4622:
6737:
4615:
3119:
3944:
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6189:
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6175:
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3125:
972:
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151:
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658:(NHM OR50100). The casts show some detail that is no longer preserved on the original specimens, which have significantly degraded due to pyrite disease. The discovery of this genus gave North American paleontologists the opportunity to observe an articulated, albeit incomplete, theropod skeleton. During the late 19th century, this genus unfortunately became a wastebasket taxon for the referral of isolated theropod elements from across North America, given that
5435:
1106:
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515:
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1114:
793:
129:
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in 1997, in light of the many different theropods discovered since Cope's time. He felt that due to some unusual features it couldn't be placed in any existing family and, like Marsh, he felt that it warranted placement in
Dryptosauridae. This phylogenetic assignment was also supported by the work of
958:
tubercle, which is approximately 1.7 times as wide as the shaft immediately distally, the presence of an ovoid fossa on the medial surface of the femoral shaft immediately proximal to the medial condyle (which is demarcated anteriorly by the mesiodistal crest and demarcated medially by a novel crest)
2210:
Dryptosaurus." In: Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, Catherine A. & Gillette, David D. & Norell, Mark A. & Olshevsky, George & Parrish, J. Michael & Weishampel, David B. The Age of
Dinosaurs. Publications International, LTD. p.
558:
was notified of the discovery and encouraged to visit the marl pit to obtain the specimen. When he arrived, he was thoroughly surprised by the skeleton's completeness and uniqueness, calling it the "finest discovery I have yet made". The skeleton was then deposited at the
959:
the presence of a proximomedially trending ridge on the anterior surface of the fibula immediately proximal to the iliofibularis tubercle, the lip on the lateral surface of the lateral condyle of the astragalus being prominent and overlapping the proximal surface of the
890:
III. The ontological maturity of the holotype individual is supported by the fact that the neurocentral sutures are closed in all of its caudal vertebrae. AMNH FARB 2438 consists of left metatarsal IV, which are likely from the same individual as the holotype.
1816:
may represent the earliest depiction of theropods as highly active and dynamic animals. Knight's artistic hand was guided by E. D. Cope and reflects their progressive opinion about theropod agility, despite their large size, as well as the opinion of
1768:
and its coeval fauna from western North
America, which was dominated by the much larger tyrannosaurids. Although certainly a carnivore, the paucity of known Cretaceous East Coast dinosaurs make ascertaining the specific diet of
625:
possibly belonging to the skeleton was found at the same locale, but this has come into question. Cope presented his description of the specimen to the
Academy on August 21, 1866, naming it a new genus and species of dinosaur,
922:/cm), and recurved. The femur is only 3% longer than the tibia. The longest manual ungual phalanx recovered measured 176 mm (6.9 in) in length. The morphology of the proximal portion of metatarsal IV suggests that
953:
can be distinguished based on the following characteristics: the combination of a reduced humerus (humerus: femur ratio = 0.375) and a large hand (phalanx I-1:femur ratio = 0.200), the strong mediolateral expansion of the
2100:
Chan-gyu Yun (2017). "Teihivenator gen. nov., a new generic name for the tyrannosauroid dinosaur "Laelaps" macropus (Cope, 1868; preoccupied by Koch, 1836)". Journal of
Zoological And Bioscience Research. 4 (2): 7–13.
470:
that could grow up to 7.5 metres (25 ft) long and weigh up to 756–1,500 kilograms (1,667–3,307 lb). Although it is now largely unknown outside of academic circles, the famous 1897 painting of the genus by
1752:
in central New Jersey. Studies suggest that the New Egypt
Formation is a marine unit and it is considered to be the deeper-water equivalent of the Tinton and Red Bank formations. This formation overlies the
2031:
Spamer, E.E., E. Daeschler, and L.G. Vostreys-Shapiro. 1995. A study of fossil vertebrate types in the
Academy of Natural Sciences of Philadelphia. Special Publication 16, Academy of Natural Sciences of
819:. Its hands, which are also relatively large, were once believed to have had three fingers. Brusatte et al. (2011), however, observed an overall similarity in the shape of the available phalanges of
2519:
Olsson, R.K. 1987. Cretaceous stratigraphy of the
Atlantic coastal plain, Atlantic highlands of New Jersey. Geological Society of America Centennial Field Guide–Northeastern Section: 87–90.
914:
had ziphodont dentition. The shape of the alveolus situated on the anterior portion of the fragment suggests that it housed a tooth that was smaller and more circular than the others. This
1789:
were also present, although they were less likely to be hunted due to their armor plating. When hunting, both the skull and hands were important for the capture and processing of prey.
803:
is estimated to have been 7.5 metres (25 ft) long and 756–1,500 kilograms (1,667–3,307 lb), although this is based on partial remains of one individual. Like its relative
2265:
Carr, T.D.; Williamson, T.E. & Schwimmer, D.R. (2005). "A new genus and species of tyrannosauroid from the Late
Cretaceous (middle Campanian) Demopolis Formation of Alabama".
1764:
During the
Maastrichtian stage, the Western Interior Seaway, which stretched in a north–south direction from the present-day Arctic Ocean down to the Gulf of Mexico, separated
963:, and metatarsal IV being observed with a flat shaft proximally (resulting in a semiovoid cross section that is much wider mediolaterally than it is long anteroposteriorly).
1821:, the curator of vertebrate paleontology at the American Museum of Natural History at the time of the painting's commission. The original painting is now preserved in the
2299:
Holtz, T.R. 2004. Tyrannosauroidea. In D. B. Weishampel, P. Dodson, and H. Osmólska (editors), The Dinosauria, 2nd ed.: 111–136. Berkeley: University of California Press.
2229:
Molnar, R.E. 1990. Problematic Theropoda. In D. B. Weishampel, P. Dodson, and H. Osmólska (editors), The Dinosauria: 306–317. Berkeley: University of California Press.
827:
may have had only two functional digits. Each of its fingers were tipped with an eight-inch long, talon-like claw. Its arm morphology suggests that arm reduction in
2220:
Russell, D.A. 1970. Tyrannosauroids from the Late Cretaceous of western Canada. National Museum of Natural Science (Ottawa) Publications in Paleontology 1: 1–34.
2115:
Theropod specimens from the Navesink Formation and their implications for the Diversity and Biogeography of Ornithomimosaurs and Tyrannosauroids on Appalachia
6886:
983:
has been classified in a number of different theropod families. Cope (1866), Leidy (1868), and Lydekker (1888) noted obvious similarities with the genus
777:
and found that only the partial tibia could be definitely classified as that of a tyrannosauroid and that the distal metatarsal could have been from an
2356:
1049:. This observation was also supported by the work of Baird and Horner (1979), but did not have wide acceptance until a new discovery was made in 2005.
1875:
Leidy, J. 1856. Notice of remains of extinct reptiles and fishes discovered by F. V. Hayden in the badlands of the Judith River, Nebraska Territory.
654:(2011) noted that well-preserved, historic casts of most of the type material from ANSP 9995 and AMNH FARB 2438 are housed in the collections of the
2360:
2018:
1717:
726:
560:
2352:
6906:
2438:"Dinosaurs from the Santonian–Campanian Atlantic coastline substantiate phylogenetic signatures of vicariance in Cretaceous North America"
910:
The fragmentary right maxilla preserves the three alveoli in full and the fourth only partially. The authors were able to ascertain that
838:
The type specimen is a fragmentary skeleton belonging to a single adult individual, ANSP 9995. ANSP 9995 consists of a fragmentary right
1672:
6814:
662:
was not recognized as a distinct group of large theropods at the time and numerous theropod species were assigned to it (often as
6901:
6896:
2063:
4621:
4614:
2602:
1977:"Redescription of the holotype of Dryptosaurus aquilunguis (Dinosauria: Theropoda) from the Upper Cretaceous of New Jersey"
6736:
1976:
6911:
2581:
725:-like characteristics of the teeth are primitive to theropods as a whole and may have been present in other large-bodied
1822:
17:
1952:
420:
2501:
Olsson, R.K. (1960). "Foraminifera of latest Cretaceous and earliest Tertiary age in the New Jersey coastal plain".
1856:
1499:
918:
tooth is common among tyrannosauroids. The disarticulated teeth recovered are transversely narrow, serrated (17–18
5409:
3943:
1889:
611:
which are now lost were also noted by Cope in addition to several sacral vertebrae which are now referred to the
527:
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2278:
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who never failed to catch what it was hunting. The genus name was occupied by a mite and replaced by the name
6891:
6615:
3922:
3118:
2337:
1727:
5427:
3124:
781:. Brownstein also placed the pedal phalanges in Ornithimimosauria, though he did not create a lectotype for
6921:
6188:
4628:
835:
may have used both its arms and its jaws and as weapons when hunting, capturing, and its processing prey.
713:
of Colorado. Five damaged partial tooth crowns from AMNH 5780, mistakenly thought to have belonged to the
1734:. The specimen was collected by quarry workers in marl and sandstone that were deposited during the late
539:
150:
3936:
2311:"The systematics of Late Jurassic tyrannosauroids (Dinosauria: Theropoda) from Europe and North America"
6181:
2131:
1936:
523:
6858:
5434:
6749:
2669:
919:
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and assigned it as an "intermediate" tyrannosauroid that is more derived than basal taxa, such as
6743:
5415:
2574:
1974:
1826:
1676:
137:
6845:
6769:
2055:
2043:
1818:
2087:
Holtz, T.R. (2004). "Tyrannosauroidea." Pp. 111-136 in Weishampel, Dodson and Osmolska (eds).
6881:
6840:
6832:
5140:
5052:
2012:
1782:
1723:
1053:
was the only large carnivore known in eastern North America until the discovery of the basal
813:
seems to have had relatively long arms when compared with more derived tyrannosaurs, such as
643:
522:
Up until 1866, theropods from the Americas were only known from isolated teeth discovered by
492:
447:
298:
255:
6801:
6701:
4829:
4797:
2540:
2449:
2390:
1988:
1932:
1738:
745:
6168:
1831:, in the late 19th century, depicted the animals as large, slow, tail-dragging behemoths.
634:", which is derived from the Greek word "hurricane" or "storm wind", was also the name of
8:
5153:
4940:
1749:
754:
694:
635:
555:
551:
488:
318:
5422:
2528:
Carr & Williamson (2002). "Evolution of basal Tyrannosauroidea from North America".
2453:
2394:
1992:
1825:
collections. By contrast, the typical illustrations of large carnivorous dinosaurs like
942:, in which the third metatarsal is "pinched" between the second and fourth metatarsals.
6608:
5129:
4989:
4845:
4785:
4220:
4130:
4013:
2877:
2567:
2478:
2413:
2376:
2282:
1913:
Cope, E.D. (1866). "Discovery of a gigantic dinosaur in the Cretaceous of New Jersey".
1754:
1722:
and was recovered in the West Jersey Marl Company Pit, in what came to be known as the
1026:
990:
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327:
145:
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2186:
2059:
1948:
1805:
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1006:
778:
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Gilmore, C.W. 1946. A new carnivorous dinosaur from the Lance Formation of Montana.
