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Mandragora (genus)

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species have a long use in traditional medicine, extracts being used for their real or supposed aphrodisiac, hypnotic, emetic, purgative, sedative, and pain-killing effects. Tropane alkaloids are known to be effective as analgesics and anaesthetics, and can be used to increase circulation and dilate
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species. More than 80 substances have been identified; their paper gives the detailed chemical structure of 37 of them. The different parts of the plant contain different proportions and concentrations of alkaloids, with the roots having the highest concentrations. Alkaloids present include
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Tuttolomondo, Teresa; Licata, Mario; Leto, Claudio; Savo, Valentina; Bonsangue, Giuseppe; Gargano, Maria Letizia; Venturella, Giuseppe & La Bella, Salvatore (2014), "Ethnobotanical investigation on wild medicinal plants in the Monti Sicani Regional Park (Sicily, Italy)",
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Tu, Tieyao; Volis, Sergei; Dillon, Michael O.; Sun, Hang & Wen, Jun (2010). "Dispersals of Hyoscyameae and Mandragoreae (Solanaceae) from the New World to Eurasia in the early Miocene and their biogeographic diversification within Eurasia".
491:) suggest that there are two clades in the genus - one based in the Mediterranean and beyond to Turkmenistan and Iran, and one in the Sino-Himalayan region. A simplified cladogram based on these studies is shown below. In one of the studies, 1002:
alkaloids and the sometimes vaguely humanoid shape of their roots have led to mandrakes being associated with a variety of myths and superstitious practices throughout history. However, the plants used in this way are not always species of
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are perennial herbaceous plants. They have large vertical tap roots, sometimes forked. Their stems are short or virtually absent. The leaves form a rosette at the base of the plant. The flowers are sometimes borne on a short stalk
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suggest that the genus belongs in the large subfamily Solanoideae, but that within this subfamily, it is one of a number of isolated genera with no immediate relatives. It has thus been placed in its own tribe, Mandragoreae.
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Jiménez-Mejías, M.E.; Montaño-Díaz, M.; López Pardo, F.; Campos Jiménez, E.; Martín Cordero, M.C.; Ayuso González, M.J. & González de la Puente, M.A. (1990-11-24), "Intoxicación atropínica por
834:). Stems sometimes present; petals dark purple or yellow; berry globose. Considerably variable in size and appearance, possibly justifying dividing the taxon into subspecies or even species. 1469:
Särkinen, Tiina; Bohs, Lynn; Olmstead, Richard G. & Knapp, Sandra (2013), "A phylogenetic framework for evolutionary study of the nightshades (Solanaceae): a dated 1000-tip tree",
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Olmstead, Richard G.; Bohs, Lynn; Migid, Hala Abdel; Santiago-Valentin, Eugenio; Garcia, Vicente F. & Collier, Sarah M. (2008), "A molecular phylogeny of the Solanaceae",
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and their scientific names. Dioscorides was among those who distinguished between "male" and "female" mandrakes, a distinction used in 1764 when Garsault published the names
418:. Using statistical analysis of morphological characters, Ungricht et al. found no distinct clusters among the specimens they examined and concluded that Linnaeus's 914: 728:. Virtually stemless; petals 1.2–6.5 cm (0.5–2.6 in) long, greenish white through blue to violet; berry globose to ovoid, yellow to orange when ripe. 1998:
Shengji, Pei; Huyin, Huai & Lixin, Yang (2009), "Medicinal Plants and Their Conservation in China with Reference to the Chinese Himalayan Region",
384:. The size and shape of the fruit and the colour and time of appearance of the flowers have been used to distinguish possible species. In the 1820s, 2238: 2277: 372:
Jackson and Berry (1979) and Ungricht et al. (1998) have documented the subsequent confusion over the number of Mediterranean species of
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Hawkes, J.G. (1972). "Mandragora". In Tutin, T.G.; Heywood, V.H.; Burges, N.A.; Valentine, D.H.; Walters, S.M. & Webb, D.A. (eds.).
