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Layia carnosa

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movement on a periodic basis support early successional communities. Movement of sand by wind is essential for the development and sustainability of a dune system. Wind is also important to beach layia specifically because it is the mechanism by which seeds are dispersed. The achenes (a small, dry, one-seeded fruit that does not open to release the seed) have pappus (feathery bristles) that allow them to be carried by wind for a short distance. Although not all seeds may land on suitable habitat, this adaptation allows the small annual to spread across the landscape into uninhabited areas. As a winter germinating annual, beach layia requires rainfall during the winter months (November through February) for germination and, although it is relatively tolerant to the droughtlike conditions of upland dunes, it does need some moisture through the spring to prevent desiccation. Moisture also reduces the risk of burial, as dry sand is more mobile and mortality caused by burial has been documented. The overall resource needs that beach layia requires in order for individuals to complete their life cycles and for populations to maintain viability are: (1) Sandy soils with sparse native vegetation cover, (2) Rainfall during the winter germination period, (3) Sunlight (full sun exposure for photosynthesis), and (4) Unknown degree of crosspollination (to add to genetic diversity).
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associated with the shifts in dune blowouts, remobilization, and natural dune stabilization that occur in the coastal dune ecosystem. Beach layia plants often occur where sparse vegetation traps wind-dispersed seeds, but causes minimal shading. Seeds are dispersed by wind mostly during late spring and summer months. Additionally, beach layia is self-compatible (i.e., able to be fertilized by its own pollen), is capable of self-pollination, and is visited by a variety of insects that may assist in cross-pollination . Although the role of pollinators is currently unclear, sexual reproduction does add to genetic diversity. Beach layia occurs in open spaces of sandy soil between the low-growing perennial plants in the
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Beach layia germinates during the rainy season between fall and midwinter, blooms in spring (March to July), and completes its life cycle before the dry season (July to September). Populations tend to be patchy and subject to large annual fluctuations in size and dynamic changes in local distribution
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Coastal dune systems are composed of a mosaic of vegetation communities of varying successional stages. Beach layia occurs in early to midsuccessional communities in areas where sand is actively being deposited or eroding. Too much sand movement prevents plants from establishing, but areas with some
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herbaceous alliance (sea lyme grass patches). Typically, the total vegetation cover in both communities is relatively sparse, and many annual species, including beach layia, colonize the space between established, tufted perennials. Beach layia can also occur in narrow bands of moderately disturbed
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This is a petite annual herb producing a short, glandular stem along the ground or somewhat upright to a maximum length of about 18 centimeters. The fleshy green leaves are oval in shape, with the larger ones roughly lobed and up to 4 or 5 centimeters in length. The
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which form a cup to hold several small white ray florets. The center is filled with yellow disc florets with purple
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habitat along the edges of trails and roads in dune systems dominated by invasive species.
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in California. On March 31, 2022, the category was changed from endangered species to
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of two types. The ray florets yield hairy, curved fruits with no
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by the US Department of the Interior Fish and Wildlife Service.
