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Pterygota

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Unlike Archaeognatha and Zygentoma, the pterygotes do not have styli or vesicles on their abdomen (also absent in some zygentomans), and with the exception of the majority of mayflies, are also missing the median terminal filament which is present in the ancestrally wingless insects.
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The oldest known representatives of the group appeared during the mid-Carboniferous, around 328–324 million years ago, and the group subsequently underwent a rapid explosive diversification. Claims that they originated substantially earlier during the
632:. The reason given is: Highly outdated; for example, "Exopterygota" is almost certainly considered paraphyletic by most phylogenetic studies in the last 10 years at least by now, as most of them instead recognise at least 552:
that includes all winged insects and the orders that are secondarily wingless (that is, insect groups whose ancestors once had wings but that have lost them as a result of subsequent evolution).
682:, and better treatments (such as dividing or dissolving the group) are presently being discussed. In addition, it is not clear how exactly the neopterans are related among each other. The 1332: 2316: 2258: 2238: 2331: 772: 2627: 1007: 2346: 1358:
Schachat, Sandra R; Goldstein, Paul Z; Desalle, Rob; Bobo, Dean M; Boyce, C Kevin; Payne, Jonathan L; Labandeira, Conrad C (2 February 2023).
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The following scheme uses finer divisions than the one above, which is not well-suited to correctly accommodating the fossil groups.
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Please help update this article to reflect recent events or newly available information.
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estimates are unlikely based on the fossil record, and are likely analytical artefacts.
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Proceedings of the Royal Entomological Society of London. Series A, General Entomology
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The pterygotan group comprises 99.9% of all insects. The orders not included are the
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seem to be very close relatives, indeed, but nonetheless appear to contain several
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insects among the Neoptera like the Palaeoptera are among insects as a whole. The
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Grundriss der vergleichenden Anatomie. Zweite verbesserte Auflage
1033: 778: 584: 576: 421: 240: 98: 73: 2501: 1413: 571:), two primitively wingless insect orders. Also not included is 1987: 1887: 1814: 1720: 1629: 1621: 1459: 1444: 971: 698: 549: 371: 185: 165: 2380: 1261:"Coxal Setal Organs in Archaeognatha and Zygentoma (Insecta)" 1357: 1217: 2075: 1077: 1027: 929: 701:
of related orders, the status of which is not agreed upon.
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Mechanoreceptors: Development, Structure, and Function
