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2210:
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1650:; they live a more solitary lifestyle than the other great apes. Bornean orangutans are generally more solitary than Sumatran orangutans. Most social bonds occur between adult females and their dependent and weaned offspring. Resident females live with their offspring in defined home ranges that overlap with those of other adult females, which may be their immediate relatives. One to several resident female home ranges are encompassed within the home range of a resident male, who is their main mating partner. Interactions between adult females range from friendly to avoidance to antagonistic. Flanged males are hostile to both other flanged males and unflanged males, while unflanged males are more peaceful towards each other.
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1876:
infant has with its mother declines. When an orangutan reaches the age of one-and-a-half years, its climbing skills improve and it will travel through the canopy holding hands with other orangutans, a behaviour known as "buddy travel". After two years of age, juvenile orangutans will begin to move away from their mothers temporarily. They reach adolescence at six or seven years of age and are able to live alone but retain some connections with their mothers. Females may nurse their offspring for up to eight years, which is more than any other mammal. Typically, orangutans live over 30 years both in the wild and in captivity.
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1995:
1342:
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1804:
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1885:
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1203:
142:
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2011:
according to the task at hand, and preference was given to oral tool use. This preference was also found in an experimental study of captive orangutans. Orangutans have been observed to use sticks to poke at catfish, causing them to leap out of the water so the orangutan can grab them. Orangutan have also been documented to keep tools for later. When building a nest, orangutans appear to be able to determine which branches would better support their body weight.
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1978:-like vocalisations in response to physical contact such as wrestling, play chasing or tickling. This suggests that laughter derived from a common origin among primate species and therefore evolved before the origin of humans. Orangutans can learn to mimic new sounds by purposely controlling the vibrations of their vocal folds, a trait that led to speech in humans.
1658:
his home range. Both resident and transient orangutans aggregate on large fruiting trees to feed. The fruits tend to be abundant, so competition is low and individuals may engage in social interactions. Orangutans will also form travelling groups with members moving between different food sources. They are often consortships between an adult male and a female.
1782:
with a series of low frequency noises known collectively as the "rolling call". When uncomfortable, an orangutan will produce a "kiss squeak", which involves sucking in air through pursed lips. Mothers produce throatscrapes to keep in contact with their offspring. Infants make soft hoots when distressed. When building a nest, orangutans will produce smacks or
1922:
2023:
relatively tolerant of each other; this creates favourable conditions for the spreading of new behaviours. Further evidence that highly social orangutans are more likely to exhibit cultural behaviours came from a study of leaf-carrying behaviours of formerly captive orangutans that were being rehabilitated on the island of Kaja in Borneo.
2600:
In May 2017, the BOS rescued an albino orangutan from captivity in a remote village in Kapuas Hulu, on the island of
Kalimantan in Indonesian Borneo. According to volunteers at BOS, albino orangutans are extremely rare (one in ten thousand). This is the first albino orangutan the organisation has seen in 25 years of activity.
2032:
differences were cultural: first, the extent of the differences increased with distance, suggesting cultural diffusion was occurring, and second, the size of the orangutans' cultural repertoire increased according to the amount of social contact present within the group. Social contact facilitates cultural transmission.
1901:
orangutan then does the same to smaller, leafier branches to create a "mattress". After this, the ape stands and braids the tips of branches into the mattress. Doing this increases the stability of the nest. Orangutans make their nests more comfortable by creating "pillows", "blankets", "roofs" and "bunk-beds".
1407:
for a strong grip on branches as they travel high in the trees. The resting configuration of the fingers is curved, creating a suspensory hook grip. With the thumb out of the way, the fingers (and hands) can grip securely around objects with a small diameter by resting the tops of the fingers against
2010:
in ex-captive populations. Orangutans in Suaq
Balimbing were recorded to develop a tool kit for use in foraging which consisted of both insect-extraction sticks for use in the hollows of trees and seed-extraction sticks for harvesting seeds from hard-husked fruit. The orangutans adjusted their tools
490:
adult males develop distinctive cheek pads or flanges and make long calls that attract females and intimidate rivals; younger subordinate males do not and more resemble adult females. Orangutans are the most solitary of the great apes: social bonds occur primarily between mothers and their dependent
2967:
Anthropometamorphosis: man transform'd: or, The artificiall changling, historically presented, in the mad and cruel gallantry, foolish bravery, ridiculous beauty, filthy finenesse, and loathsome loveliness of most nations, fashioning and altering their bodies from the mould intended by nature; with
2599:
A female orangutan was rescued from a village brothel in Kareng Pangi village, Central
Kalimantan, in 2003. The orangutan was shaved and chained for sexual purposes. Since being freed, the orangutan, named Pony, has been living with the BOS. She has been re-socialised to live with other orangutans.
2031:
the listener into believing they are larger animals. In 2003, researchers from six different orangutan field sites who used the same behavioural coding scheme compared the behaviours of the animals from each site. They found each orangutan population used different tools. The evidence suggested the
1875:
Females do most of the caring of the young. The mother will carry the infant while travelling, suckle it and sleep with it. During its first four months, the infant is almost never without physical contact and clings to its mother's belly. In the following months, the amount of physical contact the
1657:
and establish their home ranges by age 11. Females tend to live near their birth range, while males disperse farther but may still visit their birth range within their larger home range. They enter a transient phase, which lasts until a male can challenge and displace a dominant, resident male from
939:
that the
Sumatran and Bornean species diverged 4.9 to 2.9 mya. By contrast, the 2011 genome study suggested that these two species diverged as recently as circa 400,000 years ago. The study also found that orangutans evolved at a slower pace than both chimpanzees and humans. A 2017 genome
1789:
Mother orangutans and offspring also use several different gestures and expressions such as beckoning, stomping, lower lip pushing, object shaking and "presenting" a body part. These communicate goals such as "acquire object", "climb on me", "climb on you", "climb over", "move away", "play change:
1422:
Compared to their relatives in Borneo, Sumatran orangutans are more slender with paler and longer hair and a longer face. Tapanuli orangutans resemble
Sumatran orangutans more than Bornean orangutans in body build and hair colour. They have shaggier hair, smaller skulls, and flatter faces than the
1781:
Orangutans communicate with various vocals and sounds. Males will make long calls, both to attract females and to advertise themselves to other males. These calls have three components; they begin with grumbles, peak with pulses and end with bubbles. Both sexes will try to intimidate conspecifics
510:
and construct elaborate sleeping nests each night from branches and foliage. The apes' learning abilities have been studied extensively. There may be distinctive cultures within populations. Orangutans have been featured in literature and art since at least the 18th century, particularly in works
1867:
to signal fertility. A female first gives birth around 15 years of age and they have a six to nine year interbirth interval, the longest among the great apes. Gestation is around nine months long and infants are born at a weight of 1.5–2 kg (3.3–4.4 lb). Usually only a single infant is
2139:
in other parts of Borneo and in
Sumatra. While some communities hunted them for food and decoration, others placed taboos on such practices. In central Borneo, some traditional folk beliefs consider it bad luck to look an orangutan in the face. Some folk tales involve orangutans mating with and
2559:
trade and bones are secretly sold in souvenir shops in several cities in
Indonesian Borneo. Conflicts between locals and orangutans also pose a threat. Orangutans that have lost their homes often raid agricultural areas and end up being killed by villagers. Locals may also be motivated to kill
2022:
fruit. The orangutans of Suaq
Balimbing were found to be avid users of insect and seed-extraction tools when compared to other wild orangutans. The scientists suggested these differences are cultural as they do not correlate with habitat. The orangutans at Suaq Balimbing are closely spaced and
1900:
Construction of a night nest is done by following a sequence of steps. Initially, a suitable tree is located. Orangutans are choosy about sites, though nests can be found in many tree species. To establish a foundation, the ape grabs the large branches under it and bends them so they join. The
647:
variety of Malay, but the age of the Old
Javanese sources mentioned above make Old Malay a more likely origin for the term. Cribb and colleagues (2014) suggest that Bontius' account referred not to apes (as this description was from Java where the apes were not known to be from) but to humans
3536:
Locke, D. P.; Hillier, L. W.; Warren, W. C.; Worley, K. C.; Nazareth, L. V.; Muzny, D. M.; Yang, S. P.; Wang, Z.; Chinwalla, A. T.; Minx, P.; Mitreva, M.; Cook, L.; Delehaunty, K. D.; Fronick, C.; Schmidt, H.; Fulton, L. A.; Fulton, R. S.; Nelson, J. O.; Magrini, V.; Pohl, C.; Graves, T. A.;
2563:
Estimates in the 2000s found that around 6,500 Sumatran orangutans and around 54,000 Bornean orangutans remain in the wild. A 2016 study estimates a population of 14,613 Sumatran orangutans in the wild, twice that of previous population estimates, while 2016 estimates suggest 104,700 Bornean
1896:
specialised for either day or night use. These are carefully constructed; young orangutans learn from observing their mother's nest-building behaviour. In fact, nest-building allows young orangutans to become less dependent on their mother. From six months of age onwards, orangutans practise
1584:
Orangutans also include leaves in their diet, which take up 25% of their average foraging time. Leaves are eaten more when fruit is less available, but even during times of fruit abundance, orangutans will eat leaves 11–20% of the time. They appear to depend on the leaf and stem material of
1482:
Most of the day is spent feeding, resting, and travelling. They start the day feeding for two to three hours in the morning. They rest during midday, then travel in the late afternoon. When evening arrives, they prepare their nests for the night. Potential predators of orangutans include
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1439:
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van
Noordwijk, Maria A.; Sauren, Simone E.B.; Nuzuar; Abulani, Ahbam; Morrogh-Bernard, Helen C.; Atmoko, S. Suci Utami; van Schaik, Carel P. (2009). "Development of Independence". In Wich, Serge A.; Atmoko, S. Suci Utami; Setia, Tatang Mitra; van Schaik, Carel P. (eds.).
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orangutans for food or because of their perceived danger. Mother orangutans are killed so their infants can be sold as pets. Between 2012 and 2017, the Indonesian authorities, with the aid of the Orangutan Information Center, seized 114 orangutans, 39 of which were pets.
1353:; females typically stand 115 cm (45 in) tall and weigh around 37 kg (82 lb), while adult males stand 137 cm (54 in) tall and weigh 75 kg (165 lb). Compared to humans, they have proportionally long arms, a male orangutan having an
940:
study found that the Bornean and Tapanuli orangutans diverged from Sumatran orangutans about 3.4 mya, and from each other around 2.4 mya. Millions of years ago, orangutans travelled from mainland Asia to Sumatra and then Borneo as the islands were connected by
643:. He reported that Malays claimed the ape could talk, but preferred not to "lest he be compelled to labour". The word appeared in several German-language descriptions of Indonesian zoology in the 17th century. It has been argued that the word comes specifically from the
485:
of the great apes, orangutans spend most of their time in trees. They have proportionally long arms and short legs, and have reddish-brown hair covering their bodies. Adult males weigh about 75 kg (165 lb), while females reach about 37 kg (82 lb).
1750:
1721:
1692:
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Little was known about orangutan behaviour until the field studies of Birutė Galdikas, who became a leading authority on the apes. When she arrived in Borneo in 1971, Galdikas settled into a primitive bark-and-thatch hut at a site she dubbed Camp Leakey, in
2564:
orangutans exist in the wild. A 2018 study found that Bornean orangutans declined by 148,500 individuals from 1999 to 2015. Fewer than 800 Tapanuli orangutans are estimated to still exist, which puts the species among the most endangered of the great apes.
5407:
Wich, Serge A.; de Vries, Hans; Ancrenaz, Marc; Perkins, Lori; Shumaker, Robert W.; Suzuki, Akira; van Schaik, Carel P. (2009). "Orangutan life history variation". In Wich, Serge A.; Atmoko, S. Suci Utami; Setia, Tatang Mitra; van Schaik, Carel P. (eds.).
1872:. This may be because they cannot ensure they will sire a female's next offspring, because she does not immediately begin ovulating again after her infant dies. There is evidence that females with offspring under six years old generally avoid adult males.
1928:
1926:
1924:
1959:. Without any training, the orangutans succeeded in pulling off an object to get food in the first session. Over the course of 30 sessions, the apes succeeded more quickly, having learned to coordinate. An adult orangutan has been documented to pass the
2458:
Galdikas reported that her cook was sexually assaulted by a captive male orangutan. The ape may have suffered from a skewed species identity and forced copulation is a standard mating strategy for low-ranking male orangutans. American animal trafficker
2442:
groups have urged a stop to such acts, considering them abusive. Starting in the 1960s, zoos became more concerned with education and orangutans' exhibits were designed to mimic their natural environment and let them display their natural behaviours.
1436:
2147:, who observed the animals and dissected some specimens, gave the first scientific description of the orangutan. Camper mistakenly thought that flanged and unflanged male orangutans were different species, a misconception corrected after his death.
1927:
4877:
Atmoko, S. Suci Utami; Singleton, Ian; van Noordwijk, Maria A.; van Schaik, Carel P.; Setia, Tatang Mitra (2009). "Male–male relationships in orangutans". In Wich, Serge A.; Atmoko, S. Suci Utami; Setia, Tatang Mitra; van Schaik, Carel P. (eds.).
2968:
figures of those transformations. To which artificial and affected deformations are added, all the native and national monstrosities that have appeared to disfigure the humane fabrick. With a vindication of the regular beauty and honesty of nature
4675:
Zimmermann, N; Pirovino, M; Zingg, R; Clauss, M; Kaup, F. J.; Heistermann, M; Hatt, J. M.; Steinmetz, H. W. (2011). "Upper respiratory tract disease in captive orangutans (Pongo sp.): prevalence in 20 European zoos and predisposing factors".
1408:
the inside of the palm, thus creating a double-locked grip. Their feet have four long toes and an opposable big toe, giving them hand-like dexterity. The hip joints also allow for their legs to rotate similarly to their arms and shoulders.
6913:
2502:, which prohibits their unlicensed trade under international law. The Bornean orangutan range has become more fragmented, with few or no apes documented in the southeast. The largest remaining population is found in the forest around the
1832:
by not developing the distinctive cheek pads, pronounced throat pouches, long fur, or long calls until a resident dominant male is absent. The transformation from unflanged to flanged can occur quickly. Flanged males attract females in
5754:
2256:(1895), the protagonist Reuben Power travels to Borneo and captures an orangutan to train it to speak so he can "know what a beast like that might think of us". Orangutans are featured prominently in the 1963 science fiction novel
2329:
features an intelligent orangutan which serves a university professor but has sinister motives; he plots against humanity and stalks a female student assistant. Other stories have portrayed orangutans helping humans, such as
1925:
7962:
Wich, Serge A.; Singleton, Ian; Nowak, Matthew G.; Utami Atmoko, Sri Suci; Nisam, Gonda; Arif, Sugesti Mhd.; Putra, Rudi H.; Ardi, Rio; Fredriksson, Gabriella; Usher, Graham; Gaveau, David L. A.; Kühl, Hjalmar S. (2016).
4574:
Foitová, Ivona; Huffman, Michael A.; Wisnu, Nurcahyo; Olšanský, Milan (2009). "Parasites and their impacts on orangutan health". In Wich, Serge A.; Atmoko, S. Suci Utami; Setia, Tatang Mitra; van Schaik, Carel P. (eds.).
1974:: mother orangutans remain silent in the presence of a perceived threat but when it passes, the mother produces an alarm call to their offspring to teach them about the danger. Orangutans and other great apes show
3961:
Wang, Cui-Bin; Zhao, Ling-Xia; Jin, Chang-Zhu; Wang, Yuan; Qin, Da-Gong; Pan, Wen-Shi (December 2014). "New discovery of Early Pleistocene orangutan fossils from Sanhe Cave in Chongzuo, Guangxi, southern China".
1748:
1719:
1690:
7654:
Cheyne, S. M.; Thompson, C. J.; Phillips, A. C.; Hill, R. M.; Limin, S. H. (2007). "Density and population estimate of gibbons (Hylobates albibarbis) in the Sabangau catchment, Central Kalimantan, Indonesia".
4023:
Ibrahim, Yasamin Kh.; Tshen, Lim Tze; Westaway, Kira E.; Cranbrook, Earl of; Humphrey, Louise; Muhammad, Ros Fatihah; Zhao, Jian-xin; Peng, Lee Chai (December 2013). "First discovery of Pleistocene orangutan
6784:
van Schaik, C. P.; Ancrenaz, M.; Borgen, G.; Galdikas, B.; Knott, C. D.; Singleton, I.; Suzuki, A.; Utami, S. S.; Merrill, M.; et al. (2003). "Orangutan cultures and the evolution of material culture".
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2026:
Wild orangutans in Tuanan, Borneo, were reported to use tools in acoustic communication. They use leaves to amplify the kiss squeak sounds they produce. The apes may employ this method of amplification to
1749:
1720:
1691:
3248:
1475:, while drier forests away from the flooded areas have fewer apes. Population density also decreases at higher elevations. Orangutans occasionally enter grasslands, cultivated fields, gardens, young
7074:
924:
between 24.1 and 19.7 million years ago (mya), and the orangutans diverged from the African great ape lineage between 19.3 and 15.7 mya. Israfil and colleagues (2011) estimated based on
6897:
3685:"Unresolved molecular phylogenies of gibbons and siamangs (Family: Hylobatidae) based on mitochondrial, Y-linked, and X-linked loci indicate a rapid Miocene radiation or sudden vicariance event"
2018:
and biological anthropologist Cheryl D. Knott further investigated tool use in different wild orangutan populations. They compared geographic variations in tool use related to the processing of
3997:
Schwartz, J. H.; Vu The Long; Nguyen Lan Cuong; Le Trung Kha; Tattersall, I. (1995). "A review of the Pleistocene hominoid fauna of the Socialist Republic of Vietnam (excluding Hylobatidae)".
2248:. The novel satirises the class and political system of Britain. Oran's purity and status as a "natural man" stands in contrast to the immorality and corruption of the "civilised" humans. In
1329:
date to between 66 and 57,000 years ago. The range of orangutans had contracted significantly by the end of the Pleistocene, most likely because of the reduction of forest habitat during the
2228:(The Bell of Nature, 1772) is told from the point of view of a human-orangutan hybrid who calls himself the "metaphysician of the woods". Around 50 years later, the anonymously written work
6433:
Russon, A. E.; Compost, A.; Kuncoro, P.; Ferisa, A. (2014). "Orangutan Fish Eating, Primate Aquatic Fauna Eating, and Their Implications for the Origins of Ancestral Hominin Fish Eating".
897:), despite populations being six to seven times higher in Borneo. The researchers hope these data may help conservationists preserve the endangered ape, as well as learn more about human
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1844:
on her, the occurrence of which is unusually high among mammals. Females prefer to mate with the fitter flanged males, forming pairs with them and benefiting from their protection. Non-
948:
when sea levels were much lower. The present range of Tapanuli orangutans is thought to be close to where ancestral orangutans first entered what is now Indonesia from mainland Asia.
1435:
1970:
Studies in the wild indicate that flanged male orangutans plan their movements in advance and signal them to other individuals. Experiments have also suggested that orangutans can
8075:
Nater, Alexander; Mattle-Greminger, Maja P.; Nurcahyo, Anton; Nowak, Matthew G.; Manuel, Marc de; Desai, Tariq; Groves, Colin; Pybus, Marc; Sonay, Tugce Bilgin (2 November 2017).
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6944:
5509:
Scott, A. M.; Knott, C. D.; Susanto, T. W. (2019). "Are Male Orangutans a Threat to Infants? Evidence of Mother–Offspring Counterstrategies to Infanticide in Bornean Orangutans (
2103:. Animal rights groups like Great Ape Project Argentina argued the ruling should apply to all species in captivity, and legal specialists from the Argentina's Federal Chamber of
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1923:
273:
250:
1260:, which lived 5–7 mya in Thailand, is believed to be the closest known relative of the living orangutans and inhabited similar environments. The largest known primate,
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Europeans became aware of the existence of the orangutan in the 17th century. Explorers in Borneo hunted them extensively during the 19th century. In 1779, Dutch anatomist
8224:
4763:
Teboekhorst, I.; Schürmann, C.; Sugardjito, J. (1990). "Residential status and seasonal movements of wild orang-utans in the Gunung Leuser Reserve (Sumatera, Indonesia)".
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Wich, S. A.; Swartz, K. B.; Hardus, M. E.; Lameira, A. R.; Stromberg, E.; Shumaker, R. W. (2008). "A case of spontaneous acquisition of a human sound by an orangutan".
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who had been observing Rakus at a nature preserve, "Five days later the facial wound was closed, while within a few weeks it had healed, leaving only a small scar".
1741:
1712:
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Xu, X.; Arnason, U. (1996). "The mitochondrial DNA molecule of sumatran orangutan and a molecular proposal for two (Bornean and Sumatran) species of orangutan".
1415:. Compared to other great apes, they infrequently descend to the ground where they are more cumbersome. Unlike gorillas and chimpanzees, orangutans are not true
1253:, which lived in India and Pakistan from 12.5 mya until 8.5 mya. These animals likely lived in drier and cooler environments than orangutans do today.
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6308:
Fox, E. A.; Sitompul, A. F.; van Schaik, C. P. (1999). "Intelligent tool use in wild Sumatran orangutans". In Parker, S; Mitchell, R. W.; Miles, H. L. (eds.).
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582:, meaning "forest". The locals originally used the term for actual forest-dwelling humans, but the word underwent a semantic extension to include apes of the
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5311:"Female reproductive strategies in orangutans, evidence for female choice and counterstrategies to infanticide in a species with frequent sexual coercion"
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have been found in Vietnam, and multiple fossil subspecies have been described from several parts of southeastern Asia. It is unclear if these belong to
7122:
2224:
Orangutans first appeared in Western fiction in the 18th century and have been used to comment on human society. Written by the pseudonymous A. Ardra,
1952:", which involves an individual aiding another with the expectation of being paid back. Orangutans are the first nonhuman species documented to do so.
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van Schaik, C. P.; Fox, E. A.; Sitompul, A. F. (1996). "Manufacture and use of tools in wild Sumatran orangutans – implications or human evolution".
2391:. He rejected the company of other animals, aside from a dog, and preferred to be with humans. He was occasionally taken on coach rides clothed in a
2083:, orangutans, and gorillas not to be used for animal experiments. In December 2014, a court in Argentina ruled that an orangutan named Sandra at the
746:
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Mitani, J. C.; Grether, G. F.; Rodman, P. S.; Priatna, D. (1991). "Associations among wild orang-utans: sociality, passive aggregations or chance".
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2506:, but this environment is at risk. The Sumatran orangutan is found only in the northern part of Sumatra, most of the population inhabiting the
671:
are found in many languages. This spelling (and pronunciation) has remained in use in English up to the present but has come to be regarded as
2163:. Galdikas became an outspoken advocate for orangutans and the preservation of their rainforest habitat, which is rapidly being devastated by
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Russon, AE; Handayani, DP; Kuncoro, P; Ferisa, A. (2007). "Orangutan leaf-carrying for nest-building: toward unraveling cultural processes".
1552:, which can take up 57–80% of their foraging time. Even during times of scarcity, fruit is 16% of their feeding time. Fruits with soft pulp,
4065:
Liao, Wei; Harrison, Terry; Yao, Yanyan; Liang, Hua; Tian, Chun; Feng, Yuexing; Li, Sheng; Bae, Christopher J.; Wang, Wei (September 2022).
