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Newt

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1110: 1212: 565: 786: 750: 774: 1273: 1383: 762: 2782: 1234: 1527: 3190: 306: 813: 728:, during which they develop legs, and the gills are absorbed and replaced by air-breathing lungs. Some species, such as the North American newts, also become more brightly colored during this phase. Once fully metamorphosed, they leave the water and live a terrestrial life, when they are known as "efts." Only when the eft reaches adulthood will the North American species return to live in water, rarely venturing back onto the land. Conversely, most European species live their adult lives on land and only visit water to breed. 640: 1947: 136: 1156: 649: 116: 3168: 1796:
were prone to a loss of motor control: After only 22% water weight loss, newts in the aquatic phase lost their ability to remain upright and mobile. However, after adaptation to a terrestrial phase, they could lose 30% before a loss of motor control was recorded. Newts in the terrestrial phase were
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In early stages of development in amphibians, ventilator gas transport and hemoglobin gas transport are independent mechanisms and not yet coupled as they are in adulthood. In juvenile amphibians, there is no cardiovascular response in conditions of hypoxia. When newts are induced into anemia, they
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of the Pacific Northwest used the toxin to poison their enemies. However, the toxins are only dangerous if ingested or otherwise enter the body; for example, through a wound. Newts can safely live in the same ponds or streams as frogs and other amphibians or be kept as pets. The only predators of
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The main breeding season for newts (in the Northern Hemisphere) is in June and July. A single newt female can produce hundreds of eggs. For instance, the warty newt can produce 200–300 eggs (Bradford 2017). After courtship rituals of varying complexity, which take place in ponds or slow-moving
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The nomenclatural status of the nomina of amphibians and reptiles created by Garsault (1764), with a parsimonious solution to an old nomenclatural problem regarding the genus Bufo (Amphibia, Anura), comments on the taxonomy of this genus, and comments on some nomina created by Laurenti
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to a temperature range that exists outside the predator's preferred temperature range. Larvae that are in the metamorphosizing stage tend to prefer warmer temperatures than those in the stage following metamorphosis. Therefore, the larvae in this stage will undergo a much more precise
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Yazawa T, Yamamoto T, Nakayama Y, Hamada S, Abé S. Conversion from mitosis to meiosis: morphology and expression of proliferating cell nuclear antigen (PCNA) and Dmc1 during newt spermatogenesis. Dev Growth Differ. 2000 Dec;42(6):603-11. doi: 10.1046/j.1440-169x.2000.00544.x. PMID
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level of the streams and lakes where they live. Because their skin is permeable to water, they absorb oxygen and other substances they need through their skin. Scientists study the stability of the amphibian population when studying the water quality of a particular body of water.
2994:(A. Capaldo, F. Gay, S. Valiante, M. DeFalco, R. Sciarrillo, M. Maddaloni, V. Laforgia. 2012. Endocrine-disrupting effects of nonylphenol in the newt, Triturus carnifex (Amphibia, Urodela). Comparative Biochemistry and Physiology. 155(2): 352–358. doi: 10.1016/j.cbpc.2011.10.004) 1724:
The heart of newts, like that of most amphibians, consists of two atria and one ventricle. Blood flows from the anterior and posterior caval veins into the right atrium; blood that entered the heart from the left atrium is then expelled out of the ventricle. Newts do not have a
2839:(Giuliano Frangioni, Mario Santoni, Stefano Bianchi, Marco Franchi, Giancarlo Fuzzi, Stefano Marcaccini, Carlo Varlani, Gianfranco Borgioli. 2006. Function of the Hepatic Melanogenesis in the Newt, Triturus Carnifex. Journal of Experimental Zoology. 303A:123-131.) 1784:. Spleen size can increase as the temperature declines for adults – in larvae, there is no dramatic change in spleen size. During hibernation, an increase in liver pigment cells allows for storage of oxygen, as well as other important ions and free radicals. 2893:
Brown, M., Brown, S., Bisceglio, I., and Lemke, S. 1983. Breeding Condition, Temperature, and the Regulation of Salt and Water by Pituitary Hormones in the Red-Spotted Newt, Notophthalmus viridescens. General and Comparative Endocrinology 51,
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Some newt populations in Europe have decreased because of pollution or destruction of their breeding sites and terrestrial habitats, and countries such as the UK have taken steps to halt their declines. In the UK, they are protected under the
596:) with together seven species are the only representatives in North America, while most diversity is found in the Old World: In Europe and the Middle East, the group's likely origin, eight genera with roughly 30 species are found, with the 1737:: when a newt's ventricle is punctured, the heart will divert the blood directly into an ascending aorta via a duct located between the ventricle and the conus arteriosus. Newts begin to regenerate the ventricle by a thickening of the 611:, spending part of the year in the water for reproduction and the rest of the year on land. While most species prefer stagnant water bodies such as ponds, ditches, or flooded meadows for reproduction, some species such as the 2945:(R. Smolinsky, L. Gvozdik. 2009. The ontogenetic shift in thermoregulatory behavior of newt larvae: testing the “enemy-free temperature” hypothesis. Journal of Zoology. 279(2): 180-186. doi: 10.1111/j.1469-7998.2009.00603.x) 2985:(Gay, F., V. Laforgia, I. Caputo, C. Esposito, M. Lepretti, and A. Capaldo, “Chronic Exposure to Cadmium Disrupts the Adrenal Gland Activity of the Newt Triturus carnifex(Amphibia, Urodela),” BioMed Research International.) 391:, especially in composition, the larva being called "water-eft" and the mature form "land-eft" well into the 18th century, but the simplex "eft" as equivalent to "water-eft" has been in use since at least the 17th century. 1797:
found to dehydrate much quicker than newts in the aquatic phase, but conversely, during rehydration, dehydrated terrestrial animals will go through water gain 5x faster than dehydrated newts that are in the aquatic phase.
