011-08-28. 3421:(3): 427–443. 3398: 3380: 3358: 3342: 3324: 3308: 3252: 3228: 3221: 3203:Hanegraaff, H. 3194: 3192: 3191: 3185: 3166: 3160: 3138: 3115: 3096: 3093:on 2010-12-06. 3060: 3057:on 2016-06-23. 3029: 3005: 2978: 2937: 2922: 2901: 2890:(3): 239–255. 2874: 2853:(1): 209–220. 2833: 2815: 2797: 2754: 2738: 2711:(4): 870–880. 2692: 2648: 2635: 2628: 2595: 2573: 2560:10.1086/284724 2554:(3): 465–473. 2536: 2520: 2504: 2481: 2458: 2403: 2379: 2330: 2309: 2273: 2260: 2247: 2212: 2190: 2177: 2161: 2142: 2131:(2): 179–206. 2108: 2088: 2066: 2059: 2030: 2017: 2000: 1983: 1967: 1940: 1885:Rhodes, F.H.T. 1876: 1863:Poecilozonites 1846: 1811:(1): 156–167. 1787: 1769: 1743: 1716: 1714: 1713: 1705:|journal= 1663: 1655:|journal= 1582: 1581: 1546: 1544: 1541: 1539: 1538: 1533: 1528: 1523: 1518: 1513: 1508: 1503: 1498: 1493: 1488: 1482: 1480: 1477: 1472: 1469: 1446:R. M. W. Dixon 1435: 1432: 1402: 1399: 1319:Charles Darwin 1306: 1303: 1287:Robert Trivers 1263:Daniel Dennett 1261:, philosopher 1227: 1224: 1174: 1171: 1118: 1115: 1111: 1110: 1059: 1056: 1038: 1035: 1033:of evolution. 1018: 1015: 959: 956: 901: 898: 896: 893: 883:Michael Benton 861:Main article: 858: 855: 851:Pierre TrĂ©maux 813:Niles Eldredge 783:'s concept of 776: 773: 765:Charles Darwin 729:Niles Eldredge 669: 668: 666: 665: 658: 651: 643: 640: 639: 638: 637: 624: 607: 606: 603: 602: 597: 592: 587: 582: 577: 575:Social effects 572: 567: 562: 556: 553: 552: 549: 548: 545: 544: 539: 534: 529: 524: 519: 514: 509: 504: 499: 494: 489: 484: 479: 474: 469: 464: 459: 454: 449: 444: 439: 434: 429: 424: 418: 415: 414: 411: 410: 406: 405: 395: 390: 385: 380: 375: 370: 365: 358: 353: 348: 343: 337: 336: 333: 332: 329: 328: 324: 323: 318: 313: 308: 303: 301:Classification 298: 293: 288: 283: 282: 281: 271: 266: 261: 259:Common descent 256: 254:Origin of life 250: 249: 246: 245: 242: 241: 237: 236: 231: 226: 221: 216: 211: 206: 201: 196: 191: 186: 181: 176: 171: 166: 161: 156: 151: 146: 140: 139: 136: 135: 132: 131: 129: 128: 123: 118: 112: 111: 106: 101: 96: 90: 87: 86: 77: 69: 68: 62: 61: 26: 9: 6: 4: 3: 2: 7341: 7330: 7327: 7325: 7322: 7320: 7317: 7315: 7312: 7310: 7307: 7305: 7302: 7300: 7297: 7296: 7294: 7287: 7277: 7276: 7270: 7266: 7264: 7263: 7257: 7253: 7251: 7250: 7240: 7238: 7237: 7233: 7232: 7229: 7219: 7216: 7214: 7211: 7209: 7206: 7204: 7203:Neo-Darwinism 7201: 7199: 7196: 7194: 7191: 7189: 7188:Functionalism 7186: 7182: 7179: 7177: 7174: 7172: 7169: 7167: 7164: 7162: 7159: 7157: 7154: 7152: 7149: 7147: 7146:Connectionism 7144: 7142: 7139: 7138: 7137: 7136:indeterminism 7133: 7130: 7128: 7125: 7124: 7122: 7118: 7110: 7107: 7105: 7102: 7100: 7097: 7095: 7092: 7090: 7087: 7085: 7082: 7080: 7077: 7075: 7072: 7070: 7067: 7065: 7062: 7060: 7057: 7055: 7052: 7050: 7047: 7045: 7042: 7041: 7040: 7037: 7035: 7032: 7030: 7027: 7025: 7022: 7020: 7017: 7015: 7012: 7010: 7007: 7006: 7004: 7000: 6994: 6991: 6989: 6986: 6984: 6981: 6979: 6976: 6974: 6971: 6969: 6966: 6964: 6961: 6959: 6956: 6954: 6951: 6949: 6946: 6944: 6941: 6939: 6936: 6934: 6931: 6929: 6925: 6921: 6917: 6914: 6912: 6908: 6905: 6903: 6899: 6896: 6895: 6893: 6889: 6886: 6882: 6872: 6869: 6867: 6864: 6862: 6859: 6857: 6856:Schizophrenia 6854: 6852: 6849: 6847: 6844: 6842: 6841:Mental health 6839: 6837: 6834: 6832: 6829: 6827: 6824: 6822: 6819: 6817: 6814: 6812: 