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Baird, D., and J. Horner. 1979. Cretaceous dinosaurs (Reptilia) of North Carolina.
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1996:
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998:
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455:
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237:
41:
2078:
Cope, E.D., On the genus Laelaps, American Journal of Science, 1868; 2: 415-417.
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from the left hand (I-1, II-2, and an ungual), the shafts of the left and right
6622:
6585:
6566:
6528:
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6314:
6027:
5993:
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2004:
1684:
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994:
971:
343:
2377:"Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans"
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2753:
2542:- Cretaceous Dinosaurs of the Southeastern United States by David T. King Jr.
2469:
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The following cladogram containing almost all tyrannosauroids is by Loewen
985:
863:
859:
768:
740:
683:
564:
352:
2327:
2310:
1975:
Carpenter, Ken & Russell, Dale A, Donald Baird, and R. Denton (1997).
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2884:
2792:
2784:
2437:
2264:
1774:
894:
847:
689:
600:
54:
2461:
709:
was coined by Cope in 1877 for a partial dentary (now missing) from the
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6503:
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5939:
5757:
5743:
5728:
5680:
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3378:
3298:
3169:
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2898:
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1254:
1105:
915:
883:
805:
543:
463:
208:
99:
64:
2161:(Dinosauria: Theropoda) and a Review of its Tyrannosauroid Affinities"
514:
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6643:
6343:
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5602:
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4770:
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4090:
3975:
3962:
3889:
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3727:
3484:
3347:
3155:
3142:
3060:
2912:
2731:
2718:
2681:
2590:
2181:
1113:
997:. Marsh, however, examined the remains and later assigned to its own
960:
930:
foot, an advanced feature shared by derived tyrannosauroids, such as
647:
500:
467:
440:
221:
162:
104:
48:
6763:
1070:
in both body size and morphology. This discovery made it clear that
792:
128:
6786:
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5659:
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5017:
4398:
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3322:
2959:
2638:
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2172:
2054:. Bloomington and Indianapolis: Indiana University Press. pp.
1748:
The southern New Jersey Hornerstown Formation is also known as the
1680:
1394:
1197:
1180:
1088:
1029:
was the first to observe that certain anatomical features may link
1022:, though its exact placement within that group remained uncertain.
899:
851:
843:
596:
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572:
503:
480:
443:
195:
182:
94:
89:
74:
69:
59:
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5284:
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1931:
955:
867:
855:
839:
608:
604:
584:
531:
109:
84:
773:. Earlier that year, Brownstein (2017) analyzed the material of
721:
and probably belong to that genus instead. However, some of the
6443:
6131:
3575:
2608:
1034:
879:
673:
592:
484:
172:
1947:(Abridged ed.). United Kingdom: Oxford University Press.
1915:
Proceedings of the Academy of Natural Sciences of Philadelphia
1877:
Proceedings of the Academy of Natural Sciences of Philadelphia
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2279:
10.1671/0272-4634(2005)025[0119:ANGASO]2.0.CO;2
2091:(second edition). University of California Press, Berkeley.
1066:, which is known from more complete remains, is similar to
993:. Based on this line of reasoning, Cope classified it as a
538:
uncovered an incomplete theropod skeleton in a quarry near
535:
450:
approximately 67.6 million years ago during the end of the
409:
397:
2197:
1812:
The 1897 watercolor painting by Charles R. Knight titled
989:, which was known at the time from remains discovered in
388:
1001:
family, Dryptosauridae. The fossil material assigned to
765:. In 2017, it was informally given the new generic name
1785:, and they may have been a prominent part of its diet.
735:
was also coined by Cope for a partial leg found in the
1098:, but more primitive than members of the more derived
2155:
Brusatte, S.L.; Benson, R.B.J.; Norell, M.A. (2011).
421:
412:
403:
379:
2154:
1838:
823:
with those of derived tyrannosaurids and noted that
394:
391:
385:
382:
6116:
567:under the catalogue number ANSP 9995 and includes;
400:
2035:
1940:
1082:(2013) confirmed the tyrannosauroid affinities of
136:Reconstructed skeletons mounted in fighting pose,
2338:20.500.11820/31f38145-54e7-48f8-819a-262601e93f2b
761:in his contribution to the second edition of the
6873:
6218:
6084:
2435:
5575:
2150:
2148:
2146:
2144:
2142:
2140:
2050:. In Tanke, Darren; Carpenter, Kenneth (eds.).
1687:, which now includes both "Cryptotyrannus" and
1683:. Brownstein (2021) also reinstated the family
4099:
2653:
2527:
2308:
534:. During the summer of 1866, workers from the
2575:
2351:
1078:. Detailed phylogenetic analysis by Brusatte
831:may not have proceeded in a uniform fashion.
717:holotype, share many features in common with
5464:
2137:
2017:: CS1 maint: multiple names: authors list (
1745:period, approximately 67 million years ago.
4658:
4576:
3698:
1925:
6887:Late Cretaceous dinosaurs of North America
2582:
2568:
1777:are known from the same time and place as
127:
2477:
2429:
2412:
2402:
2336:
2326:
2180:
1675:", an informally named theropod from the
850:, lateral teeth, 11 middle-distal caudal
743:had referred earlier to the ornithomimid
3973:
3888:
3153:
2729:
2345:
2081:
1970:
1968:
1966:
1964:
1796:
1792:
1698:
1112:
1104:
1014:studies during the 1990s suggested that
1010:Russell (1970) and Molnar (1990). Other
970:
893:
791:
672:
513:
509:
2309:Brusatte, S.L.; Benson, R.B.J. (2013).
1906:
14:
6874:
2500:
2258:
2128:The Princeton Field Guide to Dinosaurs
2044:"On the type and referred material of
701:
446:that lived on the island continent of
6768:
6767:
6206:
6205:
5451:
4645:
3960:
3140:
2716:
2601:
2563:
2240:Smithsonian Miscellaneous Collections
2109:
2107:
2041:
1961:
1761:referred material has been reported.
906:with three fingers and lipless snout.
6859:A52387D3-0775-FFC1-FF97-FAB1B85ED2A6
1912:
1703:Diagram showing known remains in of
677:An outdated historical depiction of
554:. In August of 1866, paleontologist
6907:Taxa named by Othniel Charles Marsh
727:Morrison Formation theropod species
24:
2530:Journal of Vertebrate Paleontology
2267:Journal of Vertebrate Paleontology
2104:
2048:Cope 1877 (Dinosauria: Theropoda)"
1981:Journal of Vertebrate Paleontology
1726:in Barnsboro, Mantua Township, in
483:of its time, in spite of its poor
25:
6933:
5452:
2546:
979:Since the time of its discovery,
966:
796:Estimated size, compared to human
670:), only to be later reclassified.
6748:
6742:
6735:
6187:
6180:
6173:
6167:
5433:
5426:
5421:
5414:
5408:
4646:
4627:
4620:
4613:
3942:
3935:
3928:
3921:
3123:
3117:
2436:Doran Brownstein, Chase (2021).
1857:Timeline of tyrannosaur research
1841:
1500:Appalachiosaurus montgomeriensis
1037:Late Cretaceous tyrannosaurids,
656:Natural History Museum in London
546:with sediments belonging to the
375:
149:
52:
2513:
2494:
2302:
2293:
2245:
2232:
2223:
2214:
2120:
2117:(No. e3105v1). PeerJ Preprints.
2094:
1694:
2072:
2025:
2001:10.1080/02724634.1997.10011003
1882:
1869:
787:
757:listed it as an indeterminate
13:
1:
6902:Fossil taxa described in 1877
6897:Fossil taxa described in 1866
3961:
3141:
2717:
2589:
2315:Acta Palaeontologica Polonica
1862:
886:, and a midshaft fragment of
518:Elements of the type specimen
491:in 1866 and later renamed by
479:one of the more widely known
2404:10.1371/journal.pone.0079420
7:
1834:
1671:to be the sister taxon to "
866:bones, a fragmentary right
595:, and nearly complete left
561:Academy of Natural Sciences
487:record. First described by
10:
6938:
6912:Paleontology in New Jersey
2442:Royal Society Open Science
2132:Princeton University Press
2113:Brownstein, C. D. (2017).
2101:doi:10.24896/jzbr.2017422.
1781:, the island continent of
854:, both the left and right
27:Extinct genus of dinosaurs
6776:
6732:
6689:
6584:
6527:
6498:
6489:
6415:
6406:
6390:
6379:
6289:
6274:
6270:
6259:
6229:
6214:
6201:
6165:
6112:
6080:
6026:
5992:
5965:
5938:
5916:
5907:
5880:
5869:
5840:
5829:
5709:
5705:
5694:
5642:
5613:
5590:
5586:
5564:
5475:
5460:
5447:
5406:
5376:
5310:
5242:
5232:
5228:
5163:
5148:
5139:
5060:
5051:
4999:
4948:
4939:
4895:
4884:
4828:
4805:
4796:
4758:
4754:
4743:
4704:
4693:
4669:
4654:
4641:
4610:
4572:
4529:
4485:
4476:
4438:
4393:
4384:
4346:
4335:
4271:
4230:
4215:
4206:
4190:
4186:
4175:
4129:
4114:
4110:
4088:
3984:
3969:
3956:
3918:
3884:
3787:
3707:
3677:
3613:
3527:
3516:
3460:
3426:
3342:
3333:
3293:
3289:
3278:
3247:
3202:
3198:
3187:
3164:
3149:
3136:
3114:
3079:
3059:
2983:
2939:
2930:
2822:
2811:
2740:
2725:
2712:
2680:
2662:
2649:
2597:
2165:American Museum Novitates
2042:Chure, Daniel J. (2001).
1570:
1544:
1527:
1520:
1513:
1496:
1489:
1463:
1446:
1439:
1432:
1415:
1408:
1391:
1384:
1367:
1360:
1343:
1336:
1319:
1312:
1268:
1251:
1244:
1227:
1220:
1194:
1177:
1160:
1153:
1146:
1136:
749:, distinguishing it from
333:
326:
304:
297:
146:Scientific classification
144:
135:
126:
34:
2554:Dryptosaurus aquilunguis
2159:Dryptosaurus aquilunguis
2052:Mesozoic Vertebrate Life
1894:www.theropoddatabase.com
1667:Brownstein (2021) found
1467:Xiongguanlong baimoensis
1419:Dryptosaurus aquilunguis
1272:Stokesosaurus clevelandi
1164:Proceratosaurus bradleyi
636:a dog in Greek mythology
536:West Jersey Marl Company
311:Dryptosaurus aquilunguis
2503:Journal of Paleontology
1943:A Greek-English Lexicon
1757:, from which potential
1677:Merchantville Formation
1231:Sinotyrannus kazuoensis
642:"slashing reptile", by
506:ever known to science.
