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Parasaurolophus

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2455: 2228:. Other hadrosaurids had large heads without needing large hollow crests to serve as attachment points for supporting ligaments. Also, none of the proposals explain why the crest has such a shape, why other lambeosaurines should have crests that look much different but perform a similar function, how crestless or solid-crested hadrosaurids got along without such capabilities, or why some hadrosaurids had solid crests. These considerations particularly impact hypotheses based on increasing the capabilities of systems already present in the animal, such as the salt gland and olfaction hypotheses, and indicate that these were not primary functions of the crest. Additionally, work on the nasal cavity of lambeosaurines shows that 7821: 3019: 7816: 6906: 2472:. The skeleton shows a v-shaped gap or notch in the vertebrae at the base of the neck. Originally thought to be pathologic, Parks published a second interpretation of this, as a ligament attachment to support the head. The crest would attach to the gap via muscles or ligaments, and be used to support the head while bearing a frill, like predicted to exist in some hadrosaurids. One other possibility, is that during preparation, the specimen was damaged, creating the possible pathology. The notch, however, is still considered more likely to be a pathology, even though some illustrations of 2011: 6920: 3010: 2506: 1953: 146: 7803: 5848: 930: 2051: 6913: 171: 1893: 2404: 5860: 7827: 705: 1296: 1088: 2150: 2354:, has allowed the reconstruction of the possible sound its crest produced. The main path resonates at around 30 Hz, but the complicated sinus anatomy causes peaks and valleys in the sound. The other main behavioral theory is that the crest was used for intra-species recognition. This means that the crest could have been used for species recognition, as a warning signal, and for other, non-sexual uses. These could have been some of the reasons crests evolved in 5812: 2263: 7810: 1168: 2694: 5836: 5800: 2446:. This led to the frill being depicted in many other sources, though the advent of the now-debunked "snorkel" hypothesis, and conflation of the frill hypothesis with the idea that the crest serves as an anchor point for neck ligaments, along with lack of strong evidence for its presence, has seen it fall out of favor in most modern depictions. 2335:, of a slender stapes (reptilian ear bone) in place, which combined with a large space for an eardrum implies a sensitive middle ear. Furthermore, the hadrosaurid lagena is elongate like a crocodilian's, indicating that the auditory portion of the inner ear was well-developed. Based on the similarity of hadrosaurid 2062:
performed, but most have been discredited. It is now believed that it may have had several functions: visual display for identifying species and sex, sound amplification for communication, and thermoregulation. It is not clear which was most significant at what times in the evolution of the crest and
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was plausibly the result of the dinosaur being hit by a falling tree, perhaps during a severe storm. Based on the regrowth of bone, it is suggested that the hadrosaur survived for at least one to four months to perhaps years after being injured. None of the pathologies on the holotype individual are
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Most of these hypotheses have been discredited or rejected. For example, there is no hole at the end of the crest for a snorkeling function. There are no muscle scars for a proboscis and it is dubious that an animal with a beak would need one. As a proposed airlock, it would not have kept out water.
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of any ontogenetic stage. The endocast was reconstructed in two sections, one on the portion of the braincase articulated with the left half of the skull and the remainder on the disarticulated portion of the braincase. Their relative position was then approximated based on cranial landmarks and
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is estimated at 9.45 metres (31.0 ft), and allometry-based body mass estimates indicate that a 9 metres (30 ft) long individual would have weighed more than 5 metric tons (5.5 short tons). Gregory S. Paul estimated that an average adult individual of the type species would measure 7.5
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and Osmólska, who realized that like modern lizards, dinosaurs could have possessed salt glands, and cooled off by osmo-regulation. In 2006 Evans published an argument about the functions of lambeosaurine crests, and supported why this could be a causing factor for the evolution of the crest.
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of large Late Cretaceous ornithopods that are known for their range of bizarre head adornments, which were likely used for communication and increased hearing. This genus is known for its large, elaborate cranial crest, which forms a long curved tube projecting upwards and back from the
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Another possible pathology was noticed by Parks, and from around the notch. In the fourth, fifth, and sixth vertebrae, directly anterior to the notch, the neural spines were damaged. The fourth had an obvious fracture, with the other two possessing a swelling at the base of the break.
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habits, evidence that sight was important to these animals. If, as is commonly illustrated, a skin frill extended from the crest to the neck or back, the proposed visual display would have been even showier. As is suggested by other lambeosaurine skulls, the crest of
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form. Neither hypothesis became widely accepted. As only six good skulls, one juvenile braincase, and one recently discovered juvenile skull are known, additional material will help clear up these potential relationships. Williamson noted that in any case, juvenile
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swamps, ponds and lakes, and was bordered by highlands. The climate was wet and humid, and supported an abundant and diverse range of organisms. This formation contains one of the best and most continuous records of Late Cretaceous terrestrial life in the world.
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remains are rare in Alberta, with only one other partial skull that is possibly from the Dinosaur Park Formation and three Dinosaur Park specimens lacking their skulls that possibly belong to the genus. In some faunal lists, there is a mention of possible
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Manucci, F, Dempsey, M, Tanke, D H., et al. Description and etiology of paleopathological lesions in the type specimen of Parasaurolophus walkeri (Dinosauria: Hadrosauridae), with proposed reconstructions of the nuchal ligament J. Anat. 2020; 00: 1– 15.
1928:. Its teeth were continually being replaced and were packed into dental batteries containing hundreds of teeth, but only a relative handful of which were in use at any time. It used its beak to crop plant material, which was held in the jaws by a 2244:
of the crest also suggests a thermoregulatory function. The first to propose the cranial crests of lambeosaurines related to temperature regulation was Wheeler (1978). He proposed that there was a nerve connection between the crest and the
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species. It lived in New Mexico, where three specimens are known, and can be differentiated from its other species. It possesses a long and straight crest, with a very complex interior compared to the other species. All known specimens of
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Roberts, E.M.; Deino, A.L.; Chan, M.A. (2005). "40Ar/39Ar age of the Kaiparowits Formation, southern Utah, and correlation of contemporaneous Campanian strata and vertebrate faunas along the margin of the Western Interior Basin".
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in northern latitude fossil sites may represent faunal exchange between otherwise distinct northern and southern biomes in Late Cretaceous North America. Both taxa are uncommon outside of the southern biome, where, along with
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is often hypothesized to have used its crest as a resonating chamber to produce low frequency sounds to alert other members of a group or its species. This function was originally suggested by Wiman in 1931 when he described
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treatment in 1999 that covered aspects of its anatomy and taxonomy, as well as the functions of its crest. Williamson later published an independent review of the remains that disagreed with the taxonomic conclusions.
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Jasinski, S.E.; Sullivan, R.M. (2011). "Re-evaluation of pachycephalosaurids from the Fruitland-Kirtland transition (Kirtlandian, late Campanian), San Juan Basin, New Mexico, with a description of a new species of
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Eaton, J.G. (2002). "Multituberculate mammals from the Wahweap (Campanian, Aquilan) and Kaiparowits (Campanian, Judithian) formations, within and near Grand Staircase-Escalante National Monument, southern Utah".
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that was similar to but smaller than what is seen in adult specimens; in adults, the frontals formed a platform that supported the base of the crest. This specimen also indicates that the growth of the crest in
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Barnum Brown (1912) noted the presence of fine striations near the back of the crest that he hypothesized could be associated with the presence of a frill of skin, comparable to the one found in the modern
2343:, he also proposed that adult hadrosaurids were sensitive to high frequencies, such as their offspring might produce. According to Weishampel, this is consistent with parents and offspring communicating. 2200:
proposed it was used as a weapon in combat among members of the same species, and Andrew Milner suggested that it could be used as a foliage deflector, like the helmet crest (called a 'casque') of the
1187:. The crest was hollow, with distinct tubes leading from each nostril to the end of the crest before reversing direction and heading back down the crest and into the skull. The tubes were simplest in 4538:
Eaton, J.G.; Cifelli, R.L.; Hutchinson, J.H.; Kirkland, J.I.; Parrish, M.J. (1999). "Cretaceous vertebrate faunas from the Kaiparowits Plateau, south-central Utah". In Gillete, David D. (ed.).
2295:. He noted that the crest's internal structures are similar to those of a swan and theorized that an animal could use its elongated nasal passages to create noise. However, the nasal tubes of 5535:
Weishampel, David B.; Barrett, Paul M.; Coria, Rodolfo A.; Le Loeuff, Jean; Xu Xing; Zhao Xijin; Sahni, Ashok; Gomani, Elizabeth, M.P.; Noto, Christopher R. (2004). "Dinosaur Distribution".
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Sullivan, R.S.; Williamson, T.E. (1996). "A new skull of Parasaurolophus (long-crested form) from New Mexico: external and internal (CT scans) features and their functional implications".
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Williamson, T.E. (2000). Lucas, Spencer G.; Heckert, Andrew B. (eds.). "Dinosaurs of New Mexico: Review of Hadrosauridae (Dinosauria: Ornithischia) from the San Juan Basin, New Mexico".
2670:, may represent the remains of individuals who died while migrating through the region. They might also have had a more upland habitat where they may have nested or fed. The presence of 5347: 4061:"Description and etiology of paleopathological lesions in the type specimen of Parasaurolophus walkeri (Dinosauria: Hadrosauridae), with proposed reconstructions of the nuchal ligament" 5333:"Fossil Record 3: The first 'lambeosaurin' (Dinosauria, Hadrosauridae, Lambeosaurinae) from the Upper Cretaceous Ojo Alamo Formation (Naashoibito Member), San Juan Basin, New Mexico" 2370:
identification and communication. As a large object, the crest has clear value as a visual signal and sets this animal apart from its contemporaries. The large size of hadrosaurid
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Many early suggestions focused on adaptations for an aquatic lifestyle, following the hypothesis that hadrosaurids were amphibious, a common line of thought until the 1960s. Thus,
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being a female, many other studies have found that it is not because of the differences in age, distribution, and the large differences in the crest and its internal structure.
2007:, because of this difference in age between when their crests started developing. The crest of the juvenile is not long and tubular like the adults, but low and hemispherical. 2196:
or muscles, and helped with moving and supporting the head. This is unlikely, because in all modern archosaurs, the nuchal ligament attaches to the neck or base of the skull.
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from China, which may have been its closest relative, had a similar skull and a potentially similar crest. Visual recognition of both species and sex, acoustic resonance, and
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and corresponding sensory tissue were largely outside the portion of the nasal passages in the crest, so the expansion of the crest had little to do with the sense of smell.
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metres (25 ft) long and weigh 2.6 metric tons (2.9 short tons). Its skull is about 1.6 m (5 ft 3 in) long, including the crest, whereas the type skull of
5320:"The Kirtlandian Land-Vertebrate "Age"-Faunal Composition, Temporal Position, and Biostratigraphic Correlation in the Nonmarine Upper Cretaceous of Western North America" 2331:(interpreted as a possible female). Hopson found that there is anatomical evidence that hadrosaurids had a strong hearing. There is at least one example, in the related 2996:, Troodontidae and Tyrannosauridae. The beginning of the Kirtland Formation dates to 74.55 ± 0.22 mya, with the formation ending at around 73.05 ± 0.25 mya. 4367:
Brett-Surman, Michael K.; Wagner, Jonathan R. (2006). "Appendicular anatomy in Campanian and Maastrichtian North American hadrosaurids". In Carpenter, Kenneth (ed.).
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is over 2 m (6 ft 7 in) long, indicating it was a larger animal. Its single known arm was relatively short for a hadrosaurid, with a short but wide
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formations of Utah and New Mexico. The second specimen, the first known from the Kaiparowits Formation, was originally unassigned to a specific taxon. Of the
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is the smallest and has the most curved crest. Because of its possession of the two above features, it has often been speculated that it was a female of
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Other proposals were more mechanical in nature. William Parks, in 1922, suggested that the crest was joined to the vertebrae above the shoulders by
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growth stages, and lived approximately 75 million years ago. Even though no complete skull of the intermediate age between RAM 14000 and adult
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Like other hadrosaurids, it was able to walk on either two legs or four. It probably preferred to forage for food on four legs, but ran on two. The
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age. This occurrence is not noted by Sullivan and Williamson in their 1999 review of the genus and has not been further elaborated upon elsewhere.
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has been found yet, a partial braincase of about the right size is known. At 25% of the total adult size, the juvenile show that crest growth of
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comparison with other hadrosaurids. Because of weathering, many of the smaller neural canals and foramina could not be identified for certain.
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have been proposed as functional explanations for the crest. It is one of the rarer hadrosaurids, known from only a handful of good specimens.
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Horner, J.A.; Weishampel, D.B.; Forster, C.A. (2004). "Hadrosauridae". In Weishampel, David B.; Osmólska, Halszka; Dodson, Peter (eds.).
5648:"New Basal Hadrosauroid Dinosaur (Dinosauria: Ornithopoda) with Transitional Features from the Late Cretaceous of Henan Province, China" 749:
768, a skull and partial skeleton missing most of the tail and the back legs below the knees, which was found by a field party from the
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is known from many adult specimens, and a juvenile described in 2013, numbered RAM 140000 and nicknamed Joe, after a volunteer at the
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indicates that the Kaiparowits Formation was deposited between 76.6 and 74.5 million years ago, during the Campanian age of the Late
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Bertozzo, Filippo; Manucci, Fabio; Dempsey, Matthew; Tanke, Darren H.; Evans, David C.; Ruffell, Alastair; Murphy, Eileen (2020).
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Zanno, L.E.; Sampson, S.D. (2005). "A new oviraptorosaur (Theropoda; Maniraptora) from the Late Cretaceous (Campanian) of Utah".
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Xing, H.; Wang, D.; Han, F.; Sullivan, C.; Ma, Q.; He, Y.; Hone, D.W.E.; Yan, R.; Du, F.; Xu, X. (2014). Evans, David C. (ed.).
5359: 8363: 2116:, provides evidence that a small tubular crest was present in juveniles. This specimen preserves a small upward flaring of the 7820: 8423: 6905: 5892: 5397: 5364:, n. gen., n. sp., (Pterosauria, ?Azhdarchidae) from the Upper Cretaceous Kirtland Formation (upper Campanian) of New Mexico" 5240: 5104: 5053: 4978: 4928: 4789: 4547: 4493: 4376: 4357: 4338: 3722: 1964: 952:. All of them can be clearly distinguished from each other and have many differences. The first named species, therefore the 8368: 3047:
period. During the Late Cretaceous period, the site of the Kaiparowits Formation was located near the western shore of the
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that could reach over 9 metres (30 ft) long and weigh over 5 metric tons (5.5 short tons), and were able to move as a
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to the east. The plateau where dinosaurs lived was an ancient floodplain dominated by large channels and abundant wetland
1932:-like organ. Vegetation could have been taken from the ground up to a height of around 4 m (13 ft). As noted by 8388: 7815: 6919: 4863: 5761: 5215: 5307: 893:
or, more likely, the base of the overlying Kirtland Formation. The range of this species was described in 1979, when
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were tall, as was common in lambeosaurines. At their tallest over the hips, they increased the height of the back.