6809: 6807: 6804: 6802: 6799: 6797: 6794: 6792: 6789: 6788: 6786: 6784: 6780: 6774: 6771: 6769: 6766: 6764: 6760: 6756: 6753: 6751: 6747: 6744: 6742: 6739: 6737: 6734: 6732: 6729: 6727: 6724: 6722: 6719: 6717: 6716:Mate guarding 6714: 6712: 6709: 6707: 6704: 6702: 6699: 6697: 6694: 6692: 6689: 6687: 6684: 6682: 6679: 6677: 6676:Age disparity 6674: 6672: 6669: 6667: 6664: 6663: 6661: 6659: 6655: 6647: 6644: 6642: 6639: 6637: 6634: 6632: 6629: 6628: 6627: 6624: 6622: 6619: 6617: 6614: 6612: 6609: 6607: 6606:Schizophrenia 6604: 6602: 6599: 6597: 6594: 6592: 6589: 6587: 6584: 6582: 6579: 6577: 6574: 6572: 6569: 6565: 6562: 6560: 6557: 6555: 6552: 6550: 6547: 6545: 6542: 6541: 6540: 6537: 6536: 6534: 6532: 6531:Mental health 6527: 6526:Human factors 6523: 6517: 6516:Socialization 6514: 6512: 6509: 6507: 6504: 6502: 6499: 6497: 6494: 6492: 6489: 6487: 6484: 6482: 6479: 6477: 6476:paternal bond 6473: 6469: 6466: 6464: 6461: 6459: 6456: 6455: 6453: 6451: 6447: 6441: 6438: 6434: 6431: 6430: 6429: 6426: 6422: 6419: 6418: 6417: 6414: 6410: 6407: 6405: 6402: 6400: 6397: 6396: 6395: 6392: 6390: 6387: 6383: 6380: 6379: 6378: 6375: 6371: 6368: 6366: 6363: 6362: 6361: 6358: 6357: 6355: 6353: 6349: 6341: 6340:NaĂŻve physics 6338: 6336: 6333: 6331: 6328: 6327: 6326: 6323: 6321: 6318: 6316: 6313: 6311: 6307: 6306:Motor control 6304: 6300: 6297: 6295: 6292: 6291: 6290: 6287: 6285: 6281: 6278: 6276: 6272: 6269: 6265: 6264:Ophidiophobia 6262: 6260: 6257: 6255: 6254:Arachnophobia 6252: 6251: 6250: 6247: 6245: 6242: 6240: 6237: 6235: 6232: 6228: 6225: 6223: 6220: 6218: 6215: 6213: 6210: 6208: 6205: 6204: 6203: 6199: 6196: 6194: 6191: 6187: 6184: 6182: 6181:Display rules 6179: 6177: 6174: 6173: 6172: 6169: 6168: 6166: 6164: 6159: 6155: 6152: 6148: 6142: 6139: 6135: 6132: 6130: 6127: 6125: 6121: 6118: 6116: 6113: 6112: 6111: 6108: 6104: 6101: 6100: 6099: 6096: 6094: 6091: 6089: 6086: 6084: 6083:Kin selection 6081: 6077: 6074: 6073: 6072: 6069: 6067: 6064: 6062: 6059: 6057: 6054: 6052: 6049: 6045: 6042: 6040: 6037: 6035: 6032: 6031: 6030: 6027: 6025: 6022: 6021: 6019: 6017: 6011: 6005: 6002: 5998: 5995: 5993: 5990: 5988: 5985: 5983: 5980: 5978: 5977:Adaptationism 5975: 5974: 5973: 5970: 5966: 5963: 5962: 5961: 5958: 5957: 5954: 5950: 5943: 5938: 5936: 5931: 5929: 5924: 5923: 5920: 5908: 5904: 5900: 5898: 5890: 5889: 5886: 5880: 5877: 5875: 5872: 5870: 5867: 5865: 5862: 5858: 5855: 5854: 5853: 5852:Phylogenetics 5850: 5848: 5845: 5843: 5840: 5838: 5835: 5833: 5830: 5826: 5823: 5821: 5818: 5816: 5813: 5812: 5811: 5808: 5806: 5803: 5801: 5798: 5796: 5793: 5792: 5790: 5786: 5780: 5777: 5773: 5770: 5768: 5765: 5761: 5758: 5757: 5756: 5755:Structuralism 5753: 5751: 5748: 5746: 5743: 5741: 5738: 5736: 5733: 5731: 5730:Catastrophism 5728: 5727: 5726: 5723: 5721: 5718: 5717: 5715: 5711: 5705: 5702: 5700: 5697: 5695: 5692: 5690: 5689:Neo-Darwinism 5687: 5685: 5682: 5680: 5677: 5675: 5672: 5670: 5667: 5665: 5662: 5658: 5657: 5653: 5652: 5651: 5648: 5644: 5643: 5639: 5638: 5637: 5634: 5632: 5629: 5627: 5624: 5623: 5621: 5619: 5615: 5609: 5606: 5604: 5603:Reinforcement 5601: 5599: 5596: 5594: 5591: 5589: 5586: 5584: 5581: 5579: 5576: 5574: 5571: 5569: 5566: 5564: 5561: 5559: 5556: 5554: 5551: 5550: 5548: 5546: 5542: 5536: 5535:Catastrophism 5532: 5529: 5527: 5526:Macromutation 5523: 5522:Micromutation 5520: 