1147:; Yavorskaya, Margarita & Beutel, Rolf G. (2016). 678:. The former are nowadays strongly suspected of being 509: 473: 1246:"The Evolution and Genetics of Migration in Insects" 497: 482: 476: 226: 488: 1331:Gullan, P. J.; Cranston, P. S. (3 November 2014). 2604: 686:might be a similar assemblage of rather ancient 1218:Vincent H. Resh; Ring T. Cardé (4 April 2003). 667:Traditionally, this group was divided into the 1330: 1865: 1429: 1211: 1697: 1505: 1611: 1436: 1422: 109: 1375: 1364:Biological Journal of the Linnean Society 1172: 1122: 862:(stick insects – tentatively placed here) 775:†(extinct; sometimes included in Odonata) 765:†(extinct; sometimes included in Odonata) 759:†(extinct; sometimes included in Odonata) 753:†(extinct; sometimes included in Odonata) 707: 2157:Four most speciose orders are marked in 1747:(crickets, wetas, grasshoppers, locusts) 1398: 1295: 868:(ice-crawlers – tentatively placed here) 2628:Extant Pennsylvanian first appearances 2605: 1153:Journal of the Royal Society Interface 1138: 1136: 874:(gladiators – tentatively placed here) 793: 2409: 2408: 2195: 1490: 1417: 1334:The Insects: An Outline of Entomology 1258: 2020:(alderflies, dobsonflies, fishflies) 1296:Drummond, F. H. (25 December 1953). 614: 262:(barklice, booklice, parasitic lice) 1133: 13: 1399:Maddison, David (1 January 2002). 1318:10.1111/j.1365-3032.1953.tb00643.x 14: 2644: 1443: 1267:. Springer US. pp. 365–367. 1259:Weyda, František (25 June 1988). 2379: 1530: 619: 469: 142: 41: 1392: 559:(jumping bristletails) and the 1351: 1324: 1289: 1252: 1238: 1224:. Academic Press. p. 64. 1189: 1116: 610: 1: 1950:(sawflies, wasps, ants, bees) 1110: 1898:(cicadas, aphids, true bugs) 1273:10.1007/978-1-4899-0812-4_70 533: 7: 2196: 1298:"THE EVERSIBLE VESICLES OF 1199:. Oxford English Dictionary 10: 2649: 2088:(gnats, mosquitoes, flies) 1981:(twisted-winged parasites) 1653:(dragonflies, damselflies) 1491: 1401:"Pterygota Winged Insects" 1377:10.1093/biolinnean/blac137 1285:– via Springer Link. 980:(twisted-winged parasites) 888:(cockroaches and termites) 522: 380:(twisted-winged parasites) 289:(cockroaches and termites) 23:Pterygota (disambiguation) 20: 2417: 2373: 2206: 2202: 2191: 2153: 2098: 2056: 2041: 2000: 1969: 1960: 1942: 1911: 1861: 1834: 1797: 1764: 1708: 1693: 1670: 1641: 1620: 1607: 1582: 1566: 1539: 1528: 1501: 1497: 1486: 1454: 1347:– via Google Books. 1337:. John Wiley & Sons. 628:This section needs to be 223: 218: 139:Scientific classification 137: 117: 108: 32: 1739:(stick and leaf insects) 712:(probably paraphyletic) 649:Last update: August 2007 544:'winged') are a 2166:are paraphyletic groups 1809:(cockroaches, termites) 1573:(silverfish, firebrats) 1221:Encyclopedia of Insects 1517:(jumping bristletails) 1165:10.1098/rsif.2016.0363 708:Infraclass Palaeoptera 2562:Paleobiology Database 1320:– via CrossRef. 1123:Gegenbaur, C (1878). 1020:(scorpionflies, etc.) 408:(scorpionflies, etc.) 230:Palaeodictyopteroidea 2618:Arthropod subclasses 2119:(moths, butterflies) 1052:Proposed superorder 1006:Proposed superorder 1002:(net-veined insects) 916:(booklice, barklice) 906:Proposed superorder 878:Proposed superorder 402:(net-veined insects) 21:For other uses, see 794:Infraclass Neoptera 644:than Polyneoptera.. 