1380:
of an orangutan is underdeveloped. Females and juveniles have relatively circular skulls and thin faces while mature males have a prominent
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of about 2 m (6 ft 7 in), and short legs. They are covered in long reddish hair that starts out bright orange and darkens to
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683:. In Malay, the term was first attested in 1840, not as an indigenous name but referring to how the English called the animal. The word '
7901:
2075:
In June 2008, Spain would become the first country to recognise the rights of some non-human great apes, based on the guidelines of the
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2447:, an orangutan of the San Diego Zoo, became world famous in the 1980s for multiple escapes from his enclosures. He was nicknamed "the
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Bradon-Jones, D.; Eudey, A. A.; Geissmann, T.; Groves, C. P.; Melnick, D. J.; Morales, J. C.; Shekelle, M.; Stewart, C. B. (2004).
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By the early 19th century, orangutans were being kept in captivity. In 1817, an orangutan joined several other animals in London's
1983:
768:
3089:
Rubis, June Mary (2020). "The orang utan is not an indigenous name: knowing and naming the maias as a decolonizing epistemology".
2627:
in Sumatra. In Malaysia, conservation areas include Semenggoh Wildlife Centre and Matang Wildlife Centre also in Sarawak, and the
13222:
8495:
8138:
3128:
753:
8398:
8214:
5620:
Didik, Prasetyo; Ancrenaz, Marc; Morrogh-Bernard, Helen C.; Atmoko, S. Suci Utami; Wich, Serge A.; van Schaik, Carel P. (2009).
2827:
9243:
9141:
7931:
Wich, S A; Meijaard, E; Marshall, A J; Husson, S; et al. (2002). "Distribution and conservation status of the orang-utan (
7251:
2584:
Several organisations are working for the rescue, rehabilitation and reintroduction of orangutans. The largest of these is the
8367:
5161:"Orangutan information broadcast via consonant-like and vowel-like calls breaches mathematical models of linguistic evolution"
3216:
13240:
9151:
8339:
7388:
7163:
6317:
5633:
4887:
4439:
4352:
4176:
3692:
3389:
2948:
2869:
2245:
1388:. The cheek pads are made mostly of fatty tissue and are supported by the musculature of the face. The throat pouches act as
13227:
7203:
Studies in History and Philosophy of Science Part C: Studies in History and Philosophy of Biological and Biomedical Sciences
6281:
Galdikas, B. M. F. (1982). "Orang-Utan tool use at Tanjung Putting Reserve, Central Indonesian Borneo (Kalimantan Tengah)".
4848:
Galdikas, B. M. F. (1984). "Adult female sociality among wild orangutans at Tanjung Puting Reserve". In Small, M. F. (ed.).
620:
refers only to the apes and not forest-dwelling humans. The word is not originally Javanese, but was borrowed from an early
9831:
7040:
Race and Aesthetics in the Anthropology of Petrus Camper (1722–1789) (Studies in the History of Ideas in the Low Countries)
4982:
van Schaik, C. P.; Preuschoft, S.; Watts, D. P. (2004). "Great ape social systems". In Russon, A. E.; Begun, D. R. (eds.).
3727:
3472:
2467:
to orphaned orangutan babies in hopes of keeping them alive long enough to sell to a trader, which would be an instance of
2494:
of mammals. They are legally protected from capture, harm or killing in both Malaysia and Indonesia, and are listed under
2232:
is narrated by a pure orangutan in captivity in the US, writing a letter critiquing Boston society to her friend in Java.
2087:
must be moved to a sanctuary in Brazil to provide her "partial or controlled freedom". Sandra has since been relocated to
1986:, was recorded spontaneously whistling after hearing a caretaker. She appears to whistle without expecting a food reward.
1365:
with age, while the skin is grey-black. Though largely hairless, males' faces can develop some hair, giving them a beard.
8251:
8160:
5269:
4957:
2616:
5901:
Chalmeau, Raphaël; Lardeux, Karine; Brandibas, Pierre; Gallo, Alain (1997). "Cooperative problem solving by orangutans (
3537:
Markovic, C.; Cree, A.; Dinh, H. H.; Hume, J.; Kovar, C. L.; Fowler, G. R.; Lunter, G.; Meader, S.; et al. (2011).
2536:
when she began studying them in 1971. By the 2000s, orangutan habitats decreased rapidly because of logging, mining and
522:. Human activities have caused severe declines in populations and ranges. Threats to wild orangutan populations include
9073:
8312:
7126:
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5003:
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4861:
4592:
4558:
3664:
3155:
2636:
6967:
7452:
7425:
4201:
3789:
1944:
has a touch-screen computer on which their two Sumatran orangutans play games. A 2008 study of two orangutans at the
1786:. Orangutan calls display consonant- and vowel-like components and they maintain their meaning over great distances.
7632:
6560:
van Casteren, A.; Sellers, W. I.; Thorpe, S. K. S.; Coward, S.; Crompton, R. H.; Myatt, J. P.; Ennos, A. R. (2012).
3779:
13337:
8277:
8192:
3186:
2522:
719:
7304:
4608:
Morrogh-Bernard HC, Foitová I, Yeen Z, Wilkin P, de Martin R, Rárová L, Doležal K, Nurcahyo W, Olšanský M (2017).
115:
13347:
13253:
9083:
7012:
6622:
van Schaik, Carel P.; Knott, Cheryl D. (2001). "Geographic variation in tool use onNeesia fruits in orangutans".
4725:
4254:
1610:, which involves consuming soil and other earth substances. They will uproot soil from the ground as well as eat
820:
was confirmed as a full species based on molecular evidence published in 1996, and three distinct populations on
13322:
8800:
8453:
3296:
2940:
2468:
1849:
710:, which describes two anthropoid "monsters" named Pongo and Engeco. He is now believed to have been describing
8421:
1544:
Although orangutans may consume leaves, shoots, and bird eggs, fruit is the most important part of their diet.
885:
to have its genome sequenced. Subsequently, the Bornean species had its genome sequenced. Bornean orangutans (
570:
The name "orangutan" (also written orang-utan, orang utan, orangutang, and ourang-outang) is derived from the
438:, orangutans were originally considered to be one species. From 1996, they were divided into two species: the
13292:
9412:
2858:
Malay Words and Malay Things: Lexical Souvenirs from an Exotic Archipelago in German Publications Before 1700
2460:
2297:
2214:
2204:
1841:
5725:"Do some taxa have better domain-general cognition than others? A meta-analysis of nonhuman primate studies"
4803:
Singleton, I.; van Schaik, C. P. (2002). "The Social Organisation of a population of Sumatran orang-utans".
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2209:
491:
offspring. Fruit is the most important component of an orangutan's diet, but they will also eat vegetation,
9452:
5129:
2766:
2640:
2632:
2628:
1956:
13332:
13235:
13144:
11244:
9783:
8488:
5940:
Suárez, S. D.; Gallup, G. G. (1981). "Self-recognition in chimpanzees and orangutans, but not gorillas".
5820:
4938:
Delgado, R. A. Jr.; van Schaik, C. P. (2000). "The behavioral ecology and conservation of the orangutan (
2620:
2612:
2306:
7586:
4514:
Galdikas, Birute M. F. (1988). "Orangutan Diet, Range, and Activity at Tanjung Puting, Central Borneo".
2592:
and which operates projects such as the Nyaru Menteng Rehabilitation Program founded by conservationist
1564:
and berries. Orangutans are thought to be the sole fruit disperser for some plant species including the
13149:
9778:
9280:
9270:
9236:
9156:
9008:
6841:"Active self-treatment of a facial wound with a biologically active plant by a male Sumatran orangutan"
4991:
4765:
4030:
3139:
2485:
2258:
1971:
1868:
born; twins are a rare occurrence. Unlike many other primates, male orangutans do not seem to practise
462:), was identified definitively in 2017. The orangutans are the only surviving species of the subfamily
20:
6469:
2127:
Orangutans were known to the native people of Sumatra and Borneo for millennia. The apes are known as
515:
and they have been kept in captive facilities around the world since at least the early 19th century.
9275:
9120:
9088:
9038:
8993:
7756:
4549:
Rodman, P. S. (1988). "Diversity and consistency in ecology and behavior". In Schwartz, J. H. (ed.).
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2585:
2415:
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1945:
1893:
857:
12331:
4217:
Hilloowala, R. A.; Trent, R. B. (1988). "Supraorbital ridge and masticatory apparatus I: Primates".
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2548:
plantations in response to international demand. Hunting is also a major problem, as is the illegal
2244:
features Sir Oran Haut Ton, an orangutan who lives among English people and becomes a candidate for
507:
9824:
7544:
7413:
7151:
Centralizing Fieldwork: Critical Perspectives from Primatology, Biological, and Social Anthropology
3926:
2861:
2707:
2403:
who was dressed in human clothing and learned to drink tea. She is remembered for her meeting with
2249:
1848:
females do not usually resist copulation with unflanged males, as the chance of conception is low.
806:
by Lacépède in 1799. The populations on the two islands were suggested to be separate species when
9018:
8473:
2631:
in Sabah. Major conservation centres headquartered outside the orangutans' home countries include
2593:
1325:
that date to 60,000 years ago. The youngest remains from South China, which are teeth assigned to
141:
13053:
12089:
11907:
11314:
9265:
9114:
9093:
5619:
5133:
5107:
4066:
3280:
2685:
2302:
1949:
1468:
672:
635:
In Western sources, the first printed attestation of the word for the apes is in Dutch physician
624:
at least a thousand years ago. Hence the ultimate origin of the term "orangutan" as denoting the
559:
7562:
Nowak, Matthew G.; Rianti, Puji; Wich, Serge A.; Meijaard, Erik; Fredriksson, Gabriella (2017).
4168:
4162:
3602:
1937:
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is found as part of the faunal complex in the Pleistocene cave assemblage in Vietnam, alongside
13279:
13074:
11368:
11072:
10453:
9753:
9003:
8481:
4995:
4876:
4584:
4573:
3412:
Nater, A.; Mattle-Greminger, M. P.; Nurcahyo, A.; Nowak, M. G.; et al. (2 November 2017).
3381:
2608:
2423:
2070:
1266:, was also a member of Ponginae and lived in China, from 2 mya to 300,000 years ago.
8329:
7699:
7380:
6737:"Tool use in wild orang-utans modifies sound production: a functionally deceptive innovation?"
5624:. In Wich, Serge A.; Atmoko, S. Suci Utami; Setia, Tatang Mitra; van Schaik, Carel P. (eds.).
5621:
5309:
Knott, Cheryl Denise; Thompson, Melissa Emery; Stumpf, Rebecca M; McIntyre, Matthew H (2009).
5208:
Knox, A; Markx, J; How, E; Azis, A; Hobaiter, C; an Veen, F. J. F; Morrogh-Bernard, H (2019).
499:, insects and bird eggs. They can live over 30 years, both in the wild and in captivity.
13274:
12154:
9605:
9372:
9229:
6659:
van Schaik, CP; Van Noordwijk, MA; Wich, SA. (2006). "Innovation in wild Bornean orangutans (
6562:"Nest-building orangutans demonstrate engineering know-how to produce safe, comfortable beds"
3920:
Harrison, Terry; Jin, Changzhu; Zhang, Yingqi; Wang, Yuan; Zhu, Min (December 2014). "Fossil
3813:
Bhandari, A.; Kay, R. F.; Williams, B. A.; Tiwari, B. N.; Bajpai, S.; Heironymus, T. (2018).
3516:
2537:
2287:
2088:
1869:
1532:
as an anti-inflammatory balm. Captive animals may suffer an upper respiratory tract disease.
1488:
945:
555:
519:
13266:
7841:
Davis, J. T.; Mengersen, K.; Abram, N. K.; Ancrenaz, M.; Wells, J. A.; Meijaard, E. (2013).
7417:
7149:
3815:"First record of the Miocene hominoid Sivapithecus from Kutch, Gujarat state, western India"
3653:
Our vanishing relative: the status of wild orang-utans at the close of the twentieth century
1837:
with their characteristic long calls, which may also suppress development in younger males.
1815:
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13121:
11388:
9492:
9352:
9063:
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8958:
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6084:
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5571:
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4471:
3971:
3935:
3826:
3552:
3305:
2644:
2576:
Zoologische Gesellschaft Frankfurt Programm Director Peter Pratje works with orangutans in
2418:
and to perform tricks. Notable orangutan "character actors" include: Jacob and Rosa of the
2062:
2028:
1936:
Orangutans are among the most intelligent non-human primates. Experiments suggest they can
1654:
1614:
from tree trunks. Orangutans also visit the sides of cliffs or earth depressions for their
1540:
1372:
volume is 397 cm. The cranium is elevated relative to the face, which is incurved and
1330:
10535:
7781:
7563:
7482:
5263:
Fox, E. A. (2002). "Female tactics to reduce sexual harassment in the Sumatran orangutan (
2140:
kidnapping humans. There are even stories of hunters being captured by female orangutans.
8:
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12831:
12212:
11491:
11409:
11354:
9817:
9680:
9660:
9575:
9560:
9517:
9507:
9502:
9472:
9407:
9337:
9053:
9043:
8304:
7784:"Quantifying Killing of Orangutans and Human-Orangutan Conflict in Kalimantan, Indonesia"
6335:
4805:
4431:
4421:
4417:
4340:
4252:
Winkler, L. A. (1989). "Morphology and relationships of the orangutan fatty cheek pads".
4028:
sp.) fossils in Peninsular Malaysia: Biogeographic and paleoenvironmental implications".
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3485:
2982:
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2041:
1975:
1783:
1647:
1456:
1452:
1412:
1322:
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680:
535:
511:
that comment on human society. Field studies of the apes were pioneered by primatologist
487:
482:
7986:
7858:
7799:
7524:
Ancrenaz, M.; Gumal, M.; Marshall, A. J.; Meijaard, E.; Wich, S. A.; Husson, S. (2016).
6873:
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6798:
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6348:
6196:
6088:
6029:
5681:
5575:
5558:
Smith, Tanya M.; Austin, Christine; Hinde, Katie; Vogel, Erin R.; Arora, Manish (2017).
5019:
van Schaik, C. P. (1999). "The socioecology of fission-fusion sociality in orangutans".
4633:
4475:
3975:
3939:
3924:
from the Early Pleistocene Gigantopithecus fauna of Chongzuo, Guangxi, southern China".
3830:
3556:
3309:
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1855:
679:
and the shift from -ng to -n has been taken to suggest the term entered English through
12474:
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5851:
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3068:
3031:
2788:
2427:
2419:
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1979:
1916:
1840:
Unflanged males wander widely in search of oestrous females and upon finding one, will
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866:
838:
808:
455:
447:
349:
322:
308:
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136:
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Meijaard, E.; Buchori, B.; Hadiprakarsa, Y.; Utami-Atmoko, S. S.; et al. (2011).
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5953:
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Lameira, A. R.; Hardus, M. E.; Shumaker, R. W.; Wich, S. A.; Menken, S. B. J. (2015).
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2007:
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512:
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9512:
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9457:
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9367:
9342:
9078:
8921:
8817:
8597:
8394:
8335:
8308:
8116:
8108:
8049:
8028:"Global demand for natural resources eliminated more than 100,000 Bornean Orangutans"
8008:
7882:
7823:
7672:
7525:
7458:
7448:
7421:
7384:
7369:
7286:
7278:
7228:
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7043:
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6860:
6810:
6766:
6709:
6641:
6601:
6542:
6450:
6407:
6360:
6313:
6255:
6220:
6153:
6112:
6053:
6012:"Wild Orangutan Males Plan and Communicate Their Travel Direction One Day in Advance"
5984:
5883:
5852:"Calculated reciprocity after all: computation behind token transfers in orang-utans"
5801:
5724:
5705:
5663:
5629:
5599:
5544:
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5438:
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5410:
5381:
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4913:
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4822:
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4435:
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4385:
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and hat and even given drinks at an inn where he behaved politely for the hosts. The
2374:
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2345:
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1910:
1570:
1472:
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1350:
1274:
973:
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267:
123:
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5290:
5085:
5050:
4965:
4902:
4834:
4786:
4535:
4283:
4238:
3493:
3012:
Tan, Peter (October 1998). "Malay loan words across different dialects of English".
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12961:
12605:
12498:
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11807:
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10810:
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9555:
9382:
9377:
9177:
8947:
8371:
8098:
8088:
8061:
8039:
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7990:
7973:
7944:
7872:
7862:
7813:
7803:
7684:
7664:
7581:
7539:
7376:
7336:"The missing lynx and five other animal escapees including Ken Allen the Orangutan"
7210:
6997:
6868:
6852:
6802:
6756:
6748:
6721:
6701:
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6672:
6633:
6591:
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6532:
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6247:
6210:
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5914:
5873:
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5793:
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5695:
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5278:
5235:
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4814:
4774:
4734:
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4523:
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4389:
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4319:
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4226:
4128:
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4039:
4002:
3979:
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3709:
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2701:
2507:
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2084:
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2015:
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during times of low fruit abundance. Other food items consumed by the apes include
1528:
1510:
1476:
1464:
1460:
1389:
936:
898:
846:, came with a surprising twist: it is more closely related to the Bornean species,
603:
6267:
5776:
de Blois, S. T.; Novak, M. A.; Bond, M. (1998). "Object Permanence in Orangutans (
4705:
3102:
2410:
Zoos and circuses in the Western world would continue to use orangutans and other
1852:
has been recorded in the context of both affiliative and aggressive interactions.
1341:
877:
The Sumatran orangutan genome was sequenced in January 2011. Following humans and
13112:
12884:
12853:
12845:
12729:
12629:
12484:
12454:
12071:
11917:
11854:
11684:
11634:
11620:
11613:
11567:
11361:
11156:
10826:
10779:
10713:
10028:
9733:
9695:
9690:
9685:
9655:
9630:
9625:
9600:
9580:
9570:
9392:
9327:
9167:
8892:
8882:
8658:
8649:
7867:
7840:
7808:
6474:
6446:
6038:
5856:
5690:
5210:"Gesture use in communication between mothers and offspring in wild Orang-Utans (
4082:
4043:
3983:
3947:
3839:
3147:
3051:
2672:
2577:
2451:" and was the subject of a fan club, T-shirts, bumper stickers and a song titled
2388:
2335:
2311:
2292:
1994:
1864:
1659:
1419:, instead bending their digits and walking on the sides of their hands and feet.
1416:
1404:
1262:
913:
779:
644:
636:
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7215:
7198:
5797:
3705:
1321:
and mainland Southeast Asia and South China. Teeth of orangutans are known from
13010:
12971:
12902:
12759:
12749:
12719:
12696:
12686:
12676:
12588:
12537:
12522:
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9347:
9304:
7155:
6856:
6676:
6527:
6205:
5967:
Robert, S. (1986). "Ontogeny of mirror behavior in two species of great apes".
5741:
5526:
5230:
5209:
4642:
4427:
2648:
2503:
2404:
2152:
2036:
1964:
1863:
Unlike females of other non-human great ape species, orangutans do not exhibit
1611:
1484:
1381:
1301:
1255:
1233:
739:
727:
688:
571:
422:
315:
13126:
9442:
8370:. Australian Department of the Environment and Water Resources. Archived from
8167:
8093:
8076:
8044:
8027:
7668:
7199:"Going the whole orang: Darwin, Wallace and the natural history of orangutans"
6705:
6251:
6148:
6131:
5918:
5477:
5282:
5102:
Munn, C.; Fernandez, M. (1997). "Infant development". In Sodaro, Carol (ed.).
4484:
4459:
4324:
4307:
3683:
Israfil, H.; Zehr, S. M.; Mootnick, A. R.; Ruvolo, M.; Steiper, M. E. (2011).
3430:
3413:
3270:
3027:
2783:
1967:. Mirror tests with a 2-year-old orangutan failed to reveal self-recognition.
813:
13311:
12989:
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12544:
12514:
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12434:
12276:
12242:
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11183:
11176:
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11102:
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10645:
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9635:
9462:
9210:
8786:
8767:
8112:
7572:
7530:
7491:
7224:
7194:
6909:
6898:"Orangutan seen treating wound with medicinal herb in first for wild animals"
6864:
6432:
6386:
O'Malley, R. C.; McGrew, W. C. (2000). "Oral tool use by captive orangutans (
5406:
4303:
3775:
3014:
2533:
2491:
2448:
2439:
2435:
2263:
2218:
2144:
2092:
1999:
1627:
1592:
1501:
784:
703:
653:
598:
539:
492:
280:
66:
8077:"Morphometric, Behavioral, and Genomic Evidence for a New Orangutan Species"
7462:
6839:
Laumer, Isabelle B.; Rahman, Arif; Rahmaeti, Tri; et al. (2 May 2024).
6806:
6586:
5115:
4161:
van Schaik, C.; MacKinnon, J. (2001). "Orangutans". In MacDonald, D. (ed.).
3799:
3784:(3rd ed.). Baltimore: Johns Hopkins University Press. pp. 183–84.
3414:"Morphometric, Behavioral, and Genomic Evidence for a New Orangutan Species"
1884:
13097:
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9013:
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7886:
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6882:
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6713:
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6605:
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6411:
6259:
6224:
6157:
6116:
6097:
6057:
5988:
5980:
5887:
5869:
5709:
5603:
5584:
5495:
5385:
5360:
Fox, Elizabeth A (2001). "Homosexual behavior in wild Sumatran orangutans (
5346:
5327:
5194:
5176:
5042:
4981:
4826:
4697:
4661:
4493:
4403:
4275:
4267:
4090:
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3903:
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3723:
3582:
3449:
3124:
2695:
2624:
2589:
2282:
2192:
2180:
1615:
1385:
1333:. Though they may have survived into the Holocene in Cambodia and Vietnam.
1249:
1243:
933:
929:
714:, but in the 18th century, the terms orangutan and pongo were used for all
259:
7290:
6364:
5805:
3325:
2366:
2039:
in 2022, a male Sumatran orangutan, known to researchers as Rakus, chewed
1202:
13209:
13136:
13106:
12927:
12110:
11991:
11974:
11738:
11344:
11254:
11119:
11082:
11002:
10941:
10662:
10115:
9427:
9402:
9221:
9023:
8978:
8847:
8682:
7628:
7447:. Steven Lehrer. Lubbock, Tex.: Texas Tech University Press. p. 37.
7442:
6009:
3614:
2511:
2392:
2325:
2184:
2050:
1960:
1941:
1897:
nest-building and gain proficiency by the time they are three years old.
1526:
are reported in young individuals. Orangutans also use the plant species
1373:
1369:
1219:
The three orangutan species are the only extant members of the subfamily
1121:
941:
660:
471:
426:
418:
333:
41:
8368:"Tax Deductible Organisations (Register of Environmental Organisations)"
7700:"Endangered Orangutans: Fires, Poaching and Palm Oil: Facts and Details"
7603:"Endangered Orangutans Gain From Eco-Friendly Shifts in Palm Oil Market"
5849:
4986:
The Evolution of Thought: Evolutionary Origins of Great Ape Intelligence
4719:
Rijksen, H. D. (December 1978). "A field study on Sumatran orang utans (
4367:
3564:
3239:
3178:
2733:
2495:
1638:
13201:
12403:
12195:
11522:
11509:
11459:
11395:
11334:
11237:
11220:
11112:
11055:
11028:
10991:
10961:
10951:
10676:
10402:
10284:
9988:
9933:
9885:
9840:
9417:
9319:
9309:
9172:
8863:
8703:
8544:
8103:
7965:"Land-cover changes predict steep declines for the Sumatran orangutan (
7312:
6356:
5626:
Orangutans: Geographic Variation in Behavioral Ecology and Conservation
5412:
Orangutans: Geographic Variation in Behavioral Ecology and Conservation
5240:
5034:
4908:
Orangutans: Geographic Variation in Behavioral Ecology and Conservation
4880:
Orangutans: Geographic Variation in Behavioral Ecology and Conservation
4746:
4579:
Orangutans: Geographic Variation in Behavioral Ecology and Conservation
4527:
4230:
4140:
3894:
3873:
3440:
3317:
2690:
2549:
2396:
2316:
2281:
Orangutans are sometimes portrayed as antagonists, notably in the 1832
2271:
2172:
1773:
1600:
1578:
1557:
1377:
878:
562:
organisations are dedicated to the survival of orangutans in the wild.