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confined to northern Italy and parts of the coast of former Yugoslavia. Some more recent sources distinguish plants found in the
677: 1086:""Hexing Herbs" in Ethnobotanical Perspective: A Historical Review of the Uses of Anticholinergic Solanaceae Plants in Europe" 431:
differently. Plants from the western Mediterranean, from Turkey westwards to the Iberian peninsula and Morocco, are placed in
1176: 2256: 1918:"Common anticholinergic solanaceaous plants of temperate Europe - A review of intoxications from the literature (1966–2018)" 1724:"Common anticholinergic solanaceaous plants of temperate Europe - A review of intoxications from the literature (1966–2018)" 1032:"Common anticholinergic solanaceaous plants of temperate Europe - A review of intoxications from the literature (1966–2018)" 1192: 949:) include severe symptoms similar to those of atropine poisoning, including blurred vision, dilation of the pupils ( 953:), dryness of the mouth, difficulty in urinating, dizziness, headache, vomiting, blushing, and a rapid heart rate ( 2282: 2225: 2321: 2264: 660:
through the agency of birds, with the main split between the species occurring around 10 million years ago.
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pupils, among other effects. Hyoscine and anisodamine are used medicinally in China. Continued use of
877: 849: 2204: 1534: 1239: 1224: 780:). Stemless; petals 2.3–2.7 cm (0.9–1.1 in) long, violet; berry yellow, strongly aromatic. 252:. Individual flowers are bell-shaped, whitish through to violet, and followed by yellow or orange 2308: 2173: 1917: 1723: 1031: 827: 464: 1878:
Piccillo, Giovita A.; Mondati, Enrico G. M. & Moro, Paola A. (2002), "Six clinical cases of
1772:; Řezanka, Tomáš; Spížek, Jaroslav & Dembitsky, Valery M. (2005), "Substances isolated from 2160: 2051: 761: 487: 347: 48: 2243: 463:, being grouped together in the same tribe or subtribe as at least the first of these genera. 910: 784: 704: 481: 2295: 2370: 2134: 2089: 684:. Three species are accepted in a 1998 review of the genus and by GRIN. Other sources keep 268: 8: 2380: 1658: 1013:) is explicitly mentioned as the source of a "mandrake" or "mandragora" in some sources. 230: 1308: 2342: 1945: 1831: 1810:
Razzakov, N. A.; Aripova, S. F.; Akhmedova, E. & Karimov, A. (1998), "Alkaloids of
1751: 1612: 1493: 1212: 1113: 1059: 56: 1085: 2290: 2121: 2031: 1981: 1949: 1937: 1899: 1895: 1861: 1793: 1755: 1743: 1498: 1422: 1276: 1172: 1117: 1105: 1063: 1051: 777: 385: 341: 1835: 1789: 1696: 1616: 1136: 2347: 2126: 2007: 1973: 1929: 1891: 1847: 1823: 1785: 1735: 1604: 1488: 1478: 1452: 1414: 1097: 1043: 873: 249: 241: 2100: 1933: 1739: 1047: 238: 99: 2269: 1418: 1101: 999: 957:). Hyperactivity and hallucinations also occurred in the majority of patients. 881: 753:, one difference being that the seeds are more than twice as large as those of 673: 402: 352: 309: 301: 285: 253: 86: 1977: 1608: 475:
of the Mediterranean mandrakes. Two studies that separate plants found in the
2364: 2074: 2035: 2012: 1483: 1109: 717: 435:; plants from the eastern Mediterranean, from Syria to Israel, are placed in 336: 300:
are shorter than the petals, joined to the floral tube towards the base. The
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that make the plants poisonous. Their roots in particular have a long use in
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is suggested to have originated around 20 million years ago, arriving in
234: 155: 1630: 1563: 2191: 1827: 1456: 945: 427: 237:. Two or three further species are found eastwards into China. All are 218: 145: 2217: 936:
Clinical reports of the effects of consumption of plants described as
396:, the autumn-flowering mandrake. Identifying the former as Linnaeus's 2147: 1508: 995: 991:) has been listed as a Chinese medicinal plant needing conservation. 950: 649: 135: 2045: 2329: 2068: 890: 853: 713: 669: 264: 245: 222: 125: 112: 2152: 1442: 355:
of the genus. (Linnaeus later changed his mind and in 1759 placed
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are placed in the genus. The one or two species found around the
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Ungricht, Stefan; Knapp, Sandra & Press, John R. (1998).
