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Partula (gastropod)

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Individuals are being reintroduced to Tahiti from captive breeding programmes since 2014. In April 2023, over 5,000 individual snails from zoos in the United States and the United Kingdom were released on Tahiti and
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that were endemic to Tahiti and the nearby islands, causing all but 12 species to go extinct within a decade. Scientists were able to save 12 of these species prior to their becoming
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ceremonial wear and jewelry, these small snails (averaging about one-half to three-quarters of an inch in length) gained the attention of science when Dr.
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act as an example of the possible deleterious effects of attempted biological control. After an infestation of the introduced giant African land snails (
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Myers, P.; Espinosa, R.; Parr, C. S.; Jones, T.; Hammond, G. S. & Dewey, T. A. (2006). The Animal Diversity Web (online). Accessed at
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runs the Partula Programme Consortium which maintains a captive-breeding programme in the United Kingdom, France, and the United States.
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Studies on the variation, distribution and evolution of the genus Partula. The species of the Mariana Islands, Guam and Saipan.
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Gerlach, J. (2016) Icons of Evolution - Pacific island tree snails, family Partulidae. Phelsuma Press, Cambridge
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Studies on the variation, distribution and evolution of the genus Partula. The species inhabiting Moorea.
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Studies on the variation, distribution and evolution of the genus Partula. The species inhabiting Tahiti.
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The list may be currently missing some species. If you notice a missing species please add it.
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Proc. Joint Conference - American Malacological Society and Western Society of Malacologists
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Type population is almost certainly extinct. Remaining populations have not been surveyed.
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Currently only known from a dead subadult in one location and a live one in another.
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Lee, T.; Burch, J. B.; Jung, Y.; Coote, T.; Pearce-Kelly, P.; ó Foighil, D. (2007).
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Survives in populations above 950m altitude and in one population on Tahiti Iti.
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The species pages are currently outdated because of the IUCN 2024-1 update.
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Only known from a single location. Not seen since its discovery in 2005.
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species on Tahiti were usually found on the undersides of the leaves of
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This article incorporates public domain text from the reference.
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Could not be located after 1991. Old shells were found in 1994.
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were misidentified as the species in parentheses next to them.
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Many populations are unstable and inviable in the long-term.
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2005, 2006 and 2008 surveys found only 4 old, eroded shells.
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Little is known about this species. May already be extinct.
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Instead, the rosy wolfsnail hunted the nearly 76 species of
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Single valley endemic. A single shell was found in 1992.
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Tolerant of current levels of disturbance and predators.
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Has become much rarer but several populations persist.
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Only one population remains. Viable but vulnerable.
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Only dead shells could be found in the 1991 survey.
67:. Unsourced material may be challenged and removed. 1549:All populations appear to be tolerant of threats. 5759: 975:, though currently relatively free of predators. 811:As of the 2024-1 update released on June 27, the 5997: 5747:Carnegie Institution of Washington, 228a: 1-116. 3063:Phylogenetic analyses revealed that many of the 2448:Tolerant of habitat loss and free of predators. 2107:. Population declined severely in both islands. 5756:Carnegie Institution of Washington, 410: 1-335. 1047:The last captive individual died in the 1980s. 784:spp.), the carnivorous Florida rosy wolfsnail ( 5801:Partula evolution, diversity and conservation 5481: 2821:The list of surviving species are as follows: 1788:Survival confirmed from a single individual. 5707:Memoirs of the Museum of Comparative Zoölogy 2006:No major threats to this species are known. 653:snails on the underside of a single leaf of 645:Black-and-white photo taken in July 1920 in 927:but reasonable numbers exist in captivity. 5631:Elizabeth Claire Alberts (28 April 2023). 5380: 3071:. The resulting cladogram is shown below. 1688:The last captive individual died in 2002. 1442:The last captive individual died in 2016. 950:The last captive individual died in 1994. 723:are known under the general common names " 151: 5777: 2149:Endemic to nine islands. Eliminated from 1247:Could not be located in the 1992 survey. 