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The most noticeable feature was the cranial crest that protruded from the rear of the head and was made up of the
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Sullivan, R.S.; Jasinski, S.E.; Guenther, M.; Lucas, S.G. (2011). Sullivan, Robert S.; Lucas, Spencer G. (eds.).
2892: 2608:. It was a rare constituent of this fauna. The Dinosaur Park Formation is interpreted as a low-relief setting of 2432:
with a substantial frill of skin between the crest and neck appeared in influential paleoart including murals by
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The Illustrated Encyclopedia of Dinosaurs: An Original and Compelling Insight into Life in the Dinosaur Kingdom
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lacks the thin bony 'coxcomb' that makes up the upper portion of the crest of the other three lambeosaurines.
2083:, one of the first researchers to describe lambeosaurine crests in terms of such distinctions, suggested that 8408: 8393: 8196: 5745: 4390:, gen. et sp. nov., a new tyrannosauroid from New Mexico and the origin of deep snouts in Tyrannosauroidea". 170: 8378: 1237:. It is usually interpreted as a separate offshoot of the lambeosaurines, distinct from the helmet-crested 7826: 2852: 2751: 1308: 819: 2487:
published in December 2020, suggests the pathology to the shoulder and thoracic ribs in the holotype of
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ever found, and measures 2.5 m (8.2 ft). This individual fits neatly into the currently known
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existed, the Fruitland Formation was swampy, positioned in the lowlands, and close to the shore of the
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Some of the less common hadrosaurs in the Dinosaur Park Formation of Dinosaur Provincial Park, such as
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Benson, R.J.; Brussatte, S.J.; Anderson; Hone, D.; Parsons, K.; Xu, X.; Milner, D.; Naish, D. (2012).
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shared its habitat with other ornithischians and theropods. Specifically, its contemporaries were the
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in their crests (~70% of adult size). Even though many scientists have supported the possible fact of
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Evans, D.C. (2006). "Nasal cavity homologies and cranial crest function in lambeosaurine dinosaurs".
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A partial cranial endocast for RAM 14000 was reconstructed from CT scan data, the first ever for a
902: 5293:(Dinosauria: Hadrosauridae) from the Kirtland Formation of New Mexico and a revision of the genus" 1944:
had a diet consisting of leaves, twigs, and pine needles which would imply that it was a browser.
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Ostrom, J.H. (1961). "A New Species of Hadrosaurian Dinosaur from the Cretaceous of New Mexico".
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Diegert, C.F.; Williamson, T.E. (1998). "A digital acoustic model of the lambeosaurine hadrosaur
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skeleton except for the feet, neck, and parts of the tail. Its specific name is derived from the
767: 5389: 5383: 4841:"Does mutual sexual selection explain the evolution of head crests in pterosaurs and dinosaurs?" 4697:
Farke, A.A.; Chok, D.J.; Herrero, A.; Scolieri, B.; Werning, S. (2013). Hutchinson, John (ed.).
4485: 4479: 8330: 8239: 6351: 6203: 5152: 3248: 2923: 2914: 2904: 2873: 2311:. A large amount of material and data supports the hypothesis that the large, tubular crest of 1940:
and its relatives could feed more selectively than their broad-beaked, crestless counterparts.
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because of its superficially similar crest. However, it was soon reassessed as a member of the
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lived approximately a million years later. As noted by Thomas Williamson, the type material of
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Hopson, J.A. (1975). "The Evolution of Cranial Display Structures in Hadrosaurid Dinosaurs".
4628: 4505:"Presidential Proclamation: Establishment of the Grand Staircase-Escalante National Monument" 4330:
The Dinosaur Heresies: New Theories Unlocking the Mysteries of Dinosaurs and their Extinction
4328: 3134: 3056: 3028: 2948: 2861: 2846: 2166: 1968: 1277: 1004: 910: 906: 750: 4598:, a crested hadrosaurid dinosaur (Ornithischia) from the Dinosaur Park Formation, Alberta". 4542:. Miscellaneous Publication 99-1. Salt Lake City: Utah Geological Survey. pp. 345–353. 8286: 7581: 5878: 5864: 5659: 5567: 5457: 5072: 5008: 4960: 4883: 4819: 4677: 4644: 4566: 4427: 4399: 4299: 2442: 2213: 2186: 1152: 792: 779: 746: 449: 396: 377: 326: 7809: 4805:
n. g., n. sp., a lambeosaurine dinosaur from the Late Maastrichtian of northeastern China"
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Evans, D.C.; Larson, D.W.; Cullen, T.M.; Sullivan, R.M. (2014). Sues, Hans-Dieter (ed.). "
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The skull of RAM 14000 is almost complete, with the left side only lacking a piece of the
8: 5326:. New Mexico Museum of Natural History and Science Bulletin. Vol. 35. pp. 7–23. 4437: 4436:. New York: American Museum of Natural History, Man and Nature Publications, 14. p.  3149: 3128: 3071: 2886: 2834: 2710: 2092: 2010: 1975:, the juvenile has been identified as around only one year old when it died. Referred to 1285: 1000: 894: 890: 851: 775: 761: 5663: 5571: 5461: 5076: 5012: 4887: 4823: 4681: 4648: 4570: 4403: 4303: 3009: 2505: 2250: 2205: 7933: 7544: 6129: 5716: 5682: 5647: 5591: 5544: 5523: 5481: 5473: 5443:"Acoustic Analysis of Vocalization of Lambeosaurine Dinosaurs (Reptilia: Ornithischia)" 5429: 5123: 5084: 5028: 4907: 4899: 4731: 4698: 4615: 4582: 4415: 4315: 4087: 4060: 3693: 3685: 3096: 3040: 2954: 2805: 2170: 985: 815: 811: 420: 165: 7979: 4831: 4800: 2420:. His hypothesis was seemingly supported by skin preserved above the neck and back of 7940: 7679: 7600: 7560: 6444: 6330: 6276: 6247: 5720: 5687: 5583: 5393: 5236: 5202: 5100: 5049: 4974: 4924: 4859: 4785: 4736: 4586: 4543: 4512: 4489: 4441: 4372: 4353: 4334: 4092: 3728: 3718: 2939: 2433: 2363: 2076: 1036: 445: 5485: 4911: 4619: 4419: 3697: 3662: 1952: 8081: 8074: 7884: 7877: 7406: 7317: 7155: 7056: 7042: 6756: 6724: 6359: 6268: 6182: 6117: 5840: 5708: 5677: 5667: 5595: 5575: 5513: 5465: 5279: 5275: 5182: 5080: 5016: 4891: 4855: 4827: 4726: 4716: 4685: 4652: 4607: 4574: 4470: 4466: 4407: 4319: 4307: 4082: 4072: 3677: 3191: 3145: 2727: 1972: 1933: 1269: 764: 693: 678: 588: 475: 302: 5186: 5020: 2224:
The proposed air reservoir would have been insufficient for an animal the size of
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in 1961. It includes a partial skull with a short, rounded crest and much of the
585: 48: 4431: 2050: 8176: 8171: 8109: 8046: 7919: 7768: 7731: 7713: 7658: 7290: 7106: 7099: 7028: 6950: 6937: 6865: 6853: 6822: 6797: 6661: 6583: 6544: 6530: 6407: 6318: 6240: 6218: 6211: 5952: 5253:(1935). "Hooded hadrosaurs of the Belly River Series of the Upper Cretaceous". 5063:
Maryanska, T.; Osmólska, H. (1979). "Aspects of hadrosaurian cranial anatomy".
3154: 3101: 3086: 2598: 2588: 2582: 2375: 1917: 1347: 754: 633:, has been proposed, although it is more commonly placed in the separate genus 289: 247: 5469: 4895: 4411: 2362:
functions have become more supported as function(s) of the crest, focusing on
145: 8347: 8262: 8116: 8060: 8039: 8010: 7926: 7863: 7761: 7754: 7740: 7650: 7630: 7623: 7520: 7470: 7463: 7447: 7422: 7395: 7388: 7381: 7352: 7303: 7265: 7230: 7211: 7183: 7127: 6790: 6783: 6738: 6668: 6537: 6396: 6381: 6343: 6254: 6147: 6110: 6103: 5852: 3732: 3663:"A new method to calculate allometric length-mass relationships of dinosaurs" 3178: 3106: 3091: 2919: 2782: 2778: 2773: 2757: 2698: 2681: 2644: 2572: 2557: 2548: 2403: 2268: 2204:. Still, other proposals made housing specialized organs the major function. 2058:
Many hypotheses have been advanced as to what functions the cranial crest of
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Wheeler, P.E. (1978). "Elaborate CNS cooling structure in large dinosaurs".
4371:. Bloomington and Indianapolis: Indiana University Press. pp. 135–169. 1261:, a lambeosaurine with a similar skull (but no complete crest yet) from the 8024: 8000: 7988: 7954: 7947: 7898: 7891: 7697: 7641: 7616: 7588: 7477: 7279: 7244: 7190: 7120: 7092: 6978: 6971: 6811: 6776: 6703: 6696: 6618: 6559: 6514: 6416: 6290: 6283: 6232: 6196: 6154: 6004: 5943: 5765: 5691: 5371:
Fossil Record 3. New Mexico Museum of Natural History and Science, Bulletin
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Canada Department of Mines Geological Survey Bulletin (Geological Series)
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likely permitted both species identification (such as separating it from
2320: 2149: 1924:
with a sophisticated skull that permitted a grinding motion analogous to
1449: 1362: 1221: 996: 862: 674: 560: 392: 263: 65: 1087: 964:, along with a straighter crest and different internal structuring than 8304: 8155: 8130: 8123: 8095: 8088: 7968: 7854: 7553: 7533: 7491: 7366: 7218: 7085: 7035: 7000: 6993: 6964: 6804: 6682: 6654: 6523: 6496: 6471: 6311: 6304: 6297: 6175: 5527: 5477: 5433: 5409:
At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah
5127: 5046:
Dinosaurs Without Bones: Dinosaur Lives Revealed by Their Trace Fossils
4903: 4311: 3689: 3174: 3162: 3044: 2935: 2821: 2800:
Existing slightly later than the species from the Fruitland Formation,
2718: 2660: 2648: 2613: 2592: 2379: 2340: 2336: 2209: 2003:
adults bore such large crests, especially when compared to the related
1936:, lambeosaurines have narrower beaks than hadrosaurines, implying that 1913: 1736: 1262: 1184: 1180: 874: 827: 659:
is in fact part of the genus. The genus was first described in 1922 by
644: 596: 373: 110: 75: 31: 4721: 4077: 2071:
As for other lambeosaurines, it is believed that the cranial crest of
1132:
and is robust for its length when compared to other hadrosaurids. The
854:
of this skull, Robert Sullivan and Thomas Williamson gave the genus a
8137: 8053: 8032: 7870: 7567: 7505: 7014: 6986: 6846: 6710: 6640: 6611: 6437: 6423: 6166: 5579: 4923:(Second ed.). University of California Press. pp. 438–463. 3712: 3195: 3052: 2989: 2825: 2656: 2469: 2217: 2201: 2174: 2031: 2027: 889:, meaning "crested". The specimen was found in either the top of the 855: 758: 600: 592: 577: 182: 115: 59: 52: 8233: 5518: 5501: 4656: 4578: 3940: 2220:
of the lambeosaurines, which had no obvious defensive capabilities.
8256: 8017: 7686: 7574: 7374: 7345: 7272: 7197: 7162: 7063: 7049: 7007: 6831: 6632: 6597: 6503: 6489: 6451: 5961: 5934: 5925: 5811: 3078: 3051:, a large inland sea that split North America into two landmasses, 2531:
of prehistoric animals, including well-known dinosaurs such as the
2262: 2193: 2189:
that it served as an air reservoir for prolonged stays underwater.
1148: 1048: 743: 573: 215: 202: 105: 100: 85: 80: 70: 4481:
Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed
1195:, where some tubes were blind and others met and separated. While 1167: 1159:, showing uniform tubercle-like scales, but no larger structures. 8291: 7310: 6745: 3247:
Colbert, Edwin H. (Edwin Harris); Knight, Charles Robert (1951).
2640: 2632: 2125:
and the facial profile of juvenile individuals differed from the
2023: 2019: 1925: 1133: 1121: 823: 796: 640: 604: 120: 95: 27:
Hadrosaurid ornithopod dinosaur genus from Late Cretaceous Period
5764:. Sandia National Laboratories. December 5, 1997. Archived from 5502:"Dinosaurian Cacophony: Inferring function in extinct organisms" 4994:, a giant horned dinosaur from the Late Campanian of New Mexico" 4290:(1924). "Die neuen Dinosaurierfunde in der Oberkreide Canadas". 3962: 3960: 3413: 3411: 3409: 3407: 3405: 3403: 3401: 3399: 3397: 3395: 3382: 2808:. Numerous vertebrate groups are from this formation, including 2693: 2108:. Recent restudy of a juvenile braincase previously assigned to 7176: 6769: 5919: 5799: 4594:
Evans, D.C.; Reisz, R.R. (2007). "Anatomy and Relationships of
4120: 4118: 3813: 3496: 3494: 3380: 3378: 3376: 3374: 3372: 3370: 3368: 3366: 3364: 3362: 3187: 3170: 3166: 2982:. Unidentified taxa are known, including the crurotarsan ? 2829: 1137: 192: 5870: 4973:(Abridged ed.). United Kingdom: Oxford University Press. 3987: 3758: 2212:, and John Ostrom suggested that it housed expanded areas for 484: 7021: 6551: 4707: 4158: 4156: 4040: 3957: 3392: 3326: 3324: 3322: 3320: 3318: 2636: 2621: 2617: 2609: 2528: 2324: 2246: 1929: 1921: 1266: 1125: 878: 835: 733: 683: 664: 565: 493: 4143: 4141: 4139: 4137: 4135: 4133: 4115: 3782: 3557: 3555: 3553: 3551: 3538: 3536: 3534: 3532: 3491: 3359: 2315:
was a resonating chamber. Weishampel in 1981 suggested that
1979:
sp., the juvenile is the most complete, as well as youngest
529: 6389: 5713:
10.1671/0272-4634(2005)025[0897:ANOTMF]2.0.CO;2
4058: 3682:
10.1671/0272-4634(2001)021[0051:ANMTCA]2.0.CO;2
3644: 3642: 3640: 3638: 3636: 3567: 3158: 3153:. Paleofauna present in the Kaiparowits Formation included 3060: 2856:. Ornithischians from the formation are represented by the 2809: 2652: 2182: 1306:
The following cladogram is after the 2007 redescription of
914: 648: 581: 547: 541: 532: 514: 508: 487: 4949:
New Mexico Museum of Natural History and Science, Bulletin
4433:
The Dinosaur Book: The Ruling Reptiles and their Relatives
4204: 4180: 4153: 4103: 3519: 3517: 3515: 3513: 3511: 3509: 3315: 3268: 3266: 3264: 3262: 3260: 3250:
The dinosaur book: the ruling reptiles and their relatives
2307:
are much more variable and complicated than the airway of
502: 5618:
New Mexico Museum of Natural History and Science Bulletin
5340:
New Mexico Museum of Natural History and Science Bulletin
5300:
New Mexico Museum of Natural History and Science Bulletin
5216:"Scientists Use Digital Paleontology to Produce Voice of 4612:
10.1671/0272-4634(2007)27[373:AAROLM]2.0.CO;2
4130: 4018: 4016: 4014: 3748: 3746: 3744: 3742: 3592: 3590: 3588: 3586: 3584: 3582: 3548: 3529: 3464: 3462: 3460: 3458: 3456: 2173:, Martin Wilfarth that it was an attachment for a mobile 2014:
Reconstruction of a juvenile skeleton, based on RAM 14000
1971:
in 2009. Excavated by the joint expedition by museum and
1052:
in 2014 found that one more species could be referred to
5138:(1962). "The cranial crests of hadrosaurian dinosaurs". 4264: 4252: 4192: 3633: 3349: 3347: 3345: 3343: 3341: 3339: 5603:
Wilfarth, Martin (1947). "Rüsseltragende Dinosaurier".
