590:
2515:
117:
4355:
1661:
150:
1655:
1649:
4361:
4349:
2510:
564:
The tibia and fibula were not fused, and the forearm and lower leg bones were respectively shorter than the humerus and femur, but the hind legs were much longer than the forelegs. The second and third fingers of the hand were of similar length, and the fourth was smaller ; the fifth finger was
485:
The skull had a complete dentition, but the second and third incisors, as well as the canines and the first lower premolars, were vestigial. The first upper incisors were very large, and showed a considerable development.
889:"A poorly known rodentlike mammal (Pachyrukhinae, Hegetotheriidae, Notoungulata) from the Deseadan (Late Oligocene) of Argentina. Paleoecology, biogeography, and radiation of the rodentlike ungulates in South America"
868:
F. Ameghino. 1897. Mammiféres crétacés de l’Argentine (Deuxième contribution à la connaissance de la fauna mammalogique de couches à Pyrotherium) . Boletin
Instituto Geografico Argentino 18(4–9):406-521
807:
F. D. Seoane, S. R. Juñent, and E. Cerdeño. 2017. Phylogeny and paleobiogeography of
Hegetotheriidae (Mammalia, Notoungulata). Journal of Vertebrate Paleontology 37(1):e1278547:1-13
877:
F. Ameghino. 1901. Notices préliminaires sur des ongulés nouveaux des terrains crétacés de
Patagonie . Boletin de la Academia Nacional de Ciencias de Córdoba 16:349-429
3626:
3710:
4233:
4020:
2380:
4149:
4128:
4055:
3937:
3468:
3286:
3196:
2396:
1197:
3654:
3633:
3391:
3335:
3153:
2963:
2786:
1162:
4076:
3668:
3503:
2865:
2447:
1900:
1352:
542:
was likewise characteristic, with the trochlear nerve extant covering entirely the dorsal part of the bone, and a low and flat condylar crest. The
4034:
3823:
3788:
3489:
3482:
3475:
3461:
3419:
3356:
3279:
3132:
3062:
2823:
2816:
2705:
2177:
2060:
1807:
1736:
4142:
3930:
3872:
3765:
3696:
3661:
3405:
3139:
2691:
2684:
2603:
2417:
2163:
2074:
2009:
1995:
1693:
1043:
3816:
3517:
3377:
3370:
3342:
3307:
3293:
3146:
3118:
3040:
2844:
2830:
2809:
2726:
2403:
2142:
2023:
1213:
1082:
1067:
561:
Its body was slender, with a very short tail and strong forelegs, although much shorter than the hind legs, equipped with long metatarsals.
3398:
3272:
3182:
3104:
2368:
1800:
1482:
1155:
1133:
1119:
1089:
4445:
3363:
3314:
2610:
2433:
2251:
1075:
1053:
4470:
3689:
3160:
2991:
2487:
3758:
960:
4465:
768:
among the
Oligocene hegetotheres, but also reflects the remarkable evolutionary radiation of rodent-like ungulates during the
4475:
888:
2514:
441:
744:
The Late
Oligocene terrains of Patagonia present at least three genera of hegetotheres with hypsodont teeth :
970:
4354:
4480:
4460:
4485:
4455:
953:
1660:
389:
149:
824:"Re-assessment of the Oligocene genera Prosotherium and Propachyrucos (Hegetotheriidae, Notoungulata)"
823:
466:
were high and deep ; the mandible had a subtle coronoid process. The teeth were characterized by
1654:
1577:
946:
4417:
4380:
4348:
1648:
482:, generally on the outside part of the upper teeth and on the inside part of the lower teeth.
4412:
4360:
558:. The distal phalanx was high and narrow, flattening and widening towards the terminal part.
522:
was equipped of a small head separated from the main bone. The femur was even longer than in
2509:
938:
2051:
900:
835:
700:
This was supported by a 2019 reassessment as well as a number of other studies. This means
589:
8:
4440:
3610:
2698:
2156:
614:
381:
904:
839:
776:, and suggest environmental differences between the fauna of Patagonia and the fauna of
4121:
1600:
924:
296:
144:
526:, and had a small trochanter curiously pushed back in the posterior part. Contrary to
4450:
4219:
4069:
4062:
3580:
2765:
2756:
2624:
2002:
1953:
1921:
1621:
1563:
1524:
1510:
1425:
916:
851:
451:
928:
116:
4226:
4205:
4105:
3893:
3809:
3675:
3175:
2654:
2286:
2272:
2207:
2107:
1935:
1886:
1591:
1554:
1462:
1441:
1411:
1306:
1206:
908:
843:
765:
728:
for cranial and post-cranial characters, which are known only from the synonymized
393:
1014:
4403:
4189:
4177:
4013:
3779:
3724:
3647:
3589:
3249:
2779:
2314:
2237:
2081:
1914:
1856:
1828:
1791:
1764:
650:
598:
385:
373:
217:
132:
4083:
3921:
3907:
3900:
3886:
3596:
3349:
3300:
3189:
3125:
3090:
3054:
2984:
2872:
2772:
2675:
2647:
2633:
2569:
2545:
2389:
2321:
2307:
2293:
2149:
2121:
2088:
2016:
1979:
1966:
1614:
1607:
1570:
1533:
1489:
1448:
1373:
1359:
1315:
1148:
1126:
1103:
1000:
847:
30:
4434:
4270:
4254:
4212:
4198:
4027:
4004:
3969:
3951:
3851:
3844:
3703:
3566:
3447:
3440:
3321:
3083:
3013:
2956:
2886:
2590:
2300:
2279:
2223:
1907:
1398:
1336:
1220:
1023:
920:
855:
822:
Seoane, Federico Damián; Cerdeño, Esperanza; Singleton, Hayley (2019-09-01).
773:
761:
750:
684:
666:
503:
471:
406:
230:
74:
4314:
4298:
4240:
4114:
4041:
3990:
3944:
3865:
3837:
3802:
3737:
3433:
3263:
3235:
3226:
3214:
3069:
3047:
3027:
3006:
2970:
2949:
2928:
2921:
2912:
2900:
2858:
2640:
2532:
2342:
2230:
2030:
1928:
1893:
1879:
1872:
1835:
1821:
1814:
1540:
1345:
675:
658:
577:
418:
256:
204:
4321:
4307:
4284:
4048:
3879:
3858:
3795:
3426:
3412:
3384:
3168:
2851:
2800:
2712:
2661:
2576:
2562:
2456:
2426:
2335:
2328:
2265:
2216:
2193:
2135:
1988:
1777:
1750:
1728:
1721:
1705:
1475:
1380:
1246:
1234:
1015:
567:
547:
446:
412:
49:
4263:
4156:
3559:
3550:
3111:
3097:
3076:
2999:
2744:
2494:
2258:
2244:
2128:
2067:
1678:
1294:
1269:
1019:
756:
551:
550:
slightly concave and occupying the entire distal part of the bone. The
539:
507:
424:
94:
59:
4163:
3982:
3830:
3640:
3619:
3454:
3328:
2977:
2879:
2837:
2555:
1842:
1743:
1714:
1496:
1366:
1169:
1112:
1060:
912:
618:
543:
511:
377:
370:
161:
136:
99:
43:
4374:
887:
Reguero, Marcelo A.; Dozo, María Teresa; Cerdeño, Esperanza (2007).