762:, and behavioral aspects that illustrate 127:Learn how and when to remove this message 5694: 5692: 2794: 1922:. Could not be located in 2006 or 2017. 673: 661: 640: 5544: 5542: 14: 5998: 2550:Last captive individual died in 1992. 1897:. Little is known about this species. 1709:One population may survive in central 1131:Last captive individual died in 1996. 419:; Leptopartula is a replacement name) 5812: 5811: 5689: 5585:"IUCN Red List of Threatened Species" 5501:ITIS Standard Report Page: Partulidae 1152:One population photographed in 2004. 1018:Extirpated from its type locality in 5705:from Tahiti. A study in variation". 5605: 5579: 5577: 5539: 2816: 2325:Little is known about this species. 819:species. Of these, 32 are listed as 65:adding citations to reliable sources 36: 5494: 5325: 5096: 1943:Only known from a single specimen. 1872:Could not be located in the 1990s. 813:IUCN Red List of Threatened Species 535:accepted, alternate representation 24: 5736:Carnegie Institution of Washington 5722: 5561:"Partula snail conservation | ZSL" 5286: 5199: 2621:Collected for ex situ conservation 25: 6032: 5795: 5701:(January 1902). "Some species of 5574: 2571:Could not be located after 1980. 2529:Could not be located after 1990. 2406:Could not be located after 1990. 2346:Could not be located after 1980. 2241:Could not be located after 1980. 2220:Could not be located after 1980. 1830:Could not be located after 1980. 1809:Could not be located after 1980. 1636:All individuals are hybrids with 1421:Could not be located after 1980. 1379:Could not be located after 1992. 1358:Could not be located after 1980. 1337:Could not be located after 1980. 1089:Could not be located after 1993. 746:Once used as decorative items in 5435: 5419: 5403: 5387: 5364: 5348: 5332: 5309: 5293: 5270: 5254: 5238: 5222: 5206: 5183: 5167: 5151: 5135: 5119: 5103: 2635:breeding include the following: 2427:Dead shells were found in 1989. 923:A single population persists on 173: 41: 5671: 5650: 5624: 774:The partulids of the island of 52:needs additional citations for 5599: 5553: 5518: 5505: 5462: 2199:Could not be located in 1994. 13: 1: 6021:Molluscs of the Pacific Ocean 5524:Jung, Younghun, Taehwan Lee, 5452: 2802:snail with its shell removed. 729:Moorean viviparous tree snail 3058: 1179:. Old shells found in 2005. 866: 806:Zoological Society of London 583:(invalid: junior homonym of 544:(invalid: junior homonym of 519:(invalid: junior homonym of 503:(invalid: junior homonym of 447:(invalid: junior homonym of 428:(invalid: junior homonym of 412:(invalid: junior homonym of 396:(invalid: junior homonym of 372:(invalid: junior homonym of 7: 10: 6037: 5606:Kuta, Sarah (2 May 2023). 5489:http://animaldiversity.org 5091: 5060: 854: 769: 695:of air-breathing tropical 29: 5820: 5779:10.1016/j.cub.2007.05.006 5015: 5008: 4991: 4984: 4926: 4909: 4902: 4880: 4873: 4856: 4839: 4804: 4787: 4780: 4763: 4756: 4749: 4718: 4701: 4684: 4658: 4613: 4595: 4588: 4552: 4545: 4528: 4521: 4514: 4460: 4453: 4436: 4429: 4412: 4405: 4388: 4371: 4364: 4357: 4350: 4324: 4307: 4290: 4273: 4247: 4230: 4223: 4179: 4172: 4155: 4148: 4104: 4097: 4080: 4063: 4056: 4039: 4032: 4025: 3999: 3981: 3964: 3946: 3928: 3884: 3866: 3859: 3832: 3814: 3796: 3789: 3772: 3765: 3748: 3741: 3715: 3698: 3672: 3655: 3648: 3622: 3605: 3588: 3571: 3554: 3537: 3520: 3513: 3506: 3462: 3455: 3438: 3403: 3396: 3379: 3372: 3365: 3339: 3322: 3305: 3288: 3281: 3246: 3239: 3222: 3196: 3179: 3162: 3145: 3138: 3131: 3124: 3117: 3110: 3103: 3091: 3084: 859:Species within the genus 288: 281: 170:Scientific classification 168: 159: 150: 143: 3935:(Austral Islands, etc.) 3776:P. otaheitana otaheitana 2629:that were collected for 2490:3/4 subspecies survive. 2469:2/2 subspecies survive. 2385:Not located after 1936. 76:"Partula" gastropod 32:Partula (disambiguation) 5750:Crampton H. E. (1932). 5741:Crampton H. E. (1925). 5381:Mariana Islands species 3166:P. suturalis dendronica 1268:Isolated population on 839:and only 4 species are 508:H. & A. Adams, 1855 438:is a replacement name) 5469:Férussac A. É. d'A. de 2803: 1918:Only species found in 683: 671: 659: 458:Partula (Leptopartula) 5965:Paleobiology Database 4392:P. otaheitana affinis 3326:P. suturalis vexillum 3250:P. suturalis vexillum 3226:P. suturalis strigosa 3200:P. suturalis vexillum 3183:P. suturalis strigosa 2798: 1767:Not located in 1992. 1204:, though survives in 829:critically endangered 725:Polynesian tree snail 677: 665: 644: 383:Partula (Carolinella) 5665:University of Hawaii 5612:Smithsonian Magazine 4930:Palaopartula calypso 3676:P. taeniata nucleola 3609:P. taeniata simulans 3592:P. taeniata nucleola 3575:P. taeniata elongata 2578:Partula vanikorensis 2164:Partula pearcekellyi 1649:Partula jackieburchi 1344:Partula dolichostoma 1215:Partula crassilabris 1022:but persists in the 594:Partula (Thakombaua) 490:Partula (Melanesica) 466:Partula (Marianella) 61:improve this article 30:For other uses, see 6016:Molluscs of Oceania 6006:Partula (gastropod) 5715:, Cambridge, U.S.A. 5373:Partula navigatoria 4913:Palaopartula thetis 4767:Samoana decussatula 4532:undescribed species 3752:undescribed species 2185:Partula planilabrum 2049:Partula navigatoria 1816:Partula levistriata 1272:confirmed in 2005. 825:extinct in the wild 649:, showing numerous 569:Partula (Rennellia) 27:Genus of gastropods 5473:Journ. de Physique 5192:Partula otaheitana 2804: 2557:Partula umbilicata 2311:Partula recluziana 2114:Partula otaheitana 1795:Partula leptochila 1720:Partula lanceolata 1054:Partula auriculata 764:adaptive radiation 735:islands, from the 684: 672: 660: 539:Partula (Pasithea) 533:A. Férussac, 1821· 474:Partula (Marianna) 423:Partula (Harmonia) 303:Bulimus (Partula) 5993: 5992: 5952:Open Tree of Life 5814:Taxon identifiers 5772:(13): R502–R503. 