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stay in deeper water. Many grey reef sharks have a home range on a specific area of the reef, to which they continually return. However, they are social rather than territorial. During the day, these sharks often form groups of 5β20 individuals near coral-reef drop-offs, splitting up in the evening as the sharks begin to hunt. They are found over continental and insular shelves, preferring the leeward (away from the direction of the current) sides of coral reefs with clear water and rugged topography. They are frequently found near the drop-offs at the outer edges of the reef, and less commonly within
231:
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1906:, and electrical cues given off by potential prey. Its visual system is attuned more to movement and/or contrast than to object details. It is especially sensitive to natural and artificial low-frequency sounds in the 25β100 Hz range, which evoke struggling fish. Whitetips hunt primarily at night, when many fishes are asleep and easily taken. After dusk, a group of sharks may target the same prey item, covering every exit route from a particular coral head. Each shark hunts for itself and in competition with the others in its group. They feed mainly on bony fishes, including
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likely that a number of factors contribute. These include the rich habitat complexity and diversity inherent in coral reef ecosystems, the wide variety and temporal availability of food resources available to coral reef fishes, a host of pre and post-larval settlement processes, and as yet unresolved interactions between all these factors. The wealth of fishes on reefs is filled by tiny, bottom-dwelling reef fishes.
206:
738:) often schools with the blue-striped snapper. The yellowfins change their colouration to match that of the snapper. Presumably this is for predator protection, since goatfish are a more preferred prey than bluestripe snapper. By night the schools disperse and individual goatfish head their separate ways to loot the sands. Other nocturnal feeders shadow the active goatfish, waiting patiently for overlooked morsels.
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1513:, who are also immune to the anemone toxins. To get their meal, butterflyfish must get past these protective clownfish who, although smaller, are not intimidated. An anemone without its clownfish will quickly be eaten by butterflyfish. In return, the anemones provide the clownfish protection from their predators, who are not immune to anemone stings. As a further benefit to the anemone, waste
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912:
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The stonefish has control over whether to shoot its venom, and does so when provoked or frightened. The venom results in severe pain, paralysis and tissue death, and can be fatal if not treated. Despite its formidable defence, the stonefish does have predators. Some bottom feeding rays and sharks with crushing teeth feed on them, as does the
468:
believing that the fish will flee tail first. When escape is not possible, the butterflyfish will sometimes turn to face its aggressor, head lowered and spines fully erect, like a bull about to charge. This may serve to intimidate the other animal or may remind the predator that the butterflyfish is too spiny to make a comfortable meal.
1198:. Spinefoot rabbitfish are named for their defensive venomous spines, and they are seldom attacked by predators. Spines are a last-ditch defence. It is better to avoid predator detection in the first place, and avoid being thrust into risky spine-to-fang battles. So rabbitfish have also evolved skilful colour changing abilities.
148:. Many reef fish confine themselves to one small neighbourhood where every hiding place is known and can be immediately accessed. Others cruise the reefs for food in shoals, but return to a known area to hide when they are inactive. Resting small fish are still vulnerable to attack by crevice predators, so many fish, such as
1422:, which are up to 30 cm (12 in) long, have developed a resistance to these toxins. Saddle butterflyfish usually flutter gently rather than swim. However, in the presence of their preferred food, sea anemones, this gentleness disappears, and the butterflyfish dash in and out, ripping off the anemone tentacles.
224:
fish fauna with no natural overlap in species. Of the two regions, the richest by far in terms of reef fish diversity is the Indo-Pacific where there are an estimated 4,000β5,000 species of fishes associated with coral reef habitats. Another 500β700 species can be found in the greater
Caribbean region.
1871:, or near drop-offs to deeper water. Whitetips prefer very clear water and rarely swim far from the bottom. They spend most of the daytime resting inside caves. Unlike other requiem sharks, which usually rely on ram ventilation and must constantly swim to breathe, these sharks can pump water over their
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that sits on the bottom waiting for prey to come close. It does not swim away if disturbed, but erects 13 venomous spines along its back. For defence, it can shoot venom from each or all of these spines. Each spine is like a hypodermic needle, delivering the venom from two sacs attached to the spine.
223:
There are two major regions of coral reef development recognized; the Indo-Pacific (which includes the
Pacific and Indian Oceans as well as the Red Sea), and the tropical western Atlantic (also known as the "wider" or "greater" Caribbean). Each of these two regions contains its own unique coral reef
2048:
are usually less than 1.9 metres (6.2 ft) long. Despite their moderate size, grey reef sharks actively expel most other shark species from favored habitats. In areas where this species co-exists with the blacktip reef shark, the latter species occupy the shallow flats while the grey reef sharks
1186:
are named for their parrot-like beaks and bright colours. They are large herbivores that graze on the algae that grows on hard dead corals. Equipped with two pairs of crushing jaws and their beaks, they pulverize chunks of algae-coated coral, digesting the algae and excreting the coral as fine sand.
765:
that can vary greatly in size according to the population size and reef habitat. When no male is available, in each school the largest female shifts sex to male. If the final male disappears, changes to the largest female occur, with male behavior occurring within several hours and sperm production
163:
which feeds on benthic turf algae. They also provide cleaner services to marine turtles, by removing algal growth from their shells. They do not tolerate other fish with the same colour or shape. When alarmed, the usually placid yellow tang can erect spines in its tail and slash at its opponent with
2097:
in the reef ecosystem. Like the whitetip reef shark, they have been documented resting motionless on the sea bottom or inside caves - unusual behaviour for requiem sharks. Caribbean reef sharks play a major role in shaping
Caribbean reef communities. They are more active at night, with no evidence
2072:
separation from the blacktip reef sharks; around islands where both species occur, the blacktip occupies shallow flats, while the grey reef shark is usually found in deeper areas, but where the blacktip is absent, the grey reef shark is commonly found on the flats... complements the whitetip shark
657:
Some fishes associated with reefs are generalized carnivores that feed on a variety of animal prey. These typically have large mouths that can be rapidly expanded, thereby drawing in nearby water and any unfortunate animals contained within the inhaled water mass. The water is then expelled through
354:
landscapes of coral reefs. For this manoeuvrability is more important than straight line speed, so coral reef fish have developed bodies which optimize their ability to dart and change direction. They outwit predators by dodging into fissures in the reef or playing hide and seek around coral heads.
1769:
can die as a result of eating a trunkfish. Ciguatera toxins appear to accumulate in top predators of coral reefs. Many of the
Caribbean groupers and the barracuda for example may contain enough of this toxin to cause severe symptoms in humans who eat them. What makes the situation particularly
1753:
can only release venom when something strikes its spines. Although not native to the US coast, lionfish have appeared around
Florida and have spread up the coast to New York. They are attractive aquarium fish, sometimes used to stock ponds, and may have been washed into the sea during a hurricane.
353:
Most reef fishes have body shapes that are different from open water fishes. Open water fish are usually built for speed in the open sea, streamlined like torpedoes to minimise friction as they move through the water. Reef fish are operating in the relatively confined spaces and complex underwater
1652:
Venomous fish carry their venom in venom glands and use various delivery systems, such as spines or sharp fins, barbs or spikes, and fangs. Venomous fish tend to be either very visible, using flamboyant colours to warn enemies, or skilfully camouflaged and maybe buried in the sand. Apart from the
596:
Many reef fish species have evolved different feeding strategies accompanied by specialized mouths, jaws and teeth particularly suited to deal with their primary food sources found in coral reef ecosystems. Some species even shift their dietary habits and distributions as they mature. This is not
1998:
in the central
Pacific has found that the blacktip reef shark had a home range of about 0.55 km (0.21 sq mi), among the smallest of any shark species. The size and location of the range does not change with time of day. The blacktip reef shark swims alone or in small groups. Large
467:
thinking the fish is bigger than it is, and confusing the back end with the front end. The butterflyfish's first instinct when threatened is to flee, putting the false eyespot closer to the predator than the head. Most predators aim for the eyes, and this false eyespot tricks the predator into
219:
The mechanisms that first led to, and continue to maintain, such concentrations of fish species on coral reefs has been widely debated over the last 50 years. While many reasons have been proposed, there is no general scientific consensus on which of these is the most influential, but it seems
1009:". The bluestreak is marked with a conspicuous bright blue stripe and behaves in a stereotypical way which attracts larger fish to its cleaning station. As the bluestreak snacks on the parasites it gently tickles its client. This seems to bring the larger fish back again for regular servicing.
873:
also form schools. They are swift predators who patrol the reef in hunting packs. When they find a school of forage fish, such as cardinalfish, they surround them and herd them close to the reef. This panics the prey fish, and their schooling becomes chaotic, leaving them open to attack by the
863:, including defence against predators through better predator detection, since each fish is on the lookout. Schooling fish have developed remarkable displays of precise choreography which confuse and evade predators. For this they have evolved special pressure sensors along their sides, called
63:. Among the myriad inhabitants, the fish stand out as colourful and interesting to watch. Hundreds of species can exist in a small area of a healthy reef, many of them hidden or well camouflaged. Reef fish have developed many ingenious specialisations adapted to survival on the reefs.
1993:
are prefer deeper water. Younger sharks favour shallow sandy flats, and older sharks spend more time around reef ledges and near reef drop-offs. Blacktip reef sharks are strongly attached to their own area, where they may remain for up to several years. A tracking study off
503:
avoid predators by tucking themselves into coral crevices or partly burying themselves in sand. They continually scan for predators with eyes that swivel independently. The camouflage of the tasseled scorpionfish can prevent gobies from seeing them until it's too late.
1649:. Venomous fish do not necessarily cause poisoning if they are eaten, since the venom is often destroyed in the digestive system. By contrast, poisonous fish contain strong toxins which are not destroyed by the digestive system. This makes them poisonous to eat.
1875:
and lie still on the bottom. They have slender, lithe bodies, which allow them to wriggle into crevices and holes and extract prey inaccessible to other reef sharks. On the other hand, they are rather clumsy when attempting to take food suspended in open water.
1890:
Whitetip reef sharks do not frequent very shallow water like the blacktip reef shark, nor the outer reef like the grey reef shark. They generally remain within a highly localized area. An individual shark may use the same cave for months to years. The daytime
1016:
secretes a mucus coating which reduces drag when they swim and also protects it from some parasites. But other parasites find the mucus itself good to eat. So lizardfish visit the cleaner wrasse, which clean the parasites from the skin, gills and mouth.
447:
The patterns have different functions. Sometimes they camouflage the fish when the fish rests in places with the right background. Colouration can also be used to help species recognition during mating. Some unmistakable contrasting patterns are used to
954:
are ferocious predators on other fishes, with razor-sharp conical teeth which make it easy for them to rip their prey to shreds. Barracuda patrol the outer reef in large schools, and are extremely fast swimmers with streamlined, torpedo-shaped bodies.
1770:
dangerous is that such species may be toxic only at certain sizes or locations, making it difficult to know whether or when they are or are not safe to eat. In some locations this leads to many cases of ciguatera poisoning among tropical islanders.
911:
3071:"An annotated list of fish parasites (Isopoda, Copepoda, Monogenea, Digenea, Cestoda, Nematoda) collected from Snappers and Bream (Lutjanidae, Nemipteridae, Caesionidae) in New Caledonia confirms high parasite biodiversity on coral reef fish"
230:
66:
Coral reefs occupy less than 1% of the surface area of the world oceans, but provide a home for 25% of all marine fish species. Reef habitats are a sharp contrast to the open water habitats that make up the other 99% of the world oceans.
3030:"An annotated list of fish parasites (Copepoda, Monogenea, Digenea, Cestoda and Nematoda) collected from Emperors and Emperor Bream (Lethrinidae) in New Caledonia further highlights parasite biodiversity estimates on coral reef fish"
1866:
is a small shark usually less than 1.6 m (5.2 ft) in length. It is found almost exclusively around coral reefs where it can be encountered around coral heads and ledges with high vertical relief, or over sandy flats, in
1172:, which means they are plants synthesising food directly from solar energy and carbon dioxide and other simple nutrient molecules. Without algae, everything on the reef would die. One important algal group, the bottom dwelling (
1602:. Numerical estimates of parasite biodiversity have shown that certain coral fish species have up to 30 species of parasites. The mean number of parasites per fish species is about ten. This has a consequence in term of
926:
1847:, all have the robust, streamlined bodies typical of the requiem shark. As fast-swimming, agile predators, they feed primarily on free-swimming bony fishes and cephalopods. Other species of reef sharks include the
772:
488:
is an ambush predator that looks like part of a sea floor encrusted with coral and algae. It lies in wait on the sea floor for crustaceans and small fish, such as gobies, to pass by. Another ambush predator is the
250:
2974:"An annotated list of parasites (Isopoda, Copepoda, Monogenea, Digenea, Cestoda and Nematoda) collected in groupers (Serranidae, Epinephelinae) in New Caledonia emphasizes parasite biodiversity in coral reef fish"
477:(right) is not easily seen due to its bottom-feeding habit and its small size, reaching only about 6 cm. It feeds primarily on small crustaceans and other invertebrates, and is popular in the aquarium trade.
1653:
defence or hunting value, venom might have value for bottom dwelling fish by killing the bacteria that try to invade their skin. Few of these venoms have been studied. They are a yet to be tapped resource for
523:: from below, the white spots look like the lighted surface of the water above; and from above, the fish blends more with the coral reef below. The brightly painted yellow mouth may deter potential predators.
365:, have evolved bodies which are deep and laterally compressed like a pancake. Their pelvic and pectoral fins are designed differently, so they act together with the flattened body to optimise manoeuvrability.
2132:
1089:
2243:
Similarity and
Diversity Among Coral Reef Fish Communities: A Comparison between Tropical Western Atlantic (Virgin Islands) and Tropical Central Pacific (Marshall Islands) Patch Reefs. Gladfelter et al.
529:
270:
3625:
Papastamatiou, Y.P.; C.G. Lowe; J.E. Caselle & A.M. Friedlander (April 2009). "Scale-dependent effects of habitat on movements and path structure of reef sharks at a predator-dominated atoll".
493:(right). They lie on the bottom and wave a conspicuous worm-like lure strategically attached above their mouth. Normally about 10 cm (4 in) long, they can also inflate themselves like
961:
are medium to large sized, and are usually found swimming among or near coral reefs. They inflate their body by swallowing water, reducing potential predators to those with much bigger mouths.
561:
185:
3181:
2944:
125:
among algae, invertebrates and fish. They have become crowded and complex environments, and the fish have evolved many ingenious ways of surviving. Most fishes found on coral reefs are
1777:
buries itself in sand and can deliver electric shocks as well as venom. It is a delicacy in some cultures (the venom is destroyed when it is cooked), and can be found for sale in some
1340:
which would normally prevent close contact. The protection fire corals offer hawkfish means the hawkfish has the high ground of the reef, and can safely survey its surroundings like a
1046:
5080:
3940:
2940:
Justine, J.-L. 2010: Parasites of coral reef fish: how much do we know? With a bibliography of fish parasites in New
Caledonia. Belgian Journal of Zoology, 140 (Suppl.), 155β190.