1286:. In Hawkes, J.G.; Lester, R.N. & Skelding, A.D. (eds.). 1135:. In Wu, Zhengyi; Raven, Peter H. & Hong, Deyuan (eds.). 811: 807: 721: 668:
As of March 2015, major online plant databases (such as
293: 289: 214: 211: 73: 1768: 773: 1962: 1185: 485:) from those found in the rest of the Mediterranean area ( 271:. Mandrakes are involved in many myths and superstitions. 1514: 1468: 737:
or considered a separate species. Older sources consider
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Akhania, Hossein & Ghorbanib, Abdol-Basset (2003). "
308:). After fertilization, a yellow or orange fruit forms ( 1169:
Flora Europaea, Volume 3: Diapensiaceae to Myoporaceae
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Bulletin of the Natural History Museum, Botany Series
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to the main species found in the Mediterranean with
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is a single, variable species. Other sources divide
288:), and are solitary, with whorls of five parts. The 1877: 1691:Zhang, Zhi-Yun; Lu, Anmin & D'Arcy, William G. 1623: 1131:Zhang, Zhi-Yun; Lu, Anmin & D'Arcy, William G. 680:) accept different numbers of species in the genus 233:constitute the mandrake of ancient writers such as 1956: 1841: 1803: 1762: 1529: 1527: 1525: 1523: 1462: 1433: 1399: 1397: 1395: 1393: 1391: 1389: 1387: 1385: 1354: 1275:Jackson, Betty P. & Berry, Michael I. (1979). 1997: 1281:- taxonomy and chemistry of the European species" 2362: 2030:, trans. Ross Nichols, New York: Citadel Press, 1590: 1403: 1171:. Cambridge University Press. pp. 199–200. 868:Like many members of the Solanaceae, species of 259:Like many members of the Solanaceae, species of 1520: 1382: 1079: 1077: 1075: 1073: 442: 1991: 1871: 1651: 1301: 872:contain highly biologically active alkaloids, 471:Within the genus, studies have used different 1690: 1130: 1124: 451:has been considered to be closely related to 1274: 1270: 1268: 1266: 1264: 1262: 1260: 1070: 388:used flowering time to name two species as 1916:Fatur, Karsten; Kreft, Samo (April 2020). 1915: 1722:Fatur, Karsten; Kreft, Samo (April 2020). 1721: 1584: 1290:. London: Academic Press. pp. 505–512 1288:The biology and taxonomy of the Solanaceae 1030:Fatur, Karsten; Kreft, Samo (April 2020). 1029: 38: 2025: 2011: 1686: 1684: 1682: 1492: 1482: 1257: 296:, both in the shape of a lobed bell. The 1350: 1348: 1346: 1344: 1342: 1340: 1338: 1336: 1334: 1332: 1237: 848: 319: 2363: 2019: 1884:European Journal of Emergency Medicine 1679: 1166: 2050: 2049: 1882:poisoning: diagnosis and treatment", 1407:Molecular Phylogenetics and Evolution 1329: 1231: 1083: 392:, the spring-flowering mandrake, and 221:). Members of the genus are known as 1162: 1160: 1158: 406:listed two Mediterranean species of 217:belonging to the nightshade family ( 292:are joined at the base, as are the 263:contain highly biologically active 13: 1194:IPNI Plant Name Query Results for 14: 2392: 2028:The History and Practice of Magic 1852:: descripción de quince casos ", 1155: 979:in folk medicine was reported in 351:was described, which is thus the 