133:
474:
155:
1806:
2264:
423:
is a medium-sized mosasaur with an estimated skull length of 0.8 metres (2.6 ft) and a body length of approximately 6 metres (20 ft). Both the front and hind flippers are longer than the skull. This feature is not seen in any other mosasauroid with paddle-like limbs. Furthermore, the hind
1391:
Konishi, Takuya; Ohara, Masaaki; Misaki, Akihiro; Matsuoka, Hiroshige; Street, Hallie P.; Caldwell, Michael W. (2023). "A new derived mosasaurine (Squamata: Mosasaurinae) from south-western Japan reveals unexpected postcranial diversity among hydropedal mosasaurs".
1284:
holotype, based on related taxa, it likely had a crescent-shaped tail fin that could have aided in propulsion and fast turning. Based on the combination of all of these features and the large size of the flippers, Konishi
357:
of Japan. The specimen consists of a partial skull, a complete cervical and dorsal vertebral series with more than 40 vertebrae, ribs, both front flippers, and the left hind flipper.
2180:
321:
1345:
2314:
445:
dorsal vertebrae exhibit an abrupt change in orientation, with at least five neural spines projecting anterodorsally. This is also seen in some
1475:
1454:
1323:
1270:
was capable of rapid pitch control (rotation) and braking. Skull part around eye socket is slightly wide horizontally, which suggests that had
346:
2251:
350:
1317:
17:
442:
2375:
1546:
1255:
349:-Ge-1140240002, was discovered in sediments of the Hasegawa Muddy Sandstone Member of the Toyajo Formation near the peak of
1498:
1293:
would have been a capable hunter of large groups of small prey items that were hard to capture, like schools of fish.
1259:
453:. Also like cetaceans, the change in orientation occurs posterior to the center of gravity of the animal, where in
1262:
of the front flippers, allowing for quick maneuvering. The femoral head is well-rounded and almost circular in
2241:
1280:, this is the second case of mosasaur with binocular vision. Although the tail is not well preserved in the
273:
154:
1446:
2360:
1539:
2370:
2365:
1632:
2246:
1254:
is saddle-shaped and mediolaterally thick. This likely indicates a high range of motion for both
394:
2332:
2319:
2282:
1339:
2355:
2327:
1532:
528:
1805:
1401:
408:
is "Wakayama Soryu" (和歌山滄竜), which translates to "Wakayama blue dragon". The word "Soryu" (
1469:
461:. If present, a dorsal fin in a mosasaur could provide stability while moving underwater.
8:
364:
354:
1405:
2267:
1514:
1425:
149:
1949:
1892:
1429:
1417:
2305:
2224:
1926:
1726:
1660:
1510:
1409:
1271:
908:
477:
1413:
424:
flippers are even larger than the front flippers, which is otherwise only seen in
401:" honors Wakayama Prefecture, where the holotype was found and is currently kept.
2217:
2153:
2118:
2074:
1956:
1935:
1872:
1857:
1276:
324:(Hasegawa Muddy Sandstone Member) of Japan. The genus contains a single species,
41:
132:
2090:
2021:
1762:
1708:
473:
2349:
2194:
2166:
2124:
2097:
2056:
2012:
1983:
1742:
1701:
1421:
389:, meaning wing, referencing the unusually large wing-shaped flippers seen in
379:
370:
as a new genus and species of mosasaurine based on these fossil remains. The
83:
45:
409:
2145:
2039:
1976:
1967:
1915:
1908:
1901:
1881:
1814:
1769:
1735:
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1625:
1604:
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1354:
620:
580:
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represents the first published evidence for such a structure in mosasaurs.
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248:
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2187:
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2028:
1997:
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1834:
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1618:
1307:
1045:
963:
556:
523:
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488:
317:
58:
2063:
2049:
1850:
1783:
1687:
1667:
1639:
1334:
1329:
1020:
980:
517:
511:
450:
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426:
222:
103:
68:
2173:
1990:
1843:
1823:
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1611:
1361:
holotype, suggesting it was scavenged by these sharks after it died.
784:
688:
532:
166:
108:
52:
2276:
2299:
1715:
1585:
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1573:
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1302:
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93:
78:
73:
63:
313:
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113:
88:
1646:
1567:
176:
371:
1524:
1390:
457:, the affected vertebrae are located after the main
527:. The 50% majority-rule consensus results of their
1353:holotype site. In addition, more than one hundred
2347:
1386:
1384:
1382:
1380:
1378:
1376:
1374:
449:whales, where it corresponds to the base of a
1540:
2252:List of mosasaur-bearing stratigraphic units
1371:
1833:
1547:
1533:
330:, known from an almost complete skeleton.
