925:
3314:
2725:
133:
111:
3332:
933:
784:
382:
history involving frequent synonymy between diagnostic and undiagnostic material. This is mainly due to the fact that it is a quintessential basal phytosaur, mostly distinguished by a lack of specializations rather than unique traits. Historically, it was common practice to
624:. TMM 31025-172, a complete skull, was collected at Otis Chalk Quarry 1 (also known as SMU 122), TMM 31100-8, 101, 175, 418, 419, 453 were collected at Otis Chalk Quarry 3, and TMM 31185-11, 38 were collected at Otis Chalk Quarry 3A.
1043:, but a re-description of the species by Butler (2013) found no evidence to support the synonymy, or even a phytosaurian identification. Although it is based solely on a
593:
2407:"A new taxon of phytosaur (Archosauria: Pseudosuchia) from the Late Triassic (Norian) Sonsela Member (Chinle Formation) in Arizona, and a critical reevaluation of
676:
2466:
1342:
2253:"A new phylogenetic analysis of Phytosauria (Archosauria: Pseudosuchia) with the application of continuous and geometric morphometric character coding"
3471:
2666:
3486:
3397:
900:, it might be possible that they also share the same species of phytosaur. Even though they agreed that due to the very fragmentary nature of
1354:
855:
was suggested to represent the oldest reliably dated phytosaur, and was reassigned to various phytosaur species over the years, including
797:& Sulej (2007) assigned several skulls, partial articulated postcranial skeletons, and numerous isolated phytosaur bones in various
3466:
1401:
on the species level, due to the poor preservation of it holotype and only known specimen. Stocker and Butler (2013) suggested that
886:, Dzik & Sulej (2007) noted that its holotype "does not differ significantly from corresponding parts of the juvenile Krasiejów
3384:
3481:
2758:
2562:
Hungerbühler, A. (2001). "The status and phylogenetic relationships of "Zanclodon"arenaceus: The earliest known phytosaur?".
2779:
3456:
2492:
Long, R. A.; Murry, P. A. (1995). "Late
Triassic (Carnian and Norian) tetrapods from the southwestern United States".
2514:
Kuhn, O (1936). "Weitere
Parasuchier und Labyrinthodonten aus dem Blasensandstein des mittleren Keuper von Ebrach".
1369:, studies as of 2013 considered the synonymy between the genera to be premature. However, later studies argued that
3313:
1269:(1978), who described many complete remains of the Indian parasuchian, showed that it is not assignable either to
1345:(Opinion 2045) following the application of Chatterjee (2001). As a result, most subsequent studies referred all
2313:
592:
of the
Chugwater Group. The holotype was thoroughly described by Lees (1907). Hunt and Lucas (1991) referred
1935:
132:
2681:
1911:
1887:
1777:
2873:
2751:
394:
ancestral to later phytosaurs. More recently, these grades have been broken up into multiple genera.
1051:
was found to possess a unique combination of characters that distinguish it from all other
Triassic
1389:
890:, which is clearly a phytosaur". Furthermore, as the Feuerbacher Heide Schilfsandstein, from which
847:
565:
233:
3461:
2705:
Case, E. C. (1922). "New reptiles and stegocephalians from the Upper
Triassic of western Texas".
2349:
Kammerer, Christian F.; Butler, Richard J.; Bandyopadhyay, Saswati; Stocker, Michelle R. (2016).
1952:
1818:
1180:, although recent species-level phylogenetic analyses of phytosaurs suggest that this species is
1168:
617:
581:
396:
3428:
3352:
1652:
1326:
1074:
1011:
in a different subfamily. Hunt and Lucas (1991) considered these species to be synonymous with
491:
3402:
2667:"A primitive parasuchid (phytosaur) reptile from the Upper Triassic Maleri Formation of India"
3476:
3423:
3415:
1708:
1593:
1203:
1181:
953:
507:
440:
384:
327:
2350:
2744:
2164:
2096:
1801:
1255:
1031:
981:
961:. Subsequent researches accepted this referral. More recently, phylogenetic analysis found
924:
701:
642:
621:
569:
237:
8:
3451:
3335:
2846:
2730:
1239:
1234:
717:
589:
580:
that has been extensively reconstructed with plaster. It was collected at Squaw Creek of
2168:
2100:
2647:
2579:
2432:
2382:
2287:
2252:
2228:
2180:
2112:
1860:
1551:
1479:
1210:
1173:
1137:
1132:
1113:
1108:
987:
712:
707:
478:
406:
391:
127:
1317:
should be avoided. Nevertheless, Hunt and Lucas (1991) considered the species to be a
3410:
3245:
3068:
2427:
2406:
2374:
2292:
2274:
2232:
2184:
2116:
1676:
1385:
under ICZN rules, formally synonymizing the two genera in this more restricted sense.
1266:
1088:, more recent studies disagree with this referral. Kammerer et al. (2016) considered
1044:
660:
605:
2651:
2583:
2436:
2386:
991:
Kuhn, 1933 - Gregory (1962) and
Wastphal (1976) referred these species, as well as,
3147:
3140:
3093:
2858:
2806:
2639:
2571:
2422:
2366:
2282:
2264:
2220:
2172:
2104:
1721:
200:
187:
834:. The shape of the external mandibular fenestra of the material resembled that of
3375:
3252:
3022:
2951:
2643:
1606:
843:
726:
608:. Later, Long and Murry (1995) referred additional material, including isolated
3131:
1873:
1288:
1066:
882:
601:
2224:
2213:
Earth and
Environmental Science Transactions of the Royal Society of Edinburgh
1247:), thus he assigned the phytosaurian material to a newly named species "aff."