138:New Jersey State Museum
1819:Henry Fairfield Osborn
1809:
1708:
1118:
1110:
976:
945:According to Brusatte
907:
846:, a fragmentary right
842:, a fragmentary right
797:
698:
693:in the background. By
646:, Cope's rival in the
519:
6917:Cretaceous New Jersey
6841:Paleobiology Database
5141:Carcharodontosaurinae
5053:Carcharodontosauridae
2328:10.4202/app.2011.0141
1933:Liddell, Henry George
1800:
1793:Cultural significance
1724:Hornerstown Formation
1711:The type specimen of
1702:
1116:
1108:
974:
897:
795:
681:(center) confronting
676:
517:
510:Discovery and species
493:Othniel Charles Marsh
6892:Maastrichtian genera
6117:Dubious coelurosaurs
4830:Metriacanthosaurinae
4798:Metriacanthosauridae
3080:Dubious neotheropods
1450:Alectrosaurus olseni
1395:Raptorex kriegsteini
1181:Kileskus aristotocus
1117:Limb and other bones
1109:Postcranial elements
991:southeastern England
753:by its longer toes.
650:, in 1877. Brusatte
524:Ferdinand Van Hayden
6922:Symbols of Delaware
5154:Carcharodontosaurus
4941:Carcharodontosauria
2462:10.1098/rsos.210127
2454:2021RSOS....810127D
2395:2013PLoSO...879420L
2126:Paul, G.S. (2010).
2046:Laelaps trihedrodon
1993:1997JVPal..17..561C
1750:New Egypt Formation
1255:Juratyrant langhami
707:Laelaps trihedrodon
702:Misassigned species
695:Edward Drinker Cope
628:Laelaps aquilunguis
556:Edward Drinker Cope
552:New Egypt Formation
499:is among the first
489:Edward Drinker Cope
338:Laelaps aquilunguis
290:D. aquilunguis
6609:Archaeornithomimus
5130:Veterupristisaurus
4990:Veterupristisaurus
4846:Metriacanthosaurus
4786:Piatnitzkysauridae
4221:Eustreptospondylus
4131:Piatnitzkysauridae
4014:Chuandongocoelurus
2878:Notatesseraeraptor
1890:"Tyrannosauroidea"
1810:
1755:Navesink Formation
1709:
1371:Bagaraatan ostromi
1119:
1111:
1027:Charles W. Gilmore
977:
975:Holotype claw bone
928:arctometatarsalian
908:
798:
737:Navesink Formation
711:Morrison Formation
699:
520:
466:, ground-dwelling
18:Laelaps (dinosaur)
6869:
6868:
6828:Open Tree of Life
6770:Taxon identifiers
6761:
6760:
6757:
6756:
6730:
6729:
6726:
6725:
6722:
6721:
6718:
6717:
6685:
6684:
6681:
6680:
6491:Macrocheiriformes
6408:Ornithomimosauria
6382:Maniraptoriformes
6375:
6374:
6221:Maniraptoromorpha
6208:Maniraptoromorpha
6197:
6196:
6163:
6162:
6159:
6158:
6155:
6154:
6108:
6107:
6104:
6103:
6086:Maniraptoromorpha
6076:
6075:
6072:
6071:
6068:
6067:
6064:
6063:
6060:
6059:
5825:
5824:
5690:
5689:
5644:Proceratosauridae
5596:Chingkankousaurus
5443:
5442:
5404:
5403:
5400:
5399:
5396:
5395:
5372:
5371:
5224:
5223:
5220:
5219:
5216:
5215:
5212:
5211:
5208:
5207:
5204:
5203:
4880:
4879:
4876:
4875:
4637:
4636:
4608:
4607:
4604:
4603:
4600:
4599:
4596:
4595:
4568:
4567:
4564:
4563:
4560:
4559:
4556:
4555:
4472:
4471:
4428:Vallibonavenatrix
4331:
4330:
4327:
4326:
4171:
4170:
3952:
3951:
3916:
3915:
3912:
3911:
3908:
3907:
3880:
3879:
3876:
3875:
3872:
3871:
3868:
3867:
3864:
3863:
3512:
3511:
3508:
3507:
3274:
3273:
3132:
3131:
3112:
3111:
3108:
3107:
3104:
3103:
3055:
3054:
2708:
2707:
2704:
2703:
2700:
2699:
2065:978-0-253-33907-2
1806:Charles R. Knight
1728:Gloucester County
1664:
1663:
1655:
1654:
1646:
1645:
1637:
1636:
1628:
1627:
1619:
1618:
1610:
1609:
1601:
1600:
1592:
1591:
1583:
1582:
1559:
1558:
1548:Alioramus remotus
1478:
1477:
1301:
1300:
1292:
1291:
1283:
1282:
1209:
1208:
1143:Proceratosauridae
1074:was also a basal
783:Laelaps macropus.
528:Geological Survey
473:Charles R. Knight
366:
365:
360:
347:
278:
259:
16:(Redirected from
6929:
6862:
6861:
6849:
6848:
6836:
6835:
6823:
6822:
6810:
6809:
6797:
6796:
6795:
6765:
6764:
6752:
6746:
6739:
6595:Aepyornithomimus
6496:
6495:
6413:
6412:
6388:
6387:
6352:Sinocalliopteryx
6272:
6271:
6268:
6267:
6262:Neocoelurosauria
6227:
6226:
6216:
6215:
6203:
6202:
6191:
6184:
6177:
6171:
6114:
6113:
6082:
6081:
6051:Zhuchengtyrannus
5983:Thanatotheristes
5914:
5913:
5878:
5877:
5846:Appalachiosaurus
5838:
5837:
5707:
5706:
5703:
5702:
5697:Pantyrannosauria
5588:
5587:
5584:
5583:
5578:Tyrannosauroidea
5573:
5572:
5548:Xinjiangovenator
5473:
5472:
5462:
5461:
5449:
5448:
5437:
5430:
5425:
5418:
5412:
5240:
5239:
5230:
5229:
5146:
5145:
5066:Acrocanthosaurus
5058:
5057:
4946:
4945:
4893:
4892:
4819:Yangchuanosaurus
4803:
4802:
4756:
4755:
4752:
4751:
4702:
4701:
4683:Lourinhanosaurus
4667:
4666:
4656:
4655:
4643:
4642:
4631:
4624:
4617:
4574:
4573:
4540:Sigilmassasaurus
4483:
4482:
4440:Ceratosuchopsini
4391:
4390:
4344:
4343:
4289:Dubreuillosaurus
4213:
4212:
4196:Streptospondylus
4188:
4187:
4184:
4183:
4154:Piatnitzkysaurus
4112:
4111:
4108:
4107:
4097:
4096:
3982:
3981:
3971:
3970:
3958:
3957:
3946:
3939:
3932:
3925:
3886:
3885:
3721:Ekrixinatosaurus
3705:
3704:
3696:
3695:
3525:
3524:
3340:
3339:
3291:
3290:
3287:
3286:
3200:
3199:
3196:
3195:
3162:
3161:
3151:
3150:
3138:
3137:
3127:
3121:
3031:Procompsognathus
2937:
2936:
2820:
2819:
2738:
2737:
2727:
2726:
2714:
2713:
2660:
2659:
2651:
2650:
2644:
2643:
2599:
2598:
2584:
2577:
2570:
2561:
2560:
2537:
2520:
2517:
2511:
2510:
2498:
2492:
2491:
2481:
2433:
2427:
2426:
2416:
2406:
2349:
2343:
2342:
2340:
2330:
2306:
2300:
2297:
2291:
2290:
2262:
2256:
2249:
2243:
2236:
2230:
2227:
2221:
2218:
2212:
2208:
2195:
2194:
2184:
2157:"The Anatomy of
2152:
2135:
2124:
2118:
2111:
2102:
2098:
2092:
2085:
2079:
2076:
2070:
2069:
2039:
2033:
2029:
2023:
2022:
2016:
2008:
2003:. Archived from
1972:
1959:
1958:
1946:
1929:
1923:
1922:
1910:
1904:
1903:
1901:
1900:
1886:
1880:
1873:
1851:
1849:Dinosaurs portal
1846:
1845:
1844:
1773:very difficult.
1523:
1522:
1516:
1515:
1492:
1491:
1442:
1441:
1435:
1434:
1411:
1410:
1387:
1386:
1363:
1362:
1347:Eotyrannus lengi
1339:
1338:
1323:Dilong paradoxus
1315:
1314:
1247:
1246:
1223:
1222:
1156:
1155:
1149:
1148:
1139:
1138:
1133:Tyrannosauroidea
1129:
1128:
1064:Appalachiosaurus
1059:Appalachiosaurus
1005:was reviewed by
900:life restoration
775:Laelaps macropus
733:Laelaps macropus
660:Tyrannosauroidea
599:. However, four
587:, an incomplete
581:caudal vertebrae
530:excursions into
438:eutyrannosaurian
428:
424:
419:
418:
415:
414:
411:
408:
405:
402:
399:
396:
393:
390:
387:
384:
381:
358:
341:
313:
309:
276:
269:
254:
249:
236:
154:
153:
131:
121:
51:
40:Temporal range:
32:
31:
21:
6937:
6936:
6932:
6931:
6930:
6928:
6927:
6926:
6872:
6871:
6870:
6865:
6857:
6852:
6844:
6839:
6831:
6826:
6818:
6813:
6805:
6800:
6791:
6790:
6785:
6772:
6762:
6753:
6740:
6714:
6677:
6658:Sinornithomimus
6580:
6523:
6485:
6402:
6396:Compsognathidae
6384:
6371:
6359:Sinosauropteryx
6291:Compsognathidae
6285:
6264:
6255:
6223:
6210:
6193:
6192:
6178:
6151:
6146:Shanyangosaurus
6139:Phaedrolosaurus
6100:
6056:
6022:
5988:
5967:Daspletosaurini
5961:
5934:
5909:Tyrannosaurinae
5903:
5882:Albertosaurinae
5874:
5872:Tyrannosauridae
5865:
5834:
5832:Eutyrannosauria
5821:
5699:
5686:
5667:Proceratosaurus
5638:
5609:
5580:
5569:
5560:
5526:Richardoestesia
5510:Chilantaisaurus
5469:
5456:
5439:
5438:
5419:
5392:
5368:
5328:Australovenator
5306:
5300:Chilantaisaurus
5278:Phuwiangvenator
5200:
5165:Giganotosaurini
5159:
5135:
5047:
5010:Chilantaisaurus
4995:
4935:
4889:
4872:
4824:
4792:
4748:
4739:
4717:Monolophosaurus
4698:
4689:
4663:
4650:
4633:
4632:
4618:
4592:
4552:
4525:
4491:Camarillasaurus
4468:
4434:
4380:
4374:Ostafrikasaurus
4352:Camarillasaurus
4340:
4323:
4273:Afrovenatorinae
4267:
4226:
4202:
4180:
4167:
4125:
4104:
4102:Megalosauroidea
4093:
4084:
4079:Vectaerovenator
4072:Szechuanosaurus
4050:Monolophosaurus
4036:Kaijiangosaurus
4029:Cryolophosaurus
3990:Calamospondylus
3978:
3965:
3948:
3947:
3940:
3933:
3926:
3904:
3860:
3841:Pycnonemosaurus
3783:
3694:
3688:Majungasaurinae
3673:
3615:Majungasaurinae
3609:
3604:Xenotarsosaurus
3540:Dryptosauroides
3521:
3504:
3456:
3428:Elaphrosaurinae
3422:
3329:
3315:Ornithomimoides
3283:
3270:
3243:
3192:
3183:
3158:
3145:
3128:
3122:
3100:
3075:
3051:
2979:
2926:
2892:Shuangbaisaurus
2816:
2807:
2734:
2721:
2696:
2676:
2670:Avemetatarsalia
2655:Avemetatarsalia
2645:
2630:Avemetatarsalia
2604:
2603:
2593:
2588:
2549:
2524:
2523:
2518:
2514:
2499:
2495:
2434:
2430:
2361:Sertich, J.J.W.