1896:10.1097/00063110-200212000-00010 345:where the Mediterranean species 60: 2322:urn:lsid:ipni.org:names:38432-1 1909: 1790:10.1016/j.phytochem.2005.07.016 1715: 818:and parts of China (south-east 648:The Solanaceae are primarily a 1816:Chemistry of Natural Compounds 1023: 465:Molecular phylogenetic studies 274: 1: 1934:10.1016/j.toxicon.2020.02.005 1740:10.1016/j.toxicon.2020.02.005 1048:10.1016/j.toxicon.2020.02.005 1016: 929:are also reported to contain 1966:Journal of Ethnopharmacology 1597:Systematics and Biodiversity 1084:Fatur, Karsten (June 2020). 443:Classification and phylogeny 248:and leaves in the form of a 196: 7: 1419:10.1016/j.ympev.2010.09.007 1196:Mandragora officinarum 838: 315: 10: 2397: 1102:10.1007/s12231-020-09498-w 964: 842: 663: 335:was first used in 1753 by 18: 2058: 1978:10.1016/j.jep.2014.02.032 1609:10.1017/S1477200003001105 1357:"A revision of the genus 1244:Species Plantarum, vol. 1 919:3α,6β-ditigloyloxytropane 614: 599: 592: 568: 544: 529: 522: 515: 508: 225:. Between three and five 194: 189: 57:Scientific classification 55: 46: 37: 30: 2026:Christian, Paul (1963), 2013:10.1163/157342109X568810 1543:GRIN Taxonomy for Plants 1484:10.1186/1471-2148-13-214 1471:BMC Evolutionary Biology 967:Mandrake § Folklore 856:is a mixture of D and L 845:Mandrake § Toxicity 793:Mandragora chinghaiensis 678:GRIN Taxonomy for Plants 617:Mandragora chinghaiensis 339:in the first edition of 1693:"Mandragora caulescens" 1007:; for example, bryony ( 960: 733:may be included within 1812:Mandragora turcomanica 1593:Mandragora turcomanica 1566:Mandragora turcomanica 1515:Särkinen et al. (2013) 1311:Mandragora officinarum 865: 762:Mandragora turcomanica 697:Mandragora officinarum 571:Mandragora officinarum 532:Mandragora turcomanica 488:Mandragora officinarum 348:Mandragora officinarum 328: 325:Mandragora officinarum 49:Mandragora officinarum 1880:Mandragora autumnalis 1850:Mandragora autumnalis 1633:Mandragora caulescens 1238:Linnaeus, C. (1753). 989:M. chinghaiensis 860:, the L isomer being 852: 785:Mandragora caulescens 751:Mandragora autumnalis 712:Central and southern 705:Mandragora autumnalis 692:as separate species. 690:M. chinghaiensis 602:Mandragora caulescens 547:Mandragora autumnalis 493:M. chinghaiensis 482:Mandragora autumnalis 394:Mandragora autumnalis 323: 1659:"Search results for 1535:"Search results for 915:3α-tigloyloxytropane 880:et al. reviewed the 495:was embedded within 269:traditional medicine 19:For other uses, see 1517:, additional file 2 927:M. turcomanica 800:Mandragora tibetica 755:M. officinarum 743:M. officinarum 735:M. officinarum 720:area, eastwards to 716:and throughout the 433:M. officinarum 424:M. officinarum 420:M. officinarum 412:M. officinarum 398:M. officinarum 390:Mandragora vernalis 357:M. officinarum 327:with violet flowers 310:botanically a berry 1828:10.1007/BF02336107 1457:10.1002/tax.574010 985:M. caulescens 977:M. autumnalis 923:M. caulescens 866: 796:Kuang & A.M.Lu 739:M. autumnalis 731:M. autumnalis 686:M. autumnalis 497:M. caulescens 437:M. autumnalis 416:M. autumnalis 382:Mandragora foemina 329: 304:has two chambers ( 2376:Solanaceae genera 2358: 