2976:(Brunelli, E., E. Sperone, M. Maisano, and S. Tripepi. 2009. Morphology and ultrastructure of the gills in two Urodela species: Salamandrina terdigitata and Triturus carnifex. Italian Journal of Zoology. 76(2): 158-164) 262:, alternating between aquatic and terrestrial habitats. Not all aquatic salamanders are considered newts, however. More than 100 known species of newts are found in North America, Europe, North Africa and Asia. Newts 1804:
is released into the circulatory system, which drives the newts into the aquatic environment and reduces the active transport of sodium ions. In contrast to prolactin, which decreases osmotic permeability,
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because of their thin, sensitive skin and evidence of their presence (or absence) can serve as an indicator of the health of the environment. Most species are highly sensitive to subtle changes in the
504:) in historical literature, and "triton" remains in use as common name in some Romance languages, such as Spanish and Romanian, but as well as in Greek, Russian, and Bulgarian. The systematic name 2922:(M. Hadamova, L. Gvozdik. 2011. Seasonal acclimation of preferred body temperatures improves the opportunity for thermoregulation in newts. Physiological and Biochemical Zoology. 84(2):166-174. ) 2054: 797:
Newts lay their eggs on structures such as plants or stones under water. The larvae first develop fore- and later hindlimbs, and are strictly carnivorous. After metamorphosis, juveniles known as
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Minoru Uchiyama, Norifumi Konno, Hormonal regulation of ion and water transport in anuran amphibians, General and Comparative Endocrinology, Volume 147, Issue 1, 15 May 2006, Pages 54-61,
631:) have even adapted to life in cold, oxygen-rich mountain streams. During their terrestrial phase, newts live in humid habitats with abundant cover such as logs, rocks, or earth holes. 1844:
Reproductive females of the Italian crested newt were shown to regulate their body temperature more precisely and prefer higher temperatures than non-reproductive females and males.
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increases the permeability and is secreted during the summer months. Arginine vasotocin not only increases cutaneous water permeability, but promotes increased cutaneous blood flow.
2723:"An Extremely Peramorphic Newt (Urodela: Salamandridae: Pleurodelini) from the Latest Oligocene of Germany, and a New Phylogenetic Analysis of Extant and Extinct Salamandrids" 2505:"Amphibians - body, used, water, process, Earth, life, characteristics, form, animals, oxygen, air, plant, change, History, Characteristics, Life cycle, Three major groupings" 2884:
Lodi, G., Biciotti, M., and Viotto, B. 1981. Cutaneous Osmoregulation in Triturus cristatus carnifex (Laur.) (Urodela). General and Comparative Endocrinology 46: 452-457.
868:, some having developed a resistance to the toxin. Most newts can be safely handled, provided the toxins they produce are not ingested or allowed to come in contact with 1829:, which means they will acclimate to their surrounding environmental temperatures. When there is a large range of environmental temperatures, newts are insensitive to a 2875:
P. Walters, L. Greenwald. 1977. Physiological adaptations of aquatic newts (Notophtalmus viridescens) to a terrestrial environment. Physiological Zoology. 50(2): 88-98.
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but are distinguished by their feathery external gills, hatch out in about three weeks. After hatching, they eat algae, small invertebrates, or other amphibian larvae.
1830: 1773: 705:, which is taken up by the female. Fertilized eggs are laid singly and are usually attached to aquatic plants. This distinguishes them from the free-floating eggs of 1734: 680:, including semipermeable glandular skin, four equal-sized limbs, and a distinct tail. The newt's skin, however, is not as smooth as that of other salamanders. The 2437: 2797:(Miyachi, Y. 2011. The Unusual Circulation of the Newt Heart after Ventricular Injury and Its Implications for Regeneration. Anatomy Research International:1-7). 2848:(Frangioni, G., G. Borgioli. 1996. Variations in blood pressure of newts according to their respiratory conditions. Italian Journal of Zoology. 63:3, 201-206). 1109: 454: 3099: 1825:
cope with the changing temperatures existing in their environments. This regulation is most often achieved through behavioral thermoregulation. They are
2931:(V. Marek, L. Gvozdik. 2012. The insensitivity of thermal preferences to various thermal gradient profiles in newts. Journal of Ethology. 30(1): 35-41) 1826: 3215: 2821:(Casale. G.P, Khairallah. E. A., Grasso. J. A. 1980. An Analysis of Hemoglobin Synthesis in Erythropoietic Cells. Developmental Biology 80:107-119. ) 1753:, around two weeks after anemia is induced, the newts produced a mass of cells that helps to revitalize the already circulating red blood cell mass. 1742: 2954:(L. Gvozdik. 1995. Does reproduction influence temperature preference in newts. Canadian Journal of Zoology. 83(8): 1038-1044. doi: 10.1139/z05-096) 1769:. Lung breathing is mainly used when there is a lack of oxygen in the water, or at high activity such as during courtship, breeding, or feeding. 2482: 604:) extending to northernmost Africa. Eastern Asia, from Eastern India over Indochina to Japan, is home to five genera with more than 40 species. 2019: 2654:"A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of advanced frogs, salamanders, and caecilians" 688:, reproduce rapidly, and differentiate again to create a new limb or organ. One hypothesis is that the undifferentiated cells are related to 2830:(Eddy, F. B., P. McDonald. 1978. Aquatic respiration of The Crested Newt Triturus Cristatus. Comparative Biochemical Physiology. 59: 85-88) 423: 2857:(Frangioni, G., G Borgioli. 1997. Blood and splenic respiratory compensation in larval newts. Italian Journal of Zoology. 64:3, 221-226) 2525: 2419: 2700: 2235: 1994:. This affects especially the aquatic breeding sites they depend on, but also their land habitats. Several species, such as the 2358:
Constable's miscellany of original and selected publications in the various departments of literature, science, & the arts
2466: 713:, which are laid in clumps or in strings. Plant leaves are usually folded over and attached to the eggs to protect them. The 1910:, a heavy metal released into the environment from industrial and consumer waste, has been shown to be detrimental to the 3062: 2236:"Phylogeny and biogeography of the family Salamandridae (Amphibia: Caudata) inferred from complete mitochondrial genomes" 1039:
The term "newt" has traditionally been seen as an exclusively functional term for salamanders living in water, and not a
369:("an eft" → "a n'eft" → ...) by the early 15th century. The form "newt" appears to have arisen as a dialectal variant of 2456: 2433: 1875: 856: 2394:. Deutscher Taschenbuch Verlag, Munich 1997 (revised edition of Akademie Verlag, Munich 1989 and 1993), p. 265, s.v. 2031: 2409:"distaff", so that both animals (lizard and badger) were parallelized due to their common, spindle-shaped bodies. 3103: 1982:) in Europe, are still relatively common, populations of newts throughout their distribution range suffer from 1051:. Other, more distantly related salamander families also contain fully or in part aquatic species, such as the 274:-like bodies and return to the water every year to breed, otherwise living in humid, cover-rich land habitats. 2356:: "the water-eft, or newt, is only the larva of the land-eft, as tadpoles are of frogs". Archibald Constable, 1066:
Classification of all genera of the Pleurodelinae subfamily after Pyron and Weins, revised by Mikko Haaramo.
