113:
375:
387:
142:
310:, but their bodies are more compact and flattened, an adaptation for living and hiding under rocks. Porcelain crabs are quite fragile animals, and often shed their limbs to escape predators, hence their name. The lost
1092:
Jonas Keiler, Stefan
Richter & Christian S. Wirkner (2014). "Evolutionary morphology of the organ systems in squat lobsters and porcelain crabs (crustacea: Decapoda: Anomala): an insight into carcinization".
278:. They have flattened bodies as an adaptation for living in rock crevices. They are delicate, readily losing limbs when attacked, and use their large claws for maintaining territories. They first appeared in the
1178:"Annotated checklist of anomuran decapod crustaceans of the world (exclusive of the Kiwaoidea and families Chirostylidae and Galatheidae of the Galatheoidea) Part II – Porcellanidae"
1245:
451:
431:
739:
499:
829:
809:
669:
559:
112:
839:
795:
579:
569:
819:
599:
549:
469:
1606:
1452:
1274:"Extreme diversity and parasitism of Late Jurassic squat lobsters (Decapoda: Galatheoidea) and the oldest records of porcellanids and galatheids"
729:
589:
509:
868:
489:
1491:
679:
639:
629:
519:
749:
649:
609:
1611:
539:
529:
1177:
759:
1139:
1426:
1465:
1022:
982:
944:
908:
1601:
1470:
1047:
342:, whereby a noncrab-like animal (in this case a relative of a squat lobster) evolves into an animal that resembles a
1563:
1212:
1496:
866:
of
Austria. However, it was subsequently determined to be a true crab. With the new oldest porcelain crab being
900:
1478:
17:
463:
As of 2018, some 4723 extant species of porcelain crab had been described, divided among these 30 genera:
141:
1387:
1374:
1392:
936:
1545:
1568:
1506:
1039:
391:
1208:
1591:
1352:
1273:
1532:
1314:
410:
and shorelines, startled creatures scurrying away when a stone is lifted. They feed by combing
379:
966:
892:
1555:
1527:
1457:
1095:
1002:
928:
1519:
1361:
699:
435:, found in large numbers under rocks in the intertidal, and the red-and-white polka-dotted
419:
238:
1190:
8:
1151:
1040:"Mitochondrial gene rearrangements confirm the parallel evolution of the crab-like form"
1120:
1069:
323:
136:
1366:
1238:"Earliest known Porcellanidae (Decapoda: Anomura: Galatheoidea) (Jurassic: Tithonian)"
1514:
1379:
1295:
1147:
1112:
1074:
1038:
C. L. Morrison; A. W. Harvey; S. Lavery; K. Tieu; Y. Huang; C. W. Cunningham (2001).
1018:
1014:
978:
940:
904:
437:
355:
1258:
1124:
1596:
1285:
1254:
1104:
1064:
1056:
974:
479:
374:
619:
346:. Porcelain crabs can be distinguished from true crabs by the apparent number of
1483:
1290:
1091:
1337:
1237:
386:
350:(three instead of four pairs; the fourth pair is reduced and held against the
294:
Porcelain crabs are small, usually with body widths less than 15 millimetres (
1585:
1299:
771:
426:
339:
319:
307:
283:
70:
406:. They are common under rocks, and can often be found and observed on rocky
1116:
1078:
1060:
689:
399:
223:
183:
1037:
1439:
1346:
971:
Marine decapod
Crustacea of southern Australia: a guide to identification
659:
442:
203:
45:
1405:
1108:
860:
719:
267:
90:
55:
1235:
366:
of the porcelain crab is long and folded underneath it, free to move.
1444:
1413:
863:
856:
851:
446:
403:
347:
327:
311:
279:
173:
153:
95:
1431:
1308:
1418:
1400:
1331:
1137:
709:
411:
359:
351:
264:
193:
85:
80:
65:
60:
50:
1185:
1175:
363:
315:
213:
100:
75:
964:
441:, which lives commensally within the shells inhabited by large
163:
1138:
P. McLaughlin, S. Ahyong & J. K. Lowry (October 2, 2002).
926:
1540:
407:
1246:
Neues
Jahrbuch für Geologie und Paläontologie, Abhandlungen
415:
398:
Porcelain crabs live in all the world's oceans, except the
343:
275:
326:
struggles, but not for catching food. The fifth pair of
1272:
Robins, Cristina M; Klompmaker, Adiël A (2019-11-14).