438:(a dubious phytosaur genus from the same area), and has also been compared to
3445:
3296:
3224:
3168:
3154:
3082:
3054:
3040:
3027:
2990:
2537:
2378:
2278:
1620:
1393:
Case, 1922 - Gregory (1962) and Hunt and Lucas (1991) referred this taxon to
1275:
1223:
1189:
1153:
1078:
Dutuit, 1977b - Long and Murry (1995) reassigned this species to a new genus
966:
863:
815:
688:
334:
68:
30:
3282:
3267:
3238:
3231:
3216:
3192:
3047:
2983:
2970:
2296:
1464:
1398:
1318:
1284:
1249:
1145:
1080:
1057:
1024:
896:
770:
762:
750:
695:
647:
557:
434:
342:
245:
110:
3275:
3161:
3120:
3113:
3106:
2958:
2538:"A review of the early Late Triassic Krasiejów biotafrom Silesia, Poland"
2471:
biochron and the correlation of the non-marine Upper
Triassic of Pangaea"
1410:
1293:
1260:
1230:
1226:
878:
869:
495:
387:
all basal phytosaurs into only one or two genera, rendering those genera
43:
2318:
Lydekker, 1885 (Reptilia, Archosauria): Lectotype
Replaced By A Neotype"
410:(from Texas) were two phytosaur genera originally considered species of
3389:
3289:
3209:
3004:
2831:
2815:
2797:
2575:
2269:
830:
794:
638:
609:
285:
88:
53:
2370:
1333:
species. To avoid additional confusion, the nondiagnostic holotype of
3259:
2937:
2915:
2885:
2767:
1426:
1306:
1172:
Langston, 1949 - Hunt and Lucas (1991) considered this species to be
1149:
1062:
837:
482:, a basal phytosaur species named in 1885 from fossils discovered in
458:
422:(from Wyoming) are two more phytosaur species informally referred to
388:
330:
213:
144:
93:
37:
3346:
2600:
Gregory, J. T. (1962). "The relationships of the
American phytosaur
2176:
2084:
600:. It is an incomplete skull that was collected at Palo Duro Canyon,
3369:
3075:
3061:
2791:
2108:
1298:
1052:
798:
766:
664:
656:
573:
379:
164:
83:
78:
63:
58:
48:
2348:
1329:, stating that the Indian species can be distinguished from other
1297:). He noted that since the rhynchosaur basicranium is neither the
1136:
Long and Murry, 1995 - some studies considered this species to be
506:
have helped to re-establish it as a valid genus and species. Some
3202:
1338:
1219:
1112:
Mehl, 1928 - Hunt and Lucas (1991) considered this species to be
932:
857:
672:
668:
585:
499:
401:
365:
351:
338:
320:
174:
98:
73:
3011:
2785:
1198:
802:
604:, west Texas, from the Carnian-aged Camp Springs Member of the
467:
154:
2736:
2630:: Implications for Late Triassic vertebrate biostratigraphy".
783:
2085:"Notice of some new reptiles from the Upper Trias of Wyoming"
957:
Kuhn, 1936 - Hunt and Lucas (1991) reassigned the species as
577:
515:
483:
355:
323:
940:
Through the years, various species have been referred to as
2494:
New Mexico Museum of Natural History and Science Bulletin
1429:, from Kammerer et al., 2016, shows the relationships of
1258:(1958) designated all the Indian parasuchian material as
944:. The species that are no longer considered to belong to
2207:
Stocker, M. R. (2013). "A new taxonomic arrangement for
908:
can't be referred to it, they tentatively used the name
809:. They found some similarities between the material and
2155:
Stocker, M. R.; Butler, R. J. (2013). "Phytosauria".
1152:
suggest that this species is more closely related to
1007:, while Chatterjee (1978) placed the four species in
667:. It was collected at Ebrach Quarry, bed number 9 of
2720:
426:, though likely closer to more advanced phytosaurs.
368:, Germany, is also commonly referred to the genus.
292:
275:
252:
2404:
2251:Jones, Andrew S.; Butler, Richard J. (2018-12-10).
877:as not belonging to Phytosauria and referred it to
2150:
2148:
2146:
1237:(1940) identified the basicranium as belonging to
2664:
2535:
2144:
2142:
2140:
2138:
2136:
2134:
2132:
2130:
2128:
2126:
1325:for the diagnosable phytosaur specimens from the
1120:, while Stocker and Butler (2013) suggested that
1084:. Although several studies reassigned it back to
341:stage). The genus was named in 1904 based on the
3443:
2157:Geological Society, London, Special Publications
1273:(which is closely related to or synonymous with
1196:. This species was reassigned to its own genus,
1365:. In light of the complex taxonomic history of
2707:Carnegie Institution of Washington Publication
2555:
2531:
2529:
2123:
1397:on the genus level, and considered it to be a
894:was collected, and Krasiejów share species of
2752:
2460:
2458:
2456:
2454:
2452:
2450:
2448:
2446:
2400:
2398:
2396:
2154:
2078:
2076:
828:), as well as some apparent differences with
682:Hunt and Lucas (1991) mistakenly referred to
372:had a length of about 2.5 meters (8.2 feet).
358:in the United States. Another valid species,
2658:
2612:
2595:
2593:
2561:
1321:, and provisionally created the combination
659:BSPG 1931 X 502 a partial skull lacking the
526:. If this is the case, the valid species of
2526:
2250:
1202:by Stocker (2013), and represents the most
873:. Although Hungerbühler (2001) redescribed
2935:
2759:
2745:
2485:
2443:
2393:
2073:
534:, since that genus name has priority over
109:
2830:
2590:
2509:
2507:
2491:
2464:
2426:
2286:
2268:
2082:
1453:, owing to their close relationship with
1265:. Gregory (1962) accepted this proposal.
1055:. Thus it was reassigned to a new genus,
2202:
2200:
2198:
2196:
2194:
1441:to other phytosaurs. The authors placed
931:
923:
782:
745:as it shares unique synapomorphies with
725:was found to be more closely related to
576:FMNH UC 632, a complete but fragmentary
3472:Late Triassic reptiles of North America
2599:
2351:"Relationships of the Indian phytosaur
2206:
432:was originally considered a species of
3487:Taxa named by Samuel Wendell Williston
3444:
2618:
2504:
2308:
2306:
1233:partial snout, scutes and some teeth.
693:, and considered it and the other two
564:. It was first described and named by
466:), fragmentary phytosaur remains from
3351:
3350:
2778:
2740:
2191:
1535:Parasuchus (Paleorhinus) angustifrons
530:may instead be considered species of
2704:
2513:
2344:
2342:
2340:
2338:
2246:
2244:
2242:
1381:. This clade would receive the name
1353:in accordance with the rules of the
948:, are summarised in the list below:
2325:Bulletin of Zoological Nomenclature
2303:
776:
13:
2632:Journal of Vertebrate Paleontology
2465:Hunt, A. P.; Lucas, S. G. (1991).
