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Kingdom (biology)

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20: 5098: 5670: 5991: 453: 1590:, of Bacteria, Archaea, and Eukaryota. Combined with the five-kingdom model, this created a six-kingdom model, where the kingdom Monera is replaced by the kingdoms Bacteria and Archaea. This six-kingdom model is commonly used in recent US high school biology textbooks, but has received criticism for compromising the current scientific consensus. But the division of prokaryotes into two kingdoms remains in use with the recent 500:, for "neutral organisms" or "the kingdom of primitive forms", which were neither animal nor plant; he did not include the Regnum Lapideum in his scheme. Haeckel revised the content of this kingdom a number of times before settling on a division based on whether organisms were unicellular (Protista) or multicellular (animals and plants). 5259:
According to genetic data, although eukaryote groups such as plants, fungi, and animals may look different, they are more closely related to each other than they are to either the Eubacteria or Archaea. It was also found that the eukaryotes are more closely related to the Archaea than they are to the
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Following publication of Whittaker's system, the five-kingdom model began to be commonly used in high school biology textbooks. But despite the development from two kingdoms to five among most scientists, some authors as late as 1975 continued to employ a traditional two-kingdom system of animals and
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In 1990, the name "domain" was proposed for the highest rank. This term represents a synonym for the category of dominion (lat. dominium), introduced by Moore in 1974. Unlike Moore, Woese et al. (1990) did not suggest a Latin term for this category, which represents a further argument supporting the
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However, in the same year as the International Society of Protistologists' classification was published (2005), doubts were being expressed as to whether some of these supergroups were monophyletic, particularly the Chromalveolata, and a review in 2006 noted the lack of evidence for several of the
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On this basis, the diagram opposite (redrawn from their article) showed the real "kingdoms" (their quotation marks) of the eukaryotes. A classification which followed this approach was produced in 2005 for the International Society of Protistologists, by a committee which "worked in collaboration
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In 1998, Cavalier-Smith published a six-kingdom model, which has been revised in subsequent papers. The version published in 2009 is shown below. Cavalier-Smith no longer accepted the importance of the fundamental Eubacteria–Archaebacteria divide put forward by Woese and others and supported by
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in 2009 noted that "The deep phylogeny of eukaryotes is an extremely difficult and controversial problem." As of December 2010, there appears to be a consensus that the six supergroup model proposed in 2005 does not reflect the true phylogeny of the eukaryotes and hence how they should be
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The most recent research does not support the classification of the eukaryotes into any of the standard systems. As of April 2010, no set of kingdoms is sufficiently supported by research to attain widespread acceptance. In 2009, Andrew Roger and Alastair Simpson emphasized the need for
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At first, microscopic organisms were classified within the animal and plant kingdoms. However, by the mid–19th century, it had become clear to many that "the existing dichotomy of the plant and animal kingdoms rapidly blurred at its boundaries and outmoded".
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of London a copy of his first observations of microscopic single-celled organisms. Until then, the existence of such microscopic organisms was entirely unknown. Despite this, Linnaeus did not include any microscopic creatures in his original taxonomy.
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The remaining two kingdoms, Protista and Monera, included unicellular and simple cellular colonies. The five kingdom system may be combined with the two empire system. In the Whittaker system, Plantae included some algae. In other systems, such as
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that are not animals, plants or fungi". They held that only monophyletic groups should be accepted as formal ranks in a classification and that – while this approach had been impractical previously (necessitating "literally dozens of eukaryotic
850:. The resulting five-kingdom system, proposed in 1969 by Whittaker, has become a popular standard and with some refinement is still used in many works and forms the basis for new multi-kingdom systems. It is based mainly upon differences in 6118: 841:
and other organisms regarded as plants had long been recognised by some; Haeckel had moved the fungi out of Plantae into Protista after his original classification, but was largely ignored in this separation by scientists of his time.
