1887:
1710:
302:
685:
1091:
553:
1033:
962:
1271:
1941:
1235:
1914:
1811:
1647:
921:
887:
1320:
253:
309:
716:
1060:
585:
402:
conditions on the seafloor hindering the decomposition of organic matter as well as rapid burial of fossils, although the presence of bottom-dwelling scavengers suggests that anoxic conditions must have been relatively moderate.
118:
442:
with extremely well-preserved fossils that were deposited during the latest
Maastrichtian, around 66.5 to 66.1 million years ago, making it one of the last lagerstatten to be deposited before the
398:
from the Tethys that increased the region's planktonic productivity, with these fossilized plankton eventually turning into oil. The sites of excellent fossil preservation may have been formed by
382:
of the Afro-Arabian continent, with the
Harrana locality being deposited no deeper than 100 metres (330 ft) below the surface, based on the occurrence of depth-limited fauna such as
2186:
Rosenbach, Kierstin L.; Goodvin, Danielle M.; Albshysh, Mohammed G.; Azzam, Hassan A.; Smadi, Ahmad A.; Mustafa, Hakam A.; Zalmout, Iyad S. A.; Wilson
Mantilla, Jeffrey A. (2024-09-05).
128:
2269:
473:, who described most of the formation's taxa in a 2009 book. Many fossils from this locality are held in the Eternal River Museum of Natural History in Jordan.
2110:
Sokol, Ella V.; Kozmenko, Olga A.; Khoury, Hani N.; Kokh, Svetlana N.; Novikova, Sofya A.; Nefedov, Andrey A.; Sokol, Ivan A.; Zaikin, Pavel (2017-06-01).
2301:
Alhalabi, Wafa A.; Bardet, Nathalie; Sachs, Sven; Kear, Benjamin P.; Joude, Issam B.; Yazbek, Muhammed K.; Godoy, Pedro L.; Langer, Max C. (2024-07-01).
1834:
422:. In addition, the highly fossiliferous portion of the formation was only deposited in the span of a few hundred thousand years at the end of the
1014:
470:
2000:
1819:
1975:
1886:
2112:"Calcareous sediments of the Muwaqqar Chalk Marl Formation, Jordan: Mineralogical and geochemical evidences for Zn and Cd enrichment"
476:
The following list of biota is based primarily on
Kaddumi (2009). Unless stated otherwise, a majority are from the Harrana locality:
443:
301:
2303:"Recovering lost time in Syria: New Late Cretaceous (Coniacian-Santonian) elasmosaurid remains from the Palmyrides mountain chain"
362:, which are among the largest in the world. Some outcrops of the formation contain extremely well-preserved fossils, making it a
394:
at the same time as this depositional event, turning it into an open water ecosystem. The oil shales may have formed from cold
215:
176:
2270:"The Cretaceous–Paleogene transition in spiny-rayed fishes: surveying "Patterson's Gap" in the acanthomorph skeletal record"
418:, which may be due to a drop in sea levels at the time, with deposition continuing following a second sea level rise in the
2228:"Ammonite dating of latest Cretaceous mosasaurid reptiles (Squamata, Mosasauroidea) from Jordan—preliminary observations"
458:
mining, which started in the
Harrana region around 1995. Notable fossils from this locality include articulated fish and
2188:"New pterosaur remains from the Late Cretaceous of Afro-Arabia provide insight into flight capacity of large pterosaurs"
2394:
415:
2389:
2419:
2414:
2399:
169:
2341:
2302:
2111:
2042:
2404:
406:
While the formation as a whole lasts from the early
Maastrichtian to the end of the Paleocene based on
2227:
2187:
2409:
2384:
2086:
387:
347:
108:
2226:
Jagt, John W.M.; Jagt-Yazykova, Elena A.; Kaddumi, Hani F.; Lindgren, Johan (2018-10-02).
8:
359:
355:
1230:, the most completely known member of its genus. Likely adapted to a pelagic lifestyle.