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Larvae, with their great number of lamellae in their gills, are more susceptible to pollutants than adults.
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are not listed, the sale of either species is prohibited under the Wildlife and Countryside Act, 1981.
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layer that protrudes to allow the new vessels to form, and conclude with a regeneration of the entire
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cells, since chemicals that produce tumors in other animals will produce additional limbs in newts.
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analyses have however shown that species in the Salamandridae traditionally called newts do form a
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even at a concentrations below Italian and European thresholds, by disrupting the activity of the
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In Europe, nine newts are listed as "strictly protected fauna species" under appendix II of the
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without a licence, or to cause them harm or death, or to disturb their habitat in any way. The
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Zhang, Peng; Papenfuss, Theodore J.; Wake, Marvalee H.; Qu, Lianghu; Wake, David B. (2008).
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are some commonly seen newts in the pet trade. Some newts rarely seen in the pet trade are
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The remaining European species are listed as "protected fauna species" under appendix III.
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and the Habitat Regulations Act 1994. It is illegal to catch, possess, or handle
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The Natural History and Antiquities of Selborne, in the County of Southampton
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are regarded as separate genera by some authors, but this is not unanimous.
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species in the family, roughly 100, which are classified in sixteen genera:
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categorises the species as ‘lower risk’ Although the other UK species, the
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model for investigating the mechanism(s) controlling the transition from
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Convention on the Conservation of European Wildlife and Natural Habitats
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Phylogenetic analyses estimated the origin of the newt subfamily in the
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The Gentleman's Magazine, and Historical Chronicle, Volume 47 (1777),
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Collins, J. T.; Conant, R.; Stebbins, R. C.; Peterson, R. T. (1999).
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Newts, as with salamanders in general and other amphibians, serve as
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In the Italian crested newt, it was shown that during winter months,
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Newts are found in North America, Europe, North Africa and Asia. The
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in their skin secretions as a defence mechanism against predators.
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Newts share many of the characteristics of their salamander kin,
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is an example of a newt species that has gone extinct recently.
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To escape predators, newt larvae have been found to shift their
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An Essay Towards a Real Character, and a Philosophical Language
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thermoregulation process than those in the intermediate stage.
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in 1830 (the name meaning "having prominent ribs," formed from
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for a type of spotted newt, now used for species of the genus
3079:"Science & Nature - Wildfacts - Smooth newt, common newt" 2454: 1040: 833: 689: 665:(left) and as an adult during aquatic breeding season (right) 469: 267: 534:
in 1815. Tschudi's Pleurodelinae is based on the type genus
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newts of western North America are particularly toxic. The
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on the ventricle, due to circulation that is found in the
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through three distinct developmental life stages: aquatic
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have also been referred to the Pleurodelinae, including:
2603:"BBC Nature - Great crested newt videos, news and facts" 1749:
are able to respire without the need of blood cells. In
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Molecular Biology Program - Bioscience Graduate Studies
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common in leakage from sewers, there was a decrease in
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emerge from the water and live a terrestrial lifestyle.