551:
398:
342:
203:
86:
51:
6735:
Hardus, M. E.; Lameira, A. R.; van Schaik, C. P.; Wich, S. A. (2009).
6637:
6403:
5535:
4818:
3072:
13214:
13183:
12872:
12802:
12645:
12595:
12232:
12222:
12145:
11998:
11888:
11787:
11553:
11307:
11210:
10424:
10186:
9902:
9768:
9705:
9545:
9482:
9162:
8624:
8554:
8505:
8468:
7948:
7480:
5377:
2517:
2464:
2444:
2414:
as sources for entertainment, training them to behave like humans at
2340:
2275:
2096:
2049:
the mashed plant material to an open wound on his face. According to
1845:
1549:
1467:. Populations are more concentrated near riverside habitats, such as
1395:
1368:
Orangutans have small ears and noses; the ears are unlobed. The mean
1318:
1317:
had a far more extensive range than at present, extending throughout
1224:
1011:
882:
843:
715:
656:, who was the first to use the term in a publication a decade later.
649:
629:
402:
394:
223:
153:
91:
13068:
4132:
4006:
2885:
Dellios, Paulette (2008). "A lexical odyssey from the Malay World".
13188:
13091:
12938:
12914:
12707:
12666:
12612:
12551:
12124:
11274:
11167:
10867:
10792:
10692:
10277:
9917:
9858:
9131:
8672:
8563:
8526:
8503:
8219:
7905:
6132:"Reconstructing the Evolution of Laughter in Great Apes and Humans"
6016:
4738:
3064:
2666:
2556:
2545:
2529:
2176:
2168:
2155:. She studied orangutans for the next four years and developed her
2100:
1955:
In a 1997 study, two captive adult orangutans were tested with the
1834:
1607:
1575:
1496:
1354:
1278:
1220:
735:
612:
608:
543:
527:
523:
463:
406:
233:
173:
81:
76:
61:
56:
46:
8395:"Rehabilitation not enough to solve orangutan crisis in Indonesia"
8074:
4368:
Oishi, M.; Ogihara, N.; Endo, H.; Ichihara, N.; Asari, M. (2009).
3411:
2540:
by roads. A major factor has been the conversion of vast areas of
2115:
1403:
Orangutan hands have four long fingers but a dramatically shorter
1238:
13287:
12999:
12892:
12619:
12259:
12188:
11440:
11433:
11295:
11190:
10972:
10853:
10799:
10619:
10480:
10437:
9870:
9743:
9728:
9585:
9527:
9126:
9109:
8876:
8634:
8538:
5850:
Dufour, V.; Pelé, M.; Neumann, M.; Thierry, B.; Call, J. (2008).
4308:"Male bimaturism and reproductive success in Sumatran orangutans"
3781:
Mammal Species of the World: A Taxonomic and Geographic Reference
2528:
Birutė Galdikas wrote that orangutans were already threatened by
2352:. More comical portrayals of the orangutan include the 1996 film
2132:
1829:
1506:
1384:, large cheek pads or flanges, extensive throat pouches and long
1313:
or, in fact, represent distinct species. During the Pleistocene,
1296:
1094:
921:
902:
901:. Similarly to gorillas and chimpanzees, orangutans have 48
711:
547:
503:
467:
414:
193:
96:
71:
19:
This article is about the group of primates. For other uses, see
8215:"Orangutan shaved, made up and prostituted to men for six years"
7282:
6783:
6130:
Ross, Marina Davila; Owren, Michael J; Zimmermann, Elke (2009).
4958:
10.1002/1520-6505(2000)9:5<201::AID-EVAN2>3.0.CO;2-Y
4607:
4460:"Newly discovered orangutan species is also the most endangered"
3999:
Anthropological Papers of the American Museum of Natural History
3462:
2971:. Internet Archive (2nd ed.). London: W. Hunt. p. 437.
1828:
Males become sexually mature at around age 15. They may exhibit
1411:
Orangutans move through the trees by both vertical climbing and
12569:
12393:
12202:
12103:
11824:
11021:
10737:
10567:
10554:
10506:
9864:
9852:
9610:
9477:
9432:
9357:
9291:
8943:
8853:
8694:
8532:
8520:
8463:
7277:. Caldwell, Idaho: The Caxton Printers, Ltd. pp. 102–103.
2767:"The Word 'Orangutan': Old Malay Origin or European Concoction"
2603:
Other major conservation centres in Indonesia include those at
2411:
2320:
2080:
1761:
1732:
1703:
1646:
The social structure of the orangutan can be best described as
1623:
1619:
1358:
1045:
917:
821:
707:
410:
213:
183:
163:
13175:
7961:
6073:"Time-space–displaced responses in the orangutan vocal system"
4762:
4370:"Dimensions of forelimb muscles in orangutans and chimpanzees"
4302:
Utami, S. S.; Goossens, B.; Bruford, M. W.; de Ruiter, J. R.;
726:
for the genus in 1799. Battel's "Pongo", in turn, is from the
687:' in modern Malay and Indonesian was borrowed from English or
9896:
9809:
9650:
9595:
9540:
9299:
9192:
9182:
8834:
8733:
8448:
7843:"It's Not Just Conflict That Motivates Killing of Orangutans"
6178:
5900:
5666:"Speech-Like Rhythm in a Voiced and Voiceless Orangutan Call"
4674:
4339:
Rose, M. D. (1988). "Functional Anatomy of the Cheirdia". In
4301:
4022:
3049:
Cannon, Garland (1992). "Malay(sian) borrowings in English".
2499:
2490:
All three species are critically endangered according to the
2107:
considered the ruling applicable only to non-human hominids.
1596:
1561:
1492:
1444:
1141:
496:
475:
430:
7902:"The military family that kept a pet orangutan in Indonesia"
7557:
7555:
6689:
3539:"Comparative and demographic analysis of orang-utan genomes"
659:
The word was first attested in English in 1693 by physician
12979:
12012:
9187:
8828:
8718:
7308:
6734:
6658:
5821:"Orangutans play video games (for research) at Georgia zoo"
5308:
4160:
2160:
1565:
1553:
1146:
691:
in the 20th century—explaining the missing initial 'h' of '
607:, a ninth- or early tenth-century Javanese adaption of the
531:
8331:
A Critical Companion to Zoosemiotics: People, Paths, Ideas
7930:
6838:
5939:
5459:
5063:
2934:
12362:
12349:
12299:
9253:
8809:
7552:
3812:
3682:
3650:
3276:
Manuel de mammalogie ou Histoire naturelle des mammifères
2937:
Wild Man from Borneo: A Cultural History of the Orangutan
2567:
2156:
1757:
1728:
1699:
1228:
8026:
Voigt, M; Wich, S. A.; Ancrenaz, M; et al. (2018).
7408:
Among Orangutans: Red Apes and the Rise of Human Culture
7013:"Argentine orangutan granted unprecedented legal rights"
6181:"Orangutans show active voicing through a membranophone"
4067:"Evidence for the latest fossil Pongo in southern China"
3535:
8778:
7561:
7519:
7517:
7515:
7513:
7098:
Encyclopedia of World Scientists: From Around the World
6237:
5722:
5439:"First Sighting of Bornean Orangutan Twins in the Wild"
5315:
Proceedings of the Royal Society B: Biological Sciences
3746:"New Species of Orangutan Is Rarest Great Ape on Earth"
2860:. Frankfurter Forschungen zu Südostasien. Vol. 3.
2764:
706:, an English sailor held prisoner by the Portuguese in
6010:
van Schaik, Carel P.; Damerius, L.; Isler, K. (2013).
5723:
Deaner, R. O.; van Schaik, C. P.; Johnson, V. (2006).
5436:
4723:, Lesson 1827): Ecology, Behaviour and Conservation".
3646:
3644:
3642:
3640:
3638:
3636:
3134:. In Taylor, Andrea B.; Goldsmith, Michele L. (eds.).
2407:
who compared her reactions to those of a human child.
648:
suffering some serious medical condition (most likely
466:, which diverged genetically from the other hominids (
7476:
7474:
7472:
7371:
Demonic Males: Apes and the Origins of Human Violence
7275:
Al G. Barnes, Master Showman, as told by Al G. Barnes
6937:"Spanish parliament approves 'human rights' for apes"
6332:
6307:
5557:
3878:: a giant ape from the Pleistocene of Asia revisited"
2935:
Cribb, Robert; Gilbert, Helen; Tiffin, Helen (2014).
2006:
Tool use in orangutans was observed by primatologist
1599:, bird eggs, insects and small vertebrates including
881:, the Sumatran orangutan became the third species of
586:
genus at an early stage in the development of Malay.
13017:
13006:
12995:
12985:
12967:
12957:
12946:
12898:
12880:
12812:
12782:
12765:
12755:
12745:
12735:
12725:
12715:
12692:
12682:
12672:
12662:
12625:
12601:
12577:
12495:
12480:
12470:
12460:
12450:
12440:
12430:
12419:
12409:
12399:
12389:
12379:
12265:
12248:
12238:
12228:
12218:
12208:
12170:
12160:
12130:
12035:
12018:
11980:
11970:
11953:
11943:
11933:
11923:
11913:
11896:
11835:
11820:
11660:
11631:
11601:
11564:
11549:
11539:
11429:
11350:
11340:
11330:
11320:
11303:
11280:
11270:
11260:
11250:
11233:
11216:
11206:
11196:
11152:
11142:
11132:
11108:
11098:
11088:
11078:
11068:
11044:
11034:
11010:
10987:
10957:
10947:
10937:
10927:
10917:
10907:
10776:
10710:
10673:
10642:
10626:
10615:
10605:
10595:
10584:
10486:
10469:
10459:
10419:
10408:
10398:
9898:
8248:"Extremely Rare Albino Orangutan Found in Indonesia"
7510:
4945:
Evolutionary Anthropology: Issues, News, and Reviews
4064:
3919:
3375:
2662:
2323:
to teach him how to make fire. The 1986 horror film
1790:
decrease intensity", "resume play" and "stop that".
652:) and that his use of the word was misunderstood by
7587:
10.2305/IUCN.UK.2017-3.RLTS.T120588639A120588662.en
6129:
5460:Beaudrot, LH; Kahlenberg, SM; Marshall, AJ (2009).
4802:
3633:
8276:. Sustainable Travel International. Archived from
7481:Singleton, I.; Wich, S. A.; Griffiths, M. (2008).
7469:
7444:Bring 'em back alive : the best of Frank Buck
7405:
7368:
7123:"Palm oil frenzy threatens to wipe out orangutans"
7095:
6385:
5409:
4983:
4905:
4847:
4576:
4167:(2nd ed.). Oxford University Press. pp.
3238:
2732:
1642:Orangutans are the least social of the great apes.
1556:or seed-walls are consumed the most, particularly
1281:, consisting of teeth ascribed to extinct species
8298:
8166:. Orangutan Protection Foundation. Archived from
8025:
7899:
7731:"Hundreds of Orangutans Killed Annually for Meat"
7193:
7010:
6617:
6615:
5775:
5615:
5613:
5508:
5018:
4977:
4975:
4937:
4798:
4796:
4758:
4756:
4509:
4507:
4505:
4503:
4416:
2884:
2270:derived from it. They are typically portrayed as
13309:
8571:
7935:spp.) on Borneo and Sumatra: how many remain?".
7366:
7147:
5207:
5104:Orangutan Species Survival Plan Husbandry Manual
4933:
4931:
4929:
4850:Female Primates: Studies by Women Primatologists
3678:
3676:
3136:Gorilla Biology: A Multidisciplinary Perspective
2422:in Hamburg, Germany, in the early 20th century;
2002:using a tool to extract orange-juice concentrate
1618:. Orangutans may eat soils for their anti-toxic
1345:Adult male (left) and female Tapanuli orangutans
1247:, which lived in India from 9.2 to 8.6 mya; and
641:Historiae naturalis et medicae Indiae orientalis
7189:
7187:
7120:
6934:
6621:
6312:. Cambridge University Press. pp. 99–116.
4216:
3960:
3378:Orangutans: Behaviour, Ecology and Conservation
3048:
1793:
836:). The description in 2017 of a third species,
8245:
7545:10.2305/IUCN.UK.2016-1.RLTS.T17975A17966347.en
6777:
6728:
6612:
6500:
5659:
5657:
5610:
5560:"Cyclical nursing patterns in wild orangutans"
5437:Goossens, B; Mohd, D; Kapar; Kahar, S (2011).
5214:) from the Sabangau Peat-Swamp Forest, Borneo"
4972:
4793:
4753:
4500:
4251:
2510:. The Tapanuli orangutan is found only in the
9825:
9237:
8794:
8489:
8454:AZA's Orangutan Conservation Education Center
8250:. National Geographic Society. Archived from
7733:. National Geographic Society. Archived from
7605:. National Geographic Society. Archived from
7250:. National Geographic Society. Archived from
6503:"An Orangutan Hangs Up a Tool for Future Use"
6231:
5818:
5101:
4926:
4896:
4548:
4297:
4295:
4293:
4194:An Introduction to Human Evolutionary Anatomy
4116:
4114:
4112:
4110:
4108:
3871:
3748:. National Geographic Society. Archived from
3673:
3287:
2855:
2588:(BOS) Foundation, founded by conservationist
2463:claimed to have seen human mothers acting as
1972:communicate about things that are not present
7600:
7367:Wrangham, Richard W; Peterson, Dale (1996).
7184:
7056:
5628:. Oxford University Press. pp. 270–75.
5304:
5302:
5300:
5097:
5095:
4882:. Oxford University Press. pp. 227–29.
4156:
4154:
4152:
4150:
2887:Studia Universitatis Petru Maior. Philologia
2771:Bijdragen tot de Land-, Taal- en Volkenkunde
1376:. Compared to chimpanzees and gorillas, the
916:, within apes (superfamily Hominoidea), the
7924:
7397:
7360:
7239:
6470:"Orangutans use simple tools to catch fish"
6070:
5654:
5416:. Oxford University Press. pp. 67–68.
4553:. Oxford University Press. pp. 31–51.
4338:
4121:Groves, Colin P. (1971). "Pongo pygmaeus".
3737:
3521:: CS1 maint: DOI inactive as of May 2024 (
2817:
2760:
2758:
2756:
2647:work with the palm oil industry to improve
1948:showed orangutans may practise "calculated
1295:, though it is known only from teeth. Some
597:, in a variety of premodern sources in the
518:All three orangutan species are considered
478:) between 19.3 and 15.7 million years ago.
10565:
9832:
9818:
9251:
9244:
9230:
8801:
8787:
8496:
8482:
8327:
8019:
7403:
6965:
5966:
4453:
4451:
4290:
4191:
4105:
3263:
3117:
1622:minerals, since their diet contains toxic
1495:. The most common orangutan parasites are
502:Orangutans are among the most intelligent
332:
114:
13343:Taxa named by Bernard Germain de Lacépède
12360:
12298:
11520:
11458:
8301:Ethical Debates in Orangutan Conservation
8102:
8092:
8043:
8002:
7955:
7876:
7866:
7817:
7807:
7728:
7585:
7543:
7333:
7214:
6872:
6760:
6625:American Journal of Physical Anthropology
6595:
6585:
6536:
6526:
6467:
6214:
6204:
6147:
6106:
6096:
6071:Lameira, Adriano R.; Call, Josep (2018).
6047:
6037:
5877:
5740:
5699:
5689:
5593:
5583:
5534:
5485:
5462:"Why male orangutans do not kill infants"
5336:
5326:
5297:
5239:
5229:
5184:
5092:
4651:
4641:
4483:
4423:The Red Ape: Orangutans and Human Origins
4393:
4323:
4147:
3893:
3848:
3838:
3713:
3596:
3594:
3592:
3572:
3529:
3439:
3429:
3407:
3405:
3403:
3401:
3371:
3369:
3367:
3365:
3363:
3293:
2782:
2651:and encourages the industry to establish
2643:. Conservation organisations such as the
2639:, which was founded by Galdikas, and the
2095:, as it is the only accredited orangutan
762:
8419:
7266:
7114:
6834:
6832:
6310:The Mentality of Gorillas and Orangutans
6280:
6179:Lameira, A. R.; Shumaker, R. W. (2019).
5256:
4912:. Oxford University Press. p. 199.
4513:
4410:
4347:. Oxford University Press. p. 301.
3361:
3359:
3357:
3355:
3353:
3351:
3349:
3347:
3345:
3343:
3084:
3082:
2930:
2928:
2926:
2924:
2922:
2920:
2753:
2571:
2516:
2365:
2208:
2114:
2110:
1993:
1920:
1883:
1854:
1637:
1539:
1430:
1426:
1394:
1340:
1201:
856:
616:. In these Old Javanese texts, the word
8422:"New deal for orangutans in Kalimantan"
7272:
7062:
5158:
4718:
4448:
3872:Zhang, Yingqi; Harrison, Terry (2017).
3743:
2918:
2916:
2914:
2912:
2910:
2908:
2906:
2904:
2902:
2900:
2851:
2849:
2847:
2845:
1989:
1606:In some areas, orangutans may practise
702:, comes from a 16th-century account by
601:language. The earliest of these is the
13310:
8392:
8141:from the original on 28 September 2020
7245:
7037:
7011:Giménez, Emiliano (23 December 2014).
5960:
5933:
4185:
4120:
3770:
3589:
3398:
3279:(in French). Roret, Libraire. p.
3269:
3123:
2963:
2878:
2568:Conservation centres and organisations
2187:, Galdikas is considered to be one of
1399:Skeleton of subadult Bornean orangutan
409:. They are now found only in parts of
13073:
13072:
12347:
11507:
10552:
9883:
9813:
9225:
9137:Chimpanzee–human last common ancestor
8782:
8477:
7348:from the original on 25 December 2018
7197:; Kjærgaard, Peter C. (1 June 2015).
6978:from the original on 13 November 2012
6895:
6829:
5122:
4457:
4018:
4016:
3915:
3913:
3744:Goldman, Jason G. (2 November 2017).
3693:Molecular Phylogenetics and Evolution
3600:
3456:
3340:
3088:
3079:
2479:
850:than to its fellow Sumatran species,
13293:8BE3221A-62FE-4604-B27D-B9151BCE14DB
9147:Orangutan–human last common ancestor
7440:
5907:International Journal of Primatology
5515:International Journal of Primatology
5400:
5218:International Journal of Primatology
5159:Lameira, A. R.; et al. (2021).
4870:
4516:International Journal of Primatology
4332:
3825:(11): 10.1371/journal.pone.0206314.
3764:
3651:Rijksen H. D.; Meijaard, E. (1999).
3621:from the original on 27 January 2011
3503:from the original on 6 December 2010
3473:International Journal of Primatology
3219:from the original on 5 December 2018
3189:from the original on 5 December 2018
2897:
2842:
2830:from the original on 4 November 2012
2727:
2725:
1443:Wild orangutan in the Danum Valley (
960:Phylogeny of superfamily Hominoidea
905:chromosomes, in contrast to humans,
769:Orangutan–human last common ancestor
508:use a variety of sophisticated tools
13328:Species endangered by the pet trade
8401:from the original on 5 January 2012
8227:from the original on 3 October 2020
8185:"Ny projektledare på Nyaru Menteng"
8161:"10 Years: Nyaru Menteng 1999–2009"
7900:Jacobson, Phillip (29 March 2017).
7763:from the original on 3 January 2012
7573:IUCN Red List of Threatened Species
7531:IUCN Red List of Threatened Species
7492:IUCN Red List of Threatened Species
7334:Middleton, Duncan (7 August 2016).
7148:MacClancy, J.; Fuentes, A. (2010).
7121:McDowell, Robin (18 January 2009).
6947:from the original on 26 August 2009
5757:from the original on 19 August 2021
5466:Behavioral Ecology and Sociobiology
5359:
5270:Behavioral Ecology and Sociobiology
5262:
5140:from the original on 3 October 2020
4583:. Oxford University Press. p.
3251:from the original on 19 August 2021
3011:
2617:Bukit Baka Bukit Raya National Park
1932:An orangutan imitating human speech
1535:
812:was described by French naturalist
13:
9142:Gorilla–human last common ancestor
9074:Kinshasa Declaration on Great Apes
8449:Orangutan Foundation International
8348:from the original on 30 April 2016
7172:from the original on 30 April 2016
7125:. Associated Press. Archived from
6741:Proceedings of the Royal Society B
5642:from the original on 24 April 2016
4013:
3910:
3603:"Orang-utans join the genome gang"
3601:Singh, Ranjeet (26 January 2011).
3486:10.1023/B:IJOP.0000014647.18720.32
2799:from the original on 12 April 2021
2637:Orangutan Foundation International
2123:presenting her book about the apes
1772:Problems playing these files? See
1672:
1336:
1210:, an extinct relative of orangutan
14:
13359:
9152:Gibbon–human last common ancestor
8442:
8191:. 26 January 2012. Archived from
7912:from the original on 14 June 2020
7759:. Antara News. 7 September 2011.
7019:from the original on 3 April 2021
5786:Journal of Comparative Psychology
4610:"Self-medication by orang-utans (
3882:Yearbook of Physical Anthropology
2722:
2555:Orangutans may be killed for the
1938:track the displacement of objects
446:, with three subspecies) and the
13052:
12330:
11490:
10534:
9206:
9205:
8957:
8763:
8762:
8413:
8393:Butler, R. A. (20 August 2009).
8386:
8360:
8321:
8292:
8274:"Viewing Orangutans in the Wild"
8266:
8239:
8207:
8177:
8153:
8127:
8068:
7893:
7834:
7775:
7749:
7722:
7710:from the original on 12 May 2019
7697:
7691:
7647:
7621:
7601:Neme, Laurel (11 October 2014).
7594:
7434:
7327:
7297:
7141:
7089:
7031:
7004:
6990:
6959:
6935:Glendinning, L. (26 June 2008).
6928:
6482:from the original on 8 July 2013
5831:from the original on 2 July 2007
4690:10.1111/j.1600-0684.2011.00490.x
4614:) using bioactive properties of
4458:Reese, April (2 November 2017).
4386:10.1111/j.1469-7580.2009.01125.x
2993:from the original on 7 July 2016
2765:Sastrawan, Wayan Jarrah (2020).
2665:
2523:palm oil production in Indonesia
2399:housed a female orangutan named
1814:
1802:
1746:
1717:
1688:
1665:
1392:chambers for making long calls.
1241:in southern China and Thailand;
1214:
1048: (family Hylobatidae)
140:
39:
10553:
9084:Great Apes Survival Partnership
9034:Iowa Primate Learning Sanctuary
7635:from the original on 8 May 2020
7077:from the original on 4 May 2013
7063:de Waal, Frans (January 1995).