229:) are the two ranks immediately below kingdom. Superkingdom may be considered as an equivalent of domain or empire or as an independent rank between kingdom and domain or subdomain. In some classification systems the additional rank 5260:
Eubacteria. Although the primacy of the Eubacteria-Archaea divide has been questioned, it has been upheld by subsequent research. There is no consensus on how many kingdoms exist in the classification scheme proposed by Woese.
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Raoult, Didier; Audic, StĂ©phane; Robert, Catherine; Abergel, Chantal; Renesto, Patricia; Ogata, Hiroyuki; La Scola, Bernard; Suzan, Marie; Claverie, Jean-Michel (2004). "The 1.2 megabase genome sequence of Mimivirus".
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In the same way, his paraphyletic kingdom Protozoa includes the ancestors of Animalia, Fungi, Plantae, and Chromista. The advances of phylogenetic studies allowed Cavalier-Smith to realize that all the phyla thought to be
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The IUCN Species Survival Commission calls for the due recognition of fungi as major components of biodiversity in legislation and policy. It fully endorses the Fauna Flora Funga Initiative and asks that the phrases
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outside of a host cell. Another argument is that their placement in the tree would be problematic, since it is suspected that viruses have various evolutionary origins, and they have a penchant for harvesting
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with specialists from many societies". It divided the eukaryotes into the same six "supergroups". The published classification deliberately did not use formal taxonomic ranks, including that of "kingdom".
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Cavalier-Smith and his collaborators revised their classification in 2015. In this scheme they introduced two superkingdoms of Prokaryota and Eukaryota and seven kingdoms. Prokaryota have two kingdoms:
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Ruggiero, Michael A.; Gordon, Dennis P.; Orrell, Thomas M.; Bailly, Nicolas; Bourgoin, Thierry; Brusca, Richard C.; Cavalier-Smith, Thomas; Guiry, Michael D.; Kirk, Paul M.; Thuesen, Erik V. (2015).
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Ruggiero, Michael A.; Gordon, Dennis P.; Orrell, Thomas M.; Bailly, Nicolas; Bourgoin, Thierry; Brusca, Richard C.; Cavalier-Smith, Thomas; Guiry, Michael D.; Kirk, Paul M.; Thuesen, Erik V. (2015).
2204:; the bimembranous-unimembranous transition was thought to be far more fundamental than the long branch of genetic distance of Archaebacteria, viewed as having no particular biological significance. 8068: 685:
promoted and popularized Édouard Chatton's earlier work, particularly in their paper of 1962, "The Concept of a Bacterium"; this created, for the first time, a rank above kingdom—a
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Whittaker, R.H. (January 1969). "New concepts of kingdoms or organisms. Evolutionary relations are better represented by new classifications than by the traditional two kingdoms".
670:, had placed the blue-green algae (or Phycochromacea) in Monera; this would gradually gain acceptance, and the blue-green algae would become classified as bacteria in the phylum 8173:
Burki, Fabien; Shalchian-Tabrizi, Kamran; Minge, Marianne; Skjæveland, Åsmund; Nikolaev, Sergey I.; Jakobsen, Kjetill S. & Pawlowski, Jan (2007). Butler, Geraldine (ed.).
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and colleagues proposed the fundamental subdivision of the prokaryotes into the Eubacteria (later called the Bacteria) and Archaebacteria (later called the Archaea), based on
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While the concept of kingdoms continues to be used by some taxonomists, there has been a movement away from traditional kingdoms, as they are no longer seen as providing a
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plants, dividing the plant kingdom into subkingdoms Prokaryota (bacteria and cyanobacteria), Mycota (fungi and supposed relatives), and Chlorota (algae and land plants).