5255:
Canada Department of Mines Bulletin (Geological Series)
4228: 3621: 3506: 3278: 3257: 814:.R1250) from what is now known as the slightly younger 4011: 3739: 3579: 3453: 3443: 3441: 2181:
that it served as an airtrap to keep water out of the
2104:, that probably grew faster as individuals approached 5824: 5324:
Late Cretaceous vertebrates from the Western Interior
4633:(Ornithischia: Lambeosaurinae) from southern Alberta" 4531:
Miscellaneous Publication 02-4, UtahGeological Survey
4369:
Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs
3977: 3975: 3905: 3852: 3850: 3848: 3611: 3609: 3607: 3605: 3481: 3479: 3477: 3336: 2492:
believed to have caused or contributed to its death.
591:, about 76.9–73.5 million years ago. It was a large 550: 544: 526: 523: 517: 511: 490: 478: 4812:
Comptes Rendus de l'Académie des Sciences, Série IIA
4168: 3893: 3871: 3869: 3867: 3865: 3428: 3426: 2235: 933:
Holotype skulls of the three species arranged by age
834:, in 1931. The specific epithet is derived from the 538: 505: 499: 481: 402: 383: 364: 341: 5635:
Nova Acta Regiae Societatis Scientiarum Upsaliensis
4627:Evans, D.C.; Bavington, R.; Campione, N.E. (2009). 4240: 3999: 3917: 3835: 3833: 3831: 3770: 3438: 3305: 3303: 3301: 3299: 3297: 3295: 3293: 2952:. Turtles are fairly plentiful, and are known from 1265:region of northeastern China. The two may form the 999:. Its three known specimens have been found in the 535: 496: 5286: 5265: 4799:Godefroit, Pascal; Shuqin Zan; Liyong Jin (2000). 4484:. Bloomington: Indiana University Press. pp.  4366: 4028: 3993: 3972: 3845: 3794: 3764: 3602: 3474: 3417: 2346:Computer modeling of a well-preserved specimen of 2026:, possibly when it was resting on the bottom of a 2022:. However, the skull was split down the middle by 558:; meaning "beside crested lizard" in reference to 5534: 5214:Sandia National Laboratories (December 5, 1997). 5157:, a new genus and species of trachodont dinosaur" 5062: 4703:(Hadrosauridae) and heterochrony in hadrosaurids" 4478:Currie, Phillip J.; Koppelhus, Eva, eds. (2005). 4452: 4216: 4046: 3966: 3881: 3862: 3423: 3386: 3228: 2483:Analysis of the pathology undertaken by Bertozzo 2468:is known from one specimen which might contain a 1276:, with its short, rounded crest, may be the most 842:, meaning "trumpeter". A second, nearly complete 8345: 5415: 5235:(5 ed.). London: Cassell Ltd. p. 883. 5161:University of Toronto Studies: Geological Series 4937: 4798: 4124: 3945: 3934: 3932: 3828: 3788: 3500: 3290: 2832:. The fishes are represented by the two species 2527:, was a member of a diverse and well-documented 2177:used as a breathing tube or for food gathering, 1207:had a short crest with a more circular profile. 901:described a partial skull with a similar crest ( 850:P-25100) was found in New Mexico in 1995. Using 5330: 4839:Hone, D.W.E.; Naish, D.; Cuthill, I.C. (2011). 4477: 3573: 3330: 2395:) and sexual identification by shape and size. 2350:, with more complex air passages than those of 2144: 2087:, with its small crest, was the female form of 1219:was initially thought to be closely related to 1023:, which were all thought to be males, although 778:, the chairman of the Board of Trustees of the 5976: 5381: 5357: 5222:. Sandia National Laboratories. Archived from 5171: 5099:. New York: Crescent Books. pp. 122–127. 5035:. Geological Society of America Special Paper 4385: 4210: 4186: 4162: 4109: 2054:Diagram showing internal features of the crest 6948: 5886: 5317: 5039:. Geological Society of America. p. 229. 4918: 4784:. McFarland & Company. pp. 678–940. 4755:, with a description of the type specimen of 4347: 4147: 3929: 3561: 3542: 3246: 2735:; and some unidentified fossils belonging to 2685:, they are predominate members of the fauna. 2635:was warmer than present-day Alberta, without 1995:began sooner than in related genera, such as 757:in Alberta. These rocks are now known as the 5698: 5549:: CS1 maint: multiple names: authors list ( 5201:. University of Chicago Press. p. 491. 4959: 4696: 4663: 4626: 4537: 4270: 4258: 4198: 3648: 3284: 3272: 2358:and other hadrosaurids. Instead, social and 1075:, then the genus therefore lasted until the 5645: 5411:. Indiana University Press. pp. 1–634. 5406: 4838: 4234: 3717:. Princeton University Press. p. 341. 3627: 3523: 1956:Juvenile skeleton RAM 14000 (nicknamed Joe) 1255:. Its closest known relative appears to be 412:(Godefroit, Zan & Jin, 2000) Paul, 2010 8354:Late Cretaceous dinosaurs of North America 6083: 5893: 5879: 5615: 5499: 5440: 5027: 4668:is a troodontid (Dinosauria: Theropoda)". 4593: 4022: 3752: 3596: 3468: 3353: 1967:(RAM). The juvenile was discovered in the 1128:measures 103 cm (41 in) long in 144: 5681: 5671: 5517: 5382:Tanke, D.H.; Carpenter, K., eds. (2001). 5322:. In Lucas, S.G.; Sullivan, R.M. (eds.). 5287:Sullivan, R.S.; Williamson, T.E. (1999). 5249: 4730: 4720: 4086: 4076: 3911: 3660: 2153:Comparison drawing between the crests of 2091:. Thomas Williamson suggested it was the 1107:is incompletely known. The length of the 5633:, n. sp. aus der Kreide in New Mexico". 5602: 5416:Weishampel, D.B.; Jensen, J.A. (1979). " 5407:Titus, A.L.; Loewen, M.A., eds. (2013). 4987: 4174: 3899: 2692: 2504: 2453: 2402: 2261: 2257: 2148: 2100:probably had small, rounded crests like 2066: 2049: 2009: 1951: 1891: 1294: 1166: 1086: 928: 822:, New Mexico. This specimen was sent to 716:Meaning "near crested lizard", the name 703: 8419:Multispecific non-avian dinosaur genera 5785: 5772: 5735: 5557: 5230: 5033:Hadrosaurian Dinosaurs of North America 4747: 4699:"Ontogeny in the tube-crested dinosaur 4502: 4426: 4246: 4005: 3923: 3776: 3447: 1203:had long crests with slight curvature, 712:, showing the pathologic v-shaped notch 699: 667:and partial skeleton found in Alberta. 14: 8346: 5420:(Reptilia: Hadrosauridae) from Utah". 5213: 5134: 5113: 5091: 5043: 4873: 4386:Carr, T.D.; Williamson, T.E. (2010). " 4326: 4034: 3981: 3856: 3800: 3714:The Princeton Field Guide to Dinosaurs 3615: 3485: 3234: 984:come from the De-Na-Zin Member of the 8238: 8237: 7842: 6935: 6070: 5912: 5874: 5744:. About.Com Dinosaurs. Archived from 5628: 5388:. Indiana University Press. pp.  5358:Sullivan, R.M.; Fowler, D.W. (2011). 5193: 5150: 4556: 4527: 4222: 3887: 3875: 3432: 3309: 3253:. New York: McGraw-Hill. p. 152. 2639:, but with wetter and drier seasons. 1965:Raymond M. Alf Museum of Paleontology 1103:Like most dinosaurs, the skeleton of 940:is known from three certain species: 7843: 5318:Sullivan, R.M.; Lucas, S.G. (2006). 4771: 4286: 3951: 3839: 3710: 2804:is also found in New Mexico, in the 2398: 2137:model, in part because the crest of 5794:. Scott Hartman's Skeletal Drawing. 5781:. Scott Hartman's Skeletal Drawing. 4352:. Dorling Kindersley. p. 342. 2428:. Subsequently, reconstructions of 2378:in the eyes imply acute vision and 1903: 1071:. If this species is indeed inside 770:. William Parks named the specimen 24: 6936: 5701:Journal of Vertebrate Paleontology 5268:Journal of Vertebrate Paleontology 5085:10.1111/j.1502-3931.1979.tb01006.x 4670:Canadian Journal of Earth Sciences 4637:Canadian Journal of Earth Sciences 4600:Journal of Vertebrate Paleontology 4459:Journal of Vertebrate Paleontology 4392:Journal of Vertebrate Paleontology 3670:Journal of Vertebrate Paleontology 1079:and is known from two continents. 1063:was actually nested deeply inside 830:described it as a second species, 753:in 1920 near Sand Creek along the 629:. Additionally, a fourth species, 25: 8435: 8404:Campanian genus first appearances 5729: 5539:(2nd ed.). pp. 517–606. 3941:https://doi.org/10.1111/joa.13363 2449: 2236:Temperature regulation hypothesis 1210: 1056:. This study, led by Xing, found 865:described another good specimen ( 7825: 7819: 7814: 7808: 7801: 6918: 6911: 6904: 5858: 5846: 5834: 5810: 5798: 4860:10.1111/j.1502-3931.2011.00300.x 3017: 3008: 2946:. One pterosaur is known, named 2319:made noises ranging between the 2045: 1067:, which created the new species 474: 169: 63: 8414:Maastrichtian genus extinctions 8182:List of informal dinosaur names 6071: 5900: 4540:Vertebrate Paleontology in Utah 4333:. William Morrow. p. 194. 4052: 3806: 3704: 3654: 2893:Nodocephalosaurus kirtlandensis 2495: 1887: 1140:bones were also heavily built. 158:Field Museum of Natural History 8192:Timeline of hadrosaur research 5777:Parasaurolophus cyrtocristatus 5280:10.1080/02724634.1996.10011371 4471:10.1080/02724634.1998.10011116 3994:Sullivan & Williamson 1996 3765:Brett-Surman & Wagner 2006 3418:Sullivan & Williamson 1999 3240: 3212:Timeline of hadrosaur research 3074:with other dinosaurs, such as 3041:Argon-argon radiometric dating 1082: 975:, which is the largest of the 437:Godefroit, Zan & Jin, 2000 13: 1: 8364:Fossil taxa described in 1922 5187:10.1016/j.cretres.2005.01.002 5048:. Pegasus Books. p. 42. 5021:10.1016/j.cretres.2010.12.007 4832:10.1016/S1251-8050(00)00214-7 4047:Diegert & Williamson 1998 3967:Maryanska & Osmolska 1979 3217: 2688: 2643:were apparently the dominant 2063:its internal nasal passages. 1999:. It has been suggested that 905:2467) from the Campanian age 8424:Ornithopods of North America 5673:10.1371/journal.pone.0098821 4780:. In Glut, Donald F. (ed.). 4666:"Saurornitholestes" robustus 4292:Jahrbuch Naturwissenschaften 4279: 4125:Jasinski & Sullivan 2011 3501:Weishampel & Jensen 1979 3222: 2908:; and the pachycephalosaurs 2624:conditions over time as the 2411:with hypothetical skin frill 2145:Rejected function hypotheses 1284:species or it may represent 991:In 1961, the third species, 728:/παρα ("beside" or "near"), 607:are universally recognized: 7: 8369:Taxa named by William Parks 5095:(1985). "Hadrosaurids II". 4782:Dinosaurs: The Encyclopedia 4596:Lambeosaurus magnicristatus 3331:Currie & Koppelhus 2005 3205: 2918:. Saurischians include the 2853:Denazinosuchus kirtlandicus 2844:. The crurotarsans include 2752:Saurornitholestes langstoni 2461:with notch in the vertebrae 2240:The large surface area and 2075:changed with age and was a 1899:head with scalation detail. 1309:Lambeosaurus magnicristatus 869:P27393) from New Mexico as 810:recovered a partial skull ( 134:Possible record during the 10: 8440: 8389:Paleontology in New Mexico 5233:Cassell's Latin Dictionary 4187:Carr & Williamson 2010 4163:Sullivan & Fowler 2011 4110:Tanke & Carpenter 2001 2795:Cretaceous Interior Seaway 2500: 2079:characteristic in adults. 