131:Tribulus terrestris 39:Serpukhovian–Recent 16:Subclass of insects 2386:Insects portal 2357:Triadophlebioptera 1312:(10–12): 145–148. 763:Triadophlebioptera 2600: 2599: 2411:Taxon identifiers 2402: 2401: 2369: 2368: 2365: 2364: 2307:Palaeodictyoptera 2292: 2224: 2187: 2186: 2183: 2182: 2149: 2148: 2145: 2144: 2141: 2140: 2137: 2136: 2133: 2132: 2129: 2128: 2120: 2111: 2089: 2079: 2071: 2049: 2037: 2036: 2029: 2021: 2013: 1991: 1982: 1951: 1907: 1906: 1899: 1891: 1883: 1830: 1829: 1826: 1825: 1818: 1810: 1789: 1786:Mantophasmatodea 1781: 1770: 1756: 1748: 1740: 1732: 1724: 1716: 1666: 1665: 1662: 1661: 1654: 1633: 1622:Ephemeropteroidea 1574: 1526: 1525: 1518: 1344:978-1-118-84615-5 1282:978-1-4899-0814-8 1231:978-0-08-054605-6 723:Palaeodictyoptera 665: 664: 650: 543: 531: 462: 461: 214: 2640: 2593: 2592: 2580: 2579: 2570: 2569: 2557: 2556: 2544: 2543: 2531: 2530: 2518: 2517: 2505: 2504: 2492: 2491: 2479: 2478: 2466: 2465: 2453: 2452: 2451: 2438: 2437: 2436: 2406: 2405: 2384: 2383: 2317:Permoplectoptera 2290: 2249:Diaphanopterodea 2222: 2204: 2203: 2193: 2192: 2118: 2109: 2100:Amphiesmenoptera 2087: 2081: 2077: 2069: 2059: 2054: 2053: 2048: 2045: 2027: 2019: 2011: 1989: 1980: 1967: 1966: 1963: 1949: 1940: 1939: 1897: 1889: 1882:(barklice, lice) 1881: 1873: 1863: 1862: 1859: 1858: 1816: 1808: 1787: 1779: 1778:Grylloblattodea 1768: 1754: 1746: 1738: 1730: 1722: 1714: 1706: 1705: 1702: 1695: 1694: 1691: 1690: 1652: 1631: 1618: 1617: 1609: 1608: 1605: 1604: 1572: 1564: 1563: 1534: 1516: 1503: 1502: 1499: 1498: 1488: 1487: 1438: 1431: 1424: 1415: 1414: 1409: 1408: 1396: 1390: 1389: 1379: 1355: 1349: 1348: 1328: 1322: 1321: 1293: 1287: 1286: 1256: 1250: 1249: 1242: 1236: 1235: 1215: 1209: 1208: 1206: 1204: 1193: 1187: 1186: 1176: 1140: 1131: 1130: 1120: 1087:Neoptera orders 1054:Amphiesmenoptera 872:Mantophasmatodea 741:Diaphanopterodea 660: 657: 651: 648: 623: 622: 615: 538: 536: 526: 524: 513: 507: 506: 503: 502: 499: 496: 493: 490: 487: 484: 481: 478: 475: 345:Mantophasmatodea 323:(walking sticks) 228: 209: 147: 146: 118:Giant honey bee 113: 103: 40: 36:Temporal range: 30: 29: 2648: 2647: 2643: 2642: 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1143:Kjer, Karl M.; 1141: 1134: 1121: 1117: 1113: 866:Grylloblattodea 840:(angel insects) 796: 710: 661: 655: 652: 647: 645: 624: 620: 613: 605:molecular clock 511: 472: 468: 339:Grylloblattodea 335:(angel insects) 208: 141: 104: 102: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 38: 37: 34: 26: 17: 12: 11: 5: 2646: 2636: 2635: 2630: 2625: 2620: 2615: 2598: 2597: 2595: 2594: 2581: 2571: 2558: 2545: 2532: 2519: 2506: 2493: 2480: 2467: 2454: 2439: 2423: 2421: 2415: 2414: 2400: 2399: 2397: 2396: 2389: 2374: 2371: 2370: 2367: 2366: 2363: 2362: 2360: 2359: 2354: 2349: 2344: 2342:Protozygoptera 2339: 2337:Protorthoptera 2334: 2332:Protephemerida 2329: 2324: 2322:Protanisoptera 2319: 2314: 2309: 2304: 2299: 2294: 2291:(griffinflies) 2289:Meganisoptera 2286: 2281: 2276: 2271: 2269:Glosselytrodea 2266: 2261: 2256: 2251: 2246: 2241: 2236: 2231: 2226: 2218: 2213: 