5444:Partula langfordi 5428:Partula radiolata 5412:Partula lutaensis 5341:Partula garrettii 5279:Partula tohiveana 5247:Partula suturalis 5215:Partula mirabilis 5057: 5056: 5048: 5047: 5039: 5038: 5030: 5029: 4968: 4967: 4959: 4958: 4950: 4949: 4941: 4940: 4860:Samoana thurstoni 4828: 4827: 4819: 4818: 4791:Samoana attenuata 4733: 4732: 4673: 4672: 4647: 4646: 4638: 4637: 4629: 4628: 4577: 4576: 4568: 4567: 4503: 4502: 4494: 4493: 4485: 4484: 4476: 4475: 4339: 4338: 4262: 4261: 4212: 4211: 4203: 4202: 4194: 4193: 4137: 4136: 4128: 4127: 4119: 4118: 4014: 4013: 3917: 3916: 3908: 3907: 3899: 3898: 3848: 3847: 3730: 3729: 3687: 3686: 3637: 3636: 3495: 3494: 3486: 3485: 3477: 3476: 3427: 3426: 3418: 3417: 3354: 3353: 3270: 3269: 3261: 3260: 3211: 3210: 2817:Surviving species 2618: 2617: 2497:Partula tohiveana 2455:Partula suturalis 2434:Partula similaris 2269:Partula radiolata 2227:Partula protracta 2013:Partula mirabilis 1879:Partula lutaensis 1753:Partula langfordi 1695:Partula laevigata 1611:Wild populations 1556:Partula guamensis 1474:Partula garrettii 1279:Partula dentifera 1186:Partula cramptoni 1138:Partula compressa 1075:Partula bilineata 1002:Partula auraniana 957:Partula assimilis 760:ecological niches 680:Partula langfordi 639: 638: 632: 624: 616:(junior synonym) 615: 607:(junior synonym) 606: 598: 589: 582: 578:Partula (Sterope) 574:(junior synonym) 573: 564: 557: 550: 543: 534: 530:Partula (Partula) 525: 518: 509: 502: 494: 486: 478: 470: 462: 453: 446: 434: 427: 418: 411: 402: 395: 387: 378: 371: 367:Partula (Astraea) 363: 355: 347: 339: 331: 323: 315: 309: 300: 292:list of synonyms: 277: 162:Partula radiolata 137: 136: 129: 111: 16:(Redirected from 6028: 6011:Gastropod genera 5986: 5985: 5973: 5972: 5960: 5959: 5947: 5946: 5934: 5933: 5921: 5920: 5908: 5907: 5895: 5894: 5882: 5881: 5869: 5868: 5856: 5855: 5854: 5841: 5840: 5839: 5809: 5808: 5791: 5781: 5716: 5696: 5687: 5686: 5675: 5669: 5668: 5662: 5654: 5648: 5647: 5645: 5643: 5628: 5622: 5621: 5619: 5618: 5603: 5597: 5596: 5594: 5592: 5581: 5572: 5571: 5569: 5567: 5557: 5551: 5546: 5537: 5522: 5516: 5509: 5503: 5498: 5492: 5485: 5479: 5466: 5439: 5423: 5407: 5391: 5368: 5352: 5336: 5326:Raiatean species 5313: 5297: 5274: 5263:Partula taeniata 5258: 5242: 5231:Partula mooreana 5226: 5210: 5187: 5171: 5160:Partula incrassa 5155: 5139: 5123: 5107: 5097:Tahitian species 5011: 5010: 4987: 4986: 4905: 4904: 4884:Samoana fragilis 4876: 4875: 4783: 4782: 4759: 4758: 4752: 4751: 4591: 4590: 4548: 4547: 4524: 4523: 4517: 4516: 4456: 4455: 4432: 4431: 4408: 4407: 4367: 4366: 4360: 4359: 4353: 4352: 4226: 4225: 4175: 4174: 4151: 4150: 4100: 4099: 4059: 4058: 4035: 4034: 4028: 4027: 3862: 3861: 3792: 3791: 3768: 3767: 3744: 3743: 3651: 3650: 3516: 3515: 3509: 3508: 3458: 3457: 3399: 3398: 3375: 3374: 3368: 3367: 3284: 3283: 3242: 3241: 3141: 3140: 3134: 3133: 3127: 3126: 3120: 3119: 3113: 3112: 3106: 3105: 3094: 3093: 3087: 3086: 3075: 3074: 3067:species are not 3031:Papua New Guinea 2476:Partula taeniata 2413:Partula salifana 2206:Partula producta 2135:Partula pacifica 2031:Partula mooreana 1964:Fossil species. 1929:Partula magistri 1858:Partula lugubris 1674:Partula labrusca 1622:Partula incrassa 1449:Partula flexuosa 1386:Partula emersoni 1365:Partula dolorosa 1323:Partula diminuta 1316:Fossil species. 1300:Partula desolata 1254:Partula cytherea 1200:Extirpated from 1033:Partula aurantia 879: 878: 787:Euglandina rosea 719:Many species of 630: 622: 613: 604: 596: 587: 580: 571: 562: 555: 548: 541: 532: 523: 516: 514:Partula (Oenone) 507: 500: 492: 484: 482:Partula (Matata) 476: 468: 460: 451: 444: 442:Partula (Helena) 432: 425: 416: 409: 400: 393: 391:Partula (Clytia) 385: 376: 369: 361: 353: 345: 337: 329: 321: 313: 307: 298: 272: 178: 177: 155: 141: 140: 132: 125: 121: 118: 112: 110: 69: 45: 37: 21: 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4662:P. subgonochila 4648: 4639: 4630: 4578: 4569: 4504: 4495: 4486: 4477: 4340: 4311:P. carteriensis 4263: 4213: 4204: 4195: 4138: 4129: 4120: 4015: 3918: 3909: 3900: 3849: 3731: 3688: 3638: 3496: 3487: 3478: 3428: 3419: 3355: 3271: 3262: 3212: 3061: 3014:Solomon Islands 2819: 2623: 2536:Partula turgida 2515:Partula tristis 2392:Partula sagitta 2290:Partula radiosa 2248:Partula pyramis 1950:Partula makatea 1663:May survive on 1615:be recovering. 1595:Partula hyalina 1407:Partula eremita 1233:Partula cuneata 1176:Partula hyalina 1173:Outcompeted by 1117:Partula clarkei 907:Partula affinis 869: 857: 772: 737:Society Islands 635: 498:Partula (Nenia) 401:Lamouroux, 1812 294: 293: 271: 227:Stylommatophora 172: 133: 122: 116: 113: 70: 68: 58: 46: 35: 28: 23: 22: 18:Partula (genus) 15: 12: 11: 5: 6034: 6024: 6023: 6018: 6013: 6008: 5991: 5990: 5988: 5987: 5974: 5961: 5948: 5935: 5922: 5909: 5896: 5883: 5870: 5857: 5842: 5826: 5824: 5818: 5817: 5806: 5805: 5797: 5796:External links 5794: 5793: 5792: 5757: 5748: 5739: 5729:Crampton H. E. 5724: 5721: 5718: 5717: 5688: 5670: 5649: 5623: 5598: 5587:. 27 June 2024 5573: 5552: 5538: 5517: 5504: 5493: 5480: 5460: 5459: 5454: 5451: 5450: 5449: 5441: 5434: 5432: 5425: 5418: 5416: 5409: 5402: 5400: 5393: 5386: 5382: 5379: 5378: 5377: 5370: 5363: 5361: 5354: 5347: 5345: 5338: 5331: 5327: 5324: 5323: 5322: 5315: 5308: 5306: 5299: 5292: 5288: 5285: 5284: 5283: 5276: 5269: 5267: 5260: 5253: 5251: 5244: 5237: 5235: 5228: 5221: 5219: 5212: 5205: 5201: 5198: 5197: 5196: 5189: 5182: 5180: 5176:Partula nodosa 5173: 5166: 5164: 5157: 5150: 5148: 5141: 5134: 5132: 5125: 5118: 5116: 5109: 5102: 5098: 5095: 5093: 5090: 5062: 5059: 5055: 5054: 5051: 5050: 5046: 5045: 5042: 5041: 5037: 5036: 5033: 5032: 5028: 5027: 5024: 5023: 5014: 5009: 5007: 5004: 5003: 5000: 4999: 4990: 4985: 4983: 4975: 4974: 4971: 4970: 4966: 4965: 4962: 4961: 4957: 4956: 4953: 4952: 4948: 4947: 4944: 4943: 4939: 4938: 4935: 4934: 4925: 4922: 4921: 4918: 4917: 4908: 4903: 4901: 4893: 4892: 4889: 4888: 4879: 4874: 4872: 4869: 4868: 4865: 4864: 4855: 4852: 4851: 4848: 4847: 4843:Samoana conica 4838: 4835: 4834: 