131:
412:竜) itself is used to refer to mosasaurs.
1447:"有田川町産出のモササウルス類は新属新種！！－これまでの学説を覆す新たな発見－"
472:
1470:"モササウルスとは？ ｜ モササウルス発掘プロジェクト｜和歌山県立自然博物館"
333:
14:
2348:
2281:
2280:
1528:
1441:
1439:
1394:Journal of Systematic Palaeontology
1274:that helped it to hunt prey. After
24:
1515:10.15080/agcjchikyukagaku.75.2_147
1496:
1357:shark teeth were found around the
320:mosasaur from the Late Cretaceous
25:
2387:
1436:
468:
2263:
2262:
1804:
1349:) have also been found near the
153:
56:
1296:
1245:
1490:
1462:
415:
13:
1:
2376:Fossil taxa described in 2023
2242:Timeline of mosasaur research
1554:
1414:10.1080/14772019.2023.2277921
1364:
312:(meaning "large wing") is an
1346:Gigantocapulus problematicus
7:
1499:"博物館と発掘現場で体感する日本一のモササウルス化石"
368:Megapterygius wakayamaensis
291:Megapterygius wakayamaensis
18:Megapterygius wakayamaensis
10:
2392:
1266:view, which suggests that
27:Extinct genus of mosasaurs
2289:
2260:
2234:
2112:Related groups and genera
2111:
2073:
2038:
2011:
1966:
1925:
1891:
1871:
1813:
1802:
1752:
1725:
1677:
1594:
1562:
1103:
1088:
1081:
1066:
1059:
1042:
1035:
1017:
1010:
977:
960:
953:
946:
929:
922:
905:
898:
874:
859:
852:
845:
830:
823:
799:
781:
774:
767:
725:
710:
703:
685:
678:
666:Prognathodon waiparaensis
663:
656:
641:
634:
617:
610:
595:
577:
570:
553:
546:
483:Konishi (2023) recovered
287:
280:
150:Scientific classification
148:
139:
130:
34:
2247:List of mosasaur genera
378:", is derived from the
1308:Pachydiscus awajiensis
1260:protraction/retraction
877:Prognathodon saturator
802:Globidens alabamaensis
503:to a clade containing
480:
404:The Japanese name for
2328:Paleobiology Database
1340:Globularia izumiensis
1091:Mosasaurus hoffmannii
1069:Mosasaurus lemonnieri
862:Prognathodon overtoni
529:phylogenetic analyses
476:
385:, meaning large, and
1250:The humeral head of
598:Prognathodon solvayi
334:Discovery and naming
144:compared to a human
1406:2023JSPal..2177921K
1256:adduction/abduction
728:Clidastes liodontus
713:Clidastes propython
644:Prognathodon kianda
355:Wakayama Prefecture
1106:Mosasaurus maximus
833:Prognathodon rapax
481:
2361:Mosasaurs of Asia
2343:
2342:
2283:Taxon identifiers
2274:
2273:
2107:
2106:
2007:
2006:
1950:Plesioplatecarpus
1893:Plioplatecarpinae
1867:
1866:
1800:
1799:
1242:
1241:
1233:
1232:
1224:
1223:
1215:
1214:
1206:
1205:
1197:
1196:
1188:
1187:
1179:
1178:
1170:
1169:
1161:
1160:
1152:
1151:
1143:
1142:
1134:
1133:
1125:
1124:
1116:
1115:
999:
998:
887:
886:
812:
811:
756:
755:
747:
746:
738:
737:
531:are shown in the
360:In 2023, Konishi
305:
304:
276:
16:(Redirected from
2383:
2371:Cretaceous Japan
2366:Fossils of Japan
2336:
2335:
2323:
2322:
2310:
2309:
2308:
2278:
2277:
2266:
2265:
2235:Related articles
2225:Vallecillosaurus
1927:Plioplatecarpini
1923:
1922:
1889:
1888:
1831:
1830:
1808:
1727:Prognathodontini