1236:
Carrie E. Schweitzer & Rodney M. Feldmann (2010).
414:
and other organic particles from the water using long
425:
Some of the common species of porcelain crabs in the
899:. Encyclopedias of the Natural World, Issue 1.
790:, and a further six genera known only from fossils:
418:(feathery hair- or bristle-like structures) on the
1176:Masayuki Osawa & Patsy A. McLaughlin (2010).
891:Denny, Mark W. & Gaines, Steven Dean (2007).
1583:
1271:
965:Gary C. B. Poore & Shane T. Ahyong (2004).
849:The earliest claimed porcelain crab fossil was
927:Brian Morton & John Edward Morton (1993).
1231:
1229:
306:inch). They share the general body plan of a
1171:
1169:
890:
369:
1226:
922:
920:
897:Encyclopedia of Tidepools and Rocky Shores
111:
1289:
1278:Zoological Journal of the Linnean Society
1068:
1166:
385:
373:
358:originating on the front outside of the
996:
994:
960:
958:
956:
917:
886:
884:
774:of porcelain crabs includes species of
14:
1607:Extant Late Jurassic first appearances
1584:
1313:
1312:
1206:
1000:
1546:F304879B-FFCA-FF9D-FF35-F96E2861F97C
1507:3743b640-3f08-4c01-a1f7-9ba21ccaa120
1189:. Suppl. 23: 109–129. Archived from
991:
953:
881:
274:, which superficially resemble true
24:
1612:Taxa named by Adrian Hardy Haworth
1048:Proceedings of the Royal Society B
933:The Sea Shore Ecology of Hong Kong
338:Porcelain crabs are an example of
330:is reduced and used for cleaning.
286:epoch, 145–152 million years ago.
25:
1623:
1213:World Register of Marine Species
140:
120:Southeastern Pacific species of
43:
1265:
834:De Angeli & Garassino, 2002
824:De Angeli & Garassino, 2002
814:De Angeli & Garassino, 2002
1200:
1131:
1085:
1031:
901:University of California Press
289:
13:
1:
1140:"Porcellanidae Haworth, 1825"
967:"Porcellanidae Haworth, 1825"
875:
318:. Porcelain crabs have large
458:
333:
322:(claws), which are used for
7:
1602:Tithonian first appearances
1259:10.1127/0077-7749/2010/0096
474:Nakasone & Miyake, 1969
314:can grow back over several
10:
1628:
1013:. Periplus Nature Guides.
937:Hong Kong University Press
844:Müller & Collins, 1991
1321:
1291:10.1093/zoolinnean/zlz067
1005:Neopetrolisthes maculatus
1001:Allen, Gerald R. (1997).
392:Neopetrolisthes maculatus
270:in the widespread family
251:
246:
137:Scientific classification
135:
119:
110:
34:
872:from the same locality.
370:Biogeography and ecology
544:Ng & Nakasone, 1993
1061:10.1098/rspb.2001.1886
452:Petrolisthes japonicus
432:Petrolisthes quadratus
395:
383:
380:Porcellana platycheles
1528:Paleobiology Database
1096:Journal of Morphology
389:
377:
27:Family of crustaceans
1011:Tropical Marine Life
977:. pp. 242–246.
903:. pp. 164–176.
939:. pp. 86–125.
744:Sankarankutty, 1962
740:Pseudoporcellanella
1109:10.1002/jmor.20311
396:
384:
1579:
1578:
1515:Open Tree of Life
1315:Taxon identifiers
1154:on April 29, 2005
1148:Australian Museum
1144:Anomura: Families
1055:(1489): 345–350.
1024:978-962-593-157-9
1015:Tuttle Publishing
984:978-0-643-06906-0
946:978-962-209-027-9
910:978-0-520-25118-2
845:
835:
825:
815:
805:
765:
755:
745:
735:
725:
715:
705:
695:
685:
675:
665:
655:
645:
635:
625:
615:
605:
595:
585:
575:
565:
555:
545:
535:
525:
515:
505:
500:Capilliporcellana
495:
485:
475:
438:Porcellana sayana
258:
257:
242:
16:(Redirected from
1619:
1572:
1571:
1559:
1558:
1549:
1548:
1536:
1535:
1523:
1522:
1510:
1509:
1500:
1499:
1487:
1486:
1484:NHMSYS0021049569
1474:
1473:
1461:
1460:
1448:
1447:
1435:
1434:
1422:
1421:
1409:
1408:
1396:
1395:
1383:
1382:
1370:
1369:
1357:
1356:
1355:
1342:
1341:
1340:
1310:
1309:
1304:
1303:
1293:
1284:(4): 1131–1154.
