104:
1341:
122:
1347:
1352:
1373:
1359:
415:
291:, the head shield and trunk tergites were fused into a single plate. There are about 19 furrows indicating the original trunk segments. a small anterior sclerite articulated to the front of its head. A pair of
475:
346:
Xianguang, Hou; Ramskold, Lars; Bergstrom, Jan (October 1991). "Composition and preservation of the
Chengjiang fauna -a Lower Cambrian soft-bodied biota".
850:
895:
676:
1438:
923:
874:
1250:
1086:
978:
930:
866:
797:
192:
1259:
992:
971:
902:
783:
768:
660:
1006:
950:
858:
790:
1075:
916:
720:
1287:
1013:
999:
985:
281:
1130:
804:
1425:
776:
398:
538:
317:
1340:
103:
121:
531:
1469:
1351:
468:
1492:
1393:
843:
1443:
1464:
1456:
1358:
427:
295:
and lateral eyes located ventrally behind the anterior sclerite, lateral to the hypostome.
1346:
8:
1316:
431:
1497:
1376:
524:
497:
470:
451:
371:
359:
268:
116:
1451:
502:
443:
394:
363:
299:
reached 151 mm in length and had nearly 25 pairs of biramous limbs for walking.
292:
455:
375:
886:
492:
484:
435:
355:
263:
stage, which lasted from 521 to 514 million years ago during the early part of the
1416:
1033:
836:
669:
264:
1109:
1061:
699:
620:
439:
416:"Relationships of Cambrian Arachnata and the systematic position of Trilobita"
1486:
1102:
752:
574:
447:
367:
61:
1150:
1054:
941:
761:
506:
488:
218:
1047:
964:
828:
815:
713:
469:
Nicholas J. Minter, M. Gabriela Mángano & Jean-Bernard Caron (2011).
36:
1430:
1186:
1177:
1166:
1141:
1123:
1116:
1095:
1068:
909:
634:
260:
81:
46:
1305:
1294:
1235:
1228:
1213:
741:
727:
641:
613:
595:
580:
565:
547:
287:
256:
252:
179:
166:
153:
133:
86:
1387:
1410:
1266:
1204:
1159:
957:
734:
683:
648:
76:
71:
56:
51:
41:
391:
Arthropods of the Lower
Cambrian Chengjiang fauna, southwest China
1273:
1193:
627:
245:
91:
66:
559:
318:"A compendium of fossil marine animal genera (Trilobita entry)"
272:
143:
471:"Skimming the surface with Burgess Shale arthropod locomotion"
1221:
276:
248:
516:
345:
476:
Proceedings of the Royal
Society B: Biological Sciences
393:. Fossils and Strata. Oslo: Scandinavian Univ. Press.
389:
Hou, Xianguang; Bergstöm, Jan; Bergström, Jan (1997).
414:
Edgecombe, Gregory D.; Ramsköld, Lars (March 1999).
388:
413:
1484:
532:
539:
525:
102:
496:
593:
315:
309:
1485:
1392:
1391:
520:
232:Hou, Ramsköld & Bergström, 1991
212:Hou, Ramsköld & Bergström, 1991
13:
360:10.1111/j.1463-6409.1991.tb00303.x
322:Bulletins of American Paleontology
14:
1509:
279:. Similar to the closely-related
1372:
1371:
1357:
1350:
1345:
1339:
120:
34:
462:
407:
382:
339:
1:
546:
302:
111:diagrammatic reconstruction
7:
259:. It lived during the late
10:
1514:
16:Extinct genus of arthropod
1400:
1367:
1337:
1304:
1249:
1203:
1176:
1149:
1140:
1085:
1042:
1031:
940:
885:
823:
814:
751:
708:
697:
659:
608:
604:
589:
554:
440:10.1017/S0022336000027761
224:
217:
117:Scientific classification
115:
110:
101:
23:
420:Journal of Paleontology
316:Sepkoski, Jack (2002).
489:10.1098/rspb.2011.1986
267:. It was found in the
1465:Paleobiology Database
483:(1733): 1613–1620.