671:, southern Germany, from the late
627:
14:
3498:
2335:
2239:
1511:Parasuchus (Paleorhinus) bransoni
936:Model based on the Polish remains
637:was first described and named by
3467:Late Triassic reptiles of Europe
3331:
3330:
3312:
2723:
2428:10.1111/j.1475-4983.2010.00983.x
1192:, and thus can't be referred to
1144:, although recent species-level
919:
131:
41:
2766:
2698:
842:but it is also shared with the
2516:Palaeontographica, Abteilung A
1357:, while others kept referring
1160:and thus can't be referred to
965:to be more closely related to
928:Skull and skeleton from Poland
761:because they were found to be
757:species, were not referred to
655:is known exclusively from the
546:
1:
3482:Fossil taxa described in 1904
2066:
2644:10.1080/02724634.2013.740542
2564:Paläontologische Zeitschrift
2536:Dzik, J.; Sulej, T. (2007).
2405:Michelle R. Stocker (2010).
1936:Machaeroprosopus jablonskiae
1420:
679:of the Hassberge Formation.
7:
2680:(1): 83–127. Archived from
1377:formed a strong clade with
824:(mistakenly referred to as
448:-like Bavarian phytosaur. "
10:
3503:
3457:Prehistoric reptile genera
2665:Sankar Chatterjee (1978).
1912:Machaeroprosopus pristinus
1888:Machaeroprosopus mccauleyi
1778:Smilosuchus lithodendrorum
789:Paleorhinus" cf. arenaceus
541:
3359:
3326:
3309:
3190:
3130:
3101:
3092:
3035:
3021:
2978:
2969:
2946:
2931:
2912:
2884:
2866:
2857:
2839:
2826:
2774:
2606:American Museum Novitates
2225:10.1017/S1755691013000340
2083:Williston, S. W. (1904).
1949:
1932:
1925:
1908:
1901:
1884:
1877:
1857:
1850:
1815:
1798:
1791:
1774:
1767:
1760:
1745:
1732:
1725:
1705:
1690:
1673:
1666:
1649:
1634:
1617:
1610:
1590:
1583:
1548:
1531:
1524:
1507:
1500:
1493:
1476:
1469:
1169:"Paleorhinus" scurriensis
910:Paleorhinus cf. arenaceus
807:Paleorhinus cf. arenaceus
272:
267:
251:
244:
128:Scientific classification
126:
117:
108:
23:
16:Extinct genus of reptiles
1693:"Machaeroprosopus" zunii
1405:may not be referable to
1390:Promystriosuchus ehlersi
1375:Paleorhinus angustifrons
1218:was based on a chimeric
1124:may not be referable to
1098:P. hislopi, P. bransoni,
848:Chanaresuchus bonapartei
749:and was found to be its
737:was re-validated, while
566:Samuel Wendell Williston
490:or considered a dubious
486:, was often lumped into
361:Paleorhinus angustifrons
2619:Butler, R. J. (2013). "
2545:Palaeontologia Polonica
2359:Papers in Palaeontology
2209:Paleorhinus scurriensis
1953:Mystriosuchus westphali
1819:Smilosuchus adamanensis
1341:with approval from the
1184:to the clade formed by
904:holotype the Krasiejów
801:stages from Krasiejów,
618:Cooper Canyon Formation
582:Fremont County, Wyoming
498:fossils. More complete
397:Arganarhinus magnoculus
1748:"Phytosaurus" doughtyi
1653:Brachysuchus megalodon
1327:Lower Maleri Formation
1075:Paleorhinus magnoculus
1032:"Francosuchus" trauthi
1023:were considered to be
937:
929:
791:
769:were destroyed during
677:Blasensandstein Member
616:from the Carnian-aged
378:has had a complicated
350:, which is known from
122:holotype with CT scan
3424:Paleobiology Database
2624:Francosuchus' trauthi
1709:Rutiodon carolinensis
1594:Ebrachosuchus neukami
1313:, the suppression of
1245:Hyperodapedon huxleyi
1039:was synonymized with
954:Ebrachosuchus neukami
935:
927:
787:Life restoration of "
786:
620:of the Dockum Group,
508:phylogenetic analyses
441:Ebrachosuchus neukami
1802:Smilosuchus gregorii
1637:"Paleorhinus" sawini
1371:Paleorhinus bransoni
1256:Edwin Harris Colbert
1133:"Paleorhinus" sawini
1109:"Paleorhinus" parvus
982:Francosuchus broilii
622:Howard County, Texas
572:on the basis of the
520:Paleorhinus bransoni
420:"Paleorhinus" parvus
416:"Paleorhinus" sawini
347:Paleorhinus bransoni
257:Paleorhinus bransoni
2731:paleontology portal
2169:2013GSLSP.379...91S
2101:1904JG.....12..688W
1409:and referred it to
1323:Paleorhinus hislopi
1311:Paradapedon huxleyi
1240:Paradapedon huxleyi
1235:Friedrich von Huene
1092:to be a species of
1061:, now referable to
1013:Paleorhinus neukami
973:and thus the genus
959:Paleorhinus neukami
733:and thus the genus
718:Paleorhinus neukami
590:Popo Agie Formation
392:evolutionary grades
2576:10.1007/BF03022600
2353:Parasuchus hislopi
2316:Parasuchus Hislopi
2270:10.7717/peerj.5901
2089:Journal of Geology
1861:Pravusuchus hortus
1552:Parasuchus hislopi
1480:Wannia scurriensis
1455:Parasuchus hislopi
1379:Parasuchus hislopi
1337:was replaced by a
1211:Parasuchus hislopi
938:
930:
912:for the Krasiejów
792:
741:was reassigned as
512:Parasuchus hislopi
504:Parasuchus hislopi
479:Parasuchus hislopi
407:Wannia scurriensis
3439:
3438:
3411:Open Tree of Life
3353:Taxon identifiers
3344:
3343:
3322:
3321:
3307:
3306:
3246:Heterodontosuchus
3186:
3185:
3182:
3181:
3178:
3177:
2927:
2926:
2923:
2922:
2908:
2907:
2904:
2903:
2371:10.1002/spp2.1022
2063:
2062:
2054:
2053:
2045:
2044:
2036:
2035:
2027:
2026:
2018:
2017:
2009:
2008:
2000:
1999:
1991:
1990:
1982:
1981:
1973:
1972:
1964:
1963:
1839:
1838:
1830:
1829:
1677:Protome batalaria
1572:
1571:
1563:
1562:
1267:Sankar Chatterjee
1214:Lydekker, 1885 -
1015:. More recently,
977:was re-validated.