6916:from the original on 2 May 2024
6889:
6683:
6652:
6553:
6494:
6461:
6426:
6379:
6326:
6301:
6274:
6172:
6123:
6064:
6003:
5969:American Journal of Primatology
5894:
5843:
5819:Turner, Dorie (12 April 2007).
5812:
5769:
5716:
5551:
5502:
5453:
5430:
5366:American Journal of Primatology
5353:
5201:
5152:
5057:
5012:
4841:
4726:The Quarterly Review of Biology
4712:
4668:
4601:
4567:
4542:
4361:
4255:American Journal of Primatology
4245:
4210:
4196:. Academic Press. p. 193.
4058:
3990:
3954:
3865:
3806:
3376:Payne, J; Prundente, C (2008).
3231:
3201:
3171:
3129:"A history of gorilla taxonomy"
3042:
2474:
2361:
1904:
1349:Orangutans display significant
12348:
11508:
9884:
9839:
8246:Brady, Heather (18 May 2017).
7729:Than, Ker (16 November 2011).
7631:. Sumatran Orangutan Society.
7246:Zimmer, Carl (21 April 2015).
7100:. Facts on File. p. 260.
7038:Meijer, Miriam Claude (2014).
5136:– Department of Anthropology.
4678:Journal of Medical Primatology
3465:"Asian primate classification"
3297:Journal of Molecular Evolution
3005:
2975:
2957:
2811:
2315:added a jazzy orangutan named
2079:, which are that chimpanzees,
1888:An orangutan lying in its nest
1662:is uncommon among orangutans.
1633:
530:and retaliation for consuming
1:
9413:Kasakela chimpanzee community
8420:Marusiak, J. (28 June 2011).
8334:. Springer. pp. 218–19.
7248:"When Darwin Met Another Ape"
6295:10.1016/S0047-2484(82)80028-6
5954:10.1016/s0047-2484(81)80016-4
5622:"Nest building in orangutans"
5078:10.1016/S0003-3472(05)80603-7
4779:10.1016/S0003-3472(05)80782-1
4192:Aiello, L.; Dean, C. (1990).
3103:10.1080/09502386.2020.1780281
2716:
2378:
2350:What the Orangutan Told Alice
2298:The Murders in the Rue Morgue
2215:The Murders in the Rue Morgue
2205:Orangutans in popular culture
2198:
2191:, named after anthropologist
2056:
1024:– Sumatra south of Lake Toba)
824:were elevated to subspecies (
9453:Mitumba chimpanzee community
7868:10.1371/journal.pone.0075373
7809:10.1371/journal.pone.0027491
7757:"Stop orangutan skull trade"
6896:Davis, Nicola (2 May 2024).
6447:10.1016/j.jhevol.2014.06.007
6039:10.1371/journal.pone.0074896
5691:10.1371/journal.pone.0116136
5130:"Orangutan call repertoires"
4083:10.1016/j.jhevol.2022.103233
4044:10.1016/j.jhevol.2013.09.005
3984:10.1016/j.quaint.2014.06.020
3948:10.1016/j.quaint.2014.01.013
3840:10.1371/journal.pone.0206314
3148:10.1017/CBO9780511542558.004
2641:Australian Orangutan Project
2633:Frankfurt Zoological Society
2629:Sepilok Orang Utan Sanctuary
2605:Tanjung Puting National Park
2430:in the 1910s and 1920s; and
1957:cooperative pulling paradigm
1794:Reproduction and development
565:
7:
7629:"On the edge of extinction"
7216:10.1016/j.shpsc.2015.02.006
7096:Oaks, Elizabeth H. (2007).
6966:Singer, P. (18 July 2008).
6468:Bower, B. (18 April 2011).
5798:10.1037/0735-7036.112.2.137
4942:): a tale of two islands".
4164:The Encyclopedia of Mammals
3706:10.1016/j.ympev.2010.11.005
3247:. Oxford University Press.
2941:University of Hawai'i Press
2824:Online Etymology Dictionary
2658:
2621:Gunung Leuser National Park
2613:Gunung Palung National Park
2307:animated musical adaptation
1859:Mother orangutan with young
1269:The oldest known record of
720:Bernard Germain de Lacépède
667:, and variants ending with
593:appears in its older form,
10:
13364:
9281:List of fictional primates
9271:List of individual monkeys
9157:List of fictional primates
9009:Neanderthal genome project
8459:Orangutan Language Project
8299:Palmer, Alexandra (2020).
7580:: e.T120588639A120588662.
7404:van Schaik, Carel (2004).
6857:10.1038/s41598-024-58988-7
6677:10.1163/156853906778017944
6528:10.1038/s41598-018-31331-7
6435:Journal of Human Evolution
6283:Journal of Human Evolution
6206:10.1038/s41598-019-48760-7
5942:Journal of Human Evolution
5742:10.1177/147470490600400114
5527:10.1007/s10764-019-00097-8
5231:10.1007/s10764-019-00095-w
4992:Cambridge University Press
4643:10.1038/s41598-017-16621-w
4071:Journal of Human Evolution
4031:Journal of Human Evolution
3140:Cambridge University Press
2486:Endangerment of orangutans
2483:
2469:human–animal breastfeeding
2202:
2060:
1914:
1908:
1879:
1447:, Malaysia, Borneo island)
1237:, which occurred 8–2
920:diverged during the early
766:
37:Early Pleistocene – Recent
21:Orangutan (disambiguation)
18:
13081:
13049:
12937:
12922:
12913:
12871:
12811:
12800:
12706:
12658:
12654:
12643:
12568:
12494:
12375:
12371:
12356:
12343:
12328:
12294:
12144:
12079:
12070:
11887:
11834:
11816:
11805:
11728:
11701:
11674:
11642:
11630:
11600:
11563:
11535:
11531:
11516:
11503:
11488:
11454:
11378:
11294:
11166:
11001:
10971:
10903:
10892:
10836:
10809:
10787:
10775:
10709:
10672:
10641:
10580:
10576:
10561:
10548:
10531:
10505:
10436:
10394:
10385:
10381:
10343:
10153:
10026:
9950:
9912:
9892:
9879:
9847:
9714:
9526:
9318:
9290:
9276:Monkeys and apes in space
9260:
9201:
9102:
9089:International Primate Day
9052:
9039:Borneo Orangutan Survival
8994:Chimpanzee genome project
8966:
8955:
8816:
8758:
8716:
8680:
8671:
8632:
8623:
8562:
8553:
8515:
8504:Extant species of family
8094:10.1016/j.cub.2017.09.047
8045:10.1016/j.cub.2018.01.053
7669:10.1007/s10329-007-0063-0
6706:10.1007/s10071-006-0058-z
6252:10.1007/s10329-008-0117-y
6149:10.1016/j.cub.2009.05.028
5478:10.1007/s00265-009-0827-1
5283:10.1007/s00265-002-0495-x
4485:10.1038/nature.2017.22934
4304:van Hooff, J. A. R. A. M.
3431:10.1016/j.cub.2017.09.047
3028:10.1017/S026607840001052X
2784:10.1163/22134379-bja10016
2739:Oxford English Dictionary
2681:International Primate Day
2586:Borneo Orangutan Survival
2089:The Center for Great Apes
1940:both visible and hidden.
1588:Borassodendron borneensis
1574:which contains the toxic
1548:Orangutans are primarily
1299:described under the name
1139:
1119:
1112:
1092:
1085:
1066:
1059:
1043:
1036:
959:
953:
355:
348:
340:
331:
293:
288:
265:
258:
137:Scientific classification
135:
122:
113:
30:
8808:
8464:The Orangutan Foundation
7414:Harvard University Press
5780:) and Squirrel Monkeys (
3964:Quaternary International
3927:Quaternary International
3778:; Reeder, D. M. (eds.).
2862:Otto Harrassowitz Verlag
2438:in the 1930s and 1940s.
2278:, the science minister.
2250:Frank Challice Constable
1821:Unflanged male orangutan
842:, from Sumatra south of
775:described scientifically
773:The orangutan was first
578:, meaning "person", and
454:). A third species, the
13338:Fauna of Southeast Asia
9266:List of individual apes
9115:List of individual apes
9094:Nonhuman Rights Project
8328:Martinelli, D. (2010).
8137:. Masarang Foundation.
7065:"The Loneliest of Apes"
6968:"Of great apes and men"
6807:10.1126/science.1078004
6587:10.1073/pnas.1200902109
6501:Mulcahy, N. J. (2018).
5919:10.1023/A:1026337006136
5729:Evolutionary Psychology
5108:Chicago Zoological Park
4325:10.1093/beheco/13.5.643
3488:(inactive 3 May 2024).
2686:List of individual apes
2453:The Ballad of Ken Allen
2371:The Female Orang – Utan
2213:1870 illustration for "
1459:, particularly lowland
1208:Sivapithecus sivalensis
1069:orangutans (genus
1010:– Sumatra northwest of
998:– southwest populations
986:– northwest populations
984:Pongo pygmaeus pygmaeus
893:than in Sumatran ones (
800:Christian Emmanuel Hopp
798:in 1760 by his student
698:The name of the genus,
421:they ranged throughout
13348:Taxa described in 1799
9004:Gladys Kalema-Zikusoka
7995:10.1126/sciadv.1500789
7273:Robeson, Dave (1935).
7042:. Rodopi. p. 42.
6753:10.1098/rspb.2009.1027
6661:Pongo pygmaeus wurmbii
6098:10.1126/sciadv.aau3401
5981:10.1002/ajp.1350100202
5870:10.1098/rsbl.2008.0644
5585:10.1126/sciadv.1601517
5511:Pongo pygmaeus wurmbii
5443:Asian Primates Journal
5328:10.1098/rspb.2009.1552
5212:Pongo pygmaeus wurmbii
5177:10.1098/rsbl.2021.0302
4268:10.1002/ajp.1350170405
3876:Gigantopithecus blacki
3382:New Holland Publishers
3271:Lesson, René-Primevère
2856:Mahdi, Waruno (2007).
2609:Sebangau National Park
2581:
2525:
2384:
2254:The Curse of Intellect
2221:
2124:
2071:Great ape research ban
2003:
1982:, an orangutan at the
1933:
1889:
1860:
1809:Flanged male orangutan
1677:
1643:
1545:
1451:Orangutans are mainly
1448:
1400:
1346:
1227:also includes extinct
1211:
996:Pongo pygmaeus wurmbii
946:recent glacial periods
874:
763:Taxonomy and phylogeny
13323:Primates of Indonesia
13275:Paleobiology Database
9019:Lone Drøscher Nielsen
8087:(22): 3487–3498.e10.
7538:: e.T17975A17966347.
5362:Pongo pygmaeus abelii
5265:Pongo pygmaeus abelii
4721:Pongo pygmaeus abelii
3424:(22): 3487–3498.e10.
2964:Bulwer, John (1653).
2594:Lone Drøscher Nielsen
2575:
2520:
2369:
2288:Count Robert of Paris
2226:Tintinnabulum naturae
2212:
2119:Orangutan researcher
2118:
2111:Orangutans and humans
1997:
1931:
1909:Further information:
1887:
1858:
1676:
1641:
1543:
1479:, and shallow lakes.
1442:
1427:Ecology and behaviour
1398:
1344:
1205:
860:
675:. The loss of "h" in
520:critically endangered
8999:Human Genome Project
8607:Tapanuli orangutan (
8589:Sumatran orangutan (
8469:Orangutan Land Trust
8305:Taylor & Francis
8223:. 28 November 2018.
8173:on 15 February 2010.
7441:Buck, Frank (2000).
5564:Evolutionary Biology
3615:10.1038/news.2011.50
2645:Orangutan Land Trust
2514:forest of Sumatra.
2377:sitting in a chair)
2246:Member of Parliament
2063:Great ape personhood
1998:An orangutan at the
1990:Tool use and culture
1850:Homosexual behaviour
1830:arrested development
1331:Last Glacial Maximum
990:Pongo pygmaeus morio
718:. French naturalist
628:ape was most likely
429:. Classified in the
12213:Paradolichopithecus
9561:Charles the Gorilla
9044:Primate archaeology
8598:Bornean orangutan (
8374:on 10 February 2007
8254:on 17 February 2020
7987:2016SciA....2E0789W
7859:2013PLoSO...875373D
7800:2011PLoSO...627491M
7704:factsanddetails.com
7566:Pongo tapanuliensis
6799:2003Sci...299..102V
6578:2012PNAS..109.6873V
6519:2018NatSR...812900M
6392:Folia Primatologica
6349:1996NW.....83..186V
6336:Naturwissenschaften
6197:2019NatSR...912289L
6089:2018SciA....4.3401L
6030:2013PLoSO...874896V
5682:2015PLoSO..10k6136L
5576:2017SciA....3E1517S
4856:. pp. 217–35.
4806:Folia Primatologica
4634:2017NatSR...716653M
4476:2017Natur.551..151R
3976:2014QuInt.354...68W
3940:2014QuInt.354...59H
3831:2018PLoSO..1306314B
3565:10.1038/nature09687
3557:2011Natur.469..529L
3310:1996JMolE..43..431X
2864:. pp. 170–81.
2319:, who tries to get
2236:Thomas Love Peacock
2042:Fibraurea tinctoria
2016:Carel P. van Schaik
1648:solitary but social
1457:tropical rainforest
1423:other two species.
1323:Peninsular Malaysia
1022:Pongo tapanuliensis
802:and given the name
550:), and the illegal
536:habitat destruction
341:Range of the three
309:Pongo tapanuliensis
16:Genus of Asian apes
13333:Tool-using mammals
11839:Victoriapithecidae
11781:Plesiopliopithecus
11719:Platodontopithecus
9536:Alfred the Gorilla
9252:Notable non-human
8974:Great ape language
8932:Tapanuli orangutan
8912:Sumatran orangutan
8195:on 10 January 2014
8189:Save the Orangutan
8038:(5): P761–769.E5.
7129:on 20 January 2009
7070:The New York Times
6845:Scientific Reports
6507:Scientific Reports
6357:10.1007/BF01143062
6185:Scientific Reports
5134:Universität Zürich
5110:. pp. 59–66.
5035:10.1007/BF02557703
4622:Scientific Reports
4551:Orang-utan biology
4528:10.1007/BF02740195
4374:Journal of Anatomy
4345:Orang-utan Biology
4312:Behavioral Ecology
4231:10.1007/BF02447216
3895:10.1002/ajpa.23150
3318:10.1007/BF02337514
3142:. pp. 15–34.
2653:conservation areas
2619:in Borneo and the
2582:
2526:
2521:Deforestation for
2480:Status and threats
2428:Universal City Zoo
2420:Tierpark Hagenbeck
2385:
2338:'s fantasy novels
2259:Planet of the Apes
2222:
2125:
2004:
1934:
1917:Great ape language
1890:
1861:
1678:
1644:
1546:
1449:
1401:
1347:
1212:
1144: (genus
1124: (genus
1018:Tapanuli orangutan
1004:Sumatran orangutan
992:– east populations
954:Taxonomy of genus
914:molecular evidence
875:
816:in 1827. In 2001,
456:Tapanuli orangutan
448:Sumatran orangutan
323:Pongo weidenreichi
13303:
13302:
13262:Open Tree of Life
13075:Taxon identifiers
13066:
13065:
13062:
13061:
13047:
13046:
13043:
13042:
13039:
13038:
13035:
13034:
13031:
13030:
12796:
12795:
12639:
12638:
12339:
12338:
12326:
12325:
12322:
12321:
12318:
12317:
12290:
12289:
12286:
12285:
11801:
11800:
11797:
11796:
11703:Dionysopithecidae
11605:Propliopithecidae
11499:
11498:
11486:
11485:
11482:
11481:
11478:
11477:
11450:
11449:
10888:
10887:
10884:
10883:
10544:
10543:
10529:
10528:
10525:
10524:
10501:
10500:
10377:
10376:
9807:
9806:
9219:
9218:
9079:Great Ape Project
8922:Bornean orangutan
8776:
8775:
8754:
8753:
8750:
8749:
8746:
8745:
8727:
8688:
8659:Western gorilla (
8650:Eastern gorilla (
8643:
8619:
8618:
8582:
8509:
8341:978-90-481-9248-9
8280:on 3 October 2020
7737:on 3 October 2020
7390:978-0-395-69001-7
7165:978-1-84545-690-0
6747:(1673): 3689–94.
6638:10.1002/ajpa.1045
6404:10.1159/000021756
6319:978-0-521-03193-6
5635:978-0-19-921327-6
4889:978-0-19-921327-6
4819:10.1159/000060415
4616:Dracaena cantleyi
4441:978-0-8133-4064-7
4418:Schwartz, Jeffrey
4354:978-0-19-504371-6
4341:Schwartz, Jeffrey
4178:978-0-87196-871-5
4124:Mammalian Species
3391:978-0-262-16253-1
3167:on 26 March 2009.
2950:978-0-8248-3714-3
2871:978-3-447-05492-8
2818:Harper, Douglas.
2355:Dunston Checks In
2105:Criminal Cassatio
2077:Great Ape Project
2067:Great Ape Project
2037:field observation
1929:
1911:Primate cognition
1751:
1722:
1693:
1571:Strychnos ignatii
1529:Dracaena cantleyi
1473:peat swamp forest
1440:
1351:sexual dimorphism
1275:Early Pleistocene
1200:
1199:
1195:
1194:
1186:
1185:
1177:
1176:
1168:
1167:
1159:
1158:
1034: Hominoidea
974:Bornean orangutan
891:genetic diversity
791:. It was renamed
738:from the region:
440:Bornean orangutan
417:, but during the
388:
387:
383:
376:
369:
362:
254:
124:Bornean orangutan
13355:
13296:
13295:
13283:
13282:
13270:
13269:
13257:
13256:
13244:
13243:
13231:
13230:
13218:
13217:
13205:
13204:
13192:
13191:
13179:
13178:
13166:
13165:
13153:
13152:
13140:
13139:
13130:
13129:
13117:
13116:
13115:
13102:
13101:
13100:
13070:
13069:
13057:
13056:
13019:
13008:
12997:
12987:
12969:
12962:Australopithecus
12959:
12948:
12920:
12919:
12900:
12882:
12814:
12809:
12808:
12784:
12767:
12757:
12747:
12737:
12727:
12717:
12694:
12684:
12674:
12664:
12656:
12655:
12652:
12651:
12627:
12603:
12579:
12499:Dendropithecidae
12497:
12482:
12472:
12462:
12452:
12442:
12432:
12421:
12411:
12401:
12391:
12381:
12373:
12372:
12369:
12368:
12358:
12357:
12345:
12344:
12334:
12296:
12295:
12267:
12250:
12240:
12230:
12220:
12210:
12172:
12162:
12132:
12077:
12076:
12037:
12020:
11982:
11972:
11955:
11945:
11935:
11925:
11915:
11901:Cercopithecoides
11898:
11871:Victoriapithecus
11837:
11822:
11814:
11813:
11808:Cercopithecoidea
11662:
11640:
11639:
11633:
11603:
11566:
11551:
11541:
11533:
11532:
11529:
11528:
11518:
11517:
11505:
11504:
11494:
11456:
11455:
11431:
11352:
11342:
11332:
11322:
11305:
11282:
11272:
11262:
11252:
11235:
11218:
11208:
11198:
11154:
11144:
11134:
11110:
11100:
11090:
11080:
11070:
11046:
11036:
11012:
10989:
10959:
10949:
10939:
10929:
10919:
10909:
10901:
10900:
10811:Proteopithecidae
10785:
10784:
10778:
10712:
10675:
10644:
10628:
10617:
10607:
10597:
10586:
10578:
10577:
10574:
10573:
10563:
10562:
10550:
10549:
10539:
10538:
10488:
10471:
10461:
10421:
10410:
10400:
10392:
10391:
10383:
10382:
9910:
9909:
9900:
9894:
9893:
9881:
9880:
9834:
9827:
9820:
9811:
9810:
9784:Nonja (Malaysia)
9246:
9239:
9232:
9223:
9222:
9209:
9208:
9178:Mythic humanoids
8961:
8936:P. tapanuliensis
8803:
8796:
8789:
8780:
8779:
8766:
8765:
8726:
8723:
8687:
8678:
8677:
8642:
8639:
8630:
8629:
8609:P. tapanuliensis
8581:
8578:
8569:
8568:
8560:
8559:
8507:
8498:
8491:
8484:
8475:
8474:
8437:
8436:
8434:
8432:
8417:
8411:
8410:
8408:
8406:
8390:
8384:
8383:
8381:
8379:
8364:
8358:
8357:
8355:
8353:
8325:
8319:
8318:
8296:
8290:
8289:
8287:
8285:
8270:
8264:
8263:
8261:
8259:
8243:
8237:
8236:
8234:
8232:
8211:
8205:
8204:
8202:
8200:
8181:
8175:
8174:
8172:
8165:
8157:
8151:
8150:
8148:
8146:
8131:
8125:
8124:
8106:
8096:
8072:
8066:
8065:
8047:
8023:
8017:
8016:
8006:
7974:Science Advances
7959:
7953:
7952:
7949:10.5167/uzh-3914
7928:
7922:
7921:
7919:
7917:
7897:
7891:
7890:
7880:
7870:
7838:
7832:
7831:
7821:
7811:
7779:
7773:
7772:
7770:
7768:
7753:
7747:
7746:
7744:
7742:
7726:
7720:
7719:
7717:
7715:
7695:
7689:
7688:
7651:
7645:
7644:
7642:
7640:
7625:
7619:
7618:
7616:
7614:
7598:
7592:
7591:
7589:
7559:
7550:
7549:
7547:
7526:"Pongo pygmaeus"
7521:
7508:
7507:
7505:
7503:
7478:
7467:
7466:
7438:
7432:
7431:
7411:
7401:
7395:
7394:
7377:Houghton Mifflin
7374:
7364:
7358:
7357:
7355:
7353:
7331:
7325:
7324:
7322:
7320:
7311:. Archived from
7301:
7295:
7294:
7270:
7264:
7263:
7261:
7259:
7243:
7237:
7236:
7218:
7191:
7182:
7181:
7179:
7177:
7158:. pp. 6–7.
7145:
7139:
7138:
7136:
7134:
7118:
7112:
7111:
7093:
7087:
7086:
7084:
7082:
7060:
7054:
7053:
7035:
7029:
7028:
7026:
7024:
7008:
7002:
7001:
6994:
6988:
6987:
6985:
6983:
6963:
6957:
6956:
6954:
6952:
6932:
6926:
6925:
6923:
6921:
6893:
6887:
6886:
6876:
6836:
6827:
6826:
6793:(5603): 102–05.
6781:
6775:
6774:
6764:
6732:
6726:
6725:
6693:Animal Cognition
6687:
6681:
6680:
6656:
6650:
6649:
6619:
6610:
6609:
6599:
6589:
6557:
6551:
6550:
6540:
6530:
6498:
6492:
6491:
6489:
6487:
6465:
6459:
6458:
6430:
6424:
6423:
6383:
6377:
6376:
6330:
6324:
6323:
6305:
6299:
6298:
6278:
6272:
6271:
6235:
6229:
6228:
6218:
6208:
6176:
6170:
6169:
6151:
6127:
6121:
6120:
6110:
6100:
6083:(11): eaau3401.
