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On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection

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of the views therein set forth, that he proposed, in a letter to Sir Charles Lyell, to obtain Mr. Wallace's consent to allow the Essay to be published as soon as possible. Of this step we highly approved, provided Mr. Darwin did not withhold from the public, as he was strongly inclined to do (in favour of Mr. Wallace), the memoir which he had himself written on the same subject, and which, as before stated, one of us had perused in 1844, and the contents of which we had both of us been privy to for many years. On representing this to Mr. Darwin, he gave us permission to make what use we thought proper of his memoir, &c.; and in adopting our present course, of presenting it to the Linnean Society, we have explained to him that we are not solely considering the relative claims to priority of himself and his friend, but the interests of science generally".
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of business that had been dealt with, or possibly due to polite reluctance to speak out against a theory which the eminent Lyell and Hooker were supporting. Bentham noted that the audience appeared fatigued. Hooker later said there was "no semblance of a discussion", though "it was talked over with bated breath" at tea afterwards, and in his reminiscences many years later thought "the subject too ominous for the old school to enter the lists before armouring." Although Bell apparently disapproved, the Vice-President promptly removed all references to immutability from his own paper which was awaiting publication.
160:. This made guarded comments about evolution, and in the spring of 1856 it was noticed by Lyell who drew it to the attention of Darwin who was then working out a strategy for presenting his theory. Darwin apparently mistook Wallace's meaning, writing "nothing very new ... Uses my simile of tree, it seems all creation with him". However, he spelt out the details of Natural Selection to Lyell, who found the idea hard to accept but urged Darwin to publish to establish priority. On 14 May 1856 Darwin began what became his draft for a book titled 330:
neither of them having published his views, though Mr. Darwin has for many years past been repeatedly urged by us to do so, and both authors having now unreservedly placed their papers in our hands, we think it would best promote the interests of science that a selection from them should be laid before the Linnean Society. Taken in the order of their dates, they consist of:—
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abstract!" While Wallace had not asked for publication, Darwin would, "of course, at once write and offer to send to any journal" that Wallace chose. He sadly added that "all my originality, whatever it may amount to, will be smashed". Lyell's immediate response urged Darwin to publish his own ideas, and in his reply of 25 June Darwin agreed that he could point to his own
134:, he became an able geologist as well as collecting plant and animal specimens, and fossils of gigantic extinct mammals. By the return journey, he was connecting patterns of geographical and historical distribution, and starting to doubt the stability of Species. In September 1838 he conceived his theory of 265:
was persuaded by Hooker to step down so that the Darwin and Wallace papers were first on the agenda, followed by six other papers on botanical and zoological topics. Bell had introduced discussions at the end of meetings, but there was no discussion of natural selection, perhaps because of the amount
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in his presidential report of May 1859 wrote that "The year which has passed has not, indeed, been marked by any of those striking discoveries which at once revolutionize, so to speak, the department of science on which they bear". Years later, Darwin could only recall one review; Professor Haughton
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of 1857 showing that he was still developing the ideas, "so that I could most truly say and prove that I take nothing from Wallace. I should be extremely glad now to publish a sketch of my general views in about a dozen pages or so. But I cannot persuade myself that I can do so honourably... I would
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of the steam engine, which checks and corrects any irregularities almost before they become evident; and in like manner no unbalanced deficiency in the animal kingdom can ever reach any conspicuous magnitude, because it would make itself felt at the very first step, by rendering existence difficult
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The communication letter briefly describes the first two, and regarding Wallace's paper states that he sent to Darwin "with the expressed wish that it should be forwarded to Sir Charles Lyell, if Mr. Darwin thought it sufficiently novel and interesting. So highly did Mr. Darwin appreciate the value
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The gentlemen having, independently and unknown to one another, conceived the same very ingenious theory to account for the appearance and perpetuation of varieties and of specific forms on our planet, and both fairly claim the merit of being original thinkers in this important line of inquiry; but
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MY DEAR SIR,—The accompanying papers, which we have the honour of communicating to the Linnean Society, and which all related to the same subject, viz. the Laws which affect the Production of Varieties, Races, and Species, contain the results of the investigations of two indefatigable naturalists,