571: 353: 346: 94: 342: 230: 148: 363: 939: 292:is a species of flowering plant in the family 461:US Department of Interior Rulemaking Docket 112: 93: 69:Learn how and when to remove this message 32:This article includes a list of general 940: 401: 399: 472: 471: 364:Ecology, habitat, and resource needs 18: 396: 377:herbaceous alliance (dune mat) and 13: 38:it lacks sufficient corresponding 14: 969: 435: 886:urn:lsid:ipni.org:names:231589-1 754:urn:lsid:ipni.org:names:229111-1 341:has a base of overlapping green 135: 23: 452:Elkhorn Slough Species Profile 323: 1: 948:NatureServe imperiled species 389: 7: 958:Endemic flora of California 420:(62): 18722. March 31, 2022 10: 974: 16:Species of flowering plant 803: 480: 296:known by the common name 259: 252: 132:Scientific classification 130: 110: 101: 92: 85: 442:Jepson Manual Treatment 407:"Rules and Regulations" 53:more precise citations. 352:The fruits are small 805:Madaroglossa carnosa 375:Ambrosia chamissonis 333:is borne on a short 447:USDA Plants Profile 104:Conservation status 318:threatened species 314:endangered species 935: 934: 710:Open Tree of Life 474:Taxon identifiers 371:Abronia latifolia 285: 284: 125: 79: 78: 71: 965: 928: 927: 915: 914: 902: 901: 889: 888: 876: 875: 863: 862: 850: 849: 837: 836: 824: 823: 822: 796: 795: 783: 782: 770: 769: 757: 756: 744: 743: 731: 730: 718: 717: 705: 704: 692: 691: 679: 678: 666: 665: 653: 652: 640: 639: 627: 626: 614: 613: 601: 600: 588: 587: 575: 574: 562: 561: 552: 551: 539: 538: 526: 525: 516: 515: 514: 501: 500: 499: 469: 468: 429: 428: 426: 425: 414:Federal Register 411: 403: 265: 140: 139: 119: 116: 115: 97: 83: 82: 74: 67: 63: 60: 54: 49:this article by 40:inline citations 27: 26: 19: 973: 972: 968: 967: 966: 964: 963: 962: 938: 937: 936: 931: 923: 918: 910: 905: 897: 892: 884: 879: 871: 866: 858: 853: 845: 840: 832: 827: 818: 817: 812: 799: 791: 786: 778: 773: 765: 760: 752: 747: 739: 734: 726: 721: 713: 708: 700: 695: 687: 682: 674: 669: 661: 656: 648: 643: 635: 630: 622: 617: 609: 604: 596: 591: 583: 578: 570: 565: 557: 555: 547: 542: 534: 529: 521: 519: 510: 509: 504: 495: 494: 489: 476: 463:Regulations.gov 438: 433: 432: 423: 421: 409: 405: 404: 397: 392: 366: 326: 281: 267: 261: 248: 245:L. carnosa 134: 126: 117: 113: 106: 75: 64: 58: 55: 45:Please help to 44: 28: 24: 17: 12: 11: 5: 971: 961: 960: 955: 950: 933: 932: 930: 929: 916: 912:wfo-0000079977 903: 890: 877: 864: 851: 838: 825: 809: 807: 801: 800: 798: 797: 784: 780:wfo-0000063759 771: 758: 745: 732: 719: 706: 693: 680: 667: 654: 641: 628: 615: 602: 589: 576: 563: 553: 540: 527: 517: 502: 486: 484: 478: 477: 466: 465: 459: 454: 449: 444: 437: 436:External links 434: 431: 430: 394: 393: 391: 388: 365: 362: 325: 322: 298:beach tidytips 283: 282: 268: 257: 256: 250: 249: 242: 240: 236: 235: 228: 224: 223: 218: 214: 213: 208: 204: 203: 198: 191: 190: 185: 178: 177: 172: 165: 164: 159: 152: 151: 146: 142: 141: 128: 127: 111: 108: 107: 102: 99: 98: 90: 89: 77: 76: 31: 29: 22: 15: 9: 6: 4: 3: 2: 970: 959: 956: 954: 951: 949: 946: 945: 943: 926: 921: 917: 913: 908: 904: 900: 895: 891: 887: 882: 878: 874: 869: 865: 861: 856: 852: 848: 843: 839: 835: 830: 826: 821: 815: 811: 810: 808: 806: 802: 794: 789: 785: 781: 776: 772: 768: 763: 759: 755: 750: 746: 742: 