1229:subfamily of hadrosaurids— 1155:impressions are known for 971:The next named species is 924: 885:, meaning "shortened" and 799:, a rock unit of the late 734: 576:that lived in what is now 29: 8246: 8147: 7998: 7978: 7853: 7849: 7838: 7799: 7730: 7696: 7640: 7599: 7543: 7528: 7519: 7446: 7405: 7298: 7289: 7239: 7228: 6959: 6944: 6931: 6902: 6864: 6821: 6764: 6755: 6592: 6581: 6513: 6484: 6480: 6469: 6406: 6370: 6338: 6329: 6165: 6128: 6098: 6094: 6079: 6066: 6030: 6012: 6003: 5985: 5972: 5908: 5500:Weishampel, D.B. (1997). 5470:10.1017/S0094837300004036 5441:Weishampel, D.B. (1981). 5346:: 405–417. Archived from 4896:10.1017/S0094837300002165 4803:Charonosaurus jiayinensis 4412:10.1080/02724630903413032 4148:Sullivan & Lucas 2006 3711:Paul, Gregory S. (2016). 3141:Kosmoceratops richardsoni 3120:Gryposaurus monumentensis 2899:Pentaceratops sternbergii 2868:Naashoibitosaurus ostromi 2723:Pentaceratops sternbergii 2208:suggested that it housed 1947: 1779: 1762: 1755: 1733: 1716: 1709: 1678: 1661: 1654: 1642: 1620: 1613: 1564: 1547: 1540: 1518: 1511: 1487: 1480: 1468: 1446: 1439: 1417: 1410: 1388: 1381: 1359: 1352: 1332: 1325: 1312:(Evans and Reisz, 2007): 1031:is about 72% the size of 639:. Remains are known from 426: 419: 361: 356: 340: 333: 166:Scientific classification 164: 152: 143: 41: 5637:. Series 4 (in German). 5385:Mesozoic Vertebrate Life 4259:Zanno & Sampson 2005 3285:Liddell & Scott 1980 3031:tentatively assigned to 2910:Stegoceras novomexicanum 2732:Stegoceras novomexicanum 2169:proposed it served as a 1766:Hypacrosaurus altispinus 1162: 742:/λοφος ("crest"). It is 623:, and the short-crested 30:Not to be confused with 8384:Paleontology in Alberta 5790:Parasaurolophus walkeri 5786:Hartman, Scott (2013). 5773:Hartman, Scott (2004). 5762:"Parasaurolophus sound" 5631:Parasaurolophus tubicen 5422:Journal of Paleontology 5199:Vertebrate Paleontology 5155:Parasaurolophus walkeri 5116:Journal of Paleontology 5031:; Wright, N.E. (1942). 4992:Titanoceratops ouranous 4988:Longrich, N.R. (2011). 4970:A Greek-English Lexicon 4455:Parasaurolophus tubicen 4235:Titus & Loewen 2013 3190:were present including 3049:Western Interior Seaway 2999: 2626:Western Interior Seaway 2525:Dinosaur Park Formation 2521:Parasaurolophus walkeri 2514:Dinosaur Park Formation 2476:restore the skin flap. 1288:or female specimens of 776:Sir Byron Edmund Walker 768:Dinosaur Park Formation 346:Parasaurolophus walkeri 18:Parasaurolophus walkeri 5306:: 1–52. Archived from 5231:Simpson, D.P. (1979). 5195:Romer, Alfred Sherwood 4945:Texascephale langstoni 4943:and a reassessment of 4751:(1924). "On the genus 4690:10.1139/cjes-2014-0073 4388:Bistahieversor sealeyi 3753:Lull & Wright 1942 3661:Seebacher, F. (2001). 3387:Weishampel et al. 2004 3354:Evans & Reisz 2007 2979:Thescelus hemispherica 2924:Bistahieversor sealeyi 2915:Sphaerotholus goodwini 2905:Titanoceratops ouranos 2874:Kritosaurus navajovius 2841:Myledalphus bipartitus 2706: 2517: 2462: 2412: 2284: 2162: 2055: 2015: 1957: 1900: 1303: 1191:, and more complex in 1176: 1100: 1046:A study published in 934: 713: 651:, as well as possibly 156:skeletal mount at the 8399:Kaiparowits Formation 8326:Paleobiology Database 8162:Basturs Poble bonebed 5736:Strauss, Bob (2014). 5362:Navajodactylus boerei 5251:Sternberg, Charles M. 5044:Martin, A.J. (2014). 4961:Liddell, Henry George 4327:Bakker, R.T. (1986). 3789:Godefroit et al. 2000 3186:. A variety of early 3135:Nasutoceratops titusi 3112:armored ankylosaurids 3029:Kaiparowits Formation 2949:Navajodactylus boerei 2862:Anasazisaurus horneri 2847:Brachychampsa montana 2696: 2508: 2457: 2406: 2265: 2258:Behavioral hypotheses 2167:Alfred Sherwood Romer 2152: 2067:Differences in crests 2053: 2013: 1969:Kaiparowits Formation 1955: 1895: 1298: 1215:As its name implies, 1170: 1090: 932: 907:Kaiparowits Formation 751:University of Toronto 708:Holotype specimen of 707: 686:in its largest form. 578:western North America 8409:Ornithischian genera 8394:Paleontology in Utah 8184:(contains hadrosaur 7582:Probrachylophosaurus 5807:at Wikimedia Commons 5151:Parks, W.A. (1922). 3816:. Raymond Alf Museum 3574:Sullivan et al. 2011 3116:duckbilled hadrosaur 2973:Plastomenus robustus 2374:and the presence of 2179:Charles M. Sternberg 808:Charles H. Sternberg 793:Hell Creek Formation 780:Royal Ontario Museum 720:is derived from the 700:Discovery and naming 677:, part of a diverse 8379:Dinosaur Park fauna 5768:on October 6, 2014. 5664:2014PLoSO...998821X 5629:Wiman, C. (1931). " 5572:1978Natur.275..441W 5495:on October 6, 2014. 5462:1981Pbio....7..252W 5353:on October 6, 2014. 5226:on August 17, 2014. 5175:Cretaceous Research 5077:1979Letha..12..265M 5013:2011CrRes..32..264L 5001:Cretaceous Research 4951:. Fossil Record 3. 4888:1975Pbio....1...21H 4869:on October 6, 2014. 4824:2000CRASE.330..875G 4772:Glut, D.F. (1997). 4757:Lambeosaurus lambei 4749:Gilmore, Charles W. 4682:2014CaJES..51..730E 4649:2009CaJES..46..791E 4571:2006Pbio...32..109E 4404:2010JVPal..30....1C 4304:1924NW.....12..709A 4211:Roberts et al. 2005 3157:(sharks and rays), 3150:Hagryphus giganteus 3129:Utahceratops gettyi 2896:; the ceratopsians 2887:Ahshislepelta minor 2835:Melvius chauliodous 2711:Fruitland Formation 2561:; fellow duckbills 1665:Lambeosaurus lambei 1280:of the three known 1095:(left, violet) and 1091:Size comparison of 895:David B. Weishampel 891:Fruitland Formation 852:computed tomography 7934:Ornithopodichnites 7545:Brachylophosaurini 6130:Hypsilophodontidae 4515:on August 28, 2013 4509:September 18, 1996 4503:Clinton, William. 4312:10.1007/BF01504818 4065:Journal of Anatomy 3814:"Joe the Dinosaur" 3562:Horner et al. 2004 3543:Benson et al. 2012 3097:Ornithomimus velox 2967:Neurankylus baueri 2955:Denazinemys nodosa 2806:Kirtland Formation 2707: 2620:and influenced by 2518: 2463: 2413: 2285: 2216:and much improved 2163: 2112:, now assigned to 2077:sexually dimorphic 2056: 2016: 1958: 1920:herbivore, eating 1908:As a hadrosaurid, 1901: 1304: 1177: 1101: 986:Kirtland Formation 935: 921:crest morphology. 816:Kirtland Formation 714: 8341: 8340: 8240:Taxon identifiers 8231: 8230: 8227: 8226: 8223: 8222: 7941:Shiraminesauropus 7834: 7833: 7797: 7796: 7793: 7792: 7789: 7788: 7785: 7784: 7680:Naashoibitosaurus 7601:Austrokritosauria 7561:Brachylophosaurus 7515: 7514: 6927: 6926: 6900: 6899: 6896: 6895: 6892: 6891: 6888: 6887: 6884: 6883: 6577: 6576: 6465: 6464: 6461: 6460: 6445:Transylvanosaurus 6331:Rhabdodontomorpha 6277:Notohypsilophodon 6248:Macrogryphosaurus 6062: 6061: 6058: 6057: 6054: 6053: 6050: 6049: 5803:Media related to 5788:"Ornithischians: 5775:"Ornithischians: 5738:"Ten Facts About 5566:(5679): 441–443. 5399:978-0-253-33907-2 5313:on April 8, 2023. 5242:978-0-304-52257-6 5106:978-0-517-46890-6 5055:978-1-60598-499-5 4980:978-0-19-910207-5 4930:978-0-520-24209-8 4791:978-0-899-50917-4 4722:10.7717/peerj.182 4549:978-1-55791-634-1 4495:978-0-253-34595-0 4428:Colbert, Edwin H. 4378:978-0-253-34817-3 4359:978-0-7566-9910-9 4340:978-0-8217-2859-8 4271:Eaton et al. 1999 4199:Evans et al. 2014 4078:10.1111/joa.13363 3724:978-1-78684-190-2 3649:Farke et al. 2013 3273:Evans et al. 2009 3192:multituberculates 3146:oviraptorosaurian 3033:P. cyrtocristatus 2961:Basilemys nobilis 2940:Saurornitholestes 2715:P. cyrtocristatus 2704:P. cyrtocristatus 2616:that became more 2434:Charles R. Knight 2399:Soft tissue frill 2329:P. cyrtocristatus 2282:P. cyrtocristatus 2155:P. cyrtocristatus 2102:P. cyrtocristatus 2085:P. cyrtocristatus 1884: 1883: 1875: 1874: 1866: 1865: 1857: 1856: 1848: 1847: 1839: 1838: 1830: 1829: 1821: 1820: 1812: 1811: 1803: 1802: 1794: 1793: 1693: 1692: 1682:L. magnicristatus 1597: 1596: 1588: 1587: 1579: 1578: 1522:P. cyrtocristatus 1274:P. cyrtocristatus 1205:P. cyrtocristatus 1093:P. cyrtocristatus 1041:P. cyrtocristatus 1037:sexual dimorphism 1029:P. cyrtocristatus 1013:P. cyrtocristatus 993:P. cyrtocristatus 966:P. cyrtocristatus 950:P. cyrtocristatus 919:P. cyrtocristatus 871:P. cyrtocristatus 626:P. cyrtocristatus 465: 464: 459: 438: 413: 399: 388:P. cyrtocristatus 380: 329: 154:P. cyrtocristatus 138: 16:(Redirected from 8431: 8334: 8333: 8321: 8320: 8308: 8307: 8295: 8294: 8282: 8281: 8280: 8267: 8266: 8265: 8235: 8234: 8208:mummy S.M. R4036 8082:Microhadrosaurus 8075:Mandschurosaurus 7885:Camptosaurichnus 7878:Bonaparteichnium 7851: 7850: 7844:Related articles 7840: 7839: 7829: 7823: 7818: 7813: 7812: 7805: 7526: 7525: 7407:Parasaurolophini 7318:Angulomastacator 7296: 7295: 7237: 7236: 7156:Shuangmiaosaurus 7057:Huehuecanauhtlus 7043:Gongpoquansaurus 6957: 6956: 6946: 6945: 6933: 6932: 6922: 6916: 6915: 6908: 6762: 6761: 6757:Hadrosauriformes 6725:Ratchasimasaurus 6590: 6589: 6482: 6481: 6478: 6477: 6360:Muttaburrasaurus 6336: 6335: 6269:Muttaburrasaurus 6183:Atlascopcosaurus 6118:Thescelosauridae 6096: 6095: 6092: 6091: 6081: 6080: 6068: 6067: 6010: 6009: 5983: 5982: 5974: 5973: 5967: 5966: 5910: 5909: 5895: 5888: 5881: 5872: 5871: 5863: 5862: 5861: 5851: 5850: 5849: 5839: 5838: 5837: 5830: 5815:Data related to 5814: 5802: 5795: 5782: 5769: 5757: 5755: 5753: 5748:on April 7, 2015 5724: 5695: 5685: 5675: 5642: 5625: 5612: 5599: 5580:10.1038/275441a0 5554: 5548: 5540: 5531: 5521: 5496: 5494: 5488:. Archived from 5447: 5437: 5428:(6): 1422–1427. 5412: 5403: 5378: 5368: 5354: 5352: 5337: 5327: 5314: 5312: 5297: 5289:"A new skull of 5283: 5262: 5246: 5227: 5210: 5190: 5168: 5147: 5131: 5110: 5093:Norman, David B. 5088: 5059: 5040: 5024: 4998: 4984: 4956: 4934: 4915: 4870: 4868: 4862:. Archived from 4845: 4835: 4809: 4795: 4768: 4744: 4734: 4724: 4693: 4660: 4623: 4590: 4553: 4534: 4524: 4522: 4520: 4511:. Archived from 4499: 4474: 4449: 4423: 4382: 4363: 4350:Prehistoric Life 4344: 4323: 4274: 4268: 4262: 4256: 4250: 4244: 4238: 4232: 4226: 4220: 4214: 4208: 4202: 4196: 4190: 4184: 4178: 4172: 4166: 4160: 4151: 4145: 4128: 4122: 4113: 4107: 4101: 4100: 4090: 4080: 4071:(5): 1055–1069. 