6077:Science Advances
6068:
6062:
6061:
6051:
6041:
6007:
6001:
6000:
5964:
5958:
5957:
5937:
5931:
5930:
5898:
5892:
5891:
5881:
5847:
5841:
5840:
5838:
5836:
5816:
5810:
5809:
5782:Saimiri Sciureus
5773:
5767:
5766:
5764:
5762:
5744:
5720:
5714:
5713:
5703:
5693:
5661:
5652:
5651:
5649:
5647:
5617:
5608:
5607:
5597:
5587:
5555:
5549:
5548:
5538:
5506:
5500:
5499:
5489:
5457:
5451:
5450:
5434:
5428:
5427:
5415:
5404:
5398:
5397:
5378:10.1002/ajp.1051
5357:
5351:
5350:
5340:
5330:
5321:(1678): 105–13.
5306:
5295:
5294:
5260:
5254:
5253:
5243:
5233:
5205:
5199:
5198:
5188:
5156:
5150:
5149:
5147:
5145:
5126:
5120:
5119:
5099:
5090:
5089:
5066:Animal Behaviour
5061:
5055:
5054:
5016:
5010:
5009:
4989:
4979:
4970:
4969:
4935:
4924:
4923:
4911:
4900:
4894:
4893:
4874:
4868:
4867:
4845:
4839:
4838:
4800:
4791:
4790:
4773:(6): 1098–1109.
4766:Animal Behaviour
4760:
4751:
4750:
4716:
4710:
4709:
4672:
4666:
4665:
4655:
4645:
4628:(16653): 16653.
4605:
4599:
4598:
4582:
4571:
4565:
4564:
4546:
4540:
4539:
4511:
4498:
4497:
4487:
4455:
4446:
4445:
4414:
4408:
4407:
4397:
4365:
4359:
4358:
4336:
4330:
4329:
4327:
4299:
4288:
4287:
4249:
4243:
4242:
4214:
4208:
4207:
4189:
4183:
4182:
4158:
4145:
4144:
4118:
4103:
4102:
4062:
4056:
4055:
4020:
4011:
4010:
3994:
3988:
3987:
3958:
3952:
3951:
3917:
3908:
3907:
3897:
3869:
3863:
3862:
3852:
3842:
3810:
3804:
3803:
3768:
3762:
3761:
3759:
3757:
3741:
3735:
3734:
3732:
3726:. Archived from
3717:
3689:
3680:
3671:
3670:
3648:
3631:
3630:
3628:
3626:
3598:
3587:
3586:
3576:
3551:(7331): 529–33.
3533:
3527:
3526:
3520:
3512:
3510:
3508:
3502:
3469:
3460:
3454:
3453:
3443:
3433:
3409:
3396:
3395:
3373:
3338:
3337:
3291:
3285:
3284:
3267:
3261:
3260:
3258:
3256:
3242:
3235:
3229:
3228:
3226:
3224:
3205:
3199:
3198:
3196:
3194:
3183:Etymology Online
3175:
3169:
3168:
3166:
3160:. Archived from
3133:
3125:Groves, Colin P.
3121:
3115:
3114:
3091:Cultural Studies
3086:
3077:
3076:
3046:
3040:
3039:
3009:
3003:
3002:
3000:
2998:
2987:Alpha Dictionary
2979:
2973:
2972:
2961:
2955:
2954:
2932:
2895:
2894:
2882:
2876:
2875:
2853:
2840:
2839:
2837:
2835:
2815:
2809:
2808:
2806:
2804:
2786:
2762:
2751:
2750:
2748:
2746:
2736:
2729:
2702:Orangutan Island
2675:
2670:
2669:
2655:for orangutans.
2508:Leuser Ecosystem
2383:
2380:
2230:The Orang Outang
2175:, and unnatural
2085:Buenos Aires Zoo
2045:vine leaves and
1930:
1865:sexual swellings
1842:force copulation
1818:
1806:
1784:blow raspberries
1753:
1752:
1724:
1723:
1695:
1694:
1675:
1536:Diet and feeding
1511:Balantidium coli
1489:clouded leopards
1477:secondary forest
1465:secondary forest
1441:
1327:P. weidenreichi,
1307:P. pygmaeus
1302:P. hooijeri
1206:Fossil skull of
1115:
1114:
1088:
1087:
1062:
1061:
1039:
1038:
1031:
1030:
951:
950:
899:genetic diseases
887:P. pygmaeus
839:P. tapanuliensis
789:Homo troglodytes
622:Malayic language
604:Kakawin Ramayana
546:cultivation and
460:P. tapanuliensis
381:
374:
367:
360:
336:
249:
145:
144:
118:
108:
38:
34:Temporal range:
28:
27:
13363:
13362:
13358:
13357:
13356:
13354:
13353:
13352:
13308:
13307:
13304:
13299:
13291:
13286:
13278:
13273:
13265:
13260:
13252:
13247:
13239:
13234:
13226:
13221:
13213:
13208:
13200:
13195:
13187:
13182:
13174:
13169:
13161:
13156:
13148:
13143:
13135:
13133:
13125:
13120:
13111:
13110:
13105:
13096:
13095:
13090:
13077:
13067:
13058:
13051:
13027:
12933:
12909:
12885:Chororapithecus
12867:
12854:Pierolapithecus
12846:Hispanopithecus
12805:
12792:
12730:Gigantopithecus
12702:
12648:
12635:
12630:Yuanmoupithecus
12564:
12559:Turkanapithecus
12490:
12485:Samburupithecus
12455:Nacholapithecus
12365:
12352:
12335:
12314:
12282:
12253:Procynocephalus
12140:
12072:Cercopithecinae
12066:
11918:Dolichopithecus
11883:
11855:Nsungwepithecus
11830:
11810:
11793:
11746:Barberapithecus
11724:
11712:Dionysopithecus
11697:
11685:Epipliopithecus
11670:
11648:Krishnapithecus
11635:Pliopithecoidea
11626:
11621:Propliopithecus
11614:Aegyptopithecus
11596:
11568:Oligopithecidae
11559:
11525:
11512:
11495:
11474:
11446:
11374:
11290:
11162:
11137:Proteropithecia
10997:
10967:
10897:
10880:
10875:Ucayalipithecus
10832:
10805:
10780:Parapithecoidea
10771:
10752:Myanmarpithecus
10714:Amphipithecidae
10705:
10700:Phenacopithecus
10668:
10637:
10570:
10557:
10540:
10533:
10521:
10497:
10432:
10373:
10339:
10149:
10029:Anaptomorphinae
10022:
9946:
9906:
9888:
9875:
9843:
9838:
9808:
9803:
9779:Nonja (Austria)
9710:
9581:Guy the Gorilla
9522:
9314:
9286:
9285:
9256:
9250:
9220:
9215:
9197:
9168:Human evolution
9098:
9055:
9048:
8984:Birutė Galdikas
8962:
8953:
8893:Western gorilla
8883:Eastern gorilla
8819:
8812:
8807:
8777:
8772:
8742:
8724:
8722:
8712:
8686:
8667:
8640:
8638:
8615:
8579:
8577:
8549:
8511:
8502:
8445:
8440:
8430:
8428:
8418:
8414:
8404:
8402:
8391:
8387:
8377:
8375:
8366:
8365:
8361:
8351:
8349:
8342:
8326:
8322:
8315:
8307:. p. 127.
8297:
8293:
8283:
8281:
8272:
8271:
8267:
8257:
8255:
8244:
8240:
8230:
8228:
8213:
8212:
8208:
8198:
8196:
8183:
8182:
8178:
8170:
8163:
8159:
8158:
8154:
8144:
8142:
8133:
8132:
8128:
8081:Current Biology
8073:
8069:
8032:Current Biology
8024:
8020:
7981:(3): e1500789.
7960:
7956:
7929:
7925:
7915:
7913:
7898:
7894:
7839:
7835:
7780:
7776:
7766:
7764:
7755:
7754:
7750:
7740:
7738:
7727:
7723:
7713:
7711:
7698:Hays, Jeffrey.
7696:
7692:
7652:
7648:
7638:
7636:
7627:
7626:
7622:
7612:
7610:
7609:on 29 June 2020
7599:
7595:
7560:
7553:
7522:
7511:
7501:
7499:
7479:
7470:
7455:
7439:
7435:
7428:
7402:
7398:
7391:
7365:
7361:
7351:
7349:
7332:
7328:
7318:
7316:
7315:on 3 March 2010
7305:"Animal Actors"
7303:
7302:
7298:
7271:
7267:
7257:
7255:
7244:
7240:
7192:
7185:
7175:
7173:
7166:
7146:
7142:
7132:
7130:
7119:
7115:
7108:
7094:
7090:
7080:
7078:
7061:
7057:
7050:
7036:
7032:
7022:
7020:
7009:
7005:
6996:
6995:
6991:
6981:
6979:
6964:
6960:
6950:
6948:
6933:
6929:
6919:
6917:
6894:
6890:
6837:
6830:
6782:
6778:
6733:
6729:
6688:
6684:
6657:
6653:
6620:
6613:
6572:(18): 6873–77.
6558:
6554:
6499:
6495:
6485:
6483:
6466:
6462:
6431:
6427:
6384:
6380:
6331:
6327:
6320:
6306:
6302:
6279:
6275:
6236:
6232:
6177:
6173:
6142:(13): 1106–11.
6136:Current Biology
6128:
6124:
6069:
6065:
6008:
6004:
5965:
5961:
5938:
5934:
5899:
5895:
5857:Biology Letters
5848:
5844:
5834:
5832:
5817:
5813:
5774:
5770:
5760:
5758:
5721:
5717:
5662:
5655:
5645:
5643:
5636:
5618:
5611:
5570:(5): e1601517.
5556:
5552:
5507:
5503:
5472:(11): 1549–62.
5458:
5454:
5435:
5431:
5424:
5405:
5401:
5358:
5354:
5307:
5298:
5261:
5257:
5206:
5202:
5165:Biology Letters
5157:
5153:
5143:
5141:
5128:
5127:
5123:
5100:
5093:
5062:
5058:
5017:
5013:
5006:
4980:
4973:
4936:
4927:
4920:
4901:
4897:
4890:
4875:
4871:
4864:
4846:
4842:
4801:
4794:
4761:
4754:
4717:
4713:
4673:
4669:
4606:
4602:
4595:
4572:
4568:
4561:
4547:
4543:
4512:
4501:
4456:
4449:
4442:
4415:
4411:
4366:
4362:
4355:
4337:
4333:
4300:
4291:
4250:
4246:
4219:Human Evolution
4215:
4211:
4204:
4190:
4186:
4179:
4159:
4148:
4133:10.2307/3503852
4119:
4106:
4063:
4059:
4021:
4014:
3995:
3991:
3959:
3955:
3918:
3911:
3888:(S63): 153–77.
3870:
3866:
3811:
3807:
3792:
3769:
3765:
3755:
3753:
3752:on 23 July 2018
3742:
3738:
3733:on 10 May 2012.
3730:
3687:
3681:
3674:
3667:
3649:
3634:
3624:
3622:
3599:
3590:
3534:
3530:
3514:
3513:
3506:
3504:
3500:
3467:
3461:
3457:
3418:Current Biology
3410:
3399:
3392:
3374:
3341:
3292:
3288:
3268:
3264:
3254:
3252:
3237:
3236:
3232:
3222:
3220:
3213:Merriam-Webster
3207:
3206:
3202:
3192:
3190:
3177:
3176:
3172:
3164:
3158:
3131:
3122:
3118:
3087:
3080:
3052:American Speech
3047:
3043:
3010:
3006:
2996:
2994:
2981:
2980:
2976:
2962:
2958:
2951:
2933:
2898:
2883:
2879:
2872:
2854:
2843:
2833:
2831:
2816:
2812:
2802:
2800:
2763:
2754:
2744:
2742:
2734:"orangutan, n."
2731:
2730:
2723:
2719:
2673:Primates portal
2671:
2664:
2661:
2578:Bukit Tigapuluh
2570:
2542:tropical forest
2488:
2482:
2477:
2389:Exeter Exchange
2381:
2364:
2336:Terry Pratchett
2312:The Jungle Book
2293:Edgar Allan Poe
2268:media franchise
2207:
2201:
2189:Leakey's Angels
2121:Birutė Galdikas
2113:
2073:
2061:Main articles:
2059:
2008:Birutė Galdikas
1992:
1984:US National Zoo
1921:
1919:
1913:
1907:
1882:
1826:
1825:
1824:
1823:
1822:
1819:
1811:
1810:
1807:
1796:
1779:
1778:
1770:
1768:
1767:
1766:
1765:
1754:
1747:
1744:
1738:
1737:
1736:
1725:
1718:
1715:
1709:
1708:
1707:
1696:
1689:
1686:
1679:
1673:
1668:
1660:Social grooming
1636:
1538:
1431:
1429:
1417:knuckle-walkers
1405:opposable thumb
1339:
1337:Characteristics
1284:P. weidenreichi
1263:Gigantopithecus
1217:
1196:
1187:
1178:
1169:
1160:
780:Systema Naturae
777:in 1758 in the
771:
765:
637:Jacobus Bontius
568:
513:Birutė Galdikas
377:
370:
363:
326:
319:
312:
306:
300:
284:
271:
248:
139:
126:
109:
107:
106:
105:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
36:
35:
32:
24:
17:
12:
11:
5:
13361:
13351:
13350:
13345:
13340:
13335:
13330:
13325:
13320:
13301:
13300:
13298:
13297:
13284:
13271:
13258:
13245:
13232:
13219:
13206:
13193:
13180:
13167:
13154:
13141:
13131:
13118:
13103:
13087:
13085:
13079:
13078:
13064:
13063:
13060:
13059:
13050:
13048:
13045:
13044:
13041:
13040:
13037:
13036:
13033:
13032:
13029:
13028:
13026:
13025:
13015:
13011:Sahelanthropus
13004:
12993:
12983:
12976:
12972:Graecopithecus
12965:
12955:
12943:
12941:
12935:
12934:
12932:
12931:
12923:
12917:
12911:
12910:
12908:
12907:
12903:Nakalipithecus
12896:
12889:
12877:
12875:
12869:
12868:
12866:
12865:
12858:
12850:
12842:
12835:
12828:
12820:
12818:
12806:
12801:
12798:
12797:
12794:
12793:
12791:
12790:
12780:
12773:
12763:
12760:Lufengpithecus
12753:
12750:Khoratpithecus
12743:
12733:
12723:
12720:Ankarapithecus
12712:
12710:
12704:
12703:
12701:
12700:
12697:Ouranopithecus
12690:
12687:Nakalipithecus
12680:
12677:Graecopithecus
12670:
12659:
12649:
12644:
12641:
12640:
12637:
12636:
12634:
12633:
12623:
12616:
12609:
12599:
12592:
12585:
12574:
12572:
12566:
12565:
12563:
12562:
12555:
12548:
12541:
12538:Rangwapithecus
12534:
12526:
12523:Nyanzapithecus
12519:
12511:
12508:Dendropithecus
12503:
12501:
12492:
12491:
12489:
12488:
12478:
12468:
12458:
12448:
12445:Morotopithecus
12438:
12428:
12424:Kamoyapithecus
12417:
12414:Griphopithecus
12407:
12397:
12387:
12376:
12366:
12361:
12354:
12353:
12341:
12340:
12337:
12336:
12329:
12327:
12324:
12323:
12320:
12319:
12316:
12315:
12313:
12312:
12311:
12310:
12304:
12302:
12292:
12291:
12288:
12287:
12284:
12283:
12281:
12280:
12273:
12270:Soromandrillus
12263:
12256:
12246:
12236:
12226:
12216:
12206:
12199:
12192:
12185:
12178:
12168:
12158:
12150:
12148:
12142:
12141:
12139:
12138:
12128:
12121:
12114:
12107:
12100:
12093:
12090:Allenopithecus
12085:
12083:
12081:Cercopithecini
12074:
12068:
12067:
12065:
12064:
12061:Trachypithecus
12057:
12050:
12043:
12033:
12026:
12016:
12009:
12002:
11995:
11988:
11978:
11968:
11961:
11951:
11941:
11931:
11921:
11911:
11904:
11893:
11891:
11885:
11884:
11882:
11881:
11878:Zaltanpithecus
11874:
11867:
11859:
11851:
11843:
11841:
11832:
11831:
11829:
11828:
11817:
11811:
11806:
11803:
11802:
11799:
11798:
11795:
11794:
11792:
11791:
11784:
11777:
11770:
11763:
11756:
11749:
11742:
11734:
11732:
11726:
11725:
11723:
11722:
11715:
11707:
11705:
11699:
11698:
11696:
11695:
11688:
11680:
11678:
11676:Pliopithecidae
11672:
11671:
11669:
11668:
11658:
11651:
11643:
11637:
11628:
11627:
11625:
11624:
11617:
11609:
11607:
11598:
11597:
11595:
11594:
11587:
11580:
11572:
11570:
11561:
11560:
11558:
11557:
11547:
11536:
11526:
11521:
11514:
11513:
11501:
11500:
11497:
11496:
11489:
11487:
11484:
11483:
11480:
11479:
11476:
11475:
11473:
11472:
11471:
11470:
11464:
11462:
11452:
11451:
11448:
11447:
11445:
11444:
11437:
11427:
11420:
11417:Leontopithecus
11413:
11406:
11399:
11392:
11384:
11382:
11380:Callitrichidae
11376:
11375:
11373:
11372:
11365:
11358:
11348:
11338:
11328:
11318:
11311:
11300:
11298:
11292:
11291:
11289:
11288:
11278:
11268:
11258:
11248:
11241:
11231:
11224:
11214:
11204:
11194:
11187:
11180:
11172:
11170:
11164:
11163:
11161:
11160:
11150:
11140:
11130:
11123:
11116:
11106:
11096:
11086:
11076:
11066:
11059:
11052:
11042:
11032:
11025:
11018:
11007:
11005:
10999:
10998:
10996:
10995:
10985:
10977:
10975:
10969:
10968:
10966:
10965:
10955:
10945:
10935:
10925:
10915:
10904:
10898:
10893:
10890:
10889:
10886:
10885:
10882:
10881:
10879:
10878:
10871:
10864:
10857:
10850:
10842:
10840:
10838:Parapithecidae
10834:
10833:
10831:
10830:
10823:
10820:Proteopithecus
10815:
10813:
10807:
10806:
10804:
10803:
10796:
10788:
10782:
10773:
10772:
10770:
10769:
10762:
10755:
10748:
10741:
10734:
10726:
10718:
10716:
10707:
10706:
10704:
10703:
10696:
10689:
10681:
10679:
10670:
10669:
10667:
10666:
10659:
10651:
10649:
10639:
10638:
10636:
10635:
10624:
10613:
10603:
10593:
10581:
10571:
10566:
10559:
10558:
10546:
10545:
10542:
10541:
10532:
10530:
10527:
10526:
10523:
10522:
10520:
10519:
10518:
10517:
10511:
10509:
10503:
10502:
10499:
10498:
10496:
10495:
10484:
10477:
10467:
10464:Hesperotarsius
10457:
10450:
10442:
10440:
10434:
10433:
10431:
10430:
10417:
10406:
10395:
10389:
10379:
10378:
10375:
10374:
10372:
10371:
10364:
10357:
10349:
10347:
10341:
10340:
10338:
10337:
10330:
10323:
10316:
10309:
10302:
10295:
10288:
10281:
10274:
10267:
10260:
10253:
10246:
10239:
10232:
10225:
10222:Gunnelltarsius
10218:
10211:
10204:
10197:
10190:
10183:
10176:
10169:
10161:
10159:
10151:
10150:
10148:
10147:
10140:
10133:
10126:
10119:
10112:
10105:
10098:
10091:
10088:Pseudotetonius
10084:
10077:
10070:
10063:
10056:
10049:
10042:
10034:
10032:
10024:
10023:
10021:
10020:
10013:
10006:
9999:
9992:
9985:
9978:
9971:
9964:
9956:
9954:
9952:Microchoerinae
9948:
9947:
9945:
9944:
9937:
9930:
9922:
9913:
9907:
9897:
9890:
9889:
9877:
9876:
9874:
9873:
9867:
9861:
9855:
9848:
9845:
9844:
9837:
9836:
9829:
9822:
9814:
9805:
9804:
9802:
9801:
9796:
9791:
9786:
9781:
9776:
9771:
9766:
9761:
9756:
9751:
9746:
9741:
9736:
9731:
9726:
9720:
9718:
9712:
9711:
9709:
9708:
9703:
9698:
9693:
9688:
9683:
9678:
9673:
9668:
9663:
9658:
9653:
9648:
9643:
9638:
9633:
9628:
9623:
9618:
9613:
9608:
9603:
9598:
9593:
9588:
9583:
9578:
9573:
9568:
9563:
9558:
9553:
9548:
9543:
9538:
9532:
9530:
9524:
9523:
9521:
9520:
9515:
9510:
9505:
9500:
9495:
9490:
9485:
9480:
9475:
9470:
9465:
9460:
9455:
9450:
9448:Marquis Chimps
9445:
9440:
9435:
9430:
9425:
9420:
9415:
9410:
9405:
9400:
9395:
9390:
9385:
9380:
9375:
9370:
9365:
9360:
9355:
9350:
9348:Pierre Brassau
9345:
9340:
9335:
9330:
9324:
9322:
9316:
9315:
9313:
9312:
9307:
9302:
9296:
9294:
9288:
9287:
9284:
9283:
9278:
9273:
9268:
9262:
9261:
9258:
9257:
9249:
9248:
9241:
9234:
9226:
9217:
9216:
9214:
9213:
9202:
9199:
9198:
9196:
9195:
9190:
9185:
9180:
9175:
9170:
9165:
9160:
9154:
9149:
9144:
9139:
9134:
9129:
9124:
9118:
9112:
9106:
9104:
9100:
9099:
9097:
9096:
9091:
9086:
9081:
9076:
9071:
9066:
9060:
9058:
9050:
9049:
9047:
9046:
9041:
9036:
9031:
9026:
9021:
9016:
9011:
9006:
9001:
8996:
8991:
8986:
8981:
8976:
8970:
8968:
8964:
8963:
8956:
8954:
8952:
8951:
8950:: Hylobatidae)
8941:
8940:
8939:
8929:
8919:
8902:
8901:
8900:
8890:
8873:
8872:
8871:
8868:P. troglodytes
8861:
8844:
8843:
8842:
8824:
8822:
8814:
8813:
8806:
8805:
8798:
8791:
8783:
8774:
8773:
8771:
8770:
8759:
8756:
8755:
8752:
8751:
8748:
8747:
8744:
8743:
8741:
8740:
8730:
8728:
8714:
8713:
8711:
8710:
8706:P. troglodytes
8701:
8691:
8689:
8675:
8669:
8668:
8666:
8665:
8656:
8646:
8644:
8627:
8621:
8620:
8617:
8616:
8614:
8613:
8604:
8595:
8585:
8583:
8566:
8557:
8551:
8550:
8548:
8547:
8541:
8535:
8529:
8523:
8516:
8513:
8512:
8501:
8500:
8493:
8486:
8478:
8472:
8471:
8466:
8461:
8456:
8451:
8444:
8443:External links
8441:
8439:
8438:
8412:
8385:
8359:
8340:
8320:
8314:978-0429576638
8313:
8291:
8265:
8238:
8206:
8176:
8152:
8135:"Willie Smits"
8126:
8067:
8018:
7954:
7923:
7892:
7853:(10): e75373.
7833:
7794:(11): e27491.
7774:
7748:
7721:
7690:
7646:
7620:
7593:
7551:
7509:
7468:
7453:
7433:
7426:
7396:
7389:
7359:
7326:
7296:
7265:
7254:on 7 June 2020
7238:
7195:van Wyhe, John
7183:
7164:
7156:Berghahn Books
7140:
7113:
7107:978-0816041305
7106:
7088:
7055:
7049:978-9042004344
7048:
7030:
7003:
6989:
6958:
6927:
6888:
6828:
6776:
6727:
6700:(2): 189–202.