969:
954:
463:
383:
462:
skeletons, mosasaur skin impressions, and evidence of mosasaur carcasses scavenged by
2361:
2322:
2247:
2203:
2165:
2131:
2062:
1377:
1365:
675:
608:
447:
379:
1709:
2353:
2314:
2281:
2239:
2195:
2123:
2054:
1410:
2318:
2243:
2199:
1115:
A bizarre fish with a deeply forked tail and a stooped forehead akin to that of a
2357:
2058:
1570:
1175:
826:
796:
335:
999:
593:
358:
from north to south. It is the geological formation containing Jordan's famous
208:
2085:
Khrewesh, Amani M.; Hamad, Abdullah Abu; Abed, Abdulkader M. (December 2014).
439:
363:
2378:
2365:
2326:
2251:
2207:
2169:
2135:
2127:
2066:
1879:
1732:
1717:
1702:
1249:
1168:
1068:
951:
935:
879:
561:
423:
399:
339:
230:
217:
191:
178:
65:
30:
24:
1090:
1032:
552:
2014:
1922:
1624:
1525:
1513:
1399:
1388:
1227:
961:
799:
783:
762:
750:
684:
435:
411:
407:
391:
375:
2286:
2268:
Matt, Friedman; James V., Andrews; Hadeel, Saad; Sanaa, El-Sayed (2023).
1895:
1800:
1577:
1561:
1312:
1270:
1167:
An indeterminate marine reptile, initially described as a late-surviving
1083:
1003:
987:
772:
693:
663:
630:
618:
577:
542:
469:
The fossil potential of the
Harrana locality was documented by geologist
273:
40:
1940:
1234:
1913:
1810:
1803:
1646:
1590:
1580:
1297:
1212:
1171:
1153:
1116:
920:
901:
821:
809:
767:
708:
527:
451:
157:
85:
50:
2087:"Late Cretaceous Muwaqqar Formation Ammonites in Southeastern Jordan"
1785:
1684:
1041:
652:
640:
455:
419:
395:
343:
90:
34:
1319:
886:
1861:
1278:
1261:
864:
853:
841:
516:
504:
459:
80:
75:
60:
55:
45:
1417:
has the first known complete mosasaur skull from the Middle East.
1223:
A complete skull, some vertebrae, and digits from front flippers.
1079:
Six articulated specimens, the most complete known for the genus.
1989:
1636:
1104:
1059:
715:
446:. The preservation is akin to that of the older, more well-known
95:
70:
2225:
386:. The formation of this habitat likely originates from a major
351:
258:
2340:
Farouk, Sherif; Marzouk, Akmal M.; Ahmad, Fayez (2014-11-01).
2041:
Farouk, Sherif; Marzouk, Akmal M.; Ahmad, Fayez (2014-11-01).
584:
2185:
545:
143:
147:
2162:
Fossils of the
Harrana fauna : and the adjacent areas
977:
Poorly-preserved articulated skull & vertebral column
947:
Articulated skull, rest of the skeleton most likely lost
2300:
2267:
2109:
980:
An indeterminate ichthyodectid with very large teeth.
1705:
pterosaur, one of the most complete from the region.
1266:
A small durophagous globidensine. Remains very rare.
580:, teeth found embedded in a fossil mosasaur carcass.
308:
2232:
1480:
Cranial elements including left maxilla with teeth.
2164:. Jordan: Eternal River Museum of Natural History.
2339:
2094:Jordan Journal of Earth and Environmental Sciences
2040:
1174:, but more recently found to possibly be a marine
374:The formation appears to have been deposited in a
2084:
2376:
1406:Complete post-cranial skeleton missing skull.
1332:Hindlimbs, sacral vertebrae, preserved scales
875:Articulated partial specimens, isolated teeth
2342:"The Cretaceous/Paleogene boundary in Jordan"
2043:"The Cretaceous/Paleogene boundary in Jordan"
604:3 articulated specimens, one nearly complete
2036:
2034:
1002:-like percomorph (initially identified as a
1939:
1912:
1885:
1809:
1708:
1645:
1318:
1269:
1058:
1031:
1025:Articulated skull with associated remains
960:
714:
683:
583:
551:
2285:
2031:
1233:
1089:
919:
885:
2159:
1695:Jaws, vertebrae, and front limb bones.