4056: 4050: 4044: 4038: 4032: 4026: 4020: 4009: 4003: 3997: 3991: 3985: 3979: 3970: 3964: 3955: 3949: 3943: 3936: 3927: 3921: 3915: 3909: 3903: 3897: 3891: 3885: 3879: 3873: 3860: 3854: 3843: 3837: 3826: 3825: 3823: 3821: 3810: 3804: 3798: 3792: 3786: 3780: 3774: 3768: 3762: 3756: 3750: 3737: 3736: 3708: 3702: 3701: 3667: 3658: 3652: 3646: 3631: 3628:Xing et al. 2014 3625: 3619: 3613: 3600: 3594: 3577: 3571: 3565: 3559: 3546: 3540: 3527: 3524:Hone et al. 2011 3521: 3504: 3498: 3489: 3483: 3472: 3466: 3451: 3445: 3436: 3430: 3421: 3415: 3390: 3384: 3357: 3351: 3334: 3328: 3313: 3307: 3288: 3282: 3276: 3270: 3255: 3254: 3244: 3238: 3232: 3072:paleoenvironment 3055:to the west and 3021: 3012: 2728:pachycephalosaur 2647:plants, with an 2251:Teresa Maryańska 2230:olfactory nerves 2214:olfactory tissue 2206:Halszka Osmólska 1973:The Webb Schools 1904:Diet and feeding 1758: 1757: 1752: 1712: 1711: 1706: 1657: 1656: 1651: 1645: 1644: 1639: 1616: 1615: 1610: 1543: 1542: 1537: 1514: 1513: 1483: 1482: 1477: 1471: 1470: 1465: 1442: 1441: 1436: 1413: 1412: 1407: 1384: 1383: 1378: 1355: 1354: 1328: 1327: 1316: 1315: 1270:Parasaurolophini 791:material in the 738:("lizard"), and 737: 736: 694:thermoregulation 557: 556: 553: 552: 549: 546: 543: 540: 537: 534: 531: 528: 525: 520: 519: 516: 513: 510: 507: 504: 501: 498: 495: 492: 489: 486: 483: 480: 444: 436: 411: 404: 391: 385: 372: 366: 343: 321: 314: 303:Parasaurolophini 301: 288: 275: 262: 246: 230: 174: 173: 148: 133: 132: 62: 47:Temporal range: 39: 38: 21: 8439: 8438: 8434: 8433: 8432: 8430: 8429: 8428: 8344: 8343: 8342: 8337: 8329: 8324: 8316: 8311: 8303: 8298: 8290: 8285: 8278:Parasaurolophus 8276: 8275: 8270: 8261: 8260: 8255: 8248:Parasaurolophus 8242: 8232: 8219: 8200:mummy AMNH 5060 8167:Dakota (fossil) 8143: 7994: 7974: 7962:Tetrapodosaurus 7913:Hadrosaurichnus 7845: 7830: 7807: 7781: 7776:Shantungosaurus 7726: 7707:Augustynolophus 7692: 7636: 7610:Bonapartesaurus 7595: 7539: 7511: 7442: 7430:Parasaurolophus 7401: 7285: 7233: 7224: 7149:Qianjiangsaurus 7135:Probactrosaurus 7114:Penelopognathus 6953: 6940: 6923: 6910: 6880: 6860: 6817: 6751: 6732:Riabininohadros 6605:Bayannurosaurus 6586: 6573: 6509: 6474: 6457: 6402: 6372:Tenontosauridae 6366: 6325: 6161: 6140:Gideonmantellia 6124: 6088: 6075: 6046: 6026: 5999: 5993:Avemetatarsalia 5978:Avemetatarsalia 5968: 5915: 5914: 5904: 5899: 5869: 5859: 5857: 5847: 5845: 5835: 5833: 5825: 5817:Parasaurolophus 5805:Parasaurolophus 5760: 5751: 5749: 5740:Parasaurolophus 5732: 5727: 5542: 5541: 5519:10.2307/1313034 5492: 5445: 5418:Parasaurolophus 5400: 5366: 5350: 5335: 5310: 5295: 5291:Parasaurolophus 5243: 5218:Parasaurolophus 5136:Ostrom, John H. 5107: 5056: 4996: 4981: 4931: 4866: 4843: 4818:(12): 875–882. 4807: 4792: 4776:Parasaurolophus 4701:Parasaurolophus 4657:10.1139/E09-050 4643:(11): 791–800. 4631:Parasaurolophus 4579:10.1666/04027.1 4550: 4518: 4516: 4496: 4379: 4360: 4341: 4298:(36): 709–716. 4282: 4277: 4269: 4265: 4257: 4253: 4245: 4241: 4233: 4229: 4221: 4217: 4209: 4205: 4197: 4193: 4185: 4181: 4173: 4169: 4161: 4154: 4146: 4131: 4123: 4116: 4108: 4104: 4057: 4053: 4045: 4041: 4033: 4029: 4023:Weishampel 1997 4021: 4012: 4004: 4000: 3992: 3988: 3980: 3973: 3965: 3958: 3950: 3946: 3937: 3930: 3922: 3918: 3910: 3906: 3898: 3894: 3886: 3882: 3874: 3863: 3855: 3846: 3838: 3829: 3819: 3817: 3812: 3811: 3807: 3799: 3795: 3787: 3783: 3775: 3771: 3763: 3759: 3751: 3740: 3725: 3709: 3705: 3665: 3659: 3655: 3647: 3634: 3626: 3622: 3614: 3603: 3597:Weishampel 1981 3595: 3580: 3572: 3568: 3560: 3549: 3541: 3530: 3522: 3507: 3499: 3492: 3484: 3475: 3469:Williamson 2000 3467: 3454: 3446: 3439: 3431: 3424: 3416: 3393: 3385: 3360: 3352: 3337: 3329: 3316: 3308: 3291: 3283: 3279: 3271: 3258: 3245: 3241: 3233: 3229: 3225: 3220: 3208: 3155:chondrichthyans 3068:Parasaurolophus 3038: 3037: 3036: 3035: 3027:Skull from the 3024: 3023: 3022: 3014: 3013: 3002: 2791:Parasaurolophus 2737:Tyrannosauridae 2713:of New Mexico, 2691: 2672:Parasaurolophus 2668:Parasaurolophus 2503: 2498: 2474:Parasaurolophus 2452: 2440:animated film, 2430:Parasaurolophus 2418:basilisk lizard 2407:Restoration of 2401: 2385:Parasaurolophus 2376:sclerotic rings 2356:Parasaurolophus 2317:Parasaurolophus 2313:Parasaurolophus 2309:Parasaurolophus 2288:Parasaurolophus 2266:Restoration of 2260: 2242:vascularization 2238: 2226:Parasaurolophus 2147: 2139:Parasaurolophus 2123:Parasaurolophus 2114:Parasaurolophus 2106:sexual maturity 2098:Parasaurolophus 2073:Parasaurolophus 2069: 2060:Parasaurolophus 2048: 2039:Parasaurolophus 2001:Parasaurolophus 1993:Parasaurolophus 1989:Parasaurolophus 1985:Parasaurolophus 1981:Parasaurolophus 1977:Parasaurolophus 1961:Parasaurolophus 1950: 1942:Parasaurolophus 1938:Parasaurolophus 1910:Parasaurolophus 1906: 1890: 1885: 1876: 1867: 1858: 1849: 1840: 1831: 1822: 1813: 1804: 1795: 1753: 1750: 1707: 1704: 1694: 1652: 1649: 1640: 1637: 1611: 1608: 1598: 1589: 1580: 1538: 1535: 1506:Parasaurolophus 1478: 1475: 1466: 1463: 1437: 1434: 1408: 1405: 1379: 1376: 1299:Restoration of 1282:Parasaurolophus 1217:Parasaurolophus 1213: 1165: 1105:Parasaurolophus 1085: 1077:K-Pg extinction 1073:Parasaurolophus 1065:Parasaurolophus 1054:Parasaurolophus 1009:Parasaurolophus 977:Parasaurolophus 938:Parasaurolophus 927: 911:Garfield County 899:James A. Jensen 820:San Juan County 784:Parasaurolophus 765:Late Cretaceous 718:Parasaurolophus 702: 671:Parasaurolophus 586:Late Cretaceous 522: 477: 473: 469:Parasaurolophus 442:Paralophosaurus 435: 352: 349: 320: 317:Parasaurolophus 312: 299: 286: 273: 260: 244: 228: 168: 139: 131: 130: 129: 128: 123: 118: 113: 108: 103: 98: 93: 88: 83: 78: 73: 68: 58:76.9–73.5  57: 56: 49:Late Cretaceous 45: 43:Parasaurolophus 35: 28: 23: 22: 15: 12: 11: 5: 8437: 8427: 8426: 8421: 8416: 8411: 8406: 8401: 8396: 8391: 8386: 8381: 8376: 8374:Kirtland fauna 8371: 8366: 8361: 8359:Lambeosaurines 8356: 8339: 8338: 8336: 8335: 8322: 8309: 8296: 8283: 8268: 8252: 8250: 8244: 8243: 8229: 8228: 8225: 8224: 8221: 8220: 8218: 8217: 8210: 8202: 8194: 8189: 8179: 8177:Hypsilophodont 8174: 8172:Hadrosaur diet 8169: 8164: 8159: 8151: 8149: 8148:Other articles 8145: 8144: 8142: 8141: 8134: 8127: 8120: 8113: 8110:Stephanosaurus 8106: 8099: 8092: 8085: 8078: 8071: 8064: 8057: 8050: 8047:Fulgurotherium 8043: 8036: 8029: 8021: 8014: 8006: 8004: 7996: 7995: 7993: 7992: 7984: 7982: 7976: 7975: 7973: 7972: 7965: 7958: 7951: 7944: 7937: 7930: 7923: 7920:Iguanodontipus 7916: 7909: 7902: 7895: 7888: 7881: 7874: 7867: 7859: 7857: 7847: 7846: 7836: 7835: 7832: 7831: 7800: 7798: 7795: 7794: 7791: 7790: 7787: 7786: 7783: 7782: 7780: 7779: 7772: 7769:Laiyangosaurus 7765: 7758: 7751: 7744: 7736: 7734: 7732:Edmontosaurini 7728: 7727: 7725: 7724: 7717: 7714:Prosaurolophus 7710: 7702: 7700: 7694: 7693: 7691: 7690: 7683: 7676: 7669: 7662: 7659:Coahuilasaurus 7655: 7646: 7644: 7638: 7637: 7635: 7634: 7627: 7620: 7613: 7605: 7603: 7597: 7596: 7594: 7593: 7585: 7578: 7571: 7564: 7557: 7549: 7547: 7541: 7540: 7538: 7537: 7529: 7523: 7517: 7516: 7513: 7512: 7510: 7509: 7502: 7495: 7488: 7481: 7474: 7467: 7460: 7452: 7450: 7444: 7443: 7441: 7440: 7433: 7426: 7419: 7411: 7409: 7403: 7402: 7400: 7399: 7392: 7385: 7378: 7371: 7363: 7356: 7349: 7342: 7335: 7328: 7321: 7314: 7307: 7299: 7293: 7291:Lambeosaurinae 7287: 7286: 7284: 7283: 7276: 7269: 7262: 7255: 7248: 7240: 7234: 7229: 7226: 7225: 7223: 7222: 7215: 7208: 7201: 7194: 7187: 7180: 7173: 7166: 7159: 7152: 7145: 7138: 7131: 7124: 7117: 7110: 7107:Nanyangosaurus 7103: 7100:Nanningosaurus 7096: 7089: 7082: 7074: 7067: 7060: 7053: 7046: 7039: 7032: 7029:Gilmoreosaurus 7025: 7018: 7011: 7004: 6997: 6990: 6983: 6975: 6968: 6960: 6954: 6951:Hadrosauroidea 6949: 6942: 6941: 6938:Hadrosauroidea 6929: 6928: 6925: 6924: 6903: 6901: 6898: 6897: 6894: 6893: 6890: 6889: 6886: 6885: 6882: 6881: 6879: 6878: 6877: 6876: 6870: 6868: 6866:Hadrosauroidea 6862: 6861: 6859: 6858: 6854:Mantellisaurus 6850: 6843: 6836: 6827: 6825: 6823:Iguanodontidae 6819: 6818: 6816: 6815: 6808: 6801: 6798:Mantellisaurus 6794: 6787: 6780: 6773: 6765: 6759: 6753: 6752: 6750: 6749: 6742: 6735: 6728: 6721: 6714: 6707: 6700: 6693: 6686: 6679: 6672: 6665: 6662:Iguanacolossus 6658: 6651: 6644: 6637: 6629: 6622: 6615: 6608: 6601: 6593: 6587: 6584:Ankylopollexia 6582: 6579: 6578: 6575: 6574: 6572: 6571: 6564: 6556: 6548: 6545:Eousdryosaurus 6541: 6534: 6531:Dysalotosaurus 6527: 6519: 6517: 6511: 6510: 6508: 6507: 6500: 6493: 6485: 6475: 6470: 6467: 6466: 6463: 6462: 6459: 6458: 6456: 6455: 6448: 6441: 6434: 6427: 6420: 6412: 6410: 6408:Rhabdodontidae 6404: 6403: 6401: 6400: 6393: 6386: 6377: 6375: 6368: 6367: 6365: 6364: 6356: 6348: 6339: 6333: 6327: 6326: 6324: 6323: 6319:Weewarrasaurus 6315: 6308: 6301: 6294: 6287: 6280: 6273: 6265: 6258: 6251: 6244: 6241:Leaellynasaura 6237: 6229: 6222: 6219:Gasparinisaura 6215: 6212:Galleonosaurus 6208: 6200: 6193: 6186: 6179: 6171: 6169: 6163: 6162: 6160: 6159: 6151: 6144: 6135: 6133: 6126: 6125: 6123: 6122: 6114: 6107: 6099: 6089: 6084: 6077: 6076: 6064: 6063: 6060: 6059: 6056: 6055: 6052: 6051: 6048: 6047: 6045: 6044: 6043: 6042: 6036: 6034: 6028: 6027: 6025: 6024: 6023: 6022: 6013: 6007: 6001: 6000: 5998: 5997: 5996: 5995: 5986: 5980: 5970: 5969: 5965: 5964: 5955: 5953:Neornithischia 5946: 5937: 5928: 5922: 5906: 5905: 5898: 5897: 5890: 5883: 5875: 5868: 5867: 5855: 5843: 5821: 5820: 5819:at Wikispecies 5808: 5796: 5783: 5770: 5758: 5731: 5730:External links 5728: 5726: 5725: 5707:(4): 897–904. 5696: 5643: 5626: 5613: 5605:Orion (Munich) 5600: 5555: 5537:The Dinosauria 5532: 5512:(3): 150–155. 5497: 5456:(2): 252–261. 5438: 5413: 5404: 5398: 5379: 5355: 5328: 5315: 5284: 5263: 5247: 5241: 5228: 5211: 5191: 5181:(2): 307–318. 5169: 5148: 5132: 5122:(3): 575–577. 5111: 5105: 5089: 5071:(3): 265–273. 5060: 5054: 5041: 5025: 5007:(3): 264–276. 4985: 4979: 4957: 4935: 4929: 4921:The Dinosauria 4916: 4871: 4854:(2): 139–156. 4836: 4796: 4790: 4769: 4753:Stephanosaurus 4745: 4694: 4676:(7): 730–734. 4661: 4624: 4606:(2): 373–393. 4591: 4565:(1): 109–125. 