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Darwin wrote to Wallace to explain what had occurred, enclosing a letter which Hooker wrote at Darwin's request. His next action was to get his family away from the danger of infection, both himself and Emma exhausted by sickening fear about the safety of their children. The day after the funeral
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Wallace collected specimens and corresponded with Darwin from Borneo. In December 1857, he wrote to ask if Darwin's book would delve into human origins, to which Darwin responded that "I think I shall avoid the whole subject, as so surrounded with prejudices, though I fully admit that it is the
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in February 1858 and sent it to Darwin with a request to send it on to Lyell. Darwin received it on 18 June 1858, and wrote to Lyell that "your words have come true with a vengeance,... forestalled" and "If Wallace had my MS. sketch written out in 1842, he could not have made a better short
142:, then as well as developing his career as a geologist and writer worked privately on finding evidence and answering possible objections. He wrote out his ideas in an 1842 "pencil sketch", then in an essay written in 1844. He discussed transmutation with his friend 217:
on 10 June 1858, and was to be held on 1 July. On the afternoon of 30 June Mrs. Hooker copied out extracts from the handwritten documents they had just received from Darwin, then that evening Lyell and Hooker handed them in to the secretary with a covering letter.
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on 28 June. On the morning of the 29th he acknowledged Hooker's letters, writing "I cannot think now", then that night he read the letters, and though "quite prostrated", got his servant to take to Hooker Wallace's essay, the letter to Asa Gray and the
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far rather burn my whole book than that he or any man should think that I had behaved in a paltry spirit", also requesting that Hooker be invited to give a second opinion. Darwin was overwrought by a deepening crisis of illness of his baby son
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would observe in the 1970s that, although seeing it only as an illustration, Wallace had "probably said the most powerful thing that’d been said in the 19th century". Bateson revisited the topic in his 1979 book
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highest & most interesting problem for the naturalist". He encouraged Wallace's theorising, saying "without speculation there is no good & original observation", adding that "I go much further than you".
381:, whose judgement was "on asking myself, 'What does all this prove?' the only answer I could make was, 'A possibility'." It was commented on in some reviews and letters. The Linnean Society President 340:
consisting of a portion of a Chapter entitled, "On the Variation of Organic Beings in a state of Nature; on the Natural Means of Selection; on the Comparison of Domestic Races and true Species."
1588: 1352: 146:, who read the essay in 1847. After turning his attention to biology and completing eight years of work on barnacles, Darwin intensified work on his theory of species in 1854. 1376: 1360: 1053: 234:. Neither author was present. Darwin was attending the funeral of his son, and Wallace was still in Borneo. The meeting was chaired by the president of the society, 1583: 1303: 1440: 1320: 922:
The autobiography of Charles Darwin 1809–1882. With the original omissions restored. Edited and with appendix and notes by his grand-daughter Nora Barlow.
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and Darwin pressed on, with the facts he felt necessary to support his "abstract" expanding far beyond the thirty pages Hooker had originally suggested.
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About thirty were present, including two unnamed guests from overseas, with many there to hear an obituary notice for the former president and botanist
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dated 30 June 1858, giving information on the three sections of the papers. The introduction is shown below, along with the section headings:
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given by Lyell. Wallace's natural history agent Samuel Stevens happened to be present, while Darwin's friends there included
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On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection
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and Arthur Henfry would later support evolution, while Cuthbert Collingwood became an opponent.
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of Dublin claimed that "all that was new in them was false, and what was true was old."
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Despite illness, Darwin pressed on with writing the "abstract" of his "big book" on
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took a break between graduating and starting his career as a clergyman to study the
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of Wallace's paper together with an extract from Darwin's essay and his letter to
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The life and letters of Charles Darwin, including an autobiographical chapter.
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The foundations of The origin of species: Two essays written in 1842 and 1844.
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In a largely overlooked passage from Wallace's essay (p.62), he says of the
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The Works of Charles Darwin: An Annotated Bibliographical Handlist. 2nd edn
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On the Tendency of Varieties to depart indefinitely from the Original Type.