5518: 5514: 5510: 5507: 5506: 5504: 5502: 5498: 5492: 5489: 5485: 5482: 5480: 5477: 5475: 5472: 5470: 5467: 5465: 5462: 5461: 5460: 5457: 5455: 5452: 5450: 5447: 5445: 5442: 5440: 5437: 5435: 5432: 5430: 5429:Immune system 5427: 5425: 5422: 5420: 5417: 5415: 5412: 5410: 5407: 5403: 5400: 5399: 5398: 5395: 5393: 5390: 5388: 5385: 5383: 5380: 5376: 5373: 5371: 5368: 5367: 5366: 5363: 5362: 5360: 5358: 5353: 5345: 5342: 5340: 5337: 5335: 5332: 5330: 5327: 5325: 5322: 5321: 5319: 5315: 5312: 5310: 5307: 5305: 5302: 5300: 5297: 5295: 5292: 5290: 5289:symbiogenesis 5287: 5286: 5285: 5282: 5280: 5277: 5275: 5272: 5270: 5267: 5266: 5264: 5262: 5257: 5251: 5248: 5246: 5243: 5241: 5238: 5236: 5233: 5229: 5226: 5225: 5224: 5221: 5217: 5214: 5210: 5207: 5205: 5202: 5201: 5200: 5197: 5195: 5192: 5190: 5187: 5185: 5182: 5180: 5177: 5173: 5170: 5168: 5165: 5164: 5163: 5160: 5158: 5155: 5154: 5153: 5150: 5148: 5145: 5141: 5138: 5137: 5136: 5133: 5131: 5128: 5126: 5123: 5121: 5118: 5114: 5111: 5110: 5109: 5106: 5104: 5101: 5097: 5094: 5093: 5092: 5089: 5087: 5084: 5083: 5081: 5079: 5074: 5068: 5065: 5063: 5060: 5058: 5055: 5053: 5050: 5048: 5045: 5043: 5040: 5039: 5037: 5033: 5027: 5024: 5022: 5019: 5017: 5014: 5010: 5007: 5005: 5002: 5000: 4997: 4996: 4995: 4992: 4990: 4987: 4985: 4982: 4980: 4977: 4975: 4972: 4968: 4965: 4963: 4960: 4959: 4958: 4957:Kin selection 4955: 4953: 4952:Genetic drift 4950: 4948: 4945: 4941: 4938: 4937: 4936: 4933: 4931: 4928: 4926: 4923: 4921: 4918: 4916: 4913: 4912: 4910: 4908: 4902: 4894: 4891: 4890: 4889: 4886: 4884: 4881: 4877: 4874: 4872: 4869: 4868: 4867: 4864: 4860: 4857: 4856: 4855: 4852: 4850: 4847: 4845: 4842: 4840: 4837: 4835: 4832: 4830: 4827: 4825: 4822: 4820: 4817: 4815: 4812: 4810: 4807: 4803: 4800: 4799: 4798: 4795: 4793: 4790: 4788: 4785: 4783: 4780: 4778: 4775: 4773: 4770: 4768: 4765: 4763: 4760: 4758: 4755: 4751: 4748: 4747: 4746: 4743: 4741: 4738: 4736: 4733: 4729: 4726: 4724: 4721: 4720: 4719: 4716: 4714: 4711: 4709: 4706: 4704: 4701: 4700: 4698: 4696: 4692: 4686: 4683: 4681: 4678: 4676: 4673: 4671: 4668: 4666: 4663: 4662: 4659: 4655: 4648: 4643: 4641: 4636: 4634: 4629: 4628: 4625: 4613: 4612: 4603: 4601: 4600: 4595: 4591: 4589: 4588: 4579: 4577: 4576: 4567: 4566: 4563: 4556: 4555:Chronospecies 4552: 4548: 4544: 4540: 4537: 4535: 4532: 4530: 4527: 4525: 4522: 4520: 4517: 4516: 4514: 4510: 4504: 4501: 4499: 4496: 4493: 4489: 4488:Reinforcement 4486: 4483: 4482:Recombination 4479: 4475: 4471: 4468: 4467: 4465: 4463: 4459: 4453: 4450: 4448: 4445: 4443: 4440: 4437: 4433: 4429: 4426: 4424: 4421: 4419: 4416: 4414: 4411: 4410: 4408: 4406: 4402: 4396: 4393: 4390: 4386: 4382: 4379: 4376: 4372: 4368: 4364: 4360: 4357: 4356: 4354: 4350: 4344: 4341: 4339: 4336: 4334: 4331: 4329: 4326: 4324: 4321: 4318: 4314: 4311: 4310: 4308: 4304: 4298: 4295: 4293: 4290: 4288: 4285: 4283: 4280: 4279: 4276: 4272: 4265: 4260: 4258: 4253: 4251: 4246: 4245: 4242: 4236: 4232: 4229: 4226: 4223: 4220: 4217: 4215: 4211: 4208: 4205: 4202: 4199: 4198: 4194: 4193: 4182: 4178: 4174: 4171: 4167: 4163: 4157: 4150: 4146: 4140: 4136: 4132: 4128: 4123: 4118: 4114: 4110: 4107:(5863): 588. 