1675:
1674:
1661:Plesiotylosaurus
1549:
1542:
1535:
1526:
1525:
1519:
1518:
1494:
1488:
1487:
1485:
1484:
1466:
1460:
1459:
1451:
1443:
1434:
1433:
1388:
1272:binocular vision
1084:
1083:
1062:
1061:
1038:
1037:
1013:
1012:
990:
956:
955:
949:
948:
925:
924:
909:Plesiotylosaurus
901:
900:
855:
854:
848:
847:
826:
825:
777:
776:
770:
769:
706:
705:
681:
680:
659:
658:
637:
636:
613:
612:
573:
572:
549:
548:
539:
538:
478:Life restoration
327:M. wakayamaensis
322:Toyajo Formation
267:
260:
247:
234:
221:
158:
157:
135:
125:
55:
40:Temporal range:
32:
31:
21:
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2390:
2386:
2385:
2384:
2382:
2381:
2380:
2346:
2345:
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2331:
2326:
2318:
2313:
2304:
2303:
2298:
2285:
2275:
2270:
2256:
2230:
2218:Portunatasaurus
2154:Dolichosauridae
2119:Aigialosauridae
2103:
2075:Yaguarasaurinae
2069:
2034:
2003:
1962:
1957:Plioplatecarpus
1936:Latoplatecarpus
1921:
1887:
1873:Russellosaurina
1863:
1858:Phosphorosaurus
1829:
1809:
1796:
1748:
1721:
1673:
1590:
1558:
1553:
1523:
1522:
1497:正顕, 小原 (2021).
1495:
1491:
1482:
1480:
1474:モササウルス発掘プロジェクト｜
1468:
1467:
1463:
1449:
1445:
1444:
1437:
1389:
1372:
1367:
1299:
1289:speculate that
1277:Phosphorosaurus
1248:
1243:
1234:
1225:
1216:
1207:
1198:
1189:
1180:
1171:
1162:
1153:
1144:
1135:
1126:
1117:
1000:
984:
888:
813:
757:
748:
739:
471:
418:
336:
301:
294:
266:
258:
245:
232:
219:
152:
126:
124:
123:
122:
121:
116:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
50:
49:
42:Late Cretaceous
38:
28:
23:
22:
15:
12:
11:
5:
2389:
2379:
2378:
2373:
2368:
2363:
2358:
2341:
2340:
2338:
2337:
2324:
2311:
2295:
2293:
2287:
2286:
2272:
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2261:
2258:
2257:
2255:
2254:
2249:
2244:
2238:
2236:
2232:
2231:
2229:
2228:
2221:
2214:
2207:
2200:
2199:
2198:
2191:
2184:
2177:
2170:
2163:
2151:
2150:
2149:
2142:
2135:
2128:
2115:
2113:
2109:
2108:
2105:
2104:
2102:
2101:
2094:
2091:Russellosaurus
2087:
2079:
2077:
2071:
2070:
2068:
2067:
2060:
2053:
2044:
2042:
2036:
2035:
2033:
2032:
2025:
2022:Pannoniasaurus
2017:
2015:
2009:
2008:
2005:
2004:
2002:
2001:
1994:
1987:
1980:
1972:
1970:
1964:
1963:
1961:
1960:
1953:
1946:
1939:
1931:
1929:
1920:
1919:
1912:
1905:
1897:
1895:
1886:
1885:
1877:
1875:
1869:
1868:
1865:
1864:
1862:
1861:
1854:
1847:
1839:
1837:
1828:
1827:
1819:
1817:
1811:
1810:
1803:
1801:
1798:
1797:
1795:
1794:
1787:
1780:
1773:
1766:
1763:Bentiabasaurus
1758:
1756:
1750:
1749:
1747:
1746:
1739:
1731:
1729:
1723:
1722:
1720:
1719:
1712:
1709:Igdamanosaurus
1705:
1698:
1691:
1683:
1681:
1672:
1671:
1664:
1657:
1650:
1643:
1636:
1629:
1622:
1615:
1608:
1600:
1598:
1592:
1591:
1589:
1588:
1582:
1576:
1570:
1563:
1560:
1559:
1552:
1551:
1544:
1537:
1529:
1521:
1520:
1509:(2): 147–150.