404:
Compared to its relatives, it would have been about 20% larger than
4397:
3751:
3744:
3682:
3510:
2942:
2935:
2719:
2463:
2186:
1865:
1503:
1434:
1322:
1276:
1255:
1183:
1176:
1141:
988:
769:
669:
subfamily comprising several hare-like genera. It is possible that
479:
475:
463:
181:
89:
84:
69:
64:
54:
34:
4135:
3524:
3496:
3242:
781:
777:
724:
Despite this, until 2019, phylogenetic analyses regularly scored
495:
467:
459:
440:
Its skull was light and thin, notably in the posterior part. The
432:, in aspect and in size. Its hind legs were particularly long.
363:
104:
79:
2037:
1262:
994:
982:
704:
is known from an almost complete specimen, ie, the holotype of
654:
535:
515:
429:
191:
171:
968:
531:
519:
366:
639:, a fourth species, was described in 1901 by Ameghino. Only
631:, which was a bit larger than the type species, as well as
610:
499:
455:
581:. The metatarsals were elongated, especially the third.
546:
was relatively small, with an articulated facet for the
444:
was particularly developed, even more than its relative
821:
4295:
4251:
4186:
4174:
4102:
4001:
3966:
3918:
3776:
3721:
3607:
3577:
3547:
3260:
3223:
3211:
3024:
2909:
2897:
2797:
2753:
2741:
2672:
2621:
2587:
2444:
2414:
2377:
2365:
2204:
2174:
2104:
2048:
1976:
1963:
1950:
1853:
1788:
1761:
1702:
1690:
1588:
1551:
1521:
1459:
1422:
1395:
1333:
1303:
1291:
1243:
1231:
1194:
1100:
1040:
518:
was elongated, slender and lightly build, while the
514:
indicates that the hand was small and delicate. The
886:
760:, the latter of which is sometimes considered an
4432:
621:in Late Oligocene terrains. The type species is
880:
682:In a 1999 unpublished Ph.D. dissertation, the
954:
673:was an ancestral form or a close relative of
627:, and Ameghino described in this same paper
1242:
961:
947:
571:. The hand seems more specialized than in
454:was quite large, similar to the ears of a
115:
565:small, even smaller than in its relative
708:The 2019 study recovers the features of
617:, over fossilized remains discovered in
593:Closeup of AMNH 29574, the holotype of "
588:
764:. This implies a strict repartition of
575:, and shares similarities with that of
489:
376:notoungulate. It has been found in two
4433:
803:
801:
799:
797:
4379:
4378:
2530:
1676:
1013:
942:
712:as consistent with those of juvenile
817:
815:
813:
794:
784:, where those animals were absent.
13:
4446:Oligocene mammals of South America
478:covered by a thin layer of dental
442:tympanic part of the temporal bone
14:
4497:
4471:Taxa named by Florentino Ameghino
810:
584:
4359:
4353:
4347:
2531:
2513:
2508:
1659:
1653:
1647:
148:
47:
971:South American native ungulates
739:
554:was robust and shorter than in
538:were completely separated. The
1677:
871:
862:
450:, and it is probable that its
399:
16:Extinct genus of notoungulates
1:
4466:Fossil taxa described in 1897
787:
498:was large and thin, like the
428:. This animal was similar to
4476:Prehistoric placental genera
422:, but slightly smaller than
7:
643:is still considered valid.
390:Agua de la Piedra Formation
380:fossil formations, ie, the
10:
4502:
848:10.1016/j.crpv.2019.07.001
716:(although the holotype of
312:Prosotherium triangulidens
4387:
4345:
4294:
4250:
4185:
4173:
4101:
4000:
3977:
3965:
3917:
3775:
3732:
3720:
3606:
3576:
3546:
3259:
3222:
3210:
3035:
3023:
2908:
2896:
2796:
2752:
2740:
2671:
2620:
2598:
2586:
2543:
2539:
2526:
2506:
2477:
2443:
2413:
2376:
2364:
2203:
2173:
2116:
2102:
2047:
1975:
1961:
1949:
1852:
1787:
1772:
1760:
1701:
1689:
1685:
1672:
1645:
1587:
1550:
1520:
1470:
1458:
1421:
1406:
1394:
1332:
1302:
1290:
1230:
1193:
1099:
1039:
1035:
1031:
1009:
977:
720:was not directly examined
595:Propachyrucos ameghinorum
435:
336:Propachyrucos ameghinorum
302:
295:
281:
276:
262:
255:
145:Scientific classification
143:
129:Propachyrucos ameghinorum
123:
114:
23:
1026:, and other minor groups
893:Journal of Paleontology
609:was first described in
828:Comptes Rendus Palevol
696:were synonymized with
602:
344:Propachyrucos simpsoni
4481:Golfo San Jorge Basin
4413:Paleobiology Database
698:Prosotherium garzoni.
592:
320:Prosotherium robustum
4461:Fossils of Argentina
2052:Sparnotheriodontinae
714:Prosotherium garzoni
702:Prosotherium garzoni
679:, more specialized.