4106: 4102: 4101: 4093: 4085: 4083:9789027295118 4079: 4075: 4071: 4067: 4060: 4053: 4047: 4040: 4036: 4032: 4028: 4024: 4020: 4016: 4012: 4008: 4004: 4000: 3996: 3992: 3986: 3980: 3976: 3970: 3964: 3960: 3954: 3947: 3943: 3937: 3930: 3926: 3920: 3913: 3912: 3908: 3904: 3901: 3895: 3893: 3891: 3889: 3881: 3880: 3873: 3866: 3865: 3858: 3852: 3848: 3844: 3839: 3831: 3830:New Scientist 3827: 3820: 3806: 3802: 3795: 3787: 3783: 3779: 3775: 3768: 3760: 3756: 3752: 3748: 3744: 3740: 3733: 3731: 3715: 3711: 3705: 3698: 3695: 3691: 3688: 3684: 3679: 3672: 3671: 3664: 3657: 3656: 3649: 3642: 3641: 3634: 3627: 3626: 3621: 3616: 3610:, p. 250-251. 3609: 3608: 3601: 3593: 3589: 3585: 3581: 3577: 3573: 3569: 3565: 3559: 3552: 3551: 3544: 3535: 3528: 3527: 3522: 3517: 3510: 3505: 3498: 3497: 3490: 3483: 3482: 3477: 3472: 3465: 3459: 3448: 3444: 3440: 3436: 3432: 3428: 3424: 3420: 3416: 3409: 3402: 3396: 3392: 3391: 3384: 3377: 3373: 3370: 3369: 3362: 3355: 3353: 3346: 3339: 3338: 3331: 3329: 3321: 3317: 3312: 3298:on 2018-09-23 3294: 3290: 3286: 3282: 3278: 3274: 3270: 3263: 3256: 3248: 3247: 3242: 3238: 3232: 3224: 3222:9781418515096 3218: 3214: 3210: 3209: 3204: 3198: 3188: 3186:9780684824710 3182: 3178: 3174: 3173: 3167: 3163: 3161:9780393315707 3157: 3153: 3149: 3148: 3143: 3139: 3135: 3131: 3127: 3126: 3124: 3123:For reply see 3119: 3111: 3107: 3100: 3089: 3085: 3081: 3074: 3070: 3064: 3053: 3049: 3042: 3041: 3033: 3025: 3024: 3019: 3015: 3009: 3002: 2998: 2994: 2991: 2987: 2982: 2973: 2968: 2964: 2960: 2957:(1–4): 1–20. 2956: 2952: 2948: 2941: 2934: 2933: 2926: 2920: 2916: 2915: 2910: 2905: 2897: 2893: 2889: 2885: 2878: 2870: 2866: 2861: 2856: 2852: 2848: 2844: 2837: 2829: 2825: 2819: 2813: 2811: 2808: 2801: 2793: 2789: 2785: 2781: 2777: 2773: 2769: 2765: 2758: 2751: 2747: 2742: 2734: 2730: 2726: 2722: 2718: 2714: 2710: 2706: 2702: 2696: 2687: 2682: 2678: 2674: 2670: 2666: 2662: 2655: 2653: 2645: 2639: 2631: 2625: 2621: 2617: 2613: 2609: 2605: 2599: 2592: 2588: 2587: 2582: 2577: 2569: 2565: 2561: 2557: 2553: 2549: 2548: 2540: 2533: 2530: 2524: 2517: 2514: 2508: 2501: 2498: 2494: 2490: 2485: 2478: 2474: 2470: 2465: 2463: 2454: 2450: 2446: 2442: 2438: 2434: 2430: 2426: 2422: 2418: 2414: 2407: 2400: 2396: 2392: 2389: 2383: 2375: 2369: 2361: 2357: 2353: 2349: 2345: 2341: 2334: 2327: 2326:1-77007-148-2 2323: 2319: 2313: 2306: 2303: 2300: 2296: 2293: 2289: 2285: 2280: 2278: 2270: 2264: 2257: 2251: 2243: 2239: 2235: 2231: 2227: 2223: 2216: 2209: 2205: 2204: 2199: 2194: 2187: 2181: 2174: 2168: 2166: 2159: 2155: 2149: 2147: 2138: 2134: 2130: 2126: 2119: 2112: 2105: 2102:. Princeton: 2101: 2097: 2092: 2085: 2082: 2078: 2073: 2071: 2062: 2056: 2052: 2047: 2046: 2040: 2034: 2027: 2021: 2014: 2010: 2004: 1997: 1993: 1987: 1981: 1977: 1971: 1964: 1960: 1956: 1951: 1949: 1947: 1945: 1936: 1932: 1928: 1924: 1920: 1916: 1912: 1908: 1904: 1900: 1899: 1894: 1892: 1886: 1880: 1872: 1868: 1864: 1860: 1856: 1850: 1842: 1838: 1834: 1830: 1826: 1822: 1818: 1814: 1810: 1806: 1805: 1800: 1794: 1792: 1783: 1779: 1773: 1765: 1761: 1754: 1752: 1750: 1748: 1739: 1738: 1737:New Scientist 1733: 1729: 1723: 1721: 1710: 1697: 1688: 1683: 1678: 1673: 1669: 1664: 1660: 1647: 1638: 1633: 1628: 1623: 1619: 1614: 1613: 1604:on 2020-01-13 1600: 1593: 1592: 1585: 1580: 1579: 1576: 1572: 1568: 1564: 1558: 1556: 1554: 1552: 1547: 1537: 1534: 1532: 1529: 1527: 1524: 1522: 1519: 1517: 1514: 1512: 1509: 1507: 1504: 1502: 1499: 1497: 1494: 1492: 1489: 1487: 1484: 1483: 1476: 1468: 1466: 1462: 1457: 1455: 1451: 1447: 