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On The Tendency of Varieties to Depart Indefinitely from the Original Type
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On The Tendency of Varieties to Depart Indefinitely from the Original Type
916: 425: 67:. The article was the first announcement of the Darwin–Wallace theory of 1519: 1460: 966:
Charles Darwin's zoology notes & specimen lists from H.M.S. Beagle.
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The paper, consisting of the letter and papers, was published in the
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The Effects of Cross and Self Fertilisation in the Vegetable Kingdom
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The paper was reprinted and reviewed in several magazines including
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Presentation and analysis of these Wallace and Darwin texts on
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Works originally published in science and technology magazines
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The Formation of Vegetable Mould Through the Action of Worms
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The Different Forms of Flowers on Plants of the Same Species
1058:(for English version of the analysis, click 'à télécharger') 156:, had a paper on the "introduction" of species published in 393:; this condensed version was published in November 1859 as 205:
Lyell and Hooker had decided on a joint publication at the
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of 1844, leaving matters in the hands of Lyell and Hooker.
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The action of this principle is exactly like that of the
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The Variation of Animals and Plants Under Domestication
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of 1844 which Hooker had read in 1847, and a letter to
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of South America, an interest he had developed at the
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The Descent of Man, and Selection in Relation to Sex
766: 1329:The Expression of the Emotions in Man and Animals 1006:"On the Origin of a Theory: Science & Nature" 402: 313:The paper consisted of a communication letter by 1550: 336:Extract from an unpublished Work on Species, by 1584:Works originally published in British magazines 1265:Geological Observations on the Volcanic Islands 1000: 1345:On the Movements and Habits of Climbing Plants 1030: 644: 1249:The Structure and Distribution of Coral Reefs 1078: 848: 613: 1281:On the Tendency of Species to form Varieties 934: 625: 601: 577: 467: 370: 354:Boston, US, dated Down, September 5th, 1857. 282:. At first they stayed in the King's Hotel, 1442:Vestiges of the Natural History of Creation 326:Mr. Charles Darwin and Mr. Alfred Wallace. 308:Zoological Journal of the Linnean Society 3 41:Extract from an unpublished Work on Species 1085: 1071: 420:and extinction almost sure soon to follow. 