4831: 4830: 4826: 4825: 4822: 4821: 4817: 4816: 4813: 4812: 4808:Samoana burchi 4803: 4800: 4799: 4796: 4795: 4786: 4781: 4779: 4776: 4775: 4772: 4771: 4762: 4757: 4755: 4750: 4748: 4740: 4739: 4736: 4735: 4731: 4730: 4727: 4726: 4717: 4714: 4713: 4710: 4709: 4700: 4697: 4696: 4693: 4692: 4683: 4680: 4679: 4676: 4675: 4671: 4670: 4667: 4666: 4657: 4654: 4653: 4650: 4649: 4645: 4644: 4641: 4640: 4636: 4635: 4632: 4631: 4627: 4626: 4623: 4622: 4612: 4609: 4608: 4605: 4604: 4594: 4589: 4587: 4584: 4583: 4580: 4579: 4575: 4574: 4571: 4570: 4566: 4565: 4562: 4561: 4551: 4546: 4544: 4541: 4540: 4537: 4536: 4527: 4522: 4520: 4515: 4513: 4510: 4509: 4506: 4505: 4501: 4500: 4497: 4496: 4492: 4491: 4488: 4487: 4483: 4482: 4479: 4478: 4474: 4473: 4470: 4469: 4459: 4454: 4452: 4449: 4448: 4445: 4444: 4435: 4430: 4428: 4425: 4424: 4421: 4420: 4411: 4406: 4404: 4401: 4400: 4397: 4396: 4387: 4384: 4383: 4380: 4379: 4370: 4365: 4363: 4358: 4356: 4351: 4349: 4346: 4345: 4342: 4341: 4337: 4336: 4333: 4332: 4323: 4320: 4319: 4316: 4315: 4306: 4303: 4302: 4299: 4298: 4289: 4286: 4285: 4282: 4281: 4272: 4269: 4268: 4265: 4264: 4260: 4259: 4256: 4255: 4246: 4243: 4242: 4239: 4238: 4229: 4224: 4222: 4219: 4218: 4215: 4214: 4210: 4209: 4206: 4205: 4201: 4200: 4197: 4196: 4192: 4191: 4188: 4187: 4178: 4173: 4171: 4168: 4167: 4164: 4163: 4154: 4149: 4147: 4144: 4143: 4140: 4139: 4135: 4134: 4131: 4130: 4126: 4125: 4122: 4121: 4117: 4116: 4113: 4112: 4103: 4098: 4096: 4093: 4092: 4089: 4088: 4079: 4076: 4075: 4072: 4071: 4062: 4057: 4055: 4052: 4051: 4048: 4047: 4038: 4033: 4031: 4026: 4024: 4021: 4020: 4017: 4016: 4012: 4011: 4008: 4007: 3998: 3995: 3994: 3991: 3990: 3980: 3977: 3976: 3973: 3972: 3963: 3960: 3959: 3956: 3955: 3945: 3942: 3941: 3938: 3937: 3927: 3924: 3923: 3920: 3919: 3915: 3914: 3911: 3910: 3906: 3905: 3902: 3901: 3897: 3896: 3893: 3892: 3883: 3880: 3879: 3876: 3875: 3865: 3860: 3858: 3855: 3854: 3851: 3850: 3846: 3845: 3842: 3841: 3831: 3828: 3827: 3824: 3823: 3813: 3810: 3809: 3806: 3805: 3795: 3790: 3788: 3785: 3784: 3781: 3780: 3771: 3766: 3764: 3761: 3760: 3757: 3756: 3747: 3742: 3740: 3737: 3736: 3733: 3732: 3728: 3727: 3724: 3723: 3714: 3711: 3710: 3707: 3706: 3697: 3694: 3693: 3690: 3689: 3685: 3684: 3681: 3680: 3671: 3668: 3667: 3664: 3663: 3654: 3649: 3647: 3644: 3643: 3640: 3639: 3635: 3634: 3631: 3630: 3621: 3618: 3617: 3614: 3613: 3604: 3601: 3600: 3597: 3596: 3587: 3584: 3583: 3580: 3579: 3570: 3567: 3566: 3563: 3562: 3553: 3550: 3549: 3546: 3545: 3536: 3533: 3532: 3529: 3528: 3519: 3514: 3512: 3507: 3505: 3502: 3501: 3498: 3497: 3493: 3492: 3489: 3488: 3484: 3483: 3480: 3479: 3475: 3474: 3471: 3470: 3461: 3456: 3454: 3451: 3450: 3447: 3446: 3437: 3434: 3433: 3430: 3429: 3425: 3424: 3421: 3420: 3416: 3415: 3412: 3411: 3402: 3397: 3395: 3392: 3391: 3388: 3387: 3378: 3373: 3371: 3366: 3364: 3361: 3360: 3357: 3356: 3352: 3351: 3348: 3347: 3338: 3335: 3334: 3331: 3330: 3321: 3318: 3317: 3314: 3313: 3304: 3301: 3300: 3297: 3296: 3287: 3282: 3280: 3277: 3276: 3273: 3272: 3268: 3267: 3264: 3263: 3259: 3258: 3255: 3254: 3245: 3240: 3238: 3235: 3234: 3231: 3230: 3221: 3218: 3217: 3214: 3213: 3209: 3208: 3205: 3204: 3195: 3192: 3191: 3188: 3187: 3178: 3175: 3174: 3171: 3170: 3161: 3158: 3157: 3154: 3153: 3144: 3139: 3137: 3132: 3130: 3125: 3123: 3118: 3116: 3111: 3109: 3104: 3102: 3092: 3090: 3085: 3083: 3073: 3060: 3057: 2940:P. navigatoria 2818: 2815: 2811:P. navigatoria 2763:P. navigatoria 2622: 2619: 2616: 2615: 2612: 2609: 2606: 2598: 2597: 2590: 2587: 2584: 2581: 2573: 2572: 2569: 2566: 2563: 2560: 2552: 2551: 2548: 2545: 2542: 2539: 2531: 2530: 2527: 2526:Misidentified 2524: 2521: 2518: 2510: 2509: 2506: 2503: 2500: 2492: 2491: 2488: 2485: 2482: 2479: 2471: 2470: 2467: 2464: 2461: 2458: 2450: 2449: 2446: 2443: 2440: 2437: 2429: 2428: 2425: 2422: 2419: 2416: 2408: 2407: 2404: 2401: 2398: 2395: 2387: 2386: 2383: 2380: 2377: 2374: 2366: 2365: 2362: 2359: 2356: 2348: 2347: 2344: 2341: 2338: 2335: 2332:Partula remota 2327: 2326: 2323: 2320: 2317: 2314: 2306: 2305: 2302: 2299: 2296: 2293: 2285: 2284: 2281: 2278: 2275: 2272: 2264: 2263: 2260: 2257: 2254: 2251: 2243: 2242: 2239: 2236: 2233: 2230: 2222: 2221: 2218: 2215: 2212: 2209: 2201: 2200: 2197: 2194: 2191: 2188: 2180: 2179: 2176: 2173: 2170: 2167: 2159: 2158: 2151:Espiritu Santo 2147: 2144: 2141: 2138: 2130: 2129: 2126: 2125:Misidentified 2123: 2120: 2117: 2109: 2108: 2097: 2094: 2091: 2088: 2080: 2079: 2076: 2073: 2070: 2067:Partula nodosa 2062: 2061: 2058: 2055: 2052: 2044: 2043: 2040: 2037: 2034: 2026: 2025: 2022: 2019: 2016: 2008: 2007: 2004: 2001: 1998: 1995: 1992:Partula micans 1987: 1986: 1983: 1980: 1977: 1974: 1971:Partula meyeri 1966: 1965: 1962: 1959: 1956: 1953: 1945: 1944: 1941: 1938: 1935: 1932: 1924: 1923: 1916: 1913: 1910: 1907: 1899: 1898: 1891: 1888: 1885: 1882: 1874: 1873: 1870: 1867: 1864: 1861: 1853: 1852: 1849: 1846: 1843: 1840: 1837:Partula lirata 1832: 1831: 1828: 1825: 1822: 1819: 1811: 1810: 1807: 1804: 1801: 1798: 1790: 1789: 1786: 1783: 1780: 1777: 1774:Partula leefei 1769: 1768: 1765: 1762: 1759: 1756: 1748: 1747: 1732: 1729: 1726: 1723: 1715: 1714: 1707: 1704: 1701: 1698: 1690: 1689: 1686: 1683: 1680: 1677: 1669: 1668: 1661: 1658: 1655: 1652: 1644: 1643: 1634: 1631: 1628: 1625: 1617: 1616: 1609: 1606: 1603: 1598: 1590: 1589: 1586: 1583: 1580: 1572: 1571: 1568: 1565: 1562: 1559: 1551: 1550: 1547: 1544: 1541: 1538: 1535:Partula grisea 1530: 1529: 1506: 1503: 1500: 1497: 1489: 1488: 1485: 1482: 1477: 1469: 1468: 1465: 1462: 1457: 1452: 1444: 1443: 1440: 1437: 1434: 1431: 1423: 1422: 1419: 1416: 1413: 1410: 1402: 1401: 1398: 1395: 1392: 1389: 1381: 1380: 1377: 1374: 1371: 1368: 1360: 1359: 1356: 1353: 1350: 1347: 1339: 1338: 1335: 1332: 1329: 1326: 1318: 1317: 1314: 1311: 1308: 1303: 1295: 1294: 1291: 1290:Misidentified 1288: 1285: 1282: 1274: 1273: 1266: 1263: 1260: 1257: 1249: 1248: 1245: 1242: 1239: 1236: 1228: 1227: 1224: 1221: 1218: 1210: 1209: 