1269:
1263:
1262:
1242:
1233:
1224:
1223:
1221:
1219:
1204:
1198:
1197:
1195:
1182:
1173:
1164:
1163:
1161:
1159:
1150:. Archived from
1135:
1129:
1128:
1089:
1083:
1082:
1072:
1044:
1035:
1029:
1028:
998:
989:
988:
975:CSIRO Publishing
962:
951:
950:
929:"Boulder shores"
924:
915:
914:
888:
843:
833:
830:Lobipetrolisthes
823:
813:
810:Beripetrolisthes
799:
763:
753:
743:
733:
723:
713:
703:
693:
683:
673:
670:Parapetrolisthes
663:
653:
643:
633:
623:
613:
603:
593:
583:
573:
563:
560:Heteroporcellana
553:
543:
533:
523:
513:
503:
493:
483:
480:Allopetrolisthes
473:
354:), and the long
305:
304:
301:
298:
237:
145:
144:
126:Allopetrolisthes
115:
105:
42:
41:Tithonian–Recent
38:Temporal range:
32:
31:
21:
1627:
1626:
1622:
1621:
1620:
1618:
1617:
1616:
1592:Porcelain crabs
1582:
1581:
1580:
1575:
1567:
1562:
1554:
1552:
1544:
1539:
1531:
1526:
1518:
1513:
1505:
1503:
1495:
1490:
1482:
1477:
1469:
1464:
1456:
1451:
1443:
1438:
1430:
1425:
1417:
1412:
1404:
1399:
1391:
1386:
1378:
1373:
1365:
1360:
1351:
1350:
1345:
1336:
1335:
1330:
1317:
1307:
1270:
1266:
1240:
1234:
1227:
1217:
1215:
1209:"Porcellanidae"
1205:
1201:
1193:
1180:
1174:
1167:
1157:
1155:
1136:
1132:
1090:
1086:
1042:
1036:
1032:
1025:
999:
992:
985:
963:
954:
947:
925:
918:
911:
889:
882:
878:
840:Longoporcellana
796:Annieporcellana
768:
620:Neopetrolisthes
580:Lissoporcellana
570:Liopetrolisthes
461:
372:
336:
302:
299:
296:
295:
292:
261:Porcelain crabs
236:
139:
130:Liopetrolisthes
106:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
40:
39:
36:
35:Porcelain crabs
28:
23:
22:
15:
12:
11:
5:
1625:
1615:
1614:
1609:
1604:
1599:
1594:
1577:
1576:
1574:
1573:
1560:
1550:
1537:
1524:
1511:
1501:
1488:
1475:
1462:
1449:
1436:
1423:
1410:
1397:
1384:
1371:
1358:
1343:
1327:
1325:
1319:
1318:
1306:
1305:
1264:
1253:(2): 243–248.
1225:
1207:WoRMS (2010).
1199:
1196:on 2012-03-02.
1165:
1130:
1084:
1030:
1023:
1017:. p. 35.