6682:
6651:
6632:(4): 331–342.
6611:
6552:
6493:
6460:
6425:
6388:Pongo pygmaeus
6378:
6325:
6318:
6300:
6273:
6230:
6171:
6122:
6063:
6002:
5959:
5932:
5903:Pongo pygmaeus
5893:
5842:
5811:
5778:Pongo Pygmaeus
5768:
5715:
5676:(1): e116136.
5653:
5634:
5609:
5550:
5501:
5452:
5429:
5423:978-0199213276
5422:
5399:
5352:
5296:
5255:
5224:(3): 393–416.
5200:
5151:
5121:
5091:
5056:
5011:
5005:978-0521039925
5004:
4971:
4940:Pongo pygmaeus
4925:
4919:978-0199213276
4918:
4895:
4888:
4869:
4863:978-0845134030
4862:
4840:
4792:
4752:
4739:10.1086/410942
4711:
4684:(6): 365–375.
4667:
4612:Pongo pygmaeus
4600:
4594:978-0199213276
4593:
4566:
4560:978-0195043716
4559:
4541:
4499:
4447:
4440:
4428:Westview Press
4409:
4360:
4353:
4331:
4289:
4244:
4209:
4202:
4184:
4177:
4146:
4104:
4057:
4012:
3989:
3953:
3909:
3864:
3805:
3790:
3763:
3736:
3672:
3666:978-0792357551
3665:
3632:
3588:
3528:
3455:
3397:
3390:
3339:
3286:
3262:
3230:
3200:
3170:
3157:978-0521792813
3156:
3116:
3078:
3065:10.2307/455451
3041:
3004:
2974:
2956:
2949:
2896:
2877:
2870:
2841:
2810:
2752:
2720:
2718:
2715:
2714:
2713:
2705:
2698:
2693:
2688:
2683:
2677:
2676:
2660:
2657:
2649:sustainability
2569:
2566:
2504:Sabangau River
2484:Main article:
2481:
2478:
2476:
2473:
2405:Charles Darwin
2363:
2360:
2348:'s 2004 novel
2238:'s 1817 novel
2203:Main article:
2200:
2197:
2179:. Along with
2153:Tanjung Puting
2112:
2109:
2058:
2055:
2051:primatologists
2014:Primatologist
1991:
1988:
1965:self-awareness
1906:
1903:
1881:
1878:
1820:
1813:
1812:
1808:
1801:
1800:
1799:
1798:
1797:
1795:
1792:
1769:
1755:
1745:
1740:
1739:
1726:
1716:
1711:
1710:
1697:
1687:
1682:
1681:
1680:
1671:
1670:
1669:
1667:
1664:
1635:
1632:
1628:phenolic acids
1537:
1534:
1524:S. stercoralis
1520:S. fuelleborni
1518:, the species
1428:
1425:
1382:sagittal crest
1338:
1335:
1311:P. abelii
1293:Giganopithecus
1256:Khoratpithecus
1234:Lufengpithecus
1216:
1213:
1198:
1197:
1193:
1192:
1189:
1188:
1184:
1183:
1180:
1179:
1175:
1174:
1171:
1170:
1166:
1165:
1162:
1161:
1157:
1156:
1153:
1152:
1138:
1135:
1134:
1131:
1130:
1118:
1113:
1111:
1108:
1107:
1104:
1103:
1091:
1086:
1084:
1081:
1080:
1077:
1076:
1065:
1060:
1058:
1055:
1054:
1051:
1050:
1042:
1037:
1035:
1029:
1027:
1026:
1025:
1015:
1001:
1000:
999:
993:
987:
978:Pongo pygmaeus
962:
961:
958:
895:P. abelii
826:P. p. pygmaeus
764:
761:
722:used the term
567:
564:
560:rehabilitation
423:Southeast Asia
397:native to the
386:
385:
375:Billberg, 1828
353:
352:
346:
345:
343:extant species
338:
337:
329:
328:
316:Pongo hooijeri
297:Pongo pygmaeus
291:
290:
286:
285:
272:
268:Pongo pygmaeus
263:
262:
256:
255:
241:
237:
236:
231:
227:
226:
221:
217:
216:
211:
207:
206:
201:
197:
196:
191:
187:
186:
181:
177:
176:
171:
167:
166:
161:
157:
156:
151:
147:
146:
133:
132:
129:Pongo pygmaeus
120:
119:
111:
110:
102:
101:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
33:
15:
9:
6:
4:
3:
2:
13360:
13349:
13346:
13344:
13341:
13339:
13336:
13334:
13331:
13329:
13326:
13324:
13321:
13319:
13316:
13315:
13313:
13306:
13294:
13289:
13285:
13281:
13276:
13272:
13268:
13263:
13259:
13255:
13250:
13246:
13242:
13237:
13233:
13229:
13224:
13220:
13216:
13211:
13207:
13203:
13198:
13194:
13190:
13185:
13181:
13177:
13172:
13168:
13164:
13159:
13155:
13151:
13146:
13142:
13138:
13132:
13128:
13123:
13119:
13114:
13108:
13104:
13099:
13093:
13089:
13088:
13086:
13084:
13080:
13076:
13071:
13055:
13024:
13023:
13016:
13013:
13012:
13005:
13002:
13001:
12994:
12992:
12991:
12990:Kenyanthropus
12984:
12982:
12981:
12977:
12974:
12973:
12966:
12964:
12963:
12956:
12953:
12952:
12945:
12944:
12942:
12940:
12936:
12930:
12929:
12925:
12924:
12921:
12918:
12916:
12912:
12905:
12904:
12897:
12895:
12894:
12890:
12887:
12886:
12879:
12878:
12876:
12874:
12870:
12864:
12863:
12859:
12856:
12855:
12851:
12848:
12847:
12843:
12841:
12840:
12836:
12834:
12833:
12829:
12827:
12826:
12825:Anoiapithecus
12822:
12821:
12819:
12817:
12816:Dryopithecini
12810:
12807:
12804:
12799:
12789:
12788:
12781:
12779:
12778:
12774:
12772:
12771:
12764:
12762:
12761:
12754:
12752:
12751:
12744:
12742:
12741:
12734:
12732:
12731:
12724:
12722:
12721:
12714:
12713:
12711:
12709:
12705:
12699:
12698:
12691:
12689:
12688:
12681:
12679:
12678:
12671:
12669:
12668:
12661:
12660:
12657:
12653:
12650:
12647:
12642:
12632:
12631:
12624:
12622:
12621:
12617:
12615:
12614:
12610:
12608:
12607:
12600:
12598:
12597:
12593:
12591:
12590:
12586:
12584:
12583:
12576:
12575:
12573:
12571:
12567:
12561:
12560:
12556:
12554:
12553:
12549:
12547:
12546:
12545:Rukwapithecus
12542:
12540:
12539:
12535:
12532:
12531:
12527:
12525:
12524:
12520:
12517:
12516:
12515:Micropithecus
12512:
12510:
12509:
12505:
12504:
12502:
12500:
12493:
12487:
12486:
12479:
12477:
12476:
12469:
12467:
12466:
12465:Otavipithecus
12459:
12457:
12456:
12449:
12447:
12446:
12439:
12437:
12436:
12435:Kenyapithecus
12429:
12426:
12425:
12418:
12416:
12415:
12408:
12406:
12405:
12398:
12396:
12395:
12388:
12386:
12385:
12378:
12377:
12374:
12370:
12367:
12364:
12359:
12355:
12351:
12346:
12342:
12333:
12308:
12307:
12306:
12305:
12303:
12301:
12297:
12293:
12279:
12278:
12277:Theropithecus
12274:
12272:
12271:
12264:
12262:
12261:
12257:
12255:
12254:
12247:
12245:
12244:
12243:Procercocebus
12237:
12235:
12234:
12227:
12225:
12224:
12217:
12215:
12214:
12207:
12205:
12204:
12200:
12198:
12197:
12193:
12191:
12190:
12186:
12184:
12183:
12179:
12177:
12176:
12175:Gorgopithecus
12169:
12167:
12166:
12159:
12157:
12156:
12152:
12151:
12149:
12147:
12143:
12137:
12136:
12129:
12127:
12126:
12122:
12120:
12119:
12115:
12113:
12112:
12108:
12106:
12105:
12104:Cercopithecus
12101:
12099:
12098:
12097:Allochrocebus
12094:
12092:
12091:
12087:
12086:
12084:
12082:
12078:
12075:
12073:
12069:
12063:
12062:
12058:
12056:
12055:
12051:
12049:
12048:
12047:Semnopithecus
12044:
12042:
12041:
12034:
12032:
12031:
12030:Rhinopithecus
12027:
12025:
12024:
12017:
12015:
12014:
12010:
12008:
12007:
12003:
12001:
12000:
11996:
11994:
11993:
11989:
11987:
11986:
11985:Parapresbytis
11979:
11977:
11976:
11969:
11967:
11966:
11962:
11960:
11959:
11952:
11950:
11949:
11942:
11940:
11939:
11932:
11930:
11929:
11928:Kuseracolobus
11922:
11920:
11919:
11912:
11910:
11909:
11905:
11903:
11902:
11895:
11894:
11892:
11890:
11886:
11880:
11879:
11875:
11873:
11872:
11868:
11865:
11864:
11860:
11857:
11856:
11852:
11850:
11849:
11845:
11844:
11842:
11840:
11833:
11827:
11826:
11819:
11818:
11815:
11812:
11809:
11804:
11790:
11789:
11785:
11783:
11782:
11778:
11776:
11775:
11774:Laccopithecus
11771:
11769:
11768:
11764:
11762:
11761:
11760:Egarapithecus
11757:
11755:
11754:
11750:
11748:
11747:
11743:
11741:
11740:
11736:
11735:
11733:
11731:
11727:
11721:
11720:
11716:
11714:
11713:
11709:
11708:
11706:
11704:
11700:
11694:
11693:
11689:
11687:
11686:
11682:
11681:
11679:
11677:
11673:
11667:
11666:
11659:
11657:
11656:
11652:
11650:
11649:
11645:
11644:
11641:
11638:
11636:
11629:
11623:
11622:
11618:
11616:
11615:
11611:
11610:
11608:
11606:
11599:
11593:
11592:
11591:Talahpithecus
11588:
11586:
11585:
11584:Oligopithecus
11581:
11579:
11578:
11574:
11573:
11571:
11569:
11562:
11556:
11555:
11548:
11546:
11545:
11544:Micropithecus
11538:
11537:
11534:
11530:
11527:
11524:
11519:
11515:
11511:
11506:
11502:
11493:
11468:
11467:
11466:
11465:
11463:
11461:
11457:
11453:
11443:
11442:
11438:
11436:
11435:
11428:
11426:
11425:
11421:
11419:
11418:
11414:
11412:
11411:
11407:
11405:
11404:
11400:
11398:
11397:
11393:
11391:
11390:
11386:
11385:
11383:
11381:
11377:
11371:
11370:
11366:
11364:
11363:
11359:
11357:
11356:
11349:
11347:
11346:
11339:
11337:
11336:
11329:
11327:
11326:
11319:
11317:
11316:
11312:
11310:
11309:
11302:
11301:
11299:
11297:
11293:
11287:
11286:
11279:
11277:
11276:
11269:
11267:
11266:
11265:Protopithecus
11259:
11257:
11256:
11249:
11247:
11246:
11242:
11240:
11239:
11232:
11230:
11229:
11225:
11223:
11222:
11215:
11213:
11212:
11205:
11203:
11202:
11195:
11193:
11192:
11188:
11186:
11185:
11181:
11179:
11178:
11174:
11173:
11171:
11169:
11165:
11159:
11158:
11151:
11149:
11148:
11141:
11139:
11138:
11131:
11129:
11128:
11127:Plecturocebus
11124:
11122:
11121:
11117:
11115:
11114:
11107:
11105:
11104:
11103:Miocallicebus
11097:
11095:
11094:
11087:
11085:
11084:
11077:
11075:
11074:
11067:
11065:
11064:
11060:
11058:
11057:
11053:
11051:
11050:
11043:
11041:
11040:
11033:
11031:
11030:
11026:
11024:
11023:
11019:
11017:
11016:
11009:
11008:
11006:
11004:
11000:
10994:
10993:
10986:
10984:
10983:
10979:
10978:
10976:
10974:
10970:
10964:
10963:
10956:
10954:
10953:
10946:
10944:
10943:
10936:
10934:
10933:
10926:
10924:
10923:
10916:
10914:
10913:
10906:
10905:
10902:
10899:
10896:
10891:
10877:
10876:
10872:
10870:
10869:
10865:
10863:
10862:
10858:
10856:
10855:
10851:
10849:
10848:
10844:
10843:
10841:
10839:
10835:
10829:
10828:
10824:
10822:
10821:
10817:
10816:
10814:
10812:
10808:
10802:
10801:
10797:
10795:
10794:
10790:
10789:
10786:
10783:
10781:
10774:
10768:
10767:
10766:Siamopithecus
10763:
10761:
10760:
10756:
10754:
10753:
10749:
10747:
10746:
10742:
10740:
10739:
10735:
10732:
10731:
10730:Bugtipithecus
10727:
10725:
10724:
10723:Amphipithecus
10720:
10719:
10717:
10715:
10708:
10702:
10701:
10697:
10695:
10694:
10690:
10688:
10687:
10683:
10682:
10680:
10678:
10671:
10665:
10664:
10660:
10658:
10657:
10653:
10652:
10650:
10647:
10646:Afrotarsiidae
10640:
10633:
10632:
10625:
10622:
10621:
10614:
10612:
10611:
10610:Aseanpithecus
10604:
10602:
10601:
10594:
10591:
10590:
10583:
10582:
10579:
10575:
10572:
10569:
10564:
10560:
10556:
10551:
10547:
10537:
10515:
10514:
10513:
10512:
10510:
10508:
10504:
10493:
10492:
10485:
10483:
10482:
10478:
10476:
10475:
10468:
10466:
10465:
10458:
10456:
10455:
10454:Cephalopachus
10451:
10449:
10448:
10444:
10443:
10441:
10439:
10435:
10428:
10426:
10418:
10415:
10414:
10413:Afrotarsiidae
10407:
10405:
10404:
10397:
10396:
10393:
10390:
10388:
10384:
10380:
10370:
10369:
10365:
10363:
10362:
10358:
10356:
10355:
10351:
10350:
10348:
10346:
10345:Tarkadectinae
10342:
10336:
10335:
10331:
10329:
10328:
10324:
10322:
10321:
10317:
10315:
10314:
10310:
10308:
10307:
10303:
10301:
10300:
10296:
10294:
10293:
10289:
10287:
10286:
10282:
10280:
10279:
10275:
10273:
10272:
10268:
10266:
10265:
10261:
10259:
10258:
10254:
10252:
10251:
10247:
10245:
10244:
10240:
10238:
10237:
10233:
10231:
10230:
10226:
10224:
10223:
10219:
10217:
10216:
10215:Ekwiiyemakius
10212:
10210:
10209:
10205:
10203:
10202:
10198:
10196:
10195:
10191:
10189:
10188:
10184:
10182:
10181:
10177:
10175:
10174:
10170:
10168:
10167:
10166:Ageitodendron
10163:
10162:
10160:
10157:
10152:
10146:
10145:
10141:
10139:
10138:
10134:
10132:
10131:
10127:
10125:
10124:
10120:
10118:
10117:
10113:
10111:
10110:
10106:
10104:
10103:
10099:
10097:
10096:
10092:
10090:
10089:
10085:
10083:
10082:
10078:
10076:
10075:
10071:
10069:
10068:
10064:
10062:
10061:
10057:
10055:
10054:
10050:
10048:
10047:
10046:Anaptomorphus
10043:
10041:
10040:
10036:
10035:
10033:
10030:
10025:
10019:
10018:
10014:
10012:
10011:
10007:
10005:
10004:
10000:
9998:
9997:
9993:
9991:
9990:
9986:
9984:
9983:
9979:
9977:
9976:
9972:
9970:
9969:
9965:
9963:
9962:
9958:
9957:
9955:
9953:
9949:
9943:
9942:
9938:
9936:
9935:
9931:
9928:
9927:
9923:
9920:
9919:
9915:
9914:
9911:
9908:
9904:
9895:
9891:
9887:
9882:
9878:
9872:
9868:
9866:
9862:
9860:
9856:
9854:
9850:
9849:
9846:
9842:
9835:
9830:
9828:
9823:
9821:
9816:
9815:
9812:
9800:
9797:
9795:
9792:
9790:
9787:
9785:
9782:
9780:
9777:
9775:
9772:
9770:
9767:
9765:
9762:
9760:
9757:
9755:
9752:
9750:
9747:
9745:
9742:
9740:
9737:
9735:
9732:
9730:
9727:
9725:
9722:
9721:
9719:
9717:
9713:
9707:
9704:
9702:
9699:
9697:
9694:
9692:
9689:
9687:
9684:
9682:
9679:
9677:
9674:
9672:
9669:
9667:
9664:
9662:
9659:
9657:
9654:
9652:
9649:
9647:
9644:
9642:
9639:
9637:
9634:
9632:
9629:
9627:
9624:
9622:
9619:
9617:
9614:
9612:
9609:
9607:
9604:
9602:
9599:
9597:
9594:
9592:
9589:
9587:
9584:
9582:
9579:
9577:
9574:
9572:
9569:
9567:
9564:
9562:
9559:
9557:
9554:
9552:
9549:
9547:
9544:
9542:
9539:
9537:
9534:
9533:
9531:
9529:
9525:
9519:
9516:
9514:
9511:
9509:
9506:
9504:
9501:
9499:
9496:
9494:
9491:
9489:
9486:
9484:
9481:
9479:
9476:
9474:
9471:
9469:
9466:
9464:
9463:J. Fred Muggs
9461:
9459:
9456:
9454:
9451:
9449:
9446:
9444:
9441:
9439:
9436:
9434:
9431:
9429:
9426:
9424:
9421:
9419:
9416:
9414:
9411:
9409:
9406:
9404:
9401:
9399:
9396:
9394:
9391:
9389:
9386:
9384:
9381:
9379:
9376:
9374:
9371:
9369:
9366:
9364:
9361:
9359:
9356:
9354:
9351:
9349:
9346:
9344:
9341:
9339:
9336:
9334:
9331:
9329:
9326:
9325:
9323:
9321:
9317:
9311:
9308:
9306:
9303:
9301:
9298:
9297:
9295:
9293:
9289:
9282:
9279:
9277:
9274:
9272:
9269:
9267:
9264:
9263:
9259:
9255:
9247:
9242:
9240:
9235:
9233:
9228:
9227:
9224:
9212:
9204:
9203:
9200:
9194:
9191:
9189:
9186:
9184:
9181:
9179:
9176:
9174:
9171:
9169:
9166:
9164:
9161:
9158:
9155:
9153:
9150:
9148:
9145:
9143:
9140:
9138:
9135:
9133:
9130:
9128:
9125:
9122:
9121:Apes in space
9119:
9116:
9113:
9111:
9108:
9107:
9105:
9101:
9095:
9092:
9090:
9087:
9085:
9082:
9080:
9077:
9075:
9072:
9070:
9067:
9065:
9062:
9061:
9059:
9057:
9056:social status
9051:
9045:
9042:
9040:
9037:
9035:
9032:
9030:
9027:
9025:
9022:
9020:
9017:
9015:
9012:
9010:
9007:
9005:
9002:
9000:
8997:
8995:
8992:
8990:
8987:
8985:
8982:
8980:
8977:
8975:
8972:
8971:
8969:
8967:Study of apes
8965:
8960:
8949:
8945:
8942:
8937:
8933:
8930:
8927:
8923:
8920:
8917:
8913:
8910:
8909:
8908:
8907:
8903:
8898:
8894:
8891:
8888:
8884:
8881:
8880:
8879:
8878:
8874:
8869:
8865:
8862:
8859:
8855:
8852:
8851:
8850:
8849:
8845:
8840:
8836:
8833:
8832:
8831:
8830:
8826:
8825:
8823:
8821:
8815:
8811:
8804:
8799:
8797:
8792:
8790:
8785:
8784:
8781:
8769:
8761:
8760:
8757:
8739:
8737:
8732:
8731:
8729:
8721:
8720:
8715:
8709:
8707:
8702:
8700:
8698:
8693:
8692:
8690:
8685:
8684:
8679:
8676:
8674:
8670:
8664:
8662:
8657:
8655:
8653:
8648:
8647:
8645:
8637:
8636:
8631:
8628:
8626:
8622:
8612:
8610:
8605:
8603:
8601:
8596:
8594:
8592:
8587:
8586:
8584:
8576:
8575:
8570:
8567:
8565:
8561:
8558:
8556:
8552:
8546:
8542:
8540:
8536:
8534:
8530:
8528:
8524:
8522:
8518:
8517:
8514:
8510:
8499:
8494:
8492:
8487:
8485:
8480:
8479:
8476:
8470:
8467:
8465:
8462:
8460:
8457:
8455:
8452:
8450:
8447:
8446:
8427:
8423:
8416:
8400:
8396:
8389:
8373:
8369:
8363:
8347:
8343:
8337:
8333:
8332:
8324:
8316:
8310:
8306:
8302:
8295:
8279:
8275:
8269:
8253:
8249:
8242:
8226:
8222:
8221:
8216:
8210:
8194:
8190:
8186:
8180:
8169:
8162:
8156:
8140:
8136:
8130:
8122:
8118:
8114:
8110:
8105:
8100:
8095:
8090:
8086:
8082:
8078:
8071:
8063:
8059:
8055:
8051:
8046:
8041:
8037:
8033:
8029:
8022:
8014:
8010:
8005:
8000:
7996:
7992:
7988:
7984:
7980:
7976:
7975:
7970:
7968:
7958:
7950:
7946:
7943:(3): 329–39.
7942:
7938:
7934:
7927:
7911:
7907:
7903:
7896:
7888:
7884:
7879:
7874:
7869:
7864:
7860:
7856:
7852:
7848:
7844:
7837:
7829:
7825:
7820:
7815:
7810:
7805:
7801:
7797:
7793:
7789:
7785:
7778:
7762:
7758:
7752:
7736:
7732:
7725:
7709:
7705:
7701:
7694:
7686:
7682:
7678:
7674:
7670:
7666:
7662:
7658:
7650:
7634:
7630:
7624:
7608:
7604:
7597:
7588:
7583:
7579:
7575:
7574:
7569:
7567:
7558:
7556:
7546:
7541:
7537:
7533:
7532:
7527:
7520:
7518:
7516:
7514:
7498:
7494:
7493:
7488:
7486:
7477:
7475:
7473:
7464:
7460:
7456:
7454:0-89672-430-1
7450:
7446:
7445:
7437:
7429:
7427:9780674015777
7423:
7419:
7415:
7410:
7409:
7400:
7392:
7386:
7382:
7378:
7373:
7372:
7363:
7347:
7343:
7342:
7337:
7330:
7314:
7310:
7306:
7300:
7292:
7288:
7284:
7280:
7276:
7269:
7253:
7249:
7242:
7234:
7230:
7226:
7222:
7217:
7212:
7208:
7204:
7200:
7196:
7190:
7188:
7171:
7167:
7161:
7157:
7153:
7152:
7144:
7128:
7124:
7117:
7109:
7103:
7099:
7092:
7076:
7072:
7071:
7066:
7059:
7051:
7045:
7041:
7034:
7018:
7014:
7007:
6999:
6993:
6977:
6973:
6969:
6962:
6946:
6942:
6938:
6931:
6915:
6911:
6907:
6903:
6899:
6892:
6884:
6880:
6875:
6870:
6866:
6862:
6858:
6854:
6850:
6846:
6842:
6835:
6833:
6824:
6820:
6816:
6812:
6808:
6804:
6800:
6796:
6792:
6788:
6780:
6772:
6768:
6763:
6758:
6754:
6750:
6746:
6742:
6738:
6731:
6723:
6719:
6715:
6711:
6707:
6703:
6699:
6695:
6694:
6686:
6678:
6674:
6671:(7): 839–76.