1449:The Genetical Theory of Natural Selection 946:, London: Michael Joseph, Penguin Group, 873: 862:Zoological Journal of the Linnean Society 809:Charles Darwin: vol. 2 The Power of Place 493: 79:. This was published in November 1859 as 1273:Geological Observations on South America 968:, Cambridge: Cambridge University Press 836:, Cambridge: Cambridge University Press 1551: 1398:The Life and Letters of Charles Darwin 1257:Zoology of the Voyage of H.M.S. Beagle 959: 911: 888: 824: 803: 702: 687: 675: 663: 589: 565: 553: 538: 523: 456: 158:Annals and Magazine of Natural History 51:of 1844. The article also includes an 1066: 294:journal. They next rented a villa in 270:they moved to his sister-in-law's in 638: 479: 89:Events leading up to the publication 1385:The Autobiography of Charles Darwin 929:The Autobiography of Charles Darwin 906:The Autobiography of Charles Darwin 718: 619: 13: 1092: 875:10.1111/j.1096-3642.1858.tb02500.x 435:Mind and Nature: A Necessary Unity 14: 1600: 994: 772: 745: 480:Ball, Philip (12 December 2011). 230:on 1 July 1858, by the Secretary 1233:Extracts from Letters to Henslow 59:, and an introductory letter by 1559:Evolutionary biology literature 1456:History of evolutionary thought 1369:The Power of Movement in Plants 775:"Wallace's Unfinished Business" 739: 712: 696: 681: 669: 657: 607: 595: 1564:Works by Alfred Russel Wallace 1406:More Letters of Charles Darwin 960:Keynes, Richard (ed.) (2000), 583: 571: 559: 547: 532: 517: 473: 461: 450: 403:As an influence on cybernetics 221: 99:Publication of Darwin's theory 1: 796: 488:. Nature News & Comment. 832:, in Darwin, Francis (ed.), 444: 301: 226:The papers were read to the 7: 10: 1605: 748:"For God's Sake, Margaret" 362:By ALFRED RUSSEL WALLACE. 251:William Benjamin Carpenter 152:, a naturalist working in 92: 1423: 1222: 1100: 931:) Retrieved on 2006-12-15 908:) Retrieved on 2007-01-14 811:, London: Jonathan Cape, 653:. pp. 17: 6474–6475. 634:. pp. 17: 6357–6359. 614:Darwin & Wallace 1858 371:Impact of the publication 346:Abstract of a Letter from 228:Linnean Society of London 207:Linnean Society of London 29:Linnean Society of London 1296:Fertilisation of Orchids 1288:On the Origin of Species 1241:The Voyage of the Beagle 1155:On the Origin of Species 719:Wallace, Alfred Russel. 602:Desmond & Moore 1991 578:Desmond & Moore 1991 495:10.1038/nature.2011.9613 468:Desmond & Moore 1991 396:On the Origin of Species 171:Wallace wrote his paper 82:On the Origin of Species 1569:Works by Charles Darwin 1476:things named for Darwin 1031:Freeman, R. B. (1977). 645:Hussey, Arthur (1859). 124:University of Cambridge 120:University of Edinburgh 1415:List of described taxa 1108:Darwin–Wedgwood family 854:Wallace, Alfred Russel 422: 409:evolutionary principle 364: 257:. Amongst the others, 1148:Publication of theory 1143:Development of theory 902:, London: John Murray 626:Boyd, Thomas (1859). 413: 323: 191:Charles Waring Darwin 150:Alfred Russel Wallace 132:Principles of Geology 95:Alfred Russel Wallace 37:Alfred Russel Wallace 1337:Insectivorous Plants 1184:Insectivorous Plants 1039:. Dawson: Folkestone 1011:Smithsonian Magazine 604:, pp. 470, 471. 417:centrifugal governor 315:Joseph Dalton Hooker 310:(20 August): 46–50. 