6119:"Re:wild and IUCN SSC become first global organizations to call for the recognition of fungi as one of three kingdoms of life critical to protecting and restoring Earth" 1815:
was so great (particularly considering the genetic distance of ribosomal genes) that the prokaryotes needed to be separated into two different kingdoms. He then divided
8384:"Analysis of rare genomic changes does not support the unikont–bikont phylogeny, and suggests cyanobacterial symbiosis as the point of primary radiation of eukaryotes" 490:, a third kingdom of life composed of "all the lower creatures, or the primary organic beings"; he retained Regnum Lapideum as a fourth kingdom of minerals. In 1866, 2215:(therefore "monophyletic" in the sense he uses this term) taxa, marking important innovations of biological significance (in regard of the concept of biological 5712:
to their classifications. This can create a "superdomain" called "Acytota", also called "Aphanobionta", of non-cellular life; with the other superdomain being "
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diligence in analyzing new discoveries: "With the current pace of change in our understanding of the eukaryote tree of life, we should proceed with caution."
3298: 8290:"Large-scale phylogenomic analyses reveal that two enigmatic protist lineages, Telonemia and Centroheliozoa, are related to photosynthetic Chromalveolates" 5975:
On the other hand, there are arguments in favor of their inclusion. One of these comes from the discovery of unusually large and complex viruses, such as
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The kingdom-level classification of life is still widely employed as a useful way of grouping organisms, notwithstanding some problems with this approach:
3341: 1855:. Since then, many non-photosynthetic phyla of protists, thought to have secondarily lost their chloroplasts, were integrated into the kingdom Chromista. 4058: 3326: 3278: 3218: 2669: 2609: 1870:. As a result, these amitochondriate protists were separated from the protist kingdom, giving rise to the, at the same time, superkingdom and kingdom 4182: 4053: 3847: 3686: 3293: 3258: 3164: 3139: 2997: 8987: 4157: 4099: 3754: 3386: 3381: 3366: 3331: 3283: 3248: 3129: 3094: 3084: 3079: 3069: 2704: 2624: 2599: 2584: 2574: 2227:(i.e. primitively amitochondriate eukaryotes) had in fact secondarily lost their mitochondria, typically by transforming them into new organelles: 456:
Haeckel's original (1866) conception of the three kingdoms of life, including the new kingdom Protista. Notice the inclusion of the cyanobacterium
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International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress, Vienna, Austria, July 2005
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or Eubacteria), while textbooks in other parts of the world, such as the United Kingdom, Pakistan, Bangladesh, India, Greece, Brazil, Spain use
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Parfrey, Laura W.; Barbero, Erika; Lasser, Elyse; Dunthorn, Micah; Bhattacharya, Debashish; Patterson, David J. & Katz, Laura A. (2006).
4268: 4224: 4162: 4104: 4094: 3565: 3356: 3346: 3233: 3119: 3104: 3063: 3002: 2785: 2780: 2689: 2589: 2189: 8288:
Burki, F.; Inagaki, Y.; Brate, J.; Archibald, J. M.; Keeling, P. J.; Cavalier-Smith, T.; Sakaguchi, M.; Hashimoto, T.; et al. (2009).
6995: 5108: 4167: 4109: 3827: 3630: 3625: 3228: 3182: 3149: 2972: 2934: 2929: 2914: 2800: 2744: 5151:
thought life could be divided into three large divisions and referred to them as the "three primary kingdom" model or "urkingdom" model.
4142: 3931: 3812: 3759: 3676: 3575: 3550: 3530: 3204: 3030: 2821: 6712: 6386: 3797: 3610: 3509: 3197: 3055: 2211:("holophyletic" in his terminology) to be valid. He defines Prokaryota, Bacteria, Negibacteria, Unibacteria, and Posibacteria as valid 6343:
Scamardella, Joseph M. (1999). "Not plants or animals: a brief history of the origin of Kingdoms Protozoa, Protista and Protoctista".