450:from Lebanon. Fossils are contained in
2377:
316:Muwaqqar Chalk-Marl Formation (Jordan)
479:
444:Cretaceous-Paleogene extinction event
378:subtropical environment on the outer
119:Um Al'Rijam Chert-Limestone Formation
2263:
2261:
2221:
2219:
2217:
2181:
2179:
2155:
2153:
2151:
2149:
2147:
2145:
2105:
2103:
2080:
2078:
2076:
1289:A mosasaur of uncertain affinities.
1642:A very large cheloniid sea turtle.
1028:An ichthyodectid ichthyodectiform.
13:
2192:Journal of Vertebrate Paleontology
771:interpretation, but potentially a
14:
2431:
2258:
2214:
2176:
2142:
2100:
2073:
414:exists in some localities at the
1119:. Taxonomic identity uncertain.
307:
300:
251:
38:
2346:Journal of Asian Earth Sciences
2047:Journal of Asian Earth Sciences
1953:
1763:
1659:
1540:
1190:
1131:
728:
483:
2333:
2294:
1759:
1052:Four fully articulated skulls
830:and possibly a new species of
369:
1:
2319:10.1016/j.cretres.2024.105871
2244:10.1080/03115518.2017.1308011
2200:10.1080/02724634.2024.2385068
2025:
1655:
429:
416:Cretaceous-Paleogene boundary
410:-based dating, a significant
328:Muwaqqar Chalk-Marl Formation
19:Muwaqqar Chalk-Marl Formation
2358:10.1016/j.jseaes.2014.08.015
2059:10.1016/j.jseaes.2014.08.015
1186:
724:
651:A complete, fully-preserved
209:Approximate paleocoordinates
129:Alhisa Phosphorite Formation
7:
2160:Kaddumi, Hani Faig (2009).
1949:
1743:
1493:Lumbar vertebrae and pygal
1467:Partial dentary with teeth
1127:
916:An enchodontid aulopiform.
10:
2436:
1536:
512:Articulated pectoral wing
354:, cropping out across the
2395:Cretaceous System of Asia
1936:A sphenodiscid ammonite.
1832:
1817:
1616:A very large sea turtle.
1601:A very large sea turtle.
1408:
1386:
1310:
1295:
626:Nearly complete specimen
519:of uncertain affinities.
295:
287:
279:
269:
264:
246:
207:
168:
163:
153:
139:
134:
124:
114:
104:
23:
18:
2128:10.1016/j.gr.2017.03.008
1909:A pachydiscid ammonite.
1286:Articulated hind paddle
1796:Jebal Khuzaym, Harrana
1608:Chelonioidea indet. 2.
1055:A dercetid aulopiform.
824:aulopiform, similar to
2390:Paleontology in Jordan
1345:Front paddle, humerus
1242:
1098:
928:
913:Articulated skeletons
894:
1237:
1093:
923:
889:
792:Articulated skeleton
440:Konservat-Lagerstätte
2420:Oil shale formations
2287:10.20341/gb.2023.002
466:& other fishes.
388:marine transgression
348:geological formation
109:Geological formation
2415:Oil shale in Jordan
2400:Maastrichtian Stage
2307:Cretaceous Research
1454:Cranium with teeth
1241:, a related species
1164:Rostrum with teeth
1097:, a related species
927:, a related species
893:, a related species
758:Complete skeletons
356:Jordanian Highlands
227: /
188: /
25:Stratigraphic range
1243:
1112:Complete skeleton
1099:
1086:ichthyodectiform.
995:Complete skeleton
929:
895:
849:Complete skeleton
827:Dercetis triqueter
817:Complete skeleton
648:Complete specimen
629:A nearly complete
480:Cartilaginous fish
396:upwelling currents
332:Muwaqqar Formation
2405:Cretaceous Jordan
2274:Geologica Belgica
2116:Gondwana Research
2023:
2022:
1947:
1946:
1748:Based on Krewesh
1741:
1740:
1653:
1652:
1534:
1533:
1184:
1183:
1125:
1124:
722:
721:
448:Sannine Formation
380:continental shelf
324:
323:
2427:
2370:
2369:
2337:
2331:
2330:
2298:
2292:
2291:
2289:
2265:
2256:
2255:
2223:
2212:
2211:
2183:
2174:
2173:
2157:
2140:
2139:
2107:
2098:
2097:
2091:
2082:
2071:
2070:
2038:
1954:
1943:
1916:
1889:
1826:L. acutidorsatus
1813:
1764:
1752:(2014) and Jagt
1712:
1660:
1649:
1598:A right humerus
1541:
1441:Complete skull.