6670:
6666:
6662:
6655:
6647:
6643:
6639:
6635:
6631:
6627:
6626:
6618:
6616:
6607:
6603:
6598:
6593:
6588:
6583:
6579:
6575:
6571:
6567:
6563:
6556:
6548:
6544:
6539:
6534:
6529:
6524:
6520:
6516:
6512:
6508:
6504:
6497:
6481:
6477:
6476:
6471:
6464:
6456:
6452:
6448:
6444:
6440:
6436:
6429:
6421:
6417:
6413:
6409:
6405:
6401:
6398:(5): 334–41.
6397:
6393:
6389:
6382:
6374:
6370:
6366:
6362:
6358:
6354:
6350:
6346:
6343:(4): 186–88.
6342:
6338:
6337:
6329:
6321:
6315:
6311:
6304:
6296:
6292:
6288:
6284:
6277:
6269:
6265:
6261:
6257:
6253:
6249:
6245:
6241:
6234:
6226:
6222:
6217:
6212:
6207:
6202:
6198:
6194:
6190:
6186:
6182:
6175:
6167:
6163:
6159:
6155:
6150:
6145:
6141:
6137:
6133:
6126:
6118:
6114:
6109:
6104:
6099:
6094:
6090:
6086:
6082:
6078:
6074:
6067:
6059:
6055:
6050:
6045:
6040:
6035:
6031:
6027:
6024:(9): e74896.
6023:
6019:
6018:
6013:
6006:
5998:
5994:
5990:
5986:
5982:
5978:
5975:(2): 109–17.
5974:
5970:
5963:
5955:
5951:
5948:(2): 175–88.
5947:
5943:
5936:
5928:
5924:
5920:
5916:
5912:
5908:
5904:
5897:
5889:
5885:
5880:
5875:
5871:
5867:
5864:(2): 172–75.
5863:
5859:
5858:
5853:
5846:
5830:
5826:
5822:
5815:
5807:
5803:
5799:
5795:
5792:(2): 137–52.
5791:
5787:
5783:
5779:
5772:
5756:
5752:
5748:
5743:
5738:
5734:
5730:
5726:
5719:
5711:
5707:
5702:
5697:
5692:
5687:
5683:
5679:
5675:
5671:
5667:
5660:
5658:
5641:
5637:
5631:
5627:
5623:
5616:
5614:
5605:
5601:
5596:
5591:
5586:
5581:
5577:
5573:
5569:
5565:
5561:
5554:
5546:
5542:
5537:
5532:
5528:
5524:
5521:(3): 435–55.
5520:
5516:
5512:
5505:
5497:
5493:
5488:
5483:
5479:
5475:
5471:
5467:
5463:
5456:
5448:
5444:
5440:
5433:
5425:
5419:
5414:
5413:
5403:
5395:
5391:
5387:
5383:
5379:
5375:
5372:(3): 177–81.
5371:
5367:
5363:
5356:
5348:
5344:
5339:
5334:
5329:
5324:
5320:
5316:
5312:
5305:
5303:
5301:
5292:
5288:
5284:
5280:
5277:(2): 93–101.
5276:
5272:
5271:
5266:
5259:
5251:
5247:
5242:
5237:
5232:
5227:
5223:
5219:
5215:
5213:
5204:
5196:
5192:
5187:
5182:
5178:
5174:
5170:
5166:
5162:
5155:
5139:
5135:
5131:
5125:
5117:
5113:
5109:
5105:
5098:
5096:
5087:
5083:
5079:
5075:
5071:
5067:
5060:
5052:
5048:
5044:
5040:
5036:
5032:
5028:
5024:
5023:
5015:
5007:
5001:
4997:
4993:
4988:
4987:
4978:
4976:
4967:
4963:
4959:
4955:
4952:(1): 201–18.
4951:
4947:
4946:
4941:
4934:
4932:
4930:
4921:
4915:
4910:
4909:
4899:
4891:
4885:
4881:
4873:
4865:
4859:
4855:
4851:
4844:
4836:
4832:
4828:
4824:
4820:
4816:
4812:
4808:
4807:
4799:
4797:
4788:
4784:
4780:
4776:
4772:
4768:
4767:
4759:
4757:
4748:
4744:
4740:
4736:
4733:(4): 493–94.
4732:
4728:
4727:
4722:
4715:
4707:
4703:
4699:
4695:
4691:
4687:
4683:
4679:
4671:
4663:
4659:
4654:
4649:
4644:
4639:
4635:
4631:
4627:
4623:
4619:
4617:
4613:
4604:
4596:
4590:
4586:
4581:
4580:
4570:
4562:
4556:
4552:
4545:
4537:
4533:
4529:
4525:
4521:
4517:
4510:
4508:
4506:
4504:
4495:
4491:
4486:
4481:
4477:
4473:
4470:(7679): 151.
4469:
4465:
4461:
4454:
4452:
4443:
4437:
4433:
4429:
4425:
4424:
4419:
4413:
4405:
4401:
4396:
4391:
4387:
4383:
4380:(4): 373–82.
4379:
4375:
4371:
4364:
4356:
4350:
4346:
4342:
4335:
4326:
4321:
4318:(5): 643–52.
4317:
4313:
4309:
4305:
4298:
4296:
4294:
4285:
4281:
4277:
4273:
4269:
4265:
4262:(4): 305–19.
4261:
4257:
4256:
4248:
4240:
4236:
4232:
4228:
4225:(5): 343–50.
4224:
4220:
4213:
4205:
4203:0-12-045590-0
4199:
4195:
4188:
4180:
4174:
4170:
4166:
4165:
4157:
4155:
4153:
4151:
4142:
4138:
4134:
4130:
4126:
4125:
4117:
4115:
4113:
4111:
4109:
4100:
4096:
4092:
4088:
4084:
4080:
4076:
4072:
4068:
4061:
4053:
4049:
4045:
4041:
4038:(6): 770–97.
4037:
4033:
4032:
4027:
4019:
4017:
4008:
4004:
4000:
3993:
3985:
3981:
3977:
3973:
3969:
3965:
3957:
3949:
3945:
3941:
3937:
3933:
3929:
3928:
3923:
3916:
3914:
3905:
3901:
3896:
3891:
3887:
3883:
3879:
3877:
3868:
3860:
3856:
3851:
3846:
3841:
3836:
3832:
3828:
3824:
3820:
3816:
3809:
3801:
3797:
3793:
3791:0-801-88221-4
3787:
3783:
3782:
3777:
3776:Wilson, D. E.
3773:
3772:Groves, C. P.
3767:
3751:
3747:
3740:
3729:
3725:
3721:
3716:
3711:
3707:
3703:
3700:(3): 447–55.
3699:
3695:
3694:
3686:
3679:
3677:
3668:
3662:
3658:
3654:
3647:
3645:
3643:
3641:
3639:
3637:
3620:
3616:
3612:
3608:
3604:
3597:
3595:
3593:
3584:
3580:
3575:
3570:
3566:
3562:
3558:
3554:
3550:
3546:
3545:
3540:
3532:
3524:
3518:
3499:
3495:
3491:
3487:
3483:
3479:
3475:
3474:
3466:
3459:
3451:
3447:
3442:
3437:
3432:
3427:
3423:
3419:
3415:
3408:
3406:
3404:
3402:
3393:
3387:
3383:
3379:
3372:
3370:
3368:
3366:
3364:
3362:
3360:
3358:
3356:
3354:
3352:
3350:
3348:
3346:
3344:
3335:
3331:
3327:
3323:
3319:
3315:
3311:
3307:
3304:(5): 431–37.
3303:
3299:
3298:
3290:
3282:
3278:
3277:
3272:
3266:
3250:
3246:
3241:
3234:
3218:
3214:
3210:
3204:
3188:
3184:
3180:
3174:
3163:
3159:
3153:
3149:
3145:
3141:
3137:
3130:
3126:
3120:
3112:
3108:
3104:
3100:
3097:(5): 811–30.
3096:
3092:
3085:
3083:
3074:
3070:
3066:
3062:
3059:(2): 134–62.
3058:
3054:
3053:
3045:
3037:
3033:
3029:
3025:
3021:
3017:
3016:
3015:English Today
3008:
2992:
2988:
2984:
2978:
2970:
2969:
2960:
2952:
2946:
2942:
2938:
2931:
2929:
2927:
2925:
2923:
2921:
2919:
2917:
2915:
2913:
2911:
2909:
2907:
2905:
2903:
2901:
2892:
2888:
2881:
2873:
2867:
2863:
2859:
2852:
2850:
2848:
2846:
2829:
2825:
2821:
2814:
2798:
2794:
2790:
2785:
2780:
2777:(4): 532–41.
2776:
2772:
2768:
2761:
2759:
2757:
2740:
2735:
2728:
2726:
2721:
2712:
2710:
2706:
2704:
2703:
2699:
2697:
2694:
2692:
2689:
2687:
2684:
2682:
2679:
2678:
2674:
2668:
2663:
2656:
2654:
2650:
2646:
2642:
2638:
2634:
2630:
2626:
2622:
2618:
2614:
2610:
2606:
2601:
2597:
2595:
2591:
2587:
2579:
2574:
2565:
2561:
2558:
2553:
2551:
2547:
2543:
2539:
2538:fragmentation
2535:
2534:deforestation
2531:
2524:
2519:
2515:
2513:
2509:
2505:
2501:
2497:
2493:
2492:IUCN Red List
2487:
2472:
2470:
2466:
2462:
2456:
2454:
2450:
2449:hairy Houdini
2446:
2441:
2440:Animal rights
2437:
2436:San Diego Zoo
2433:
2429:
2425:
2421:
2417:
2413:
2408:
2406:
2402:
2398:
2394:
2390:
2376:
2372:
2368:
2359:
2357:
2356:
2351:
2347:
2343:
2342:
2337:
2333:
2332:The Librarian
2328:
2327:
2322:
2318:
2314:
2313:
2308:
2304:
2300:
2299:
2294:
2291:and the 1841
2290:
2289:
2284:
2279:
2277:
2273:
2269:
2265:
2264:Pierre Boulle
2261:
2260:
2255:
2251:
2247:
2243:
2242:
2237:
2233:
2231:
2227:
2220:
2219:Daniel Vierge
2216:
2211:
2206:
2196:
2194:
2190:
2186:
2182:
2178:
2174:
2171:plantations,
2170:
2166:
2162:
2158:
2154:
2148:
2146:
2145:Petrus Camper
2141:
2138:
2134:
2130:
2122:
2117:
2108:
2106:
2102:
2098:
2094:
2093:United States
2090:
2086:
2082:
2078:
2072:
2068:
2064:
2054:
2052:
2048:
2044:
2043:
2038:
2033:
2030:
2024:
2021:
2017:
2012:
2009:
2001:
2000:San Diego Zoo
1996:
1987:
1985:
1981:
1977:
1973:
1968:
1966:
1963:, indicating
1962:
1958:
1953:
1951:
1947:
1943:
1939:
1918:
1912:
1902:
1898:
1895:
1886:
1877:
1873:
1871:
1866:
1857:
1853:
1851:
1847:
1843:
1838:
1836:
1831:
1817:
1805:
1791:
1787:
1785:
1777:
1775:
1763:
1759:
1756:Throatscrape
1743:
1734:
1730:
1714:
1705:
1701:
1685:
1666:Communication
1663:
1661:
1656:
1651:
1649:
1640:
1631:
1629:
1625:
1621:
1617:
1616:mineral licks
1613:
1612:shelter tubes
1609:
1604:
1602:
1598:
1594:
1590:
1589:
1582:
1580:
1577:
1573:
1572:
1567:
1563:
1559:
1555:
1551:
1542:
1533:
1531:
1530:
1525:
1521:
1517:
1516:Strongyloides
1513:
1512:
1508:
1504:
1503:
1502:Strongyloides
1499:of the genus
1498:
1494:
1490:
1486:
1480:
1478:
1474:
1470:
1466:
1462:
1458:
1454:
1446:
1424:
1420:
1418:
1414:
1409:
1406:
1397:
1393:
1391:
1387:
1383:
1379:
1375:
1371:
1366:
1364:
1360:
1356:
1352:
1343:
1334:
1332:
1328:
1324:
1320:
1316:
1312:
1308:
1304:
1303:
1298:
1294:
1290:
1286:
1285:
1280:
1276:
1272:
1267:
1265:
1264:
1259:
1257:
1252:
1251:
1246:
1245:
1240:
1236:
1235:
1230:
1226:
1222:
1215:Fossil record
1209:
1204:
1191:
1190:
1182:
1181:
1173:
1172:
1164:
1163:
1155:
1154:
1151:
1149:
1148:
1143:
1137:
1136:
1133:
1132:
1129:
1127:
1123:
1117:
1116:
1110:
1109:
1106:
1105:
1102:
1100:
1097: (genus
1096:
1090:
1089:
1083:
1082:
1079:
1078:
1075:
1074:
1072:
1064:
1063:
1057:
1056:
1053:
1052:
1049:
1047:
1041:
1040:
1033:
1032:
1028:
1023:
1019:
1016:
1013:
1009:
1005:
1002:
997:
994:
991:
988:
985:
982:
981:
979:
975:
972:
971:
970:
969:
964:
963:
957:
952:
949:
947:
943:
938:
935:
931:
927:
926:mitochondrial
923:
919:
915:
912:According to
910:
908:
907:which have 46
904:
900:
896:
892:
888:
884:
880:
872:
868:
864:
861:Flanged male
859:
855:
853:
849:
845:
841:
840:
835:
834:P. p. wurmbii
831:
827:
823:
819:
815:
811:
810:
805:
801:
797:
796:
790:
786:
785:Carl Linnaeus
782:
781:
776:
770:
760:
758:
755:
751:
748:
744:
741:
737:
733:
729:
725:
721:
717:
713:
709:
705:
704:Andrew Battel
701:
696:
694:
690:
686:
682:
678:
674:
670:
666:
662:
657:
655:
654:Nicolaes Tulp
651:
646:
642:
638:
633:
631:
627:
623:
619:
615:
614:
610:
606:
605:
600:
596:
592:
587:
585:
581:
577:
573:
563:
561:
557:
553:
549:
545:
541:
540:deforestation
537:
533:
529:
525:
521:
516:
514:
509:
505:
500:
498:
494:
489:
484:
479:
477:
473:
469:
465:
461:
457:
453:
449:
445:
441:
437:
436:
432:
428:
424:
420:
416:
412:
408:
404:
400:
396:
392:
384:
380:
373:
368:Fischer, 1813
366:
359:
354:
351:
347:
344:
339:
335:
330:
327:
325:
324:
318:
317:
311:
310:
305:
304:
299:
298:
292:
287:
282:
279:
278:Simia satyrus
275:
270:
269:
264:
261:
257:
252:
247:
246:
242:
239:
238:
235:
232:
229:
228:
225:
222:
219:
218:
215:
212:
209:
208:
205:
202:
199:
198:
195:
192:
189:
188:
185:
182:
179:
178:
175:
172:
169:
168:
165:
162:
159:
158:
155:
152:
149:
148:
143:
138:
134:
130:
125:
121:
117:
112:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
29:
26:
22:
13305:
13082:
13022:Paranthropus
13020:
13009:
12998:
12988:
12978:
12970:
12960:
12951:Ardipithecus
12949:
12926:
12901:
12891:
12883:
12862:Rudapithecus
12860:
12852:
12844:
12839:Dryopithecus
12837:
12830:
12823:
12787:Sivapithecus
12785:
12776:
12775:
12770:Meganthropus
12768:
12758:
12748:
12740:Indopithecus
12738:
12728:
12718:
12695:
12685:
12675:
12665:
12628:
12620:Symphalangus
12618:
12611:
12604:
12594:
12587:
12582:Bunopithecus
12580:
12557:
12550:
12543:
12536:
12530:Oreopithecus
12528:
12521:
12513:
12506:
12483:
12473:
12463:
12453:
12443:
12433:
12422:
12412:
12402:
12392:
12384:Afropithecus
12382:
12275:
12268:
12258:
12251:
12241:
12231:
12221:
12211:
12201:
12194:
12187:
12180:
12173:
12165:Dinopithecus
12163:
12153:
12135:Nanopithecus
12133:
12123:
12118:Erythrocebus
12116:
12109:
12102:
12095:
12088:
12059:
12052:
12045:
12038:
12028:
12023:Rhinocolobus
12021:
12011:
12004:
11997:
11992:Piliocolobus
11990:
11983:
11973:
11963:
11958:Microcolobus
11956:
11948:Mesopithecus
11946:
11938:Libypithecus
11936:
11926:
11916:
11906:
11899:
11876:
11869:
11863:Prohylobates
11861:
11853:
11848:Noropithecus
11846:
11823:
11786:
11779:
11772:
11765:
11758:
11751:
11744:
11737:
11730:Crouzeliidae
11717:
11710:
11692:Pliopithecus
11690:
11683:
11663:
11653:
11646:
11619:
11612:
11589:
11582:
11577:Catopithecus
11575:
11552:
11542:
11439:
11432:
11422:
11415:
11408:
11401:
11394:
11387:
11367:
11360:
11353:
11343:
11333:
11325:Dolichocebus
11323:
11313:
11306:
11283:
11273:
11263:
11253:
11243:
11236:
11226:
11219:
11209:
11199:
11189:
11182:
11175:
11155:
11145:
11135:
11125:
11118:
11111:
11101:
11093:Mazzonicebus
11091:
11081:
11071:
11061:
11054:
11049:Cebupithecia
11047:
11037:
11027:
11020:
11015:Antillothrix
11013:
10990:
10980:
10960:
10950:
10940:
10930:
10922:Perupithecus
10920:
10910:
10873:
10866:
10861:Parapithecus
10859:
10852:
10845:
10825:
10818:
10798:
10791:
10764:
10757:
10750:
10743:
10736:
10728:
10721:
10698:
10691:
10684:
10661:
10654:
10631:Phileosimias
10629:
10618:
10608:
10598:
10589:Altiatlasius
10587:
10491:Xanthorhysis
10489:
10479:
10474:Oligotarsius
10472:
10462:
10452:
10445:
10422:
10411:
10401:
10387:Tarsiiformes
10366:
10359:
10352:
10332:
10325:
10318:
10311:
10304:
10297:
10290:
10283:
10276:
10269:
10262:
10255:
10250:Macrotarsius
10248:
10241:
10234:
10227:
10220:
10213:
10206:
10199:
10192:
10185:
10178:
10171:
10164:
10142:
10135:
10128:
10121:
10114:
10107:
10102:Strigorhysis
10100:
10095:Sphacorhysis
10093:
10086:
10079:
10074:Chlororhysis
10072:
10065:
10058:
10051:
10044:
10037:
10015:
10008:
10001:
9994:
9987:
9980:
9975:Microchoerus
9973:
9966:
9959:
9939:
9932:
9926:Altiatlasius
9924:
9916:
9715:
9468:Nim Chimpsky
9069:Research ban
9029:Elgin Center
9014:Willie Smits
8989:Jane Goodall
8935:
8925:
8915:
8905:
8904:
8896:
8886:
8875:
8867:
8857:
8846:
8838:
8827:
8735:
8717:
8705:
8704:Chimpanzee (
8696:
8681:
8660:
8651:
8633:
8608:
8599:
8590:
8580:(Orangutans)
8573:
8572:
8508:(great apes)
8429:. Retrieved
8426:Eco-Business
8425:
8415:
8403:. Retrieved
8397:. Mongabay.
8388:
8376:. Retrieved
8372:the original
8362:
8350:. Retrieved
8330:
8323:
8300:
8294:
8282:. Retrieved
8278:the original
8268:
8256:. Retrieved
8252:the original
8241:
8229:. Retrieved
8218:
8209:
8197:. Retrieved
8193:the original
8188:
8179:
8168:the original
8155:
8143:. Retrieved
8129:
8084:
8080:
8070:
8035:
8031:
8021:
7978:
7972:
7967:Pongo abelii
7966:
7957:
7940:
7936:
7932:
7926:
7914:. Retrieved
7895:
7850:
7846:
7836:
7791:
7787:
7777:
7765:. Retrieved
7751:
7739:. Retrieved
7735:the original
7724:
7712:. Retrieved
7703:
7693:
7663:(1): 50–56.
7660:
7656:
7649:
7637:. Retrieved
7623:
7611:. Retrieved
7607:the original
7596:
7577:
7571:
7565:
7535:
7529:
7500:. Retrieved
7496:
7490:
7485:Pongo abelii
7484:
7443:
7436:
7407:
7399:
7370:
7362:
7350:. Retrieved
7341:BBC Newsbeat
7339:
7329:
7317:. Retrieved
7313:the original
7299:
7274:
7268:
7256:. Retrieved
7252:the original
7241:
7206:
7202:
7174:. Retrieved
7150:
7143:
7131:. Retrieved
7127:the original
7116:
7097:
7091:
7079:. Retrieved
7068:
7058:
7039:
7033:
7021:. Retrieved
7006:
6992:
6980:. Retrieved
6972:The Guardian
6971:
6961:
6949:. Retrieved
6941:The Guardian
6940:
6930:
6918:. Retrieved
6902:The Guardian
6901:
6891:
6848:
6844:
6790:
6786:
6779:
6744:
6740:
6730:
6697:
6691:
6685:
6668:
6664:
6660:
6654:
6629:
6623:
6569:
6565:
6555:
6510:
6506:
6496:
6484:. Retrieved
6473:
6463:
6438:
6434:
6428:
6395:
6391:
6387:
6381:
6340:
6334:
6328:
6309:
6303:
6286:
6282:
6276:
6246:(1): 56–64.
6243:
6239:
6233:
6191:(1): 12289.
6188:
6184:
6174:
6139:
6135:
6125:
6080:
6076:
6066:
6021:
6015:
6005:
5972:
5968:
5962:
5945:
5941:
5935:
5913:(1): 23–32.
5910:
5906:
5902:
5896:
5861:
5855:
5845:
5833:. Retrieved
5824:
5814:
5789:
5785:
5781:
5777:
5771:
5759:. Retrieved
5732:
5728:
5718:
5673:
5669:
5644:. Retrieved
5625:
5567:
5563:
5553:
5518:
5514:
5510:
5504:
5469:
5465:
5455:
5446:
5442:
5432:
5411:
5402:
5369:
5365:
5361:
5355:
5318:
5314:
5274:
5268:
5264:
5258:
5221:
5217:
5211:
5203:
5168:
5164:
5154:
5142:. Retrieved
5124:
5103:
5072:(1): 33–46.
5069:
5065:
5059:
5029:(1): 69–86.