9014: 5114: 4147: 3946: 2952: 2831: 8978: 8341:"Phylogenomic analysis supports the monophyly of cryptophytes and haptophytes and the association of Rhizaria with chromalveolates" 6067: 5106: 2891: 5111: 6659:
Balch, W.E.; Magrum, L.J.; Fox, G.E.; Wolfe, C.R. & Woese, C.R. (August 1977). "An ancient divergence among the bacteria".
5116: 6640: 5166:, stressing that there was as much genetic difference between these two groups as between either of them and all eukaryotes. 8969: 5289:") – it had now become possible to divide the eukaryotes into "just a few major groups that are probably all monophyletic". 166:
into biology in 1735, the highest rank was given the name "kingdom" and was followed by four other main or principal ranks:
5689:, so that Rhizaria is not one of the main eukaryote groups. Beyond this, there does not appear to be a consensus. Rogozin 5117: 8017: 5703: 5127: 5109: 2266:, the diagram below is an 'organization chart', not an 'ancestor chart', and does not represent an evolutionary tree. 8554: 7533: 6551: 5115: 288: 7721: 5107: 7610: 6260: 5126: 5681:
As of 2010, there is widespread agreement that the Rhizaria belong with the Stramenopiles and the Alveolata, in a
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can be included in the tree of life. The arguments against include the fact that they are obligate intracellular
5112: 4343:). The Eukaryota have five kingdoms: Protozoa, Chromista, Plantae, Fungi, and Animalia. In this classification a 5128: 5125: 5118: 5110: 2235:, according to the significance of the term given by Cavalier-Smith. Some of the members of the defunct kingdom 1858:
Finally, some protists lacking mitochondria were discovered. As mitochondria were known to be the result of the
7296:; Balch, W.E.; Magrum, L.J.; Fox, G.E.; Wolfe, R.S. (August 1977). "An ancient divergence among the bacteria". 6661: 6153: 5120: 658:
of prokaryotic organisms; as a revised phylum Monera of the Protista, it included organisms now classified as
9007: 8715: 5105: 25: 5119: 9408: 8388: 8294: 7247: 7106:
Systemae Naturae, sive regna tria naturae, systematics proposita per classes, ordines, genera & species
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Systemae Naturae, sive regna tria naturae, systematics proposita per classes, ordines, genera & species
4653: 3168:                Superorder 3163:                Superorder 843: 29: 8947: 8769:
Moreira, David; PurificaciĂłn LĂłpez-GarcĂ­a (2009). "Ten reasons to exclude viruses from the tree of life".
7936: 9403: 8848: 8771: 1866:, it was thought that these amitochondriate eukaryotes were primitively so, marking an important step in 24: 6433: 5103: 28: 27: 6906: 6345: 5124: 5104: 7223: 6505: 5158:
Woese divided the prokaryotes (previously classified as the Kingdom Monera) into two groups, called
5101: 31: 9000: 6582: 327: 8339:
Hackett, J.D.; Yoon, H.S.; Li, S.; Reyes-Prieto, A.; Rummele, S.E. & Bhattacharya, D. (2007).
5121: 5102: 8815: 7978: 1902:, which has since been abandoned; later schemes did not include the Archezoa–Metakaryota divide. 467: 299: 26: 7414:"Towards a natural system of organisms: proposal for the domains Archaea, Bacteria, and Eucarya" 3778:              Infraphylum 3773:              Infraphylum 3763:              Infraphylum 3758:              Infraphylum 3723:              Infraphylum 3718:              Infraphylum 3713:              Infraphylum 3695:              Infraphylum 3690:              Infraphylum 3629:              Infraphylum 3624:              Infraphylum 3579:              Infraphylum 3574:              Infraphylum 3569:              Infraphylum 9254: 8655:"Eocytes: A new ribosome structure indicates a kingdom with a close relationship to eukaryotes" 7660: 6987: 6500: 5123: 1832: 1824: 1594:
scheme of Thomas Cavalier-Smith, although it primarily differs in that Protista is replaced by
447: 5122: 30: 8010:"The new higher-level classification of eukaryotes with emphasis on the taxonomy of protists" 7717: 7656: 7606: 7509: 7471: 6934: 4679: 3158:              Subclass 3153:              Subclass 3143:              Subclass 3138:              Subclass 1844: 1804: 483: 6724: 8897: 8668: 8607: 8452: 8188: 7782: 7425: 7363: 7305: 7259: 6822: 6765: 6720: 6670: 6492: 6205: 6158:(electronic ed.). Vienna: International Association for Plant Taxonomy. Archived from 5969: 5271: 4348: 1118: 1112: 631:
revealed important distinctions between those organisms whose cells do not have a distinct
8: 5749: 5140: 5088: 4728: 1899: 8975: 8901: 8672: 8611: 8456: 8192: 8008:
Adl SM, Simpson AGB, Farmer MA, Andersen RA, Anderson OR, Barta JR, et al. (2005).