1070:
654:. Competition among carnivores is intense, resulting in a treacherous environment for their prey. Hungry predators lurk in ambush or patrol every part of the reef, night and day.
216:
contain the most diverse fish assemblages to be found anywhere on earth, with perhaps as many as 6,000β8,000 species dwelling within coral reef ecosystems of the world's oceans.
7645:
5572:
624:, are generalized feeders with more standard jaw and mouth structures that allow them to forage on a wide range of animal prey types, including small fishes and invertebrates.
129:, known for the characteristic sharp, bony rays and spines in their fins. These spines provide formidable defences, and when erected they can usually be locked in place or are
612:
and other small invertebrate animals. Their mouths protrude like forceps, and are equipped with fine teeth that allow them to nip off such exposed body parts of their prey.
2102:. Juveniles tend to remain in a localized area throughout the year, while adults range over a wider area. The Caribbean reef shark feeds on a wide variety of reef-dwelling
7298:
807:
1212:
for larger predators. They are small, typically five centimetres (two inches) long. Many species are aggressive towards other fishes which also graze on algae, such as
1734:
2275:
Tiny fishes fuel life on coral reefs typically evoke clear, turquoise waters and a staggering number of colourful fishes. But what supports such an abundance of life?
577:
440:
Coral reef fishes exhibit a huge variety of dazzling and sometimes bizarre colours and patterns. This is in marked contrasts to open water fishes which are usually
2810:
2073:
as it is far more adapt at catching off-bottom fish than the whitetip, but the later is far more competent in extracting prey from crevices and holes in reefs."
7039:
5075:
3571:
Papastamatiou, Y.P.; J.E. Caselle; A.M. Friedlander & C.G. Lowe (16 September 2009). "Distribution, size frequency, and sex ratios of blacktip reef sharks
1714:
3552:
Yano, K.; H. Mori; K. Minamikawa; S. Ueno; S. Uchida; K. Nagai; M. Toda & M. Masuda (June 2000). "Behavioral response of sharks to electric stimulation".
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toxin from glands on its skin when touched. The toxin is only dangerous when ingested, so there's no immediate harm to divers. However, predators as large as
2031:, and members of their own species. At Palmyra Atoll, adult blacktip reef sharks avoid patrolling tiger sharks by staying out of the central, deeper lagoon.
1694:
545:
2903:
Porat, D.; Chadwick-Furman, N. E. (2005). "Effects of anemonefish on giant sea anemones: ammonium uptake, zooxanthella content and tissue regeneration".
1610:
suggest that extinction of a coral reef fish species of average size would eventually result in the co-extinction of at least ten species of parasites.
170:
3909:
1895:
of a whitetip reef shark is limited to about 0.05 km (0.019 sq mi); at night this range increases to 1 km (0.39 sq mi).
944:
can move relatively large rocks when feeding and is often followed by smaller fishes that feed on leftovers. They also use a jet of water to uncover
1440:
1270:
1190:
Smaller parrotfish are relatively defenceless herbivores, poorly defended against predators like barracuda. They have evolved to find protection by
459:
gets its name from a large dark spot on the rear portion of each side of the body. This spot is surrounded by a brilliant white ring, resembling an
6820:
3695:"Distribution and diet of four species of carcharhinid shark in the Hawaiian Islands: evidence for resource partitioning and competitive exclusion"
1428:
1254:
337:, are usually streamlined for straightline speed, with a deeply forked tail and a smooth body shaped like a spindle tapered at both ends. They are
433:. The colouration of triggerfish fade into drabness when they sleep or are being submissive, and return to vividness when they are not threatened.
7655:
7281:
5105:
4317:
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hunt cardinalfish in packs and herd them against the reef. When the cardinalfish panic and break school formation, the trevally pick them off.
1641:. Both types of fish contain strong toxins, but the difference is in the way the toxin is delivered. Venomous fish deliver their toxins (called
639:, are generalized reef feeders with standard jaw and mouth structures that allow them to eat almost anything, though they prefer small fish and
3157:
896:
7328:
4183:
3815:
4597:
3542:
Nelson, D.R. and R.H. Johnson. (1970). Acoustic studies on sharks: Rangiroa Atoll, July 1969. ONR Technical Report 2, No. N00014-68-C-0138.
3316:
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Mixed schooling and its possible significance in a tropical western
Atlantic parrotfish and surgeonfish. WS Alevizon, Copeia 1976:797β798.
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5679:
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2481:
880:
4920:
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Porat, D.; Chadwick-Furman, N. E. (2004). "Effects of anemonefish on giant sea anemones: expansion behavior, growth, and survival".
2542:
7403:
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792:
317:, have a body flattened like a pancake, with pectoral and pelvic fins that act with the flattened body to maximize manoeuvrability.
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718:(whiskers) protruding from their chins to rifle through the sediments in search of a meal. Like goats, they seek anything edible:
7564:
5095:
4880:
1977:
is typically about 1.6 m (5.2 ft) long. It is usually found over reef ledges and sandy flats, though it can also enter
3131:
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poisoning if eaten. Outbreaks of ciguatera poisoning in the 11th to 15th centuries from large, carnivorous reef fish, caused by
1222:
4262:
1023:
516:
512:
4414:
3838:
3738:, at Fernando de Noronha Archipelago, Brazil: the potential of marine protected areas for conservation of a nursery ground".
2706:
FAO Species
Catalogue Vol. 6: Snappers of the World; An Annotated and Illustrated Catalogue of Lutjanid Species Known to Date
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2369:
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2118:. Young sharks feed on small fishes, shrimps, and crabs. In turn, young sharks are preyed on by larger sharks such as the
2643:
Clark, Nicholas; Russ, G (2012). "Ontogenetic shifts in the habitat associations of butterflyfishes (F. Chaetodontidae)".
1328:. Thanks to their large, skinless pectoral fins, hawkfish can perch on fire corals without harm. Fire corals are not true
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6051:
5208:
5070:
3242:
3069:
Justine, J-L.; Beveridge, I.; Boxshall, GA.; Bray, RA.; Miller, TL.; Moravec, F.; Trilles, JP.; Whittington, ID. (2012).
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Analysis of the influence of substrate variables on coral reef fish communities. Luckhurst, B.E. and K. Luckhurst, 1978.
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where other fish can come to have their parasites nibbled away. The "resident fish doctor and dentist on the reef is the
3734:
Garla, R.C.; Chapman, D.D.; Wetherbee, B.M. & Shivji, M. (2006). "Movement patterns of young Caribbean reef sharks,
3230:
1363:. The pearlfish enters the sea cucumber through its anus, and spends the day safely protected inside the sea cucumber's
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are the most diverse of feeding types among coral reef fishes. There are many more carnivore species on the reefs than
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7393:
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5100:
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3896:
3869:
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Springer, S. (1967), "Social organization of shark populations", in Gilbert, P.W.; R.F. Mathewson; D.P. Rail (eds.),
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Justine, J-L.; Beveridge, I.; Boxshall, GA.; Bray, RA.; Moravec, F.; Trilles, JP.; Whittington, ID. (November 2010).
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3662:"A review of shark agonistic displays: comparison of display features and implications for shark-human interactions"
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4457:
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3987:
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Depczynski, Martial; Bellwood, David R (2005), "Shortest recorded vertebrate lifespan found in a coral reef fish",
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859:. The forage fish are, in turn, eaten by larger fish, such as the bigeye trevally. Fish receive many benefits from
2823:
114:. If the triggerfish wants to hide, it will squeeze into a coral crevice and lock itself in place with its spines.
102:
have mouths that crush shells. Orange-lined triggerfish are particularly aggressive. The black and white fish are
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6806:
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Lionfish can aggressively dart at scuba divers and attempt to puncture the facemask with their venomous spines.
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5562:
2328:
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inflate themselves by swallowing water or air, which restricts potential predators to those with bigger mouths.
620:
growing on reef surfaces, utilizing mouths like beaks well adapted to scrape off their food. Other fish, like
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79:
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2757:"Interspecific Communicative and Coordinated Hunting between Groupers and Giant Moray Eels in the Red Sea"
2345:
2142:
uses its hammer both to locate electrical signatures of stingrays buried in the sand and to pin them down.
1099:
secretes a mucus coating which reduces drag when they swim. But some parasites find the mucus good to eat.
463:. A black vertical bar on the head runs through the true eye, making it hard to see. This can result in a
5720:
5577:
4583:
4342:
4302:
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2690:
2525:
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141:
1124:. This may be a mechanism for overwhelming the highly aggressive defence responses of small territorial
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6199:
6169:
6056:
5665:
4885:
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Kline, R.J. (2010) "Hormonal correlates of coloration and sexual change in the hermaphroditic grouper,
1594:(digeneans). The high biodiversity of coral reefs increases the complexity of the interactions between
1352:
as they pass by. They are mostly solitary, although some species form pairs and share a head of coral.
1216:. Surgeonfish sometimes use schooling as a countermeasure to defensive attacks by solitary damselfish.
7739:
7734:
7711:
7620:
7451:
7426:
6745:
6738:
6705:
6447:
6119:
5800:
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5087:
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3028:
Justine, J.-L.; Beveridge, I.; Boxshall, G. A.; Bray, R. A.; Moravec, F.; Whittington, I. D. (2010).
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17:
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95:
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1800:, could be a reason why Polynesians migrated to Easter Island, New Zealand, and possibly Hawaii.
1509:. This gives the sea anemones a second line of defence. They are guarded by fiercely territorial
480:
Just as some prey species evolved cryptic colouration and patterns to help avoid predators, some
241:
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6519:
6504:
6327:
6046:
5935:
5173:
5163:
4865:
4850:
4424:
4277:
1461:
1191:
1121:
887:
860:
840:
473:
386:
378:
145:
7054:
3876:
3859:
1305:
refers to two species that have a close relationship with each other. The relationship can be
511:
has strong jaws for crushing and eating sea urchins, crustaceans and hard-shell molluscs. Its
7524:
7398:
7024:
6919:
6725:
6710:
6219:
6036:
5592:
5519:
5454:
5319:
5294:
5269:
5153:
4900:
4680:
4419:
4383:
4347:
4189:
4177:
4155:
2853:
1579:
1535:, parasites of coral reef fish show tremendous variety. Parasites of coral reef fish include
813:
Yellowfin goatfish change their colouration so they can school with the blue-striped snapper.
539:
has false eyes on its back end, confusing predators about which is the front end of the fish.
536:
485:
456:
334:
7773:
7768:
7383:
7363:
7323:
7293:
7240:
6720:
6016:
5327:
5289:
4620:
4529:
4467:
4272:
3980:
3960:
3706:
2615:
2090:
2082:
1840:
1797:
1493:
1476:
1446:
1419:
568:
390:
is one of only two animal species known to have blue colouring because of cellular pigment.
3498:
Sharks of the World: An Annotated and Illustrated Catalogue of Shark Species Known to Date
8:
7544:
7534:
7467:
7436:
7421:
7248:
7244:
7208:
6964:
6934:
6509:
6452:
6252:
6139:
5928:
5549:
5203:
5012:
4970:
4740:
4700:
4447:
4242:
4217:
1986:
1974:
1966:
1883:
1863:
1824:
1820:
1813:
1781:
with the electric organ removed. They have been called "the meanest things in creation".
1306:
1176:) algae, grows over dead coral and other inert surfaces, and provides grazing fields for
1135:
feed on plants. The four largest groups of coral reef fishes that feed on plants are the
240:
species, small coral reef-dwelling fishes, is the world's shortest lived vertebrate, the
103:
3710:
3624:
3570:
3029:
586:
are very cautious, yet they can fail to see a tassled scorpionfish until it is too late.
7729:
7505:
7348:
7128:
6091:
5469:
5464:
5332:
5274:
5168:
5017:
4935:
4820:
4800:
4710:
4653:
4393:
4378:
4327:
4267:
3755:
3097:
3070:
2920:
2885:
2828:
2783:
2756:
2660:
2456:
2312:
1096:
1013:
798:
659:
636:
134:
3692:
3414:
Voyages of discovery or necessity? Fish poisoning may be why Polynesians left paradise
7610:
7595:
7078:
6999:
6929:
6874:
6859:
6843:
6437:
6342:
6247:
6159:
6154:
6101:
5908:
5715:
5705:
5479:
5357:
5304:
5223:
4890:
4855:
4795:
4755:
4725:
4695:
4606:
4398:
4247:
3954:
3892:
3865:
3848:
3834:
3819:
3774:
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3592:
3588:
3501:
3399:
3382:
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3102:
2995:
2788:
2735:
2710:
2623:
2417:
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1852:
1758:
1701:
1670:
1169:
1053:
1030:
941:
658:
the gills with the mouth closed, thereby trapping the helpless prey For example, the
597:
surprising, given the huge variety in the types of prey on offer around coral reefs.
508:
490:
460:
430:
405:
126:
4204:
3759:
2924:
2704:
2664:
2460:
2316:
195:
can erect spines in its tail and slash at its opponent with rapid sideways movements
7691:
7495:
7490:
7198:
7118:
6914:
6899:
6894:
6179:
6124:
6086:
6041:
5999:
5923:
5710:
5587:
5279:
5183:
5135:
5047:
4940:
4638:
4559:
4534:
4150:
4107:
4031:
3792:
3747:
3714:
3673:
3634:
3584:
3394:
3092:
3082:
3044:
3004:
2985:
2912:
2889:
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2768:
2652:
2446:
2296:
2139:
2061:
1856:
1774:
1741:
1364:
1355:
A more bizarre example of commensalism occurs between the slim, eel-shaped pinhead
1081:
1002:
989:
715:
663:
152:, squeeze into a small hiding place and wedge themselves by erecting their spines.
71:
3861:
Coral Reef Fishes: Dynamics and Diversity in a Complex Ecosystem by Peter F. Sale
7696:
7554:
7343:
7133:
7073:
6974:
6396:
6204:
6191:
6174:
6129:
6011:
5950:
5874:
5869:
5805:
5792:
5762:
5557:
5524:
5337:
5213:
5188:
4997:
4950:
4875:
4720:
4705:
4554:
4524:
4373:
4227:
3973:
3935:
3733:
3320:
3161:
2948:
2773:
2115:
2045:
2039:
1990:
1955:
1848:
1828:
1665:
1618:
1540:
917:
870:
761:
605:
552:
481:
413:
330:
314:
144:. Small reef fish get protection from predators by hiding in reef crevices or by
107:
82:, are threatening the survival of the coral reefs and the associated reef fish.