721:. More recently,
606:Tecovas Formation
494:of phytosaur and
418:(from Texas) and
308:
307:
302:
289:
240:
36:221.5–205.6
3494:
3432:
3431:
3419:
3418:
3406:
3405:
3393:
3392:
3380:
3379:
3378:
3348:
3347:
3334:
3333:
3317:
3316:
3148:Machaeroprosopus
3141:Angistorhinopsis
3099:
3098:
3094:Leptosuchomorpha
3033:
3032:
2976:
2975:
2944:
2943:
2933:
2932:
2874:Archosauromorpha
2864:
2863:
2859:Archosauromorpha
2837:
2836:
2828:
2827:
2821:
2820:
2807:Archosauromorpha
2776:
2775:
2761:
2754:
2747:
2738:
2737:
2733:
2728:
2727:
2726:
2715:
2714:
2702:
2696:
2695:
2693:
2692:
2686:
2671:
2662:
2656:
2655:
2623:
2616:
2610:
2609:
2597:
2588:
2587:
2559:
2553:
2552:
2542:
2533:
2524:
2523:
2511:
2502:
2501:
2489:
2483:
2482:
2462:
2441:
2440:
2430:
2402:
2391:
2390:
2346:
2333:
2332:
2331:: 174–175. 2003.
2322:
2310:
2301:
2300:
2290:
2272:
2248:
2237:
2236:
2219:(3–4): 251–263.
2204:
2189:
2188:
2152:
2121:
2120:
2080:
1928:
1927:
1904:
1903:
1880:
1879:
1853:
1852:
1794:
1793:
1770:
1769:
1763:
1762:
1728:
1727:
1722:Leptosuchomorpha
1669:
1668:
1613:
1612:
1586:
1585:
1527:
1526:
1503:
1502:
1496:
1495:
1472:
1471:
1461:
1460:
1206:known phytosaur.
778:P. cf. arenaceus
753:. The other two
645:as a species of
470:, may represent
300:
294:
283:
277:
254:
232:
225:
212:
201:Archosauriformes
188:Archosauromorpha
136:
135:
113:
103:
40:
29:Temporal range:
21:
20:
3502:
3501:
3497:
3496:
3495:
3493:
3492:
3491:
3442:
3441:
3440:
3435:
3427:
3422:
3414:
3409:
3401:
3396:
3388:
3383:
3374:
3373:
3368:
3355:
3345:
3340:
3318:
3311:
3303:
3253:Mesorhinosuchus
3174:
3126:
3088:
3025:
3023:Mystriosuchinae
3017:
2965:
2952:Diandongosuchus
2940:
2919:
2900:
2880:
2853:
2822:
2781:
2780:
2770:
2765:
2729:
2724:
2722:
2719:
2718:
2703:
2699:
2690:
2688:
2684:
2669:
2663:
2659:
2621:
2617:
2613:
2598:
2591:
2560:
2556:
2540:
2534:
2527:
2512:
2505:
2490:
2486:
2463:
2444:
2421:(5): 997–1022.
2403:
2394:
2355:Lydekker, 1885"
2347:
2336:
2320:
2312:
2311:
2304:
2249:
2240:
2205:
2192:
2177:10.1144/SP379.5
2153:
2124:
2081:
2074:
2069:
2064:
2055:
2046:
2037:
2028:
2019:
2010:
2001:
1992:
1983:
1974:
1965:
1840:
1831:
1607:Mystriosuchinae
1573:
1564:
1449:into the genus
1447:P. angustifrons
1435:P. angustifrons
1423:
1287:, probably the
1102:P. angustifrons
997:P. angustifrons
922:
844:proterochampsid
822:F. angustifrons
781:
759:P. angustifrons
743:P. angustifrons
739:F. angustifrons
727:Mystriosuchinae
684:F. angustifrons
653:P. angustifrons
635:P. angustifrons
632:
629:P. angustifrons
551:
544:
430:P. angustifrons
333:known from the
301:Williston, 1904
280:P. angustifrons
263:
262:Williston, 1904
260:
231:
223:
210:
130:
120:P. angustifrons
104:
102:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
27:
17:
12:
11:
5:
3500:
3490:
3489:
3484:
3479:
3474:
3469:
3464:
3462:Carnian genera
3459:
3454:
3437:
3436:
3434:
3433:
3420:
3407:
3394:
3381:
3365:
3363:
3357:
3356:
3342:
3341:
3339:
3338:
3327:
3324:
3323:
3320:
3319:
3310:
3308:
3305:
3304:
3302:
3301:
3293:
3286:
3279:
3272:
3264:
3256:
3249:
3242:
3235:
3228:
3221:
3213:
3206:
3198:
3196:
3188:
3187:
3184:
3183:
3180:
3179:
3176:
3175:
3173:
3172:
3165:
3158:
3151:
3144:
3136:
3134:
3132:Mystriosuchini
3128:
3127:
3125:
3124:
3117:
3110:
3102:
3096:
3090:
3089:
3087:
3086:
3079:
3072:
3065:
3058:
3051:
3044:
3036:
3030:
3019:
3018:
3016:
3015:
3008:
3001:
2994:
2987:
2979:
2973:
2967:
2966:
2964:
2963:
2955:
2947:
2941:
2936:
2929:
2928:
2925:
2924:
2921:
2920:
2913:
2910:
2909:
2906:
2905:
2902:
2901:
2899:
2898:
2897:
2896:
2890:
2888:
2882:
2881:
2879:
2878:
2877:
2876:
2867:
2861:
2855:
2854:
2852:
2851:
2850:
2849:
2840:
2834:
2824:
2823:
2819:
2818:
2809:
2800:
2794:
2788:
2772:
2771:
2764:
2763:
2756:
2749:
2741:
2735:
2734:
2717:
2716:
2697:
2657:
2638:(4): 858–864.