7786: 7429: 7367: 7309: 7263: 6826: 6769: 6674: 6496: 6209: 6173: 6074: 23: 9183: 9127: 9112: 9094: 8921: 8796: 8630: 8593: 8572: 8527: 8475: 8438: 8410: 8383: 8316: 8289: 8265: 8238: 8211: 8174: 8135: 8106: 8044: 7959: 7917: 7861: 7805: 7770: 7746: 7688: 7638: 7626: 7329: 7201: 7078: 7051: 7017: 6963: 6938: 6898: 6846: 6788: 6751: 6694: 6599: 6458: 6408: 6381: 6291: 6279: 6266: 6019: 5047: 4336: 3645: 3376: 2168: 1531: 1269: 303: 264: 35: 8691: 8654: 8009: 7583: 7558: 7386: 7351: 7163: 7137: 4639: 644: 9244: 9198: 9193: 8913: 8888: 8867: 8788: 8696: 8635: 8560: 8550: 8519: 8480: 8434: 8415: 8364: 8321: 8270: 8216: 8140: 8087: 8036: 8031: 7921: 7909: 7853: 7810: 7751: 7693: 7630: 7588: 7539: 7529: 7491: 7487: 7453: 7448: 7413: 7391: 7321: 7275: 7141: 7083: 7022: 6968: 6890: 6838: 6793: 6686: 6636: 6625: 6547: 6518: 6483: 6450: 6413: 6354: 6314: 6283: 6233: 6228: 6193: 5790: 5766: 5744: 5717: 5709: 5277: 5132: 4741: 4736: 4723: 4718: 4713: 4708: 4340: 3361: 3243: 694: 8925: 8531: 7865: 7333: 7205: 6698: 6462: 5940:
uses the taxonomic rank "kingdom" in the classification of viruses (with the suffix
4272:            Infraphylum 4267:            Infraphylum 3988:            Infraphylum 3983:            Infraphylum 3970:            Infraphylum 3965:            Infraphylum 3945:            Infraphylum 3940:            Infraphylum 3917:            Infraphylum 3912:            Infraphylum 3902:            Infraphylum 3897:            Infraphylum 3846:            Infraphylum 3841:            Infraphylum 3831:            Infraphylum 3826:            Infraphylum 3806:            Infraphylum 3801:            Infraphylum 2668:            Infraphylum 2663:            Infraphylum 2548:            Superphylum 697:
of prokaryotes and eukaryotes. The two-empire system would later be expanded to the
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introduced the terms "prokaryote" and "eukaryote" to differentiate these organisms.
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Hegde, Nagendra; Maddur, Mohan S.; Kaveri, Srini V. & Bayry, Jagadeesh (2009).