7686:
7615:
7529:
7178:
7158:
7138:
7113:
7063:
6994:
6984:
6944:
6884:
6847:
6672:
6645:
6537:
6529:
6462:
6432:
6371:
6353:
6307:
6297:
5901:
5854:
5421:
5375:
5370:
5342:
5309:
5178:
5037:
4965:
4955:
4805:
4750:
4668:
4663:
4658:
4539:
4307:
4252:
4056:
3458:
Randall, J.E. (1977). "Contribution to the Biology of the Whitetip Reef Shark (
2262:
2099:
1836:
1685:
1661:
1654:
1634:
1599:
1583:
701:
material. Diet varies with age, location and the prevalent prey items locally.
520:
441:
338:
276:
261:
110:
damselfish. If the triggerfish attacks, the damselfish will hide in the nearby
7235:
3751:
3678:
3661:
3048:
2916:
2881:
2656:
2300:
2077:
7762:
7706:
7701:
7605:
7368:
7266:
7236:
7123:
7108:
7093:
6969:
6774:
6677:
6591:
6411:
6386:
6381:
6337:
6332:
6287:
6282:
6262:
6134:
5913:
5832:
5635:
5529:
5494:
5395:
5390:
5365:
4895:
4860:
4835:
4685:
4648:
4492:
4212:
4162:
4051:
4026:
3930:
3783:
3772:
3353:
3286:
3258:
3200:
3127:
2868:
2580:
2538:
2477:
2365:
2341:
2094:
1995:
1943:
1844:
1638:
1607:
1603:
1572:
958:
932:
756:
601:
424:
362:
358:
130:
3413:
2755:
Bshary, Redouan; Hohner, Andrea; Ait-el-Djoudi, Karim; Fricke, Hans (2006).
90:
7188:
7103:
7083:
7029:
6939:
6909:
6667:
6491:
6406:
6366:
6267:
6209:
6149:
6076:
6071:
6061:
5977:
5967:
5896:
5879:
5786:
5747:
5740:
5621:
5400:
5380:
5284:
5022:
4905:
4825:
4785:
4775:
4745:
4505:
4322:
4097:
4089:
4041:
4008:
3880:
3646:
3596:
3106:
3087:
2999:
2792:
2308:
2107:
2069:
2054:
1911:
1766:
1664:. It has a remarkable ability to camouflage itself amongst rocks. It is an
1646:
1576:
1568:
1532:
1360:
1349:
1310:
1061:
998:
902:
864:
852:
731:
712:
284:
60:
3693:
Papastamatiou, Y.P.; Wetherbee, B.M.; Lowe, C.G. & Crow, G.L. (2006).
1276:
Schooling spinefoot rabbitfish are often joined by defenceless parrotfish.
7441:
7353:
7203:
7183:
7168:
7153:
7148:
7098:
6879:
6798:
6650:
6638:
6564:
6322:
6317:
6292:
6272:
6214:
6109:
6004:
5982:
5972:
5945:
5815:
5767:
5534:
5509:
5504:
5474:
5257:
5233:
5198:
5007:
4910:
4830:
4790:
4643:
4498:
4287:
4222:
4036:
4021:
4016:
3965:
3417:
3383:"Did ciguatera prompt the late Holocene Polynesian voyages of discovery?"
2990:
2973:
2161:
2119:
2028:
1951:
1931:
1927:
1879:
1785:
1778:
1721:
1403:
1213:
1209:
1205:
1156:
1152:
1148:
1113:
945:
856:
848:
844:
779:
753:
690:
449:
307:
192:
156:
149:
122:
111:
99:
75:
2730:
Johnson, G.D.; Gill, A.C. (1998). Paxton, J.R.; Eschmeyer, W.N. (eds.).
1808:
7665:
7373:
7163:
7143:
6700:
6499:
6467:
6401:
6391:
6312:
6302:
5987:
5955:
5940:
5886:
5859:
5842:
5514:
5449:
5238:
5218:
5002:
4975:
4760:
4673:
4564:
4549:
4472:
4462:
4452:
4167:
4145:
4127:
4112:
4102:
4000:
3719:
3694:
3323:. "Rosenstiel School of Marine and Atmospheric Science, Univ. of Miami"
2451:
2434:
2413:
2123:
2012:
2008:
1982:
1923:
1919:
1915:
1892:
1789:
1744:
buries itself in sand and can deliver electric shocks as well as venom.
1411:
1345:
1337:
1325:
1314:
1261:
1229:
1201:
1195:
1183:
1144:
1140:
1136:
1125:
783:
687:
683:
679:
640:
621:
613:
609:
494:
409:
398:
213:
205:
118:
52:
37:
7308:
5630:
3638:
2053:. On occasion, this shark may venture several kilometers out into the
7635:
7600:
7213:
7193:
6457:
6442:
6427:
6257:
5918:
5864:
5849:
5827:
5810:
5757:
5539:
5444:
5228:
4992:
4985:
4960:
4870:
4575:
4544:
2169:
2111:
2103:
2016:
2004:
1903:
1899:
1832:
1793:
1762:
1725:
1705:
1552:
1518:
1510:
1506:
1488:
1356:
1302:
1245:
1177:
1132:
1117:
951:
826:
741:
723:
651:
647:
515:(lower) surface has large, white spots on a dark background, and its
257:
160:
56:
3436:
Hobson, E.S. (1963). "Feeding Behavior in Three Species of Sharks".
7485:
7019:
7009:
6959:
6660:
6601:
6547:
6542:
6361:
6229:
5891:
5837:
5752:
5489:
5299:
4845:
4840:
4765:
4232:
4080:
4070:
4046:
3551:
3366:
3299:
3271:
3213:
3140:
2593:
2551:
2490:
2473:
2378:
2350:
2000:
1978:
1935:
1629:
Many reef fish are toxic. Toxic fish are fish which contain strong
1595:
1544:
1536:
1407:
1367:. At night it emerges the same way and feeds on small crustaceans.
1333:
1321:
1295:
1291:
867:, that let them feel each other's movements and stay synchronized.
704:
464:
32:
1961:
1418:, and are an effective deterrent against most predators. However,
1168:
can drape reefs in kaleidoscopes of colours and shapes. Algae are
7681:
7650:
7580:
7480:
7014:
6829:
6633:
6628:
6606:
6579:
6574:
6569:
6114:
6066:
5960:
5730:
5725:
5499:
5459:
5429:
5407:
5027:
4815:
4715:
4332:
4117:
3910:
Featured Creatures in Coral Reefs at the Smithsonian Ocean Portal
3773:
Rosa, R.S.; Mancini, P.; Caldas, J.P. & Graham, R.T. (2006).
3243:
Reef Stonefish, Synanceia verrucosa (Bloch & Schneider, 1801)
2253:
Food Habits of Reef Fishes of the West Indies, Randall, J.E. 1967
2034:
2024:
2020:
1939:
1750:
1623:
1591:
1564:
1548:
1514:
1498:
1173:
749:
727:
708:
280:
5248:
4618:
1434:
The tentacles of sea anemones bristle with tiny toxic harpoons.
70:
However, loss and degradation of coral reef habitat, increasing
6552:
5484:
4810:
4357:
4064:
3949:
3888:
2856:(2010). In Encyclopædia Britannica. Retrieved January 30, 2010.
2754:
2050:
1868:
1835:
of coral reefs in the Indo-Pacific. Coral reefs in the western
1560:
1077:
1037:
786:, while their juveniles mostly eat crustaceans such as prawns.
671:
583:
500:
3923:
3500:. Rome: Food and Agricultural Organization. pp. 459β461.
519:(upper) surface has black spots on yellow. This is a form of
323:
6655:
6584:
5994:
4135:
3996:
3915:
Coral Reef Fishes: Adaptations, Diversity, and Economic Value
1999:
social aggregations have also been observed. They are active
1642:
1630:
1587:
1556:
1521:
1505:
There is a mutualistic relationship between sea anemones and
1415:
1329:
1313:, when one species benefits and the other is unaffected, and
1165:
1159:
1034:
698:
694:
632:
617:
571:
camouflaged to look like part of a coral encrusted sea floor.
3068:
3027:
2971:
1128:
that vigorously guard small patches of algae on coral reefs.
6557:
6144:
5697:
5688:
4980:
3877:"The structure and dynamics of coral reef fish communities"
1947:
1872:
1341:
1286:
719:
675:
667:
237:
48:
27:
Fish which live amongst or in close relation to coral reefs
1985:
environments. This species likes shallow water, while the
1344:. Hawkfish usually stay motionless, but dart out and grab
6596:
6164:
5042:
5032:
4140:
2809:". Doctoral dissertation, University of Texas at Austin.
1950:. Important predators of the whitetip reef shark include
1907:
484:
evolved camouflage that lets them ambush their prey. The
176:
Most coral reef fish have spines in their fins like this
6842:
3554:
Bulletin of Seikai National Fisheries Research Institute
2435:"Blue Chromatophores in Two Species of Callionymid Fish"
2093:
is up to 3 metres (10 ft) long, one of the largest
2064:
comments on the relationship between the three species.
1792:. Like many other apex reef fish, it is likely to cause
155:
As an example of the adaptations made by reef fish, the
5657:
2729:
997:
Fish can not groom themselves. Some fish specialise as
748:) are known to cooperate with each other when hunting.
279:
often inhabit reefs. Male toadfish "sing" at up to 100
1317:, when one species benefits, and the other is harmed.
452:
that the fish has venomous spines or poisonous flesh.
3618:
3182:
Venom Runs Thick in Fish Families, Researchers Learn
2902:
2865:
2151:
1264:
have venomous spines which they erect if threatened.
3575:at Palmyra Atoll: a predator-dominated ecosystem".
3564:
3380:
3155:
Poisonous vs. Venomous fish: What's the difference?
3023:
3021:
3019:
2967:
2965:
2963:
2832:. December 4, 1984. Retrieved on December 26, 2011.
1309:, when both species benefit from the relationship,
3614:, Baltimore: Johns Hopkins Press, pp. 149β174
3064:
3062:
3060:
3058:
2286:
1531:. Since coral reef fish are characterized by high
260:, often found in reefs. The slowest of these, the
3231:"The Stonefish β The Deadliest Fish in The World"
300:Contrast between coral reef and open water fishes
7760:
3845:"Fishes of the Great Barrier Reef and Coral Sea"
3495:
3016:
2960:
2432:
2038:Coral reef drop-offs are a favoured habitat for
1898:The whitetip reef shark is highly responsive to
1749:Unlike the stonefish which can shoot venom, the
1633:in their bodies. There is a distinction between
905:swim in schools for protection against trevally.
7656:International Seafood Sustainability Foundation
7282:United Nations Convention on the Law of the Sea
3816:"Coral Reef Fishes: Indo-Pacific and Caribbean"
3226:
3224:
3055:
2840:
2838:
1886:spend much of the day lying still on the bottom
1645:) by biting, stinging, or stabbing, causing an
3666:Marine and Freshwater Behaviour and Physiology
2936:
2934:
2209:
2207:
2197:
2195:
1606:. Results obtained for the coral reef fish of
1320:An example of commensalism occurs between the
6814:
5673:
4881:Stable isotope analysis in aquatic ecosystems
4591:
4184:The Structure and Distribution of Coral Reefs
3981:
3686:
3457:
3195:
3193:
1194:, sometimes with other species like shoaling
555:disguised to look like an algae-covered stone
200:
3798:10.2305/IUCN.UK.2006.RLTS.T60217A12323052.en
3727:
3659:
3609:
3435:
3221:
2835:
2642:
2221:
2219:
1410:of sea anemones bristle with tiny harpoons (
7432:List of commercially important fish species
7277:Illegal, unreported and unregulated fishing
4946:Freshwater environmental quality parameters
3885:Theory and management of tropical fisheries
3491:
3489:
3487:
3485:
3483:
3481:
3479:
3477:
2931:
2408:
2406:
2404:
2402:
2400:
2398:
2204:
2192:
1761:is a reef fish which secretes a colourless
51:which live amongst or in close relation to
6828:
6821:
6807:
5680:
5666:
4598:
4584:
3995:
3988:
3974:
3453:
3451:
3190:
2734:. San Diego: Academic Press. p. 186.
2573:
2522:Coral Reef Connections: Predators and Prey
2277:by Amit Malewar. Published on May 24, 2019
1527:As with all fish, coral reef fish harbour
886:Many small reef fishes gain advantages by
782:hunt a variety of reef fish, particularly
6225:Tradeoffs for locomotion in air and water
3796:
3718:
3677:
3521:
3519:
3517:
3431:
3429:
3427:
3398:
3381:Rongo, T; Bush, M; van Woesik, R (2009).
3148:
3096:
3086:
2989:
2782:
2772:
2611:The Macmillan book of the Marine Aquarium
2450:
2216:
834:
627:
7404:Regional fishery management organisation
3939:) is being considered for deletion. See
3474:
3176:
3174:
3172:
3170:
2696:
2414:"Pisces Guide to Caribbean Reef Ecology"
2395:
2322:
2177:β most of these fish are coral reef fish
2076:
2033:
2023:. Blacktip reef sharks are preyed on by
1960:
1878:
1816:almost exclusively inhabits coral reefs.
1807:
1617:
1483:
1449:are resistant to the sea anemone toxin.
1285:
1116:are among the most common of coral reef
1108:
631:
600:For example, the primary food source of
292:
204:
89:
31:
3831:Fishes, An Introduction to Ichthyology.
3653:
3448:
3352:
3285:
3257:
3199:
3164:Reef Biosearch. Retrieved 17 July 2009.
3126:
2682:
2680:
2678:
2676:
2674:
2579:
2537:
2476:
2364:
2340:
1803:
855:, move around the reef feeding on tiny
825:Coral grouper sometimes cooperate with
121:are the result of millions of years of
14:
7761:
4605:
4263:Florida Keys National Marine Sanctuary
3514:
3424:
2517:
2515:
2513:
2511:
2509:
2507:
2505:
2503:
2501:
1384:Pearlfish emerging from a sea cucumber
7234:
6841:
6802:
5661:
5134:
5133:
4617:
4579:
4415:Environmental issues with coral reefs
3969:
3167:
2702:
2607:
2601:
1575:. Some of these fish parasites have
1567:, and various microorganisms such as
1208:and algae, and are an important reef
734:are staples. The yellowfin goatfish (
591:
7287:United Nations Fish Stocks Agreement
3766:
2671:
2175:List of marine aquarium fish species
2085:cruises a coral reef in the Bahamas.
1720:Like many other apex reef fish, the
1684:The most venomous known fish is the
1660:The most venomous known fish is the
1204:are a group of species that feed on
244:, which lives for less than 60 days.