2611:
2589:
2554:
2525:
2503:
2484:
2442:
2392:
2334:
2302:
2238:
2190:
2122:
2109:10.1086/621190
2095:(8): 688–697.
2071:
2070:
2068:
2065:
2061:
2060:
2057:
2056:
2052:
2051:
2048:
2047:
2043:
2042:
2039:
2038:
2034:
2033:
2030:
2029:
2025:
2024:
2021:
2020:
2016:
2015:
2012:
2011:
2007:
2006:
2003:
2002:
1998:
1997:
1994:
1993:
1989:
1988:
1985:
1984:
1980:
1979:
1976:
1975:
1971:
1970:
1967:
1966:
1962:
1961:
1958:
1957:
1948:
1945:
1944:
1941:
1940:
1931:
1926:
1924:
1921:
1920:
1917:
1916:
1907:
1902:
1900:
1897:
1896:
1893:
1892:
1883:
1878:
1876:
1874:Mystriosuchini
1870:
1869:
1866:
1865:
1856:
1851:
1849:
1846:
1845:
1842:
1841:
1837:
1836:
1833:
1832:
1828:
1827:
1824:
1823:
1814:
1811:
1810:
1807:
1806:
1797:
1792:
1790:
1787:
1786:
1783:
1782:
1773:
1768:
1766:
1761:
1759:
1756:
1755:
1752:
1751:
1744:
1741:
1740:
1737:
1736:
1734:TMM 31173-120
1731:
1726:
1724:
1718:
1717:
1714:
1713:
1704:
1701:
1700:
1697:
1696:
1689:
1686:
1685:
1682:
1681:
1672:
1667:
1665:
1662:
1661:
1658:
1657:
1648:
1645:
1644:
1641:
1640:
1633:
1630:
1629:
1626:
1625:
1616:
1611:
1609:
1603:
1602:
1599:
1598:
1589:
1584:
1582:
1579:
1578:
1575:
1574:
1570:
1569:
1566:
1565:
1561:
1560:
1557:
1556:
1547:
1544:
1543:
1540:
1539:
1530:
1525:
1523:
1520:
1519:
1516:
1515:
1506:
1501:
1499:
1494:
1492:
1489:
1488:
1485:
1484:
1475:
1470:
1468:
1459:
1425:The following
1422:
1419:
1418:
1417:
1414:incertae sedis
1386:
1289:senior synonym
1243:(now known as
1224:rhynchosaurian
1207:
1165:
1129:
1105:
1071:
1067:incertae sedis
1035:Huene, 1939 -
1028:
978:
921:
918:
902:"Z." arenaceus
892:"Z." arenaceus
883:incertae sedis
875:"Z." arenaceus
853:"Z." arenaceus
780:
775:
631:
626:
602:Randall County
550:
545:
543:
540:
326:of widespread
306:
305:
304:
303:
290:
270:
269:
265:
264:
261:
249:
248:
242:
241:
221:
217:
216:
208:
204:
203:
198:
191:
190:
185:
178:
177:
172:
168:
167:
162:
158:
157:
152:
148:
147:
142:
138:
137:
124:
123:
115:
114:
106:
105:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
28:
15:
9:
6:
4:
3:
2:
3499:
3488:
3485:
3483:
3480:
3478:
3475:
3473:
3470:
3468:
3465:
3463:
3460:
3458:
3455:
3453:
3450:
3449:
3447:
3430:
3425:
3421:
3417:
3412:
3408:
3404:
3399:
3395:
3391:
3386:
3382:
3377:
3371:
3367:
3366:
3364:
3362:
3358:
3354:
3349:
3337:
3329:
3328:
3325:
3315:
3299:
3298:
3297:Termatosaurus
3294:
3292:
3291:
3287:
3285:
3284:
3280:
3278:
3277:
3273:
3270:
3269:
3265:
3262:
3261:
3257:
3255:
3254:
3250:
3248:
3247:
3243:
3241:
3240:
3236:
3234:
3233:
3229:
3227:
3226:
3225:Coburgosuchus
3222:
3219:
3218:
3214:
3212:
3211:
3207:
3205:
3204:
3200:
3199:
3197:
3195:
3194:
3189:
3171:
3170:
3169:Redondasaurus
3166:
3164:
3163:
3159:
3157:
3156:
3155:Mystriosuchus
3152:
3150:
3149:
3145:
3143:
3142:
3138:
3137:
3135:
3133:
3129:
3123:
3122:
3118:
3116:
3115:
3111:
3109:
3108:
3104:
3103:
3100:
3097:
3095:
3091:
3085:
3084:
3083:Volcanosuchus
3080:
3078:
3077:
3073:
3071:
3070:
3066:
3064:
3063:
3059:
3057:
3056:
3055:Colossosuchus
3052:
3050:
3049:
3045:
3043:
3042:
3041:Angistorhinus
3038:
3037:
3034:
3031:
3029:
3028:Phytosauridae
3024:
3020:
3014:
3013:
3009:
3007:
3006:
3002:
3000:
2999:
2995:
2993:
2992:
2991:Ebrachosuchus
2988:
2986:
2985:
2981:
2980:
2977:
2974:
2972:
2968:
2961:
2960:
2956:
2954:
2953:
2949:
2948:
2945:
2942:
2939:
2934:
2930:
2918:
2917:
2911:
2894:
2893:
2892:
2891:
2889:
2887:
2883:
2875:
2871:
2870:
2869:
2868:
2865:
2862:
2860:
2856:
2848:
2844:
2843:
2842:
2841:
2838:
2835:
2833:
2829:
2825:
2817:
2813:
2810:
2808:
2804:
2801:
2799:
2795:
2793:
2789:
2787:
2783:
2782:
2777:
2773:
2769:
2762:
2757:
2755:
2750:
2748:
2743:
2742:
2739:
2732:
2721:
2712:
2708:
2701:
2687:on 2013-08-10
2683:
2679:
2675:
2674:Palaeontology
2668:
2661:
2653:
2649:
2645:
2641:
2637:
2633:
2629:
2625:
2615:
2608:(2095): 1–22.