665:was a member of the
653:, a clade of light,
649:was a member of the
641:Prosotherium garzoni
624:Prosotherium garzoni
490:Postcranial skeleton
328:Prosotherium quartum
266:Prosotherium garzoni
125:Prosotherium garzoni
4486:Sarmiento Formation
4456:Paleogene Argentina
4234:Pseudohegetotherium
3611:Oldfieldthomasiidae
2699:Edvardotrouessartia
2157:Pseudobrachytherium
905:2007JPal...81.1301R
840:2019CRPal..18..643S
657:-like or hare-like
615:Florentino Ameghino
382:Sarmiento Formation
4122:Archaeotypotherium
4021:Caraguatypotherium
1601:Granastrapotherium
603:
131:). Collected from
4428:
4427:
4381:Taxon identifiers
4372:
4371:
4368:
4367:
4343:
4342:
4339:
4338:
4335:
4334:
4331:
4330:
4220:Hemihegetotherium
4097:
4096:
4093:
4092:
4070:Pseudotypotherium
4063:Plesiotypotherium
3961:
3960:
3581:Archaeopithecidae
3542:
3541:
3538:
3537:
3534:
3533:
3206:
3205:
2757:Homalodotheriidae
2736:
2735:
2625:Henricosborniidae
2522:
2521:
2504:
2503:
2473:
2472:
2381:Amilnedwardsiidae
2360:
2359:
2356:
2355:
2352:
2351:
2098:
2097:
2003:Guilielmofloweria
1954:Proterotherioidea
1945:
1944:
1922:Scalabrinitherium
1668:
1667:
1643:
1642:
1639:
1638:
1635:
1634:
1631:
1630:
1622:Xenastrapotherium
1564:Astrapothericulus
1525:Albertogaudryinae
1511:Parastrapotherium
1426:Trigonostylopidae
1390:
1389:
1286:
1285:
766:ecological niches
470:pointing inward,
355:
354:
251:
4493:
4421:
4420:
4408:
4407:
4406:
4376:
4375:
4363:
4357:
4351:
4297:
4292:
4291:
4253:
4227:Prohegetotherium
4206:Hegetotheriopsis
4188:
4183:
4182:
4176:
4150:Protarchaeohyrax
4129:Bryanpattersonia
4106:Archaeohyracidae
4104:
4099:
4098:
4056:Microtypotherium
4003:
3975:
3974:
3968:
3938:Guilielmoscottia
3920:
3894:Progaleopithecus
3810:Brucemacfaddenia
3778:
3730:
3729:
3723:
3676:Oldfieldthomasia
3609:
3579:
3574:
3573:
3549:
3469:Posnanskytherium
3287:Calchaquitherium
3262:
3225:
3220:
3219:
3213:
3197:Trimerostephanos
3033:
3032:
3026:
2911:
2906:
2905:
2899:
2799:
2755:
2750:
2749:
2743:
2674:
2655:Peripantostylops
2623:
2596:
2595:
2589:
2541:
2540:
2528:
2527:
2517:
2512:
2446:
2416:
2397:Ernestohaeckelia
2379:
2374:
2373:
2367:
2287:Paramacrauchenia
2273:Neobrachytherium
2208:Proterotheriinae
2206:
2176:
2114:
2113:
2108:Proterotheriidae
2106:
2050:
1978:
1973:
1972:
1965:
1959:
1958:
1952:
1936:Xenorhinotherium
1887:Macraucheniopsis
1855:
1790:
1770:
1769:
1763:
1704:
1699:
1698:
1692:
1687:
1686:
1674:
1673:
1663:
1657:
1651:
1592:Uruguaytheriinae
1590:
1555:Astrapotheriinae
1553:
1523:
1468:
1467:
1463:Astrapotheriidae
1461:
1442:Tetragonostylops
1424:
1412:Eoastrapostylops
1404:
1403:
1397:
1335:
1307:Colombitheriidae
1305:
1300:
1299:
1293:
1245:
1240:
1239:
1233:
1207:Asmithwoodwardia
1198:Protolipternidae
1196:
1102:
1042:
1037:
1036:
1033:
1032:
1011:
1010:
963:
956:
949:
940:
939:
933:
932:
913:10.1666/05-100.1
899:(6): 1301–1307.
884:
878:
875:
869:
866:
860:
859:
819:
808:
805:
706:Pr. ameghinorum.
629:P. triangulidens
597:". Collected in
394:Mendoza Province
348:
340:
332:
324:
316:
289:
249:
242:
229:
216:
203:
153:
152:
119:
109:
46:
29:Temporal range:
21:
20:
4501:
4500:
4496:
4495:
4494:
4492:
4491:
4490:
4431:
4430:
4429:
4424:
4416:
4411:
4402:
4401:
4396:
4383:
4373:
4364:
4327:
4290:
4246:
4190:Hegetotheriinae
4178:Hegetotheriidae
4169:
4089:
4014:Altitypotherium
3996:
3957:
3913:
3780:Interatheriinae
3771:
3725:Interatheriidae
3716:
3655:Itaboraitherium
3648:Dolichostylodon
3634:Camargomendesia
3627:Brachystephanus
3602:
3590:Archaeopithecus
3572:
3530:
3392:Neoadinotherium
3336:Haplodontherium
3255:
3250:Proadinotherium
3202:
3019:
2964:Henricofilholia
2892:
2792:
2787:Trigonolophodon
2780:Homalodotherium
2732:
2667:
2616:
2582:
2535:
2518:
2500:
2469:
2439:
2409:
2348:
2315:Tetramerorhinus
2238:Diplasiotherium
2199:
2169:
2110:
2094:
2082:Sparnotheriodon
2043:
1969:
1941:
1915:Promacrauchenia
1857:Macraucheniinae
1848:
1829:Phoenixauchenia
1792:Cramaucheniinae
1783:
1765:Macraucheniidae
1756:
1681:
1664:
1627:
1583:
1546:
1516:
1454:
1417:
1386:
1328:
1282:
1226:
1189:
1163:Ricardocifellia
1095:
1027:
1005:
973:
967:
937:
936:
885:
881:
876:
872:
867:
863:
820:
811:
806:
795:
790:
742:
651:Hegetotheriidae
599:Chubut province
587:
492:
438:
402:
386:Chubut Province
351:
346:
338:
330:
322:
314:
308:
307:
287:
272:
269:
248:
240:
227:
218:Hegetotheriidae
214:
201:
147:
133:Chubut province
110:
108:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
41:
40:
38:
27:
17:
12:
11:
5:
4499:
4489:
4488:
4483:
4478:
4473:
4468:
4463:
4458:
4453:
4448:
4443:
4426:
4425:
4423:
4422:
4409:
4393:
4391:
4385:
4384:
4370:
4369:
4366:
4365:
4346:
4344:
4341:
4340:
4337:
4336:
4333:
4332:
4329:
4328:
4326:
4325:
4318:
4311:
4303:
4301:
4289:
4288:
4281:
4274:
4267:
4259:
4257:
4248:
4247:
4245:
4244:
4237:
4230:
4223:
4216:
4209:
4202:
4194:
4192:
4180:
4171:
4170:
4168:
4167:
4160:
4153:
4146:
4139:
4132:
4125:
4118:
4110:
4108:
4095:
4094:
4091:
4090:
4088:
4087:
4084:Typotheriopsis
4080:
4077:Rusconitherium
4073:
4066:
4059:
4052:
4045:
4038:
4031:
4024:
4017:
4009:
4007:
3998:
3997:
3995:
3994:
3987:
3978:
3972:
3963:
3962:
3959:
3958:
3956:
3955:
3948:
3941:
3934:
3926:
3924:
3922:Notopithecinae
3915:
3914:
3912:
3911:
3908:Santiagorothia
3904:
3901:Protypotherium
3897:
3890:
3887:Proargyrohyrax
3883:
3876:
3869:
3862:
3855:
3848:
3841:
3834:
3827:
3820:
3813:
3806:
3799:
3792:
3784:
3782:
3773:
3772:
3770:
3769:
3762:
3755:
3748:
3741:
3733:
3727:
3718:
3717:
3715:
3714:
3707:
3700:
3693:
3686:
3679:
3672:
3669:Maxschlosseria
3665:
3658:
3651:
3644:
3637:
3630:
3623:
3615:
3613:
3604:
3603:
3601:
3600:
3597:Teratopithecus
3593:
3585:
3583:
3571:
3570:
3563:
3555:
3553:
3544:
3543:
3540:
3539:
3536:
3535:
3532:
3531:
3529:
3528:
3521:
3514:
3507:
3504:Toxodontherium
3500:
3493:
3486:
3479:
3472:
3465:
3458:
3451:
3444:
3437:
3430:
3423:
3416:
3409:
3402:
3395:
3388:
3381:
3374:
3367:
3360:
3353:
3350:Hoffstetterius
3346:
3339:
3332:
3325:
3318:
3311:
3304:
3301:Charruatoxodon
3297:
3290:
3283:
3276:
3268:
3266:
3257:
3256:
3254:
3253:
3246:
3239:
3231:
3229:
3217:
3208:
3207:
3204:
3203:
3201:
3200:
3193:
3190:Teushentherium
3186:
3179:
3172:
3165:
3157:
3150:
3143:
3136:
3129:
3126:Pascualihippus
3122:
3115:
3108:
3101:
3094:
3091:Moqueguahippus
3087:
3080:
3073:
3066:
3059:
3055:Deuterotherium
3051:
3044:
3036:
3030:
3021:
3020:
3018:
3017:
3010:
3003:
2996:
2988:
2985:Martinmiguelia
2981:
2974:
2967:
2960:
2953:
2946:
2939:
2932:
2925:
2917:
2915:
2903:
2894:
2893:
2891:
2890:
2883:
2876:
2873:Pleurostylodon
2869:
2866:Pleurocoelodon
2862:
2855:
2848:
2841:
2834:
2827:
2820:
2813:
2805:
2803:
2794:
2793:
2791:
2790:
2783:
2776:
2773:Chasicotherium
2769:
2761:
2759:
2747:
2738:
2737:
2734:
2733:
2731:
2730:
2723:
2716:
2709:
2702:
2695:
2688:
2680:
2678:
2676:Notostylopidae
2669:
2668:
2666:
2665:
2658:
2651:
2648:Othnielmarshia
2644:
2637:
2634:Henricosbornia
2629:
2627:
2618:
2617:
2615:
2614:
2607:
2599:
2593:
2584:
2583:
2581:
2580:
2573:
2570:Carolodarwinia
2566:
2559:
2551:
2549:
2546:incertae sedis
2537:
2536:
2524:
2523:
2520:
2519:
2507:
2505:
2502:
2501:
2499:
2498:
2491:
2483:
2481:
2479:Incertae sedis
2475:
2474:
2471:
2470:
2468:
2467:
2460:
2452:
2450:
2448:Notonychopidae
2441:
2440:
2438:
2437:
2430:
2422:
2420:
2411:
2410:
2408:
2407:
2400:
2393:
2390:Amilnedwardsia
2385:
2383:
2371:
2362:
2361:
2358:
2357:
2354:
2353:
2350:
2349:
2347:
2346:
2339:
2332:
2325:
2322:Thoatheriopsis
2318:
2311:
2308:Proterotherium
2304:
2297:
2294:Picturotherium
2290:
2283:
2276:
2269:
2262:
2255:
2248:
2241:
2234:
2227:
2220:
2212:
2210:
2201:
2200:
2198:
2197:
2190:
2182:
2180:
2171:
2170:
2168:
2167:
2160:
2153:
2150:Prothoatherium
2146:
2139:
2132:
2125:
2122:Mesolicaphrium
2117:
2111:
2103:
2100:
2099:
2096:
2095:
2093:
2092:
2089:Victorlemoinea
2085:
2078:
2071:
2064:
2056:
2054:
2045:
2044:
2042:
2041:
2034:
2027:
2020:
2017:Paranisolambda
2013:
2006:
1999:
1992:
1984:
1982:
1980:Anisolambdinae
1970:
1967:Anisolambdidae
1962:
1956:
1947:
1946:
1943:
1942:
1940:
1939:
1932:
1925:
1918:
1911:
1904:
1901:Oxyodontherium
1897:
1890:
1883:
1876:
1869:
1861:
1859:
1850:
1849:
1847:
1846:
1839:
1832:
1825:
1818:
1811:
1804:
1796:
1794:
1785:
1784:
1782:
1781:
1773:
1767:
1758:
1757:
1755:
1754:
1747:
1740:
1733:
1725:
1718:
1710:
1708:
1696:
1683:
1682:
1670:
1669:
1666:
1665:
1646:
1644:
1641:
1640:
1637:
1636:
1633:
1632:
1629:
1628:
1626:
1625:
1618:
1615:Uruguaytherium
1611:
1608:Hilarcotherium
1604:
1596:
1594:
1585:
1584:
1582:
1581:
1574:
1571:Astrapotherium
1567:
1559:
1557:
1548:
1547:
1545:
1544:
1537:
1534:Albertogaudrya
1529:
1527:
1518:
1517:
1515:
1514:
1507:
1500:
1493:
1490:Comahuetherium
1486:
1479:
1471:
1465:
1456:
1455:
1453:
1452:
1449:Trigonostylops
1445:
1438:
1430:
1428:
1419:
1418:
1416:
1415:
1407:
1401:
1392:
1391:
1388:
1387:
1385:
1384:
1377:
1374:Propyrotherium
1370:
1363:
1360:Carolozittelia
1356:
1353:Berracotherium
1349:
1341:
1339:
1330:
1329:
1327:
1326:
1319:
1316:Colombitherium
1311:
1309:
1297:
1288:
1287:
1284:
1283:
1281:
1280:
1273:
1266:
1259:
1251:
1249:
1237:
1228:
1227:
1225:
1224:
1217:
1210:
1202:
1200:
1191:
1190:
1188:
1187:
1180:
1173:
1166:
1159:
1152:
1149:Paulogervaisia
1145:
1138:
1130:
1127:Ernestokokenia
1123:
1116:
1108:
1106:
1104:Didolodontidae
1097:
1096:
1094:
1093:
1086:
1079:
1072:
1064:
1057:
1049:
1047:
1029:
1028:
1007:
1006:
1004:
1003:
1001:Laurasiatheria
997:
991:
985:
978:
975:
974:
966:
965:
958:
951:
943:
935:
934:
879:
870:
861:
834:(6): 643–662.