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This article is about the animal. For other uses, see
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During the subsequent few months, the larvae undergo
2714: 2597: 2595: 2233: 2229: 2227: 2225: 2720: 2528:. Lnr.cambridge.gov.uk. 2005-07-25. Archived from 2379:An etmological dictionary of the Scottish language 1821:, describes the major mechanisms of how newts, as 2647: 2645: 2643: 2592: 2222: 3202: 2526:"Cambridge City Council: Local Nature Reserves" 2458:Peterson First Guide to Reptiles and Amphibians 1817:Thermoregulation, in combination with seasonal 684:at the site of an injury have the ability to un 318:, is the type genus of subfamily Pleurodelinae. 2703:. New York: American Museum of Natural History 2640: 1901: 493:, like the English term of unknown etymology. 394:Dialectal English and Scots also has the word 250:. The terrestrial juvenile phase is called an 3125:"Annexe II: Strictly protected fauna species" 2962: 2960: 2721:Marjanović, David; Witzmann, Florian (2015). 347:; this word was transformed irregularly into 3025:10.2305/IUCN.UK.2004.RLTS.T59445A11942589.en 2549: 2398:. Pfeifer gives the second element as germ. 559: 475: 615:can also occur in slow-flowing rivers. The 551: 545: 444: 3004:Yang Datong; Michael Wai Neng Lau (2004). 2957: 2651: 2621: 2619: 508:was introduced alongside Pleurodelinae by 114: 26:"Eft" redirects here. For other uses, see 3023: 2764: 2746: 2575: 1714: 16:Salamander in the subfamily Pleurodelinae 3216:Extant Late Cretaceous first appearances 2941: 2939: 2937: 2627:"Caudata Culture Articles - Newt Toxins" 2448: 1945: 811: 563: 304: 2918: 2916: 2793: 2791: 2616: 2392:Etymologisches Wörterbuch des Deutschen 512:in 1838, based on the type genus named 3203: 2878: 2842: 2817: 2815: 2805: 2803: 1941: 3145:"Annexe III: Protected fauna species" 2934: 2925: 2887: 2860: 2833: 2698: 2661:Molecular Phylogenetics and Evolution 2243:Molecular Phylogenetics and Evolution 254:. Unlike other members of the family 2997: 2970: 2913: 2869: 2851: 2824: 2788: 2489:. University of Utah. Archived from 2418:Henry George Liddell, Robert Scott, 1938:and important for stress response. 3011:IUCN Red List of Threatened Species 2988: 2979: 2948: 2897: 2812: 2800: 2130: 2018:) are considered threatened by the 1958:Although some species, such as the 1812: 701:streams, the male newt transfers a 13: 2507:. Scienceclarified.com. 2009-10-13 2368:of Hay, (the water-newt, or eft)". 1847: 779:Larva shortly before metamorphosis 671: 500:(viz., named for the mythological 408:in Scots) used for both newts and 338:(of unknown origin), resulting in 14: 3227: 3160: 2652:Pyron, R.A.; Weins, J.J. (2011). 2550:Brockes, J; Kumar, A (Jan 2005). 2032:Wildlife and Countryside Act 1981 1852:The newt is regarded as an ideal 1787: 1776:is the ability to release stored 855:to kill an adult human, and some 443:is ultimately cognate with Greek 3188: 3166: 2780: 2304:"Online etymological dictionary" 2301: 1648: 1525: 1381: 1271: 1232: 1210: 1154: 1108: 784: 772: 760: 748: 647: 638: 134: 46: 3137: 3117: 3092: 3053: 3032: 2543: 2518: 2497: 2475: 1780:when needed, for example under 828:, are known for their toxicity. 2425: 2412: 2384: 2371: 2331: 2318: 2291: 1882:auxiliary protein involved in 1756: 1719: 875: 695: 572:lives in small streams in the 1: 3006:"Hypselotriton wolterstorffi" 2461:. Houghton Mifflin Harcourt. 2431:Alain Dubois and Roger Bour, 2216: 468:. Ancient Greek had the name 3042:. Fresc.usgs.gov. 2013-01-30 2748:10.1371/journal.pone.0137068 2445:, Zootaxa 2447 (2010), 1–52. 300: 7: 3060:UK Biodiversity Action Plan 2673:10.1016/j.ympev.2011.06.012 2255:10.1016/j.ympev.2008.08.020 2173:Chuxiong fire-bellied newts 1934:, hormones produced by the 1902:Susceptibility to pollution 1792:In experiments, dehydrated 807: 10: 3232: 2147: 1972:Notophthalmus viridescens) 851:produces more than enough 470: 367:provection (juncture loss) 25: 18: 3065:October 24, 2007, at the 2577:10.1016/j.cub.2004.12.049 2298:Oxford English Dictionary 2169:Japanese fire belly newts 1540: 1518: 1511: 1494: 1487: 1470: 1463: 1445: 1438: 1421: 1414: 1374: 1357: 1350: 1324: 1307: 1300: 1293: 1286: 1264: 1257: 1225: 1203: 1196: 1188: 