1003:"Anemone crab
990:
983:
952:
945:
916:
909:
879:
877:
874:
847:
846:
836:
826:
820:Eopetrolisthes
816:
806:
767:
766:
764:Glassell, 1938
756:
754:Stimpson, 1858
746:
736:
726:
716:
714:Stimpson, 1858
706:
696:
694:Stimpson, 1858
686:
676:
666:
664:Stimpson, 1858
656:
654:Glassell, 1936
646:
636:
626:
616:
614:Stimpson, 1858
606:
604:Stimpson, 1858
600:Megalobrachium
596:
586:
576:
566:
556:
550:Heteropolyonyx
546:
536:
534:Stimpson, 1860
526:
516:
506:
496:
486:
476:
470:Aliaporcellana
465:
460:
457:
371:
368:
335:
332:
291:
288:
256:
255:
249:
248:
244:
243:
231:
227:
226:
221:
217:
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181:
177:
176:
171:
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157:
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147:
146:
133:
132:
117:
116:
108:
107:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
37:
26:
9:
6:
4:
3:
2:
1624:
1613:
1610:
1608:
1605:
1603:
1600:
1598:
1595:
1593:
1590:
1589:
1587:
1570:
1565:
1561:
1557:
1551:
1547:
1542:
1538:
1534:
1529:
1525:
1521:
1516:
1512:
1508:
1502:
1498:
1493:
1489:
1485:
1480:
1476:
1472:
1467:
1463:
1459:
1454:
1450:
1446:
1441:
1437:
1433:
1428:
1424:
1420:
1415:
1411:
1407:
1402:
1398:
1394:
1389:
1385:
1381:
1380:Porcellanidae
1376:
1372:
1368:
1367:Porcellanidae
1363:
1359:
1354:
1353:Porcellanidae
1348:
1344:
1339:
1333:
1329:
1328:
1326:
1324:
1323:Porcellanidae
1320:
1316:
1311:
1301:
1297:
1292:
1287:
1283:
1279:
1275:
1268:
1260:
1256:
1252:
1248:
1247:
1239:
1232:
1230:
1214:
1210:
1203:
1192:
1188:
1187:
1179:
1172:
1170:
1153:
1149:
1145:
1141:
1134:
1126:
1122:
1118:
1114:
1110:
1106:
1102:
1098:
1097:
1088:
1080:
1076:
1071:
1066:
1062:
1058:
1054:
1050:
1049:
1041:
1034:
1026:
1020:
1016:
1012:
1008:
1006:
997:
995:
986:
980:
976:
972:
968:
961:
959:
957:
948:
942:
938:
934:
930:
923:
921:
912:
906:
902:
898:
894:
887:
885:
880:
873:
871:
870:
865:
862:
858:
854:
853:
842:
841:
837:
832:
831:
827:
822:
821:
817:
812:
811:
807:
803:
798:
797:
793:
792:
791:
789:
785:
781:
777:
773:
772:fossil record
762:
761:
757:
752:
751:
747:
742:
741:
737:
732:
731:
730:Porcellanella
727:
724:Lamarck, 1801
722:
721:
717:
712:
711:
707:
702:
701:
697:
692:
691:
687:
682:
681:
677:
672:
671:
667:
662:
661:
657:
652:
651:
647:
642:
641:
637:
632:
631:
627:
622:
621:
617:
612:
611:
607:
602:
601:
597:
592:
591:
590:Madarateuchus
587:
582:
581:
577:
572:
571:
567:
562:
561:
557:
552:
551:
547:
542:
541:
537:
532:
531:
527:
524:Johnson, 1970
522:
521:
517:
512:
511:
510:Clastotoechus
507:
502:
501:
497:
492:
491:
487:
482:
481:
477:
472:
471:
467:
466:
464:
456:
454:
453:
448:
444:
440:
439:
434:
433:
428:
427:Caribbean Sea
423:
421:
417:
413:
409:
405:
401:
394:
393:
388:
382:
381:
376:
367:
365:
361:
357:
353:
349:
345:
341:
340:carcinisation
331:
329:
325:
321:
317:
313:
309:
308:squat lobster
287:
285:
284:Late Jurassic
281:
277:
273:
272:Porcellanidae
269:
266:
262:
254:
250:
245:
240:
235:
234:Porcellanidae
232:
229:
228:
225:
222:
220:Superfamily:
219:
218:
215:
212:
209:
208:
205:
202:
199:
198:
195:
192:
189:
188:
185:
182:
179:
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172:
169:
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159:
158:
155:
152:
149:
148:
143:
138:
134:
131:
127:
123:
118:
114:
109:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
33:
30:
19:
18:Porcellanidae
1322:
1281:
1277:
1267:
1250:
1244:
1218:November 13,
1216:. Retrieved
1202:
1191:the original
1184:
1158:February 10,
1156:. Retrieved
1152:the original
1143:
1133:
1100:
1094:
1087:
1052:
1046:
1033:
1010:
1004:
970:
932:
896:
869:Vibrissalana
867:
850:
848:
838:
828:
818:
808:
801:
794:
787:
783:
779:
775:
769:
758:
748:
738:
728:
718:
708:
698:
690:Petrolisthes
688:
678:
668:
658:
648:
638:
628:
624:Miyake, 1937
618:
608:
598:
594:Harvey, 1999
588:
578:
568:
558:
548:
538:
528:
518:
508:
498:
490:Ancylocheles
488:
478:
468:
462:
450:
443:hermit crabs
436:
430:
424:
400:Arctic Ocean
397:
390:
378:
348:walking legs
337:
293:
271:
260:
259:
252:
233:
224:Galatheoidea
210:Infraorder:
184:Malacostraca
129:
125:
122:Petrolisthes
121:
29:
1440:iNaturalist
1347:Wikispecies
1103:(1): 1–21.