1441: 1431: 1429: 1428: 1423: 1418: 1414: 1408: 1398: 1394: 1389: 1387: 1383: 1378: 1376: 1371: 1368: 1364: 1363: 1358: 1353: 1352: 1346: 1344: 1340: 1339:Charles Lyell 1336: 1332: 1328: 1324: 1323:catastrophism 1320: 1316: 1312: 1302: 1300: 1295: 1293: 1288: 1283: 1279: 1277: 1276: 1271: 1270: 1264: 1260: 1259: 1253: 1249: 1246: 1241: 1237: 1233: 1223: 1220: 1216: 1212: 1208: 1204: 1203: 1198: 1194: 1192: 1188: 1184: 1180: 1170: 1168: 1167:Sewall Wright 1163: 1159: 1155: 1154:genetic drift 1151: 1147: 1143: 1139: 1135: 1131: 1127: 1123: 1114: 1108: 1104: 1100: 1099: 1098: 1095: 1093: 1089: 1085: 1081: 1077: 1069: 1068:macromutation 1064: 1055: 1053: 1049: 1045: 1034: 1032: 1028: 1024: 1014: 1012: 1011: 1006: 1002: 997: 992: 990: 989: 984: 979: 977: 973: 969: 965: 955: 953: 949: 943: 938: 936: 931: 927: 922: 918: 913: 911: 907: 892: 890: 889: 884: 879: 878:meta-analysis 875: 871: 864: 854: 852: 847: 844: 840: 835: 831: 826: 824: 820: 819: 814: 809: 806: 805:morphological 802: 798: 794: 790: 786: 782: 772: 770: 766: 762: 758: 754: 753:developmental 750: 746: 742: 738: 734: 730: 725: 723: 719: 714: 712: 708: 704: 700: 696: 695:fossil record 692: 688: 684: 681:(also called 680: 676: 664: 659: 657: 652: 650: 645: 644: 642: 641: 635: 625: 622: 617: 611: 610: 609: 608: 601: 598: 596: 593: 591: 588: 586: 583: 581: 578: 576: 573: 571: 568: 566: 563: 561: 558: 557: 551: 550: 543: 540: 538: 535: 533: 530: 528: 525: 523: 520: 518: 515: 513: 510: 508: 507:Phylogenetics 505: 503: 500: 498: 495: 493: 490: 488: 485: 483: 480: 478: 475: 473: 470: 468: 465: 463: 460: 458: 455: 453: 450: 448: 445: 443: 440: 438: 435: 433: 430: 428: 425: 423: 420: 419: 413: 412: 403: 399: 396: 394: 391: 389: 386: 384: 381: 379: 376: 374: 371: 369: 366: 364: 363: 359: 357: 354: 352: 351:Before Darwin 349: 347: 344: 342: 339: 338: 331: 330: 322: 319: 317: 314: 312: 309: 307: 304: 302: 299: 297: 294: 292: 289: 287: 284: 280: 277: 276: 275: 272: 270: 267: 265: 262: 260: 257: 255: 252: 251: 244: 243: 235: 232: 230: 227: 225: 222: 220: 217: 215: 212: 210: 207: 205: 202: 200: 197: 195: 192: 190: 187: 185: 182: 180: 179:Genetic drift 177: 175: 172: 170: 167: 165: 162: 160: 157: 155: 152: 150: 147: 145: 142: 141: 134: 133: 127: 124: 122: 119: 117: 114: 113: 110: 107: 105: 102: 100: 97: 95: 92: 91: 89: 88: 84: 80: 75: 71: 70: 67: 64: 63: 59: 58: 52: 48: 47:morphological 43: 37: 33: 19: 7286: 7273: 7260: 7247: 7234: 7083: 6993:Sociobiology 6851:Neuroscience 6831:Intelligence 6377:Anthropology 6330:Color vision 6315:Multitasking 6294:Flynn effect 6289:Intelligence 6271:Folk biology 6014:Evolutionary 5864:Polymorphism 5847:Astrobiology 5795:Biogeography 5750:Saltationism 5740:Orthogenesis 5725:Alternatives 5654: 5640: 5573:Cospeciation 5568:Cladogenesis 5517:Saltationism 5512: 5474:Mating types 5397:Color vision 5382:Avian flight 5304:mitochondria 5042:Canalisation 4920:Biodiversity 4665:Introduction 4609: 4597: 4585: 4573: 4546: 4389:Ring species 4338:Cospeciation 4333:Cladogenesis 4282:Introduction 4234: 4196: 4180: 4172: 4168:, 15, 2012; 4165: 4156: 4148: 4104: 4098: 4092: 4065: 4059: 4051: 4046: 4006: 4002: 3985: 3979:pp. 121-122. 3974: 3969: 3963:pp. 120-121. 