1206:Bellona Island 1202:Rennell Island 1198: 1195: 1192: 1189: 1181: 1180: 1171: 1168: 1165: 1162: 1159:Partula cootei 1154: 1153: 1150: 1147: 1144: 1141: 1133: 1132: 1129: 1126: 1123: 1120: 1112: 1111: 1108: 1105: 1102: 1099: 1091: 1090: 1087: 1084: 1081: 1078: 1070: 1069: 1066: 1063: 1060: 1057: 1049: 1048: 1045: 1042: 1039: 1036: 1028: 1027: 1024:Torres Islands 1016: 1013: 1010: 1005: 997: 996: 993: 990: 985: 982:Partula atilis 977: 976: 971:Threatened by 969: 966: 963: 960: 952: 951: 948: 945: 942: 937: 934:Partula arguta 929: 928: 921: 918: 915: 910: 902: 901: 898: 893: 888: 883: 877: 876: 873: 868: 865: 856: 853: 771: 768: 752:Henry Crampton 712:in the family 637: 636: 634: 633: 625: 617: 608: 599: 591: 575: 566: 536: 527: 511: 495: 487: 479: 471: 463: 455: 439: 420: 407:Partula (Echo) 404: 388: 380: 364: 356: 348: 340: 332: 324: 316: 310: 304: 301: 291: 290: 289: 286: 285: 279: 278: 264: 260: 259: 254: 250: 249: 244: 240: 239: 234: 230: 229: 224: 220: 219: 214: 210: 209: 204: 200: 199: 194: 190: 189: 184: 180: 179: 166: 165: 157: 156: 148: 147: 135: 134: 49: 47: 40: 26: 9: 6: 4: 3: 2: 6033: 6022: 6019: 6017: 6014: 6012: 6009: 6007: 6004: 6003: 6001: 5984: 5979: 5975: 5971: 5966: 5962: 5958: 5953: 5949: 5945: 5940: 5936: 5932: 5927: 5923: 5919: 5914: 5910: 5906: 5901: 5897: 5893: 5888: 5884: 5880: 5875: 5871: 5867: 5862: 5858: 5853: 5847: 5843: 5838: 5832: 5828: 5827: 5825: 5823: 5819: 5815: 5810: 5804: 5803:Partula Pages 5800: 5799: 5789: 5785: 5780: 5775: 5771: 5767: 5763: 5758: 5755: 5754: 5749: 5746: 5745: 5740: 5738:, 228: 1-311. 5737: 5734: 5730: 5727: 5726: 5714: 5712: 5708: 5704: 5700: 5695: 5693: 5684: 5680: 5674: 5666: 5659: 5653: 5638: 5634: 5627: 5613: 5609: 5602: 5586: 5580: 5578: 5562: 5556: 5550: 5545: 5543: 5535: 5531: 5527: 5521: 5514: 5508: 5502: 5497: 5490: 5484: 5477: 5474: 5471:(June 1821). 5470: 5465: 5461: 5458: 5446: 5445: 5438: 5433: 5430: 5429: 5422: 5417: 5414: 5413: 5406: 5401: 5398: 5397: 5396:Partula gibba 5390: 5385: 5384: 5375: 5374: 5367: 5362: 5359: 5358: 5351: 5346: 5343: 5342: 5335: 5330: 5329: 5320: 5319: 5318:Partula varia 5312: 5307: 5304: 5303: 5302:Partula rosea 5296: 5291: 5290: 5281: 5280: 5273: 5268: 5265: 5264: 5257: 5252: 5249: 5248: 5241: 5236: 5233: 5232: 5225: 5220: 5217: 5216: 5209: 5204: 5203: 5194: 5193: 5186: 5181: 5178: 5177: 5170: 5165: 5162: 5161: 5154: 5149: 5146: 5145: 5138: 5133: 5130: 5129: 5128:Partula clara 5122: 5117: 5114: 5113: 5106: 5101: 5100: 5089: 5087: 5083: 5082: 5077: 5073: 5072: 5067: 5053: 5052: 5044: 5043: 5035: 5034: 5026: 5025: 5022: 5021: 5020: 5013: 5012: 5006: 5005: 5002: 5001: 4998: 4997: 4996: 4989: 4988: 4982: 4981: 4977: 4976: 4973: 4972: 4964: 4963: 4955: 4954: 4946: 4945: 4937: 4936: 4933: 4932: 4931: 4924: 4923: 4920: 4919: 4916: 4915: 4914: 4907: 4906: 4900: 4899: 4895: 4894: 4891: 4890: 4887: 4886: 4885: 4878: 4877: 4871: 4870: 4867: 4866: 4863: 4862: 4861: 4854: 4853: 4850: 4849: 4846: 4845: 4844: 4837: 4836: 4833: 4832: 4824: 4823: 4815: 4814: 4811: 4810: 4809: 4802: 4801: 4798: 4797: 4794: 4793: 4792: 4785: 4784: 4778: 4777: 4774: 4773: 4770: 4769: 4768: 4761: 4760: 4754: 4753: 4747: 4746: 4742: 4741: 4738: 4737: 4729: 4728: 4725: 4724: 4723: 4716: 4715: 4712: 4711: 4708: 4707: 4706: 4699: 4698: 4695: 4694: 4691: 4690: 4689: 4682: 4681: 4678: 4677: 4669: 4668: 4665: 4664: 4663: 4656: 4655: 4652: 4651: 4643: 4642: 4634: 4633: 4625: 4624: 4621: 4620: 4618: 4611: 4610: 4607: 4606: 4603: 4602: 4600: 4593: 4592: 4586: 4585: 4582: 4581: 4573: 4572: 4564: 4563: 4560: 4559: 4557: 4556:P. otaheitana 4550: 4549: 4543: 4542: 4539: 4538: 4535: 4533: 4526: 4525: 4519: 4518: 4512: 4511: 4508: 4507: 4499: 4498: 4490: 4489: 4481: 4480: 4472: 4471: 4468: 4467: 4465: 4464:P. otaheitana 4458: 4457: 4451: 4450: 4447: 4446: 4443: 4442: 4441: 4440:P. otaheitana 4434: 4433: 4427: 4426: 4423: 4422: 4419: 4418: 4417: 4410: 4409: 4403: 4402: 4399: 4398: 4395: 4394: 4393: 4386: 4385: 4382: 4381: 4378: 4377: 4376: 4375:P. otaheitana 4369: 4368: 4362: 4361: 4355: 4354: 4348: 4347: 4344: 4343: 4335: 4334: 4331: 4330: 4329: 4322: 4321: 4318: 4317: 4314: 4313: 4312: 4305: 4304: 4301: 4300: 4297: 4296: 4295: 4288: 4287: 4284: 4283: 4280: 4279: 4278: 4271: 4270: 4267: 4266: 4258: 4257: 4254: 4253: 4252: 4245: 4244: 4241: 4240: 4237: 4236: 4235: 4228: 4227: 4221: 4220: 4217: 4216: 4208: 4207: 4199: 4198: 4190: 4189: 4186: 4185: 4184: 4177: 4176: 4170: 4169: 4166: 4165: 4162: 4161: 4160: 4153: 4152: 4146: 4145: 4142: 4141: 4133: 4132: 4124: 4123: 4115: 4114: 4111: 4110: 4109: 4102: 4101: 4095: 4094: 4091: 4090: 4087: 4086: 4085: 4078: 4077: 4074: 4073: 4070: 4069: 4068: 4061: 4060: 4054: 4053: 4050: 4049: 4046: 4045: 4044: 4037: 4036: 4030: 4029: 4023: 4022: 4019: 4018: 4010: 4009: 4006: 4005: 4004: 3997: 3996: 3993: 3992: 3989: 3988: 3986: 3979: 3978: 3975: 3974: 3971: 3970: 3969: 3962: 3961: 3958: 3957: 3954: 3952: 3951: 3944: 3943: 3940: 3939: 3936: 3934: 3933: 3926: 3925: 3922: 3921: 3913: 3912: 3904: 3903: 3895: 3894: 3891: 3890: 3889: 3882: 3881: 3878: 3877: 3874: 3873: 3871: 3870:P. otaheitana 3864: 3863: 3857: 3856: 3853: 3852: 3844: 3843: 3840: 3839: 3837: 3836:P. otaheitana 3830: 3829: 3826: 3825: 3822: 3821: 3819: 3818:P. otaheitana 3812: 3811: 3808: 3807: 3804: 3803: 3801: 3800:P. otaheitana 3794: 3793: 3787: 3786: 3783: 3782: 3779: 3778: 3777: 3770: 3769: 3763: 3762: 3759: 3758: 3755: 3753: 3746: 3745: 3739: 3738: 3735: 3734: 3726: 3725: 3722: 3721: 3720: 3713: 3712: 3709: 3708: 3705: 3704: 3703: 3696: 3695: 3692: 3691: 3683: 3682: 3679: 3678: 3677: 3670: 3669: 3666: 3665: 3662: 3661: 3660: 3653: 3652: 3646: 3645: 3642: 3641: 3633: 3632: 3629: 3628: 3627: 3620: 3619: 3616: 3615: 3612: 3611: 3610: 3603: 3602: 3599: 3598: 3595: 3594: 3593: 3586: 3585: 3582: 3581: 3578: 3577: 3576: 3569: 3568: 3565: 3564: 3561: 3560: 3559: 3552: 3551: 3548: 3547: 3544: 3543: 3542: 3535: 3534: 3531: 3530: 3527: 3526: 3525: 3518: 3517: 3511: 3510: 3504: 3503: 3500: 3499: 