2350:
2346:
2307:
2303:
2298:
2294:
2263:
2259:
2250:
2246:
2237:
2233:
2228:
2224:
2219:
2215:
2209:
2198:
2153:
2138:
2125:
2121:
2112:
2105:
2099:
2095:
2086:
2082:
2077:
2073:
2066:
2040:
2036:
2030:
2026:
2010:
2009:
1973:
1962:
1955:
1930:
1926:
1911:
1907:
1898:
1896:
1888:
1887:
1883:
1874:
1870:
1865:
1847:
1842:
1840:
1837:
1814:Leaping Laelaps
1802:Leaping Laelaps
1795:
1743:Late Cretaceous
1697:
1665:
1656:
1647:
1638:
1629:
1620:
1611:
1602:
1593:
1584:
1573:Tyrannosauridae
1560:
1531:Alioramus altai
1479:
1302:
1293:
1284:
1210:
1198:Guanlong wucaii
1100:Tyrannosauridae
969:
858:, three manual
829:tyrannosauroids
790:
779:ornithomimosaur
755:Thomas R. Holtz
704:
617:Pneumatoarthrus
512:
456:Late Cretaceous
426:
422:
378:
374:
322:
315:
307:
306:
293:
275:
267:
253:
247:
238:Eutyrannosauria
234:
148:
122:
120:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
46:
45:
42:Late Cretaceous
38:
28:
23:
22:
15:
12:
11:
5:
6935:
6925:
6924:
6919:
6914:
6909:
6904:
6899:
6894:
6889:
6884:
6867:
6866:
6864:
6863:
6850:
6837:
6824:
6811:
6798:
6782:
6780:
6774:
6773:
6759:
6758:
6755:
6754:
6734:
6733:
6731:
6728:
6727:
6724:
6723:
6720:
6719:
6716:
6715:
6713:
6712:
6711:
6710:
6704:
6695:
6693:
6687:
6686:
6683:
6682:
6679:
6678:
6676:
6675:
6668:
6661:
6654:
6647:
6640:
6633:
6626:
6623:Dromiceiomimus
6619:
6612:
6605:
6598:
6590:
6588:
6586:Ornithomimidae
6582:
6581:
6579:
6578:
6571:
6567:Paraxenisaurus
6563:
6555:
6548:
6541:
6533:
6531:
6529:Deinocheiridae
6525:
6524:
6522:
6521:
6518:Shenzhousaurus
6514:
6507:
6499:
6493:
6487:
6486:
6484:
6483:
6476:
6468:
6461:
6454:
6447:
6440:
6432:
6424:
6416:
6410:
6404:
6403:
6401:
6400:
6391:
6385:
6380:
6377:
6376:
6373:
6372:
6370:
6369:
6362:
6355:
6348:
6340:
6333:
6326:
6319:
6315:Beipiaognathus
6311:
6304:
6295:
6293:
6287:
6286:
6284:
6283:
6275:
6265:
6260:
6257:
6256:
6254:
6253:
6246:
6239:
6230:
6224:
6219:
6212:
6211:
6199:
6198:
6195:
6194:
6186:
6172:
6166:
6164:
6161:
6160:
6157:
6156:
6153:
6152:
6150:
6149:
6142:
6135:
6128:
6120:
6118:
6110:
6109:
6106:
6105:
6102:
6101:
6099:
6098:
6097:
6096:
6090:
6088:
6078:
6077:
6074:
6073:
6070:
6069:
6066:
6065:
6062:
6061:
6058:
6057:
6055:
6054:
6047:
6040:
6032:
6030:
6028:Tyrannosaurini
6024:
6023:
6021:
6020:
6013:
6006:
5998:
5996:
5994:Teratophoneini
5990:
5989:
5987:
5986:
5979:
5976:Daspletosaurus
5971:
5969:
5963:
5962:
5960:
5959:
5956:Qianzhousaurus
5952:
5944:
5942:
5936:
5935:
5933:
5932:
5925:
5917:
5911:
5905:
5904:
5902:
5901:
5894:
5886:
5884:
5875:
5870:
5867:
5866:
5864:
5863:
5856:
5853:Bistahieversor
5849:
5841:
5835:
5830:
5827:
5826:
5823:
5822:
5820:
5819:
5811:
5804:
5797:
5789:
5782:
5775:
5768:
5761:
5754:
5747:
5740:
5733:
5725:
5718:
5710:
5700:
5695:
5692:
5691:
5688:
5687:
5685:
5684:
5677:
5670:
5663:
5656:
5648:
5646:
5640:
5639:
5637:
5636:
5628:
5620:
5618:
5611:
5610:
5608:
5607:
5599:
5591:
5581:
5576:
5570:
5567:Tyrannoraptora
5565:
5562:
5561:
5559:
5558:
5551:
5544:
5537:
5529:
5522:
5514:
5506:
5499:
5491:
5484:
5476:
5470:
5465:
5458:
5457:
5445:
5444:
5441:
5440:
5432:
5413:
5407:
5405:
5402:
5401:
5398:
5397:
5394:
5393:
5391:
5390:
5389:
5388:
5382:
5380:
5374:
5373:
5370:
5369:
5367:
5366:
5359:
5352:
5345:
5338:
5331:
5324:
5316:
5314:
5312:Megaraptoridae
5308:
5307:
5305:
5304:
5296:
5288:
5281:
5274:
5267:
5259:
5251:
5243:
5237:
5226:
5225:
5222:
5221:
5218:
5217:
5214:
5213:
5210:
5209:
5206:
5205:
5202:
5201:
5199:
5198:
5191:
5184:
5177:
5174:Giganotosaurus
5169:
5167:
5161:
5160:
5158:
5157:
5149:
5143:
5137:
5136:
5134:
5133:
5126:
5119:
5112:
5105:
5098:
5091:
5084:
5077:
5069:
5061:
5055:
5049:
5048:
5046:
5045:
5037:
5029:
5022:
5014:
5005:
5003:
5001:Neovenatoridae
4997:
4996:
4994:
4993:
4986:
4983:Ulughbegsaurus
4979:
4971:
4964:
4957:
4949:
4943:
4937:
4936:
4934:
4933:
4925:
4917:
4909:
4901:
4899:
4890:
4885:
4882:
4881:
4878:
4877:
4874:
4873:
4871:
4870:
4863:
4856:
4849:
4842:
4834:
4832:
4826:
4825:
4823:
4822:
4815:
4811:Xuanhanosaurus
4806:
4800:
4794:
4793:
4791:
4790:
4782:
4778:Xuanhanosaurus
4774:
4767:
4764:Asfaltovenator
4759:
4749:
4744:
4741:
4740:
4738:
4737:
4729:
4725:Megalosauridae
4721:
4713:
4705:
4699:
4694:
4691:
4690:
4688:
4687:
4679:
4670:
4664:
4659:
4652:
4651:
4639:
4638:
4635:
4634:
4626:
4612:
4611:
4609:
4606:
4605:
4602:
4601:
4598:
4597:
4594:
4593:
4591:
4590:
4589:
4588:
4582:
4580:
4570:
4569:
4566:
4565:
4562:
4561:
4558:
4557:
4554:
4553:
4551:
4550:
4543:
4535:
4533:
4527:
4526:
4524:
4523:
4516:
4509:
4502:
4499:Ichthyovenator
4495:
4486:
4480:
4474:
4473:
4470:
4469:
4467:
4466:
4459:
4452:
4444:
4442:
4436:
4435:
4433:
4432:
4424:
4416:
4409:
4402:
4394:
4388:
4382:
4381:
4379:
4378:
4370:
4363:
4360:Cristatusaurus
4356:
4347:
4341:
4336:
4333:
4332:
4329:
4328:
4325:
4324:
4322:
4321:
4317:Poekilopleuron
4313:
4310:Piveteausaurus
4306:
4299:
4292:
4285:
4277:
4275:
4269:
4268:
4266:
4265:
4258:
4251:
4244:
4236:
4234:
4232:Megalosaurinae
4228:
4227:
4225:
4224:
4216:
4210:
4208:Megalosauridae
4204:
4203:
4201:
4200:
4191:
4181:
4176:
4173:
4172:
4169:
4168:
4166:
4165:
4161:Xuanhanosaurus
4157:
4150:
4143:
4135:
4133:
4127:
4126:
4124:
4123:
4120:Yunyangosaurus
4115:
4105:
4100:
4094:
4089:
4086:
4085:
4083:
4082:
4075:
4068:
4060:
4057:Pandoravenator
4053:
4046:
4043:Kayentavenator
4039:
4032:
4025:
4017:
4010:
4002:
3994:
3985:
3979:
3974:
3967:
3966:
3954:
3953:
3950:
3949:
3941:
3934:
3927:
3920:
3919:
3917:
3914:
3913:
3910:
3909:
3906:
3905:
3903:
3902:
3901:
3900:
3894:
3892:
3882:
3881:
3878:
3877:
3874:
3873:
3870:
3869:
3866:
3865:
3862:
3861:
3859:
3858:
3851:
3844:
3837:
3830:
3822:
3815:
3808:
3801:
3793:
3791:
3785:
3784:
3782:
3781:
3774:
3771:Skorpiovenator
3767:
3759:
3752:
3745:
3738:
3731:
3724:
3717:
3708:
3702:
3693:
3692:
3683:
3681:
3675:
3674:
3672:
3671:
3664:
3656:
3649:
3642:
3635:
3627:
3619:
3617:
3611:
3610:
3608:
3607:
3600:
3593:
3586:
3583:Spectrovenator
3579:
3572:
3565:
3558:
3551:
3544:
3536:
3528:
3522:
3517:
3514:
3513:
3510:
3509:
3506:
3505:
3503:
3502:
3495:
3488:
3481:
3474:
3466:
3464:
3458:
3457:
3455:
3454:
3447:
3440:
3432:
3430:
3424:
3423:
3421:
3420:
3417:Spinostropheus
3413:
3406:
3399:
3391:
3383:
3375:
3367:
3360:
3352:
3343:
3337:
3331:
3330:
3328:
3327:
3319:
3311:
3303:
3294:
3284:
3281:Abelisauroidea
3279:
3276:
3275:
3272:
3271:
3269:
3268:
3261:
3253:
3251:
3249:Ceratosauridae
3245:
3244:
3242:
3241:
3238:Saltriovenator
3234:
3226:
3219:
3215:Fosterovenator
3211:
3203:
3193:
3188:
3185:
3184:
3182:
3181:
3177:Lophostropheus
3173:
3165:
3159:
3154:
3147:
3146:
3134:
3133:
3130:
3129:
3116:
3115:
3113:
3110:
3109:
3106:
3105:
3102:
3101:
3099:
3098:
3091:
3083:
3081:
3077:
3076:
3074:
3073:
3072:
3071:
3065:
3063:
3057:
3056:
3053:
3052:
3050:
3049:
3042:
3038:Pterospondylus
3034:
3027:
3019:
3011:
3008:Lucianovenator
3004:
2997:
2989:
2987:
2981:
2980:
2978:
2977:
2970:
2963:
2956:
2949:
2940:
2934:
2932:Coelophysoidea
2928:
2927:
2925:
2924:
2917:
2909:
2902:
2895:
2888:
2881:
2874:
2871:Lophostropheus
2867:
2860:
2853:
2846:
2839:
2831:
2823:
2817:
2812:
2809:
2808:
2806:
2805:
2797:
2789:
2781:
2773:
2770:Erythrovenator
2766:
2758:
2750:
2741:
2735:
2730:
2723:
2722:
2710:
2709:
2706:
2705:
2702:
2701:
2698:
2697:
2695:
2694:
2693:
2692:
2686:
2684:
2678:
2677:
2675:
2674:
2673:
2672:
2663:
2657:
2647:
2646:
2642:
2641:
2632:
2623:
2617:
2611:
2595:
2594:
2587:
2586:
2579:
2572:
2564:
2558:
2557:
2548:
2547:External links
2545:
2544:
2543:
2538:
2522:
2521:
2512:
2493:
2428:
2389:(11): e79420.