255:William Henry Fitton 144:Joseph Dalton Hooker 61:Joseph Dalton Hooker 53:Abstract of a Letter 1209:Portraits of Darwin 1138:Inception of theory 428:and anthropologist 232:John Joseph Bennett 984:has generic name ( 781:on 29 January 2009 773:Smith, Charles H. 253:and the geologist 1546: 1545: 1312:Natural Selection 1059: 925:, London: Collins 568:, pp. 40, 41 556:, pp. 35, 40 391:Natural Selection 348:C. DARWIN, Esq., 288:Natural Selection 162:Natural Selection 136:natural selection 77:Natural Selection 73:natural selection 25:scientific papers 1596: 1087: 1080: 1073: 1064: 1063: 1057: 1048: 1046: 1044: 1027: 1025: 1023: 1014:. Archived from 989: 983: 979: 977: 969: 962:"Specimen Lists" 956: 926: 903: 885: 884: 882: 877: 845:(published 1909) 844: 843: 841: 821: 805:Browne, E. Janet 791: 790: 788: 786: 770: 764: 763: 761: 759: 746:Brand, Stewart. 743: 737: 736: 734: 732: 727:on 29 April 2007 723:. Archived from 716: 710: 700: 694: 685: 679: 678:, pp. 40–42 673: 667: 661: 655: 654: 642: 636: 635: 623: 617: 611: 605: 599: 593: 592:, pp. 42–47 587: 581: 575: 569: 563: 557: 551: 545: 536: 530: 521: 515: 514: 512: 510: 497: 477: 471: 465: 459: 454: 338:C. DARWIN, Esq., 138:as the cause of 126:. Influenced by 31:on 1 July 1858: 1604: 1603: 1599: 1598: 1597: 1595: 1594: 1593: 1549: 1548: 1547: 1542: 1508:Alternatives to 1466:Darwin Industry 1419: 1218: 1194:Religious views 1118:Josiah Wedgwood 1096: 1091: 1042: 1040: 1021: 1019: 1018:on 5 April 2009 1002:Richard Conniff 997: 992: 981: 980: 971: 970: 954: 936:Desmond, Adrian 913:Darwin, Charles 890:Darwin, Charles 880: 878: 850:Darwin, Charles 839: 837: 830:"Essay of 1844" 826:Darwin, Charles 819: 799: 794: 784: 782: 771: 767: 757: 755: 754:on 15 June 1998 744: 740: 730: 728: 717: 713: 701: 697: 686: 682: 674: 670: 662: 658: 643: 639: 624: 620: 612: 608: 600: 596: 588: 584: 576: 572: 564: 560: 552: 548: 537: 533: 522: 518: 508: 506: 478: 474: 466: 462: 455: 451: 447: 430:Gregory Bateson 405: 373: 357: 356: 352:Prof. ASA GRAY, 343: 342: 333: 332: 331: 304: 224: 116:natural history 101: 93:Main articles: 91: 55:from Darwin to 17: 16:Journal article 12: 11: 5: 1602: 1592: 1591: 1586: 1581: 1579:1858 documents 1576: 1574:Biology papers 1571: 1566: 1561: 1544: 1543: 1541: 1540: 1535: 1530: 1525: 1517: 1516: 1515: 1510: 1505: 1500: 1495: 1490: 1480: 1479: 1478: 1468: 1463: 1458: 1453: 1445: 1438: 1433: 1427: 1425: 1421: 1420: 1418: 1417: 1412: 1411: 1410: 1402: 1392:Correspondence 1389: 1381: 1373: 1365: 1357: 1349: 1341: 1333: 1325: 1317: 1316: 1315: 1300: 1292: 1284: 1277: 1269: 1261: 1253: 1245: 1237: 1228: 1226: 1220: 1219: 1217: 1216: 1206: 1201: 1196: 1191: 1180: 1173:Descent of Man 1169: 1158: 1150: 1145: 1140: 1135: 1130:Voyage on HMS 1127: 1122: 1121: 1120: 1115: 1113:Erasmus Darwin 1104: 1102: 1098: 1097: 1094:Charles Darwin 1090: 1089: 1082: 1075: 1067: 1061: 1060: 1049: 1028: 996: 995:External links 993: 991: 990: 957: 952: 932: 909: 886: 846: 822: 817: 800: 798: 795: 793: 792: 765: 738: 711: 695: 680: 668: 656: 637: 618: 606: 594: 582: 580:, p. 470. 570: 558: 546: 531: 516: 472: 470:, p. 438. 460: 448: 446: 443: 439:systems theory 404: 401: 372: 369: 303: 300: 263:George Bentham 223: 220: 193:, who died of 112:Charles Darwin 105:voyage of the 90: 87: 45:Charles Darwin 15: 9: 6: 4: 3: 2: 1601: 1590: 1587: 1585: 1582: 1580: 1577: 1575: 1572: 1570: 1567: 1565: 1562: 1560: 1557: 1556: 1554: 1539: 1538:Huxley family 1536: 1534: 1533:Darwin Awards 1531: 1529: 1526: 1524: 1523: 1518: 1514: 1511: 1509: 1506: 1504: 1501: 1499: 1496: 1494: 1491: 1489: 1486: 1485: 1484: 1481: 1477: 1474: 1473: 1472: 1471:Commemoration 1469: 1467: 1464: 1462: 1459: 1457: 1454: 1451: 1450: 1446: 1444: 1443: 1439: 1437: 1434: 1432: 1429: 