5026:
5020:
5014:
4985:
4949:
4943:
4939:
4907:
4898:
4879:
4872:
4854:Alan R. Liss
4849:
4843:
4810:
4804:
4770:
4764:
4730:
4724:
4720:
4714:
4681:
4677:
4670:
4625:
4621:
4615:
4611:
4603:
4578:
4569:
4550:
4544:
4519:
4515:
4467:
4463:
4422:
4412:
4377:
4373:
4363:
4344:
4334:
4315:
4311:
4259:
4253:
4247:
4222:
4218:
4212:
4193:
4187:
4163:
4122:
4074:
4070:
4060:
4035:
4029:
4025:
4001:(76): 1–24.
3998:
3992:
3967:
3963:
3956:
3931:
3925:
3921:
3885:
3881:
3875:
3867:
3822:
3818:
3808:
3780:
3766:
3754:. Retrieved
3750:the original
3739:
3728:the original
3697:
3691:
3652:
3623:. Retrieved
3606:
3548:
3542:
3531:
3517:cite journal
3505:. Retrieved
3477:
3471:
3458:
3421:
3417:
3377:
3301:
3295:
3289:
3275:
3265:
3253:. Retrieved
3244:
3240:"pongo, n.1"
3233:
3221:. Retrieved
3212:
3203:
3191:. Retrieved
3182:
3173:
3162:the original
3135:
3119:
3094:
3090:
3056:
3050:
3044:
3022:(4): 44–50.
3019:
3013:
3007:
2995:. Retrieved
2986:
2977:
2966:
2959:
2936:
2893:(4): 141–44.
2890:
2886:
2880:
2857:
2832:. Retrieved
2823:
2813:
2801:. Retrieved
2774:
2770:
2743:. Retrieved
2741:. March 2022
2738:
2709:Skullduggery
2708:
2700:
2696:Orang Pendek
2625:Bukit Lawang
2602:
2598:
2590:Willie Smits
2583:
2580:, Indonesia.
2562:
2554:
2527:
2489:
2475:Conservation
2457:
2452:
2409:
2386:
2382: 1830s
2370:
2362:In captivity
2353:
2349:
2339:
2324:
2310:
2296:
2295:short story
2286:
2283:Walter Scott
2280:
2257:
2253:
2239:
2234:
2229:
2225:
2223:
2193:Louis Leakey
2181:Jane Goodall
2177:forest fires
2149:
2142:
2136:
2128:
2126:
2074:
2040:
2034:
2025:
2019:
2013:
2005:
1969:
1954:
1935:
1905:Intelligence
1899:
1891:
1874:
1862:
1839:
1827:
1788:
1780:
1771:
1742:Throatscrape
1698:Nest smack,
1652:
1645:
1605:
1601:slow lorises
1586:
1583:
1569:
1550:fruit-eaters
1547:
1527:
1523:
1519:
1515:
1509:
1500:
1481:
1455:and inhabit
1450:
1421:
1410:
1402:
1367:
1348:
1326:
1314:
1310:
1306:
1300:
1292:
1288:
1282:
1273:is from the
1270:
1268:
1261:
1254:
1250:Sivapithecus
1248:
1244:Indopithecus
1242:
1232:
1218:
1207:
1145:
1140:
1125:
1120:
1098:
1093:
1070:
1068:
1067:
1044:
1021:
1008:Pongo abelii
1007:
995:
989:
983:
977:
967:
965:
955:
942:land bridges
911:
894:
889:) have less
886:
876:
851:
847:
837:
833:
829:
825:
817:
807:
803:
792:
788:
778:
772:
756:
749:
742:
731:
723:
699:
697:
692:
684:
676:
668:
665:Orang-Outang
664:
663:in the form
658:
640:
634:
625:
617:
611:
602:
599:Old Javanese
594:
590:
588:
583:
579:
575:
569:
556:conservation
517:
501:
480:
459:
451:
443:
434:
433:
390:
389:
382:Lesson, 1840
378:
371:
364:
357:
356:
321:
314:
307:
303:Pongo abelii
301:
295:
294:
277:
266:
260:Type species
244:
243:
210:Infraorder:
128:
25:
13210:iNaturalist
13107:Wikispecies
12570:Hylobatidae
12260:Rungwecebus
12125:Miopithecus
12111:Chlorocebus
12040:Sawecolobus
11975:Paracolobus
11739:Anapithecus
11655:Paidopithex
11410:Leontocebus
11345:Panamacebus
11255:Paralouatta
11191:Brachyteles
11083:Insulacebus
11003:Pitheciidae
10942:Canaanimico
10895:Platyrrhini
10847:Abuqatrania
10663:Afrotarsius
10568:Simiiformes
10555:Simiiformes
10507:Simiiformes
10361:Tarkadectes
10201:Diablomomys
10180:Brontomomys
10116:Teilhardina
10060:Arapahovius
10053:Anemorhysis
10017:Vectipithex
10010:Quercyloris
10003:Pseudoloris
9982:Nannopithex
9968:Melaneremia
9606:John Daniel
9443:Macaco Tião
9428:Little Mama
9403:Jo Mendi II
9320:Chimpanzees
9159:(non-human)
9123:(non-human)
9117:(non-human)
9024:Ian Redmond
8979:Dian Fossey
8926:P. pygmaeus
8887:G. beringei
8858:P. paniscus
8820:ape species
8697:P. paniscus
8652:G. beringei
8600:P. pygmaeus
8352:16 December
8104:10230/34400
7352:25 December
7258:24 February
7176:16 December
7081:26 February
6982:10 November
6951:10 November
6851:(1): 8932.
5646:16 December
5449:(1): 10–12.
5241:10023/18054
4994:. pp.
4813:(1): 1–20.
4522:(1): 1–35.
4430:. pp.
3441:10230/34400
2997:20 December
2983:"Orangutan"
2820:"Orangutan"
2711:(1970 film)
2512:Batang Toru
2416:tea parties
2393:smock-frock
2272:bureaucrats
2185:Dian Fossey
2173:gold miners
2159:thesis for
1961:mirror test
1950:reciprocity
1946:Leipzig Zoo
1942:Zoo Atlanta
1894:build nests
1892:Orangutans
1870:infanticide
1653:Orangutans
1634:Social life
1461:dipterocarp
1374:prognathous
1370:endocranial
1122:chimpanzees
944:during the
879:chimpanzees
848:P. pygmaeus
830:P. p. morio
814:René Lesson
661:John Bulwer
472:chimpanzees
444:P. pygmaeus
427:South China
419:Pleistocene
399:rainforests
230:Subfamily:
214:Simiiformes
13318:Orangutans
13312:Categories
12404:Equatorius
12363:Hominoidea
12350:Hominoidea
12309:see below↓
12300:Hominoidea
12196:Mandrillus
12182:Lophocebus
12155:Cercocebus
12006:Procolobus
11767:Fanchangia
11523:Catarrhini
11510:Catarrhini
11469:see below↓
11460:Catarrhini
11396:Callithrix
11335:Killikaike
11238:Mohanamico
11221:Chilecebus
11147:Soriacebus
11113:Nuciruptor
11073:Homunculus
11063:Chiropotes
11056:Cheracebus
11039:Carlocebus
11029:Callicebus
10992:Tremacebus
10962:Lagonimico
10952:Branisella
10932:Szalatavus
10759:Pondaungia
10677:Eosimiidae
10516:see below↓
10403:Archicebus
10285:Shoshonius
10229:Hemiacodon
10208:Dyseolemur
10194:Chumashius
10137:Trogolemur
10130:Tetonoides
10039:Absarokius
9989:Necrolemur
9934:Bownomomys
9886:Haplorhini
9841:Haplorhini
9754:Joe Martin
9716:Orangutans
9418:Kokomo Jr.
9310:Panbanisha
9173:Monkey Day
9163:Great apes
9064:Personhood
8897:G. gorilla
8864:Chimpanzee
8839:H. sapiens
8736:H. sapiens
8661:G. gorilla
8641:(Gorillas)
8545:Haplorhini
8543:Suborder:
8506:Hominidae
8378:16 January
7767:16 January
7576:. 2017.3.
7502:28 January
7416:. p.
7379:. p.
7319:28 January
7133:18 January
6513:(1): 1–6.
5735:: 149–96.
5536:2144/39274
4127:(4): 1–6.
4077:: 103233.
3756:6 November
3625:27 January
3480:: 97–164.
3245:OED Online
3193:4 December
2717:References
2691:Monkey Day
2496:Appendix I
2465:wet nurses
2461:Frank Buck
2424:Joe Martin
2397:London Zoo
2346:Dale Smith
2317:King Louie
2241:Melincourt
2199:In fiction
2057:Personhood
1915:See also:
1774:media help
1727:Raspberry
1684:Nest smack
1579:strychnine
1469:freshwater
1413:suspension
1378:brow ridge
873:orangutans
767:See also:
716:great apes
681:Portuguese
554:. Several
395:great apes
391:Orangutans
372:Macrobates
361:Oken, 1816
204:Haplorhini
200:Suborder:
31:Orangutans
12873:Gorillini
12803:Homininae
12646:Hominidae
12596:Hylobates
12475:Proconsul
12233:Pliopapio
12223:Parapapio
12146:Papionini
12013:Pygathrix
11999:Presbytis
11889:Colobinae
11788:Pliobates
11753:Crouzelia
11554:Saadanius
11389:Callimico
11308:Acrecebus
11285:Stirtonia
11228:Lagothrix
11211:Cartelles
11157:Xenothrix
10912:Parvimico
10438:Tarsiidae
10425:Omomyidae
10320:Washakius
10306:Uintanius
10257:Nesomomys
10187:Chipetaia
10173:Asiomomys
10156:Omomyinae
10109:Tatmanius
10067:Aycrossia
9996:Paraloris
9961:Indusomys
9903:Omomyidae
9851:Kingdom:
9769:Ken Allen
9706:Willie B.
9681:Snowflake
9661:Pattycake
9576:Gargantua
9546:Binti Jua
9483:Panpanzee
9054:Legal and
8916:P. abelii
8906:Orangutan
8625:Homininae
8591:P. abelii
8555:Hominidae
8519:Kingdom:
8113:0960-9822
7225:1369-8486
7209:: 53–63.
6910:0261-3077
6865:2045-2322
6665:Behaviour
6441:: 50–63.
6289:: 19–33.
5825:USA Today
5545:198190605
5250:195329265
4099:251879262
3970:: 68–74.
3934:: 59–67.
3255:4 October
3223:4 October
3111:221192990
3036:144326996
2793:228828226
2550:pet trade
2445:Ken Allen
2341:Discworld
2276:Dr. Zaius
2097:sanctuary
2035:During a
1846:ovulating
1713:Raspberry
1560:but also
1497:nematodes
1493:wild dogs
1390:resonance
1363:chocolate
1319:Sundaland
1225:subfamily
1012:Lake Toba
883:great ape
852:P. abelii
844:Lake Toba
818:P. abelii
809:P. abelii
734:or other
685:orangutan
673:incorrect
650:cretinism
645:Banjarese
630:Old Malay
618:urangutan
595:urangutan
591:orangutan
589:The word
566:Etymology
552:pet trade
481:The most
452:P. abelii
403:Indonesia
224:Hominidae
160:Kingdom:
154:Eukaryota
13241:12100803
13134:BioLib:
13092:Wikidata
12939:Hominina
12915:Hominini
12832:Danuvius
12708:Ponginae
12667:Buronius
12613:Nomascus
12552:Simiolus
11441:Saguinus
11403:Cebuella
11355:Patasola
11275:Solimoea
11177:Alouatta
11168:Atelidae
11120:Pithecia
10868:Qatrania
10793:Arsinoea
10693:Eosimias
10327:Wyomomys
10292:Steinius
10278:Rooneyia
10236:Jemezius
10144:Walshina
10123:Tetonius
10081:Gazinius
9941:Kohatius
9918:Altanius
9871:Primates
9865:Mammalia
9859:Chordata
9857:Phylum:
9853:Animalia
9528:Gorillas
9373:Gregoire
9211:Category
9132:Bushmeat
8768:Category
8725:(Humans)
8695:Bonobo (
8673:Hominini
8564:Ponginae
8539:Primates
8533:Mammalia
8527:Chordata
8525:Phylum:
8521:Animalia
8405:26 March
8399:Archived
8346:Archived
8258:29 April
8231:29 April
8225:Archived
8220:The Week
8139:Archived
8121:29103940
8054:29456144
8013:26973868
7910:Archived
7906:Mongabay
7887:24130707
7847:PLOS ONE
7828:22096582
7788:PLOS ONE
7761:Archived
7708:Archived
7677:17899314
7657:Primates
7633:Archived
7463:43207125
7346:Archived
7283:35012032
7233:25861859
7170:Archived
7075:Archived
7017:Archived
6998:"Sandra"
6976:Archived
6945:Archived
6914:Archived
6883:38698007
6874:11066025
6823:25139547
6815:12511649
6771:19656794
6714:17160669
6646:11275962
6606:22509022
6547:30150738
6486:5 August
6480:Archived
6455:25038033
6420:19354930
6412:11093037
6373:27180148
6260:19052691
6240:Primates
6225:31444387
6166:17892549
6158:19500987
6117:30443595
6058:24040357
6017:PLoS One
5997:85330986
5989:31979488
5927:44204663
5888:19126529
5835:12 April
5829:Archived
5755:Archived
5751:16702785
5710:25569211
5670:PLOS ONE
5640:Archived
5604:28560319
5496:19701484
5394:21561581
5386:11746281
5347:19812079
5291:13583879
5195:34582737
5144:23 April
5138:Archived
5116:40349739
5086:40824300
5051:13366732
5043:23179533
5022:Primates
4966:84778685
4835:13557435
4827:12065937
4787:54322235
4698:21770970
4662:29192145
4536:40513842
4494:29120449
4420:(1987).
4404:19619166
4306:(2002).
4284:83661029
4276:31964053
4239:83923845
4091:36030625
4052:24210657
4007:2246/259
3904:28105715
3859:30427876
3819:PLOS ONE
3800:62265494
3774:(2005).
3724:21074627
3657:Springer
3619:Archived
3583:21270892
3498:Archived
3494:29045930
3450:29103940
3273:(1827).
3249:Archived
3217:Archived
3187:Archived
3127:(2002).
2991:Archived
2828:Archived
2803:12 April
2797:Archived
2745:4 August
2659:See also
2557:bushmeat
2546:palm oil
2530:poaching
2305:'s 1967
2266:and the
2169:palm oil
2101:Americas
1976:laughter
1835:oestrous
1655:disperse
1608:geophagy
1576:alkaloid
1568:species
1514:. Among
1505:and the
1463:and old
1453:arboreal
1355:arm span
1279:Chongzuo
1231:such as
1221:Ponginae
1095:gorillas
934:X-linked
930:Y-linked
871:Tapanuli
867:Sumatran
795:pygmaeus
757:yimpungu
736:cognates
712:gorillas
613:Ramayana
609:Sanskrit
544:palm oil
528:bushmeat
524:poaching
504:primates
488:Dominant
483:arboreal
468:gorillas
464:Ponginae
407:Malaysia
365:Lophotus
350:Synonyms
289:Species
281:Linnaeus
276:, 1799 (
274:Lacépède
251:Lacépède
234:Ponginae
220:Family:
194:Primates
184:Mammalia
174:Chordata
170:Phylum:
164:Animalia
150:Domain:
13288:ZooBank
13202:5219531
13000:Orrorin
12893:Gorilla
12589:Hoolock
11965:Nasalis
11908:Colobus
11434:Micodon
11369:Sapajus
11362:Saimiri
11296:Cebidae
11245:Oreonax
11201:Caipora
11022:Cacajao
10973:Aotidae
10854:Apidium
10827:Serapia
10800:Biretia
10686:Bahinia
10656:Afrasia
10620:Nosmips
10600:Amamria
10481:Tarsius
10447:Carlito
10368:Tarkops
10334:Yaquius
10299:Stockia
10271:Ourayia
10243:Loveina
9869:Order:
9863:Class:
9744:Chantek
9729:Ah Meng
9676:Shabani
9641:Michael
9586:Harambe
9493:Santino
9353:Bubbles
9292:Bonobos
9127:Bigfoot
9110:Primate
9103:Related
8877:Gorilla
8734:Human (
8635:Gorilla
8537:Order:
8531:Class:
8431:3 April
8199:2 April
8062:3756682
8004:4783118
7983:Bibcode
7878:3793980
7855:Bibcode
7819:3214049
7796:Bibcode
7685:1792717
7023:15 June
7015:. CNN.
6795:Bibcode
6787:Science
6762:2817314
6722:6875716
6597:3344992
6574:Bibcode
6538:6110832
6515:Bibcode
6365:8643126
6345:Bibcode
6216:6707206
6193:Bibcode
6108:6235548
6085:Bibcode
6049:3770631
6026:Bibcode
5879:2665816
5806:9642783
5761:19 June
5701:4287529
5678:Bibcode
5595:5435413
5572:Bibcode
5487:2728907
5338:2842634
5186:8478518
4747:2826733
4653:5709421
4630:Bibcode
4472:Bibcode
4395:2766055
4343:(ed.).
4141:3503852
3972:Bibcode
3936:Bibcode
3850:6235281
3827:Bibcode
3715:3046308
3574:3060778
3553:Bibcode
3334:3355899
3326:8875856
3306:Bibcode
3209:"pongo"
3179:"pongo"
2434:of the
2412:simians
2344:and in
2165:loggers
2133:Sarawak
2099:in the
2091:in the
2081:bonobos
2047:applied
2029:deceive
1880:Nesting
1764:format.
1735:format.
1706:format.
1624:tannins
1507:ciliate
1386:canines
1297:fossils
1258:piriyai
1223:. This
1099:Gorilla
1046:gibbons
922:Miocene
918:gibbons
903:diploid
863:Bornean
639:' 1631
548:logging
506:. They
415:Sumatra
379:Satyrus
283:, 1760)
240:Genus:
190:Order:
180:Class:
103:↓
13267:417949
13228:572839
13189:1PONGG
13098:Q41050
12394:Ekembo
12189:Macaca
12054:Simias
11825:Alophe
11184:Ateles
10745:Krabia
10738:Ganlea
10313:Utahia
10264:Omomys
9794:Sandra
9739:Bonnie
9671:Samson
9621:Kokomo
9611:Jumoke
9556:Bokito
9518:Washoe
9508:Travis
9503:Sultan
9478:Pankun
9473:Oliver
9433:Loulis
9408:Julius
9358:Cheeta
9338:Azalea
8948:family
8944:Gibbon
8854:Bonobo
8818:Extant
8338:
8311:
8284:10 May
8145:11 May
8119:
8111:
8060:
8052:
8011:
8001:
7916:11 May
7885:
7875:
7826:
7816:
7741:11 May
7714:12 May
7683:
7675:
7639:1 July
7613:1 July
7461:
7451:
7424:
7387:
7291:598387
7289:
7281:
7231:
7223:
7162:
7104:
7046:
6908:
6881:
6871:
6863:
6821:
6813:
6769:
6759:
6720:
6712:
6644:
6604:
6594:
6545:
6535:
6453:
6418:
6410:
6371:
6363:
6316:
6268:708682
6266:
6258:
6223:
6213:
6164:
6156:
6115:
6105:
6056:
6046:
5995:
5987:
5925:
5886:
5876:
5804:
5749:
5708:
5698:
5632:
5602:
5592:
5543:
5494:
5484:
5420:
5392:
5384:
5345:
5335:
5289:
5248:
5193:
5183:
5114:
5084:
5049:
5041:
5002:
4996:193–94
4964:
4916:
4886:
4860:
4833:
4825:
4785:
4745:
4706:997385
4704:
4696:
4660:
4650:
4591:
4557:
4534:
4492:
4464:Nature
4438:
4402:
4392:
4351:
4282:
4274:
4237:
4200:
4175:
4169:420–23
4139:
4097:
4089:
4050:
3902:
3857:
3847:
3798:
3788:
3722:
3712:
3663:
3607:Nature
3581:
3571:
3544:Nature
3492:
3448:
3388:
3332:
3324:
3154:
3109:
3073:455451
3071:
3034:
2947:
2868:
2791:
2321:Mowgli
2303:Disney
2285:novel
2069:, and
2020:Neesia
1980:Bonnie
1762:Vorbis
1733:Vorbis
1704:Vorbis
1620:kaolin
1562:drupes
1485:tigers
1359:maroon
1142:humans
966:Genus
932:, and
822:Borneo
793:Simia
750:mpungu
732:mpongi
708:Angola
574:words
476:humans
474:, and
411:Borneo
358:Faunus
253:, 1799
13280:40897
13215:43581
13176:42005
13163:63NZX
13137:32343
13127:Pongo
13113:Pongo
13083:Pongo
12777:Pongo
12606:Junzi
12203:Papio
11315:Cebus
10982:Aotus
10354:Tarka
9799:Tonda
9774:Manis
9764:Karta
9759:Karen
9749:Jenny
9724:Abang
9696:Trudy
9691:Titus
9686:Timmy
9656:Ozzie
9651:Ndume
9631:Massa
9626:Louis
9601:Jenny
9596:Jambo
9571:Fatou
9541:Babec
9498:Sarah
9393:Jimmy
9388:Jiggs
9363:Congo
9343:Bonzo
9333:Ayumu
9305:Nyota
9300:Kanzi
9193:Yowie
9183:Yeren
8835:Human
8574:Pongo
8171:(PDF)
8164:(PDF)
8058:S2CID
7933:Pongo
7681:S2CID
6920:2 May
6819:S2CID
6718:S2CID
6475:Wired
6416:S2CID
6369:S2CID
6264:S2CID
6162:S2CID
5993:S2CID
5923:S2CID
5747:S2CID
5541:S2CID
5390:S2CID
5287:S2CID
5246:S2CID
5171:(9).
5082:S2CID
5047:S2CID
4962:S2CID
4831:S2CID
4783:S2CID
4743:JSTOR
4702:S2CID
4532:S2CID
4280:S2CID
4235:S2CID
4137:JSTOR
4095:S2CID
4026:Pongo
3922:Pongo
3731:(PDF)
3688:(PDF)
3507:1 May
3501:(PDF)
3490:S2CID
3468:(PDF)
3330:S2CID
3165:(PDF)
3132:(PDF)
3107:S2CID
3069:JSTOR
3032:S2CID
2834:4 May
2789:S2CID
2500:CITES
2432:Jiggs
2401:Jenny
2375:Jenny
2274:like
2217:" by
2137:mawas
2129:maias
1597:honey
1554:arils
1445:Sabah
1315:Pongo
1289:Pongo
1271:Pongo
1071:Pongo
968:Pongo
956:Pongo
804:Pongo
754:Yombi
752:, or
743:pungu
740:Lumbu
730:word
728:Kongo
724:Pongo
700:Pongo
693:hutan
689:Dutch
677:hutan
626:Pongo
584:Pongo
580:hutan
576:orang
572:Malay
542:(for
532:crops
526:(for
497:honey
435:Pongo
431:genus
245:Pongo
13254:9599
13249:NCBI
13223:ITIS
13197:GBIF
13184:EPPO
13150:4537
13145:BOLD
12980:Homo
11665:Kapi
11424:Mico
9701:Toto
9666:Pogo
9646:Nico
9616:Koko
9591:Ivan
9566:Colo
9551:Bobo
9513:Viki
9488:Sami
9458:Moja
9438:Lucy
9423:Lana
9398:Jinx
9368:Enos
9254:apes
9188:Yeti
8829:Homo
8810:Apes
8719:Homo
8433:2012
8407:2012
8380:2014
8354:2015
8336:ISBN
8309:ISBN
8286:2020
8260:2020
8233:2020
8201:2012
8147:2020
8117:PMID
8109:ISSN
8050:PMID
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