1147: 1130: 1123: 1101: 1094: 1086: 1074: 560:Distribution and habitats 552: 546: 445: 218: 213: 131:Scientific classification 129: 122: 113: 37: 3102:. ARKive. Archived from 2390:Wolfgang Pfeifer (ed.), 2157:Chinese fire belly newts 1974:in North America or the 1831:thermal gradient profile 1774:compensatory respiration 880:Newts form one of three 496:Newts are also known as 383:iv.1). The regular form 277:Newts are threatened by 2420:A Greek-English Lexicon 2195:red-tailed knobby newts 2078:Euproctus platycephalus 1735:ventricular penetration 872:or breaks in the skin. 755:Embryo in jelly capsule 625:European mountain newts 540:(ribbed newt) named by 361: 355: 349: 343: 334: 328: 326:name of the animal was 2203:Kaiser's spotted newts 2092:Lissotriton montandoni 1955: 1954:is considered extinct. 1894:protein, a marker for 1715:Anatomy and physiology 829: 717:, which resemble fish 577: 476: 458:). Latin had the name 319: 44:Late Cretaceous–Recent 3018:: e.T59445A11942589. 2004:Kaiser's spotted newt 1949: 1896:genetic recombination 1761:Adult crested newts ( 896:. They comprise most 815: 736:northern crested newt 567: 308: 21:Newt (disambiguation) 2699:Frost, D.R. (2016). 2360:, Volume 45 (1829), 2211:yellow-spotted newts 2187:Spanish ribbed newts 2085:Lissotriton italicus 1980:Lissotriton vulgaris 1912:Italian crested newt 1880:DNA polymerase delta 791:Terrestrial juvenile 617:European brook newts 520:in 1768. Laurenti's 2866:(Barni et al. 1999) 2739:2015PLoSO..1037068M 2568:2005CBio...15..R42B 2366:salamandra aquatica 2199:rough-skinned newts 2153:Chinese warty newts 2113:Triturus dobrogicus 2036:great crested newts 2012:Montseny brook newt 2000:Pleurodeles poireti 1942:Conservation status 1924:endocrine disruptor 1871:Cynopa pyrrhogaster 1868:. In the male newt 832:Many newts produce 734:Development in the 613:Danube crested newt 570:Pyrenean brook newt 530:("Triton-tail") by 451:Proto-Indo-European 412:, from Old English 316:Iberian ribbed newt 2483:"Shannon Odelberg" 2440:2014-01-09 at the 2404:Middle High German 2193:genes exist), and 2120:Triturus karelinii 2106:Triturus cristatus 2071:Euproctus montanus 2016:Calotriton arnoldi 1996:Edough ribbed newt 1960:rough-skinned newt 1956: 1663:fossil salamanders 1049:monophyletic group 842:rough-skinned newt 830: 740:Triturus cristatus 578: 482:Middle High German 320: 3081:. BBC. 2012-04-27 2468:978-0-395-97195-6 2402:, relating it to 2348:. Gilbert White, 2099:Triturus carnifex 2008:Neurergus kaiseri 1964:Taricha granulosa 1707:Palaeopleurodeles 1645: 1644: 1636: 1635: 1627: 1626: 1618: 1617: 1609: 1608: 1600: 1599: 1591: 1590: 1582: 1581: 1573: 1572: 1564: 1563: 1555: 1554: 1403: 1402: 1394: 1393: 1339: 1338: 1245: 1244: 1176: 1175: 1167: 1166: 849:Pacific Northwest 845:Taricha granulosa 824:), including the 439:, both "lizard;" 233: 232: 228:genera, see text 3223: 3193: 3192: 3191: 3184: 3171:Data related to 3170: 3155: 3154: 3152: 3151: 3141: 3135: 3134: 3132: 3131: 3121: 3115: 3114: 3112: 3111: 3096: 3090: 3089: 3087: 3086: 3075: 3069: 3057: 3051: 3050: 3048: 3047: 3036: 3030: 3029: 3027: 3001: 2995: 2992: 2986: 2983: 2977: 2974: 2968: 2964: 2955: 2952: 2946: 2943: 2932: 2929: 2923: 2920: 2911: 2901: 2895: 2891: 2885: 2882: 2876: 2873: 2867: 2864: 2858: 2855: 2849: 2846: 2840: 2837: 2831: 2828: 2822: 2819: 2810: 2807: 2798: 2795: 2786: 2785: 2784: 2778: 2768: 2750: 2718: 2712: 2711: 2709: 2708: 2696: 2685: 2684: 2658: 2649: 2638: 2637: 2635: 2634: 2623: 2614: 2613: 2611: 2610: 2599: 2590: 2589: 2579: 2547: 2541: 2540: 2538: 2537: 2522: 2516: 2515: 2513: 2512: 2501: 2495: 2494: 2479: 2473: 2472: 2452: 2446: 2429: 2423: 2416: 2410: 2388: 2382: 2375: 2369: 2335: 2329: 2322: 2316: 2315: 2313: 2311: 2306:. Douglas Harper 2295: 2289: 2288: 2286: 2285: 2279: 2273:. Archived from 2240: 2231: 2165:paddletail newts 2131:As bioindicators 2064:Calotriton asper 2024:Yunnan lake newt 1952:Yunnan lake newt 1827:thermoconformers 1813:Thermoregulation 1763:Triturus cristus 1731:conus arteriosus 1529: 1514: 1513: 1490: 1489: 1466: 1465: 1441: 1440: 1417: 1416: 1385: 1353: 1352: 1303: 1302: 1296: 1295: 1289: 1288: 1275: 1260: 1259: 1236: 1214: 1199: 1198: 1191: 1190: 1158: 1126: 1125: 1112: 1097: 1096: 1089: 