3491: 3490: 3482: 3481: 3473: 3472: 3469: 3468: 3467: 3466:P. hebe bella 3460: 3459: 3453: 3452: 3449: 3448: 3445: 3444: 3443: 3436: 3435: 3432: 3431: 3423: 3422: 3414: 3413: 3410: 3409: 3408: 3401: 3400: 3394: 3393: 3390: 3389: 3386: 3385: 3384: 3377: 3376: 3370: 3369: 3363: 3362: 3359: 3358: 3350: 3349: 3346: 3345: 3344: 3337: 3336: 3333: 3332: 3329: 3328: 3327: 3320: 3319: 3316: 3315: 3312: 3311: 3310: 3303: 3302: 3299: 3298: 3295: 3294: 3293: 3286: 3285: 3279: 3278: 3275: 3274: 3266: 3265: 3257: 3256: 3253: 3252: 3251: 3244: 3243: 3237: 3236: 3233: 3232: 3229: 3228: 3227: 3220: 3219: 3216: 3215: 3207: 3206: 3203: 3202: 3201: 3194: 3193: 3190: 3189: 3186: 3185: 3184: 3177: 3176: 3173: 3172: 3169: 3168: 3167: 3160: 3159: 3156: 3155: 3152: 3151: 3150: 3143: 3142: 3136: 3135: 3129: 3128: 3122: 3121: 3115: 3114: 3108: 3107: 3101: 3100: 3096: 3095: 3089: 3088: 3082: 3081: 3077: 3076: 3072: 3070: 3066: 3056: 3055: 3054: 3049: 3045: 3044: 3043: 3038: 3037: 3032: 3028: 3027: 3026: 3021: 3020: 3015: 3011: 3010: 3009: 3004: 3003: 2998: 2994: 2993: 2992: 2987: 2986: 2981: 2977: 2976: 2975: 2970: 2969: 2964: 2963: 2958: 2957: 2952: 2948: 2947: 2945: 2941: 2936: 2935: 2930: 2929: 2924: 2922: 2918: 2913: 2909: 2908: 2907: 2902: 2901: 2896: 2892: 2891: 2890: 2886: 2882: 2878: 2873: 2872: 2867: 2863: 2862: 2861: 2860:P. otaheitana 2856: 2855: 2850: 2849: 2844: 2843: 2838: 2837: 2832: 2831: 2826: 2822: 2814: 2812: 2808: 2801: 2797: 2793: 2792: 2791: 2786: 2785: 2781: 2777: 2776: 2775: 2770: 2768: 2764: 2759: 2758: 2753: 2751: 2747: 2742: 2741: 2737: 2733: 2732: 2731: 2726: 2725: 2720: 2719: 2715: 2711: 2710: 2709: 2704: 2703: 2698: 2697: 2692: 2691: 2686: 2685: 2680: 2679: 2675: 2671: 2670: 2669: 2668:P. otaheitana 2664: 2663: 2658: 2657: 2652: 2651: 2646: 2645: 2640: 2636: 2634: 2633: 2628: 2613: 2610: 2607: 2605: 2604: 2603:Partula varia 2600: 2599: 2595: 2591: 2588: 2585: 2582: 2580: 2579: 2575: 2574: 2570: 2567: 2564: 2561: 2559: 2558: 2554: 2553: 2549: 2546: 2543: 2540: 2538: 2537: 2533: 2532: 2528: 2525: 2522: 2519: 2517: 2516: 2512: 2511: 2507: 2504: 2501: 2499: 2498: 2494: 2493: 2489: 2486: 2483: 2480: 2478: 2477: 2473: 2472: 2468: 2465: 2462: 2459: 2457: 2456: 2452: 2451: 2447: 2444: 2441: 2438: 2436: 2435: 2431: 2430: 2426: 2423: 2420: 2417: 2415: 2414: 2410: 2409: 2405: 2402: 2399: 2396: 2394: 2393: 2389: 2388: 2384: 2381: 2378: 2375: 2373: 2372: 2368: 2367: 2363: 2360: 2357: 2355: 2354: 2353:Partula rosea 2350: 2349: 2345: 2342: 2339: 2336: 2334: 2333: 2329: 2328: 2324: 2321: 2318: 2315: 2313: 2312: 2308: 2307: 2303: 2300: 2297: 2294: 2292: 2291: 2287: 2286: 2282: 2279: 2276: 2273: 2271: 2270: 2266: 2265: 2261: 2258: 2255: 2252: 2250: 2249: 2245: 2244: 2240: 2237: 2234: 2231: 2229: 2228: 2224: 2223: 2219: 2216: 2213: 2210: 2208: 2207: 2203: 2202: 2198: 2195: 2192: 2189: 2187: 2186: 2182: 2181: 2177: 2174: 2171: 2168: 2166: 2165: 2161: 2160: 2156: 2152: 2148: 2145: 2142: 2139: 2137: 2136: 2132: 2131: 2127: 2124: 2121: 2118: 2116: 2115: 2111: 2110: 2106: 2102: 2098: 2095: 2092: 2089: 2087: 2086: 2085:Partula obesa 2082: 2081: 2077: 2074: 2071: 2069: 2068: 2064: 2063: 2059: 2056: 2053: 2051: 2050: 2046: 2045: 2041: 2038: 2035: 2033: 2032: 2028: 2027: 2023: 2020: 2017: 2015: 2014: 2010: 2009: 2005: 2002: 1999: 1996: 1994: 1993: 1989: 1988: 1984: 1981: 1978: 1975: 1973: 1972: 1968: 1967: 1963: 1960: 1957: 1954: 1952: 1951: 1947: 1946: 1942: 1939: 1936: 1933: 1931: 1930: 1926: 1925: 1921: 1917: 1914: 1911: 1908: 1906: 1905: 1904:Partula lutea 1901: 1900: 1896: 1892: 1889: 1886: 1883: 1881: 1880: 1876: 1875: 1871: 1868: 1865: 1862: 1860: 1859: 1855: 1854: 1850: 1847: 1844: 1841: 1839: 1838: 1834: 1833: 1829: 1826: 1823: 1820: 1818: 1817: 1813: 1812: 1808: 1805: 1802: 1799: 1797: 1796: 1792: 1791: 1787: 1784: 1781: 1778: 1776: 1775: 1771: 1770: 1766: 1763: 1760: 1757: 1755: 1754: 1750: 1749: 1745: 1741: 1737: 1733: 1730: 1727: 1724: 1722: 1721: 1717: 1716: 1712: 1708: 1705: 1702: 1699: 1697: 1696: 1692: 1691: 1687: 1684: 1681: 1678: 1676: 1675: 1671: 1670: 1666: 1662: 1659: 1656: 1653: 1651: 1650: 1646: 1645: 1641: 1640: 1639:Partula clara 1635: 1632: 1629: 1626: 1624: 1623: 1619: 1618: 1614: 1610: 1607: 1604: 1602: 1599: 1597: 1596: 1592: 1591: 1587: 1584: 1581: 1579: 1578: 1574: 1573: 1569: 1566: 1563: 1560: 1558: 1557: 1553: 1552: 1548: 1545: 1542: 1539: 1537: 1536: 1532: 1531: 1527: 1524:. Extinct on 1523: 1519: 1515: 1511: 1507: 1504: 1501: 1498: 1496: 1495: 1494:Partula gibba 1491: 1490: 1486: 1483: 1481: 1478: 1476: 1475: 1471: 1470: 1466: 1463: 1461: 1458: 1456: 1453: 1451: 1450: 1446: 1445: 1441: 1438: 1435: 1432: 1430: 1429: 1425: 1424: 1420: 1417: 1414: 1411: 1409: 1408: 1404: 1403: 1399: 1396: 1393: 1390: 1388: 1387: 1383: 1382: 1378: 1375: 1372: 1369: 1367: 1366: 1362: 1361: 1357: 1354: 1351: 1348: 1346: 1345: 1341: 1340: 1336: 1333: 1330: 1327: 1325: 1324: 1320: 1319: 1315: 1312: 1309: 1307: 1304: 1302: 1301: 1297: 1296: 1292: 1289: 1286: 1283: 1281: 1280: 1276: 1275: 1271: 1267: 1264: 1261: 1258: 1256: 1255: 1251: 1250: 1246: 1243: 1240: 1237: 1235: 1234: 1230: 1229: 1225: 1222: 1219: 1217: 1216: 1212: 1211: 1207: 1203: 1199: 1196: 1193: 1190: 1188: 1187: 1183: 1182: 1178: 1177: 1172: 1169: 1166: 1163: 1161: 1160: 1156: 1155: 1151: 1148: 1145: 1142: 1140: 1139: 1135: 1134: 1130: 1127: 1124: 1121: 1119: 1118: 1114: 1113: 1109: 1106: 1103: 1100: 1098: 1097: 1096:Partula clara 1093: 1092: 1088: 1085: 1082: 1079: 1077: 1076: 1072: 1071: 1067: 1064: 1061: 1058: 1056: 1055: 1051: 1050: 1046: 1043: 1040: 1037: 1035: 1034: 1030: 1029: 1025: 1021: 1017: 1014: 1011: 1009: 1006: 1004: 1003: 999: 998: 994: 991: 989: 986: 984: 983: 979: 978: 974: 970: 967: 964: 961: 959: 958: 954: 953: 949: 946: 943: 941: 938: 936: 935: 931: 930: 926: 922: 919: 916: 914: 911: 909: 908: 904: 903: 899: 897: 894: 892: 889: 887: 884: 881: 880: 874: 871: 870: 864: 862: 852: 850: 844: 842: 841:least concern 838: 834: 830: 826: 822: 818: 814: 809: 807: 