386:
movement on a periodic basis support early successional communities. Movement of sand by wind is essential for the development and sustainability of a dune system. Wind is also important to beach layia specifically because it is the mechanism by which seeds are dispersed. The achenes (a small, dry, one-seeded fruit that does not open to release the seed) have pappus (feathery bristles) that allow them to be carried by wind for a short distance. Although not all seeds may land on suitable habitat, this adaptation allows the small annual to spread across the landscape into uninhabited areas. As a winter germinating annual, beach layia requires rainfall during the winter months (November through
February) for germination and, although it is relatively tolerant to the droughtlike conditions of upland dunes, it does need some moisture through the spring to prevent desiccation. Moisture also reduces the risk of burial, as dry sand is more mobile and mortality caused by burial has been documented. The overall resource needs that beach layia requires in order for individuals to complete their life cycles and for populations to maintain viability are: (1) Sandy soils with sparse native vegetation cover, (2) Rainfall during the winter germination period, (3) Sunlight (full sun exposure for photosynthesis), and (4) Unknown degree of crosspollination (to add to genetic diversity).
369:
associated with the shifts in dune blowouts, remobilization, and natural dune stabilization that occur in the coastal dune ecosystem. Beach layia plants often occur where sparse vegetation traps wind-dispersed seeds, but causes minimal shading. Seeds are dispersed by wind mostly during late spring and summer months. Additionally, beach layia is self-compatible (i.e., able to be fertilized by its own pollen), is capable of self-pollination, and is visited by a variety of insects that may assist in cross-pollination . Although the role of pollinators is currently unclear, sexual reproduction does add to genetic diversity. Beach layia occurs in open spaces of sandy soil between the low-growing perennial plants in the
25:
137:
114:
95:
368:
Beach layia germinates during the rainy season between fall and midwinter, blooms in spring (March to July), and completes its life cycle before the dry season (July to
September). Populations tend to be patchy and subject to large annual fluctuations in size and dynamic changes in local distribution
385:
Coastal dune systems are composed of a mosaic of vegetation communities of varying successional stages. Beach layia occurs in early to midsuccessional communities in areas where sand is actively being deposited or eroding. Too much sand movement prevents plants from establishing, but areas with some
381:
herbaceous alliance (sea lyme grass patches). Typically, the total vegetation cover in both communities is relatively sparse, and many annual species, including beach layia, colonize the space between established, tufted perennials. Beach layia can also occur in narrow bands of moderately disturbed
328:
This is a petite annual herb producing a short, glandular stem along the ground or somewhat upright to a maximum length of about 18 centimeters. The fleshy green leaves are oval in shape, with the larger ones roughly lobed and up to 4 or 5 centimeters in length. The
644:
312:, where it lives in coastal habitat. It is known from several areas of mostly fragmented or relict sand dune habitat, and it was listed as an
696:
735:
841:
592:
456:
854:
670:
605:
947:
957:
675:
68:
46:
345:
which form a cup to hold several small white ray florets. The center is filled with yellow disc florets with purple
39:
919:
787:
701:
867:
631:
543:
121:
885:
753:
584:
522:
657:
859:
610:
451:
880:
748:
441:
406:
924:
792:
357:
33:
688:
462:
136:
740:
511:
649:
579:
473:
50:
253:
714:
566:
382:
habitat along the edges of trails and roads in dune systems dominated by invasive species.
374:
8:
103:
906:
774:
317:
313:
131:
727:
316:
in
California. On March 31, 2022, the category was changed from endangered species to
828:
709:
530:
370:
334:
446:
911:
833:
779:
535:
548:
819:
174:
952:
722:
496:
270:
161:
941:
378:
338:
330:
662:
898:
766:
683:
618:
505:
274:
558:
846:
597:
309:
293:
220:
623:
210:
467:
360:, while the disc florets yield fruits with a long, hairlike pappus.
893:
813:
761:
490:
305:
278:
200:
187:
872:
636:
356:
of two types. The ray florets yield hairy, curved fruits with no
320:
by the US Department of the
Interior Fish and Wildlife Service.