140:Reef fish have also evolved complex
7462:Future of Marine Animal Populations
7339:Fishery Resources Monitoring System
7329:Monitoring control and surveillance
6781:
6052:Electroreception and electrogenesis
5209:Oceanic physical-biological process
5071:List of freshwater ecoregions (WWF)
3784:IUCN Red List of Threatened Species
3531:ReefQuest Centre for Shark Research
2723:
2498:
2098:of seasonal changes in activity or
1497:(arrows) on the gill filament of a
264:, attains about five feet per hour.
133:. Many reef fish have also evolved
24:
3332:Lieske, E. and Myers, R.F. (2004)
2334:
1524:found in the anemone's tentacles.
990:Porcupinefish with cleaner wrasses
256:The slowest-moving fishes are the
25:
7795:
7661:Sea Shepherd Conservation Society
7394:National Marine Fisheries Service
7389:European Fisheries Control Agency
3957:, an online virtual reef for kids
3943:to help reach a consensus. βΊ
3903:
3533:. Retrieved on February 14, 2009.
2106:and cephalopods, as well as some
1788:is a reef fish at the top of the
1657:to find drugs with medical uses.
1336:possessing stinging cells called
7307:
7053:
6870:Population dynamics of fisheries
6780:
6769:
6768:
6751:
6750:
5776:
5642:
5641:
5629:
5615:
5247:
4919:
4736:Colored dissolved organic matter
4458:International Coral Reef Society
4298:Mesoamerican Barrier Reef System
4079:
3589:10.1111/j.1095-8649.2009.02329.x
3400:10.1111/j.1365-2699.2009.02139.x
3112:
3008:
2952:
2687:Coral Reef Connections: Partners
2181:List of reef fish of the Red Sea
2154:
2131:
1733:
1713:
1693:
1677:
1559:, parasitic crustaceans such as
1469:
1454:
1439:
1427:
1390:
1378:
1269:
1253:
1237:
1221:
1162:growing on or near coral reefs.
1088:
1069:
1045:
1022:
984:
978:Giant moray with cleaner wrasses
972:
925:
910:
895:
879:
818:
806:
791:
771:
608:themselves or the appendages of
576:
560:
544:
528:
423:
397:
377:
322:
306:
269:
249:
229:
184:
169:
7591:Aquaculture Stewardship Council
7447:Environmental impact of fishing
5736:Environmental impact of fishing
5081:Latin America and the Caribbean
4389:Stony coral tissue loss disease
3961:List of aquarium saltwater fish
3814:Lieske, E and R. Myers. (2001)
3603:
3545:
3536:
3407:
3374:
3346:
3326:
3307:
3279:
3251:
3236:
3233:, Virginia Wells, Petplace.com.
3120:
2896:
2859:
2847:
2824:SEX CHANGE IN FISH FOUND COMMON
2816:
2799:
2748:
2645:Environmental Biology of Fishes
2636:
2559:
2531:
2467:
2426:
2386:
2358:
1843:. These sharks, all species of
1598:and their various and numerous
1479:guarding their sea anemone home
567:Another ambush predator is the
7565:U.S. Fish and Wildlife Service
6925:Otolith microchemical analysis
5563:Ecological values of mangroves
5106:North Pacific Subtropical Gyre
3829:Moyle, PB and Cech, JJ (2003)
3699:Marine Ecology Progress Series
2329:Guinness Book of World Records
2280:
2268:
2256:
2247:
2237:
2228:
1708:toxin from glands on its skin.
1396:Clownfish swimming among coral
1248:prey in schools on parrotfish.
801:will eat just about anything.
711:feeders, using a pair of long
368:
13:
1:
7560:European Environmental Agency
7550:Grey nurse shark conservation
7457:Destructive fishing practices
6955:Pop-up satellite archival tag
4318:Northwestern Hawaiian Islands
3928:
3847:University of Hawaii Press.
3808:
2213:Moyle and Cech, 2003, p. 561.
2201:Moyle and Cech, 2003, p. 555.
1622:Head-on view of the venomous
1104:
1080:servicing a larger fish at a
348:
80:destructive fishing practices
7540:Marine conservation activism
6990:Climate change and fisheries
5821:intramembranous ossification
5583:Marine conservation activism
5568:Fisheries and climate change
3891:Conference Proceedings (9),
3818:Princeton University Press.
2774:10.1371/journal.pbio.0040431
2693:. Retrieved 16 January 2010.
2616:Macmillan Publishing Company
2528:. Retrieved 16 January 2010.
1359:and a particular species of
1281:
7:
5578:Human impact on marine life
5455:Davidson Seamount Β§ Ecology
4691:Aquatic population dynamics
4343:Southeast Asian coral reefs
3833:5th Ed, Benjamin Cummings.
3660:Martin, R.A. (March 2007).
2691:Public Broadcasting Service
2526:Public Broadcasting Service
2433:Goda, M.; R. Fujii (2009).
2392:Moyle and Cech 2003, pp. 4.
2147:
1613:
766:occurring within ten days.
736:Mulloidichthys vanicolensis
209:Distribution of coral reefs
106:and the unstriped fish are
85:
55:. Coral reefs form complex
10:
7800:
7586:Marine Stewardship Council
7379:EU Common Fisheries Policy
6200:Fin and flipper locomotion
6170:Sequential hermaphroditism
6057:Jamming avoidance response
5774:
4303:New Caledonia barrier reef
2709:. Rome: FAO. p. 207.
1298:, surveys its surroundings
313:Many reef fish, like this
201:Diversity and distribution
164:rapid sideways movements.
7740:Glossary of fishery terms
7735:Index of fishing articles
7725:
7674:
7573:
7517:
7452:Fishing down the food web
7427:Maximum sustainable yield
7414:
7316:
7305:
7259:
7255:
7230:
7062:
7051:
6985:Match/mismatch hypothesis
6858:
6854:
6837:
6764:
6688:
6621:
6528:
6490:
6481:
6420:
6351:
6238:
6190:
6100:
6025:
5785:
5695:
5609:
5548:
5420:
5356:
5318:
5265:
5256:
5245:
5194:Marine primary production
5146:
5142:
5129:
5088:List of marine ecoregions
5063:
4928:
4917:
4631:
4627:
4613:
4517:
4485:
4438:
4407:
4366:
4203:
4195:Spur and groove formation
4126:
4088:
4077:
4007:
3752:10.1007/s00227-005-0201-4
3679:10.1080/10236240601154872
3573:Carcharhinus melanopterus
3496:Compagno, L.J.V. (1984).
3334:Coral reef guide; Red Sea
3248:. Retrieved 21 July 2009.
3049:10.11646/zootaxa.2691.1.1
2917:10.1080/10236240500057929
2882:10.1007/s10750-004-2688-y
2657:10.1007/s10641-011-9964-2
2382:. September 2009 version.
2354:. September 2009 version.
2301:10.1016/j.cub.2005.04.016
1406:are common on reefs. The
1389:
1377:
1372:
1007:bluestreak cleaner wrasse
983:
971:
966:
746:Plectropomus pessuliferus
7626:Save Our Seas Foundation
7359:Individual fishing quota
7334:Vessel monitoring system
6980:Hydrographic containment
5687:
5440:Coastal biogeomorphology
5435:Marine coastal ecosystem
4258:East African coral coast
3941:templates for discussion
3922:: Centre of Excellence
3612:Sharks, Skates, and Rays
3356:; Pauly, Daniel (eds.).
3289:; Pauly, Daniel (eds.).
3261:; Pauly, Daniel (eds.).
3203:; Pauly, Daniel (eds.).
3130:; Pauly, Daniel (eds.).
2905:Mar. Freshw. Behav. Phys
2807:Epinephelus adscensionis
2587:Balistoides conspicillum
2583:; Pauly, Daniel (eds.).
2541:; Pauly, Daniel (eds.).
2480:; Pauly, Daniel (eds.).
2416:Gulf Publishing Company
2368:; Pauly, Daniel (eds.).
2344:; Pauly, Daniel (eds.).
2186:
1969:often patrol reef ledges
1582:(i.e. they have several
1517:from the clownfish feed
1151:. All feed primarily on
1058:orange-lined triggerfish
357:Many reef fish, such as
96:orange-lined triggerfish
94:In the foreground is an
40:are numerous and diverse
7272:Exclusive economic zone
6950:GIS and aquatic science
6905:Standard weight in fish
6716:Glossary of ichthyology
6278:Diel vertical migration
5348:Paradox of the plankton
5159:Diel vertical migration
5053:Freshwater swamp forest
4771:GIS and aquatic science
4619:General components and
3577:Journal of Fish Biology
3387:Journal of Biogeography
2978:Folia Parasitol (Praha)
2597:. January 2010 version.
2555:. January 2010 version.
2346:"Family Batrachoididae"
2263:Coral Reef Fish Ecology
1491:parasites of the genus
1228:Relatively defenceless
1064:that feed on leftovers.
242:seven-figure pygmy goby
104:three-stripe damselfish
7501:Turtle excluder device
7174:Shoaling and schooling
6082:Surface wave detection
6047:Hydrodynamic reception
5721:Diseases and parasites
5174:Large marine ecosystem
4866:Shoaling and schooling
4425:Coral reef restoration
3336:London, HarperCollins
3265:Uranoscopus sulphureus
3088:10.1186/2046-9063-8-22
2732:Encyclopedia of Fishes
2225:Liske and Myers, 2001.
2086:
2075:
2042:
1970:
1887:
1817:
1626:
1590:(certain cestodes) or
1502:
1299:
1129:
835:Specialised carnivores
644:
628:Generalized carnivores
474:Synchiropus splendidus
444:with silvery colours.
387:Synchiropus splendidus
210:
146:shoaling and schooling
137:to confuse predators.
115:
41:
36:The fish that inhabit
7525:Marine protected area
7399:Regional fishery body
7025:Census of Marine Life
6920:Catch per unit effort
6220:Undulatory locomotion
6037:Ampullae of Lorenzini
5593:Marine protected area
5520:Salt pannes and pools
5295:Marine larval ecology
5270:Census of Marine Life
5154:Deep scattering layer
5111:San Francisco Estuary
5076:Africa and Madagascar
4901:Underwater camouflage
4681:Aquatic biomonitoring
4621:freshwater ecosystems
4420:Coral reef protection
4384:Skeletal eroding band
4190:Catlin Seaview Survey
4178:Census of Coral Reefs
4156:mesophotic coral reef
3883:and Murphy GI (eds.)
3791:: e.T60217A12323052.
3360:Gymnothorax javanicus
3293:Lactophrys bicaudalis
2484:Chaetodon capistratus
2080:
2066:
2037:
1964:
1882:
1839:are dominated by the
1811:
1621:
1487:
1289:
1112:
635:
537:foureye butterflyfish
486:tasseled scorpionfish
457:foureye butterflyfish
404:The well camouflaged
341:with silvery colours.
335:Atlantic bluefin tuna
293:Reef fish adaptations
208:
93:
78:including the use of
35:
7364:Minimum landing size
7324:Fisheries management
7299:MagnusonβStevens Act
7294:Fisheries Convention
7045:Fisheries scientists
6448:Genetically modified
5328:Marine bacteriophage
5290:Marine invertebrates
4530:Aquaculture of coral
4313:Non-tropical regions
3527:Caribbean Reef Shark
3370:. July 2009 version.
3303:. July 2009 version.
3275:. July 2009 version.
3217:. July 2009 version.
3144:. July 2009 version.
2991:10.14411/fp.2010.032
2703:Allen, G.R. (1985).
2608:Dakin, Nick (1992).
2567:Antennarius striatus
2545:Antennarius striatus
2494:. July 2009 version.
2091:Caribbean reef shark
2083:Caribbean reef shark
2027:, grey reef sharks,
1967:blacktip reef sharks
1884:Whitetip reef sharks
1841:Caribbean reef shark
1804:Reef sharks and rays
1798:harmful algal blooms
1728:poisoning if eaten.
1494:Pseudorhabdosynochus
1462:Yellowtail clownfish
1447:Saddle butterflyfish
1420:saddle butterflyfish
1294:, safely perched on
1180:such as parrotfish.
744:and coral groupers (
569:tassled scorpionfish
7641:Defying Ocean's End
7545:Salmon conservation
7535:Marine conservation
7475:The End of the Line
7468:The Sunken Billions
7437:Sustainable seafood
7422:Sustainable fishery
7035:Fisheries databases
6965:Age class structure
6935:Fisheries acoustics
6253:Aquatic respiration
6140:Life history theory
5204:Ocean fertilization
5013:Trophic state index
4971:Lake stratification
4701:Aquatic respiration
4448:Coral Reef Alliance
4338:Solomon Archipelago
4243:Belize Barrier Reef
4218:African coral reefs
3843:Randall, J. (1997)
3777:Carcharhinus perezi
3736:Carcharhinus perezi
3711:2006MEPS..320..239P
3207:Synanceja verrucosa
2412:Alevizon WS (1994)
2015:, and also feed on
1975:blacktip reef shark
1864:whitetip reef shark
1814:whitetip reef shark
1120:, often feeding in
861:schooling behaviour
551:The frogfish is an
191:The usually placid
142:adaptive behaviours
98:displaying spines.
7730:Outline of fishing
7349:Fisheries observer
7000:Aquatic ecosystems
6092:Weberian apparatus
5470:Intertidal wetland
5465:Intertidal ecology
5333:Marine prokaryotes
5275:Deep-sea community
5169:Iron fertilization
5092:Specific examples
5018:Upland and lowland
4936:Freshwater biology
4801:Microbial food web
4711:Aquatic toxicology
4654:Aquatic adaptation
4607:Aquatic ecosystems
4394:White band disease
4379:Black band disease
4348:Tuvalu Archipelago
4328:Raja Ampat Islands
4268:Great Barrier Reef
3924:Coral Reef Studies
3720:10.3354/meps320239
3319:2010-02-04 at the
3185:The New York Times
3160:2009-10-30 at the
2947:2016-03-07 at the
2829:The New York Times
2452:10.2108/zsj.12.811
2439:Zoological Science
2087:
2043:
1971:
1888:
1818:
1627:
1503:
1300:
1130:
1076:Two small cleaner
1056:followed by small
799:Bluestripe snapper
660:bluestripe snapper
645:
592:Feeding strategies
287:to attract mates.