4554: 4548: 4535: 4525: 4500: 4494: 4475: 4450: 4424: 4383: 4377: 4364: 4358: 4345: 4339: 4324: 4283: 4281: 4278: 4276: 4275: 4263: 4251: 4239: 4227: 4215: 4203: 4191: 4179: 4167: 4152: 4129: 4114: 4102: 4051: 4039: 4027: 4010: 3998: 3986: 3971: 3956: 3944: 3928: 3916: 3912:Sternberg 1935 3904: 3892: 3880: 3861: 3844: 3827: 3805: 3793: 3781: 3769: 3757: 3738: 3723: 3703: 3653: 3632: 3620: 3601: 3578: 3566: 3547: 3528: 3505: 3490: 3473: 3452: 3437: 3422: 3391: 3358: 3335: 3314: 3289: 3277: 3256: 3239: 3226: 3224: 3221: 3219: 3216: 3215: 3214: 3207: 3204: 3102:tyrannosaurids 3087:Talos sampsoni 3026: 3025: 3016: 3015: 3007: 3006: 3005: 3004: 3003: 3001: 2998: 2818:ornithischians 2690: 2687: 2631:westward. The 2599:Euoplocephalus 2583:Daspletosaurus 2573:tyrannosaurids 2502: 2499: 2497: 2494: 2451: 2450:Paleopathology 2448: 2400: 2397: 2259: 2256: 2237: 2234: 2218:sense of smell 2146: 2143: 2068: 2065: 2047: 2044: 1949: 1946: 1905: 1902: 1889: 1886: 1882: 1881: 1878: 1877: 1873: 1872: 1869: 1868: 1864: 1863: 1860: 1859: 1855: 1854: 1851: 1850: 1846: 1845: 1842: 1841: 1837: 1836: 1833: 1832: 1828: 1827: 1824: 1823: 1819: 1818: 1815: 1814: 1810: 1809: 1806: 1805: 1801: 1800: 1797: 1796: 1792: 1791: 1788: 1787: 1778: 1775: 1774: 1771: 1770: 1761: 1756: 1754: 1749: 1746: 1745: 1742: 1741: 1732: 1729: 1728: 1725: 1724: 1715: 1710: 1708: 1703: 1700: 1699: 1696: 1695: 1691: 1690: 1687: 1686: 1677: 1674: 1673: 1670: 1669: 1660: 1655: 1653: 1648: 1643: 1641: 1636: 1633: 1632: 1629: 1628: 1619: 1614: 1612: 1607: 1604: 1603: 1600: 1599: 1595: 1594: 1591: 1590: 1586: 1585: 1582: 1581: 1577: 1576: 1573: 1572: 1563: 1560: 1559: 1556: 1555: 1546: 1541: 1539: 1534: 1531: 1530: 1527: 1526: 1517: 1512: 1510: 1500: 1499: 1496: 1495: 1486: 1481: 1479: 1474: 1469: 1467: 1462: 1459: 1458: 1455: 1454: 1445: 1440: 1438: 1433: 1430: 1429: 1426: 1425: 1416: 1411: 1409: 1404: 1401: 1400: 1397: 1396: 1387: 1382: 1380: 1375: 1372: 1371: 1368: 1367: 1358: 1353: 1351: 1348:Lambeosaurinae 1343: 1342: 1339: 1338: 1331: 1326: 1324: 1314: 1212: 1211:Classification 1209: 1175:beak and teeth 1164: 1161: 1122:shoulder blade 1084: 1081: 926: 923: 755:Red Deer River 701: 698: 631:P. jiayinensis 572:"duck-billed" 463: 462: 461: 460: 439: 424: 423: 417: 416: 415: 414: 407:P. jiayinensis 400: 381: 359: 358: 357:Other species 354: 353: 350: 338: 337: 331: 330: 310: 306: 305: 297: 293: 292: 290:Lambeosaurinae 284: 280: 279: 271: 267: 266: 258: 251: 250: 248:Neornithischia 242: 235: 234: 226: 219: 218: 213: 206: 205: 200: 196: 195: 190: 186: 185: 180: 176: 175: 162: 161: 150: 149: 141: 140: 126: 125: 124: 119: 114: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 46: 26: 9: 6: 4: 3: 2: 8436: 8425: 8422: 8420: 8417: 8415: 8412: 8410: 8407: 8405: 8402: 8400: 8397: 8395: 8392: 8390: 8387: 8385: 8382: 8380: 8377: 8375: 8372: 8370: 8367: 8365: 8362: 8360: 8357: 8355: 8352: 8351: 8349: 8332: 8327: 8323: 8319: 8314: 8310: 8306: 8301: 8297: 8293: 8288: 8284: 8279: 8273: 8269: 8264: 8258: 8254: 8253: 8251: 8249: 8245: 8241: 8236: 8216: 8215: 8211: 8209: 8207: 8206:Edmontosaurus 8203: 8201: 8199: 8198:Edmontosaurus 8195: 8193: 8190: 8187: 8183: 8180: 8178: 8175: 8173: 8170: 8168: 8165: 8163: 8160: 8158: 8157: 8153: 8152: 8150: 8146: 8140: 8139: 8135: 8133: 8132: 8128: 8126: 8125: 8121: 8119: 8118: 8117:Syngonosaurus 8114: 8112: 8111: 8107: 8105: 8104: 8100: 8098: 8097: 8093: 8091: 8090: 8086: 8084: 8083: 8079: 8077: 8076: 8072: 8070: 8069: 8065: 8063: 8062: 8061:Koutalisaurus 8058: 8056: 8055: 8051: 8049: 8048: 8044: 8042: 8041: 8040:Eucercosaurus 8037: 8035: 8034: 8030: 8027: 8026: 8022: 8020: 8019: 8015: 8013: 8012: 8011:Arstanosaurus 8008: 8007: 8005: 8003: 8002: 7997: 7991: 7990: 7986: 7985: 7983: 7981: 7977: 7971: 7970: 7966: 7964: 7963: 7959: 7957: 7956: 7952: 7950: 7949: 7945: 7943: 7942: 7938: 7936: 7935: 7931: 7929: 7928: 7927:Orcauichnites 7924: 7922: 7921: 7917: 7915: 7914: 7910: 7908: 7907: 7903: 7901: 7900: 7896: 7894: 7893: 7889: 7887: 7886: 7882: 7880: 7879: 7875: 7873: 7872: 7868: 7866: 7865: 7864:Amblydactylus 7861: 7860: 7858: 7856: 7852: 7848: 7841: 7837: 7828: 7824: 7822: 7817: 7811: 7804: 7778: 7777: 7773: 7771: 7770: 7766: 7764: 7763: 7762:Kundurosaurus 7759: 7757: 7756: 7755:Kerberosaurus 7752: 7750: 7749: 7745: 7743: 7742: 7741:Edmontosaurus 7738: 7737: 7735: 7733: 7729: 7723: 7722: 7718: 7716: 7715: 7711: 7709: 7708: 7704: 7703: 7701: 7699: 7695: 7689: 7688: 7684: 7682: 7681: 7677: 7675: 7674: 7670: 7668: 7667: 7663: 7661: 7660: 7656: 7653: 7652: 7651:Anasazisaurus 7648: 7647: 7645: 7643: 7639: 7633: 7632: 7631:Secernosaurus 7628: 7626: 7625: 7624:Kelumapusaura 7621: 7619: 7618: 7614: 7612: 7611: 7607: 7606: 7604: 7602: 7598: 7591: 7590: 7586: 7584: 7583: 7579: 7577: 7576: 7572: 7570: 7569: 7565: 7563: 7562: 7558: 7556: 7555: 7551: 7550: 7548: 7546: 7542: 7536: 7535: 7531: 7530: 7527: 7524: 7522: 7521:Saurolophinae 7518: 7508: 7507: 7503: 7501: 7500: 7496: 7494: 7493: 7489: 7487: 7486: 7482: 7480: 7479: 7475: 7473: 7472: 7471:Hypacrosaurus 7468: 7466: 7465: 7464:Corythosaurus 7461: 7459: 7458: 7454: 7453: 7451: 7449: 7448:Lambeosaurini 7445: 7439: 7438: 7434: 7432: 7431: 7427: 7425: 7424: 7423:Charonosaurus 7420: 7418: 7417: 7413: 7412: 7410: 7408: 7404: 7398: 7397: 7396:Tsintaosaurus 7393: 7391: 7390: 7389:Pararhabdodon 7386: 7384: 7383: 7382:Nipponosaurus 7379: 7377: 7376: 7372: 7369: 7368: 7364: 7362: 7361: 7357: 7355: 7354: 7353:Jaxartosaurus 7350: 7348: 7347: 7343: 7341: 7340: 7336: 7334: 7333: 7329: 7327: 7326: 7322: 7320: 7319: 7315: 7313: 7312: 7308: 7306: 7305: 7304:Adynomosaurus 7301: 7300: 7297: 7294: 7292: 7288: 7282: 7281: 7277: 7275: 7274: 7270: 7268: 7267: 7266:Lapampasaurus 7263: 7261: 7260: 7256: 7254: 7253: 7249: 7247: 7246: 7242: 7241: 7238: 7235: 7232: 7231:Hadrosauridae 7227: 7221: 7220: 7216: 7214: 7213: 7212:Zhanghenglong 7209: 7207: 7206: 7202: 7200: 7199: 7195: 7193: 7192: 7188: 7186: 7185: 7184:Telmatosaurus 7181: 7179: 7178: 7174: 7172: 7171: 7167: 7165: 7164: 7160: 7158: 7157: 7153: 7151: 7150: 7146: 7144: 7143: 7139: 7137: 7136: 7132: 7130: 7129: 7128:Portellsaurus 7125: 7123: 7122: 7118: 7116: 7115: 7111: 7109: 7108: 7104: 7102: 7101: 7097: 7095: 7094: 7090: 7088: 7087: 7083: 7080: 7079: 7075: 7073: 7072: 7068: 7066: 7065: 7061: 7059: 7058: 7054: 7052: 7051: 7047: 7045: 7044: 7040: 7038: 7037: 7033: 7031: 7030: 7026: 7024: 7023: 7019: 7017: 7016: 7012: 7010: 7009: 7005: 7003: 7002: 6998: 6996: 6995: 6991: 6989: 6988: 6984: 6981: 6980: 6976: 6974: 6973: 6969: 6967: 6966: 6962: 6961: 6958: 6955: 6952: 6947: 6943: 6939: 6934: 6930: 6921: 6917: 6914: 6907: 6874: 6873: 6872: 6871: 6869: 6867: 6863: 6856: 6855: 6851: 6849: 6848: 6844: 6842: 6841: 6837: 6834: 6833: 6829: 6828: 6826: 6824: 6820: 6814: 6813: 6809: 6807: 6806: 6802: 6800: 6799: 6795: 6793: 6792: 6791:Jinzhousaurus 6788: 6786: 6785: 6784:Hypselospinus 6781: 6779: 6778: 6774: 6772: 6771: 6767: 6766: 6763: 6760: 6758: 6754: 6748: 6747: 6743: 6741: 6740: 6739:Theiophytalia 6736: 6734: 6733: 6729: 6727: 6726: 6722: 6720: 6719: 6715: 6713: 6712: 6708: 6706: 6705: 6701: 6699: 6698: 6694: 6692: 6691: 6687: 6685: 6684: 6680: 6678: 6677: 6673: 6671: 6670: 6669:Lanzhousaurus 6666: 6664: 6663: 6659: 6657: 6656: 6652: 6650: 6649: 6645: 6643: 6642: 6638: 6635: 6634: 6630: 6628: 6627: 6623: 6621: 6620: 6616: 6614: 6613: 6609: 6607: 6606: 6602: 6600: 6599: 6595: 6594: 6591: 6588: 6585: 6580: 6570: 6569: 6565: 6562: 6561: 6557: 6554: 6553: 6549: 6547: 6546: 6542: 6540: 6539: 6538:Elrhazosaurus 6535: 6533: 6532: 6528: 6526: 6525: 6521: 6520: 6518: 6516: 6512: 6506: 6505: 6501: 6499: 6498: 6494: 6492: 6491: 6487: 6486: 6483: 6479: 6476: 6473: 6468: 6454: 6453: 6449: 6447: 6446: 6442: 6440: 6439: 6435: 6433: 6432: 6428: 6426: 6425: 6421: 6419: 6418: 6414: 6413: 6411: 6409: 6405: 6399: 6398: 6397:Tenontosaurus 6394: 6392: 6391: 6387: 6384: 6383: 6382:Convolosaurus 6379: 6378: 6376: 6373: 6369: 6362: 6361: 6357: 6354: 6353: 6349: 6346: 6345: 6344:Ampelognathus 6341: 6340: 6337: 6334: 6332: 6328: 6321: 6320: 6316: 6314: 6313: 6309: 6307: 6306: 6302: 6300: 6299: 6295: 6293: 6292: 6288: 6286: 6285: 6281: 6279: 6278: 6274: 6271: 6270: 6266: 6264: 6263: 6259: 6257: 6256: 6255:Mahuidacursor 6252: 6250: 6249: 6245: 6243: 6242: 6238: 6235: 6234: 6230: 6228: 6227: 6223: 6221: 6220: 6216: 6214: 6213: 6209: 6206: 6205: 6201: 6199: 6198: 6194: 6192: 6191: 6187: 6185: 6184: 6180: 6178: 6177: 6173: 6172: 6170: 6168: 6164: 6157: 6156: 6152: 6150: 6149: 6148:Hypsilophodon 6145: 6142: 6141: 6137: 6136: 6134: 6131: 6127: 6120: 6119: 6115: 6113: 6112: 6111:Callovosaurus 6108: 6106: 6105: 6104:Burianosaurus 6101: 6100: 6097: 6093: 6090: 6087: 6082: 6078: 6074: 6069: 6065: 6040: 6039: 6038: 6037: 6035: 6033: 6029: 6021: 6017: 6016: 6015: 6014: 6011: 6008: 6006: 6002: 5994: 5990: 5989: 5988: 5987: 5984: 5981: 5979: 5975: 5971: 5963: 5959: 5956: 5954: 5950: 5947: 5945: 5941: 5938: 5936: 5932: 5929: 5927: 5923: 5921: 5917: 5916: 5911: 5907: 5903: 5896: 5891: 5889: 5884: 5882: 5877: 5876: 5873: 5866: 5865:United States 5856: 5854: 5844: 5842: 5832: 5831: 5828: 5823: 5818: 5813: 5809: 5806: 5801: 5797: 5793: 5791: 5784: 5780: 5778: 5771: 5767: 5763: 5759: 5747: 5743: 5741: 5734: 5733: 5722: 5718: 5714: 5710: 5706: 5702: 5697: 5693: 5689: 5684: 5679: 5674: 5669: 5665: 5661: 5658:(6): e98821. 5657: 5653: 5649: 5644: 5640: 5636: 5632: 5627: 5623: 5619: 5614: 5610: 5607:(in German). 5606: 5601: 5597: 5593: 5589: 5585: 5581: 5577: 5573: 5569: 5565: 5561: 5556: 5552: 5546: 5538: 5533: 5529: 5525: 5520: 5515: 5511: 5507: 5503: 5498: 5491: 5487: 5483: 5479: 5475: 5471: 5467: 5463: 5459: 5455: 5451: 5444: 5439: 5435: 5431: 5427: 5423: 5419: 5414: 5410: 5405: 5401: 5395: 5391: 5387: 5386: 5380: 5376: 5372: 5365: 5363: 5356: 5349: 5345: 5341: 5334: 5329: 5325: 5321: 5316: 5309: 5305: 5301: 5294: 5292: 5285: 5281: 5277: 5273: 5269: 5264: 5260: 5256: 5252: 5248: 5244: 5238: 5234: 5229: 5225: 5221: 5219: 5212: 5208: 5204: 5200: 5196: 5192: 5188: 5184: 5180: 5176: 5170: 5166: 5162: 5158: 5156: 5149: 5145: 5141: 5137: 5133: 5129: 5125: 5121: 5117: 5112: 5108: 5102: 5098: 5094: 5090: 5086: 5082: 5078: 5074: 5070: 5066: 5061: 5057: 5051: 5047: 5042: 5038: 5034: 5030: 5026: 5022: 5018: 5014: 5010: 5006: 5002: 4995: 4993: 4986: 4982: 4976: 4972: 4971: 4966: 4962: 4958: 4954: 4950: 4946: 4942: 4936: 4932: 4926: 4922: 4917: 4913: 4909: 4905: 4901: 4897: 4893: 4889: 4885: 4881: 4877: 4872: 4865: 4861: 4857: 4853: 4849: 4842: 4837: 4833: 4829: 4825: 4821: 4817: 4813: 4806: 4804: 4797: 4793: 4787: 4783: 4779: 4777: 4770: 4766: 4762: 4758: 4754: 4750: 4746: 4742: 4738: 4733: 4728: 4723: 4718: 4714: 4710: 4709: 4704: 4702: 4695: 4691: 4687: 4683: 4679: 4675: 4671: 4667: 4662: 4658: 4654: 4650: 4646: 4642: 4638: 4634: 4632: 4625: 4621: 4617: 4613: 4609: 4605: 4601: 4597: 4592: 4588: 4584: 4580: 4576: 4572: 4568: 4564: 4560: 4555: 4551: 4545: 4541: 4536: 4532: 4526: 4514: 4510: 4506: 4501: 4497: 4491: 4487: 4483: 4482: 4476: 4472: 4468: 4464: 4460: 4456: 4451: 4447: 4443: 4439: 4435: 4434: 4429: 4425: 4421: 4417: 4413: 4409: 4405: 4401: 4397: 4393: 4389: 4384: 4380: 4374: 4370: 4365: 4361: 4355: 4351: 4346: 4342: 4336: 4332: 4331: 4325: 4321: 4317: 4313: 4309: 4305: 4301: 4297: 4294:(in German). 