2357: 2291:Open Tree of Life 2052:Taxon identifiers 1784:(20): 2408–2417, 1178:978-0-521-08489-5 874:tropane alkaloids 804: 797: 790: 778:Golestan Province 767: 710: 701: 645: 644: 636: 635: 627: 626: 581: 580: 557: 556: 386:Antonio Bertoloni 342:Species Plantarum 242:herbaceous plants 203: 202: 185: 2388: 2351: 2350: 2338: 2337: 2325: 2324: 2312: 2311: 2299: 2298: 2286: 2285: 2273: 2272: 2270:NHMSYS0020823397 2260: 2259: 2247: 2246: 2234: 2233: 2221: 2220: 2208: 2207: 2195: 2194: 2182: 2181: 2169: 2168: 2156: 2155: 2143: 2142: 2130: 2129: 2117: 2116: 2104: 2103: 2094: 2093: 2092: 2079: 2078: 2077: 2047: 2046: 2040: 2038: 2023: 2017: 2016: 2015: 1995: 1989: 1988: 1960: 1954: 1953: 1913: 1907: 1906: 1875: 1869: 1868: 1854:Medicina Clínica 1845: 1839: 1838: 1807: 1801: 1800: 1766: 1760: 1759: 1719: 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1806: 1799: 1795: 1791: 1787: 1783: 1779: 1775: 1771: 1765: 1757: 1753: 1749: 1745: 1741: 1737: 1733: 1729: 1725: 1718: 1704:. eFloras.org 1703: 1702: 1699: 1694: 1687: 1685: 1683: 1668: 1664: 1662: 1654: 1640: 1636: 1634: 1626: 1618: 1614: 1610: 1606: 1602: 1598: 1594: 1587: 1573: 1569: 1567: 1559: 1544: 1540: 1538: 1530: 1528: 1526: 1524: 1516: 1511: 1504: 1500: 1495: 1490: 1485: 1480: 1476: 1472: 1465: 1458: 1454: 1450: 1446: 1439: 1437: 1428: 1424: 1420: 1416: 1412: 1408: 1400: 1398: 1396: 1394: 1392: 1390: 1388: 1386: 1370: 1366: 1362: 1361:(Solanaceae)" 1360: 1351: 1349: 1347: 1345: 1343: 1341: 1339: 1337: 1335: 1333: 1318: 1314: 1312: 1304: 1289: 1282: 1280: 1271: 1269: 1267: 1265: 1263: 1261: 1246:. p. 181 1245: 1241: 1234: 1226: 1214: 1200:. Vol. 1 1199: 1198: 1195: 1188: 1180: 1174: 1170: 1163: 1161: 1159: 1144:. eFloras.org 1143: 1142: 1139: 1134: 1127: 1119: 1115: 1111: 1107: 1103: 1099: 1095: 1091: 1087: 1080: 1078: 1076: 1074: 1065: 1061: 1057: 1053: 1049: 1045: 1041: 1037: 1033: 1026: 1022: 1014: 1012: 1011: 1006: 1001: 997: 992: 990: 986: 982: 978: 973: 968: 958: 956: 952: 948: 947: 943: 939: 938:M. autumnalis 934: 932: 928: 924: 920: 916: 912: 908: 904: 900: 896: 892: 887: 883: 879: 875: 871: 863: 859: 855: 851: 846: 833: 830:, north-west 829: 825: 821: 817: 813: 809: 801: 794: 787: 786: 782: 779: 775: 771: 764: 763: 759: 756: 752: 748: 744: 740: 736: 732: 727: 723: 719: 718:Mediterranean 715: 707: 706: 698: 695: 694: 693: 691: 687: 683: 679: 675: 671: 661: 659: 655: 651: 641: 640: 632: 631: 623: 622: 619: 618: 612: 611: 608: 607: 604: 603: 597: 596: 590: 589: 586: 585: 577: 576: 573: 572: 566: 565: 562: 561: 553: 552: 549: 548: 542: 541: 538: 537: 534: 533: 527: 526: 520: 519: 513: 512: 506:Mandragoreae 505: 504: 500: 498: 494: 490: 489: 484: 483: 478: 474: 469: 466: 462: 461: 456: 455: 450: 440: 438: 434: 430: 429: 425: 421: 417: 413: 409: 405: 404: 399: 395: 391: 387: 383: 379: 375: 370: 368: 367:A. mandragora 364: 363: 359:in the genus 358: 354: 350: 349: 344: 343: 338: 337:Carl Linnaeus 334: 326: 322: 313: 311: 307: 303: 299: 295: 291: 287: 282: 272: 270: 266: 262: 257: 255: 251: 247: 244:, with large 243: 240: 236: 232: 231:Mediterranean 228: 224: 220: 216: 213: 209: 208: 199: 198: 193: 188: 184: 179: 178: 174: 171: 170: 167: 