1088: 1077: 1076: 1070: 1069: 1053:mole salamanders 870:mucous membranes 857:Native Americans 788: 776: 764: 752: 651: 642: 555: 554: 549: 548: 479: 473: 472: 448: 447: 364: 358: 352: 346: 337: 331: 314:, including the 291:Yunnan lake newt 139: 138: 118: 108: 45: 41:Temporal range: 35: 34: 3231: 3230: 3226: 3225: 3224: 3222: 3221: 3220: 3201: 3200: 3199: 3189: 3187: 3179: 3163: 3158: 3149: 3147: 3143: 3142: 3138: 3129: 3127: 3123: 3122: 3118: 3109: 3107: 3098: 3097: 3093: 3084: 3082: 3077: 3076: 3072: 3067:Wayback Machine 3058: 3054: 3045: 3043: 3038: 3037: 3033: 3002: 2998: 2993: 2989: 2984: 2980: 2975: 2971: 2965: 2958: 2953: 2949: 2944: 2935: 2930: 2926: 2921: 2914: 2902: 2898: 2892: 2888: 2883: 2879: 2874: 2870: 2865: 2861: 2856: 2852: 2847: 2843: 2838: 2834: 2829: 2825: 2820: 2813: 2808: 2801: 2796: 2789: 2779: 2733:(9): e0137068. 2719: 2715: 2706: 2704: 2697: 2688: 2656: 2650: 2641: 2632: 2630: 2625: 2624: 2617: 2608: 2606: 2601: 2600: 2593: 2556:Current Biology 2548: 2544: 2535: 2533: 2524: 2523: 2519: 2510: 2508: 2503: 2502: 2498: 2481: 2480: 2476: 2469: 2453: 2449: 2442:Wayback Machine 2430: 2426: 2417: 2413: 2389: 2385: 2377:John Jamieson, 2376: 2372: 2336: 2332: 2323: 2319: 2309: 2307: 2296: 2292: 2283: 2281: 2277: 2238: 2232: 2223: 2219: 2150: 2133: 2125: 1944: 1904: 1884:DNA replication 1866:spermatogenesis 1850: 1848:Spermatogenesis 1815: 1790: 1759: 1743:myocardial wall 1727:coronary artery 1722: 1717: 1712: 1655:Late Cretaceous 1651: 1646: 1637: 1628: 1619: 1610: 1601: 1592: 1583: 1574: 1565: 1556: 1404: 1395: 1340: 1246: 1177: 1168: 1034:Liangshantriton 1027: 1022:Liangshantriton 894:Salamandrininae 878: 826:California newt 810: 805: 804: 803: 802: 794: 793: 792: 789: 781: 780: 777: 769: 768: 765: 757: 756: 753: 744: 743: 698: 674: 672:Characteristics 669: 668: 667: 666: 661:as terrestrial 654: 653: 652: 644: 643: 562: 524:was renamed to 303: 133: 109: 107: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 43: 42: 39: 31: 24: 17: 12: 11: 5: 3229: 3219: 3218: 3213: 3198: 3197: 3177: 3176: 3175:at Wikispecies 3162: 3161:External links 3159: 3157: 3156: 3136: 3116: 3091: 3070: 3052: 3031: 2996: 2987: 2978: 2969: 2956: 2947: 2933: 2924: 2912: 2896: 2886: 2877: 2868: 2859: 2850: 2841: 2832: 2823: 2811: 2799: 2787: 2713: 2686: 2667:(2): 543–853. 2639: 2615: 2591: 2562:(2): R42–R44. 2542: 2517: 2496: 2493:on 2006-09-02. 2474: 2467: 2447: 2424: 2411: 2383: 2370: 2330: 2317: 2290: 2249:(2): 586–597. 2220: 2218: 2215: 2149: 2146: 2132: 2129: 2124: 2123: 2116: 2109: 2102: 2095: 2088: 2081: 2074: 2067: 2059: 1943: 1940: 1928:corticosterone 1903: 1900: 1849: 1846: 1814: 1811: 1789: 1788:Osmoregulation 1786: 1758: 1755: 1721: 1718: 1716: 1713: 1711: 1710: 1703: 1696: 1689: 1686:Carpathotriton 1682: 1675: 1667: 1650: 1647: 1643: 1642: 1639: 1638: 1634: 1633: 1630: 1629: 1625: 1624: 1621: 1620: 1616: 1615: 1612: 1611: 1607: 1606: 1603: 1602: 1598: 1597: 1594: 1593: 1589: 1588: 1585: 1584: 1580: 1579: 1576: 1575: 1571: 1570: 1567: 1566: 1562: 1561: 1558: 1557: 1553: 1552: 1549: 1548: 1544:Paramesotriton 1539: 1536: 1535: 1532: 1531: 1517: 1512: 1510: 1507: 1506: 1503: 1502: 1493: 1488: 1486: 1483: 1482: 1479: 1478: 1469: 1464: 1462: 1458: 1457: 1454: 1453: 1444: 1439: 1437: 1434: 1433: 1430: 1429: 1420: 1415: 1413: 1410: 1409: 1406: 1405: 1401: 1400: 1397: 1396: 1392: 1391: 1388: 1387: 1373: 1370: 1369: 1366: 1365: 1356: 1351: 1349: 1346: 1345: 1342: 1341: 1337: 1336: 1333: 1332: 1323: 1320: 1319: 1316: 1315: 1306: 1301: 1299: 1294: 1292: 1287: 1285: 1282: 1281: 1278: 1277: 1263: 1258: 1256: 1252: 1251: 1248: 1247: 1243: 1242: 1239: 1238: 1224: 1221: 1220: 1217: 1216: 1202: 1197: 1195: 1189: 1187: 1183: 1182: 1179: 1178: 1174: 1173: 1170: 1169: 1165: 1164: 1161: 1160: 1146: 1143: 1142: 1139: 1138: 1129: 1124: 1122: 1119: 1118: 1115: 1114: 1100: 1095: 1093: 1087: 1085: 1075: 1073: 1068: 1026: 1025: 1013: 1006: 999: 992: 989:Paramesotriton 985: 978: 971: 964: 957: 950: 943: 936: 929: 922: 910: 902: 884:in the family 877: 874: 809: 806: 796: 795: 790: 783: 782: 778: 771: 770: 766: 759: 758: 754: 747: 746: 745: 733: 732: 731: 730: 697: 694: 673: 670: 656: 655: 646: 645: 637: 636: 635: 634: 633: 561: 558: 449:"snake," from 420:Proto-Germanic 340:Middle