4293: 4289: 4288:Abel, Othenio 4285: 4284: 4272: 4267: 4260: 4255: 4248: 4243: 4236: 4231: 4224: 4219: 4212: 4207: 4200: 4195: 4188: 4183: 4176: 4175:Longrich 2011 4171: 4164: 4159: 4157: 4149: 4144: 4142: 4140: 4138: 4136: 4134: 4126: 4121: 4119: 4111: 4106: 4098: 4094: 4089: 4084: 4079: 4074: 4070: 4066: 4062: 4055: 4048: 4043: 4036: 4031: 4024: 4019: 4017: 4015: 4007: 4002: 3995: 3990: 3983: 3978: 3976: 3968: 3963: 3961: 3953: 3948: 3942: 3935: 3933: 3925: 3920: 3913: 3908: 3901: 3900:Wilfarth 1947 3896: 3889: 3884: 3877: 3872: 3870: 3868: 3866: 3858: 3853: 3851: 3849: 3841: 3836: 3834: 3832: 3815: 3809: 3802: 3797: 3790: 3785: 3778: 3773: 3766: 3761: 3754: 3749: 3747: 3745: 3743: 3734: 3730: 3726: 3720: 3716: 3715: 3707: 3699: 3695: 3691: 3687: 3683: 3679: 3675: 3671: 3664: 3657: 3650: 3645: 3643: 3641: 3639: 3637: 3629: 3624: 3617: 3612: 3610: 3608: 3606: 3598: 3593: 3591: 3589: 3587: 3585: 3583: 3575: 3570: 3563: 3558: 3556: 3554: 3552: 3544: 3539: 3537: 3535: 3533: 3525: 3520: 3518: 3516: 3514: 3512: 3510: 3502: 3497: 3495: 3487: 3482: 3480: 3478: 3470: 3465: 3463: 3461: 3459: 3457: 3449: 3444: 3442: 3434: 3429: 3427: 3419: 3414: 3412: 3410: 3408: 3406: 3404: 3402: 3400: 3398: 3396: 3388: 3383: 3381: 3379: 3377: 3375: 3373: 3371: 3369: 3367: 3365: 3363: 3355: 3350: 3348: 3346: 3344: 3342: 3340: 3332: 3327: 3325: 3323: 3321: 3319: 3311: 3306: 3304: 3302: 3300: 3298: 3296: 3294: 3286: 3281: 3274: 3269: 3267: 3265: 3263: 3261: 3252: 3251: 3243: 3236: 3231: 3227: 3213: 3210: 3209: 3203: 3201: 3200:insectivorans 3197: 3193: 3189: 3185: 3184: 3180: 3179:apex predator 3176: 3172: 3168: 3164: 3160: 3156: 3152: 3151: 3147: 3143: 3142: 3137: 3136: 3131: 3130: 3126: 3122: 3121: 3117: 3113: 3109: 3108: 3107:Teratophoneus 3103: 3099: 3098: 3093: 3092:ornithomimids 3089: 3088: 3084: 3080: 3077: 3076:dromaeosaurid 3073: 3069: 3065: 3062: 3058: 3054: 3050: 3046: 3042: 3034: 3030: 3020: 3011: 2997: 2995: 2994:Struthiomimus 2991: 2987: 2986: 2981: 2980: 2975: 2974: 2969: 2968: 2963: 2962: 2957: 2956: 2951: 2950: 2945: 2941: 2937: 2934:sp.; and the 2933: 2930: 2926: 2925: 2921: 2920:tyrannosaurid 2917: 2916: 2911: 2907: 2906: 2901: 2900: 2895: 2894: 2889: 2888: 2884: 2883:ankylosaurids 2880: 2876: 2875: 2870: 2869: 2864: 2863: 2859: 2855: 2854: 2849: 2848: 2843: 2842: 2837: 2836: 2831: 2827: 2823: 2819: 2815: 2811: 2807: 2803: 2798: 2796: 2792: 2788: 2784: 2783:Hadrosauridae 2780: 2779:Hadrosaurinae 2776: 2775: 2774:Corythosaurus 2770: 2769: 2764: 2760: 2759: 2758:Struthiomimus 2754: 2753: 2748: 2744: 2743: 2738: 2734: 2733: 2729: 2725: 2724: 2720: 2716: 2712: 2705: 2701: 2700: 2699:Teratophoneus 2695: 2686: 2684: 2683: 2682:Pentaceratops 2677: 2673: 2669: 2664: 2662: 2658: 2654: 2650: 2646: 2642: 2638: 2634: 2630: 2627: 2623: 2619: 2615: 2611: 2607: 2606: 2601: 2600: 2595: 2594: 2590: 2586: 2584: 2579: 2578: 2574: 2570: 2569:Corythosaurus 2566: 2565: 2560: 2559: 2558:Struthiomimus 2555: 2554:ornithomimids 2551: 2550: 2549:Styracosaurus 2545: 2544: 2539: 2538: 2534: 2530: 2526: 2522: 2515: 2511: 2507: 2493: 2490: 2486: 2481: 2477: 2475: 2471: 2467: 2460: 2456: 2447: 2445: 2444: 2439: 2435: 2431: 2427: 2426:Edmontosaurus 2423: 2422:Corythosaurus 2419: 2410: 2405: 2396: 2394: 2390: 2389:Corythosaurus 2386: 2381: 2377: 2373: 2369: 2365: 2361: 2360:physiological 2357: 2353: 2349: 2344: 2342: 2338: 2334: 2333:Corythosaurus 2330: 2326: 2322: 2318: 2314: 2310: 2306: 2302: 2301:Corythosaurus 2298: 2297:Hypacrosaurus 2294: 2289: 2283: 2279: 2275: 2271: 2270: 2269:Charonosaurus 2264: 2255: 2252: 2248: 2243: 2233: 2231: 2227: 2221: 2219: 2215: 2211: 2207: 2203: 2199: 2195: 2190: 2188: 2184: 2180: 2176: 2172: 2168: 2160: 2156: 2151: 2142: 2140: 2136: 2132: 2131:Hypacrosaurus 2128: 2127:Corythosaurus 2124: 2119: 2118:frontal bones 2115: 2111: 2107: 2103: 2099: 2094: 2090: 2086: 2082: 2078: 2074: 2064: 2061: 2052: 2046:Cranial crest 2043: 2040: 2035: 2033: 2029: 2025: 2021: 2012: 2008: 2006: 2005:Corythosaurus 2002: 1998: 1997:Corythosaurus 1994: 1990: 1986: 1982: 1978: 1974: 1970: 1966: 1962: 1954: 1945: 1943: 1939: 1935: 1934:Robert Bakker 1931: 1927: 1923: 1919: 1915: 1911: 1898: 1894: 1880: 1879: 1871: 1870: 1862: 1861: 1853: 1852: 1844: 1843: 1835: 1834: 1826: 1825: 1817: 1816: 1808: 1807: 1799: 1798: 1790: 1789: 1786: 1785: 1784: 1783:H. stebingeri 1777: 1776: 1773: 1772: 1769: 1768: 1767: 1760: 1759: 1748: 1747: 1744: 1743: 1740: 1739: 1738: 1731: 1730: 1727: 1726: 1723: 1722: 1721: 1720:Corythosaurus 1714: 1713: 1702: 1701: 1698: 1697: 1689: 1688: 1685: 1684: 1683: 1676: 1675: 1672: 1671: 1668: 1667: 1666: 1659: 1658: 1647: 1646: 1635: 1634: 1631: 1630: 1627: 1626: 1625: 1624:Nipponosaurus 1618: 1617: 1606: 1605: 1602: 1601: 1593: 1592: 1584: 1583: 1575: 1574: 1571: 1570: 1569: 1562: 1561: 1558: 1557: 1554: 1553: 1552: 1545: 1544: 1533: 1532: 1529: 1528: 1525: 1524: 1523: 1516: 1515: 1508: 1507: 1502: 1501: 1498: 1497: 1494: 1493: 1492: 1491:Charonosaurus 1485: 1484: 1473: 1472: 1461: 1460: 1457: 1456: 1453: 1452: 1451: 1444: 1443: 1432: 1431: 1428: 1427: 1424: 1423: 1422: 1421:Jaxartosaurus 1415: 1414: 1403: 1402: 1399: 1398: 1395: 1394: 1393: 1392:Tsintaosaurus 1386: 1385: 1374: 1373: 1370: 1369: 1366: 1365: 1364: 1357: 1356: 1349: 1345: 1344: 1341: 1340: 1337: 1336: 1335:Hadrosaurinae 1330: 1329: 1322: 1321:Hadrosauridae 1318: 1317: 1313: 1311: 1310: 1302: 1297: 1293: 1291: 1287: 1283: 1279: 1275: 1271: 1268: 1264: 1260: 1259: 1258:Charonosaurus 1254: 1253: 1248: 1247: 1246:Hypacrosaurus 1242: 1241: 1240:Corythosaurus 1236: 1232: 1228: 1227:lambeosaurine 1224: 1223: 1218: 1208: 1206: 1202: 1198: 1194: 1190: 1186: 1182: 1174: 1169: 1160: 1158: 1154: 1150: 1146: 1145:neural spines 1141: 1139: 1135: 1131: 1127: 1123: 1119: 1114: 1110: 1109:type specimen 1106: 1099:(right, blue) 1098: 1094: 1089: 1080: 1078: 1074: 1070: 1066: 1062: 1060: 1059:Charonosaurus 1055: 1051: 1050: 1044: 1042: 1038: 1034: 1030: 1026: 1022: 1018: 1014: 1010: 1006: 1002: 998: 995:was named by 994: 989: 987: 983: 978: 974: 969: 967: 963: 959: 955: 951: 947: 943: 939: 931: 922: 920: 916: 912: 908: 904: 900: 896: 892: 888: 884: 880: 876: 872: 868: 864: 860: 857: 853: 849: 845: 841: 837: 833: 829: 825: 821: 817: 813: 809: 804: 802: 801:Maastrichtian 798: 794: 790: 785: 781: 777: 773: 769: 766: 763: 760: 756: 752: 748: 745: 741: 731: 727: 723: 719: 711: 706: 697: 695: 691: 690: 689:Charonosaurus 685: 680: 676: 672: 668: 666: 662: 661:William Parks 658: 657:Charonosaurus 654: 650: 646: 642: 638: 637: 636:Charonosaurus 632: 628: 627: 622: 621: 616: 612: 611: 606: 602: 598: 594: 590: 587: 583: 580:and possibly 579: 575: 571: 567: 563: 562: 555: 471: 470: 457: 456: 455:lapsus calami 451: 447: 443: 440: 433: 432: 431:Charonosaurus 428: 427: 425: 422: 418: 409: 408: 401: 398: 394: 390: 389: 382: 379: 375: 371: 370: 363: 362: 360: 355: 348: 347: 339: 336: 332: 328: 324: 319: 318: 311: 308: 307: 304: 298: 295: 294: 291: 285: 282: 281: 278: 277:Hadrosauridae 272: 269: 268: 265: 259: 256: 253: 252: 249: 243: 240: 237: 236: 233: 227: 224: 221: 220: 217: 214: 211: 208: 207: 204: 201: 198: 197: 194: 191: 188: 187: 184: 181: 178: 177: 172: 167: 163: 159: 155: 151: 147: 142: 137: 136:Maastrichtian 122: 117: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 61: 54: 50: 44: 40: 37: 33: 19: 8247: 8212: 8205: 8197: 8185: 8154: 8136: 8129: 8122: 8115: 8108: 8101: 8094: 8087: 8080: 8073: 8066: 8059: 8052: 8045: 8038: 8031: 8025:Claorhynchus 8023: 8016: 8009: 8001:Nomina dubia 7999: 7989:Guegoolithus 7987: 7967: 7960: 7955:Staurichnium 7953: 7948:Sousaichnium 7946: 7939: 7932: 7925: 7918: 7911: 7904: 7899:Dakotasaurus 7897: 7892:Caririchnium 7890: 7883: 7876: 7869: 7862: 7806: 7774: 7767: 7760: 7753: 7746: 7739: 7719: 7712: 7705: 7698:Saurolophini 7685: 7678: 7671: 7664: 7657: 7649: 7642:Kritosaurini 7629: 7622: 7617:Huallasaurus 7615: 7608: 7589:Wulagasaurus 7587: 7580: 7573: 7566: 7559: 7552: 7532: 7504: 7497: 7490: 7483: 7478:Lambeosaurus 7476: 7469: 7462: 7455: 7435: 7429: 7428: 7421: 7414: 7394: 7387: 7380: 7373: 7365: 7358: 7351: 7344: 7337: 7330: 7323: 7316: 7309: 7302: 7280:Yamatosaurus 7278: 7271: 7264: 7257: 7250: 7245:Aquilarhinus 7243: 7217: 7210: 7203: 7196: 7191:Tethyshadros 7189: 7182: 7175: 7168: 7161: 7154: 7147: 7140: 7133: 7126: 7121:Plesiohadros 7119: 7112: 7105: 7098: 7093:Lophorhothon 7091: 7084: 7076: 7069: 7062: 7055: 7048: 7041: 7034: 7027: 7020: 7013: 7006: 6999: 6992: 6985: 6979:Batyrosaurus 6977: 6972:Bactrosaurus 6970: 6963: 6909: 6852: 6845: 6838: 6830: 6812:Ouranosaurus 6810: 6803: 6796: 6789: 6782: 6777:Brighstoneus 6775: 6768: 6744: 6737: 6730: 6723: 6716: 6709: 6704:Osmakasaurus 6702: 6697:Oblitosaurus 6695: 6688: 6681: 6674: 6667: 6660: 6653: 6646: 6639: 6631: 6624: 6619:Camptosaurus 6617: 6610: 6603: 6596: 6566: 6560:Kangnasaurus 6558: 6550: 6543: 6536: 6529: 6522: 6515:Dryosauridae 6502: 6495: 6488: 6450: 6443: 6436: 6429: 6422: 6417:Matheronodon 6415: 6395: 6388: 6380: 6358: 6350: 6342: 6317: 6310: 6303: 6296: 6291:Sektensaurus 6289: 6284:Qantassaurus 6282: 6275: 6267: 6260: 6253: 6246: 6239: 6233:Kangnasaurus 6231: 6224: 6217: 6210: 6202: 6197:Diluvicursor 6195: 6188: 6181: 6174: 6155:Vectidromeus 6153: 6146: 6138: 6116: 6109: 6102: 6020:Ornithischia 6005:Ornithischia 5957: 5948: 5944:Ornithischia 5939: 5930: 5822: 5789: 5776: 5766:the original 5750:. Retrieved 5746:the original 5739: 5704: 5700: 5655: 5651: 5638: 5634: 5630: 5621: 5617: 5608: 5604: 5563: 5559: 5536: 5509: 5505: 5490:the original 5453: 5450:Paleobiology 5449: 5425: 5421: 5417: 5408: 5384: 5374: 5370: 5361: 5348:the original 5343: 5339: 5323: 5308:the original 5303: 5299: 5290: 5271: 5267: 5258: 5254: 5232: 5224:the original 5217: 5198: 5178: 5174: 5164: 5160: 5154: 5143: 5139: 5119: 5115: 5096: 5068: 5064: 5045: 5036: 5032: 5004: 5000: 4991: 4969: 4965:Robert Scott 4952: 4948: 4944: 4940: 4920: 4882:(1): 21–43. 