432:1999JPal...73..263E
269:Maotianshan Shales
227:Saperion glumaceum
1480:
1479:
1452:Open Tree of Life
1394:Taxon identifiers
1385:
1384:
1335:
1334:
1331:
1330:
1327:
1326:
1245:
1244:
1027:
1026:
1023:
1022:
693:
692:
400:978-82-00-37693-4
348:Zoologica Scripta
237:
236:
213:
1505:
1473:
1472:
1460:
1459:
1447:
1446:
1434:
1433:
1421:
1420:
1419:
1389:
1388:
1375:
1374:
1362:
1361:
1354:
1349:
1343:
1147:
1146:
1040:
1039:
887:Tremaglaspididae
821:
820:
706:
705:
606:
605:
602:
601:
591:
590:
541:
534:
527:
518:
517:
511:
510:
500:
466:
460:
459:
411:
405:
404:
386:
380:
379:
343:
337:
336:
334:
333:
313:
211:
204:
191:
178:
165:
125:
124:
106:
96:
33:
32:Cambrian Stage 3
29:Temporal range:
21:
20:
1513:
1512:
1508:
1507:
1506:
1504:
1503:
1502:
1483:
1482:
1481:
1476:
1468:
1463:
1455:
1450:
1442:
1437:
1429:
1424:
1415:
1414:
1409:
1396:
1386:
1381:
1363:
1356:
1323:
1300:
1241:
1199:
1172:
1136:
1081:
1036:
1034:Trilobitomorpha
1019:
936:
881:
851:Chacharejocaris
837:Australaglaspis
810:
747:
702:
689:
670:Acanthomeridion
655:
598:
585:
550:
545:
515:
514:
467:
463:
412:
408:
401:
387:
383:
344:
340:
331:
329:
314:
310:
305:
265:Cambrian Period
233:
230:
210:
202:
189:
180:Trilobitomorpha
176:
163:
119:
97:
95:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
31:
30:
27:
17:
12:
11:
5:
1511:
1501:
1500:
1495:
1478:
1477:
1475:
1474:
1461:
1448:
1435:
1422:
1406:
1404:
1398:
1397:
1383:
1382:
1380:
1379:
1368:
1365:
1364:
1338:
1336:
1333:
1332:
1329:
1328:
1325:
1324:
1322:
1321:
1320:
1319:
1310:
1308:
1302:
1301:
1299:
1298:
1291:
1284:
1277:
1270:
1263:
1255:
1253:
1247:
1246:
1243:
1242:
1240:
1239:
1232:
1225:
1218:
1209:
1207:
1201:
1200:
1198:
1197:
1190:
1182:
1180:
1174:
1173:
1171:
1170:
1163:
1155:
1153:
1144:
1138:
1137:
1135:
1134:
1127:
1120:
1113:
1110:Phytophilaspis
1106:
1099:
1091:
1089:
1083:
1082:
1080:
1079:
1072:
1065:
1062:Pygmaclypeatus
1058:
1051:
1043:
1037:
1032:
1029:
1028:
1025:
1024:
1021:
1020:
1018:
1017:
1010:
1003:
996:
989:
982:
975:
968:
961:
954:
946:
944:
938:
937:
935:
934:
927:
920:
913:
906:
899:
896:Brachyaglaspis
891:
889:
883:
882:
880:
879:
871:
863:
855:
847:
840:
833:
824:
818:
812:
811:
809:
808:
801:
794:
787:
780:
773:
765:
757:
755:
749:
748:
746:
745:
738:
731:
724:
717:
709:
703:
700:Vicissicaudata
698:
695:
694:
691:
690:
688:
687:
680:
677:Australimicola
673:
665:
663:
657:
656:
654:
653:
645:
638:
631:
624:
621:Falcatamacaris
617:
609:
599:
594:
587:
586:
584:
583:
577:
568:
562:
555:
552:
551:
544:
543:
536:
529:
521:
513:
512:
461:
426:(2): 263–287.
406:
399:
381:
354:(4): 395–411.