164: 161: 160: 157: 154: 151: 150: 147: 144: 141: 140: 137: 134: 131: 130: 127: 124: 121: 118: 117: 114: 111: 108: 105: 104: 101: 98: 95: 92: 91: 88: 87:Tracheophytes 85: 82: 79: 78: 75: 72: 69: 68: 63: 58: 54: 51: 50: 45: 41: 36: 33: 29: 26: 22: 2059: 2027: 2021: 2003: 1999: 1993: 1969: 1965: 1958: 1925: 1921: 1911: 1887: 1883: 1879: 1873: 1857: 1853: 1849: 1843: 1819: 1815: 1811: 1805: 1781: 1777: 1773: 1764: 1731: 1727: 1717: 1706:. Retrieved 1701: 1697: 1670:. Retrieved 1666: 1660: 1653: 1642:. Retrieved 1638: 1635:C.B. Clarke" 1632: 1625: 1600: 1596: 1592: 1586: 1575:. Retrieved 1571: 1565: 1558: 1547:. Retrieved 1542: 1536: 1510: 1474: 1470: 1464: 1448: 1444: 1410: 1406: 1373:. Retrieved 1368: 1364: 1358: 1320:. Retrieved 1316: 1310: 1303: 1292:. Retrieved 1287: 1278: 1248:. Retrieved 1243: 1240:"Mandragora" 1233: 1202:. Retrieved 1197: 1193: 1187: 1168: 1146:. Retrieved 1141: 1137: 1133:"Mandragora" 1126: 1093: 1089: 1039: 1035: 1025: 1008: 1004: 993: 988: 984: 976: 971: 970: 944: 942:M. offinarum 941: 937: 935: 926: 922: 913:(hyoscine), 903:cuscohygrine 899:belladonnine 885: 869: 867: 799: 792: 783: 770:Turkmenistan 760: 754: 750: 742: 738: 734: 730: 703: 696: 689: 685: 681: 667: 653: 647: 616: 615: 601: 600: 570: 569: 546: 545: 531: 530: 496: 492: 486: 480: 470: 458: 452: 448: 446: 436: 432: 426: 423: 419: 415: 411: 407: 401: 397: 393: 389: 381: 377: 373: 371: 366: 360: 356: 353:type species 346: 340: 332: 330: 324: 280: 278: 260: 258: 206: 205: 204: 195: 176: 175: 166:Mandragoreae 165: 119: 106: 93: 80: 47: 31: 25: 2371:Solanoideae 2213:iNaturalist 2084:Wikispecies 955:tachycardia 931:anisodamine 911:scopolamine 907:hyoscyamine 895:apoatropine 862:hyoscyamine 791:(including 279:Species of 275:Description 235:Dioscorides 156:Solanoideae 152:Subfamily: 100:Angiosperms 2381:Oneirogens 2365:Categories 2090:Mandragora 2060:Mandragora 1776:species", 1774:Mandragora 1708:2015-03-30 1672:2015-03-29 1661:Mandragora 1644:2015-03-29 1577:2015-03-29 1549:2015-03-29 1537:Mandragora 1477:(1): 214, 1375:2015-03-31 1371:(1): 17–40 1359:Mandragora 1322:2015-04-02 1294:2015-03-29 1279:Mandragora 1250:2015-03-29 1204:2015-03-29 1148:2015-03-30 1017:References 1005:Mandragora 972:Mandragora 946:sensu lato 886:Mandragora 870:Mandragora 843:See also: 789:C.B.Clarke 682:Mandragora 654:Mandragora 449:Mandragora 428:sensu lato 408:Mandragora 374:Mandragora 333:Mandragora 331:The genus 281:Mandragora 261:Mandragora 219:Solanaceae 207:Mandragora 177:Mandragora 146:Solanaceae 32:Mandragora 21:Mandragora 2036:301220772 1950:213559151 1928:: 52–88. 1756:213559151 1734:: 52–88. 1223:ignored ( 1213:cite book 1118:220844064 1110:0013-0001 1064:213559151 1042:: 52–88. 996:deliriant 983:in 2014. 951:mydriasis 652:family. 650:New World 265:alkaloids 246:tap roots 239:perennial 223:mandrakes 136:Solanales 70:Kingdom: 2335:40013146 2330:Tropicos 2069:Wikidata 1986:24632020 1942:32217234 1904:12501035 1836:27501840 1798:16137728 1748:32217234 1700:(online) 1639:Tropicos 1617:85903719 1572:Tropicos 1503:24283922 1427:20858548 1140:(online) 1056:32217234 891:atropine 854:Atropine 839:Toxicity 714:Portugal 670:Tropicos 316:Taxonomy 197:See text 190:Species 142:Family: 126:Asterids 113:Eudicots 2244:1101638 2231:38432-1 2192:2928728 2075:Q212742 1922:Toxicon 1866:2087109 1728:Toxicon 1494:3850475 1036:Toxicon 1010:Bryonia 858:isomers 826:, east 824:Sichuan 822:, west 820:Qinghai 709:Bertol. 