1411:prognathodontine
1378:plioplatecarpine
1366:Plioplatecarpini
1322:
1273:
1191:
1132:
1062:
1035:
964:
955:ichthyodectiform
729:
718:
687:
674:Nearly complete
587:
555:
484:
311:
310:
304:
257:
255:
254:
242:
241:
239:
238:
237:
232:
228:
225:
224:
223:
220:
203:
202:
200:
199:
198:
193:
189:
186:
185:
184:
181:
100:
37:
16:
15:
2435:
2434:
2430:
2429:
2428:
2426:
2425:
2424:
2410:Marl formations
2375:
2374:
2373:
2338:
2334:
2299:
2295:
2266:
2259:
2224:
2215:
2184:
2177:
2158:
2143:
2108:
2101:
2089:
2083:
2074:
2039:
2032:
2028:
1952:
1762:
1746:
1658:
1571:pectoral girdle
1539:
1219:C. palistinicus
1189:
1130:
1021:P. harranaensis
970:Ichthyodectidae
871:E. harranaensis
727:
700:S. pristodontus
534:C. biauriculata
482:
432:
372:
336:Late Cretaceous
320:
319:
318:
317:
314:
313:
312:
252:
250:
235:
233:
229:
226:
221:
218:
216:
214:
213:
196:
194:
190:
187:
182:
179:
177:
175:
174:
99:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
29:
28:
12:
11:
5:
2433:
2423:
2422:
2417:
2412:
2407:
2402:
2397:
2392:
2387:
2372:
2371:
2332:
2293:
2257:
2238:(4): 587–596.
2213:
2175:
2141:
2099:
2072:
2029:
2027:
2024:
2021:
2020:
2018:
2011:
2009:
2004:
1996:
1995:
1993:
1986:
1984:
1982:L. mayereymari
1979:
1971:
1970:
1967:
1964:
1961:
1958:
1951:
1948:
1945:
1944:
1937:
1934:
1933:Jebal Khuzaym
1931:
1926:
1918:
1917:
1910:
1907:
1904:
1899:
1891:
1890:
1883:
1876:
1873:
1865:
1857:
1856:
1854:
1853:Jebal Khuzaym
1851:
1841:
1840:
1838:
1831:
1828:
1823:
1815:
1814:
1807:
1797:
1794:
1789:
1781:
1780:
1777:
1774:
1771:
1768:
1761:
1758:
1745:
1742:
1739:
1738:
1736:
1729:
1726:
1723:
1721:
1714:
1713:
1706:
1699:
1696:
1693:
1688:
1680:
1679:
1676:
1673:
1670:
1667:
1664:
1657:
1654:
1651:
1650:
1643:
1640:
1633:
1628:
1620:
1619:
1617:
1614:
1611:
1609:
1605:
1604:
1602:
1599:
1596:
1594:
1587:
1586:
1584:
1574:
1567:
1565:
1558:
1557:
1554:
1551:
1548:
1545:
1538:
1535:
1532:
1531:
1529:
1522:
1521:Front paddle.
1519:
1517:
1510:
1509:
1507:
1504:
1497:
1496:
1494:
1491:
1484:
1483:
1481:
1478:
1471:
1470:
1468:
1465:
1458:
1457:
1455:
1452:
1445:
1444:
1442:
1439:
1433:
1432:
1430:
1429:Left dentary.