8800: 8780: 8686: 8676: 8625: 8615: 8511: 8470: 8460: 8405: 8397: 8354: 8311: 8303: 8260: 8252: 8206: 8196: 8130: 8120: 8077: 8048: 8026: 7951: 7899: 7843: 7800: 7790: 7741: 7733: 7683: 7675: 7642: 7622: 7578: 7574: 7570: 7521: 7483: 7443: 7433: 7381: 7371: 7313: 7267: 7229: 7219: 7193: 7181: 7073: 7065: 7012: 7004: 6958: 6950: 6902: 6882: 6873: 6850: 6830: 6813: 6783: 6773: 6678: 6591: 6510: 6442: 6403: 6399: 6395: 6295: 6275: 6223: 6213: 5964: 5945: 5694:
classified, although there is no agreement as to the model which should replace it.
5659: 5503: 5092: 4646: 3074: 2895:            Superclass 2890:            Superclass 2216: 698: 651: 207: 195: 63: 43: 8515: 8239:"Phylogenomics reveals a new 'megagroup' including most photosynthetic eukaryotes" 6194:"Towards a natural systs: proposal for the domains Archaea, Bacteria, and Eucarya" 3768:            Subphylum 3753:            Subphylum 3738:            Subphylum 3733:            Subphylum 3708:            Subphylum 3700:            Subphylum 3685:            Subphylum 3644:            Subphylum 3639:            Subphylum 3634:            Subphylum 3619:            Subphylum 3594:            Subphylum 3589:            Subphylum 3564:            Subphylum 3559:            Subphylum 3544:            Subphylum 3539:            Subphylum 3529:            Subphylum 3524:            Subphylum 9344: 9330: 9297: 9286: 9280: 9234: 9229: 9224: 9218: 9212: 9207: 9173: 9168: 9163: 9158: 9152: 9146: 8982: 8491: 8465: 8243: 8201: 8125: 8064:"On the monophyly of chromalveolates using a six-protein phylogeny of eukaryotes" 7890: 7834: 7795: 7271: 6778: 6537: 6514: 3966: 3817: 3749: 3744: 3701: 3385:            Subclass 3380:            Subclass 3370:            Subclass 3365:            Subclass 3360:            Subclass 3223: 2684: 2679: 2604: 1863: 1843:
kingdom. Indeed, the chloroplast of the chromists is located in the lumen of the
1429: 1414: 1297: 191: 187: 171: 167: 73:
Traditionally, some textbooks from the United States and Canada used a system of
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Lake, James A.; Henderson, Eric; Oakes, Melanie; Clark, Michael W. (June 1984).
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A short history of biology, a general introduction to the study of living things
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Proceedings of the National Academy of Sciences of the United States of America
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Proceedings of the National Academy of Sciences of the United States of America
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Proceedings of the National Academy of Sciences of the United States of America
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In 2004, a review article by Simpson and Roger noted that the Protista were "a
5214: 4852: 4842: 3807: 2977: 2962: 2851: 2815: 2739: 2724: 2714: 2654: 2564: 2307: 2197: 2188:) was subdivided into two sub-kingdoms according to their membrane topologies: 1955: 1867: 1835:). Technological advances in electron microscopy allowed the separation of the 1812: 1656: 1435: 1305: 682: 258:
The classification of living things into animals and plants is an ancient one.
59: 39: 7904: 7885: 7848: 7829: 7525: 3148:            Class 3133:            Class 3123:            Class 3118:            Class 9397: 9373: 8439:"EEF2 analysis challenges the monophyly of Archaeplastida and Chromalveolata" 8111: 7119: 7101: 6009: 5831: 4632: 4625: 4084: 4065: 3989: 3883: 3878: 3873: 3854: 3832: 3792: 3434: 3429: 3424: 3336: 3310: 3273: 3176: 2919: 2903: 2664: 2644: 2639: 2614: 2554: 2240: 2228: 1859: 1852: 1583: 1543: 1499: 1441: 1365: 1323: 1263: 1194: 1130: 871: 678: 671: 491: 471: 295: 242: 182:. Later two further main ranks were introduced, making the sequence kingdom, 159: 8909: 8681: 8620: 8564: 8499: 8359: 8340: 7722:"Kingdoms Protozoa and Chromista and the eozoan root of the eukaryotic tree" 7438: 7376: 6939:"Kingdoms Protozoa and Chromista and the eozoan root of the eukaryotic tree" 6811:
Cavalier-Smith, Thomas (March 26, 1987). "Eucaryotes with no mitochondria".