2373:Evans, David C
2369:Sampson, S. D.
2344:
2301:
2292:
2273:(1): 119–143.
2257:
2244:
2231:
2222:
2213:
2196:
2173:10.1206/3717.2
2167:(3717): 1–53.
2136:
2119:
2103:
2093:
2089:The Dinosauria
2080:
2071:
2064:
2034:
2024:
2007:on 2007-03-10.
1987:(3): 561–573.
1960:
1953:
1924:
1905:
1881:
1867:
1866:
1864:
1861:
1860:
1859:
1853:
1852:
1836:
1833:
1794:
1791:
1696:
1693:
1685:Dryptosauridae
1673:Cryptotyrannus
1662:
1661:
1658:
1657:
1653:
1652:
1649:
1648:
1644:
1643:
1640:
1639:
1635:
1634:
1631:
1630:
1626:
1625:
1622:
1621:
1617:
1616:
1613:
1612:
1608:
1607:
1604:
1603:
1599:
1598:
1595:
1594:
1590:
1589:
1586:
1585:
1581:
1580:
1577:
1576:
1569:
1566:
1565:
1562:
1561:
1557:
1556:
1553:
1552:
1543:
1540:
1539:
1536:
1535:
1526:
1521:
1519:
1514:
1512:
1509:
1508:
1505:
1504:
1495:
1490:
1488:
1485:
1484:
1481:
1480:
1476:
1475:
1472:
1471:
1462:
1459:
1458:
1455:
1454:
1445:
1440:
1438:
1433:
1431:
1428:
1427:
1424:
1423:
1414:
1409:
1407:
1404:
1403:
1400:
1399:
1390:
1385:
1383:
1380:
1379:
1376:
1375:
1366:
1361:
1359:
1356:
1355:
1352:
1351:
1342:
1337:
1335:
1332:
1331:
1328:
1327:
1318:
1313:
1311:
1308:
1307:
1304:
1303:
1299:
1298:
1295:
1294:
1290:
1289:
1286:
1285:
1281:
1280:
1277:
1276:
1267:
1264:
1263:
1260:
1259:
1250:
1245:
1243:
1240:
1239:
1236:
1235:
1226:
1221:
1219:
1216:
1215:
1212:
1211:
1207:
1206:
1203:
1202:
1193:
1190:
1189:
1186:
1185:
1176:
1173:
1172:
1169:
1168:
1159:
1154:
1152:
1147:
1145:
1137:
1135:
1127:
1076:tyrannosauroid
1055:tyrannosauroid
968:
967:Classification
965:
789:
786:
759:tyrannosauroid
715:L. trihedrodon
703:
700:
511:
508:
364:
363:
362:
361:
348:
331:
330:
324:
323:
316:
302:
301:
295:
294:
286:
284:
280:
279:
265:
261:
260:
251:Dryptosauridae
245:
241:
240:
232:
225:
224:
219:
212:
211:
206:
199:
198:
193:
186:
185:
180:
176:
175:
170:
166:
165:
160:
156:
155:
142:
141:
133:
132:
124:
123:
115:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
39:
26:
9:
6:
4:
3:
2:
6934:
6923:
6920:
6918:
6915:
6913:
6910:
6908:
6905:
6903:
6900:
6898:
6895:
6893:
6890:
6888:
6885:
6883:
6880:
6879:
6877:
6860:
6855:
6851:
6847:
6842:
6838:
6834:
6829:
6825:
6821:
6816:
6812:
6808:
6803:
6799:
6794:
6788:
6784:
6783:
6781:
6779:
6775:
6771:
6766:
6751:
6747:
6745:
6738:
6709:
6705:
6703:
6699:
6698:
6697:
6696:
6694:
6692:
6688:
6674:
6673:
6669:
6667:
6666:
6665:Struthiomimus
6662:
6660:
6659:
6655:
6653:
6652:
6648:
6646:
6645:
6641:
6639:
6638:
6634:
6632:
6631:
6627:
6625:
6624:
6620:
6618:
6617:
6616:"Coelosaurus"
6613:
6611:
6610:
6606:
6604:
6603:
6599:
6597:
6596:
6592:
6591:
6589:
6587:
6583:
6577:
6576:
6572:
6569:
6568:
6564:
6561:
6560:
6556:
6554:
6553:
6549:
6547:
6546:
6542:
6540:
6539:
6535:
6534:
6532:
6530:
6526:
6520:
6519:
6515:
6513:
6512:
6511:Pelecanimimus
6508:
6506:
6505:
6501:
6500:
6497:
6494:
6492:
6488:
6482:
6481:
6477:
6474:
6473:
6472:Thecocoelurus
6469:
6467:
6466:
6462:
6460:
6459:
6455:
6453:
6452:
6451:Kinnareemimus
6448:
6446:
6445:
6441:
6438:
6437:
6433:
6430:
6429:
6425:
6423:
6422:
6418:
6417:
6414:
6411:
6409:
6405:
6398:
6397:
6393:
6392:
6389:
6386:
6383:
6378:
6368:
6367:
6363:
6361:
6360:
6356:
6354:
6353:
6349:
6346:
6345:
6341:
6339:
6338:
6334:
6332:
6331:
6330:Huaxiagnathus
6327:
6325:
6324:
6323:Compsognathus
6320:
6317:
6316:
6312:
6310:
6309:
6305:
6302:
6301:
6297:
6296:
6294:
6292:
6288:
6282:
6281:
6277:
6276:
6273:
6269:
6266:
6263:
6258:
6252:
6251:
6250:Ornitholestes
6247:
6245:
6244:
6240:
6237:
6236:
6232:
6231:
6228:
6225:
6222:
6217:
6213:
6209:
6204:
6200:
6190:
6185:
6183:
6176:
6170:
6148:
6147:
6143:
6141:
6140:
6136:
6134:
6133:
6129:
6127:
6126:
6122:
6121:
6119:
6115:
6111:
6094:
6093:
6092:
6091:
6089:
6087:
6083:
6079:
6053:
6052:
6048:
6046:
6045:
6044:Tyrannosaurus
6041:
6039:
6038:
6034:
6033:
6031:
6029:
6025:
6019:
6018:
6017:Teratophoneus
6014:
6012:
6011:
6007:
6005:
6004:
6000:
5999:
5997:
5995:
5991:
5985:
5984:
5980:
5978:
5977:
5973:
5972:
5970:
5968:
5964:
5958:
5957:
5953:
5951:
5950:
5946:
5945:
5943:
5941:
5937:
5931:
5930:
5926:
5924:
5923:
5919:
5918:
5915:
5912:
5910:
5906:
5900:
5899:
5895:
5893:
5892:
5891:Albertosaurus
5888:
5887:
5885:
5883:
5879:
5876:
5873:
5868:
5862:
5861:
5857:
5855:
5854:
5850:
5848:
5847:
5843:
5842:
5839:
5836:
5833:
5828:
5817:
5816:
5812:
5810:
5809:
5808:Xiongguanlong
5805:
5803:
5802:
5798:
5795:
5794:
5790:
5788:
5787:
5786:Suskityrannus
5783:
5781:
5780:
5779:Stokesosaurus
5776:
5774:
5773:
5772:Santanaraptor
5769:
5767:
5766:
5762:
5760:
5759:
5755:
5753:
5752:
5748:
5746:
5745:
5741:
5739:
5738:
5734:
5731:
5730:
5726:
5724:
5723:
5719:
5717:
5716:
5715:Alectrosaurus
5712:
5711:
5708:
5704:
5701:
5698:
5693:
5683:
5682:
5678:
5676:
5675:
5671:
5669:
5668:
5664:
5662:
5661:
5657:
5655:
5654:
5650:
5649:
5647:
5645:
5641:
5634:
5633:
5632:Tanycolagreus
5629:
5627:
5626:
5622:
5621:
5619:
5616:
5612:
5605:
5604:
5600:
5598:
5597:
5593:
5592:
5589:
5585:
5582:
5579:
5574:
5571:
5568:
5563:
5557:
5556:
5552:
5550:
5549:
5545:
5543:
5542:
5538:
5535:
5534:
5530:
5528:
5527:
5523:
5520:
5519:
5515:
5512:
5511:
5507:
5505:
5504:
5500:
5497:
5496:
5495:Bahariasaurus
5492:
5490:
5489:
5485:
5483:
5482:
5478:
5477:
5474:
5471:
5468:
5467:Coelurosauria
5463:
5459:
5455:
5454:Coelurosauria
5450:
5446:
5436:
5431:
5429:
5424:
5417:
5411:
5386:
5385:
5384:
5383:
5381:
5379:
5378:Coelurosauria
5375:
5365:
5364:
5360:
5358:
5357:
5353:
5351:
5350:
5346:
5344:
5343:
5339:
5337:
5336:
5332:
5330:
5329:
5325:
5323:
5322:
5318:
5317:
5315:
5313:
5309:
5303:
5301:
5297:
5295:
5293:
5289:
5287:
5286:
5282:
5280:
5279:
5275:
5273:
5272:
5268:
5265:
5264:
5260:
5257:
5256:
5255:Bahariasaurus
5252:
5250:
5249:
5245:
5244:
5241:
5238:
5235:
5231:
5227:
5197:
5196:
5192:
5190:
5189:
5185:
5183:
5182:
5178:
5176:
5175:
5171:
5170:
5168:
5166:
5162:
5156:
5155:
5151:
5150:
5147:
5144:
5142:
5138:
5132:
5131:
5127:
5125:
5124:
5120:
5118:
5117:
5113:
5111:
5110:
5106:
5104:
5103:
5099:
5097:
5096:
5095:Kelmayisaurus
5092:
5090:
5089:
5085:
5083:
5082:
5078:
5075:
5074:
5070:
5068:
5067:
5063:
5062:
5059:
5056:
5054:
5050:
5043:
5042:
5038:
5035:
5034:
5030:
5028:
5027:
5023:
5020:
5019:
5015:
5012:
5011:
5007:
5006:
5004:
5002:
4998:
4992:
4991:
4987:
4985:
4984:
4980:
4977:
4976:
4972:
4970:
4969:
4965:
4963:
4962:
4958:
4956:
4955:
4951:
4950:
4947:
4944:
4942:
4938:
4931:
4930:
4929:Saurophaganax
4926:
4923:
4922:
4918:
4915:
4914:
4910:
4908:
4907:
4903:
4902:
4900:
4898:
4894:
4891:
4888:
4883:
4869:
4868:
4864:
4862:
4861:
4860:Siamotyrannus
4857:
4855:
4854:
4850:
4848:
4847:
4843:
4841:
4840:
4836:
4835:
4833:
4831:
4827:
4821:
4820:
4816:
4813:
4812:
4808:
4807:
4804:
4801:
4799:
4795:
4788:
4787:
4783:
4780:
4779:
4775:
4773:
4772:
4768:
4766:
4765:
4761:
4760:
4757:
4753:
4750:
4747:
4746:Allosauroidea
4742:
4735:
4734:
4733:Spinosauridae
4730:
4727:
4726:
4722:
4719:
4718:
4714:
4712:
4711:
4707:
4706:
4703:
4700:
4697:
4692:
4685:
4684:
4680:
4677:
4676:
4672:
4671:
4668:
4665:
4662:
4657:
4653:
4649:
4644:
4640:
4630:
4625:
4623:
4616:
4586:
4585:
4584:
4583:
4581:
4579:
4575:
4571:
4549:
4548:
4544:
4542:
4541:
4537:
4536:
4534:
4532:
4528:
4522:
4521:
4517:
4515:
4514:
4510:
4508:
4507:
4503:
4501:
4500:
4496:
4493:
4492:
4488:
4487:
4484:
4481:
4479:
4478:Spinosaurinae
4475:
4465:
4464:
4460:
4458:
4457:
4456:Riparovenator
4453:
4451:
4450:
4449:Ceratosuchops
4446:
4445:
4443:
4441:
4437:
4430:
4429:
4425:
4422:
4421:
4417:
4415:
4414:
4413:Riojavenatrix
4410:
4408:
4407:
4403:
4401:
4400:
4396:
4395:
4392:
4389:
4387:
4386:Baryonychinae
4383:
4376:
4375:
4371:
4369:
4368:
4364:
4362:
4361:
4357:
4354:
4353:
4349:
4348:
4345:
4342:
4339:
4338:Spinosauridae
4334:
4319:
4318:
4314:
4312:
4311:
4307:
4305:
4304:
4300:
4298:
4297:
4293:
4291:
4290:
4286:
4284:
4283:
4279:
4278:
4276:
4274:
4270:
4264:
4263:
4262:Wiehenvenator
4259:
4257:
4256:
4252:
4250:
4249:
4245:
4243:
4242:
4238:
4237:
4235:
4233:
4229:
4223:
4222:
4218:
4217:
4214:
4211:
4209:
4205:
4198:
4197:
4193:
4192:
4189:
4185:
4182:
4179:
4174:
4163:
4162:
4158:
4156:
4155:
4151:
4149:
4148:
4144:
4142:
4141:
4137:
4136:
4134:
4132:
4128:
4122:
4121:
4117:
4116:
4113:
4109:
4106:
4103:
4098:
4095:
4092:
4087:
4081:
4080:
4076:
4074:
4073:
4069:
4066:
4065:
4061:
4059:
4058:
4054:
4052:
4051:
4047:
4045:
4044:
4040:
4038:
4037:
4033:
4031:
4030:
4026:
4023:
4022:
4018:
4016:
4015:
4011:
4008:
4007:
4003:
4000:
3999:
3998:Chienkosaurus
3995:
3992:
3991:
3987:
3986:
3983:
3980:
3977:
3972:
3968:
3964:
3959:
3955:
3945:
3938:
3931:
3924:
3898:
3897:
3896:
3895:
3893:
3891:
3887:
3883:
3857:
3856:
3852:
3850:
3849:
3845:
3843:
3842:
3838:
3836:
3835:
3831:
3828:
3827:
3823:
3821:
3820:
3816:
3814:
3813:
3809:
3807:
3806:
3802:
3800:
3799:
3795:
3794:
3792:
3790:
3789:Furileusauria
3786:
3780:
3779:
3775:
3773:
3772:
3768:
3765:
3764:
3763:Rahiolisaurus
3760:
3758:
3757:
3753:
3751:
3750:
3746:
3744:
3743:
3739:
3737:
3736:
3732:
3730:
3729:
3725:
3723:
3722:
3718:
3715:
3714:
3710:
3709:
3706:
3703:
3701:
3697:
3690:
3689:
3685:
3684:
3682:
3680:
3679:Carnotaurinae
3676:
3670:
3669:
3665:
3662:
3661:
3660:Rahiolisaurus
3657:
3655:
3654:
3653:Majungasaurus
3650:
3648:
3647:
3643:
3641:
3640:
3636:
3633:
3632:
3628:
3626:
3625:
3621:
3620:
3618:
3616:
3612:
3606:
3605:
3601:
3599:
3598:
3594:
3592:
3591:
3590:Tarascosaurus
3587:
3585:
3584:
3580:
3578:
3577:
3573:
3571:
3570:
3566:
3564:
3563:
3559:
3557:
3556:
3552:
3550:
3549:
3548:Eoabelisaurus
3545:
3542:
3541:
3537:
3535:
3534:
3533:Chenanisaurus
3530:
3529:
3526:
3523:
3520:
3519:Abelisauridae
3515:
3501:
3500:
3496:
3494:
3493:
3489:
3487:
3486:
3482:
3480:
3479:
3478:Masiakasaurus
3475:
3473:
3472:
3468:
3467:
3465:
3463:
3459:
3453:
3452:
3448:
3446:
3445:
3444:Huinculsaurus
3441:
3439:
3438:
3437:Elaphrosaurus
3434:
3433:
3431:
3429:
3425:
3419:
3418:
3414:
3412:
3411:
3407:
3405:
3404:
3400:
3397:
3396:
3392:
3389:
3388:
3384:
3381:
3380:
3376:
3373:
3372:
3368:
3366:
3365:
3361:
3358:
3357:
3356:Austrocheirus
3353:
3350:
3349:
3345:
3344:
3341:
3338:
3336:
3332:
3325:
3324:
3320:
3317:
3316:
3312:
3309:
3308:
3304:
3301:
3300:
3296:
3295:
3292:
3288:
3285:
3282:
3277:
3267:
3266:
3262:
3260:
3259:
3255:
3254:
3252:
3250:
3246:
3240:
3239:
3235:
3232:
3231:
3227:
3225:
3224:
3220:
3217:
3216:
3212:
3210:
3209:
3208:Berberosaurus
3205:
3204:
3201:
3197:
3194:
3191:
3186:
3179:
3178:
3174:
3172:
3171:
3167:
3166:
3163:
3160:
3157:
3152:
3148:
3144:
3139:
3135:
3126:
3120:
3097:
3096:
3095:Halticosaurus
3092:
3090:
3089:
3088:Teinurosaurus
3085:
3084:
3082:
3078:
3069:
3068:
3067:
3066:
3064:
3062:
3058:
3048:
3047:
3043:
3040:
3039:
3035:
3033:
3032:
3028:
3025:
3024:
3020:
3017:
3016:
3015:Megapnosaurus
3012:
3010:
3009:
3005:
3003:
3002:
2998:
2996:
2995:
2991:
2990:
2988:
2986:
2985:Coelophysidae
2982:
2976:
2975:
2974:Powellvenator
2971:
2969:
2968:
2964:
2962:
2961:
2957:
2955:
2954:
2950:
2947:
2946:
2942:
2941:
2938:
2935:
2933:
2929:
2923:
2922:
2918:
2915:
2914:
2910:
2908:
2907:
2903:
2901:
2900:
2896:
2894:
2893:
2889:
2887:
2886:
2882:
2880:
2879:
2875:
2873:
2872:
2868:
2866:
2865:
2864:Liliensternus
2861:
2859:
2858:
2854:
2852:
2851:
2847:
2845:
2844:
2840:
2837:
2836:
2835:Dolichosuchus
2832:
2830:
2829:
2828:Dilophosaurus
2825:
2824:
2821:
2818:
2815:
2810:
2803:
2802:
2801:Herrerasauria
2798:
2795:
2794:
2790:
2787:
2786:
2782:
2779:
2778:
2774:
2772:
2771:
2767:
2764:
2763:
2759:
2756:
2755:
2754:Daemonosaurus
2751:
2748:
2747:
2743:
2742:
2739:
2736:
2733:
2728:
2724:
2720:
2715:
2711:
2690:
2689:
2688:
2687:
2685:
2683:
2679:
2671:
2667:
2666:
2665:
2664:
2661:
2658:
2656:
2652:
2648:
2640:
2636:
2633:
2631:
2627:
2624:
2622:
2618:
2616:
2612:
2610:
2606:
2605:
2600:
2596:
2592:
2585:
2580:
2578:
2573:
2571:
2566:
2565:
2562:
2556:
2555:
2551:
2550:
2541:
2539:
2535:
2531:
2526:
2525:
2516:
2508:
2504:
2497:
2489:
2485:
2480:
2475:
2471:
2467:
2463:
2459:
2455:
2451:
2448:(8): 210127.
2447:
2443:
2439:
2432:
2424:
2420:
2415:
2410:
2405:
2400:
2396:
2392:
2388:
2384:
2383:
2378:
2374:
2370:
2366:
2365:Currie, P. J.
2362:
2358:
2354:
2348:
2339:
2334:
2329:
2324:
2320:
2316:
2312:
2305:
2296:
2288:
2284:
2280:
2276:
2272:
2268:
2261:
2254:
2248:
2241:
2235:
2226:
2217:
2207:
2205:
2203:
2201:
2192:
2188:
2183:
2178:
2174:
2170:
2166:
2162:
2160:
2151:
2149:
2147:
2145:
2143:
2141:
2133:
2129:
2123:
2116:
2110:
2108:
2097:
2090:
2084:
2075:
2067:
2061:
2057:
2053:
2049:
2047:
2038:
2032:Philadelphia.