1427:
1421:
1420:
1418:
1407:
1404:
1392:
1384:
1383:
1381:
1374:
1371:
1369:
1362:
1361:
1359:
1356:
1349:
1348:
1346:
1343:
1336:
1335:
1333:
1330:
1324:
1323:
1316:
1309:
1306:
1301:
1293:
1292:
1290:
1287:
1284:
1282:
1275:
1274:
1267:
1264:
1258:
1253:
1245:
1244:
1231:
1224:
1221:
1216:
1208:
1207:
1204:
1201:
1198:
1195:
1188:
1185:
1182:
1181:
1179:
1176:crocodylomorph
1165:
1162:
1157:
1149:
1148:
1145:
1142:
1139:
1136:
1129:
1126:
1123:
1122:
1120:
1113:
1110:
1108:
1101:
1100:
1095:S. lanciformis
1087:
1080:
1077:
1075:S. longicorpus
1072:
1064:
1063:
1056:
1053:
1050:
1045:
1037:
1036:
1029:
1026:
1023:
1018:
1010:
1009:
1007:
996:
993:
991:
984:
983:
981:
978:
975:
973:
966:
965:
958:
948:
945:
939:
931:
930:
917:
914:
911:
905:
897:
896:
883:
876:
873:
868:
860:
859:
857:
852:A relative of
850:
847:
845:
838:
837:
835:
818:
815:
813:
806:
805:
803:
793:
790:
788:
779:
778:
776:
759:
756:
754:
746:
745:
742:
739:
736:
733:
726:
723:
720:
719:
712:
705:
702:
697:
689:
688:
681:
678:
672:
667:
659:
658:
656:
649:
646:
644:
637:
636:
634:
631:mackerel shark
627:
624:
622:
615:
614:
612:
605:
602:
597:
594:Harranahynchus
589:
588:
581:
574:
571:
565:
557:
556:
549:
539:
536:
531:
523:
522:
520:
513:
510:
508:
501:
500:
497:
494:
491:
488:
481:
478:
434:A locality in
431:
428:
371:
368:
322:
321:
315:
306:
305:
299:
298:
297:
296:
293:
292:
289:
285:
284:
281:
277:
276:
271:
267:
266:
262:
261:
248:
244:
243:
211:
205:
204:
172:
166:
165:
161:
160:
155:
151:
150:
141:
137:
136:
132:
131:
126:
122:
121:
116:
112:
111:
106:
102:
101:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
21:
20:
9:
6:
4:
3:
2:
2432:
2421:
2418:
2416:
2413:
2411:
2408:
2406:
2403:
2401:
2398:
2396:
2393:
2391:
2388:
2386:
2383:
2382:
2380:
2367:
2363:
2359:
2355:
2351:
2347:
2343:
2336:
2328:
2324:
2320:
2316:
2312:
2308:
2304:
2297:
2288:
2283:
2279:
2275:
2271:
2264:
2262:
2253:
2249:
2245:
2241:
2237:
2233:
2229:
2222:
2220:
2218:
2209:
2205:
2201:
2197:
2193:
2189:
2182:
2180:
2171:
2167:
2163:
2156:
2154:
2152:
2150:
2148:
2146:
2137:
2133:
2129:
2125:
2121:
2117:
2113:
2106:
2104:
2095:
2088:
2081:
2079:
2077:
2068:
2064:
2060:
2056:
2052:
2048:
2044:
2037:
2035:
2030:
2019:
2016:
2012:
2010:
2008:
2005:
2003:
2002:
1998:
1997:
1994:
1991:
1987:
1985:
1983:
1980:
1978:
1977:
1973:
1972:
1968:
1965:
1962:
1959:
1956:
1955:
1942:
1938:
1935:
1932:
1930:
1927:
1925:
1924:
1920:
1919:
1915:
1911:
1908:
1905:
1903:
1902:P. dossantosi
1900:
1898:
1897:
1893:
1892:
1888:
1884:
1881:
1877:
1874:
1872:
1871:fresvillensis
1869:
1866:
1864:
1863:
1859:
1858:
1855:
1852:
1850:
1846:
1843:
1842:
1839:
1836:
1829:
1827:
1824:
1822:
1821:
1816:
1812:
1808:
1805:
1802:
1798:
1795:
1793:
1790:
1788:
1787:
1783:
1782:
1778:
1775:
1772:
1769:
1766:
1765:
1757:
1755:
1751:
1737:
1734:
1730:
1727:
1724:
1722:
1719:
1718:Nyctosauridae
1716:
1715:
1711:
1707:
1704:
1700:
1697:
1694:
1692:
1689:
1687:
1686:
1682:
1681:
1677:
1674:
1671:
1668:
1665:
1662:
1661:
1648:
1644:
1641:
1638:
1634:
1632:
1629:
1627:
1626:
1622:
1621:
1618:
1615:
1613:Front paddle
1612:
1610:
1607:
1606:
1603:
1600:
1597:
1595:
1592:
1589:
1588:
1585:
1582:
1579:
1576:A very large
1575:
1572:
1568:
1566:
1563:
1560:
1559:
1555:
1552:
1549:
1546:
1543:
1542:
1530:
1527:
1523:
1520:
1518:
1515:
1512:
1511:
1508:
1505:
1502:
1499:
1498:
1495:
1492:
1489:
1486:
1485:
1482:
1479:
1476:
1473:
1472:
1469:
1466:
1463:
1460:
1459:
1456:
1453:
1450:
1447:
1446:
1443:
1440:
1438:
1435:
1434:
1431:
1428:
1426:
1423:
1422:
1419:
1416:
1412:
1405:
1402:
1401:
1396:
1393:
1391:
1390:
1385:
1382:
1379:
1375:
1372:
1370:
1367:
1364:
1363:
1360:
1357:
1354:
1351:
1350:
1347:
1344:
1341:
1338:
1337:
1334:
1331:
1329:
1326:
1325:
1321:
1317:
1314:
1307:
1305:
1302:
1300:
1299:
1294:
1291:
1288:
1285:
1283:
1280:
1277:
1276:
1272:
1268:
1265:
1263:
1259:
1257:
1254:
1252:
1251:
1250:Harranasaurus
1247:
1246:
1240:
1236:
1232:
1229:
1225:
1222:
1220:
1217:
1215:
1214:
1210:
1209:
1205:
1202:
1199:
1196:
1193:
1192:
1180:
1177:
1173:
1170:
1166:
1163:
1161:
1160:R. singularis
1158:
1156:
1155:
1151:
1150:
1146:
1143:
1140:
1137:
1134:
1133:
1121:
1118:
1114:
1111:
1109:
1106:
1103:
1102:
1096:
1092:
1088:
1085:
1081:
1078:
1076:
1073:
1071:
1070:
1069:Saurocephalus
1066:
1065:
1061:
1057:
1054:
1051:
1049:
1046:
1044:
1043:
1039:
1038:
1034:
1030:
1027:
1024:
1022:
1019:
1017:
1016:
1012:
1011:
1008:
1005:
1001:
997:
994:
992:
989:
986:
985:
982:
979:
976:
974:
971:
968:
967:
963:
959:
956:
953:
952:ichthyodectid
949:
946:
943:
940:
938:
937:
936:Ichthyodectes
933:
932:
926:
922:
918:
915:
912:
909:
906:
904:
903:
899:
898:
892:
888:
884:
881:
877:
874:
872:
869:
867:
866:
862:
861:
858:
855:
851:
848:
846:
843:
840:
839:
836:
833:
829:
828:
823:
819:
816:
814:
811:
808:
807:
804:
801:
798:
797:cimolichthyid
794:
791:
789:
786:
785:
781:
780:
777:
774:
770:
769:
764:
760:
757:
755:
752:
748:
747:
743:
740:
737:
734:
731:
730:
717:
713:
710:
706:
703:
701:
698:
696:
695:
691:
690:
686:
682:
679:
677:
673:
671:
668:
666:
665:
661:
660:
657:
654:
650:
647:
645:
642:
639:
638:
635:
632:
628:
625:
623:
620:
617:
616:
613:
610:
606:
603:
601:
600:H. minutadens
598:
596:
595:
591:
590:
586:
582:
579:
575:
572:
569:
566:
564:
563:
562:Ginglymostoma
559:
558:
554:
550:
547:
544:
540:
537:
535:
532:
530:
529:
525:
524:
521:
518:
514:
511:
509:
506:
503:
502:
498:
495:
492:
489:
486:
485:
477:
474:
472:
467:
465:
461:
457:
453:
449:
445:
441:
437:
427:
425:
424:Maastrichtian
421:
417:
413:
409:
404:
401:
397:
393:
389:
385:
381:
377:
367:
365:
361:
357:
353:
349:
345:
341:
340:Maastrichtian
337:
333:
329:
303:
294:
290:
286:
282:
278:
275:
272:
268:
263:
260:
249:
245:
240:
231:14.9°N 29.8°E
212:
210:
206:
201:
192:31.7°N 36.5°E
173:
171:
167:
162:
159:
156:
152:
149:
145:
142:
138:
133:
130:
127:
123:
120:
117:
113:
110:
107:
103:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
36:
32:
31:Maastrichtian
26:
22:
17:
2385:Lagerstätten
2349:
2345:
2335:
2310:
2306:
2296:
2277:
2273:
2235:
2231:
2191:
2161:
2119:
2115:
2093:
2050:
2046:
2006:
1999:
1981:
1974:
1928:
1923:Sphenodiscus
1921:
1901:
1894:
1870:
1867:
1860:
1848:
1844:
1835:sphenodiscid
1825:
1818:
1791:
1784:
1753:
1749:
1747:
1725:Wing bones.