6218: 8917: 8871: 8862: 8843: 8823: 8792: 8639: 8523: 8484: 8419: 8368: 8325: 8274: 8256: 8220: 8144: 8107:"Evaluating support for the current classification of eukaryotic diversity" 8091: 8040: 7986: 7913: 7857: 7814: 7755: 7737: 7697: 7679: 7233: 7087: 7026: 7008: 6972: 6954: 6894: 6797: 6557: 6454: 6417: 6358: 6318: 5663: 5594: 5524: 5488: 4288: 4192: 4187: 4152: 4132: 3908: 3868: 3779: 3663: 3525: 3486: 3444: 3253: 3109: 2896: 2861: 2856: 2795: 2734: 2729: 2709: 2699: 2694: 2629: 2619: 2579: 2559: 2539: 2201: 2193: 1828: 1820: 1549: 1537: 1525: 1232: 1136: 1094: 1074: 1068: 632: 269: 163: 150:(for fungi) are also used for life present in a particular region or time. 8700: 8082: 8063: 7634: 7592: 7543: 7495: 7457: 7279: 6842: 6522: 6287: 6237: 8401: 8307: 7395: 7325: 7069: 6690: 6632: 6431:
Stanier, R.Y. & Van Neil, C.B. (1962). "The concept of a bacterium".
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contains one or more kingdoms. Intermediate minor rankings are not shown.
8784: 7352:"Phylogenetic structure of the prokaryotic domain: the primary kingdoms" 6603: 6539:
Kingdoms and Domains: An Illustrated Guide to the Phyla of Life on Earth
5666:. Plants are thought to be more distantly related to animals and fungi. 4191:          Infraphylum 4186:          Infraphylum 4176:          Infraphylum 4171:          Infraphylum 4166:          Infraphylum 4161:          Infraphylum 4146:          Infraphylum 4141:          Infraphylum 4108:          Infraphylum 4103:          Infraphylum 4093:          Infraphylum 4088:          Infraphylum 2885:          Infraphylum 2880:          Infraphylum 2743:          Infraphylum 2738:          Infraphylum 2713:          Infraphylum 2708:          Infraphylum 9356: 7963: 7409: 7347: 7317: 7293: 6682: 6446: 6004: 5960: 5836: 5673:
One hypothesis of eukaryotic relationships depicted by Alastair Simpson
5605: 5569: 5565: 5553: 5528: 5449: 5193: 5148: 5072: 5055: 4825: 4815: 4800: 4660: 4540: 4508: 4374: 4311: 4305: 4258: 4116: 4048: 4042: 3979: 3893: 3837: 3739: 3681: 3635: 3620: 3585: 3540: 3413: 3212: 3128:          Superclass 3113:          Superclass 3035: 2992: 2924: 2909: 2775: 2549: 2444: 2412: 2289: 2252: 2185: 1939: 1925: 1816: 1808: 1639: 1625: 1579: 1517: 1475: 1465: 1447: 1408: 1394: 1317: 1311: 1212: 1186: 1180: 1172: 1166: 1148: 1106: 1051: 898: 863: 717: 636: 628: 113: 8237:
Burki, Fabien; Shalchian-Tabrizi, Kamran & Pawlowski, Jan (2008).