2028:
2020:
2014:
2006:
2002:
1998:
1994:
1990:
1986:
1982:
1978:
1971:
1969:
1967:
1965:
1956:
1954:0-19-910207-4
1950:
1945:
1944:
1938:
1934:
1928:
1920:
1916:
1909:
1895:
1891:
1885:
1878:
1872:
1868:
1858:
1855:
1854:
1850:
1839:
1832:
1830:
1829:
1824:
1820:
1815:
1807:
1803:
1799:
1790:
1788:
1784:
1780:
1776:
1772:
1767:
1762:
1760:
1756:
1751:
1746:
1744:
1740:
1737:
1736:Maastrichtian
1733:
1729:
1725:
1721:
1719:
1714:
1706:
1701:
1692:
1690:
1686:
1682:
1678:
1674:
1670:
1660:
1659:
1651:
1650:
1642:
1641:
1633:
1632:
1624:
1623:
1615:
1614:
1606:
1605:
1597:
1596:
1588:
1587:
1579:
1578:
1575:
1574:
1568:
1567:
1564:
1563:
1555:
1554:
1551:
1550:
1549:
1542:
1541:
1538:
1537:
1534:
1533:
1532:
1525:
1524:
1518:
1517:
1511:
1510:
1507:
1506:
1503:
1502:
1501:
1494:
1493:
1487:
1486:
1483:
1482:
1474:
1473:
1470:
1469:
1468:
1461:
1460:
1457:
1456:
1453:
1452:
1451:
1444:
1443:
1437:
1436:
1430:
1429:
1426:
1425:
1422:
1421:
1420:
1413:
1412:
1406:
1405:
1402:
1401:
1398:
1397:
1396:
1389:
1388:
1382:
1381:
1378:
1377:
1374:
1373:
1372:
1365:
1364:
1358:
1357:
1354:
1353:
1350:
1349:
1348:
1341:
1340:
1334:
1333:
1330:
1329:
1326:
1325:
1324:
1317:
1316:
1310:
1309:
1306:
1305:
1297:
1296:
1288:
1287:
1279:
1278:
1275:
1274:
1273:
1266:
1265:
1262:
1261:
1258:
1257:
1256:
1249:
1248:
1242:
1241:
1238:
1237:
1234:
1233:
1232:
1225:
1224:
1218:
1217:
1214:
1213:
1205:
1204:
1201:
1200:
1199:
1192:
1191:
1188:
1187:
1184:
1183:
1182:
1175:
1174:
1171:
1170:
1167:
1166:
1165:
1158:
1157:
1151:
1150:
1144:
1141:
1140:
1134:
1131:
1130:
1126:
1124:
1115:
1107:
1103:
1101:
1097:
1096:
1091:
1090:
1085:
1081:
1077:
1073:
1069:
1065:
1061:
1060:
1056:
1052:
1048:
1047:
1046:Tyrannosaurus
1042:
1041:
1040:Albertosaurus
1036:
1032:
1028:
1023:
1021:
1017:
1013:
1008:
1007:Ken Carpenter
1004:
1000:
996:
992:
988:
987:
982:
973:
964:
962:
957:
952:
948:
943:
941:
940:
939:Tyrannosaurus
935:
934:
933:Albertosaurus
929:
925:
921:
917:
913:
905:
901:
896:
892:
889:
885:
881:
877:
873:
869:
865:
861:
857:
853:
849:
845:
841:
836:
834:
830:
826:
822:
818:
817:
816:Tyrannosaurus
812:
808:
807:
802:
794:
785:
784:
780:
776:
772:
770:
764:
760:
756:
752:
748:
747:
742:
738:
734:
730:
728:
724:
720:
716:
712:
708:
696:
692:
691:
686:
685:
680:
675:
671:
669:
665:
661:
657:
653:
649:
645:
644:Othniel Marsh
641:
640:Dryptosaurus,
637:
633:
629:
624:
620:
618:
614:
610:
606:
602:
598:
594:
590:
586:
582:
578:
574:
570:
566:
562:
557:
553:
549:
548:Maastrichtian
545:
541:
537:
533:
529:
525:
516:
507:
505:
502:
498:
494:
490:
486:
482:
478:
474:
469:
465:
462:was a large,
461:
457:
453:
452:Maastrichtian
449:
445:
442:
439:
435:
431:
430:
417:
372:
371:
357:
356:
354:
349:
345:
340:
339:
335:
334:
332:
329:
325:
320:
314:
312:
303:
300:
299:Binomial name
296:
292:
291:
285:
282:
281:
274:
273:
266:
263:
262:
257:
252:
246:
243:
242:
239:
233:
230:
227:
226:
223:
220:
217:
214:
213:
210:
207:
204:
201:
200:
197:
194:
191:
188:
187:
184:
181:
178:
177:
174:
171:
168:
167:
164:
161:
158:
157:
152:
147:
143:
139:
134:
130:
125:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
50:
47:67.6–66
43:
37:
33:
30:
19:
6882:Tyrannosaurs
6778:Dryptosaurus
6777:
6741:
6670:
6663:
6656:
6649:
6642:
6637:Ornithomimus
6635:
6628:
6621:
6614:
6607:
6600:
6593:
6575:Tyrannomimus
6573:
6565:
6557:
6550:
6545:Deinocheirus
6543:
6536:
6516:
6509:
6502:
6478:
6470:
6465:Nqwebasaurus
6463:
6458:Nedcolbertia
6456:
6449:
6442:
6436:Calamosaurus
6434:
6428:Aviatyrannis
6426:
6419:
6394:
6364:
6357:
6350:
6342:
6335:
6328:
6321:
6313:
6308:Aristosuchus
6306:
6300:Aniksosaurus
6298:
6280:Aniksosaurus
6278:
6248:
6241:
6233:
6179:
6144:
6137:
6130:
6123:
6049:
6042:
6035:
6015:
6008:
6003:Dynamoterror
6001:
5981:
5974:
5954:
5947:
5927:
5922:Asiatyrannus
5920:
5896:
5889:
5860:Dryptosaurus
5859:
5858:
5851:
5844:
5813:
5806:
5799:
5791:
5784:
5777:
5770:
5763:
5756:
5751:Jinbeisaurus
5749:
5742:
5735:
5727:
5722:Aviatyrannis
5720:
5713:
5679:
5674:Sinotyrannus
5672:
5665:
5658:
5651:
5630:
5623:
5601:
5594:
5553:
5546:
5539:
5531:
5524:
5516:
5508:
5503:Bicentenaria
5501:
5493:
5488:Asiamericana
5486:
5479:
5420:
5361:
5354:
5347:
5340:
5333:
5326:
5319:
5298:
5290:
5283:
5276:
5269:
5263:Deltadromeus
5261:
5253:
5246:
5195:Tyrannotitan
5193:
5186:
5179:
5172:
5152:
5128:
5123:Taurovenator
5121:
5114:
5107:
5102:Lajasvenator
5100:
5093:
5086:
5081:Concavenator
5079:
5071:
5064:
5039:
5031:
5024:
5016:
5008:
4988:
4981:
4973:
4966:
4959:
4952:
4927:
4919:
4911:
4904:
4897:Allosauridae
4865:
4858:
4853:Shidaisaurus
4851:
4844:
4837:
4817:
4809:
4784:
4776:
4769:
4762:
4731:
4723:
4715:
4708:
4681:
4673:
4661:Avetheropoda
4648:Avetheropoda
4619:
4578:Avetheropoda
4545:
4538:
4531:Spinosaurini
4518:
4511:
4504:
4497:
4489:
4461:
4454:
4447:
4426:
4418:
4411:
4406:Protathlitis
4404:
4397:
4372:
4365:
4358:
4350:
4315:
4308:
4301:
4296:Leshansaurus
4294:
4287:
4280:
4260:
4253:
4248:Megalosaurus
4246:
4241:Duriavenator
4239:
4219:
4194:
4178:Megalosauria
4159:
4152:
4147:Marshosaurus
4145:
4140:Condorraptor
4138:
4118:
4077:
4070:
4062:
4055:
4048:
4041:
4034:
4027:
4021:Cruxicheiros
4019:
4012:
4004:
3996:
3988:
3853:
3848:Quilmesaurus
3846:
3839:
3832:
3824:
3817:
3810:
3803:
3796:
3776:
3769:
3761:
3754:
3747:
3740:
3733:
3726:
3719:
3711:
3700:Brachyrostra
3686:
3666:
3658:
3651:
3644:
3637:
3629:
3622:
3602:
3597:Tralkasaurus
3595:
3588:
3581:
3574:
3567:
3560:
3553:
3546:
3538:
3531:
3499:Vespersaurus
3497:
3492:Velocisaurus
3490:
3483:
3476:
3469:
3449:
3442:
3435:
3415:
3408:
3401:
3393:
3387:Deltadromeus
3385:
3377:
3371:Compsosuchus
3369:
3362:
3354:
3346:
3321:
3313:
3307:Lametasaurus
3305:
3297:
3263:
3258:Ceratosaurus
3256:
3236:
3228:
3221:
3213:
3206:
3190:Ceratosauria
3175:
3168:
3093:
3086:
3044:
3036:
3029:
3021:
3013:
3006:
2999:
2992:
2972:
2967:Podokesaurus
2965:
2958:
2951:
2943:
2919:
2911:
2904:
2897:
2890:
2883:
2876:
2869:
2862:
2855:
2850:Gojirasaurus
2848:
2843:Dracovenator
2841:
2833:
2826:
2814:Neotheropoda
2799:
2791:
2783:
2777:Guaibasaurus
2775:
2768:
2760:
2752:
2746:Chindesaurus
2744:
2634:
2625:
2553:
2533:
2529:
2515:
2506:
2502:
2496:
2445:
2441:
2431:
2386:
2380:
2353:Loewen, M.A.