3958: 3953: 3941: 3936: 3924: 3919: 3909: 3877: 3872: 3862: 3857: 3846: 3843:Gould, S. J. 3838: 3829: 3819: 3808:. Retrieved 3804: 3794: 3777: 3773: 3767: 3742: 3738: 3718:. Retrieved 3714:the original 3704: 3696: 3689: 3683:Gould, S. J. 3678: 3668: 3663: 3653: 3648: 3638: 3633: 3623: 3615: 3605: 3600: 3575: 3571: 3558: 3548: 3543: 3534: 3524: 3516: 3504: 3494: 3489: 3479: 3471: 3463: 3458: 3447:the original 3418: 3414: 3401: 3395:pp. 205-206. 3388: 3383: 3367: 3361: 3351: 3345: 3335: 3319: 3311: 3300:. Retrieved 3293:the original 3272: 3268: 3255: 3244: 3239:(May 1981). 3231: 3207: 3197: 3171: 3146: 3133: 3122: 3118: 3105: 3099: 3088:the original 3083: 3079: 3063: 3052:the original 3039: 3032: 3021: 3008: 3000: 2981: 2954: 2950: 2940: 2930: 2925: 2912: 2904: 2887: 2883: 2877: 2850: 2846: 2836: 2827: 2818: 2809: 2806: 2800: 2770:(1): 15–35. 2767: 2763: 2757: 2749: 2741: 2708: 2704: 2695: 2668: 2664: 2643: 2638: 2611: 2608:Gould, S. J. 2598: 2584: 2576: 2551: 2545: 2539: 2531: 2523: 2515: 2507: 2499: 2484: 2472: 2420: 2416: 2406: 2398: 2382: 2368:cite journal 2343: 2333: 2317: 2312: 2304: 2302:Paleobiology 2301: 2268: 2263: 2255: 2250: 2225: 2221: 2215: 2201: 2193: 2185: 2180: 2172: 2158:pp. 123-124. 2153: 2128: 2124: 2111: 2099: 2091: 2083: 2044: 2039:Gould, S. J. 2033: 2025: 2020: 2012: 2003: 1995: 1986: 1975: 1970: 1962: 1902: 1896: 1890: 1879: 1870: 1866: 1862: 1858: 1849: 1808: 1802: 1781: 1778:Lerner, I.M. 1772: 1763: 1735: 1696:cite journal 1646:cite journal 1606:. Retrieved 1599:the original 1590: 1584:Reprinted in 1583: 1574: 1474: 1458: 1454:Pama–Nyungan 1443: 1425: 1410: 1396: 1391: 1385: 1381: 1379: 1374: 1372: 1366: 1360: 1356: 1349: 1347: 1342: 1308: 1296: 1284: 1280: 1273: 1267: 1256: 1255:In his book 1254: 1250: 1243: 1218: 1214: 1206: 1200: 1195: 1176: 1161: 1120: 1112: 1096: 1088:creationists 1073: 1058:Saltationism 1044:saltationism 1040: 1020: 1008: 996:Kim Sterelny 993: 986: 980: 961: 945: 940: 926:homogenizing 914: 909: 903: 886: 873: 866: 848: 842: 838: 827: 816: 810: 778: 749:I. M. Lerner 743:'s model of 736: 726: 715: 711:cladogenesis 702: 682: 678: 672: 527:Sociobiology 512:Paleontology 360: 296:Biogeography 291:Biodiversity 209:Coextinction 199:Co-operation 174:Polymorphism 99:Introduction 7132:Determinism 7044:Coevolution 6988:Primatology 6826:Gender role 6731:Orientation 6611:Screen time 6468:Affectional 6450:Development 6129:Mate choice 6056:By-products 6024:Adaptations 5987:Cognitivism 5874:Systematics 5745:Mutationism 5563:Catagenesis 5491:Snake venom 5424:Eusociality 5402:in primates 5392:Cooperation 5320:In animals 5140:butterflies 5113:Cephalopods 5103:Brachiopods 5035:Development 5009:Mate choice 4762:Convergence 4745:Coevolution 4703:Abiogenesis 4611:WikiProject 4371:Centrifugal 3316:S. J. Gould 3237:Gould, S.J. 3142:Dawkins, R. 3130:Gould, S.J. 3014:Gould, S.J. 2986:S. J. Gould 2909:S. J. Gould 2644:Time Frames 2581:S. J. Gould 2493:S. J. Gould 2469:Mayr, Ernst 2284:S. J. Gould 2198:S. J. Gould 2077:S. J. Gould 1955:Mayr, Ernst 1855:Gould, S.J. 1728:Gould, S.J. 1591:Time Frames 1567:Gould, S.J. 1526:Koinophilia 1521:Gene orders 1150:synergistic 1042:ideas like 976:koinophilia 843:Time Frames 537:Systematics 346:Renaissance 224:Convergence 214:Contingency 204:Coevolution 7324:Speciation 7293:Categories 7079:Population 7074:Lamarckism 6920:behavioral 6898:Behavioral 6846:Narcissism 6791:Aggression 6581:Hypophobia 6571:Depression 6458:Attachment 6440:Universals 6404:Psychology 6382:Biological 