338:
307:
306:
304:
301:
235:
234:
231:
222:
221:
215:
214:
200:
196:
195:
187:
183:
182:
174:
170:
169:
161:
157:
156:
151:
147:
146:
141:
137:
136:
131:
127:
126:
113:
112:
108:
107:
99:
98:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
35:
28:
15:
9:
6:
4:
3:
2:
1510:
1499:
1496:
1494:
1493:Cambrian life
1491:
1490:
1488:
1471:
1466:
1462:
1458:
1453:
1449:
1445:
1440:
1436:
1432:
1427:
1423:
1418:
1412:
1408:
1407:
1405:
1403:
1399:
1395:
1390:
1378:
1370:
1369:
1366:
1360:
1355:
1353:
1348:
1342:
1318:
1314:
1313:
1312:
1311:
1309:
1307:
1303:
1297:
1296:
1292:
1290:
1289:
1285:
1283:
1282:
1278:
1276:
1275:
1271:
1269:
1268:
1264:
1262:
1261:
1257:
1256:
1254:
1252:
1248:
1238:
1237:
1233:
1231:
1230:
1226:
1224:
1223:
1219:
1216:
1215:
1211:
1210:
1208:
1206:
1202:
1196:
1195:
1191:
1189:
1188:
1184:
1183:
1181:
1179:
1175:
1169:
1168:
1164:
1162:
1161:
1157:
1156:
1154:
1152:
1148:
1145:
1143:
1139:
1133:
1132:
1128:
1126:
1125:
1121:
1119:
1118:
1114:
1112:
1111:
1107:
1105:
1104:
1103:Luohuilinella
1100:
1098:
1097:
1093:
1092:
1090:
1088:
1084:
1078:
1077:
1073:
1071:
1070:
1066:
1064:
1063:
1059:
1057:
1056:
1052:
1050:
1049:
1045:
1044:
1041:
1038:
1035:
1030:
1016:
1015:
1011:
1009:
1008:
1004:
1002:
1001:
997:
995:
994:
990:
988:
987:
983:
981:
980:
976:
974:
973:
969:
967:
966:
962:
960:
959:
955:
953:
952:
948:
947:
945:
943:
939:
933:
932:
928:
926:
925:
924:Quasimodaspis
921:
919:
918:
914:
912:
911:
907:
905:
904:
900:
898:
897:
893:
892:
890:
888:
884:
877:
876:
875:Rozhkovocaris
872:
869:
868:
864:
861:
860:
856:
853:
852:
848:
846:
845:
841:
839:
838:
834:
831:
830:
826:
825:
822:
819:
817:
813:
807:
806:
802:
800:
799:
795:
793:
792:
788:
786:
785:
781:
779:
778:
774:
771:
770:
766:
764:
763:
759:
758:
756:
754:
753:Cheloniellida
750:
744:
743:
739:
737:
736:
732:
730:
729:
725:
723:
722:
718:
716:
715:
711:
710:
707:
704:
701:
696:
686:
685:
681:
679:
678:
674:
672:
671:
667:
666:
664:
662:
658:
651:
650:
646:
644:
643:
639:
637:
636:
632:
630:
629:
625:
623:
622:
618:
616:
615:
611:
610:
607:
603:
600:
597:
592:
588:
582:
578:
576:
575:Panarthropoda
572:
569:
567:
564:Superphylum:
563:
561:
557:
556:
553:
549:
542:
537:
535:
530:
528:
523:
522:
519:
508:
504:
499:
494:
490:
486:
482:
478:
477:
472:
465:
457:
453:
449:
445:
441:
437:
433:
429:
425:
421:
417:
410:
402:
396:
392:
385:
377:
373:
369:
365:
361:
357:
353:
349:
342:
327:
323:
319:
312:
308:
300:
298:
294:
290:
289:
284:
283:
278:
274:
270:
266:
262:
258:
254:
250:
247:
243:
242:
229:
228:
223:
220:
216:
209:
208:
201:
198:
197:
194:
188:
185:
184:
181:
175:
172:
171:
168:
162:
159:
158:
155:
152:
149:
148:
145:
142:
139:
138:
135:
132:
129:
128:
123:
118:
114:
109:
105:
100:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
26:
22:
19:
1401:
1344:
1293:
1286:
1280:
1279:
1272:
1265:
1258:
1251:Conciliterga
1234:
1227:
1220:
1212:
1192:
1185:
1165:
1158:
1151:Emucarididae
1129:
1122:
1115:
1108:
1101:
1094:
1087:Xandarellida
1074:
1067:
1060:
1055:Kwanyinaspis
1053:
1046:
1012:
1005:
998:
991:
984:
979:Glypharthrus
977:
970:
963:
956:
949:
942:Aglaspididae
931:Tremaglaspis
929:
922:
915:
908:
901:
894:
873:
867:Obrutschewia
865:
857:
849:
842:
835:
827:
803:
798:Pseudarthron
796:
789:
782:
775:
767:
762:Cheloniellon
760:
740:
733:
726:
719:
712:
682:
675:
668:
647:
640:
633:
626:
619:
612:
570:
480:
474:
464:
423:
419:
409:
390:
384:
351:
347:
341:
330:. Retrieved
325:
321:
311:
296:
286:
280:
240:
239:
238:
226:
225:
219:Type species
206:
205:
193:Conciliterga
160:(unranked):
24:
18:
1260:Haifengella
1048:Arthroaspis
993:Hesselbonia
972:Chraspedops
965:Aglaspoides
903:Chlupacaris
829:Angarocaris
816:Aglaspidida
784:Neostrabops
769:Drabovaspis
714:Emeraldella
661:Protosutura
173:Subphylum:
1487:Categories
1187:Misszhouia
1178:Naraoiidae
1167:Kangacaris
1142:Nektaspida
1124:Xandarella
1117:Sinoburius
1096:Cindarella
1069:Retifacies
1007:Tuboculops
951:Aglaspella
910:Cyclopites
859:Girardevia
844:Beckwithia
791:Paraduslia
635:Squamacula
581:Arthropoda
332:2008-01-12
303:References
261:Atdabanian
186:Subclass:
154:Arthropoda
1498:Artiopoda
1417:Q11305050
1317:Trilobita
1306:Trilobita
1295:Tegopelte
1236:Tariccoia
1229:Soomaspis
1214:Buenaspis
1076:Tonglaiia
917:Flobertia
742:Carimersa
728:Kodymirus
721:Eozetetes
642:Thulaspis
614:Bailongia
596:Artiopoda
566:Ecdysozoa
558:Kingdom:
548:Artiopoda
448:0022-3360
368:0300-3256
288:Tegopelte
257:arthropod
253:trilobite
167:Artiopoda
140:Kingdom:
134:Eukaryota
1411:Wikidata
1402:Saperion
1377:Category
1288:Skioldia
1281:Saperion
1267:Helmetia
1205:Liwiidae
1160:Emucaris
1014:Uarthrus
1000:Setaspis
986:Gogglops
958:Aglaspis
735:Sidneyia
684:Zhiwenia
649:Urokodia
579:Phylum:
560:Animalia
507:22072605
456:84029615
376:85077111
297:Saperion
293:antennae
282:Skioldia
241:Saperion
207:Saperion
150:Phylum:
144:Animalia
130:Domain:
25:Saperion
1457:4709441
1444:1064044
1431:4639611
1274:Kuamaia
1194:Naraoia
1131:Zhugeia
805:Triopus
628:Molaria
498:3282348
428:Bibcode
246:extinct
199:Genus:
1470:142093
777:Duslia
505:
495:
454:
446:
397:
374:
366:
273:Yunnan
255:-like
244:is an
1439:IRMNG
1222:Liwia
571:Clade
452:S2CID
372:S2CID
328:: 560
277:China
249:genus
1426:GBIF
1315:see
503:PMID
444:ISSN
395:ISBN
364:ISSN
285:and
37:PreꞒ
493:PMC
485:doi
481:279
436:doi
356:doi
326:364
271:of
251:of
1489::
1467::
1454::
1441::
1428::
1413::
573::
501:.
491:.
479:.
473:.
450:.
442:.
434:.
424:73
422:.
418:.
370:.
362:.
352:20
350:.
324:.
320:.
275:,
87:Pg
1217:?
878:?
870:?
862:?
854:?
832:?
772:?
652:?
540:e
533:t
526:v
509:.
487::
458:.
438::
430::
403:.
378:.
358::
335:.
203:†
190:†
177:†
164:†
92:N
82:K
77:J
72:T
67:P
62:C
57:D
52:S
47:O
42:Ꞓ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