1690:
1683:
1630:
1625:Gigantatypus
1623:
1591:Chelonioidea
1569:Articulated
1514:Tylosaurinae
1500:
1487:
1474:
1461:
1448:
1436:
1424:
1414:
1400:Tenerasaurus
1398:
1394:
1389:Prognathodon
1387:
1373:Hind paddle
1352:
1339:
1327:
1303:
1296:
1279:Mosasauridae
1255:
1248:
1238:
1228:globidensine
1218:
1211:
1159:
1152:
1094:
1074:
1067:
1047:
1040:
1020:
1015:Postredectes
1013:
941:
934:
925:E. boissieri
924:
907:
900:
890:
882:aulopiform.
870:
863:
831:
825:
784:Cimolichthys
782:
766:
765:" under the
751:Beryciformes
699:
692:
680:A sawskate.
669:
662:
641:Myliobatidae
599:
592:
567:
560:
533:
526:
475:
471:Hani Kaddumi
468:
464:nurse sharks
433:
412:uncomformity
408:foraminifera
405:
392:Tethys Ocean
384:nurse sharks
373:
342:) and early
331:
327:
325:
288:Year defined
265:Type section
2352:: 113–125.
2122:: 204–226.
2053:: 113–125.
2007:T. argentea
2001:Tenuipteria
1896:Pachydiscus
1880:pachydiscid
1820:Libycoceras
1760:Cephalopoda
1735:pterosaur.
1733:nyctosaurid
1703:azhdarchoid
1698:Tal Inab 6
1691:I. alarabia
1562:Cheloniidae
1526:tylosaurine
1313:mosasaurine
1304:M. hoffmani
1256:H. khuludae
1239:C. belgicus
1169:polycotylid
1117:dolphinfish
1084:saurodontid
1048:S. apicalis
988:Percomorpha
891:E. petrosus
880:enchodontid
694:Squalicorax
670:S. stromeri
664:Schizorhiza
619:Lamniformes
578:nurse shark
515:A gigantic
454:exposed by
452:concretions
370:Environment
364:lagerstätte
274:Al-Muwaqqar
234: /
195: /
170:Coordinates
2379:Categories
2313:: 105871.
2026:References
2015:inoceramid
1976:Lyropecten
1929:S. lobatus
1882:ammonite.
1837:ammonite.
1656:Pterosaurs
1581:sea turtle
1506:Two teeth
1395:P. hashimi
1328:M. sevciki
1298:Mosasaurus
1213:Carinodens
1172:plesiosaur
1154:Rarosaurus
902:Eurypholis
810:Dercetidae
800:aulopiform
768:sensu lato
763:beryciform
709:crow shark
528:Cretalamna
430:Paleobiota
360:oil shales
236:14.9; 29.8
197:31.7; 36.5
158:Oil shales
2366:1367-9120
2327:0195-6671
2252:0311-5518
2208:0272-4634
2170:709582892
2136:1342-937X
2067:1367-9120
1963:Material
1801:baculitid
1792:B. ovatus
1786:Baculites
1773:Material
1685:Inabtanin
1672:Locality
1669:Material
1631:G. salahi
1593:indet. 1
1578:cheloniid
1550:Material
1437:P. primus
1415:P. primus
1358:Vertebra
1200:Material
1187:Mosasaurs
1141:Material
1105:Teleostei
1042:Stratodus
1004:perciform
1000:moonyfish
773:veliferid
753:' indet.