1489:
1461:
1435:
1369:
1368:
1366:
1363:
1298:
1295:
1247:
1244:
1240:
1239:
1236:
1235:
1231:
1230:
1227:
1226:
1222:
1221:
1218:
1217:
1213:
1212:
1209:
1208:
1204:
1203:
1200:
1199:
1195:
1194:
1191:
1190:
1186:
1185:
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1181:
1177:
1176:
1173:
1172:
1168:
1167:
1164:
1163:
1159:
1158:
1155:
1154:
1150:
1149:
1146:
1145:
1141:
1140:
1137:
1136:
1132:
1131:
1128:
1127:
1123:
1122:
1119:
1118:
1114:
1113:
1110:
1109:
1102:
1099:
1098:
1095:
1094:
1087:
1082:
1080:
1077:
1076:
1073:
1072:
1065:
1060:
1058:
1055:
1054:
1051:
1050:
1041:
1036:
1034:
1031:
1030:
1027:
1026:
1016:
1011:
1009:
1006:
1005:
1002:
1001:
997:
996:
993:
992:
976:
973:
972:
969:
968:
959:
954:
952:
947:
945:
942:
941:
938:
937:
928:
923:
921:
918:
917:
914:
913:
904:
899:
897:
894:
893:
890:
889:
885:
884:
881:
880:
873:
870:
869:
866:
865:
858:
853:
851:
846:
844:
841:
840:
837:
836:
829:
824:
822:
819:
818:
815:
814:
810:
809:
806:
805:
798:
795:
794:
791:
790:
780:
775:
773:
768:
766:
763:
762:
759:
758:
754:
753:
750:
749:
745:
744:
741:
740:
736:
735:
732:
731:
724:
721:
720:
717:
716:
709:
704:
702:
699:
698:
695:
694:
691:moorevillensis
684:
679:
677:
674:
673:
670:
669:
662:
657:
655:
652:
651:
648:
647:
640:
635:
633:
630:
629:
626:
625:
616:
611:
609:
606:
605:
602:
601:
594:
591:
590:
587:
586:
576:
571:
569:
566:
565:
562:
561:
552:
547:
545:
537:
491:member of the
470:
469:Classification
467:
417:
414:
335:
332:
303:
302:
295:
285:
284:
278:
277:
256:
252:
251:
243:
239:
238:
230:
226:
225:
217:
210:
209:
204:
200:
199:
194:
190:
189:
184:
180:
179:
174:
170:
169:
164:
160:
159:
146:
145:
137:
136:
128:
127:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
39:
26:
9:
6:
4:
3:
2:
2388:
2377:
2374:
2372:
2369:
2367:
2364:
2362:
2359:
2357:
2354:
2353:
2351:
2334:
2329:
2325:
2321:
2316:
2312:
2307:
2301:
2297:
2296:
2294:
2292:
2291:Megapterygius
2288:
2284:
2279:
2269:
2259:
2253:
2250:
2248:
2245:
2243:
2240:
2239:
2237:
2233:
2227:
2226:
2222:
2220:
2219:
2215:
2213:
2212:
2208:
2206:
2205:
2201:
2197:
2196:
2195:Tetrapodophis
2192:
2190:
2189:
2185:
2183:
2182:
2178:
2176:
2175:
2171:
2169:
2168:
2167:Dolichosaurus
2164:
2162:
2161:
2157:
2156:
2155:
2152:
2148:
2147:
2143:
2141:
2140:
2136:
2134:
2133:
2129:
2127:
2126:
2125:Aigialosaurus
2122:
2121:
2120:
2117:
2116:
2114:
2110:
2100:
2099:
2098:Yaguarasaurus
2095:
2093:
2092:
2088:
2086:
2085:
2081:
2080:
2078:
2076:
2072:
2066:
2065:
2061:
2059:
2058:
2057:Taniwhasaurus
2054:
2052:
2051:
2046:
2045:
2043:
2041:
2037:
2031:
2030:
2026:
2024:
2023:
2019:
2018:
2016:
2014:
2013:Tethysaurinae
2010:
2000:
1999:
1995:
1993:
1992:
1988:
1986:
1985:
1984:Goronyosaurus
1981:
1979:
1978:
1974:
1973:
1971:
1969:
1965:
1959:
1958:
1954:
1952:
1951:
1947:
1945:
1944:
1940:
1938:
1937:
1933:
1932:
1930:
1928:
1924:
1918:
1917:
1913:
1911:
1910:
1906:
1904:
1903:
1899:
1898:
1896:
1894:
1890:
1884:
1883:
1879:
1878:
1876:
1874:
1870:
1860:
1859:
1855:
1853:
1852:
1848:
1846:
1845:
1841:
1840:
1838:
1836:
1832:
1826:
1825:
1821:
1820:
1818:
1816:
1812:
1807:
1793:
1792:
1788:
1786:
1785:
1781:
1779:
1778:
1774:
1772:
1771:
1767:
1765:
1764:
1760:
1759:
1757:
1755:
1751:
1745:
1744:
1743:Thalassotitan
1740:
1738:
1737:
1733:
1732:
1730:
1728:
1724:
1718:
1717:
1713:
1711:
1710:
1706:
1704:
1703:
1702:Harranasaurus
1699:
1697:
1696:
1692:
1690:
1689:
1685:
1684:
1682:
1680:
1676:
1670:
1669:
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1662:
1658:
1656:
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1654:Megapterygius
1651:
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1479:(in Japanese)
1478:
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1465:
1457:
1456:
1448:
1442:
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1431:
1427:
1423:
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1399:
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1387:
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1383:
1381:
1379:
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1375:
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1362:
1360:
1359:Megapterygius
1356:
1352:
1351:Megapterygius
1348:
1347:
1342:
1341:
1336:
1332:
1331:
1326:
1325:
1320:
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1314:
1310:
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1304:
1294:
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1291:Megapterygius
1288:
1283:
1282:Megapterygius
1279:
1278:
1273:
1269:
1268:Megapterygius
1265:
1261:
1257:
1253:
1252:Megapterygius
1238:
1237:
1229:
1228:
1220:
1219:
1211:
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1202:
1201:
1193:
1192:
1184:
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1047:
1040:
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1033:
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1025:
1024:
1023:missouriensis
1022:
1015:
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1007:
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1003:
995:
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991:
988:
983:
982:
975:
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965:
958:
957:
951:
950:
944:
943:
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939:
936:
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933:Megapterygius
927:
926:
920:
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916:
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902:
896:
895:
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513:
508:
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485:Megapterygius
479:
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466:
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463:Megapterygius
460:
456:
455:Megapterygius
452:
448:
444:
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439:Megapterygius
436:
435:neural spines
431:
429:
428:
422:
421:Megapterygius
413:
411:
407:
406:Megapterygius
402:
400:
399:wakayamaensis
396:
395:specific name
392:
391:Megapterygius
388:
384:
381:
380:Ancient Greek
377:
376:Megapterygius
373:
369:
366:
363:
358:
356:
352:
348:
344:
341:
340:Megapterygius
331:
329:
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315:
311:
310:
309:Megapterygius
299:
293:
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286:
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279:
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265:
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263:Megapterygius
257:
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172:
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168:
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147:
143:
142:Megapterygius
138:
134:
129:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
54:
47:
46:Maastrichtian
43:
37:
36:Megapterygius
33:
30:
19:
2356:Mosasaurines
2290:
2223:
2216:
2209:
2202:
2193:
2186:
2179:
2172:
2165:
2158:
2146:Opetiosaurus
2144:
2137:
2130:
2123:
2096:
2089:
2082:
2062:
2055:
2048:
2040:Tylosaurinae
2027:
2020:
1996:
1989:
1982:
1977:Gavialimimus
1975:
1968:Selmasaurini
1955:
1948:
1941:
1934:
1916:Sarabosaurus
1914:
1909:Ectenosaurus
1907:
1902:Angolasaurus