211:
135:cryptic coloration
116:
108:blue-green chromis
42:
7756:
7755:
7752:
7751:
7748:
7747:
7712:Threatened sharks
7611:Ocean Conservancy
7596:Friend of the Sea
7226:
7225:
7222:
7221:
7134:Lobster fisheries
7079:Diversity of fish
6875:Shifting baseline
6844:Fisheries science
6796:
6795:
6706:Fish common names
6617:
6616:
6248:Aquatic predation
6072:Capacity for pain
5801:Age determination
5655:
5654:
5636:Oceans portal
5605:
5604:
5601:
5600:
5480:Hydrothermal vent
5416:
5415:
5305:Seashore wildlife
5136:Marine ecosystems
5125:
5124:
5121:
5120:
4891:Thermal pollution
4856:Ramsar Convention
4796:Microbial ecology
4756:Fisheries science
4696:Aquatic predation
4573:
4572:
4481:
4480:
4399:White pox disease
4248:Coral Sea Islands
3839:978-0-13-100847-2
3639:10.1890/08-0491.1
3460:Triaenodon obesus
3246:Australian Museum
2060:Shark researcher
1853:tawny nurse shark
1759:spotted trunkfish
1702:spotted trunkfish
1401:
1400:
1170:primary producers
1054:titan triggerfish
1031:Emperor angelfish
1003:cleaning stations
995:
994:
942:titan triggerfish
509:clown triggerfish
491:striated frogfish
431:Clown triggerfish
406:striated frogfish
127:ray-finned fishes
112:cauliflower coral
16:(Redirected from
7791:
7692:Marine pollution
7496:Cetacean bycatch
7491:Incidental catch
7311:
7257:
7256:
7232:
7231:
7199:Humboldt current
7057:
6975:Trophic cascades
6915:Data storage tag
6900:Fish measurement
6895:Stock assessment
6856:
6855:
6839:
6838:
6823:
6816:
6809:
6800:
6799:
6784:
6783:
6772:
6771:
6754:
6753:
6488:
6487:
5780:
5711:Ethnoichthyology
5682:
5675:
5668:
5659:
5658:
5645:
5644:
5638:
5634:
5633:
5624:
5622:Lakes portal
5620:
5619:
5618:
5588:Marine pollution
5280:Deep-water coral
5263:
5262:
5251:
5184:Marine chemistry
5144:
5143:
5131:
5130:
5048:Freshwater marsh
4941:Freshwater biome
4923:
4639:Acoustic ecology
4629:
4628:
4615:
4614:
4600:
4593:
4586:
4577:
4576:
4560:Deep-water coral
4535:Coral dermatitis
4436:
4435:
4151:Deep-water coral
4083:
3990:
3983:
3976:
3967:
3966:
3864:Academic Press.
3803:
3802:
3800:
3770:
3764:
3763:
3731:
3725:
3724:
3722:
3690:
3684:
3683:
3681:
3657:
3651:
3650:
3622:
3616:
3615:
3607:
3601:
3600:
3568:
3562:
3561:
3549:
3543:
3540:
3534:
3523:
3512:
3511:
3493:
3472:
3471:
3455:
3446:
3445:
3433:
3422:
3411:
3405:
3404:
3402:
3393:(8): 1423β1432.
3378:
3372:
3371:
3350:
3344:
3330:
3324:
3311:
3305:
3304:
3283:
3277:
3276:
3255:
3249:
3240:
3234:
3228:
3219:
3218:
3197:
3188:
3178:
3165:
3152:
3146:
3145:
3134:Pterois volitans
3124:
3118:
3117:
3116:
3110:
3100:
3090:
3066:
3053:
3052:
3034:
3025:
3014:
3013:
3012:
3003:
2993:
2969:
2958:
2957:
2956:
2938:
2929:
2928:
2900:
2894:
2893:
2876:(1β3): 513β520.
2863:
2857:
2851:
2845:
2842:
2833:
2820:
2814:
2803:
2797:
2796:
2786:
2776:
2752:
2746:
2745:
2727:
2721:
2720:
2700:
2694:
2684:
2669:
2668:
2640:
2634:
2633:
2605:
2599:
2598:
2577:
2571:
2570:www.frogfish.ch.
2563:
2557:
2556:
2535:
2529:
2519:
2496:
2495:
2471:
2465:
2464:
2454:
2430:
2424:
2410:
2393:
2390:
2384:
2383:
2362:
2356:
2355:
2338:
2332:
2326:
2320:
2319:
2284:
2278:
2272:
2266:
2260:
2254:
2251:
2245:
2241:
2235:
2232:
2226:
2223:
2214:
2211:
2202:
2199:
2164:
2159:
2158:
2157:
2140:great hammerhead
2135:
2116:yellow stingrays
2062:Leonard Compagno
2046:Grey reef sharks
2040:grey reef sharks
1956:Galapagos sharks
1829:grey reef sharks
1737:
1717:
1697:
1681:
1671:Stokes' seasnake
1477:Common clownfish
1473:
1464:with sea anemone
1458:
1443:
1431:
1394:
1393:
1382:
1381:
1370:
1369:
1365:alimentary tract
1348:and other small
1273:
1257:
1241:
1225:
1092:
1082:cleaning station
1073:
1049:
1026:
1001:, and establish
988:
987:
976:
975:
964:
963:
948:buried in sand.
929:
914:
899:
883:
822:
810:
795:
775:
759:, who school in
730:and other small
580:
564:
548:
532:
482:ambush predators
471:The psychedelic
427:
401:
384:The psychedelic
381:
326:
310:
273:
253:
233:
188:
173:
59:with tremendous
21:
7799:
7798:
7794:
7793:
7792:
7790:
7789:
7788:
7784:Fish by habitat
7759:
7758:
7757:
7744:
7721:
7717:Threatened rays
7697:Mercury in fish
7670:
7569:
7555:Shark sanctuary
7513:
7410:
7344:Catch reporting
7312:
7303:
7251:
7218:
7124:Krill fisheries
7089:Coral reef fish
7074:Ocean fisheries
7065:
7058:
7049:
6890:Condition index
6861:
6850:
6833:
6827:
6797:
6792:
6760:
6684:
6613:
6524:
6477:
6416:
6347:
6240:
6234:
6186:
6130:Ichthyoplankton
6096:
6028:
6021:
6017:Digital Library
6012:Teleost leptins
5951:Shark cartilage
5875:pharyngeal slit
5870:pharyngeal arch
5806:Anguilliformity
5791:
5789:
5781:
5772:
5691:
5686:
5656:
5651:
5628:
5627:
5616:
5614:
5613:
5597:
5558:Coral bleaching
5544:
5525:Seagrass meadow
5422:Marine habitats
5412:
5386:Coral reef fish
5352:
5338:Marine protists
5314:
5252:
5243:
5214:Ocean turbidity
5189:Marine food web
5138:
5117:
5059:
4998:River ecosystem
4951:Freshwater fish
4924:
4915:
4721:Bioluminescence
4706:Aquatic science
4623:
4609:
4604:
4574:
4569:
4555:Coralline algae
4525:Artificial reef
4513:
4486:Symbiotic algae
4477:
4434:
4430:Reef resilience
4403:
4374:Coral bleaching
4362:
4228:Andros, Bahamas
4199:
4173:Coral reef fish
4122:
4084:
4075:
4003:
3994:
3944:
3906:
3875:Sale PF (1982)
3858:Sale PF (2006)
3811:
3806:
3771:
3767:
3732:
3728:
3691:
3687:
3658:
3654:
3633:(4): 996β1008.
3623:
3619:
3608:
3604:
3569:
3565:
3550:
3546:
3541:
3537:
3524:
3515:
3508:
3494:
3475:
3464:Pacific Science
3456:
3449:
3438:Pacific Science
3434:
3425:
3412:
3408:
3379:
3375:
3351:
3347:
3331:
3327:
3321:Wayback Machine
3312:
3308:
3284:
3280:
3256:
3252:
3241:
3237:
3229:
3222:
3198:
3191:
3187:22 August 2006.
3179:
3168:
3162:Wayback Machine
3153:
3149:
3125:
3121:
3111:
3067:
3056:
3032:
3026:
3017:
3007:
2970:
2961:
2951:
2949:Wayback Machine
2939:
2932:
2901:
2897:
2864:
2860:
2852:
2848:
2843:
2836:
2821:
2817:
2804:
2800:
2753:
2749:
2742:
2728:
2724:
2717:
2701:
2697:
2685:
2672:
2641:
2637:
2630:
2606:
2602:
2578:
2574:
2564:
2560:
2536:
2532:
2520:
2499:
2472:
2468:
2431:
2427:
2411:
2396:
2391:
2387:
2363:
2359:
2339:
2335:
2327:
2323:
2289:Current Biology
2285:
2281:
2273:
2269:
2261:
2257:
2252:
2248:
2242:
2238:
2233:
2229:
2224:
2217:
2212:
2205:
2200:
2193:
2189:
2160:
2155:
2153:
2150:
2143:
2136:
1991:grey reef shark
1849:Galapagos shark
1806:
1745:
1738:
1729:
1718:
1709:
1698:
1689:
1682:
1666:ambush predator
1616:
1541:Platyhelminthes
1480:
1474:
1465:
1459:
1450:
1444:
1435:
1432:
1391:
1379:
1373:External images
1284:
1277:
1274:
1265:
1258:
1249:
1242:
1233:
1226:
1107:
1100:
1097:reef lizardfish
1093:
1084:
1074:
1065:
1050:
1041:
1027:
1014:reef lizardfish
985:
973:
936:
930:
921:
918:Bigeye trevally
915:
906:
900:
891:
884:
871:Bigeye trevally
837:
830:
823:
814:
811:
802:
796:
787:
776:
630:
602:butterflyfishes
594:
587:
581:
572:
565:
556:
553:ambush predator
549:
540:
533:
438:
437:
436:
435:
434:
428:
419:
418:
417:
414:ambush predator
408:, a species of
402:
393:
392:
391:
382:
371:
351:
346:
345:
344:
343:
342:
331:open water fish
327:
319:
318:
315:queen angelfish
311:
302:
301:
295:
288:
274:
265:
254:
245:
234:
203:
196:
189:
180:
174:
88:
45:Coral reef fish
28:
23:
22:
15:
12:
11:
5:
7797:
7787:
7786:
7781:
7776:
7771:
7754:
7753:
7750:
7749:
7746:
7745:
7743:
7742:
7737:
7732:
7726:
7723:
7722:
7720:
7719:
7714:
7709:
7704:
7699:
7694:
7689:
7687:Fish slaughter
7684:
7678:
7676:
7675:Related issues
7672:
7671:
7669:
7668:
7663:
7658:
7653:
7648:
7643:
7638:
7633:
7628:
7623:
7618:
7616:Ocean Outcomes
7613:
7608:
7603:
7598:
7593:
7588:
7583:
7577:
7575:
7571:
7570:
7568:
7567:
7562:
7557:
7552:
7547:
7542:
7537:
7532:
7530:Marine reserve
7527:
7521:
7519:
7515:
7514:
7512:
7511:
7503:
7498:
7493:
7488:
7483:
7478:
7471:
7464:
7459:
7454:
7449:
7444:
7439:
7434:
7429:
7424:
7418:
7416:
7415:Sustainability
7412:
7411:
7409:
7408:
7407:
7406:
7396:
7391:
7386:
7381:
7376:
7371:
7366:
7361:
7356:
7351:
7346:
7341:
7336:
7331:
7326:
7320:
7318:
7314:
7313:
7306:
7304:
7302:
7301:
7296:
7291:
7290:
7289:
7279:
7274:
7269:
7263:
7261:
7253:
7252:
7245:sustainability
7228:
7227:
7224:
7223:
7220:
7219:
7217:
7216:
7211:
7206:
7201:
7196:
7191:
7186:
7181:
7179:Marine habitat
7176:
7171:
7166:
7161:
7156:
7151:
7146:
7141:
7139:Shrimp fishery
7136:
7131:
7129:Kelp fisheries
7126:
7121:
7116:
7114:Crab fisheries
7111:
7106:
7101:
7096:
7091:
7086:
7081:
7076:
7070:
7068:
7060:
7059:
7052:
7050:
7048:
7047:
7042:
7037:
7032:
7027:
7022:
7017:
7012:
7007:
7002:
6997:
6995:Marine biology
6992:
6987:
6982:
6977:
6972:
6967:
6962:
6957:
6952:
6947:
6945:Coded wire tag
6942:
6937:
6932:
6927:
6922:
6917:
6912:
6907:
6902:
6897:
6892:
6887:
6885:Fish mortality
6882:
6877:
6872:
6866:
6864:
6852:
6851:
6848:wild fisheries
6835:
6834:
6832:science topics
6826:
6825:
6818:
6811:
6803:
6794:
6793:
6791:
6790:
6778:
6765:
6762:
6761:
6759:
6758:
6748:
6743:
6742:
6741:
6736:
6728:
6723:
6718:
6713:
6708:
6703:
6698:
6692:
6690:
6686:
6685:
6683:
6682:
6681:
6680:
6675:
6665:
6664:
6663:
6658:
6653:
6643:
6642:
6641:
6636:
6625:
6623:
6619:
6618:
6615:
6614:
6612:
6611:
6610:
6609:
6604:
6599:
6589:
6588:
6587:
6582:
6577:
6572:
6562:
6561:
6560:
6555:
6550:
6545:
6534:
6532:
6530:Wild fisheries
6526:
6525:
6523:
6522:
6517:
6512:
6507:
6502:
6496:
6494:
6485:
6479:
6478:
6476:
6475:
6470:
6465:
6460:
6455:
6453:Hallucinogenic
6450:
6445:
6440:
6435:
6430:
6424:
6422:
6418:
6417:
6415:
6414:
6409:
6404:
6399:
6394:
6389:
6384:
6379:
6374:
6369:
6364:
6358:
6356:
6349:
6348:
6346:
6345:
6340:
6335:
6330:
6328:Schooling fish
6325:
6320:
6315:
6310:
6305:
6300:
6295:
6290:
6288:Filter feeders
6285:
6280:
6275:
6270:
6265:
6263:Bottom feeders
6260:
6255:
6250:
6244:
6242:
6236:
6235:
6233:
6232:
6227:
6222:
6217:
6212:
6207:
6202:
6196:
6194:
6188:
6187:
6185:
6184:
6183:
6182:
6172:
6167:
6162:
6157:
6152:
6147:
6142:
6137:
6132:
6127:
6122:
6117:
6112:
6106:
6104:
6098:
6097:
6095:
6094:
6089:
6084:
6079:
6074:
6069:
6064:
6059:
6054:
6049:
6044:
6039:
6033:
6031:
6023:
6022:
6020:
6019:
6014:
6009:
6008:
6007:
6002:
5992:
5991:
5990:
5985:
5975:
5970:
5965:
5964:
5963:
5953:
5948:
5943:
5938:
5933:
5932:
5931:
5921:
5916:
5911:
5909:Leydig's organ
5906:
5905:
5904:
5902:pharyngeal jaw
5899:
5889:
5884:
5883:
5882:
5877:
5872:
5867:
5862:
5857:
5855:branchial arch
5847:
5846:
5845:
5835:
5830:
5825:
5824:
5823:
5818:
5808:
5803:
5797:
5795:
5783:
5782:
5775:
5773:
5771:
5770:
5765:
5760:
5755:
5750:
5745:
5744:
5743:
5738:
5733:
5723:
5718:
5713:
5708:
5702:
5700:
5693:
5692:
5685:
5684:
5677:
5670:
5662:
5653:
5652:
5650:
5649:
5639:
5625:
5610:
5607:
5606:
5603:
5602:
5599:
5598:
5596:
5595:
5590:
5585:
5580:
5575:
5570:
5565:
5560:
5554:
5552:
5546:
5545:
5543:
5542:
5537:
5532:
5527:
5522:
5517:
5512:
5507:
5502:
5497:
5492:
5487:
5482:
5477:
5472:
5467:
5462:
5457:
5452:
5447:
5442:
5437:
5432:
5426:
5424:
5418:
5417:
5414:
5413:
5411:
5410:
5405:
5404:
5403:
5398:
5393:
5388:
5383:
5376:Saltwater fish
5373:
5371:Marine reptile
5368:
5362:
5360:
5354:
5353:
5351:
5350:
5345:
5343:Marine viruses
5340:
5335:
5330:
5324:
5322:
5320:Microorganisms
5316:
5315:
5313:
5312:
5310:Wild fisheries
5307:
5302:
5297:
5292:
5287:
5282:
5277:
5272:
5266:
5260:
5254:
5253:
5246:
5244:
5242:
5241:
5236:
5231:
5226:
5224:Thorson's rule
5221:
5216:
5211:
5206:
5201:
5196:
5191:
5186:
5181:
5179:Marine biology
5176:
5171:
5166:
5161:
5156:
5150:
5148:
5140:
5139:
5127:
5126:
5123:
5122:
5119:
5118:
5116:
5115:
5114:
5113:
5108:
5103:
5098:
5090:
5085:
5084:
5083:
5078:
5067:
5065:
5061:
5060:
5058:
5057:
5056:
5055:
5050:
5045:
5040:
5038:Brackish marsh
5035:
5025:
5020:
5015:
5010:
5005:
5000:
4995:
4990:
4989:
4988:
4978:
4973:
4968:
4966:Lake ecosystem
4963:
4958:
4956:Hyporheic zone
4953:
4948:
4943:
4938:
4932:
4930:
4926:
4925:
4918:
4916:
4914:
4913:
4908:
4903:
4898:
4893:
4888:
4883:
4878:
4873:
4868:
4863:
4858:
4853:
4848:
4843:
4838:
4833:
4828:
4823:
4818:
4813:
4808:
4806:Microbial loop
4803:
4798:
4793:
4788:
4783:
4778:
4773:
4768:
4763:
4758:
4753:
4751:Eutrophication
4748:
4743:
4738:
4733:
4731:Cascade effect
4728:
4723:
4718:
4713:
4708:
4703:
4698:
4693:
4688:
4683:
4678:
4677:
4676:
4671:
4666:
4659:Aquatic animal
4656:
4651:
4646:
4641:
4635:
4633:
4625:
4624:
4611:
4610:
4603:
4602:
4595:
4588:
4580:
4571:
4570:
4568:
4567:
4562:
4557:
4552:
4547:
4542:
4540:Precious coral
4537:
4532:
4527:
4521:
4519:
4515:
4514:
4512:
4511:
4510:
4509:
4502:
4489:
4487:
4483:
4482:
4479:
4478:
4476:
4475:
4470:
4465:
4460:
4455:
4450:
4444:
4442:
4433:
4432:
4427:
4422:
4417:
4411:
4409:
4405:
4404:
4402:
4401:
4396:
4391:
4386:
4381:
4376:
4370:
4368:
4367:Coral diseases
4364:
4363:
4361:
4360:
4355:
4353:Virgin Islands
4350:
4345:
4340:
4335:
4330:
4325:
4320:
4315:
4310:
4305:
4300:
4295:
4290:
4285:
4280:
4275:
4270:
4265:
4260:
4255:
4253:Coral Triangle
4250:
4245:
4240:
4235:
4230:
4225:
4220:
4215:
4209:
4207:
4201:
4200:
4198:
4197:
4192:
4187:
4180:
4175:
4170:
4165:
4160:
4159:
4158:
4148:
4143:
4138:
4132:
4130:
4124:
4123:
4121:
4120:
4115:
4110:
4105:
4100:
4094:
4092:
4086:
4085:
4078:
4076:
4074:
4073:
4067:
4061:
4060:
4059:
4054:
4044:
4039:
4034:
4029:
4024:
4019:
4013:
4011:
4005:
4004:
3993:
3992:
3985:
3978:
3970:
3964:
3963:
3958:
3952:
3926:
3917:
3912:
3905:
3904:External links
3902:
3901:
3900:
3873:
3856:
3841:
3827:
3810:
3807:
3805:
3804:
3765:
3746:(2): 189β199.
3740:Marine Biology
3726:
3685:
3652:
3617:
3602:
3583:(3): 647β654.
3563:
3544:
3535:
3513:
3506:
3473:
3447:
3423:
3421:, 18 May 2009.
3406:
3373:
3354:Froese, Rainer
3345:
3325:
3306:
3287:Froese, Rainer
3278:
3259:Froese, Rainer
3250:
3235:
3220:
3201:Froese, Rainer
3189:
3180:Grady, Denise
3166:
3147:
3128:Froese, Rainer
3119:
3054:
3015:
2959:
2930:
2895:
2858:
2846:
2834:
2815:
2798:
2747:
2740:
2722:
2715:
2695:
2670:
2651:(4): 579β590.
2635:
2628:
2600:
2581:Froese, Rainer
2572:
2558:
2539:Froese, Rainer
2530:
2497:
2478:Froese, Rainer
2466:
2445:(6): 811β813.
2425:
2394:
2385:
2366:Froese, Rainer
2357:
2342:Froese, Rainer
2333:
2321:
2279:
2267:
2265:, Buchheim, J.
2255:
2246:
2236:
2227:
2215:
2203:
2190:
2188:
2185:
2184:
2183:
2178:
2172:
2166:
2165:
2149:
2146:
2145:
2144:
2137:
2130:
2095:apex predators
1944:spiny lobsters
1912:squirrelfishes
1837:Atlantic Ocean
1805:
1802:
1747:
1746:
1739:
1732:
1730:
1719:
1712:
1710:
1699:
1692:
1690:
1686:reef stonefish
1683:
1676:
1662:reef stonefish
1655:bioprospecting
1635:poisonous fish
1615:
1612:
1586:) among which
1482:
1481:
1475:
1468:
1466:
1460:
1453:
1451:
1445:
1438:
1436:
1433:
1426:
1414:) primed with
1399:
1398:
1387:
1386:
1375:
1374:
1311:commensalistic
1283:
1280:
1279:
1278:
1275:
1268:
1266:
1259:
1252:
1250:
1243:
1236:
1234:
1232:feed on algae.
1227:
1220:
1106:
1103:
1102:
1101:
1094:
1087:
1085:
1075:
1068:
1066:
1051:
1044:
1042:
1028:
1021:
993:
992:
981:
980:
969:
968:
967:External image
938:
937:
931:
924:
922:
916:
909:
907:
901:
894:
892:
885:
878:
836:
833:
832:
831:
824:
817:
815:
812:
805:
803:
797:
790:
788:
777:
770:
757:hermaphrodites
629:
626:
604:are the coral
593:
590:
589:
588:
582:
575:
573:
566:
559:
557:
550:
543:
541:
534:
527:
521:countershading
450:warn predators
429:
422:
421:
420:
403:
396:
395:
394:
383:
376:
375:
374:
373:
372:
370:
367:
350:
347:
328:
321:
320:
312:
305:
304:
303:
299:
298:
297:
296:
294:
291:
290:
289:
275:
268:
266:
262:dwarf seahorse
255:
248:
246:
235:
228:
202:
199:
198:
197:
190:
183:
181:
175:
168:
87:
84:
26:
9:
6:
4:
3:
2:
7796:
7785:
7782:
7780:
7779:Tropical fish
7777:
7775:
7772:
7770:
7767:
7766:
7764:
7741:
7738:
7736:
7733:
7731:
7728:
7727:
7724:
7718:
7715:
7713:
7710:
7708:
7707:Shark finning
7705:
7703:
7702:Shark culling
7700:
7698:
7695:
7693:
7690:
7688:
7685:
7683:
7680:
7679:
7677:
7673:
7667:
7664:
7662:
7659:
7657:
7654:
7652:
7649:
7647:
7644:
7642:
7639:
7637:
7634:
7632:
7631:Sea Around Us
7629:
7627:
7624:
7622:
7619:
7617:
7614:
7612:
7609:
7607:
7606:Seafood Watch
7604:
7602:
7599:
7597:
7594:
7592:
7589:
7587:
7584:
7582:
7579:
7578:
7576:
7572:
7566:
7563:
7561:
7558:
7556:
7553:
7551:
7548:
7546:
7543:
7541:
7538:
7536:
7533:
7531:
7528:
7526:
7523:
7522:
7520:
7516:
7510:
7508:
7507:Shrimp-Turtle
7504:
7502:
7499:
7497:
7494:
7492:
7489:
7487:
7484:
7482:
7479:
7477:
7476:
7472:
7470:
7469:
7465:
7463:
7460:
7458:
7455:
7453:
7450:
7448:
7445:
7443:
7440:
7438:
7435:
7433:
7430:
7428:
7425:
7423:
7420:
7419:
7417:
7413:
7405:
7402:
7401:
7400:
7397:
7395:
7392:
7390:
7387:
7385:
7382:
7380:
7377:
7375:
7372:
7370:
7369:Pulse fishing
7367:
7365:
7362:
7360:
7357:
7355:
7352:
7350:
7347:
7345:
7342:
7340:
7337:
7335:
7332:
7330:
7327:
7325:
7322:
7321:
7319:
7315:
7310:
7300:
7297:
7295:
7292:
7288:
7285:
7284:
7283:
7280:
7278:
7275:
7273:
7270:
7268:
7267:Fisheries law
7265:
7264:
7262:
7258:
7254:
7250:
7246:
7242:
7238:
7237:Fisheries law
7233:
7229:
7215:
7212:
7210:
7207:
7205:
7202:
7200:
7197:
7195:
7192:
7190:
7187:
7185:
7182:
7180:
7177:
7175:
7172:
7170:
7167:
7165:
7162:
7160:
7157:
7155:
7152:
7150:
7147:
7145:
7142:
7140:
7137:
7135:
7132:
7130:
7127:
7125:
7122:
7120:
7119:Eel fisheries
7117:
7115:
7112:
7110:
7109:Cod fisheries
7107:
7105:
7102:
7100:
7097:
7095:
7094:Demersal fish
7092:
7090:
7087:
7085:
7082:
7080:
7077:
7075:
7072:
7071:
7069:
7067:
7061:
7056:
7046:
7043:
7041:
7038:
7036:
7033:
7031:
7028:
7026:
7023:
7021:
7018:
7016:
7013:
7011:
7008:
7006:
7003:
7001:
6998:
6996:
6993:
6991:
6988:
6986:
6983:
6981:
6978:
6976:
6973:
6971:
6970:Trophic level
6968:
6966:
6963:
6961:
6958:
6956:
6953:
6951:
6948:
6946:
6943:
6941:
6938:
6936:
6933:
6931:
6928:
6926:
6923:
6921:
6918:
6916:
6913:
6911:
6908:
6906:
6903:
6901:
6898:
6896:
6893:
6891:
6888:
6886:
6883:
6881:
6878:
6876:
6873:
6871:
6868:
6867:
6865:
6863:
6857:
6853:
6849:
6845:
6840:
6836:
6831:
6824:
6819:
6817:
6812:
6810:
6805:
6804:
6801:
6789:
6788:
6779:
6777:
6776:
6767:
6766:
6763:
6757:
6756:more lists...