2607:
2603:
2596:
2594:
2585:
2581:
2577:
2573:
2569:
2565:
2558:
2550:
2546:
2539:
2532:
2530:
2521:
2517:
2510:
2508:
2499:
2495:
2488:
2481:(2): 487–501.
2480:
2476:
2475:Palaeontology
2472:
2470:
2461:
2459:
2457:
2455:
2453:
2451:
2449:
2447:
2438:
2434:
2429:
2424:
2420:
2416:
2415:Palaeontology
2412:
2410:
2401:
2399:
2397:
2388:
2384:
2380:
2376:
2372:
2368:
2364:
2360:
2356:
2354:
2345:
2343:
2341:
2339:
2330:
2326:
2319:
2317:
2309:
2307:
2298:
2294:
2289:
2284:
2280:
2276:
2271:
2266:
2262:
2258:
2254:
2247:
2245:
2243:
2234:
2230:
2226:
2222:
2218:
2214:
2210:
2203:
2201:
2199:
2197:
2195:
2186:
2182:
2178:
2174:
2170:
2166:
2163:(1): 91–117.
2162:
2158:
2151:
2149:
2147:
2145:
2143:
2141:
2139:
2137:
2135:
2133:
2131:
2129:
2127:
2118:
2114:
2110:
2106:
2102:
2098:
2094:
2090:
2086:
2079:
2077:
2072:
2059:
2058:
2050:
2049:
2041:
2040:
2032:
2031:
2023:
2022:
2014:
2013:
2005:
2004:
1996:
1995:
1987:
1986:
1978:
1977:
1969:
1968:
1960:
1959:
1956:
1955:
1954:
1947:
1946:
1943:
1942:
1939:
1938:
1937:
1930:
1929:
1923:
1922:
1919:
1918:
1915:
1914:
1913:
1906:
1905:
1899:
1898:
1895:
1894:
1891:
1890:
1889:
1882:
1881:
1875:
1872:
1871:
1868:
1867:
1864:
1863:
1862:
1855:
1854:
1848:
1847:
1844:
1843:
1835:
1834:
1826:
1825:
1822:
1821:
1820:
1813:
1812:
1809:
1808:
1805:
1804:
1803:
1796:
1795:
1789:
1788:
1785:
1784:
1781:
1780:
1779:
1772:
1771:
1765:
1764:
1758:
1757:
1754:
1753:
1750:
1749:
1743:
1742:
1739:
1738:
1735:
1730:
1729:
1723:
1720:
1719:
1716:
1715:
1712:
1711:
1710:
1703:
1702:
1699:
1698:
1695:
1694:
1688:
1687:
1684:
1683:
1680:
1679:
1678:
1671:
1670:
1664:
1663:
1660:
1659:
1656:
1655:
1654:
1647:
1646:
1643:
1642:
1639:
1638:
1632:
1631:
1628:
1627:
1624:
1623:
1622:
1621:Angistorhinus
1615:
1614:
1608:
1605:
1604:
1601:
1600:
1597:
1596:
1595:
1588:
1587:
1581:
1580:
1577:
1576:
1568:
1567:
1559:
1558:
1555:
1554:
1553:
1546:
1545:
1542:
1541:
1538:
1537:
1536:
1529:
1528:
1522:
1521:
1518:
1517:
1514:
1513:
1512:
1505:
1504:
1498:
1497:
1491:
1490:
1487:
1486:
1483:
1482:
1481:
1474:
1473:
1466:
1463:
1462:
1458:
1456:
1452:
1448:
1444:
1440:
1436:
1432:
1428:
1415:
1412:
1408:
1404:
1400:
1396:
1392:
1391:
1387:
1384:
1380:
1376:
1372:
1368:
1364:
1360:
1356:
1352:
1348:
1344:
1340:
1336:
1332:
1328:
1324:
1320:
1316:
1312:
1308:
1304:
1300:
1296:
1295:
1290:
1286:
1285:doubtful name
1282:
1278:
1277:
1276:Angistorhinus
1272:
1268:
1264:
1262:
1257:
1253:
1251:
1246:
1242:
1241:
1236:
1232:
1228:
1225:
1222:material - a
1221:
1217:
1213:
1212:
1208:
1205:
1201:
1200:
1195:
1191:
1190:Phytosauridae
1187:
1183:
1179:
1175:
1171:
1170:
1166:
1163:
1159:
1155:
1154:Phytosauridae
1151:
1147:
1143:
1139:
1135:
1134:
1130:
1127:
1123:
1119:
1115:
1111:
1110:
1106:
1103:
1099:
1095:
1091:
1090:P. magnoculus
1087:
1083:
1082:
1077:
1076:
1072:
1069:
1068:
1064:
1060:
1059:
1054:
1050:
1046:
1042:
1038:
1034:
1033:
1029:
1026:
1022:
1018:
1014:
1010:
1006:
1002:
998:
994:
990:
989:
984:
983:
979:
976:
975:Ebrachosuchus
972:
968:
967:Phytosauridae
964:
960:
956:
955:
951:
950:
949:
947:
943:
934:
926:
920:Other species
917:
915:
911:
907:
903:
899:
898:
893:
889:
885:
884:
880:
876:
872:
871:
866:
865:
864:Mystriosuchus
860:
859:
854:
850:
849:
845:
841:
839:
833:
832:
827:
823:
819:
817:
816:Ebrachosuchus
812:
808:
804:
800:
796:
790:
785:
779:
774:
772:
768:
764:
760:
756:
752:
748:
744:
740:
736:
735:Ebrachosuchus
732:
728:
724:
720:
719:
714:
710:
709:
704:
703:
698:
697:
692:
690:
689:Ebrachosuchus
685:
680:
678:
674:
670:
666:
662:
658:
654:
650:
649:
644:
640:
636:
630:
625:
623:
619:
615:
611:
607:
603:
599:
595:
591:
587:
583:
579:
575:
571:
567:
563:
559:
555:
549:
539:
537:
533:
529:
525:
521:
517:
513:
510:suggest that
509:
505:
501:
497:
493:
489:
485:
481:
480:
475:
473:
469:
465:
461:
460:
455:
451:
447:
443:
442:
437:
436:
431:
427:
425:
421:
417:
413:
409:
408:
403:
399:
398:
393:
390:
386:
381:
377:
373:
371:
367:
363:
362:
357:
353:
349:
348:
344:
340:
336:
335:Late Triassic
332:
329:
325:
322:
318:
314:
313:
299:
298:
291:
287:
282:
281:
274:
273:
271:
266:
259:
258:
250:
247:
243:
239:
235:
230:
229:
222:
219:
218:
215:
209:
206:
205:
202:
199:
196:
193:
192:
189:
186:
183:
180:
179:
176:
173:
170:
169:
166:
163:
160:
159:
156:
153:
150:
149:
146:
143:
140:
139:
134:
129:
125:
121:
118:Skull of the
116:
112:
107:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
31:Late Triassic
26:
22:
19:
3477:Chinle fauna
3360:
3295:
3288:
3283:Rileyasuchus
3281:
3274:
3268:Palaeoctonus
3266:
3258:
3251:
3244:
3239:Francosuchus
3237:
3232:Compsosaurus
3230:
3223:
3217:Clepsysaurus
3215:
3208:
3201:
3193:Nomina dubia
3191:
3167:
3160:
3153:
3146:
3139:
3119:
3112:
3105:
3081:
3074:
3067:
3060:
3053:
3048:Brachysuchus
3046:
3039:
3010:
3003:
2997:
2996:
2989:
2984:Arganarhinus
2982:
2971:Parasuchidae
2957:
2950:
2914:
2811:
2802:
2710:
2706:
2700:
2689:. Retrieved
2682:the original
2677:
2673:
2660:
2635:
2631:
2627:
2620:
2614:
2605:
2601:
2567:
2563:
2557:
2548:
2544:
2519:
2515:
2497:
2493:
2487:
2478:
2474:
2468:
2418:
2414:
2408:
2362:
2358:
2352:
2328:
2324:
2315:
2260:
2256:
2216:
2212:
2208:
2160:
2156:
2092:
2088:
1951:
1950:
1934:
1933:
1910:
1909:
1886:
1885:
1859:
1858:
1817:
1816:
1800:
1799:
1776:
1775:
1747:
1746:
1733:
1707:
1706:
1692:
1691:
1675:
1674:
1651:
1650:
1636:
1635:
1619:
1618:
1592:
1591:
1550:
1549:
1534:
1533:
1532:
1510:
1509:
1508:
1478:
1477:
1465:Parasuchidae
1454:
1450:
1446:
1442:
1438:
1434:
1430:
1424:
1413:
1406:
1402:
1399:nomen dubium
1394:
1388:
1382:
1378:
1374:
1370:
1366:
1362:
1358:
1350:
1346:
1334:
1330:
1322:
1319:nomen dubium
1314:
1310:
1302:
1292:
1280:
1274:
1271:Brachysuchus
1270:
1259:
1250:Brachysuchus
1248:
1244:
1238:
1231:phytosaurian
1215:
1209:
1197:
1193:
1185:
1177:
1167:
1161:
1157:
1148:analyses of
1146:phylogenetic
1141:
1131:
1125:
1121:
1117:
1107:
1101:
1097:
1093:
1089:
1085:
1081:Arganarhinus
1079:
1073:
1065:
1058:Dolerosaurus
1056:
1049:"F." trauthi
1048:
1040:
1037:"F." trauthi
1036:
1030:
1025:nomina dubia
1020:
1016:
1012:
1009:Francosuchus
1008:
1005:Francosuchus
1004:
1000:
996:
992:
986:
980:
974:
970:
962:
958:
952:
945:
941:
939:
913:
909:
905:
901:
897:Metoposaurus
895:
891:
887:
881:
874:
868:
862:
856:
852:
846:
835:
829:
825:
821:
814:
810:
806:
793:
788:
777:
771:World War II
763:nomina dubia
758:
755:Francosuchus
754:
751:sister taxon
746:
742:
738:
734:
730:
722:
716:
706:
700:
696:Francosuchus
694:
691:angustifrons
687:
683:
681:
652:
648:Francosuchus
646:
634:
633:
628:
613:
612:remains, to
597:
561:
558:type species
553:
552:
547:
535:
531:
527:
524:angustifrons
523:
519:
511:
503:
502:fossils for
487:
477:
476:
471:
463:
457:
453:
450:Paleorhinus"
449:
445:
439:
435:Francosuchus
433:
429:
428:
423:
419:
415:
411:
405:
395:
389:paraphyletic
375:
374:
369:
360:
359:
346:
345:
343:type species
316:
311:
310:
309:
296:
295:
279:
278:
256:
255:
246:Type species
227:
226:
194:
181:
119:
24:
18:
3361:Paleorhinus
3276:Phytosaurus
3162:Nicrosaurus
3121:Smilosuchus
3114:Pravusuchus
3107:Leptosuchus
2998:Paleorhinus
2959:Heteropelta
2938:Phytosauria
2916:Phytosauria
2886:Phytosauria
2768:Phytosauria
2628:Paleorhinus
2469:Paleorhinus
2411:Case, 1922"
2409:Leptosuchus
2365:(1): 1–23.
1443:P. bransoni
1439:"P." sawini
1431:P. bransoni
1411:Phytosauria
1407:Paleorhinus
1395:Paleorhinus
1367:Paleorhinus
1363:Paleorhinus
1349:species to
1347:Paleorhinus
1331:Paleorhinus
1294:Nicrosaurus
1281:Phytosaurus
1263:maleriensis
1261:Phytosaurus
1252:maleriensis
1229:mixed with
1227:basicranium
1194:Paleorhinus
1186:P. bransoni
1178:P. bransoni
1162:Paleorhinus
1158:P. bransoni
1142:P. bransoni
1126:Paleorhinus
1122:"P." parvus
1118:P. bransoni
1094:Parasuchus,
1086:Paleorhinus
1041:Paleorhinus
1001:Paleorhinus
971:Paleorhinus
946:Paleorhinus
942:Paleorhinus
914:Paleorhinus
906:Paleorhinus
888:Paleorhinus
879:Archosauria
870:Phytosaurus
840:" arenaceus
811:P. bransoni
799:ontogenetic
747:P. bransoni
731:Paleorhinus
614:P. bransoni
610:postcranial
598:P. bransoni
584:, from the
562:Paleorhinus
554:P. bransoni
548:P. bransoni
536:Paleorhinus
528:Paleorhinus
496:rhynchosaur
488:Paleorhinus
472:Paleorhinus
456:(formerly "
446:Paleorhinus
424:Paleorhinus
412:Paleorhinus
376:Paleorhinus
370:Paleorhinus
312:Paleorhinus
297:P. bransoni
228:Paleorhinus
214:Phytosauria
25:Paleorhinus
3452:Phytosaurs
3446:Categories
3290:Suchoprion
3210:Centemodon
3005:Parasuchus
2895:see below↓
2847:Sauropsida
2832:Sauropsida
2816:Crurotarsi
2798:Sauropsida
2691:2013-06-25
2570:: 97–112.