802: 800: 796: 791: 789: 788: 783: 782: 777: 767: 765: 761: 757: 753: 749: 744: 742: 738: 734: 733:Pacific Ocean 730: 726: 722: 717: 715: 711: 708: 705: 702: 698: 694: 690: 689: 682: 681: 676: 670: 669: 668:Partula gibba 664: 658: 657: 652: 648: 643: 631:Hartman, 1881 629: 626: 623:Iredale, 1941 621: 618: 612: 609: 605:Hartman, 1881 603: 600: 597:Pilsbry, 1909 595: 592: 588:Goodsir, 1845 586: 581:Hartman, 1881 579: 576: 570: 567: 561: 556:Lamarck, 1812 554: 547: 542:Hartman, 1881 540: 537: 531: 528: 524:Lamarck, 1818 522: 517:Hartman, 1881 515: 512: 506: 501:Hartman, 1881 499: 496: 493:Pilsbry, 1909 491: 488: 485:Hartman, 1881 483: 480: 477:Pilsbry, 1909 475: 472: 469:Pilsbry, 1909 467: 464: 461:Pilsbry, 1909 459: 456: 450: 445:Hartman, 1881 443: 440: 437: 433:Mulsant, 1846 431: 426:Hartman, 1881 424: 421: 415: 410:Hartman, 1881 408: 405: 399: 394:Hartman, 1881 392: 389: 386:Pilsbry, 1909 384: 381: 375: 370:Hartman, 1881 368: 365: 362:Hartman, 1881 360: 357: 354:Hartman, 1881 352: 349: 346:Hartman, 1881 344: 341: 338:Pilsbry, 1909 336: 333: 330:Hartman, 1881 328: 325: 322:Hartman, 1881 320: 317: 314:Hartman, 1881 311: 308:Hartman, 1881 305: 302: 299:Hartman, 1881 296: 295: 287: 284: 280: 275: 270: 269: 265: 262: 261: 258: 255: 252: 251: 248: 245: 243:Superfamily: 242: 241: 238: 235: 232: 231: 228: 225: 222: 221: 218: 215: 212: 211: 208: 205: 202: 201: 198: 195: 192: 191: 188: 185: 182: 181: 176: 171: 167: 164: 163: 158: 154: 149: 146: 142: 139: 131: 128: 120: 109: 106: 102: 99: 95: 92: 88: 85: 81: 78: –  77: 73: 72:Find sources: 66: 62: 56: 55: 50:This article 48: 44: 39: 38: 33: 19: 5821: 5769: 5765: 5752: 5743: 5732: 5710: 5706: 5702: 5682: 5673: 5664: 5658:"Tentacle 3" 5652: 5640:. Retrieved 5636: 5626: 5615:. Retrieved 5611: 5601: 5589:. Retrieved 5564:. Retrieved 5555: 5533: 5529: 5520: 5512: 5507: 5496: 5483: 5475: 5472: 5464: 5456: 5442: 5426: 5410: 5394: 5371: 5357:Partula hebe 5355: 5339: 5316: 5300: 5277: 5261: 5245: 5229: 5213: 5190: 5174: 5158: 5142: 5126: 5110: 5079: 5069: 5065: 5064: 5017: 5016: 4993: 4992: 4978: 4928: 4927: 4911: 4910: 4898:Palaopartula 4896: 4882: 4881: 4858: 4857: 4841: 4840: 4806: 4805: 4789: 4788: 4765: 4764: 4743: 4720: 4719: 4703: 4702: 4688:P. auraniana 4686: 4685: 4660: 4659: 4615: 4614: 4597: 4596: 4554: 4553: 4529: 4462: 4461: 4438: 4437: 4414: 4413: 4390: 4389: 4373: 4372: 4328:P. similaris 4326: 4325: 4309: 4308: 4294:P. radiolata 4292: 4291: 4275: 4274: 4249: 4248: 4234:P. assimilis 4232: 4231: 4181: 4180: 4157: 4156: 4106: 4105: 4082: 4081: 4067:P. dentifera 4065: 4064: 4043:P. dentifera 4041: 4040: 4003:P. tohiveana 4001: 4000: 3985:P. mirabilis 3983: 3982: 3966: 3965: 3948: 3947: 3930: 3929: 3886: 3885: 3868: 3867: 3834: 3833: 3816: 3815: 3798: 3797: 3774: 3773: 3749: 3717: 3716: 3700: 3699: 3674: 3673: 3657: 3656: 3624: 3623: 3607: 3606: 3590: 3589: 3573: 3572: 3556: 3555: 3539: 3538: 3522: 3521: 3464: 3463: 3440: 3439: 3405: 3404: 3381: 3380: 3341: 3340: 3324: 3323: 3309:P. mirabilis 3307: 3306: 3290: 3289: 3248: 3247: 3224: 3223: 3198: 3197: 3181: 3180: 3164: 3163: 3147: 3146: 3098: 3097: 3078: 3069:monophyletic 3064: 3062: 3053:P. assimilis 3051: 3048:Cook Islands 3047: 3046: 3042:P. similaris 3040: 3036:P. auraniana 3034: 3030: 3029: 3023: 3019:P. cramptoni 3017: 3013: 3012: 3006: 3000: 2996: 2995: 2989: 2983: 2979: 2978: 2974:P. radiolata 2972: 2968:P. lutaensis 2966: 2962:P. langfordi 2960: 2954: 2950: 2949: 2944:P. dentifera 2938: 2932: 2926: 2917:P. garrettii 2915: 2911: 2910: 2904: 2898: 2894: 2893: 2889:P. tohiveana 2881:P. suturalis 2874: 2871:P. mirabilis 2870: 2865: 2864: 2858: 2852: 2846: 2840: 2834: 2828: 2824: 2823: 2820: 2810: 2807:P. garrettii 2806: 2805: 2799: 2790:P. langfordi 2788: 2782: 2779: 2778: 2772: 2767:P. dentifera 2761: 2755: 2746:P. garrettii 2744: 2738: 2735: 2734: 2728: 2722: 2716: 2713: 2712: 2708:P. tohiveana 2706: 2700: 2696:P. suturalis 2694: 2688: 2684:P. mirabilis 2682: 2676: 2673: 2672: 2666: 2660: 2654: 2648: 2642: 2638: 2637: 2630: 2626: 2624: 2601: 2576: 2555: 2534: 2513: 2495: 2474: 2453: 2432: 2411: 2390: 2371:Partula rufa 2369: 2351: 2330: 2309: 2288: 2267: 2246: 2225: 2204: 2183: 2162: 2133: 2112: 2083: 2065: 2047: 2029: 2011: 1990: 1969: 1948: 1927: 1902: 1877: 1856: 1835: 1814: 1793: 1772: 1751: 1718: 1693: 1672: 1647: 1637: 1620: 1612: 1593: 1577:Partula hebe 1575: 1554: 1533: 1508:Persists in 1492: 1472: 1447: 1428:Partula faba 1426: 1405: 1384: 1363: 1342: 1321: 1298: 1277: 1252: 1231: 1213: 1184: 1174: 1157: 1136: 1115: 1094: 1073: 1052: 1031: 1000: 980: 973:habitat loss 955: 932: 905: 860: 858: 845: 816: 815:contains 73 810: 803: 794: 792: 785: 779: 773: 756:Yoshio Kondo 754:(along with 745: 720: 718: 687: 686: 685: 678: 666: 654: 650: 627: 619: 614:Clench, 1941 610: 601: 593: 584: 577: 572:Clench, 1941 568: 559: 552: 545: 538: 529: 520: 513: 504: 497: 489: 481: 473: 465: 457: 448: 441: 435: 429: 422: 413: 406: 397: 390: 382: 377:Röding, 1798 373: 366: 358: 350: 342: 334: 326: 318: 267: 266: 233:Infraorder: 160: 144: 138: 123: 114: 104: 97: 90: 83: 71: 59:Please help 54:verification 51: 5913:iNaturalist 5846:Wikispecies 5699:Mayer A. G. 5526:Burch J. B. 5019:Eua zebrina 4995:Eua globosa 3968:P. mooreana 3888:P. producta 3838:sinistrorsa 3820:sinistralis 3702:P. mooreana 3626:P. taeniata 3558:P. mooreana 3149:P. aurantia 2985:P. emersoni 2885:P. taeniata 2877:P. mooreana 2848:P. incrassa 2702:P. taeniata 2690:P. mooreana 2678:P. aurantia 2592:Endemic to 2099:Endemic to 1893:Endemic to 1734:Endemic to 1020:Aore Island 804:Today, the 701:terrestrial 697:land snails 620:Scilistylus 452:Risso, 1826 417:Selys, 1853 247:Pupilloidea 237:Pupilloidei 6000:Categories 5617:2023-05-27 5453:References 4705:P. turneri 4619:intermedia 4416:P. affinis 4251:P. luteola 4183:P. turgida 4084:P. tristis 3950:P. hyalina 3932:P. hyalina 3719:P. tohivea 3343:P. hyalina 3080:Partulidae 2980:Micronesia 2921:P. tristis 2842:P. hyalina 2830:P. affinis 