2347:
2318:
2314:
2304:
2295:
2270:
2266:
2260:
2252:
2247:
2239:
2234:
2225:
2216:
2164:
2158:
2127:
2122:
2114:
2096:
2088:
2083:
2074:
2051:
2045:
2037:
2027:
2013:cite journal
2005:the original
1984:
1980:
1942:
1937:Robert Scott
1927:
1918:
1914:
1908:
1897:. Retrieved
1893:
1884:
1876:
1871:
1828:Megalosaurus
1827:
1813:
1811:
1801:
1779:Dryptosaurus
1778:
1775:Hadrosaurids
1771:Dryptosaurus
1770:
1766:Dryptosaurus
1765:
1763:
1759:Dryptosaurus
1758:
1747:
1716:
1713:Dryptosaurus
1712:
1710:
1705:Dryptosaurus
1704:
1695:Paleoecology
1689:Dryptosaurus
1688:
1669:Dryptosaurus
1668:
1666:
1571:
1546:
1545:
1529:
1528:
1498:
1497:
1465:
1464:
1448:
1447:
1418:
1417:
1416:
1393:
1392:
1369:
1368:
1345:
1344:
1321:
1320:
1270:
1269:
1253:
1252:
1229:
1228:
1196:
1195:
1179:
1178:
1162:
1161:
1122:
1120:
1093:
1087:
1084:Dryptosaurus
1083:
1079:
1072:Dryptosaurus
1071:
1068:Dryptosaurus
1067:
1063:
1057:
1051:Dryptosaurus
1050:
1044:
1038:
1031:Dryptosaurus
1030:
1024:
1016:Dryptosaurus
1015:
1012:phylogenetic
1003:Dryptosaurus
1002:
995:megalosaurid
986:Megalosaurus
984:
981:Dryptosaurus
980:
978:
951:Dryptosaurus
950:
946:
944:
937:
931:
924:Dryptosaurus
923:
912:Dryptosaurus
911:
909:
904:Dryptosaurus
903:
898:An outdated
837:
833:Dryptosaurus
832:
825:Dryptosaurus
824:
821:Dryptosaurus
820:
814:
811:Dryptosaurus
810:
804:
801:Dryptosaurus
800:
799:
782:
774:
769:Teihivenator
766:
762:
751:Dryptosaurus
750:
744:
741:Joseph Leidy
732:
731:
722:
718:
714:
706:
705:
688:
684:Elasmosaurus
682:
679:Dryptosaurus
678:
667:
663:
651:
639:
631:
627:
621:An isolated
615:
591:, a partial
565:Philadelphia
521:
497:Dryptosaurus
496:
477:Dryptosaurus
476:
460:Dryptosaurus
459:
370:Dryptosaurus
369:
368:
367:
359:(Cope, 1866)
353:Megalosaurus
351:
350:
342:Cope, 1866 (
337:
336:
310:
305:
289:
288:
272:Dryptosaurus
271:
270:
250:
228:
215:
202:
189:
36:Dryptosaurus
35:
29:
6702:Maniraptora
6691:Maniraptora
6672:Tototlmimus
6602:Anserimimus
6552:Garudimimus
6538:Beishanlong
6480:Valdoraptor
6421:Arkansaurus
6366:Xunmenglong
6243:Juravenator
6037:Tarbosaurus
5929:Nanuqsaurus
5898:Gorgosaurus
5815:Megaraptora
5801:Timurlengia
5533:Sciurumimus
5481:Aratasaurus
5349:Murusraptor
5271:Fukuiraptor
5234:Megaraptora
5116:Shaochilong
5088:Eocarcharia
5041:Megaraptora
4961:Lusovenator
4839:Alpkarakush
4696:Carnosauria
4547:Spinosaurus
4520:Siamosaurus
4420:Suchosaurus
4367:Iberospinus
4303:Magnosaurus
4282:Afrovenator
4255:Torvosaurus
4006:Chilesaurus
3819:Carnotaurus
3812:Caletodraco
3798:Abelisaurus
3624:Arcovenator
3471:Laevisuchus
3462:Noasaurinae
3364:Berthasaura
3335:Noasauridae
3265:Genyodectes
3230:Lukousaurus
3223:Jubbulpuria
3023:Panguraptor
3001:Coelophysis
2994:Camposaurus
2953:Panguraptor
2945:Dracoraptor
2921:Zupaysaurus
2906:Tachiraptor
2885:Sarcosaurus
2785:Nhandumirim
2357:Irmis, R.B.
1020:coelurosaur
882:, the left
878:, the left
874:, the left
870:, the left
788:Description
746:Coelosaurus
690:Hadrosaurus
687:, with two
613:protostegid
575:fragments,
454:age of the
355:aquilunguis
344:preoccupied
277:Marsh, 1877
6876:Categories
6630:Gallimimus
6559:Harpymimus
6504:Harpymimus
6125:Iliosuchus
6095:see below↓
5940:Alioramini
5758:Juratyrant
5744:Eotyrannus
5729:Bagaraatan
5681:Yutyrannus
5615:Coeluridae
5541:Vayuraptor
5387:see below↓
5363:Tratayenia
5356:Orkoraptor
5342:Megaraptor
5248:Aoniraptor
5181:Mapusaurus
5109:Sauroniops
5073:Altispinax
5026:Neovenator
4968:Siamraptor
4954:Datanglong
4921:Epanterias
4913:Antrodemus
4906:Allosaurus
4887:Allosauria
4710:Altispinax
4675:Gasosaurus
4587:see below↓
4463:Suchomimus
4064:Sinosaurus
3899:see below↓
3855:Viavenator
3805:Aucasaurus
3742:Ilokelesia
3713:Dahalokely
3668:Rajasaurus
3646:Indosaurus
3639:Genusaurus
3631:Dahalokely
3555:Indosuchus
3451:Limusaurus
3410:Ligabueino
3403:Kiyacursor
3395:Genusaurus
3379:Dahalokely
3299:Betasuchus
3170:Dornraptor
3070:see below↓
3046:Segisaurus
2899:Sinosaurus
2762:Eodromaeus
2691:see below↓
2639:Dinosauria
2621:Sauropsida
2253:Brimleyana
2242:106: 1–19.
1921:: 275–279.
1899:2024-08-08
1863:References
1783:Appalachia
1732:New Jersey
916:incisiform
888:metatarsal
884:astragalus
848:surangular
806:Eotyrannus
763:Dinosauria
723:Allosaurus
719:Allosaurus
623:metatarsal
544:New Jersey
448:Appalachia
209:Saurischia
196:Dinosauria
6706:includes
6651:Rativates
6644:Qiupalong
6344:Scipionyx
6337:Mirischia
6010:Lythronax
5949:Alioramus
5603:Labocania
5321:Aerosteon
4867:Sinraptor
4771:Erectopus
4506:Irritator
4091:Orionides
3976:Tetanurae
3963:Tetanurae
3890:Tetanurae
3826:Elemgasem
3749:Llukalkan
3728:Elemgasem
3485:Noasaurus
3348:Afromimus
3156:Averostra
3143:Averostra
3061:Averostra
2913:Velocipes
2732:Theropoda
2719:Theropoda
2682:Theropoda
2607:Kingdom:
2591:Theropoda
2536:(3): 41A.
2470:2054-5703
2321:: 47–54.
2191:0003-0082
2182:2246/6117
2134:, p. 100.
1879:8: 72–73.
1787:Nodosaurs
1062:in 2005.
1025:In 1946,
999:monotypic
961:calcaneum
920:denticles
860:phalanges
852:vertebrae
648:Bone Wars
540:Barnsboro
504:dinosaurs
495:in 1877,
481:dinosaurs
468:carnivore
436:of basal
283:Species:
222:Theropoda
169:Kingdom:
163:Eukaryota
6807:52570689
6787:Wikidata
6235:Coelurus
5660:Kileskus
5653:Guanlong
5625:Coelurus
5518:Gualicho
5018:Gualicho
4399:Baryonyx
3735:Guemesia
3562:Kryptops
3323:Ozraptor
2960:Pendraig
2615:Chordata
2613:Phylum:
2609:Animalia
2488:34457333
2423:24223179
2382:PLoS ONE
2371:(2013).
2287:86243316
2255:2: 1–18.
1939:(1980).
1835:See also
1707:in brown
1681:Delaware
1125:(2013).
1089:Guanlong
949:(2011),
607:, and a
601:chevrons
597:hindlimb
501:theropod
458:period.
444:dinosaur
441:theropod
328:Synonyms
321:, 1866 )
244:Family:
183:Chordata
179:Phylum:
173:Animalia
159:Domain:
6833:4946657
6820:3238991
6793:Q131634
5793:Timimus
5555:Zuolong
5285:Rapator
5188:Meraxes
4513:Oxalaia
3834:Koleken
2857:Lepidus
2619:Class:
2509:: 1–58.
2479:8385347
2450:Bibcode
2414:3819173
2391:Bibcode
2375:(ed.).
2211:112-113
1989:Bibcode
1741:of the
956:ischial
926:had an
868:ischium
844:dentary
840:maxilla
668:Laelaps
632:Laelaps
609:scapula
605:sternum
573:dentary
532:Montana
526:during
464:bipedal
432:) is a
308:†
287:†
264:Genus:
116:↓
6444:Hexing
6132:Kakuru
5737:Dilong
3778:Thanos
3756:Niebla
3576:Rugops
3569:Kurupi
2486:
2476:
2468:
2421:
2411:
2285:
2189:
2062:
1951:
1935:&
1808:, 1897
1123:et al.
1095:Dilong
1080:et al.
1043:, and
1035:coeval
1018:was a
947:et al.
880:fibula
856:humeri
697:, 1869
664:Lælaps
652:et al.
593:pelvis
585:humeri
550:-aged
485:fossil
258:, 1890
6854:Plazi
6846:38605
6708:birds
5765:Moros
5292:Siats
5033:Siats
4975:Siats
2635:Clade
2626:Clade
2283:S2CID
2056:10–18
1739:stage
1033:with
876:tibia
872:femur
864:pubic
739:that
589:manus
577:teeth
569:skull
475:made
434:genus
425:-toh-
256:Marsh
229:Clade
216:Clade
203:Clade
190:Clade
6815:GBIF
6700:see
5335:Maip
2793:Tawa
2668:see
2484:PMID
2466:ISSN
2419:PMID
2187:ISSN
2060:ISBN
2019:link
1949:ISBN
1823:AMNH
1720:9995
1718:ANSP
1092:and
936:and
603:, a
571:and
423:DRIP
319:Cope
55:PreꞒ
6802:EoL
2474:PMC
2458:doi
2409:PMC
2399:doi
2333:hdl
2323:doi
2275:doi
2177:hdl
2169:doi
1997:doi
1804:by
1715:is
1679:of
902:of
666:or
630:. "
563:in
429:-əs
427:SOR
407:ɔːr
6878::
6856::
6843::
6830::
6817::
6804::
6789::
2637::
2628::
2534:22
2532:.
2507:34
2505:.
2482:.
2472:.
2464:.
2456:.
2444:.
2440:.
2417:.
2407:.
2397:.
2385:.
2379:.
2367:;
2363:;
2359:;
2355:;
2331:.
2319:58
2317:.
2313:.
2281:.
2271:25
2269:.
2199:^
2185:.
2175:.
2163:.
2139:^
2130:,
2106:^
2058:.
2015:}}
2011:{{
1995:.
1985:17
1983:.
1979:.
1963:^
1919:18
1917:.
1892:.
1730:,
1691:.
1102:.
809:,
729:.
583:,
579:,
542:,
398:oʊ
231::
218::
205::
192::
140:.
105:Pg
49:Ma
44:,
6570:?
6562:?
6475:?
6439:?
6431:?
6399:?
6347:?
6318:?
6303:?
6238:?
5818:?
5796:?
5732:?
5635:?
5617:?
5606:?
5536:?
5521:?
5513:?
5498:?
5302:?
5294:?
5266:?
5258:?
5236:?
5076:?
5044:?
5036:?
5021:?
5013:?
4978:?
4932:?
4924:?
4916:?
4814:?
4789:?
4781:?
4736:?
4728:?
4720:?
4686:?
4678:?
4494:?
4431:?
4423:?
4377:?
4355:?
4320:?
4199:?
4164:?
4067:?
4024:?
4009:?
4001:?
3993:?
3829:?
3766:?
3716:?
3691:?
3663:?
3634:?
3543:?
3398:?
3390:?
3382:?
3374:?
3359:?
3351:?
3326:?
3318:?
3310:?
3302:?
3233:?
3218:?
3180:?
3041:?
3026:?
3018:?
2948:?
2916:?
2838:?
2804:?
2796:?
2788:?
2780:?
2765:?
2757:?
2749:?
2583:e
2576:t
2569:v
2490:.
2460::
2452::
2446:8
2425:.
2401::
2393::
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