2207: 2200: 2199: 2189: 2188: 2185: 2184: 2181: 2180: 2178: 2177: 2167: 2161: 2154: 2151: 2150: 2147: 2146: 2143: 2142: 2139: 2138: 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31: 28: 24: 19: 2418: 2391: 2377: 2347:Syntonoptera 2229:Caloneurodea 2221:Blattoptera 2172: 2163: 2158: 2117:Lepidoptera 2115: 2108:Trichoptera 2084: 2066: 2018:Megaloptera 2012:(snakeflies) 2002:Neuropterida 1986: 1971:Coleopterida 1948:Hymenoptera 1946: 1868:Paraneoptera 1866: 1788:(gladiators) 1737:Phasmatodea 1715:(stoneflies) 1700:Polyneoptera 1643:Odonatoptera 1583: 1507:Monocondylia 1472:Pancrustacea 1470:(unranked): 1405:Tree of Life 1404: 1394: 1367: 1363: 1353: 1333: 1326: 1309: 1305: 1302:(THYSANURA)" 1299: 1291: 1264: 1254: 1240: 1220: 1213: 1203:17 September 1201:. Retrieved 1191: 1159:(121): 121. 1156: 1152: 1145:Simon, Chris 1125: 1118: 1088: 1086: 1085: 1051: 1044:Protodiptera 1024:Siphonaptera 1005: 986:(snakeflies) 978:Strepsiptera 948: 947: 926:Phthiraptera 920:Thysanoptera 908:Paraneoptera 905: 877: 854:grasshoppers 828:(stoneflies) 808:Caloneurodea 801:Exopterygota 798: 797: 711: 703: 684:Exopterygota 680:paraphyletic 669:infraclasses 666: 653: 642:Holometabola 638:Paraneoptera 634:Polyneoptera 629: 593: 589: 565:silverfishes 554: 465: 463: 412:Siphonaptera 386:(snakeflies) 378:Strepsiptera 353:Holometabola 347:(gladiators) 315:grasshoppers 307:(stoneflies) 282:Polyneoptera 266:Thysanoptera 205: 192:(unranked): 129: 121:Apis dorsata 119: 27: 18: 2523:iNaturalist 2443:Wikispecies 2393:Wikispecies 2352:Titanoptera 2284:Lapeyriidae 2279:Hypoperlida 2254:Eoblattodea 2223:(roachoids) 2216:Archodonata 2058:Antliophora 2026:Neuroptera 1988:Coleoptera 1799:Dictyoptera 1769:(Xenonomia) 1745:Orthoptera 1729:Embioptera 1721:Dermaptera 1713:Plecoptera 1613:Palaeoptera 1476:Subphylum: 1197:"Pterygota" 1074:butterflies 1070:Lepidoptera 1064:caddisflies 1060:Trichoptera 1012:Antliophora 990:Megaloptera 958:Hymenoptera 949:Superorder 898:Alienoptera 880:Dictyoptera 860:Phasmatodea 814:Titanoptera 799:Superorder 787:damselflies 783:dragonflies 747:Protodonata 735:Archodonata 611:Systematics 446:butterflies 442:Lepidoptera 436:caddisflies 432:Trichoptera 390:Megaloptera 358:Hymenoptera 321:Phasmatodea 249:damselflies 245:dragonflies 126:Hymenoptera 2623:Dicondylia 2607:Categories 2476:Pterygotes 2302:Miomoptera 2068:Mecoptera 1896:Hemiptera 1817:(mantises) 1807:Blattodea 1753:Zoraptera 1632:(mayflies) 1571:Zygentoma 1466:Arthropoda 1111:References 1106:†(extinct) 1104:Miomoptera 1100:†(extinct) 1046:†(extinct) 1000:Neuroptera 994:alderflies 972:Coleoptera 914:Psocoptera 900:†(extinct) 850:Orthoptera 844:Dermaptera 832:Embioptera 826:Plecoptera 822:†(extinct) 816:†(extinct) 810:†(extinct) 743:†(extinct) 737:†(extinct) 731:†(extinct) 725:†(extinct) 719:(mayflies) 672:Paleoptera 581:Collembola 573:Entognatha 534:pterugōtós 523:πτερυγωτός 400:Neuroptera 394:alderflies 372:Coleoptera 327:Embioptera 311:Orthoptera 305:Plecoptera 299:Dermaptera 237:(mayflies) 202:Subclass: 196:Dicondylia 176:Arthropoda 2613:Pterygota 2463:Pterygota 2449:Pterygota 2419:Pterygota 2274:Heraridea 2264:Geroptera 2169:Based on 2043:Panorpida 