809:
792:
791:
789:
786:
741:
738:
730:P. ameghinorum
718:P. ameghinorum
710:P. ameghinorum
690:P. ameghinorum
586:
585:Classification
583:
491:
488:
437:
434:
401:
398:
353:
352:
350:
349:
341:
333:
331:Ameghino, 1901
325:
323:Ameghino, 1897
317:
315:Ameghino, 1897
305:
304:
303:
300:
299:
293:
292:
291:
290:
288:Ameghino, 1897
279:
278:
274:
273:
271:Ameghino, 1897
270:
260:
259:
253:
252:
250:Ameghino, 1897
238:
234:
233:
225:
221:
220:
212:
208:
207:
199:
195:
194:
189:
185:
184:
179:
175:
174:
169:
165:
164:
159:
155:
154:
141:
140:
121:
120:
112:
111:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
31:Late Oligocene
28:
15:
9:
6:
4:
3:
2:
4498:
4487:
4484:
4482:
4479:
4477:
4474:
4472:
4469:
4467:
4464:
4462:
4459:
4457:
4454:
4452:
4449:
4447:
4444:
4442:
4439:
4438:
4436:
4419:
4414:
4410:
4405:
4399:
4395:
4394:
4392:
4390:
4386:
4382:
4377:
4362:
4358:
4356:
4350:
4324:
4323:
4319:
4317:
4316:
4312:
4310:
4309:
4305:
4304:
4302:
4300:
4293:
4287:
4286:
4282:
4280:
4279:
4275:
4273:
4272:
4271:Propachyrucos
4268:
4266:
4265:
4261:
4260:
4258:
4256:
4255:Pachyrukhinae
4249:
4243:
4242:
4238:
4236:
4235:
4231:
4229:
4228:
4224:
4222:
4221:
4217:
4215:
4214:
4213:Hegetotherium
4210:
4208:
4207:
4203:
4201:
4200:
4199:Ethegotherium
4196:
4195:
4193:
4191:
4184:
4181:
4179:
4172:
4166:
4165:
4161:
4159:
4158:
4154:
4152:
4151:
4147:
4145:
4144:
4140:
4138:
4137:
4133:
4131:
4130:
4126:
4124:
4123:
4119:
4117:
4116:
4112:
4111:
4109:
4107:
4100:
4086:
4085:
4081:
4079:
4078:
4074:
4072:
4071:
4067:
4065:
4064:
4060:
4058:
4057:
4053:
4051:
4050:
4046:
4044:
4043:
4039:
4037:
4036:
4035:Eutypotherium
4032:
4030:
4029:
4028:Eotypotherium
4025:
4023:
4022:
4018:
4016:
4015:
4011:
4010:
4008:
4006:
4005:Mesotheriinae
3999:
3993:
3992:
3988:
3985:
3984:
3980:
3979:
3976:
3973:
3971:
3970:Mesotheriidae
3964:
3954:
3953:
3952:Transpithecus
3949:
3947:
3946:
3942:
3940:
3939:
3935:
3933:
3932:
3928:
3927:
3925:
3923:
3916:
3910:
3909:
3905:
3903:
3902:
3898:
3896:
3895:
3891:
3889:
3888:
3884:
3882:
3881:
3877:
3875:
3874:
3870:
3868:
3867:
3863:
3861:
3860:
3856:
3854:
3853:
3852:Interatherium
3849:
3847:
3846:
3845:Federicoanaya
3842:
3840:
3839:
3835:
3833:
3832:
3828:
3826:
3825:
3824:Choichephilum
3821:
3819:
3818:
3814:
3812:
3811:
3807:
3805:
3804:
3800:
3798:
3797:
3793:
3791:
3790:
3789:Archaeophylus
3786:
3785:
3783:
3781:
3774:
3768:
3767:
3763:
3761:
3760:
3756:
3754:
3753:
3749:
3747:
3746:
3742:
3740:
3739:
3735:
3734:
3731:
3728:
3726:
3719:
3713:
3712:
3711:Xenostephanus
3708:
3706:
3705:
3704:Ultrapithecus
3701:
3699:
3698:
3694:
3692:
3691:
3687:
3685:
3684:
3680:
3678:
3677:
3673:
3671:
3670:
3666:
3664:
3663:
3659:
3657:
3656:
3652:
3650:
3649:
3645:
3643:
3642:
3638:
3636:
3635:
3631:
3629:
3628:
3624:
3622:
3621:
3617:
3616:
3614:
3612:
3605:
3599:
3598:
3594:
3592:
3591:
3587:
3586:
3584:
3582:
3575:
3569:
3568:
3567:Griphotherion
3564:
3562:
3561:
3557:
3556:
3554:
3552:
3545:
3527:
3526:
3522:
3520:
3519:
3515:
3513:
3512:
3508:
3506:
3505:
3501:
3499:
3498:
3494:
3492:
3491:
3490:Stereotoxodon
3487:
3485:
3484:
3483:Stenotephanos
3480:
3478:
3477:
3476:Prototrigodon
3473:
3471:
3470:
3466:
3464:
3463:
3462:Plesiotoxodon
3459:
3457:
3456:
3452:
3450:
3449:
3448:Piauhytherium
3445:
3443:
3442:
3441:Pericotoxodon
3438:
3436:
3435:
3431:
3429:
3428:
3424:
3422:
3421:
3420:Ocnerotherium
3417:
3415:
3414:
3410:
3408:
3407:
3403:
3401:
3400:
3396:
3394:
3393:
3389:
3387:
3386:
3382:
3380:
3379:
3375:
3373:
3372:
3368:
3366:
3365:
3361:
3359:
3358:
3357:Hyperoxotodon
3354:
3352:
3351:
3347:
3345:
3344:
3340:
3338:
3337:
3333:
3331:
3330:
3326:
3324:
3323:
3322:Falcontoxodon
3319:
3317:
3316:
3312:
3310:
3309:
3305:
3303:
3302:
3298:
3296:
3295:
3291:
3289:
3288:
3284:
3282:
3281:
3280:Andinotoxodon
3277:
3275:
3274:
3270:
3269:
3267:
3265:
3258:
3252:
3251:
3247:
3245:
3244:
3240:
3238:
3237:
3233:
3232:
3230:
3228:
3221:
3218:
3216:
3209:
3199:
3198:
3194:
3192:
3191:
3187:
3185:
3184:
3180:
3178:
3177:
3173:
3171:
3170:
3166:
3163:
3162:
3158:
3156:
3155:
3151:
3149:
3148:
3144:
3142:
3141:
3137:
3135:
3134:
3133:Patagonhippus
3130:
3128:
3127:
3123:
3121:
3120:
3116:
3114:
3113:
3109:
3107:
3106:
3102:
3100:
3099:
3095:
3093:
3092:
3088:
3086:
3085:
3084:Mendozahippus
3081:
3079:
3078:
3074:
3072:
3071:
3067:
3065:
3064:
3063:Edvardocopeia
3060:
3057:
3056:
3052:
3050:
3049:
3045:
3043:
3042:
3038:
3037:
3034:
3031:
3029:
3022:
3016:
3015:
3014:Termastherium
3011:
3009:
3008:
3004:
3002:
3001:
2997:
2994:
2993:
2989:
2987:
2986:
2982:
2980:
2979:
2975:
2973:
2972:
2968:
2966:
2965:
2961:
2959:
2958:
2954:
2952:
2951:
2947:
2945:
2944:
2940:
2938:
2937:
2933:
2931:
2930:
2926:
2924:
2923:
2919:
2918:
2916:
2914:
2907:
2904:
2902:
2895:
2889:
2888:
2887:Thomashuxleya
2884:
2882:
2881:
2877:
2875:
2874:
2870:
2868:
2867:
2863:
2861:
2860:
2856:
2854:
2853:
2849:
2847:
2846:
2842:
2840:
2839:
2835:
2833:
2832:
2828:
2826:
2825:
2824:Distylophorus
2821:
2819:
2818:
2817:Coelostylodon
2814:
2812:
2811:
2807:
2806:
2804:
2802:
2795:
2789:
2788:
2784:
2782:
2781:
2777:
2775:
2774:
2770:
2768:
2767:
2763:
2762:
2760:
2758:
2751:
2748:
2746:
2739:
2729:
2728:
2724:
2722:
2721:
2717:
2715:
2714:
2710:
2708:
2707:
2706:Homalostylops
2703:
2701:
2700:
2696:
2694:
2693:
2689:
2687:
2686:
2682:
2681:
2679:
2677:
2670:
2664:
2663:
2659:
2657:
2656:
2652:
2650:
2649:
2645:
2643:
2642:
2638:
2636:
2635:
2631:
2630:
2628:
2626:
2619:
2613:
2612:
2608:
2606:
2605:
2601:
2600:
2597:
2594:
2592:
2591:Notioprogonia
2585:
2579:
2578:
2574:
2572:
2571:
2567:
2565:
2564:
2560:
2558:
2557:
2553:
2552:
2550:
2548:
2547:
2542:
2538:
2534:
2529:
2525:
2516:
2511:
2497:
2496:
2492:
2490:
2489:
2485:
2484:
2482:
2480:
2476:
2466:
2465:
2461:
2459:
2458:
2454:
2453:
2451:
2449:
2442:
2436:
2435:
2431:
2429:
2428:
2424:
2423:
2421:
2419:
2412:
2406:
2405:
2401:
2399:
2398:
2394:
2392:
2391:
2387:
2386:
2384:
2382:
2375:
2372:
2370:
2363:
2345:
2344:
2340:
2338:
2337:
2333:
2331:
2330:
2326:
2324:
2323:
2319:
2317:
2316:
2312:
2310:
2309:
2305:
2303:
2302:
2301:Prolicaphrium
2298:
2296:
2295:
2291:
2289:
2288:
2284:
2282:
2281:
2280:Neolicaphrium
2277:
2275:
2274:
2270:
2268:
2267:
2263:
2261:
2260:
2256:
2254:
2253:
2249:
2247:
2246:
2242:
2240:
2239:
2235:
2233:
2232:
2228:
2226:
2225:
2224:Brachytherium
2221:
2219:
2218:
2214:
2213:
2211:
2209:
2202:
2196:
2195:
2191:
2189:
2188:
2184:
2183:
2181:
2179:
2178:Megadolodinae
2172:
2166:
2165:
2161:
2159:
2158:
2154:
2152:
2151:
2147:
2145:
2144:
2140:
2138:
2137:
2133:
2131:
2130:
2126:
2124:
2123:
2119:
2118:
2115:
2112:
2109:
2101:
2091:
2090:
2086:
2084:
2083:
2079:
2077:
2076:
2072:
2070:
2069:
2065:
2063:
2062:
2061:Heteroglyphis
2058:
2057:
2055:
2053:
2046:
2040:
2039:
2035:
2033:
2032:
2028:
2026:
2025:
2021:
2019:
2018:
2014:
2012:
2011:
2007:
2005:
2004:
2000:
1998:
1997:
1993:
1991:
1990:
1986:
1985:
1983:
1981:
1974:
1971:
1968:
1960:
1957:
1955:
1948:
1938:
1937:
1933:
1931:
1930:
1926:
1924:
1923:
1919:
1917:
1916:
1912:
1910:
1909:
1908:Paranauchenia
1905:
1903:
1902:
1898:
1896:
1895:
1891:
1889:
1888:
1884:
1882:
1881:
1877:
1875:
1874:
1870:
1868:
1867:
1863:
1862:
1860:
1858:
1851:
1845:
1844:
1840:
1838:
1837:
1833:
1831:
1830:
1826:
1824:
1823:
1819:
1817:
1816:
1812:
1810:
1809:
1808:Coniopternium
1805:
1803:
1802:
1798:
1797:
1795:
1793:
1786:
1780:
1779:
1775:
1774:
1771:
1768:
1766:
1759:
1753:
1752:
1748:
1746:
1745:
1741:
1739:
1738:
1737:Proheptaconus
1734:
1731:
1730:
1726:
1724:
1723:
1719:
1717:
1716:
1712:
1711:
1709:
1707:
1700:
1697:
1695:
1688:
1684:
1680:
1675:
1671:
1662:
1658:
1656:
1650:
1624:
1623:
1619:
1617:
1616:
1612:
1610:
1609:
1605:
1603:
1602:
1598:
1597:
1595:
1593:
1586:
1580:
1579:
1575:
1573:
1572:
1568:
1566:
1565:
1561:
1560:
1558:
1556:
1549:
1543:
1542:
1538:
1536:
1535:
1531:
1530:
1528:
1526:
1519:
1513:
1512:
1508:
1506:
1505:
1501:
1499:
1498:
1494:
1492:
1491:
1487:
1485:
1484:
1480:
1478:
1477:
1473:
1472:
1469:
1466:
1464:
1457:
1451:
1450:
1446:
1444:
1443:
1439:
1437:
1436:
1432:
1431:
1429:
1427:
1420:
1414:
1413:
1409:
1408:
1405:
1402:
1400:
1399:Astrapotheria
1393:
1383:
1382:
1378:
1376:
1375:
1371:
1369:
1368:
1364:
1362:
1361:
1357:
1355:
1354:
1350:
1348:
1347:
1343:
1342:
1340:
1338:
1337:Pyrotheriidae
1331:
1325:
1324:
1320:
1318:
1317:
1313:
1312:
1310:
1308:
1301:
1298:
1296:
1289:
1279:
1278:
1274:
1272:
1271:
1267:
1265:
1264:
1260:
1258:
1257:
1253:
1252:
1250:
1248:
1241:
1238:
1236:
1229:
1223:
1222:
1221:Protolipterna
1218:
1216:
1215:
1211:
1209:
1208:
1204:
1203:
1201:
1199:
1192:
1186:
1185:
1181:
1179:
1178:
1174:
1172:
1171:
1167:
1165:
1164:
1160:
1158:
1157:
1153:
1151:
1150:
1146:
1144:
1143:
1139:
1136:
1135:
1131:
1129:
1128:
1124:
1122:
1121:
1117:
1115:
1114:
1110:
1109:
1107:
1105:
1098:
1092:
1091:
1087:
1085:
1084:
1080:
1078:
1077:
1073:
1070:
1069:
1065:
1063:
1062:
1058:
1056:
1055:
1051:
1050:
1048:
1045:
1038:
1034:
1030:
1025:
1024:Astrapotheria
1021:
1017:
1012:
1008:
1002:
998:
996:
992:
990:
986:
984:
980:
979:
976:
972:
964:
959:
957:
952:
950:
945:
944:
941:
930:
926:
922:
918:
914:
910:
906:
902:
898:
894:
890:
883:
874:
865:
857:
853:
849:
845:
841:
837:
833:
829:
825:
818:
816:
814:
804:
802:
800:
798:
793:
785:
783:
779:
775:
774:South America
771:
767:
763:
762:interatheriid
759:
758:
753:
752:
751:Propachyrucos
747:
737:
735:
731:
727:
726:Propachyrucos
723:
719:
715:
711:
707:
703:
699:
695:
691:
687:
686:
685:Propachyrucos
680:
678:
677:
672:
668:
667:Pachyrukhinae
664:
660:
656:
652:
648:
644:
642:
638:
634:
630:
626:
625:
620:
616:
612:
608:
600:
596:
591:
582:
580:
579:
574:
570:
569:
562:
559:
557:
553:
549:
545:
541:
537:
533:
529:
525:
521:
517:
513:
509:
505:
501:
497:
487:
483:
481:
477:
473:
469:
465:
461:
457:
453:
449:
448:
443:
433:
431:
427:
426:
421:
420:
415:
414:
409:
408:
407:Propachyrucos
397:
395:
391:
387:
383:
379:
375:
374:hegetotheriid
372:
368:
365:
361:
360:
347:Chaffee, 1952
345:
342:
339:Simpson, 1945
337:
334:
329:
326:
321:
318:
313:
310:
309:
301:
298:
294:
286:
283:
282:
280:
275:
268:
267:
261:
258:
254:
247:
246:
239:
236:
235:
232:
231:Pachyrukhinae
226:
223:
222:
219:
213:
210:
209:
206:
200:
197:
196:
193:
190:
187:
186:
183:
180:
177:
176:
173:
170:
167:
166:
163:
160:
157:
156:
151:
146:
142:
138:
134:
130:
126:
122:
118:
113:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
45:
36:
32:
26:
22:
19:
4389:Prosotherium
4388:
4352:
4320:
4315:Paedotherium
4313:
4306:
4299:Pachyrukhini
4283:
4278:Prosotherium
4277:
4276:
4269:
4262:
4241:Sallatherium
4239:
4232:
4225:
4218:
4211:
4204:
4197:
4162:
4155:
4148:
4143:Pascualhyrax
4141:
4134:
4127:
4120:
4115:Archaeohyrax
4113:
4082:
4075:
4068:
4061:
4054:
4047:
4042:Hypsitherium
4040:
4033:
4026:
4019:
4012:
3991:Trachytherus
3989:
3981:
3950:
3945:Notopithecus
3943:
3936:
3931:Antepithecus
3929:
3906:
3899:
3892:
3885:
3878:
3873:Neoicochilus
3871:
3866:Miocochilius
3864:
3857:
3850:
3843:
3838:Eopachyrucos
3836:
3829:
3822:
3815:
3808:
3803:Boleatherium
3801:
3794:
3787:
3766:Punapithecus
3764:
3757:
3750:
3743:
3738:Antofagastia
3736:
3709:
3702:
3697:Tsamnichoria
3695:
3688:
3681:
3674:
3667:
3662:Kibenikhoria
3660:
3653:
3646:
3639:
3632:
3625:
3618:
3595:
3588:
3565:
3558:
3523:
3516:
3509:
3502:
3495:
3488:
3481:
3474:
3467:
3460:
3453:
3446:
3439:
3434:Paratrigodon
3432:
3425:
3418:
3411:
3406:Nesodonopsis
3404:
3397:
3390:
3383:
3376:
3369:
3362:
3355:
3348:
3341:
3334:
3327:
3320:
3313:
3306:
3299:
3292:
3285:
3278:
3271:
3264:Toxodontinae
3248:
3241:
3236:Adinotherium
3234:
3227:Nesodontinae
3215:Toxodontidae
3195:
3188:
3181:
3174:
3167:
3159:
3152:
3145:
3140:Perhippidion
3138:
3131:
3124:
3117:
3110:
3103:
3096:
3089:
3082:
3075:
3070:Eomorphippus
3068:
3061:
3053:
3048:Argyrohippus
3046:
3039:
3028:Notohippidae
3012:
3007:Taubatherium
3005:
2998:
2990:
2983:
2976:
2971:Huilatherium
2969:
2962:
2955:
2950:Elmerriggsia
2948:
2941:
2934:
2929:Ancylocoelus
2927:
2922:Anayatherium
2920:
2913:Leontiniidae
2901:Eutoxodontia
2885:
2878:
2871:
2864:
2859:Periphragnis
2857:
2850:
2843:
2836:
2829:
2822:
2815:
2808:
2785:
2778:
2771:
2764:
2725:
2718:
2711:
2704:
2697:
2692:Chilestylops
2690:
2685:Boreastylops
2683:
2660:
2653:
2646:
2641:Nanolophodon
2639:
2632:
2609:
2604:Satshatemnus
2602:
2575:
2568:
2561:
2554:
2544:
2533:Notoungulata
2493:
2486:
2478:
2462:
2455:
2432:
2425:
2418:Indaleciidae
2402:
2395:
2388:
2343:Villarroelia
2341:
2334:
2327:
2320:
2313:
2306:
2299:
2292:
2285:
2278:
2271:
2264:
2257:
2250:
2243:
2236:
2231:Diadiaphorus
2229:
2222:
2215:
2192:
2185:
2164:Salladolodus
2162:
2155:
2148:
2141:
2134:
2127:
2120:
2087:
2080:
2075:Phoradiadius
2073:
2066:
2059:
2036:
2031:Protheosodon
2029:
2022:
2015:
2010:Lambdaconops
2008:
2001:
1996:Eolicaphrium
1994:
1987:
1934:
1929:Windhausenia
1927:
1920:
1913:
1906:
1899:
1894:Micrauchenia
1892:
1885:
1880:Macrauchenia
1878:
1873:Huayqueriana
1871:
1864:
1841:
1836:Pternoconius
1834:
1827:
1822:Llullataruca
1820:
1815:Cramauchenia
1813:
1806:
1799:
1776:
1749:
1742:
1735:
1727:
1720:
1713:
1694:Eulitopterna
1652:
1620:
1613:
1606:
1599:
1576:
1569:
1562:
1541:Astraponotus
1539:
1532:
1509:
1502:
1495:
1488:
1481:
1474:
1447:
1440:
1433:
1410:
1379:
1372:
1365:
1358:
1351:
1346:Baguatherium
1344:
1321:
1314:
1275:
1268:
1261:
1254:
1219:
1212:
1205:
1182:
1175:
1168:
1161:
1154:
1147:
1140:
1132:
1125:
1118:
1111:
1088:
1081:
1074:
1066:
1059:
1052:
1044:Kollpaniidae
999:Superorder:
896:
892:
882:
873:
864:
831:
827:
755:
749:
746:Prosotherium
745:
743:
740:Paleobiology
733:
729:
725:
721:
717:
713:
709:
705:
701:
697:
693:
689:
683:
681:
676:Paedotherium
674:
671:Prosotherium
670:
663:Prosotherium
662:
659:Notoungulata
647:Prosotherium
646:
645:
640:
636:
632:
628:
623:
622:
607:Prosotherium
606:
604:
601:, Argentina.