English 302: 299: 231: 230: 216: 215: 211: 210: 205: 201: 200: 195: 191: 190: 185: 181: 180: 175: 171: 170: 165: 161: 160: 155: 151: 150: 145: 141: 140: 127: 126: 120: 119: 111: 110: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 40: 15: 9: 6: 4: 3: 2: 3228: 3217: 3214: 3212: 3209: 3208: 3206: 3196: 3186: 3185: 3182: 3174: 3173:Pleurodelinae 3169: 3165: 3164: 3146: 3140: 3126: 3120: 3106:on 2008-12-01 3105: 3101: 3095: 3080: 3074: 3068: 3064: 3061: 3056: 3041: 3035: 3026: 3021: 3017: 3013: 3012: 3007: 3000: 2991: 2982: 2973: 2963: 2961: 2951: 2942: 2940: 2938: 2928: 2919: 2917: 2910: 2906: 2900: 2890: 2881: 2872: 2863: 2854: 2845: 2836: 2827: 2818: 2816: 2806: 2804: 2794: 2792: 2783: 2776: 2772: 2767: 2762: 2758: 2754: 2749: 2744: 2740: 2736: 2732: 2728: 2724: 2717: 2702: 2695: 2693: 2691: 2682: 2678: 2674: 2670: 2666: 2662: 2655: 2648: 2646: 2644: 2629:. Caudata.org 2628: 2622: 2620: 2604: 2598: 2596: 2587: 2583: 2578: 2573: 2569: 2565: 2561: 2557: 2553: 2546: 2532:on 2013-07-23 2531: 2527: 2521: 2506: 2500: 2492: 2488: 2484: 2478: 2470: 2464: 2460: 2459: 2451: 2444: 2443: 2439: 2436: 2428: 2421: 2415: 2408: 2405: 2401: 2397: 2393: 2387: 2380: 2374: 2367: 2363: 2359: 2355: 2351: 2347: 2343: 2339: 2334: 2327: 2321: 2305: 2299: 2294: 2280:on 2017-02-02 2276: 2272: 2268: 2264: 2260: 2256: 2252: 2248: 2244: 2237: 2230: 2228: 2226: 2221: 2214: 2212: 2208: 2204: 2200: 2196: 2192: 2188: 2184: 2183:emperor newts 2180: 2179: 2174: 2170: 2166: 2162: 2161:eastern newts 2158: 2154: 2145: 2142: 2138: 2137:bioindicators 2128: 2122: 2121: 2117: 2115: 2114: 2110: 2108: 2107: 2103: 2101: 2100: 2096: 2094: 2093: 2089: 2087: 2086: 2082: 2080: 2079: 2075: 2073: 2072: 2068: 2066: 2065: 2061: 2060: 2058: 2056: 2051: 2049: 2045: 2041: 2040:IUCN Red List 2037: 2033: 2027: 2025: 2021: 2017: 2013: 2009: 2005: 2001: 1997: 1993: 1989: 1988:fragmentation 1985: 1981: 1977: 1973: 1969: 1965: 1961: 1953: 1948: 1939: 1937: 1936:adrenal gland 1933: 1929: 1925: 1921: 1917: 1916:adrenal gland 1913: 1909: 1899: 1897: 1893: 1890:, as well as 1889: 1885: 1881: 1877: 1873: 1872: 1867: 1863: 1859: 1855: 1845: 1842: 1839: 1834: 1832: 1828: 1824: 1820: 1810: 1808: 1803: 1798: 1795: 1794:eastern newts 1785: 1783: 1779: 1775: 1770: 1768: 1767:buccal cavity 1764: 1754: 1752: 1746: 1744: 1740: 1736: 1732: 1728: 1709: 1708: 1704: 1702: 1701: 1697: 1695: 1694: 1690: 1688: 1687: 1683: 1681: 1680: 1676: 1674: 1673: 1672:Archaeotriton 1669: 1668: 1666: 1664: 1660: 1656: 1649:Phylogenetics 1641: 1640: 1632: 1631: 1623: 1622: 1614: 1613: 1605: 1604: 1596: 1595: 1587: 1586: 1578: 1577: 1569: 1568: 1560: 1559: 1551: 1550: 1547: 1546: 1545: 1538: 1537: 1534: 1533: 1530: 1528: 1524: 1523: 1516: 1515: 1509: 1508: 1505: 1504: 1501: 1500: 1499: 1492: 1491: 1485: 1484: 1481: 1480: 1477: 1476: 1475: 1468: 1467: 1460: 1459: 1456: 1455: 1452: 1451: 1450: 1443: 1442: 1436: 1435: 1432: 1431: 1428: 1427: 1426: 1419: 1418: 1412: 1411: 1408: 1407: 1399: 1398: 1390: 1389: 1386: 1384: 1380: 1379: 1372: 1371: 1368: 1367: 1364: 1363: 1362: 1355: 1354: 1348: 1347: 1344: 1343: 1335: 1334: 1331: 1330: 1329: 1322: 1321: 1318: 1317: 1314: 1313: 1312: 1305: 1304: 1298: 1297: 1291: 1290: 1284: 1283: 1280: 1279: 1276: 1274: 1270: 1269: 1262: 1261: 1254: 1253: 1250: 1249: 1241: 1240: 1237: 1235: 1231: 1230: 1223: 1222: 1219: 1218: 1215: 1213: 1209: 1208: 1207:Notophthalmus 1201: 1200: 1193: 1192: 1185: 1184: 1181: 1180: 1172: 1171: 1163: 1162: 1159: 1157: 1153: 1152: 1145: 1144: 1141: 1140: 1137: 1136: 1135: 1128: 1127: 1121: 1120: 1117: 1116: 1113: 1111: 1107: 1106: 1099: 1098: 1092:Pleurodelini 1091: 1090: 1084: 1083: 1082:Pleurodelinae 1079: 1078: 1072: 1071: 1067: 1064: 1062: 1058: 1054: 1050: 1046: 1042: 1037: 1035: 1031: 1030:Hypselotriton 1023: 1019: 1018: 1014: 1012: 1011: 1007: 1005: 1004: 1000: 998: 997: 993: 991: 990: 986: 984: 983: 979: 977: 976: 972: 970: 969: 968:Notophthalmus 965: 963: 962: 958: 956: 955: 951: 949: 948: 944: 942: 941: 937: 935: 934: 930: 928: 927: 923: 920: 919:Hypselotriton 916: 915: 911: 909: 908: 904: 903: 901: 899: 895: 891: 890:Salamandrinae 887: 886:Salamandridae 883: 873: 871: 867: 866:garter snakes 863: 858: 854: 850: 846: 843: 839: 835: 827: 823: 819: 818:Pacific newts 814: 800: 