1990:(beetles) 1880:Psocodea 1815:Mantodea 1766:Notoptera 1723:(earwigs) 1458:Kingdom: 1386:0024-4066 1018:Mecoptera 974:(beetles) 936:Hemiptera 894:(mantids) 886:Blattodea 846:(earwigs) 838:Zoraptera 603:based on 569:firebrats 561:Zygentoma 528:romanized 466:Pterygota 406:Mecoptera 374:(beetles) 333:Zoraptera 301:(earwigs) 295:(mantids) 287:Blattodea 272:Hemiptera 211:Gegenbaur 206:Pterygota 162:Kingdom: 156:Eukaryota 33:Pterygota 2428:Wikidata 2312:Paoliida 2086:Diptera 1890:(thrips) 1651:Odonata 1478:Hexapoda 1464:Phylum: 1460:Animalia 1300:CAMPODEA 1183:27558853 922:(thrips) 892:Mantodea 676:Neoptera 601:Devonian 597:Silurian 546:subclass 293:Mantodea 268:(thrips) 260:Psocodea 255:Neoptera 172:Phylum: 166:Animalia 152:Domain: 2577:5791287 2515:9502664 2502:2634372 2197:Extinct 2171:Sasaki 2078:(fleas) 1872: * 1174:5014063 1034:Diptera 996:, etc.) 968:, etc.) 856:, etc.) 779:Odonata 630:updated 585:Diplura 577:Protura 550:insects 542:  530::  422:Diptera 396:, etc.) 368:, etc.) 241:Odonata 219:Orders 186:Insecta 182:Class: 124:(order 2590:151152 2574:uBio: 2541:100500 2528:184884 2489:518860 2434:Q22708 2175:(2013) 2173:et al. 2164:Italic 1492:Extant 1448:orders 1445:Insect 1384:  1341:  1279:  1228:  1181:  1171:  1036:(true 938:(true 699:clades 583:, and 510:terrə- 424:(true 317:, etc) 274:(true 213:, 1878 2585:WoRMS 2567:94228 1078:moths 1038:flies 1028:fleas 450:moths 426:flies 416:fleas 128:) on 2554:7496 2549:NCBI 2536:ITIS 2510:GBIF 2484:BOLD 2159:bold 1382:ISSN 1339:ISBN 1277:ISBN 1226:ISBN 1205:2024 1179:PMID 966:bees 962:ants 940:bugs 930:lice 785:and 674:and 567:and 540:lit. 464:The 366:bees 362:ants 276:bugs 247:and 44:PreꞒ 2497:EoL 2471:AFD 2458:ADW 1372:doi 1368:138 1314:doi 1269:doi 1169:PMC 1161:doi 771:or 749:or 599:or 548:of 514:-tə 512:GOH 2609:: 2587:: 2564:: 2551:: 2538:: 2525:: 2512:: 2499:: 2486:: 2473:: 2460:: 2445:: 2430:: 2074:+ 1403:. 1380:. 1366:. 1362:. 1310:28 1308:. 1304:. 1275:. 1177:. 1167:. 1157:13 1155:. 1151:. 1135:^ 1076:, 964:, 587:. 579:, 537:, 525:, 521:: 495:oʊ 448:, 364:, 94:Pg 1935:a 1933:l 1931:o 1929:b 1927:a 1925:t 1923:e 1921:m 1919:o 1917:l 1915:o 1913:H 1854:a 1852:l 1850:o 1848:b 1846:a 1844:t 1842:e 1840:m 1838:u 1836:E 1686:a 1684:r 1682:e 1680:t 1678:p 1676:o 1674:e 1672:N 1600:a 1598:t 1596:o 1594:g 1592:y 1590:r 1588:e 1586:t 1584:P 1559:a 1557:i 1555:l 1553:y 1551:d 1549:n 1547:o 1545:c 1543:i 1541:D 1437:e 1430:t 1423:v 1407:. 1388:. 1374:: 1316:: 1271:: 1248:. 1234:. 1207:. 1185:. 1163:: 1080:) 1072:( 1066:) 1062:( 1040:) 1030:) 1026:( 1010:/ 992:( 960:( 942:) 932:) 928:( 852:( 789:) 781:( 658:) 654:( 563:( 517:( 504:/ 501:ə 498:t 492:ɡ 489:ˈ 486:ə 483:r 480:ɛ 477:t 474:ˌ 471:/ 452:) 444:( 438:) 434:( 428:) 418:) 414:( 392:( 360:( 313:( 278:) 251:) 243:( 227:† 99:N 89:K 84:J 79:T 74:P 69:C 64:D 59:S 54:O 49:Ꞓ 25:.

Index

Pterygota (disambiguation)
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Apis dorsata
Hymenoptera
Tribulus terrestris
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Dicondylia
Pterygota
Gegenbaur
Palaeodictyopteroidea
Ephemeroptera
Odonata
dragonflies

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