6749:
6747:
6744:
6740:
6737:
6735:
6732:
6731:
6729:
6727:
6724:
6722:
6719:
6717:
6714:
6712:
6711:Fish families
6709:
6707:
6704:
6702:
6699:
6697:
6696:Aquarium life
6694:
6693:
6691:
6687:
6679:
6678:fleshy-finned
6676:
6674:
6671:
6670:
6669:
6666:
6662:
6659:
6657:
6654:
6652:
6649:
6648:
6647:
6646:Cartilaginous
6644:
6640:
6637:
6635:
6632:
6631:
6630:
6627:
6626:
6624:
6620:
6608:
6605:
6603:
6600:
6598:
6595:
6594:
6593:
6590:
6586:
6583:
6581:
6578:
6576:
6573:
6571:
6568:
6567:
6566:
6563:
6559:
6556:
6554:
6551:
6549:
6546:
6544:
6541:
6540:
6539:
6536:
6535:
6533:
6531:
6527:
6521:
6518:
6516:
6513:
6511:
6508:
6506:
6503:
6501:
6498:
6497:
6495:
6493:
6489:
6486:
6484:
6480:
6474:
6471:
6469:
6466:
6464:
6461:
6459:
6456:
6454:
6451:
6449:
6446:
6444:
6441:
6439:
6436:
6434:
6431:
6429:
6426:
6425:
6423:
6419:
6413:
6410:
6408:
6405:
6403:
6400:
6398:
6395:
6393:
6390:
6388:
6385:
6383:
6380:
6378:
6375:
6373:
6370:
6368:
6365:
6363:
6360:
6359:
6357:
6355:
6350:
6344:
6341:
6339:
6336:
6334:
6331:
6329:
6326:
6324:
6321:
6319:
6316:
6314:
6311:
6309:
6306:
6304:
6301:
6299:
6296:
6294:
6291:
6289:
6286:
6284:
6283:Electric fish
6281:
6279:
6276:
6274:
6271:
6269:
6266:
6264:
6261:
6259:
6256:
6254:
6251:
6249:
6246:
6245:
6243:
6237:
6231:
6228:
6226:
6223:
6221:
6218:
6216:
6213:
6211:
6208:
6206:
6203:
6201:
6198:
6197:
6195:
6193:
6189:
6181:
6178:
6177:
6176:
6173:
6171:
6168:
6166:
6163:
6161:
6158:
6156:
6153:
6151:
6148:
6146:
6143:
6141:
6138:
6136:
6133:
6131:
6128:
6126:
6123:
6121:
6118:
6116:
6113:
6111:
6108:
6107:
6105:
6103:
6099:
6093:
6090:
6088:
6085:
6083:
6080:
6078:
6075:
6073:
6070:
6068:
6065:
6063:
6060:
6058:
6055:
6053:
6050:
6048:
6045:
6043:
6040:
6038:
6035:
6034:
6032:
6030:
6024:
6018:
6015:
6013:
6010:
6006:
6003:
6001:
5998:
5997:
5996:
5993:
5989:
5986:
5984:
5981:
5980:
5979:
5976:
5974:
5971:
5969:
5966:
5962:
5959:
5958:
5957:
5954:
5952:
5949:
5947:
5944:
5942:
5939:
5937:
5934:
5930:
5927:
5926:
5925:
5922:
5920:
5917:
5915:
5914:Mauthner cell
5912:
5910:
5907:
5903:
5900:
5898:
5895:
5894:
5893:
5890:
5888:
5885:
5881:
5878:
5876:
5873:
5871:
5868:
5866:
5863:
5861:
5858:
5856:
5853:
5852:
5851:
5848:
5844:
5841:
5840:
5839:
5836:
5834:
5833:Chromatophore
5831:
5829:
5826:
5822:
5819:
5817:
5814:
5813:
5812:
5809:
5807:
5804:
5802:
5799:
5798:
5796:
5794:
5788:
5784:
5779:
5769:
5766:
5764:
5761:
5759:
5756:
5754:
5751:
5749:
5746:
5742:
5739:
5737:
5734:
5732:
5729:
5728:
5727:
5724:
5722:
5719:
5717:
5714:
5712:
5709:
5707:
5704:
5703:
5701:
5699:
5694:
5690:
5683:
5678:
5676:
5671:
5669:
5664:
5663:
5660:
5648:
5640:
5637:
5632:
5626:
5623:
5612:
5611:
5608:
5594:
5591:
5589:
5586:
5584:
5581:
5579:
5576:
5574:
5571:
5569:
5566:
5564:
5561:
5559:
5556:
5555:
5553:
5551:
5547:
5541:
5538:
5536:
5533:
5531:
5530:Sponge ground
5528:
5526:
5523:
5521:
5518:
5516:
5513:
5511:
5508:
5506:
5503:
5501:
5498:
5496:
5495:Marine biomes
5493:
5491:
5488:
5486:
5483:
5481:
5478:
5476:
5473:
5471:
5468:
5466:
5463:
5461:
5458:
5456:
5453:
5451:
5448:
5446:
5443:
5441:
5438:
5436:
5433:
5431:
5428:
5427:
5425:
5423:
5419:
5409:
5406:
5402:
5399:
5397:
5396:Demersal fish
5394:
5392:
5391:Deep-sea fish
5389:
5387:
5384:
5382:
5379:
5378:
5377:
5374:
5372:
5369:
5367:
5366:Marine mammal
5364:
5363:
5361:
5359:
5355:
5349:
5346:
5344:
5341:
5339:
5336:
5334:
5331:
5329:
5326:
5325:
5323:
5321:
5317:
5311:
5308:
5306:
5303:
5301:
5298:
5296:
5293:
5291:
5288:
5286:
5283:
5281:
5278:
5276:
5273:
5271:
5268:
5267:
5264:
5261:
5259:
5255:
5250:
5240:
5237:
5235:
5232:
5230:
5227:
5225:
5222:
5220:
5217:
5215:
5212:
5210:
5207:
5205:
5202:
5200:
5197:
5195:
5192:
5190:
5187:
5185:
5182:
5180:
5177:
5175:
5172:
5170:
5167:
5165:
5162:
5160:
5157:
5155:
5152:
5151:
5149:
5145:
5141:
5137:
5132:
5128:
5112:
5109:
5107:
5104:
5102:
5099:
5097:
5094:
5093:
5091:
5089:
5086:
5082:
5079:
5077:
5074:
5073:
5072:
5069:
5068:
5066:
5062:
5054:
5051:
5049:
5046:
5044:
5041:
5039:
5036:
5034:
5031:
5030:
5029:
5026:
5024:
5021:
5019:
5016:
5014:
5011:
5009:
5006:
5004:
5001:
4999:
4996:
4994:
4991:
4987:
4984:
4983:
4982:
4979:
4977:
4974:
4972:
4969:
4967:
4964:
4962:
4959:
4957:
4954:
4952:
4949:
4947:
4944:
4942:
4939:
4937:
4934:
4933:
4931:
4927:
4922:
4912:
4909:
4907:
4904:
4902:
4899:
4897:
4896:Trophic level
4894:
4892:
4889:
4887:
4884:
4882:
4879:
4877:
4874:
4872:
4869:
4867:
4864:
4862:
4861:Sediment trap
4859:
4857:
4854:
4852:
4849:
4847:
4844:
4842:
4839:
4837:
4836:Phytoplankton
4834:
4832:
4829:
4827:
4824:
4822:
4819:
4817:
4814:
4812:
4809:
4807:
4804:
4802:
4799:
4797:
4794:
4792:
4789:
4787:
4784:
4782:
4779:
4777:
4774:
4772:
4769:
4767:
4764:
4762:
4759:
4757:
4754:
4752:
4749:
4747:
4744:
4742:
4739:
4737:
4734:
4732:
4729:
4727:
4724:
4722:
4719:
4717:
4714:
4712:
4709:
4707:
4704:
4702:
4699:
4697:
4694:
4692:
4689:
4687:
4686:Aquatic plant
4684:
4682:
4679:
4675:
4672:
4670:
4667:
4665:
4662:
4661:
4660:
4657:
4655:
4652:
4650:
4649:Anoxic waters
4647:
4645:
4642:
4640:
4637:
4636:
4634:
4630:
4626:
4622:
4616:
4612:
4608:
4601:
4596:
4594:
4589:
4587:
4582:
4581:
4578:
4566:
4563:
4561:
4558:
4556:
4553:
4551:
4548:
4546:
4543:
4541:
4538:
4536:
4533:
4531:
4528:
4526:
4523:
4522:
4520:
4516:
4508:
4507:
4503:
4501:
4500:
4496:
4495:
4494:
4493:Zooxanthellae
4491:
4490:
4488:
4484:
4474:
4471:
4469:
4466:
4464:
4463:Project AWARE
4461:
4459:
4456:
4454:
4451:
4449:
4446:
4445:
4443:
4441:
4440:Organizations
4437:
4431:
4428:
4426:
4423:
4421:
4418:
4416:
4413:
4412:
4410:
4406:
4400:
4397:
4395:
4392:
4390:
4387:
4385:
4382:
4380:
4377:
4375:
4372:
4371:
4369:
4365:
4359:
4356:
4354:
4351:
4349:
4346:
4344:
4341:
4339:
4336:
4334:
4331:
4329:
4326:
4324:
4321:
4319:
4316:
4314:
4311:
4309:
4308:Ningaloo Reef
4306:
4304:
4301:
4299:
4296:
4294:
4291:
4289:
4286:
4284:
4281:
4279:
4276:
4274:
4271:
4269:
4266:
4264:
4261:
4259:
4256:
4254:
4251:
4249:
4246:
4244:
4241:
4239:
4236:
4234:
4231:
4229:
4226:
4224:
4221:
4219:
4216:
4214:
4213:List of reefs
4211:
4210:
4208:
4206:
4205:Coral regions
4202:
4196:
4193:
4191:
4188:
4186:
4185:
4181:
4179:
4176:
4174:
4171:
4169:
4166:
4164:
4161:
4157:
4154:
4153:
4152:
4149:
4147:
4144:
4142:
4139:
4137:
4134:
4133:
4131:
4129:
4125:
4119:
4116:
4114:
4111:
4109:
4106:
4104:
4101:
4099:
4096:
4095:
4093:
4091:
4087:
4082:
4072:
4068:
4066:
4062:
4058:
4055:
4053:
4050:
4049:
4048:
4045:
4043:
4040:
4038:
4035:
4033:
4030:
4028:
4025:
4023:
4020:
4018:
4015:
4014:
4012:
4010:
4006:
4002:
3998:
3991:
3986:
3984:
3979:
3977:
3972:
3971:
3968:
3962:
3959:
3956:
3953:
3951:
3947:
3942:
3938:
3937:
3932:
3927:
3925:
3921:
3918:
3916:
3913:
3911:
3908:
3907:
3898:
3897:971-0400-22-3
3894:
3890:
3886:
3882:
3878:
3874:
3871:
3870:0-12-373609-9
3867:
3863:
3862:
3857:
3854:
3853:0-8248-1895-4
3850:
3846:
3842:
3840:
3836:
3832:
3828:
3825:
3824:0-691-08995-7
3821:
3817:
3813:
3812:
3799:
3794:
3790:
3786:
3785:
3780:
3778:
3769:
3761:
3757:
3753:
3749:
3745:
3741:
3737:
3730:
3721:
3716:
3712:
3708:
3704:
3700:
3696:
3689:
3680:
3675:
3671:
3667:
3663:
3656:
3648:
3644:
3640:
3636:
3632:
3628:
3621:
3613:
3606:
3598:
3594:
3590:
3586:
3582:
3578:
3574:
3567:
3559:
3555:
3548:
3539:
3532:
3528:
3525:Martin, R.A.
3522:
3520:
3518:
3509:
3507:92-5-101384-5
3503:
3499:
3492:
3490:
3488:
3486:
3484:
3482:
3480:
3478:
3470:(2): 143β164.
3469:
3465:
3461:
3454:
3452:
3443:
3439:
3432:
3430:
3428:
3420:
3419:
3415:
3410:
3401:
3396:
3392:
3388:
3384:
3377:
3369:
3368:
3363:
3361:
3355:
3349:
3343:
3342:0-00-715986-2
3339:
3335:
3329:
3322:
3318:
3315:
3310:
3302:
3301:
3296:
3294:
3288:
3282:
3274:
3273:
3268:
3266:
3260:
3254:
3247:
3244:
3239:
3232:
3227:
3225:
3216:
3215:
3210:
3208:
3202:
3196:
3194:
3186:
3183:
3177:
3175:
3173:
3171:
3163:
3159:
3156:
3151:
3143:
3142:
3137:
3135:
3129:
3123:
3115:
3108:
3104:
3099:
3094:
3089:
3084:
3080:
3076:
3075:Aquat Biosyst
3072:
3065:
3063:
3061:
3059:
3050:
3046:
3042:
3038:
3031:
3024:
3022:
3020:
3011:
3006:
3001:
2997:
2992:
2987:
2984:(4): 237β62.
2983:
2979:
2975:
2968:
2966:
2964:
2955:
2950:
2946:
2943:
2937:
2935:
2926:
2922:
2918:
2914:
2910:
2906:
2899:
2891:
2887:
2883:
2879:
2875:
2871:
2870:
2869:Hydrobiologia
2862:
2855:
2850:
2841:
2839:
2831:
2830:
2825:
2819:
2812:
2808:
2802:
2794:
2790:
2785:
2780:
2775:
2770:
2766:
2762:
2758:
2751:
2743:
2741:0-12-547665-5
2737:
2733:
2726:
2718:
2716:92-5-102321-2
2712:
2708:
2707:
2699:
2692:
2688:
2683:
2681:
2679:
2677:
2675:
2666:
2662:
2658:
2654:
2650:
2646:
2639:
2631:
2629:0-02-897108-6
2625:
2621:
2617:
2613:
2612:
2604:
2596:
2595:
2590:
2588:
2582:
2576:
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2568:
2562:
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2475:
2470:
2462:
2458:
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2448:
2444:
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2436:
2429:
2423:
2422:1-55992-077-7
2419:
2415:
2409:
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2403:
2401:
2399:
2389:
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2380:
2375:
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2343:
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2325:
2318:
2314:
2310:
2306:
2302:
2298:
2295:(8): R288-9,
2294:
2290:
2283:
2276:
2271:
2264:
2259:
2250:
2240:
2231:
2222:
2220:
2210:
2208:
2198:
2196:
2191:
2182:
2179:
2176:
2173:
2171:
2168:
2167:
2163:
2152:
2141:
2134:
2129:
2128:
2127:
2125:
2121:
2117:
2113:
2109:
2108:elasmobranchs
2105:
2101:
2096:
2092:
2084:
2079:
2074:
2071:
2065:
2063:
2058:
2056:
2052:
2047:
2041:
2036:
2032:
2030:
2026:
2022:
2018:
2014:
2010:
2006:
2002:
1997:
1996:Palmyra Atoll
1992:
1988:
1984:
1980:
1976:
1968:
1963:
1959:
1957:
1953:
1949:
1945:
1941:
1938:, as well as
1937:
1933:
1932:triggerfishes
1929:
1928:surgeonfishes
1925:
1921:
1917:
1913:
1909:
1905:
1901:
1896:
1894:
1885:
1881:
1877:
1874:
1870:
1865:
1860:
1858:
1854:
1850:
1846:
1845:requiem shark
1842:
1838:
1834:
1831:dominate the
1830:
1826:
1822:
1815:
1810:
1801:
1799:
1795:
1791:
1787:
1782:
1780:
1776:
1771:
1768:
1764:
1760:
1755:
1752:
1743:
1736:
1731:
1727:
1723:
1716:
1711:
1707:
1703:
1696:
1691:
1687:
1680:
1675:
1674:
1673:
1672:
1667:
1663:
1658:
1656:
1650:
1648:
1644:
1640:
1639:venomous fish
1636:
1632:
1625:
1620:
1611:
1609:
1608:New Caledonia
1605:
1604:co-extinction
1601:
1597:
1593:
1589:
1585:
1581:
1578:
1574:
1573:microsporidia
1570:
1566:
1562:
1558:
1554:
1550:
1546:
1542:
1538:
1534:
1530:
1525:
1523:
1520:
1516:
1512:
1508:
1500:
1496:
1495:
1490:
1486:
1478:
1472:
1467:
1463:
1457:
1452:
1448:
1442:
1437:
1430:
1425:
1424:
1423:
1421:
1417:
1413:
1409:
1405:
1397:
1388:
1385:
1376:
1371:
1368:
1366:
1362:
1358:
1353:
1351:
1350:invertebrates
1347:
1343:
1339:
1335:
1331:
1327:
1323:
1318:
1316:
1312:
1308:
1304:
1297:
1293:
1288:
1272:
1267:
1263:
1256:
1251:
1247:
1240:
1235:
1231:
1224:
1219:
1218:
1217:
1215:
1211:
1207:
1203:
1199:
1197:
1193:
1188:
1185:
1181:
1179:
1175:
1171:
1167:
1163:
1161:
1158:
1154:
1150:
1149:surgeonfishes
1146:
1142:
1138:
1134:
1127:
1123:
1119:
1115:
1111:
1098:
1091:
1086:
1083:
1079:
1072:
1067:
1063:
1059:
1055:
1048:
1043:
1039:
1036:
1032:
1025:
1020:
1019:
1018:
1015:
1010:
1008:
1004:
1000:
991:
982:
979:
970:
965:
962:
960:
959:Porcupinefish
956:
953:
949:
947:
943:
934:
933:Porcupinefish
928:
923:
919:
913:
908:
904:
898:
893:
889:
882:
877:
876:
875:
872:
868:
866:
865:lateral lines
862:
858:
854:
850:
846:
842:
828:
821:
816:
809:
804:
800:
794:
789:
785:
781:
774:
769:
768:
767:
764:
763:
758:
755:
751:
747:
743:
739:
737:
733:
732:invertebrates
729:
725:
721:
717:
714:
710:
707:are tireless
706:
702:
700:
696:
693:, as well as
692:
689:
685:
681:
677:
673:
669:
666:, feeding on
665:
662:has a varied
661:
655:
653:
649:
642:
638:
634:
625:
623:
619:
615:
611:
607:
603:
598:
585:
579:
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570:
563:
558:
554:
547:
542:
538:
531:
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525:
524:
522:
518:
514:
510:
505:
502:
498:
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492:
487:
483:
478:
476:
475:
469:
466:
462:
458:
453:
451:
445:
443:
442:countershaded
432:
426:
415:
411:
407:
400:
389:
388:
380:
366:
364:
360:
359:butterflyfish
355:
340:
339:countershaded
336:
332:
329:In contrast,
325:
316:
309:
286:
285:swim bladders
282:
278:
272:
267:
263:
259:
252:
247:
243:
239:
232:
227:
226:
225:
221:
217:
215:
207:
194:
187:
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172:
167:
166:
165:
162:
158:
153:
151:
147:
143:
138:
136:
132:
128:
124:
120:
113:
109:
105:
101:
97:
92:
83:
81:
77:
73:
68:
64:
62:
58:
54:
50:
46:
39:
34:
30:
19:
7518:Conservation
7506:
7473:
7466:
7249:conservation
7204:Algal blooms
7189:Water column
7104:Pelagic fish
7088:
7084:Coastal fish
7030:OSTM/Jason-2
7005:Bioeconomics
6940:Acoustic tag
6910:Fish counter
6785:
6773:
6673:spiny-finned
6622:Major groups
6376:
6343:Intelligence
6323:Scale eaters
6268:Cleaner fish
6150:Mouthbrooder
6102:Reproduction
6077:Schreckstoff
6062:Lateral line
5978:Swim bladder
5968:Spiral valve
5897:hyomandibula
5880:pseudobranch
5763:Hypoxia in -
5550:Conservation
5401:Pelagic fish
5385:
5381:Coastal fish
5285:Marine fungi
5023:Water garden
4906:Water column
4851:Productivity
4826:Pelagic zone
4786:Macrobenthos
4776:Hydrobiology
4746:Ecohydrology
4506:Symbiodinium
4504:
4497:
4408:Conservation
4323:Pulley Ridge
4182:
4172:
4090:Octocorallia
4009:Hexacorallia
3934:
3884:
3860:
3830:
3788:
3782:
3776:
3768:
3743:
3739:
3735:
3729:
3702:
3698:
3688:
3669:
3665:
3655:
3630:
3626:
3620:
3611:
3605:
3580:
3576:
3572:
3566:
3557:
3553:
3547:
3538:
3530:
3497:
3467:
3463:
3459:
3441:
3437:
3416:
3409:
3390:
3386:
3376:
3365:
3359:
3348:
3333:
3328:
3309:
3298:
3292:
3281:
3270:
3264:
3253:
3238:
3212:
3206:
3184:
3150:
3139:
3133:
3122:
3078:
3074:
3040:
3036:
2981:
2977:
2908:
2904:
2898:
2873:
2867:
2861:
2849:
2827:
2818:
2806:
2801:
2767:(12): e431.