776:Pachycheles
734:White, 1852
704:Leach, 1820
684:Miers, 1876
680:Petrocheles
660:Pachycheles
644:Osawa, 1998
640:Novorostrum
630:Neopisosoma
554:Osawa, 2001
520:Enosteoides
455:is common.
324:territorial
290:Description
282:age of the
268:crustaceans
204:Pleocyemata
1586:Categories
876:References
861:Ernstbrunn
788:Porcellana
750:Raphidopus
720:Porcellana
674:Haig, 1962
650:Orthochela
634:Haig, 1960
610:Minyocerus
584:Haig, 1978
574:Haig, 1960
564:Haig, 1978
514:Haig, 1960
504:Haig, 1978
494:Haig, 1978
484:Haig, 1960
420:mouthparts
328:pereiopods
200:Suborder:
174:Arthropoda
1300:0024-4082
864:Limestone
857:Tithonian
855:from the
852:Jurellana
540:Eulenaios
530:Euceramus
459:Diversity
447:Hong Kong
404:Antarctic
360:eyestalks
344:true crab
334:Evolution
312:appendage
280:Tithonian
253:See text
160:Kingdom:
154:Eukaryota
1401:BugGuide
1338:Q1435521
1332:Wikidata
1125:26260996
1117:25156549
1079:11886621
800:Fraaije
784:Polyonyx
710:Polyonyx
412:plankton
402:and the
356:antennae
352:carapace
230:Family:
194:Decapoda
170:Phylum:
164:Animalia
150:Domain:
1597:Anomura
1556:7073261
1186:Zootaxa
1070:1690904
893:"Crabs"
780:Pisidia
760:Ulloaia
700:Pisidia
408:beaches
364:abdomen
265:decapod
247:Genera
239:Haworth
214:Anomura
190:Order:
180:Class:
1569:106734
1553:uBio:
1520:211908
1504:NZOR:
1458:110288
1419:1PRCEF
1406:396789
1298:
1123:
1115:
1077:
1067:
1021:
981:
943:
907:
804:, 2008
802:et al.
362:. The
320:chelae
316:moults
241:, 1825
1564:WoRMS
1541:Plazi
1533:89027
1497:84660
1471:98058
1453:IRMNG
1445:47805
1241:(PDF)
1194:(PDF)
1181:(PDF)
1121:S2CID
1043:(PDF)
859:aged
445:. In
416:setae
276:crabs
1492:NCBI
1466:ITIS
1432:8673
1427:GBIF
1414:EPPO
1393:1509
1388:BOLD
1296:ISSN
1220:2010
1160:2005
1113:PMID
1075:PMID
1019:ISBN
979:ISBN
941:ISBN
905:ISBN
770:The
429:are
263:are
128:and
46:PreꞒ
1479:NBN
1375:AFD
1362:ADW
1286:doi
1282:187
1255:doi
1251:258
1105:doi
1101:276
1065:PMC
1057:doi
1053:269
1588::
1566::
1543::
1530::
1517::
1494::
1481::
1468::
1455::
1442::
1429::
1416::
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1390::
1377::
1364::
1349::
1334::
1294:.
1280:.
1276:.
1249:.
1243:.
1228:^
1211:.
1183:.
1168:^
1146:.
1142:.
1119:.
1111:.
1099:.
1073:.
1063:.
1051:.
1045:.
1009:.
993:^
973:.
969:.
955:^
935:.
931:.
919:^
895:.
883:^
786:,
782:,
778:,
449:,
422:.
303:16
124:,
96:Pg
1302:.
1288::
1261:.
1257::
1222:.
1162:.
1127:.
1107::
1081:.
1059::
1027:.
1007:"
987:.
949:.
913:.
300:/
297:9
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:Ꞓ
20:)
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