6370:Musicology 6360:Aesthetics 6259:Basophobia 6066:Exaptation 6044:Reciprocal 5735:Lamarckism 5713:Philosophy 5636:David Hume 5598:Peripatric 5593:Parapatric 5578:Ecological 5558:Anagenesis 5553:Allopatric 5545:Speciation 5509:Gradualism 5434:Metabolism 5294:chromosome 5284:Eukaryotes 5062:Modularity 4979:Population 4905:Population 4866:Speciation 4844:Panspermia 4797:Extinction 4792:Exaptation 4767:Divergence 4740:Cladistics 4728:Reciprocal 4708:Adaptation 4474:Polyploidy 4436:Allochrony 4413:Adaptation 4381:Parapatric 4363:Peripatric 4359:Allopatric 4328:Anagenesis 4271:Speciation 4122:1885/33371 3810:2023-02-26 3720:2022-02-16 3302:2007-04-22 3275:(1): 104. 2228:(2): 212. 2081:"Opus 200" 1873:: 407–532. 1608:2018-01-27 1543:References 1486:Speciation 1438:See also: 1405:See also: 1335:gradualism 1230:See also: 1179:gradualism 1080:geneticist 1001:phenotypic 935:hypothesis 789:allopatric 781:Ernst Mayr 741:Ernst Mayr 722:anagenesis 311:Cladistics 234:Extinction 219:Divergence 189:Speciation 169:Adaptation 83:John Gould 6924:cognitive 6916:Affective 6801:Cognition 6755:Sexuality 6741:Pair bond 6501:Education 6158:Cognition 6076:Inclusive 6016:processes 6004:Criticism 5869:Protocell 5720:Darwinism 5608:Sympatric 5357:processes 5245:Tetrapods 5194:Kangaroos 5120:Dinosaurs 5057:Inversion 5026:Variation 4947:Gene flow 4940:Inclusive 4750:Mutualism 4695:Evolution 4395:Sympatric 4175:1, 2013; 3759:144947503 3658:, p. 251. 3643:, p. 250. 3628:, p. 241. 3572:Evolution 3553:, p. 240. 3529:, p. 245. 3415:Evolution 3134:Structure 2869:213493738 2812:, 401-409 2705:Evolution 2360:2315-7712 2173:Evolution 2106:, p. 120. 1825:2246/6568 1804:Evolution 1687:2246/6567 1637:2246/6564 1471:Mythology 1331:evolution 1226:Criticism 917:gene flow 818:Evolution 771:species. 570:Dysgenics 286:Phylogeny 184:Gene flow 154:Diversity 149:Variation 7193:Memetics 6953:Ethology 6911:genetics 6746:Physical 6711:Jealousy 6666:Activity 6472:maternal 6428:Religion 6416:Morality 6394:Language 6275:taxonomy 6088:Mismatch 6034:Cheating 6029:Altruism 5897:Category 5772:Vitalism 5767:Theistic 5760:Spandrel 5444:Morality 5439:Monogamy 5314:plastids 5279:Flagella 5235:Reptiles 5216:sea cows 5199:primates 5108:Molluscs 5086:Bacteria 4974:Mutation 4907:genetics 4883:Taxonomy 4829:Mismatch 4809:Homology 4723:Cheating 4718:Altruism 4575:Category 4492:evidence 4297:Glossary 4235:Newsweek 4149:PLoS ONE 4139:29740420 4131:18239118 3997:(1993), 3903:Archived 3685:(1997). 3622:(1996). 3478:(1996). 3443:28568042 3372:Archived 3246:Discover 3205:(1998). 3144:(1996). 3132:(2002). 3071:(1983). 2993:Archived 2826:(1989). 2748:(1992). 2733:28555824 2568:83546424 2495:(1997). 2471:(1963). 2453:38248823 2445:24653037 2391:Archived 2295:Archived 2290:(1977). 2286:, & 2137:19203015 2079:(1991). 2041:(2002). 1957:(1992). 1935:32953263 1887:(1983). 1841:28562952 1780:(1954). 1569:(1972). 1479:See also 1134:families 964:putative 912:(1963). 