664:Species 658:Eurasia 306:locules 298:stamens 254:berries 250:rosette 227:species 172:Genus: 162:Tribe: 132:Order: 74:Plantae 2304:PLANTS 2296:442498 2257:500625 2218:337455 2179:119600 2166:119600 2114:438840 2101:192388 2098:APDB: 2034:  1984:  1948:  1940:  1902:  1864:  1834:  1796:  1754:  1746:  1615:  1545:. USDA 1501:  1491:  1425:  1175:  1116:  1108:  1062:  1054:  981:Sicily 917:, and 832:Yunnan 816:Bhutan 803:Grubov 747:Levant 726:Jordan 676:, and 477:Levant 460:Lycium 454:Atropa 362:Atropa 294:petals 290:sepals 2309:MANDR 2283:24647 2239:IRMNG 2153:1MNDG 2140:61639 2127:62ZLN 1946:S2CID 1832:S2CID 1752:S2CID 1613:S2CID 1568:Mizg" 1445:Taxon 1284:(PDF) 1277:"39. 1114:S2CID 1060:S2CID 878:Hanuš 812:Nepal 808:India 766:Mizg. 722:Syria 302:ovary 286:scape 215:genus 212:plant 210:is a 120:Clade 107:Clade 94:Clade 81:Clade 2317:POWO 2278:NCBI 2252:ITIS 2226:IPNI 2205:7226 2200:GRIN 2187:GBIF 2148:EPPO 2109:BOLD 2032:OCLC 1982:PMID 1938:PMID 1900:PMID 1862:PMID 1794:PMID 1744:PMID 1499:PMID 1423:PMID 1225:help 1173:ISBN 1106:ISSN 1052:PMID 998:and 987:(as 961:Uses 925:and 774:Iran 724:and 702:and 688:and 457:and 414:and 380:and 2343:WFO 2265:NBN 2174:FoC 2161:FNA 2135:EoL 2122:CoL 2008:doi 1974:doi 1970:153 1930:doi 1926:177 1892:doi 1824:doi 1814:", 1786:doi 1736:doi 1732:177 1605:doi 1489:PMC 1479:doi 1453:doi 1415:doi 1098:doi 1044:doi 1040:177 884:of 749:as 369:.) 365:as 312:). 2367:: 2345:: 2332:: 2319:: 2306:: 2293:: 2280:: 2267:: 2254:: 2241:: 2228:: 2215:: 2202:: 2189:: 2176:: 2163:: 2150:: 2137:: 2124:: 2111:: 2086:: 2071:: 2002:, 1980:, 1968:, 1944:. 1936:. 1924:. 1920:. 1898:, 1886:, 1858:95 1856:, 1830:, 1820:34 1818:, 1792:, 1782:66 1780:, 1750:. 1742:. 1730:. 1726:. 1681:^ 1665:. 1637:. 1611:. 1599:. 1570:. 1541:. 1522:^ 1497:, 1487:, 1475:13 1473:, 1449:57 1447:, 1435:^ 1421:. 1411:57 1409:. 1384:^ 1369:28 1367:. 1363:. 1331:^ 1315:. 1259:^ 1242:. 1217:: 1215:}} 1211:{{ 1157:^ 1112:. 1104:. 1094:74 1092:. 1088:. 1072:^ 1058:. 1050:. 1038:. 1034:. 933:. 921:. 909:, 905:, 901:, 897:, 893:, 814:, 810:, 798:, 772:, 700:L. 672:, 499:. 439:. 410:: 256:. 183:L. 122:: 109:: 96:: 83:: 2010:: 2004:5 1976:: 1952:. 1932:: 1894:: 1888:9 1826:: 1788:: 1758:. 1738:: 1711:. 1675:. 1663:" 1647:. 1631:" 1619:. 1607:: 1601:1 1580:. 1564:" 1552:. 1539:" 1481:: 1455:: 1429:. 1417:: 1378:. 1325:. 1313:" 1309:" 1297:. 1253:. 1227:) 1207:. 1181:. 1151:. 1120:. 1100:: 1066:. 1046:: 940:( 864:. 805:) 776:( 757:. 479:( 284:( 23:.

Index

Mandragora

Mandragora officinarum
Scientific classification
Edit this classification
Plantae
Tracheophytes
Angiosperms
Eudicots
Asterids
Solanales
Solanaceae
Solanoideae
Mandragoreae
Mandragora
L.
See text
plant
genus
Solanaceae
mandrakes
species
Mediterranean
Dioscorides
perennial
herbaceous plants
tap roots
rosette
berries
alkaloids

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