4879: 4876:Paleobiology 4875: 4864:the original 4851: 4847: 4815: 4811: 4802: 4781: 4775: 4767:(43): 29–48. 4764: 4760: 4756: 4752: 4712: 4706: 4700: 4673: 4669: 4665: 4640: 4636: 4630: 4603: 4599: 4595: 4562: 4559:Paleobiology 4558: 4539: 4530: 4517:. Retrieved 4513:the original 4508: 4480: 4462: 4458: 4454: 4432: 4395: 4391: 4387: 4368: 4349: 4329: 4295: 4291: 4266: 4254: 4247:Clinton 1996 4242: 4230: 4218: 4206: 4194: 4182: 4170: 4105: 4068: 4064: 4054: 4042: 4030: 4006:Wheeler 1978 4001: 3989: 3947: 3924:Colbert 1945 3919: 3907: 3895: 3883: 3818:. Retrieved 3808: 3796: 3784: 3777:Gilmore 1924 3772: 3760: 3713: 3706: 3676:(1): 51–60. 3673: 3669: 3656: 3623: 3569: 3448:Simpson 1979 3280: 3249: 3242: 3230: 3181: 3175:crocodilians 3148: 3139: 3133: 3127: 3125:ceratopsians 3118: 3105: 3095: 3085: 3067: 3066: 3039: 3032: 2993: 2985:Leidyosuchus 2983: 2977: 2971: 2965: 2959: 2953: 2947: 2943: 2932:Ornithomimus 2931: 2929:ornithomimid 2922: 2913: 2909: 2903: 2897: 2891: 2885: 2878: 2872: 2866: 2860: 2858:hadrosaurids 2851: 2845: 2839: 2833: 2822:saurischians 2814:crurotarsans 2801: 2799: 2790: 2787:Ceratopsidae 2772: 2768:Chasmosaurus 2766: 2756: 2750: 2747:Troodontidae 2742:Ornithomimus 2740: 2730: 2721: 2714: 2708: 2703: 2702:attacking a 2697: 2680: 2675: 2671: 2667: 2665: 2629:transgressed 2605:Dyoplosaurus 2603: 2597: 2591: 2581: 2575: 2568: 2562: 2556: 2547: 2543:Chasmosaurus 2541: 2537:Centrosaurus 2535: 2520: 2519: 2509: 2496:Paleoecology 2488: 2484: 2482: 2478: 2473: 2465: 2464: 2458: 2441: 2429: 2425: 2421: 2414: 2408: 2393:Lambeosaurus 2392: 2388: 2384: 2355: 2351: 2347: 2345: 2339:to those of 2332: 2328: 2316: 2312: 2308: 2305:Lambeosaurus 2304: 2300: 2296: 2292: 2287: 2286: 2281: 2277: 2273: 2267: 2239: 2225: 2222: 2198:Othenio Abel 2191: 2164: 2158: 2157:(above) and 2154: 2138: 2135:Lambeosaurus 2134: 2130: 2126: 2122: 2113: 2110:Lambeosaurus 2109: 2101: 2097: 2088: 2084: 2081:James Hopson 2072: 2070: 2059: 2057: 2038: 2036: 2017: 2004: 2000: 1996: 1992: 1988: 1984: 1980: 1976: 1960: 1959: 1941: 1937: 1912:was a large 1909: 1907: 1896: 1888:Paleobiology 1781: 1780: 1764: 1763: 1735: 1734: 1718: 1717: 1680: 1679: 1663: 1662: 1622: 1621: 1567: 1566: 1565: 1550: 1549: 1548: 1521: 1520: 1519: 1505: 1504: 1489: 1488: 1448: 1447: 1419: 1418: 1390: 1389: 1361: 1360: 1333: 1307: 1305: 1300: 1289: 1281: 1273: 1256: 1252:Lambeosaurus 1250: 1244: 1238: 1235:hadrosaurine 1230: 1220: 1216: 1214: 1204: 1200: 1196: 1192: 1188: 1178: 1172: 1156: 1142: 1129: 1117: 1112: 1104: 1102: 1096: 1092: 1072: 1069:P. jiayensis 1068: 1064: 1057: 1053: 1047: 1045: 1040: 1032: 1028: 1024: 1020: 1016: 1012: 1008: 992: 990: 981: 976: 972: 970: 965: 961: 957: 949: 945: 941: 937: 936: 918: 886: 882: 870: 861: 843: 839: 831: 805: 788: 783: 774:in honor of 771: 739: 729: 725: 717: 715: 709: 687: 670: 669: 656: 653:Heilongjiang 634: 630: 625: 624: 619: 618: 615:type species 609: 608: 559: 468: 467: 466: 453: 441: 429: 405: 387: 386: 368: 367: 345: 344: 335:Type species 316: 315: 254: 238: 232:Ornithischia 222: 209: 153: 42: 36: 8272:Wikispecies 8214:Willinakaqe 8186:nomina nuda 8103:Siluosaurus 8068:Loncosaurus 7906:Dinehichnus 7748:Kamuysaurus 7721:Saurolophus 7673:Kritosaurus 7666:Gryposaurus 7499:Sahaliyania 7485:Magnapaulia 7457:Amurosaurus 7437:Tlatolophus 7416:Adelolophus 7360:Kazaklambia 7339:Blasisaurus 7332:Arenysaurus 7325:Aralosaurus 7259:Hadrosaurus 7252:Eotrachodon 7205:Yunganglong 7170:Sirindhorna 7142:Protohadros 7078:Koshisaurus 7071:Jintasaurus 6840:Comptonatus 6690:Napaisaurus 6676:Lurdusaurus 6648:Fukuisaurus 6626:Cedrorestes 6568:Valdosaurus 6431:Pareisactus 6262:Morrosaurus 6226:Isasicursor 6190:Chakisaurus 6086:Ornithopoda 6073:Ornithopoda 6032:Ornithopoda 5902:Ornithopoda 5274:(3): 1–68. 5261:(52): 1–37. 4519:November 9, 4398:(1): 1–16. 4035:Sandia 1997 3982:Ostrom 1962 3857:Norman 1985 3801:Bakker 1986 3616:Hopson 1975 3486:Ostrom 1961 3235:Martin 2014 3183:Deinosuchus 3163:salamanders 3070:shared its 2763:Ornithopoda 2719:ceratopsian 2676:Kritosaurus 2661:angiosperms 2614:floodplains 2577:Gorgosaurus 2564:Gryposaurus 2523:, from the 2516:environment 2438:Walt Disney 2436:and in the 2372:eye sockets 2323:55 and 720 2321:frequencies 2210:salt glands 2187:Ned Colbert 1918:quadrupedal 1450:Amurosaurus 1363:Aralosaurus 1231:Saurolophus 1222:Saurolophus 1185:nasal bones 1171:Closeup of 1083:Description 1005:Kaiparowits 997:John Ostrom 875:postcranial 863:John Ostrom 856:monographic 675:hadrosaurid 584:during the 570:hadrosaurid 561:Saurolophus 351:Parks, 1922 283:Subfamily: 264:Ornithopoda 8348:Categories 8156:Arkharavia 8131:Tichosteus 8124:Thespesius 8096:Pteropelyx 8089:Orthomerus 7969:Wintonopus 7554:Acristavus 7534:Barsboldia 7492:Olorotitan 7367:Latirhinus 7219:Zuoyunlong 7086:Levnesovia 7036:Gobihadros 7001:Datonglong 6994:Claosaurus 6965:Altirhinus 6875:see below↓ 6805:Morelladon 6683:Magnamanus 6655:Hippodraco 6524:Dryosaurus 6497:Hesperonyx 6472:Dryomorpha 6312:Trinisaura 6305:Tietasaura 6298:Talenkauen 6176:Anabisetia 6041:see below↓ 5935:Dinosauria 5752:October 5, 5641:(5): 1–11. 5624:: 191–213. 5611:: 525–532. 5506:BioScience 5377:: 393–404. 5029:Lull, R.S. 4955:: 202–215. 4941:Stegoceras 4759:, Parks". 4465:(3): 38A. 4223:Eaton 2002 3888:Romer 1933 3876:Evans 2006 3433:Wiman 1931 3310:Parks 1922 3218:References 3196:marsupials 3083:troodontid 3057:Appalachia 3045:Cretaceous 2988:, and the 2936:troodontid 2879:P. tubicen 2826:pterosaurs 2802:P. tubicen 2689:New Mexico 2657:tree ferns 2649:understory 2593:Edmontonia 2510:P. walkeri 2489:P. walkeri 2466:P. walkeri 2459:P. walkeri 2409:P. walkeri 2352:P. walkeri 2348:P. tubicen 2341:crocodiles 2337:inner ears 2293:P. tubicen 2278:P. walkeri 2274:P. tubicen 2159:P. walkeri 2089:P. tubicen 2034:in shape. 1897:P. walkeri 1737:Olorotitan 1568:P. walkeri 1551:P. tubicen 1301:P. walkeri 1290:P. tubicen 1201:P. tubicen 1197:P. walkeri 1193:P. tubicen 1189:P. walkeri 1181:premaxilla 1173:P. walkeri 1157:P. walkeri 1130:P. walkeri 1118:P. tubicen 1113:P. walkeri 1097:P. walkeri 1033:P. tubicen 1025:P. tubicen 1021:P. tubicen 1017:P. walkeri 982:P. tubicen 973:P. tubicen 962:P. tubicen 958:P. walkeri 946:P. tubicen 942:P. walkeri 844:P. tubicen 832:P. tubicen 828:Carl Wiman 789:P. walkeri 772:P. walkeri 710:P. walkeri 645:New Mexico 620:P. tubicen 610:P. walkeri 369:P. tubicen 216:Dinosauria 32:Parasaurus 8138:Trachodon 8054:Glishades 8033:Diclonius 7871:Anomoepus 7855:Ichnotaxa 7568:Maiasaura 7506:Velafrons 7015:Equijubus 6987:Choyrodon 6847:Iguanodon 6711:Planicoxa 6641:Dakotadon 6612:Calvarius 6438:Rhabdodon 6424:Mochlodon 6167:Elasmaria 5918:Kingdom: 5841:Dinosaurs 5721:131302174 5545:cite book 5220:Dinosaur" 4587:198152630 4280:Citations 3952:Abel 1924 3840:Glut 1997 3820:March 31, 3733:985402380 3223:Footnotes 3177:like the 3079:theropods 3053:Laramidia 2990:theropods 2470:pathology 2202:cassowary 2194:ligaments 2175:proboscis 2032:trapezoid 2028:river bed 1149:vertebrae 1134:upper arm 1126:thighbone 1061:jiayensis 1011:species, 1001:Fruitland 887:cristatus 806:In 1921, 759:Campanian 601:quadruped 593:herbivore 189:Kingdom: 183:Eukaryota 53:Campanian 8292:47459915 8257:Wikidata 8018:Cionodon 7687:Rhinorex 7575:Ornatops 7375:Minqaria 7346:Canardia 7273:Malefica 7198:Xuwulong 7163:Siamodon 7064:Jeyawati 7050:Gonkoken 7008:Eolambia 6832:Barilium 6633:Cumnoria 6598:Barilium 6504:Owenodon 6490:Draconyx 6452:Zalmoxes 6352:Fostoria 6204:Fostoria 5962:Cerapoda 5926:Chordata 5924:Phylum: 5920:Animalia 5692:24901454 5652:PLOS ONE 5486:89109302 5197:(1933). 5140:Postilla 4967:(1980). 4912:88689241 4741:24167777 4715:: e182. 4620:86070917 4430:(1945). 4420:54029279 4097:33289113 3698:53446536 3206:See also 3144:and the 2944:robustus 2771:, ? 2765:, ? 2755:, ? 2749:, ? 2745:, ? 2739:, ? 2641:Conifers 2443:Fantasia 2368:auditory 2093:juvenile 1286:subadult 1049:PLoS ONE 826:, where 744:based on 603:. Three 574:dinosaur 421:Synonyms 270:Family: 203:Chordata 199:Phylum: 193:Animalia 179:Domain: 8318:1019174 8305:4823227 8263:Q203583 7311:Ajnabia 6746:Uteodon 5827:Portals 5683:4047018 5660:Bibcode 5596:4160470 5568:Bibcode 5528:1313034 5478:2400478 5458:Bibcode 5434:1304144 5390:206–328 5207:1186563 5167:: 5–32. 5146:: 1–29. 5128:1301139 5073:Bibcode 5065:Lethaia 5009:Bibcode 4904:2400327 4884:Bibcode 4848:Lethaia 4820:Bibcode 4732:3807589 4678:Bibcode 4645:Bibcode 4567:Bibcode 4533:: 1–66. 4486:312–348 4400:Bibcode 4320:1133858 4300:Bibcode 4088:8053592 3690:4524171 3188:mammals 3171:lizards 3167:turtles 2992: ? 2830:turtles 2789:. When 2709:In the 2633:climate 2589:armored 2501:Alberta 2485:et al., 2380:diurnal 2171:snorkel 2161:(below) 2024:erosion 2020:maxilla 1926:chewing 1914:bipedal 1751:unnamed 1705:unnamed 1650:unnamed 1638:unnamed 1609:unnamed 1536:unnamed 1509:  1476:unnamed 1464:unnamed 1435:unnamed 1406:unnamed 1377:unnamed 1350:  1323:  1147:of the 925:Species 846:skull ( 840:tǔbǐcěn 824:Uppsala 797:Montana 663:from a 641:Alberta 605:species 564:) is a 309:Genus: 296:Tribe: 127:↓ 7980:Ootaxa 7177:Tanius 6770:Bolong 5853:Canada 5719:  5690:  5680:  5594:  5588:692723 5586:  5560:Nature 5526:  5484:  5476:  5432:  5396:  5239:  5205:  5126:  5103:  5052:  4977:  4927:  4910:  4902:  4788:  4739:  4729:  4618:  4585:  4546:  4492:  4446:691246 4444:  4418:  4375:  4356:  4337:  4318:  4095:  4085:  3731:  3721:  3696:  3688:  3198:, and 3123:, the 3114:, the 3081:, the 2927:; the 2881:; the 2877:, and 2828:, and 2810:fishes 2785:, and 2726:; the 2659:, and 2645:canopy 2622:marine 2618:swampy 2610:rivers 2546:, and 2533:horned 2364:visual 2303:, and 2280:, and 2185:, and 1948:Growth 1922:plants 1503:  1346:  1319:  1249:, and 1138:pelvic 1124:. 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Index

Parasaurolophus walkeri
Parasaurus
Late Cretaceous
Campanian
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N
Maastrichtian
P. cyrtocristatus skeleton
Field Museum of Natural History
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Ornithischia
Neornithischia
Ornithopoda
Hadrosauridae

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