594:
578:Paedotherium
576:
572:
566:
563:
560:
555:
527:
523:
493:
484:
445:
439:
423:
419:Paedotherium
417:
411:
405:
403:
359:Prosotherium
358:
357:
356:
343:
335:
327:
319:
311:
284:
265:
264:
257:Type species
245:Prosotherium
244:
243:
205:Notoungulata
128:
124:
25:Prosotherium
24:
18:
4322:Tremacyllus
4308:Pachyrukhos
4285:Tremacyllus
4049:Mesotherium
3880:Patriarchus
3859:Juchuysillu
3817:Caenophilus
3796:Argyrohyrax
3518:Trigodonops
3427:Palyeidodon
3413:Nonotherium
3385:Mixotoxodon
3378:Minitoxodon
3371:Mesotoxodon
3343:Hemixotodon
3308:Dinotoxodon
3294:Ceratoxodon
3169:Rhynchippus
3147:Plexotemnus
3119:Pampahippus
3041:Acoelohyrax
2852:Pampatemnus
2845:Lophocoelus
2831:Hedralophus
2810:Anisotemnus
2801:Isotemnidae
2727:Parastylops
2713:Notostylops
2662:Simpsonotus
2577:Perutherium
2563:Archaeogaia
2457:Notonychops
2427:Adiantoides
2404:Rutimeyeria
2336:Uruguayodon
2329:Thoatherium
2266:Lambdaconus
2217:Anisolophus
2194:Megadolodus
2143:Promylophis
2136:Olisanophus
2024:Polyacrodon
1989:Anisolambda
1778:Polymorphis
1751:Tricoelodus
1729:Proectocion
1722:Proadiantus
1706:Adianthidae
1476:Antarctodon
1381:Pyrotherium
1247:Carodniidae
1235:Xenungulata
1214:Miguelsoria
1083:Simoclaenus
1068:Pascualodus
1016:Xenungulata
694:P. simpsoni
633:P. robustum
573:Pachyrukhos
568:Pachyrukhos
556:Pachyrukhos
548:cuboid bone
528:Pachyrukhos
524:Pachyrukhos
508:Metacarpals
447:Pachyrukhos
425:Medistyllus
400:Description
378:Argentinian
224:Subfamily:
42:29–21
4441:Typotheres
4435:Categories
4264:Medistylus
4157:Pseudhyrax
3560:Campanorco
3551:Typotheria
3399:Neotoxodon
3273:Abothrodon
3183:Stilhippus
3112:Notohippus
3105:Nesohippus
3098:Morphippus
3077:Eurygenium
3000:Scarrittia
2745:Toxodontia
2495:Proacrodon
2369:Notopterna
2259:Epitherium
2245:Eoauchenia
2129:Neodolodus
2068:Notiolofos
1801:Caliphrium
1679:Litopterna
1483:Astrapodon
1295:Pyrotheria
1270:Notoetayoa
1156:Raulvaccia
1134:Escribania
1120:Enneoconus
1090:Tiuclaenus
1020:Pyrotheria
969:Genera of
788:References
757:Medistylus
734:P. simpson
637:P. quartum
605:The genus
552:metatarsal
540:talus bone
413:Pachrukhos
388:, and the
285:P. garzoni
4404:Q60978547
4164:Punahyrax
3983:Fiandraia
3831:Cochilius
3641:Colbertia
3620:Allalmeia
3455:Pisanodon
3364:Mesenodon
3329:Gyrinodon
3315:Eutomodus
2978:Leontinia
2880:Rhyphodon
2838:Isotemnus
2611:Seudenius
2556:Acoelodus
2434:Indalecia
2252:Epecuenia
1843:Theosodon
1744:Thadanius
1715:Adianthus
1497:Liarthrus
1367:Griphodon
1170:Saltaodus
1113:Didolodus
1076:Pucanodus
1061:Molinodus
1054:Andinodus
981:Kingdom:
921:0022-3360
856:1631-0683
619:Patagonia
544:calcaneus
512:phalanges
476:premolars
371:Oligocene
168:Kingdom:
162:Eukaryota
137:Argentina
4451:Deseadan
4398:Wikidata
3752:Johnbell
3745:Ignigena
3690:Suniodon
3683:Paginula
3511:Trigodon
3161:Purperia
2992:Purperia
2943:Coquenia
2936:Colpodon
2766:Asmodeus
2720:Otrhonia
2488:Kyraodus
2464:Requisia
2187:Bounodus
1866:Cullinia
1504:Maddenia
1435:Shecenia
1323:Proticia
1277:Rodcania
1256:Carodnia
1184:Xesmodon
1177:Umayodus
1142:Lamegoia
995:Mammalia
989:Chordata
987:Phylum:
983:Animalia
929:55259241
770:Cenozoic
688:species
480:cementum
468:incisors
464:mandible
462:and the
369:of late
297:Synonyms
277:Species
211:Family:
192:Mammalia
182:Chordata
178:Phylum:
172:Animalia
158:Domain:
35:Deseadan
4136:Eohyrax
3759:Munizia
3525:Xotodon
3497:Toxodon
3243:Nesodon
3176:Rosendo
1578:Scaglia
993:Class:
901:Bibcode
836:Bibcode
782:Bolivia
778:Uruguay
496:humerus
460:maxilla
452:auricle
430:rabbits
364:extinct
237:Genus:
198:Order:
188:Class:
3154:Puelia
2957:Gualta
2038:Wainka
1263:Etayoa
927:
919:
854:
655:rodent
536:fibula
530:, the
516:pelvis
504:radius
472:molars
458:. The
436:Crania
416:, and
362:is an
4418:43447
925:S2CID
532:tibia
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