737: 733: 729: 724: 720: 716: 711: 707: 703: 698: 694: 690: 685: 681: 677: 672: 668: 664: 659: 655: 651: 646: 642: 638: 633: 629: 625: 620: 616: 612: 607: 603: 599: 594: 590: 586: 581: 577: 573: 568: 564: 560: 554: 550: 545: 541: 537: 532: 528: 524: 518: 513: 512:Layia carnosa 507: 503: 498: 492: 488: 487: 485: 483: 482:Layia carnosa 479: 475: 470: 464: 460: 458: 457:Photo gallery 455: 453: 450: 448: 445: 443: 440: 439: 419: 415: 408: 402: 400: 395: 387: 383: 380: 379:Leymus mollis 376: 372: 361: 359: 355: 350: 348: 344: 340: 336: 332: 331:inflorescence 321: 319: 315: 311: 307: 303: 299: 295: 291: 290: 289:Layia carnosa 280: 276: 272: 266: 264: 263:Layia carnosa 258: 255: 254:Binomial name 251: 247: 246: 241: 238: 237: 234: 233: 229: 226: 225: 222: 219: 216: 215: 212: 209: 206: 205: 202: 199: 196: 193: 192: 189: 186: 183: 180: 179: 176: 173: 170: 167: 166: 163: 162:Tracheophytes 160: 157: 154: 153: 150: 147: 144: 143: 138: 133: 129: 123: 109: 105: 100: 96: 91: 88: 87:Layia carnosa 84: 81: 73: 70: 62: 52: 48: 42: 41: 35: 30: 21: 20: 804: 481: 422:. Retrieved 417: 413: 384: 367: 351: 327: 301: 297: 288: 287: 286: 262: 260: 244: 243: 231: 194: 181: 168: 155: 86: 80: 65: 56: 37: 684:NatureServe 619:iNaturalist 506:Wikispecies 339:flower head 324:Description 302:beach layia 175:Angiosperms 122:NatureServe 51:introducing 942:Categories 723:Plant List 520:Calflora: 424:2022-04-09 390:References 343:phyllaries 310:California 294:Asteraceae 221:Asteraceae 118:Imperiled 59:April 2022 34:references 820:Q95930818 728:gcc-20723 585:250067059 239:Species: 211:Asterales 145:Kingdom: 899:50245112 894:Tropicos 873:231589-1 814:Wikidata 762:Tropicos 689:2.139784 650:10198407 637:229111-1 497:Q6505561 491:Wikidata 335:peduncle 304:. It is 217:Family: 201:Asterids 188:Eudicots 925:1178086 847:3110073 793:1087884 767:2710616 598:5392987 354:achenes 347:anthers 306:endemic 279:A. Gray 227:Genus: 207:Order: 149:Plantae 120: ( 47:improve 860:319838 736:PLANTS 715:509069 702:349392 663:120815 611:319837 572:395498 556:ECOS: 358:pappus 337:. The 277:& 36:, but 953:Layia 920:WoRMS 834:6R3HW 788:WoRMS 741:LACA4 676:37856 645:IRMNG 624:60971 536:3SNXT 410:(PDF) 300:, or 275:Torr. 271:Nutt. 232:Layia 195:Clade 182:Clade 169:Clade 156:Clade 881:POWO 868:IPNI 855:GRIN 842:GBIF 749:POWO 697:NCBI 671:ITIS 632:IPNI 606:GRIN 593:GBIF 559:6728 544:CNPS 523:4635 907:WFO 829:CoL 775:WFO 658:ISC 580:FNA 567:EoL 549:960 531:CoL 308:to 944:: 922:: 909:: 896:: 883:: 870:: 857:: 844:: 831:: 816:: 790:: 777:: 764:: 751:: 738:: 725:: 712:: 699:: 686:: 673:: 660:: 647:: 634:: 621:: 608:: 595:: 582:: 569:: 546:: 533:: 508:: 493:: 418:87 416:. 412:. 398:^ 349:. 273:) 197:: 184:: 171:: 158:: 427:. 373:— 269:( 124:) 72:) 66:( 61:) 57:( 43:.

Index

references
inline citations
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introducing
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Conservation status
NatureServe
Scientific classification
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Plantae
Tracheophytes
Angiosperms
Eudicots
Asterids
Asterales
Asteraceae
Layia
Binomial name
Nutt.
Torr.
A. Gray
Asteraceae
endemic
California
endangered species
threatened species
inflorescence
peduncle
flower head

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