874:horotely 634:Category 560:Eugenics 402:timeline 383:Evo-devo 341:Overview 159:Mutation 121:Evidence 116:Glossary 7094:Species 6866:Suicide 6701:Fantasy 6681:Arousal 6463:Bonding 6352:Culture 6176:Display 6163:Emotion 6071:Fitness 5960:History 5788:Related 5618:History 5479:Meiosis 5414:Empathy 5409:Emotion 5309:nucleus 5250:Viruses 5240:Spiders 5152:Mammals 5135:Insects 4935:Fitness 4871:Species 4670:Outline 4587:Commons 4539:Fossils 4529:Insects 4478:Klepton 4367:Quantum 4313:Species 4287:History 4100:Science 4039:4253816 4031:8232582 4011:Bibcode 3592:2410098 3523:(1996) 3435:2408092 3318:(2007) 3289:4293004 3177:288–289 3152:230-236 2988:(1992) 2959:Bibcode 2792:2200930 2772:Bibcode 2725:2408397 2673:Bibcode 2532:Science 2500:Science 2425:Bibcode 2417:Science 2269:Biology 2230:Bibcode 2098:(1985) 1980:97-116. 1927:6353241 1907:Bibcode 1833:2406508 1362:Lingula 1357:Lingula 1311:species 1146:species 1142:classes 1027:classes 948:viruses 775:History 691:species 685:) is a 126:History 109:Outline 7272:  7259:  7246:  6836:Memory 6796:Autism 6763:female 6696:Desire 6433:Origin 6409:Speech 6399:Origin 6171:Affect 5907:Portal 5583:Hybrid 5419:Ethics 5261:organs 5223:Plants 5209:lemurs 5204:humans 5189:horses 5179:hyenas 5167:wolves 5162:canids 5096:origin 4534:Plants 4385:Clines 4137:  4129:  4080:  4037:  4029:  4003:Nature 3946:p. 551 3929:p. 313 3757:  3590:  3441:  3433:  3287:  3219:  3183:  3158:  2911:2002. 2867:  2790:  2731:  2723:  2626:  2583:2002. 2566:  2451:  2443:  2358:  2324:  2200:2002. 2135:  2057:  1933:  1925:  1898:Nature 1839:  1831:  1386:Origin 1238:, and 1140:, and 1138:orders 1050:, and 1003:mean. 988:Amalda 958:Stasis 769:fossil 703:stasis 687:theory 632:  356:Darwin 6806:Crime 6389:Crime 6320:Sleep 6310:skill 6150:Areas 5370:Death 5365:Aging 5344:brain 5130:Fungi 5091:Birds 5004:Fungi 4802:Event 4685:Index 4519:Birds 4135:S2CID 4035:S2CID 3851:p. 6. 3755:S2CID 3588:JSTOR 3450:(PDF) 3431:JSTOR 3411:(PDF) 3296:(PDF) 3285:S2CID 3265:(PDF) 3213:40-45 3091:(PDF) 3076:(PDF) 3055:(PDF) 3044:(PDF) 2865:S2CID 2721:JSTOR 2564:S2CID 2449:S2CID 2121:(PDF) 1931:S2CID 1829:JSTOR 1602:(PDF) 1595:(PDF) 1382:Essay 1343:Essay 94:Index 6759:male 6120:Male 5857:Tree 5329:hair 5269:Cell 5172:dogs 5157:cats 5147:Life 5125:Fish 5078:taxa 4524:Fish 4127:PMID 4078:ISBN 4027:PMID 3439:PMID 3217:ISBN 3181:ISBN 3156:ISBN 2919:872. 2788:PMID 2729:PMID 2624:ISBN 2491:and 2441:PMID 2374:link 2356:ISSN 2322:ISBN 2133:PMID 2055:ISBN 1923:PMID 1837:PMID 1709:help 1659:help 1325:and 755:and 731:and 104:Main 6658:Sex 6335:Eye 5355:Of 5324:eye 5274:DNA 5259:Of 5076:Of 4117:hdl 4109:doi 4105:319 4070:doi 4019:doi 4007:366 3782:doi 3747:doi 3580:doi 3423:doi 3277:doi 3110:216 3048:617 2967:doi 2955:127 2892:doi 2855:doi 2810:177 2780:doi 2768:144 2713:doi 2681:doi 2669:127 2616:doi 2556:doi 2552:130 2433:doi 2421:343 2348:doi 2238:doi 2226:101 2208:875 2051:775 2011:In 1915:doi 1903:305 1871:138 1821:hdl 1813:doi 1682:hdl 1672:doi 1632:hdl 1622:doi 1007:'s 787:by 724:). 673:In 81:by 7295:: 4553:· 4549:· 4545:· 4480:· 4476:· 4434:· 4387:· 4373:· 4369:· 4365:· 4179:. 4164:. 4147:, 4133:. 4125:. 4115:. 4103:. 4076:. 4033:, 4025:, 4017:, 4005:, 4001:, 3993:; 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Index

Stasis (biology)
Punctuated equilibrium in social theory
Punctuated Equilibrium (album)

morphological
phyletic gradualism
Evolutionary biology

Darwin's finches
John Gould
Index
Introduction
Main
Outline
Glossary
Evidence
History
Population genetics
Variation
Diversity
Mutation
Natural selection
Adaptation
Polymorphism
Genetic drift
Gene flow
Speciation
Adaptive radiation
Co-operation
Coevolution

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