738:Material
725:Bony fish
653:eagle ray
543:otodontid
493:Material
456:limestone
420:Paleocene
344:Paleogene
270:Named for
135:Lithology
115:Underlies
35:Paleocene
1960:Species
1950:Bivalvia
1906:Harrana
1875:Harrana
1862:Menuites
1830:Harrana
1804:ammonite
1770:Species
1756:(2017):
1744:Mollusca
1728:Harrana
1666:Species
1635:A right
1547:Species
1425:P. hudae
1197:Species
1138:Species
1128:Reptiles
865:Enchodus
854:ladyfish
842:Elopidae
832:Dercetis
822:dercetid
735:Species
609:sawskate
505:Batoidea
490:Species
460:mosasaur
283:M. Masri
280:Named by
164:Location
125:Overlies
33:to late
2280:(1–2).
1990:scallop
1969:Images
1849:ismaeli
1779:Images
1720:indet.
1678:Images
1637:humerus
1564:indet.
1556:Images
1537:Turtles
1516:indet.
1368:indet.
1281:indet.
1262:dentary
1206:Images
1147:Images
1107:indet.
990:indet.
972:indet.
844:indet.
812:indet.
744:Images
676:rostrum
643:indet.
621:indet.
507:indet.
499:Images
436:Harrana
390:of the
376:pelagic
247:Country
222:29°48′E
219:14°54′N
183:36°30′E
180:31°42′N
140:Primary
2364:
2325:
2250:
2206:
2168:
2134:
2065:
1966:Notes
1957:Genus
1776:Notes
1767:Genus
1675:Notes
1663:Genus
1553:Notes
1544:Genus
1503:sp. 5
1490:sp. 4
1477:sp. 3
1464:sp. 2
1451:sp. 1
1355:sp. 2
1342:sp. 1
1308:Teeth
1260:Right
1203:Notes
1194:Genus
1144:Notes
1135:Genus
741:Notes
732:Genus
704:Teeth
573:Teeth
538:Teeth
496:Notes
487:Genus
400:anoxic
352:Jordan
346:-aged
259:Jordan
256:
2090:(PDF)
1754:et al
1750:et al
546:shark
438:is a
334:is a
154:Other
144:Chalk
2362:ISSN
2323:ISSN
2248:ISSN
2204:ISSN
2166:OCLC
2132:ISSN
2063:ISSN
1847:cf.
944:sp.
910:sp.
787:sp.
570:sp.
326:The
291:1963
148:marl
105:Type
41:Preęž’
2354:doi
2315:doi
2311:159
2282:doi
2240:doi
2196:doi
2124:doi
2055:doi
2013:An
1701:An
1006:).
950:An
878:An
761:A "
541:An
517:ray
350:in
330:or
2381::
2360:.
2350:94
2348:.
2344:.
2321:.
2309:.
2305:.
2278:26
2276:.
2272:.
2260:^
2246:.
2236:42
2234:.
2230:.
2216:^
2202:.
2194:.
2190:.
2178:^
2144:^
2130:.
2120:46
2118:.
2114:.
2102:^
2092:.
2075:^
2061:.
2051:94
2049:.
2045:.
2033:^
2017:.
1992:.
1988:A
1878:A
1868:M.
1845:L.
1833:A
1806:.
1799:A
1731:A
1639:.
1583:.
1573:.
1528:.
1524:A
1501:P.
1488:P.
1475:P.
1462:P.
1449:P.
1413:.
1409:A
1403:)
1397:(=
1380:.
1376:A
1353:M.
1340:M.
1315:.
1311:A
1226:A
1178:.
1082:A
998:A
957:.
942:I.
908:E.
856:.
834:.
820:A
802:.
795:A
775:.
711:.
707:A
655:.
633:.
611:.
607:A
576:A
568:G.
548:.
426:.
366:.
146:,
91:Pg
27::
2368:.
2356::
2329:.
2317::
2290:.
2284::
2254:.
2242::
2210:.
2198::
2172:.
2138:.
2126::
2096:.
2069:.
2057::
749:'
338:(
96:N
86:K
81:J
76:T
71:P
66:C
61:D
56:S
51:O
46:ęž’
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