2551:(1): 3–27.
2067:References
1451:Parasuchus
1403:P. ehlersi
1383:Parasuchus
1359:P. hislopi
1351:Parasuchus
1335:P. hislopi
1315:P. hislopi
1305:, nor the
1303:P. hislopi
1216:P. hislopi
1174:synonymous
1150:phytosaurs
1138:synonymous
1114:synonymous
1096:alongside
1047:fragment,
1017:F. broilii
993:E. neukami
963:E. neukami
831:Parasuchus
723:P. neukami
702:F. broilii
639:Oskar Kuhn
532:Parasuchus
444:, another
317:"Old Nose"
3376:Q21711013
3260:Omosaurus
2784:Kingdom:
2379:2056-2802
2279:2167-8359
2263:: e5901.
2233:130567475
2185:219192243
2117:140167568
1427:cladogram
1421:Phylogeny
1307:lectotype
1279:), or to
1254:. Later,
1063:Tetrapoda
1053:tetrapods
1003:subgenus
838:Zanclodon
826:E. broili
813:and also
767:holotypes
765:as their
699:species,
665:mandibles
474:fossils.
464:arenaceus
459:Zanclodon
454:arenaceus
380:taxonomic
331:phytosaur
234:Williston
151:Kingdom:
145:Eukaryota
3370:Wikidata
3336:Category
3076:Rutiodon
3062:Jupijkam
2792:Chordata
2790:Phylum:
2786:Animalia
2652:86386615
2602:Rutiodon
2584:84966794
2522:: 61–98.
2500:: 1–254.
2437:83536253
2387:83780331
2297:30581656
1299:holotype
1188:and the
1156:than to
1021:F. latus
988:F. latus
969:than to
729:than to
713:synonyms
711:, to be
708:F. latus
657:holotype
596:P217 to
574:holotype
514:forms a
319:) is an
315:(Greek:
288:, 1936 )
268:Species
175:Reptilia
165:Chordata
161:Phylum:
155:Animalia
141:Domain:
3416:4128961
3403:1348480
3390:4817565
3203:Belodon
3069:Protome
2796:Class:
2713:: 1–84.
2626:is not
2288:6292387
2165:Bibcode
2097:Bibcode
1467:
1339:neotype
1220:syntype
1045:rostrum
858:Belodon
818:neukami
673:Carnian
669:Bavaria
661:rostrum
586:Carnian
556:is the
542:Species
500:neotype
492:chimera
402:Morocco
366:Bavaria
352:Wyoming
339:Carnian
321:extinct
220:Genus:
207:Order:
171:Class:
3012:Wannia
2650:
2582:
2435:
2385:
2377:
2295:
2285:
2277:
2231:
2183:
2115:
1437:, and
1199:Wannia
803:Poland
675:-aged
588:-aged
468:Poland
404:) and
400:(from
337:(late
3429:58928
3398:IRMNG
2812:Clade
2803:Clade
2685:(PDF)
2670:(PDF)
2648:S2CID
2622:'
2580:S2CID
2541:(PDF)
2467:"The
2433:S2CID
2383:S2CID
2321:(PDF)
2257:PeerJ
2229:S2CID
2181:S2CID
2113:S2CID
1204:basal
1182:basal
1176:with
1140:with
1116:with
578:skull
518:with
516:clade
484:India
364:from
356:Texas
328:basal
324:genus
195:Clade
182:Clade
3385:GBIF
2872:see
2845:see
2375:ISSN
2293:PMID
2275:ISSN
1445:and
1373:and
1355:ICZN
1343:ICZN
1100:and
1019:and
995:and
985:and
867:and
795:Dzik
705:and
663:and
643:1936
570:1904
522:and
452:cf.
385:lump
354:and
286:Kuhn
238:1904
44:PreꞒ
2711:321
2640:doi
2604:".
2572:doi
2423:doi
2367:doi
2283:PMC
2265:doi
2221:doi
2217:103
2211:".
2173:doi
2161:379
2105:doi
1361:to
1309:of
1301:of
1291:of
1283:(a
999:to
820:or
805:to
715:of
686:as
641:in
594:PPM
568:in
560:of
3448::
3426::
3413::
3400::
3387::
3372::
3026:/
2814::
2805::
2709:.
2678:21
2676:.
2672:.
2646:.
2636:33
2634:.
2592:^
2578:.
2568:75
2566:.
2549:64
2547:.
2543:.
2528:^
2520:83
2518:.
2506:^
2496:.
2479:34
2477:.
2473:.
2445:^
2431:.
2419:53
2417:.
2413:.
2395:^
2381:.
2373:.
2361:.
2357:.
2337:^
2329:60
2327:.
2323:.
2305:^
2291:.
2281:.
2273:.
2259:.
2255:.
2241:^
2227:.
2215:.
2193:^
2179:.
2171:.
2159:.
2125:^
2111:.
2103:.
2093:12
2091:.
2087:.
2075:^
1457:.
1433:,
916:.
861:,
851:.
773:.
651:.
538:.
462:"
414:.
236:,
197::
184::
94:Pg
38:Ma
33:,
3300:?
3271:?
3263:?
3220:?
2962:?
2760:e
2753:t
2746:v
2694:.
2654:.
2642::
2586:.
2574::
2498:4
2439:.
2425::
2389:.
2369::
2363:2
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2299:.
2267::
2261:6
2235:.
2223::
2187:.
2175::
2167::
2119:.
2107::
2099::
1416:.
1164:.
1128:.
1104:.
1070:.
1027:.
836:"
293:†
284:(
276:†
253:†
224:†
211:†
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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