2764:
2761:PLOS Biology
2760:
2750:
2731:
2725:
2705:
2698:
2648:
2644:
2638:
2614:. New York:
2610:
2603:
2592:
2586:
2575:
2566:
2561:
2550:
2544:
2533:
2489:
2483:
2469:
2442:
2438:
2428:
2388:
2377:
2372:Opsanus beta
2371:
2360:
2349:
2336:
2324:
2292:
2288:
2282:
2270:
2258:
2249:
2239:
2230:
2088:
2070:microhabitat
2067:
2059:
2044:
2029:tiger sharks
1972:
1952:tiger sharks
1924:parrotfishes
1920:damselfishes
1897:
1889:
1861:
1819:
1783:
1779:fish markets
1772:
1767:nurse sharks
1756:
1748:
1659:
1651:
1647:envenomation
1628:
1577:heteroxenous
1569:myxosporidia
1533:biodiversity
1526:
1504:
1492:
1404:Sea anemones
1402:
1361:sea cucumber
1354:
1319:
1315:parasitistic
1301:
1200:
1189:
1182:
1164:
1145:rabbitfishes
1141:damselfishes
1137:parrotfishes
1131:
1126:damselfishes
1062:moorish idol
1011:
999:cleaner fish
996:
957:
950:
946:sand dollars
939:
903:Cardinalfish
869:
853:cardinalfish
838:
827:giant morays
760:
745:
740:
735:
713:chemosensory
703:
656:
646:
614:Parrotfishes
599:
595:
506:
499:
479:
472:
470:
454:
446:
439:
385:
356:
352:
222:
218:
212:
154:
139:
117:
69:
65:
61:biodiversity
44:
43:
29:
7774:Marine fish
7769:Coral reefs
7442:Overfishing
7354:Catch share
7184:Marine snow
7169:Sardine run
7154:Fish screen
7149:Fish ladder
7099:Forage fish
6787:WikiProject
6746:Prehistoric
6730:Threatened
6421:Other types
6318:Sardine run
6293:Forage fish
6273:Corallivory
6125:Development
6110:Bubble nest
5983:physoclisti
5973:Suckermouth
5946:Root effect
5768:Ichthyology
5535:Sponge reef
5510:Rocky shore
5505:Oyster reef
5475:Kelp forest
5358:Vertebrates
5258:Marine life
5234:Viral shunt
5199:Marine snow
5101:Maharashtra
5008:Stream pool
4911:Zooplankton
4831:Photic zone
4791:Meiobenthos
4644:Algal bloom
4499:Amphidinium
4288:Lakshadweep
4223:Amazon Reef
4128:Coral reefs
4001:coral reefs
3946:Coral Reefs
3929:βΉ The
3705:: 239β251.
3672:(1): 3β34.
3418:PhysOrg.com
2618:. pp.
2162:Fish portal
2120:tiger shark
2104:bony fishes
2068:" ...shows
2013:crustaceans
2009:cephalopods
2005:bony fishes
1857:hammerheads
1786:giant moray
1722:giant moray
1704:secretes a
1580:life cycles
1553:monogeneans
1412:nematocysts
1346:crustaceans
1338:nematocysts
1307:mutualistic
1214:surgeonfish
1210:forage fish
1206:zooplankton
1157:macroscopic
1153:microscopic
1114:Surgeonfish
857:zooplankton
849:surgeonfish
845:forage fish
829:in hunting.
780:coral trout
754:protogynous
724:crustaceans
691:crustaceans
684:cephalopods
680:stomatopods
641:crustaceans
637:Red snapper
610:polychaetes
369:Colouration
363:angelfishes
283:with their
214:Coral reefs
193:yellow tang
157:yellow tang
150:triggerfish
123:coevolution
119:Coral reefs
100:Triggerfish
76:overfishing
53:coral reefs
38:coral reefs
7763:Categories
7666:Greenpeace
7374:Slot limit
7317:Management
7241:management
7209:Dead zones
7164:Salmon run
7144:Eel ladder
7040:Institutes
6880:Fish stock
6402:Groundfish
6397:Freshwater
6392:Euryhaline
6377:Coral reef
6313:Salmon run
6303:Paedophagy
6205:Amphibious
6192:Locomotion
6000:pharyngeal
5988:physostome
5941:Photophore
5887:Glossohyal
5860:gill raker
5843:dorsal fin
5793:physiology
5515:Salt marsh
5450:Coral reef
5239:Whale fall
5219:Photophore
5096:Everglades
5064:Ecoregions
5003:Stream bed
4976:Macrophyte
4929:Freshwater
4761:Food chain
4674:Water bird
4565:Fire coral
4550:Coral sand
4473:Reef Check
4453:Green Fins
4168:Microatoll
4108:Organ pipe
4032:Hermatypic
3809:References
3444:: 171β194.
2124:bull shark
2112:eagle rays
2055:open ocean
2017:sea snakes
1983:freshwater
1936:goatfishes
1893:home range
1833:ecosystems
1790:food chain
1724:can cause
1489:Monogenean
1334:hydrozoans
1332:, but are
1326:fire coral
1262:rabbitfish
1244:Ferocious
1230:parrotfish
1202:Damselfish
1196:rabbitfish
1184:Parrotfish
1178:herbivores
1133:Herbivores
1118:herbivores
1105:Herbivores
874:trevally.
847:, such as
784:damselfish
742:Moray eels
688:planktonic
652:herbivores
648:Carnivores
410:anglerfish
349:Body shape
333:like this
258:sea horses
178:damselfish
57:ecosystems
7636:WorldFish
7601:SeaChoice
7214:Fish kill
7194:Upwelling
7159:Migration
7066:fisheries
6860:Fisheries
6651:chimaeras
6538:Predatory
6515:Salmonids
6473:Whitefish
6463:Poisonous
6438:Diversity
6372:Coldwater
6308:Predatory
6298:Migratory
6258:Bait ball
6241:behaviour
6160:Pregnancy
6155:Polyandry
5929:papillare
5924:Operculum
5919:Meristics
5865:gill slit
5828:Cleithrum
5758:Fish kill
5748:Fear of -
5741:- as food
5731:Fisheries
5716:Evolution
5706:Diversity
5540:Tide pool
5445:Cold seep
5229:Upwelling
4993:Rheotaxis
4986:Fish pond
4961:Limnology
4886:Substrate
4871:Siltation
4741:Dead zone
4545:Coral rag
4468:Reef Ball
4238:Baa Atoll
3314:Ciguatera
3081:(1): 22.
3005:Free PDF
2942:Free PDF
2911:: 43β51.
2854:Pearlfish
2170:Anthiinae
2100:migration
2003:of small
2001:predators
1900:olfactory
1794:ciguatera
1775:stargazer
1763:ciguatera
1742:stargazer
1726:ciguatera
1706:ciguatera
1596:parasites
1549:digeneans
1537:nematodes
1529:parasites
1519:symbiotic
1511:clownfish
1507:clownfish
1408:tentacles
1357:pearlfish
1303:Symbiosis
1282:Symbiosis
1246:barracuda
1192:schooling
1033:feeds on
952:Barracuda
888:schooling
161:herbivore
72:pollution
18:Reef fish
7646:HERMIONE
7574:Advocacy
7486:Discards
7020:FishBase
7010:EconMult
6960:EcoSCOPE
6775:Category
6726:Smallest
6639:lampreys
6602:flatfish
6592:Demersal
6548:mackerel
6543:billfish
6483:Commerce
6412:Tropical
6387:Demersal
6382:Deep-sea
6338:Venomous
6230:RoboTuna
6180:triggers
6175:Spawning
6135:Juvenile
6120:Egg case
5753:FishBase
5647:Category
5573:HERMIONE
5490:Mangrove
5300:Seagrass
4846:Pleuston
4841:Plankton
4821:Particle
4766:Food web
4293:Maldives
4283:Kiribati
4233:Apo Reef
4163:Fringing
4118:Sea pens
4113:Sea fans
4071:Tabulate
4057:Staghorn
4027:Elegance
3931:template
3760:85922851
3647:19449694
3597:20738562
3560:: 13β30.
3367:FishBase
3317:Archived
3300:FishBase
3272:FishBase
3214:FishBase
3158:Archived
3141:FishBase
3107:22947621
3043:: 1β40.
3000:21344838
2945:Archived
2925:53051081
2793:17147471
2665:18103407
2594:FishBase
2552:FishBase
2491:FishBase
2474:FishBase
2461:86385679
2379:FishBase
2351:FishBase
2317:22684907
2309:15854891
2148:See also
2122:and the
2110:such as
2025:groupers
2021:seabirds
1989:and the
1987:whitetip
1979:brackish
1916:snappers
1904:acoustic
1825:blacktip
1821:Whitetip
1751:lionfish
1624:lionfish
1614:Toxicity
1592:molluscs
1565:copepods
1545:cestodes
1322:hawkfish
1296:Acropora
1292:hawkfish
728:molluscs
705:Goatfish
465:predator
412:, is an
281:decibels
277:Toadfish
131:venomous
86:Overview
7682:CalCOFI
7651:PROFISH
7581:FishAct
7481:Bycatch
7015:Ecopath
6930:Biomass
6862:science
6830:Fishery
6721:Largest
6634:hagfish
6629:Jawless
6607:pollock
6580:sardine
6575:herring
6570:anchovy
6520:Tilapia
6510:Octopus
6505:Catfish
6492:Farming
6407:Pelagic
6367:Coastal
6354:habitat
6210:Walking
6115:Clasper
6067:Otolith
6029:systems
6027:Sensory
5961:ganoine
5936:Papilla
5787:Anatomy
5726:Fishing
5500:Mudflat
5460:Estuary
5430:Bay mud
5408:Seabird
5164:f-ratio
5147:General
5028:Wetland
4816:Neuston
4781:Hypoxia
4726:Biomass
4716:Benthos
4632:General
4333:Red Sea
4278:Jamaica
4052:Elkhorn
4037:Chalice
3955:WhyReef
3933:below (
3881:Pauly D
3707:Bibcode
3627:Ecology
3098:3507714
3037:Zootaxa
2890:2251533
2784:1750927
2051:lagoons
1940:octopus
1869:lagoons
1561:isopods
1557:leeches
1515:ammonia
1499:grouper
1174:benthic
1078:wrasses
1038:sponges
841:schools
750:Grouper
716:barbels
709:benthic
672:shrimps
622:snapper
513:ventral
495:puffers
461:eyespot
7621:Oceana
7384:EU MLS
6739:sharks
6656:sharks
6585:sprats
6565:Forage
6553:salmon
6433:Coarse
6215:Flying
6087:Vision
6042:Barbel
5956:Scales
5816:dermal
5696:About
5485:Lagoon
4811:Nekton
4669:Mammal
4664:Insect
4358:Yabiji
4098:Bamboo
4065:Rugose
4042:Pillar
3997:Corals
3950:Curlie
3936:Curlie
3895:
3889:ICLARM
3868:
3851:
3837:
3822:
3758:
3645:
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2011:, and
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1934:, and
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1631:toxins
1588:sharks
1551:, and
1416:toxins
1330:corals
1260:Coral
1147:, and
1122:shoals
839:Large
778:Adult
762:harems
668:fishes
606:polyps
584:Gobies
517:dorsal
501:Gobies
236:Among
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6689:Lists
6468:Rough
6333:Sleep
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4876:Spawn
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4146:Coral
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4047:Table
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4017:Black
3756:S2CID
3033:(PDF)
2921:S2CID
2886:S2CID
2661:S2CID
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2187:Notes
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1873:gills
1643:venom
1600:hosts
1584:hosts
1522:algae
1166:Algae
1160:algae
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695:plant
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6661:rays
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