787: 775: 763: 751: 741: 737: 729: 727: 726:metamorphosis 722: 720: 716: 712: 708: 704: 703:spermatophore 693: 691: 687: 686:differentiate 683: 679: 664: 660: 650: 641: 632: 630: 626: 622: 618: 614: 610: 605: 603: 599: 595: 594:Notophthalmus 591: 590:Eastern newts 587: 583: 582:Pacific newts 575: 571: 566: 557: 543: 539: 538: 533: 529: 528: 523: 519: 515: 511: 507: 503: 499: 494: 492: 491: 486: 483: 478: 467: 466: 461: 457: 456: 452: 442: 438: 437: 432: 431: 426: 425: 421: 417: 416: 411: 407: 403: 399: 398: 392: 390: 386: 382: 381: 376: 375:Staffordshire 372: 368: 363: 357: 351: 345: 341: 336: 330: 325: 317: 313: 312: 307: 298: 296: 293:, has become 292: 288: 284: 283:fragmentation 280: 275: 273: 269: 265: 261: 257: 256:Salamandridae 253: 249: 248:Pleurodelinae 246: 242: 238: 229: 227: 223: 217: 212: 209: 208:Pleurodelinae 206: 203: 202: 199: 198:Salamandridae 196: 193: 192: 189: 186: 183: 182: 179: 176: 173: 172: 169: 166: 163: 162: 159: 156: 153: 152: 149: 146: 143: 142: 137: 132: 128: 125: 121: 117: 112: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 36: 33: 29: 22: 3148:. Retrieved 3139: 3128:. Retrieved 3119: 3108:. Retrieved 3104:the original 3094: 3083:. Retrieved 3073: 3055: 3044:. Retrieved 3034: 3015: 3009: 2999: 2990: 2981: 2972: 2950: 2927: 2899: 2889: 2880: 2871: 2862: 2853: 2844: 2835: 2826: 2730: 2726: 2716: 2705:. Retrieved 2664: 2660: 2631:. Retrieved 2607:. Retrieved 2559: 2555: 2545: 2534:. Retrieved 2530:the original 2520: 2509:. Retrieved 2499: 2491:the original 2486: 2477: 2457: 2450: 2432: 2427: 2414: 2406: 2399: 2391: 2386: 2378: 2373: 2365: 2357: 2349: 2341: 2338:John Wilkins 2333: 2320: 2308:. Retrieved 2293: 2282:. Retrieved 2275:the original 2246: 2242: 2207:banded newts 2176: 2151: 2134: 2126: 2118: 2111: 2104: 2097: 2090: 2083: 2076: 2069: 2062: 2052: 2048:palmate newt 2028: 2015: 2007: 1999: 1984:habitat loss 1979: 1971: 1968:Eastern newt 1963: 1957: 1905: 1869: 1851: 1843: 1838:microhabitat 1835: 1816: 1799: 1791: 1778:erythrocytes 1771: 1762: 1760: 1750: 1747: 1723: 1705: 1698: 1691: 1684: 1679:Brachycormus 1677: 1670: 1652: 1542: 1541: 1520: 1519: 1496: 1495: 1472: 1471: 1449:Ichthyosaura 1447: 1446: 1423: 1422: 1376: 1375: 1359: 1358: 1328:Ommatotriton 1326: 1325: 1309: 1308: 1266: 1265: 1227: 1226: 1205: 1204: 1151:Tylototriton 1149: 1148: 1134:Echinotriton 1132: 1131: 1103: 1102: 1080: 1065: 1045:Phylogenetic 1038: 1033: 1029: 1028: 1021: 1017:Tylototriton 1015: 1008: 1001: 994: 987: 980: 975:Ommatotriton 973: 966: 959: 952: 945: 940:Ichthyosaura 938: 931: 926:Echinotriton 924: 918: 912: 905: 879: 861: 853:tetrodotoxin 844: 837: 831: 821: 798: 739: 723: 699: 675: 662: 659:eastern newt 628: 620: 606: 601: 598:ribbed newts 593: 585: 579: 535: 525: 521: 513: 505: 497: 495: 488: 484: 463: 459: 453: 440: 434: 428: 422: 413: 410:wall lizards 405: 401: 395: 393: 388: 384: 378: 370: 321: 309: 279:habitat loss 276: 264:metamorphose 258:, newts are 251: 247: 236: 234: 219: 207: 32: 2605:. Bbc.co.uk 2044:smooth newt 1976:smooth newt 1932:aldosterone 1920:nonylphenol 1819:acclimation 1757:Respiration 1751:T. carnifex 1720:Circulation 1693:Chelotriton 1498:Pachytriton 1268:Lissotriton 1105:Pleurodeles 996:Pleurodeles 982:Pachytriton 954:Lissotriton 882:subfamilies 876:Systematics 767:Young larva 696:Development 609:semiaquatic 602:Pleurodeles 550:"ribs" and 542:Michahelles 537:Pleurodeles 424:*agiþahsijǭ 324:Old English 311:Pleurodeles 260:semiaquatic 204:Subfamily: 124:Alpine newt 3205:Categories 3195:Amphibians 3150:2013-06-06 3130:2013-06-06 3110:2013-06-06 3085:2013-06-06 3046:2013-06-06 2707:2019-10-11 2633:2013-06-06 2609:2013-06-06 2536:2013-06-06 2511:2013-06-06 2400:*þahsjō(n) 2362:p. 63 2284:2016-08-29 2217:References 2022:, and the 2010:), or the 1898:activity. 1888:DNA repair 1854:vertebrate 1823:ectotherms 1772:A form of 1739:epicardial 1700:Koalliella 1661:. 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Index

Newt (disambiguation)
EFT
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Alpine newt
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Amphibia
Urodela
Salamandridae
Pleurodelinae
extant
fossil
salamander
subfamily
Salamandridae

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