571:
353:
346:
94:
342:
230:
148:
363:
939:
292:is a species of flowering plant in the family
461:US Department of Interior Rulemaking Docket
112:
93:
69:Learn how and when to remove this message
32:This article includes a list of general
940:
401:
399:
472:
471:
364:Ecology, habitat, and resource needs
18:
396:
377:herbaceous alliance (dune mat) and
13:
38:it lacks sufficient corresponding
14:
969:
435:
886:urn:lsid:ipni.org:names:231589-1
754:urn:lsid:ipni.org:names:229111-1
341:has a base of overlapping green
135:
23:
452:Elkhorn Slough Species Profile
323:
1:
948:NatureServe imperiled species
389:
7:
958:Endemic flora of California
420:(62): 18722. March 31, 2022
10:
974:
16:Species of flowering plant
803:
480:
296:known by the common name
259:
252:
132:Scientific classification
130:
110:
101:
92:
85:
442:Jepson Manual Treatment
407:"Rules and Regulations"
53:more precise citations.
352:The fruits are small
805:Madaroglossa carnosa
375:Ambrosia chamissonis
333:is borne on a short
447:USDA Plants Profile
104:Conservation status
318:threatened species
314:endangered species
935:
934:
710:Open Tree of Life
474:Taxon identifiers
371:Abronia latifolia
285:
284:
125:
79:
78:
71:
965:
928:
927:
915:
914:
902:
901:
889:
888:
876:
875:
863:
862:
850:
849:
837:
836:
824:
823:
822:
796:
795:
783:
782:
770:
769:
757:
756:
744:
743:
731:
730:
718:
717:
705:
704:
692:
691:
679:
678:
666:
665:
653:
652:
640:
639:
627:
626:
614:
613:
601:
600:
588:
587:
575:
574:
562:
561:
552:
551:
539:
538:
526:
525:
516:
515:
514:
501:
500:
499:
469:
468:
429:
428:
426:
425:
414:Federal Register
411:
403:
265:
140:
139:
119:
116:
115:
97:
83:
82:
74:
67:
63:
60:
54:
49:this article by
40:inline citations
27:
26:
19:
973:
972:
968:
967:
966:
964:
963:
962:
938:
937:
936:
931:
923:
918:
910:
905:
897:
892:
884:
879:
871:
866:
858:
853:
845:
840:
832:
827:
818:
817:
812:
799:
791:
786:
778:
773:
765:
760:
752:
747:
739:
734:
726:
721:
713:
708:
700:
695:
687:
682:
674:
669:
661:
656:
648:
643:
635:
630:
622:
617:
609:
604:
596:
591:
583:
578:
570:
565:
557:
555:
547:
542:
534:
529:
521:
519:
510:
509:
504:
495:
494:
489:
476:
463:Regulations.gov
438:
433:
432:
423:
421:
409:
405:
404:
397:
392:
366:
326:
281:
267:
261:
248:
245:L. carnosa
134:
126:
117:
113:
106:
75:
64:
58:
55:
45:Please help to
44:
28:
24:
17:
12:
11:
5:
971:
961:
960:
955:
950:
933:
932:
930:
929:
916:
912:wfo-0000079977
903:
890:
877:
864:
851:
838:
825:
809:
807:
801:
800:
798:
797:
784:
780:wfo-0000063759
771:
758:
745:
732:
719:
706:
693:
680:
667:
654:
641:
628:
615:
602:
589:
576:
563:
553:
540:
527:
517:
502:
486:
484:
478:
477:
466:
465:
459:
454:
449:
444:
437:
436:External links
434:
431:
430:
394:
393:
391:
388:
365:
362:
325:
322:
298:beach tidytips
283:
282:
268:
257:
256:
250:
249:
242:
240:
236:
235:
228:
224:
223:
218:
214:
213:
208:
204:
203:
198:
191:
190:
185:
178:
177:
172:
165:
164:
159:
152:
151:
146:
142:
141:
128:
127:
111:
108:
107:
102:
99:
98:
90:
89:
77:
76:
31:
29:
22:
15:
9:
6:
4:
3:
2:
970:
959:
956:
954:
951:
949:
946:
945:
943:
926:
921:
917:
913:
908:
904:
900:
895:
891:
887:
882:
878:
874:
869:
865:
861:
856:
852:
848:
843:
839:
835:
830:
826:
821:
815:
811:
810:
808:
806:
802:
794:
789:
785:
781:
776:
772:
768:
763:
759:
755:
750:
746:
742:
737:
733:
729:
724:
720:
716:
711:
707:
703:
698:
694:
690:
685:
681:
677:
672:
668:
664:
659:
655:
651:
646:
642:
638:
633:
629:
625:
620:
616:
612:
607:
603:
599:
594:
590:
586:
581:
577:
573:
568:
564:
560:
554:
550:
545:
541:
537:
532:
528:
524:
518:
513:
512:Layia carnosa
507:
503:
498:
492:
488:
487:
485:
483:
482:Layia carnosa
479:
475:
470:
464:
460:
458:
457:Photo gallery
455:
453:
450:
448:
445:
443:
440:
439:
419:
415:
408:
402:
400:
395:
387:
383:
380:
379:Leymus mollis
376:
372:
361:
359:
355:
350:
348:
344:
340:
336:
332:
331:inflorescence
321:
319:
315:
311:
307:
303:
299:
295:
291:
290:
289:Layia carnosa
280:
276:
272:
266:
264:
263:Layia carnosa
258:
255:
254:Binomial name
251:
247:
246:
241:
238:
237:
234:
233:
229:
226:
225:
222:
219:
216:
215:
212:
209:
206:
205:
202:
199:
196:
193:
192:
189:
186:
183:
180:
179:
176:
173:
170:
167:
166:
163:
162:Tracheophytes
160:
157:
154:
153:
150:
147:
144:
143:
138:
133:
129:
123:
109:
105:
100:
96:
91:
88:
87:Layia carnosa
84:
81:
73:
70:
62:
52:
48:
42:
41:
35:
30:
21:
20:
804:
481:
422:. Retrieved
417:
413:
384:
367:
351:
327:
301:
297:
288:
287:
286:
262:
260:
244:
243:
231:
194:
181:
168:
155:
86:
80:
65:
56:
37:
684:NatureServe
619:iNaturalist
506:Wikispecies
339:flower head
324:Description
302:beach layia
175:Angiosperms
122:NatureServe
51:introducing
942:Categories
723:Plant List
520:Calflora:
424:2022-04-09
390:References
343:phyllaries
310:California
294:Asteraceae
221:Asteraceae
118:Imperiled
59:April 2022
34:references
820:Q95930818
728:gcc-20723
585:250067059
239:Species:
211:Asterales
145:Kingdom:
899:50245112
894:Tropicos
873:231589-1
814:Wikidata
762:Tropicos
689:2.139784
650:10198407
637:229111-1
497:Q6505561
491:Wikidata
335:peduncle
304:. It is
217:Family:
201:Asterids
188:Eudicots
925:1178086
847:3110073
793:1087884
767:2710616
598:5392987
354:achenes
347:anthers
306:endemic
279:A. Gray
227:Genus:
207:Order:
149:Plantae
120: (
47:improve
860:319838
736:PLANTS
715:509069
702:349392
663:120815
611:319837
572:395498
556:ECOS:
358:pappus
337:. The
277:&
36:, but
953:Layia
920:WoRMS
834:6R3HW
788:WoRMS
741:LACA4
676:37856
645:IRMNG
624:60971
536:3SNXT
410:(PDF)
300:, or
275:Torr.
271:Nutt.
232:Layia
195:Clade
182:Clade
169:Clade
156:Clade
881:POWO
868:IPNI
855:GRIN
842:GBIF
749:POWO
697:NCBI
671:ITIS
632:IPNI
606:GRIN
593:GBIF
559:6728
544:CNPS
523:4635
907:WFO
829:CoL
775:WFO
658:ISC
580:FNA
567:EoL
549:960
531:CoL
308:to
944::
922::
909::
896::
883::
870::
857::
844::
831::
816::
790::
777::
764::
751::
738::
725::
712::
699::
686::
673::
660::
647::
634::
621::
608::
595::
582::
569::
546::
533::
508::
493::
418:87
416:.
412:.
398:^
349:.
273:)
197::
184::
171::
158::
427:.
373:—
269:(
124:)
72:)
66:(
61:)
57:(
43:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.