1900:
1882:Haasiasaurus
1880:
1856:
1849:
1842:
1822:
1815:Halisaurinae
1789:
1782:
1775:
1770:Eremiasaurus
1768:
1761:
1741:
1736:Prognathodon
1734:
1714:
1707:
1700:
1693:
1686:
1679:Globidensini
1666:
1659:
1653:
1652:
1645:
1638:
1631:
1626:Gnathomortis
1624:
1617:
1610:
1605:Amphekepubis
1603:
1596:Mosasaurinae
1506:
1502:
1492:
1481:. Retrieved
1473:
1464:
1453:
1397:
1393:
1358:
1350:
1344:
1338:
1328:
1324:Apiotrigonia
1322:
1316:
1306:
1300:
1297:Paleoecology
1290:
1286:
1281:
1275:
1267:
1251:
1249:
1246:Paleobiology
1105:
1104:
1090:
1089:
1068:
1067:
1044:
1043:
1019:
1018:
986:
979:
978:
962:
961:
932:
931:
930:
907:
906:
876:
875:
861:
860:
832:
831:
801:
800:
783:
782:
727:
726:
712:
711:
687:
686:
665:
664:
643:
642:
621:Eremiasaurus
619:
618:
597:
596:
581:Prognathodon
579:
578:
555:
554:
543:Mosasaurinae
522:
516:
510:
504:
501:sister taxon
497:Mosasauridae
484:
482:
462:
454:
438:
432:
425:
420:
419:
405:
403:
398:
390:
386:
382:
375:
372:generic name
367:
361:
359:
339:
337:
326:
325:
308:
307:
306:
297:
290:
289:
282:Type species
269:
262:
261:
249:Mosasaurinae
236:Mosasauridae
213:
141:
35:
29:
2211:Judeasaurus
2204:Adriosaurus
2188:Pontosaurus
2160:Coniasaurus
2139:Komensaurus
2132:Carsosaurus
2084:Romeosaurus
2029:Tethysaurus
1998:Selmasaurus
1943:Platecarpus
1835:Halisaurini
1791:Plotosaurus
1777:Moanasaurus
1754:Mosasaurini
1633:Jormungandr
1619:Dallasaurus
1301:Fossils of
1046:Plotosaurus
987:Moanasaurus
964:Moanasaurus
557:Dallasaurus
524:Plotosaurus
506:Moanasaurus
489:mosasaurine
416:Description
318:mosasaurine
242:Subfamily:
2350:Categories
2306:Q123766172
2181:Primitivus
2064:Tylosaurus
2050:Kaikaifilu
1851:Halisaurus
1784:Mosasaurus
1688:Carinodens
1668:Stelladens
1640:Kourisodon
1483:2023-12-13
1476:和歌山県立自然博物館
1455:和歌山県立自然博物館
1365:References
1355:squaliform
1335:gastropods
1333:sp.), and
1330:Inoceramus
1021:Mosasaurus
981:Rikisaurus
787:dakotensis
521:spp., and
518:Mosasaurus
512:Rikisaurus
451:dorsal fin
447:delphinoid
427:Tylosaurus
351:Mt. Toyajo
345:specimen,
223:Mosasauria
2174:Kaganaias
1991:Khinjaria
1844:Eonatator
1824:Pluridens
1695:Globidens
1612:Clidastes
1566:Kingdom:
1556:Mosasaurs
1430:266284597
1422:1477-2019
1318:Nanonavis
1303:ammonites
785:Globidens
689:Clidastes
533:cladogram
499:, as the
443:posterior
441:holotype
387:pterygion
365:described
316:genus of
173:Kingdom:
167:Eukaryota
2320:11934700
2300:Wikidata
2268:Category
1716:Xenodens
1586:Squamata
1580:Reptilia
1574:Chordata
1572:Phylum:
1568:Animalia
1327:sp. and
1313:bivalves
1264:proximal
493:squamate
459:rib cage
343:holotype
296:Konishi
268:Konishi
229:Family:
207:Squamata
197:Reptilia
187:Chordata
183:Phylum:
177:Animalia
163:Domain:
140:Size of
51:72
1584:Order:
1578:Class:
1458:. 2023.
1402:Bibcode
535:below:
437:of the
314:extinct
300:., 2023
255:Genus:
203:Order:
193:Class:
120:↓
2333:483187
1647:Liodon
1428:
1420:
1287:et al.
583:currii
495:clade
393:. The
362:et al.
2315:IRMNG
1450:(PDF)
1426:S2CID
1400:(1).
1321:sp.,
487:as a
383:mégas
298:et al
270:et al
214:Clade
1503:地球科学
1418:ISSN
1258:and
433:The
347:WMNH
338:The
274:2023
59:PreꞒ
1511:doi
1410:doi
1311:),
374:, "
353:in
272:.,
48:),
2352::
2330::
2317::
2302::
1507:75
1505:.
1501:.
1472:.
1452:.
1438:^
1424:.
1416:.
1408:.
1398:21
1396:.
1373:^
1343:,
515:,
509:,
430:.
216::
109:Pg
53:Ma
2047:?
1548:e
1541:t
1534:v
1517:.
1513::
1486:.
1432:.
1412::
1404::
1337:(
1315:(
1305:(
989:)
985:(
410:滄
397:"
288:†
259:†
246:†
233:†
220:†
114:N
104:K
99:J
94:T
89:P
84:C
79:D
74:S
69:O
64:Ꞓ
44:(
20:)
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