2780:Marianas - 2774:P. turgida 2750:P. tristis 2656:P. hyalina 2644:P. affinis 925:Tahiti Iti 837:vulnerable 833:endangered 748:Polynesian 741:New Guinea 714:Partulidae 549:Oken, 1807 436:; Marianna 257:Partulidae 217:Gastropoda 87:newspapers 5683:d-nb.info 5447:(extinct) 4722:P. lirata 4617:P. nodosa 4601:composita 4599:P. nodosa 4466:rubescens 4108:P. meyeri 3987:propinqua 3953:(Tahiti) 3541:P. exigua 3059:Cladogram 3025:P. micans 3008:P. lirata 3002:P. leefei 2934:P. meyeri 2854:P. nodosa 2736:Raiatea - 2718:P. arguta 2714:Huahine - 2662:P. nodosa 2547:Formerly 1920:Bora Bora 1764:Formerly 1685:Formerly 1665:Mt. Aorai 1505:Formerly 1439:Formerly 1270:Mt. Marau 1149:Formerly 1128:Formerly 1107:Formerly 1044:Formerly 947:Formerly 867:Full list 863:include: 827:, 17 are 823:, 10 are 707:gastropod 704:pulmonate 611:Rennellia 563:Lea, 1833 193:Kingdom: 187:Eukaryota 117:June 2024 5837:Q6296067 5831:Wikidata 5788:17610827 5731:(1916). 5637:Mongabay 5086:turmeric 5081:Dracaena 5076:plantain 5071:Caladium 4277:P. gibba 3802:lignaria 3659:P. clara 3524:P. clara 3407:P. rosea 3383:P. varia 3292:P. clara 2956:P. gibba 2951:Marianas 2906:P. varia 2900:P. rosea 2836:P. clara 2784:P. gibba 2730:P. varia 2724:P. rosea 2674:Moorea - 2650:P. clara 2594:Vanikoro 1526:Aguiguan 882:Species 835:, 2 are 831:, 8 are 781:Achatina 710:mollusks 656:Caladium 602:Pasithea 560:Pasithea 553:Pasithea 546:Pasithea 430:Harmonia 335:Marianna 319:Harmonia 297:Astraea 283:Synonyms 274:Férussac 253:Family: 207:Mollusca 203:Phylum: 197:Animalia 183:Domain: 5931:1349766 5905:4597417 5892:4925260 5852:Partula 5822:Partula 5703:Partula 5591:28 June 5566:28 June 5530:Partula 5513:Partula 5092:Gallery 5066:Partula 5061:Ecology 4745:Samoana 4159:P. faba 3442:P. hebe 3099:Partula 3065:Partula 2991:P. rufa 2928:P. hebe 2912:Raiatea 2895:Huahine 2800:Partula 2757:P. hebe 2740:P. faba 2632:ex situ 2627:Partula 1514:Sarigan 861:Partula 855:Species 849:Mo'orea 821:extinct 817:Partula 799:extinct 795:Partula 770:Decline 727:" and " 721:Partula 688:Partula 651:Partula 628:Sterope 585:Sterope 374:Astraea 306:Clytia 268:Partula 263:Genus: 223:Order: 213:Class: 145:Partula 101:scholar 5983:861628 5970:324349 5957:362162 5944:145553 5786:  5642:17 May 4558:crassa 3872:crassa 2866:Moorea 2825:Tahiti 2639:Tahiti 2105:Futuna 1711:Tahiti 1522:Saipan 900:Notes 776:Tahiti 647:Saipan 521:Oenone 449:Helena 398:Clytia 359:Oenone 343:Matata 327:Helena 276:, 1821 103:  96:  89:  82:  74:  5978:WoRMS 5926:IRMNG 5918:90863 5879:7PBZ7 5661:(PDF) 2143:6-10 2122:5-10 2101:Alofi 1744:Naiau 1740:Cicia 1518:Pagan 693:genus 691:is a 505:Nenia 351:Nenia 312:Echo 108:JSTOR 94:books 5939:NCBI 5900:GBIF 5866:3012 5861:BOLD 5784:PMID 5711:XXVI 5679:"34" 5644:2023 5593:2024 5568:2024 5084:and 5074:and 2997:Fiji 2809:and 2625:The 2614:Yes 2508:Yes 2487:Yes 2466:Yes 2364:Yes 2316:N/A 2298:1-2 2256:4-5 2155:Aore 2153:and 2103:and 2078:Yes 2060:Yes 2042:Yes 2024:Yes 1955:N/A 1895:Rota 1884:N/A 1742:and 1736:Mago 1608:Yes 1605:1-5 1588:Yes 1520:and 1510:Guam 1487:Yes 920:Yes 558:and 414:Echo 80:news 5887:EoL 5874:CoL 5774:doi 5713:(2) 5532:". 4980:Eua 2608:EW 2589:No 2583:CR 2568:No 2562:EX 2541:EX 2520:EX 2502:EW 2481:CR 2460:EW 2445:No 2439:LC 2424:No 2418:EX 2403:No 2397:EX 2382:No 2376:PE 2358:EW 2343:No 2337:EX 2322:No 2301:No 2295:EN 2280:No 2274:EN 2259:No 2253:EN 2238:No 2232:EX 2217:No 2211:EX 2196:No 2190:EX 2175:No 2169:EX 2146:No 2140:VU 2119:EN 2096:No 2090:CR 2072:EW 2054:EW 2036:EW 2018:EW 2003:No 1997:LC 1982:No 1976:PE 1961:No 1940:No 1934:EX 1915:No 1909:EX 1890:No 1869:No 1863:EX 1848:No 1845:11 1842:EN 1827:No 1821:EX 1806:No 1800:EX 1785:No 1779:CR 1758:EX 1731:No 1725:EN 1706:No 1700:PE 1679:EX 1660:No 1654:PE 1633:No 1627:CR 1613:may 1582:EW 1567:No 1561:EX 1546:No 1540:LC 1499:EN 1464:No 1433:EX 1418:No 1412:EX 1397:No 1391:CR 1376:No 1370:EX 1355:No 1349:EX 1334:No 1328:EX 1313:No 1306:N/A 1284:EX 1265:No 1259:PE 1244:No 1238:EX 1226:No 1220:EX 1197:No 1191:CR 1170:No 1164:EX 1143:PE 1122:EX 1104:20 1101:CR 1086:No 1080:EX 1065:No 1059:EX 1038:EX 1015:No 995:No 968:No 962:CR 739:to 716:. 63:by 6002:: 5980:: 5967:: 5954:: 5941:: 5928:: 5915:: 5902:: 5889:: 5876:: 5863:: 5848:: 5833:: 5782:. 5770:17 5768:. 5764:. 5691:^ 5681:. 5663:. 5635:. 5610:. 5576:^ 5541:^ 5476:92 5088:. 4534:) 3754:) 3050:- 3039:, 3033:- 3022:, 3016:- 3005:, 2999:- 2988:, 2982:- 2971:, 2965:, 2959:, 2953:- 2937:, 2931:, 2925:, 2914:- 2903:, 2897:- 2887:, 2883:, 2879:, 2875:, 2868:- 2857:, 2851:, 2845:, 2839:, 2833:, 2827:- 2787:, 2771:, 2760:, 2754:, 2743:, 2727:, 2721:, 2705:, 2699:, 2693:, 2687:, 2681:, 2665:, 2659:, 2653:, 2647:, 2641:- 2611:0 2596:. 2586:1 2565:0 2544:0 2523:0 2505:0 2484:2 2463:0 2442:? 2421:0 2400:0 2379:1 2361:0 2340:0 2319:? 2277:1 2235:0 2214:0 2193:0 2172:0 2093:2 2075:0 2057:0 2039:0 2021:0 2000:? 1979:1 1958:0 1937:0 1912:0 1887:? 1866:0 1824:0 1803:0 1782:1 1761:0 1746:. 1738:, 1728:3 1713:. 1703:1 1682:0 1667:. 1657:1 1642:. 1630:1 1601:VU 1585:0 1564:0 1543:? 1528:. 1516:, 1512:, 1502:4 1484:0 1480:EW 1455:LC 1436:0 1415:0 1394:2 1373:0 1352:0 1331:0 1310:0 1287:0 1262:1 1241:0 1223:0 1208:. 1194:1 1167:0 1146:1 1125:0 1083:0 1062:0 1041:0 1026:. 1012:5 1008:EN 992:0 988:EX 965:1 944:0 940:PE 917:1 913:CR 851:. 843:. 801:. 766:. 743:. 699:, 590:) 565:) 551:, 526:) 510:) 454:) 403:) 379:) 5790:. 5776:: 5685:. 5667:. 5646:. 5620:. 5595:. 5570:. 5536:. 5491:. 4530:( 3750:( 2946:) 2942:( 2923:) 2919:( 2769:) 2765:( 2752:) 2748:( 1460:? 896:C 891:P 886:S 130:) 124:( 119:) 115:( 105:· 98:· 91:· 84:· 57:. 34:. 20:)

Index

Partula (genus)
Partula (disambiguation)

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Partula radiolata
Scientific classification
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Eukaryota
Animalia
Mollusca
Gastropoda
Stylommatophora
Pupilloidei
Pupilloidea
Partulidae
Partula
Férussac
Synonyms

Saipan

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