1428: 1426: 1422: 1416: 1413: 1408: 1407: 1403: 1400: 1399: 1395: 1394: 1393: 1390: 1387: 1386: 1382: 1379: 1378: 1374: 1371: 1370: 1366: 1363: 1362: 1358: 1355: 1354: 1350: 1347: 1346: 1342: 1339: 1338: 1334: 1331: 1330: 1326: 1323: 1322: 1318: 1314: 1313: 1309: 1308: 1306: 1305: 1301: 1298: 1297: 1293: 1290: 1289: 1285: 1282: 1278: 1275: 1274: 1270: 1267: 1266: 1262: 1259: 1258: 1254: 1251: 1250: 1246: 1243: 1242: 1238: 1235: 1234: 1230: 1229: 1227: 1225: 1221: 1214: 1210: 1207: 1205: 1202: 1200: 1197: 1195: 1192: 1190: 1189: 1185: 1181: 1179: 1178: 1174: 1170: 1168: 1167: 1163: 1159: 1157: 1156: 1153:Reactions to 1151: 1149: 1146: 1144: 1141: 1139: 1136: 1134: 1133: 1128: 1126: 1123: 1119: 1116: 1114: 1111: 1110: 1109: 1106: 1105: 1103: 1099: 1095: 1088: 1083: 1081: 1076: 1074: 1069: 1068: 1065: 1056: 1055: 1050: 1038: 1034: 1029: 1017: 1013: 1012: 1007: 1004:(June 2008). 1003: 999: 998: 987: 982:|given1= 975: 967: 963: 958: 955: 953:0-7181-3430-3 949: 945: 941: 937: 933: 930: 924: 923: 918: 914: 910: 907: 901: 900: 895: 891: 887: 876: 871: 867: 863: 859: 855: 851: 847: 835: 831: 827: 823: 820: 818:0-7126-6837-3 814: 810: 806: 802: 801: 780: 776: 769: 753: 749: 742: 726: 722: 715: 708: 704: 699: 693: 689: 684: 677: 672: 665: 660: 652: 651:The Zoologist 648: 641: 633: 632:The Zoologist 629: 622: 615: 610: 603: 598: 591: 586: 579: 574: 567: 562: 555: 550: 544: 540: 535: 529: 525: 520: 505: 501: 496: 491: 487: 483: 476: 469: 464: 458: 453: 449: 442: 440: 436: 431: 427: 426:cybernetician 421: 418: 412: 410: 400: 398: 397: 392: 387: 384: 380: 379: 378:The Zoologist 368: 363: 361: 355: 351: 347: 341: 337: 327: 322: 320: 319:Charles Lyell 316: 311: 309: 299: 297: 293: 289: 285: 281: 280:Isle of Wight 277: 274:, then on to 273: 267: 264: 260: 259:Daniel Oliver 256: 252: 248: 243: 241: 237: 233: 229: 219: 216: 212: 208: 203: 201: 196: 195:scarlet fever 192: 187: 183: 178: 174: 169: 165: 163: 159: 155: 151: 147: 145: 141: 137: 133: 129: 128:Charles Lyell 125: 121: 117: 113: 109: 108: 100: 96: 86: 84: 83: 78: 74: 70: 66: 65:Charles Lyell 62: 58: 54: 50: 46: 42: 38: 34: 30: 26: 22: 1528:Darwin Medal 1521: 1447: 1441: 1404: 1396: 1383: 1375: 1367: 1359: 1351: 1343: 1335: 1327: 1319: 1310: 1302: 1294: 1286: 1280: 1271: 1263: 1255: 1247: 1239: 1231: 1187: 1183: 1176: 1172: 1165: 1161: 1154: 1131: 1052: 1041:. Retrieved 1036: 1020:. Retrieved 1016:the original 1009: 965: 943: 940:Moore, James 921: 898: 879:, retrieved 868:(9): 46–62, 865: 861: 838:, retrieved 833: 808: 783:. Retrieved 779:the original 768: 756:. Retrieved 752:the original 741: 729:. Retrieved 725:the original 714: 698: 683: 671: 666:, p. 48 659: 640: 621: 609: 597: 585: 573: 561: 549: 534: 519: 507:. 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Index

scientific papers
Linnean Society of London
Alfred Russel Wallace
Charles Darwin
Asa Gray
Joseph Dalton Hooker
Charles Lyell
evolution
natural selection
On the Origin of Species
Alfred Russel Wallace
Publication of Darwin's theory
voyage of the Beagle
Charles Darwin
natural history
University of Edinburgh
University of Cambridge
Charles Lyell
natural selection
evolution
Joseph Dalton Hooker
Alfred Russel Wallace
Borneo
Ternate
Asa Gray
Charles Waring Darwin
scarlet fever
Linnean Society of London
Asa Gray
Robert Brown

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