5669: 4297:          Subphylum 4292:          Subphylum 4287:          Subphylum 4277:          Subphylum 4262:          Subphylum 4252:          Subphylum 4247:          Subphylum 3978:          Subphylum 3960:          Subphylum 3950:          Subphylum 3935:          Subphylum 3907:          Subphylum 3892:          Subphylum 3887:          Subphylum 3877:          Subphylum 3872:          Subphylum 3836:          Subphylum 3821:          Subphylum 3811:          Subphylum 3796:          Subphylum 3006:          Subphylum 3001:          Subphylum 2996:          Subphylum 2981:          Subphylum 2976:          Subphylum 2971:          Subphylum 2961:          Subphylum 2956:          Subphylum 2673:          Subphylum 2658:          Subphylum 2648:          Subphylum 2643:          Subphylum 2633:          Subphylum 2628:          Subphylum 2623:          Subphylum 2613:          Subphylum 2608:          Subphylum 2588:          Subphylum 2583:          Subphylum 2568:          Subphylum 2563:          Subphylum 874:'s system of five kingdoms, the plants included just the land plants ( 9266: 9137: 9132: 9123: 8172: 8157: 6834: 6595: 6039: 5976: 5956: 5897: 5721: 5577: 5463: 5430: 5281: 5235: 5231: 4923: 4918: 4913: 4884: 4864: 4488: 4484: 4464: 4460: 4442: 4298: 4278: 4273: 4243: 4238: 4211: 4203: 4127: 4021: 3971: 3961: 3925: 3918: 3913: 3898: 3888: 3842: 3769: 3719: 3714: 3670: 3650: 3605: 3600: 3595: 3570: 3535: 3493: 3460: 3330:          Subclass 3325:          Subclass 3297:          Subclass 3292:          Subclass 3282:          Subclass 3277:          Subclass 3267:          Subclass 3262:          Subclass 3257:          Subclass 3242:          Subclass 3237:          Subclass 3083:          Subclass 3078:          Subclass 3014: 2941: 2886: 2866: 2846: 2836: 2790: 2392: 2388: 2368: 2364: 2346: 2263: 2212: 2208: 2078: 1895: 1836: 1695: 1599: 1487: 1459: 1451: 1402: 1376: 1257: 1142: 934: 855: 851: 753: 654:
proposed a four-kingdom classification by creating the novel Kingdom
640: 332: 259: 121: 7955: 6886: 5075:
classification, where there is emphasis in arranging organisms into
1878:
superkingdom, grouping together the five other eukaryotic kingdoms (
9313: 9292: 8443: 8179: 7197: 7052:"Genome Networks Root the Tree of Life between Prokaryotic Domains" 6756: 6576: 6377: 6159: 6043: 5990: 5867: 5862: 5852: 5847: 5609: 5532: 5517: 5471: 5441: 5337: 5320: 5189: 5159: 4906: 4899: 4894: 4889: 4869: 4835: 4830: 4820: 4536: 4512: 4468: 4456: 4388: 4328: 4263: 4248: 3956: 3951: 3941: 3936: 3903: 3802: 3786: 3774: 3748:          Phylum 3729: 3728:          Phylum 3680:          Phylum 3649:          Phylum 3615: 3614:          Phylum 3604:          Phylum 3584:          Phylum 3560: 3555: 3554:          Phylum 3534:          Phylum 3519:          Phylum 3473: 3467: 3455: 3450: 3419: 3397: 3025: 3020: 2967: 2841: 2826: 2769: 2761: 2533: 2440: 2416: 2372: 2360: 2303: 2256: 2248: 2236: 2224: 2181: 2062: 2008: 1993: 1883: 1879: 1871: 1709: 1595: 1555: 1511: 1493: 1420: 1388: 1359: 1337: 1288: 1280: 1124: 1086: 997: 948: 800: 767: 659: 560: 527: 496: 452: 129: 98: 8992: 3375:          Class 3355:          Class 3345:          Class 3103:          Class 1807:
supported the consensus at that time, that the difference between
9350: 8740: 8382:
Rogozin, I.B.; Basu, M.K.; Csürös, M. & Koonin, E.V. (2009).
8069:
International Journal of Systematic and Evolutionary Microbiology
5952: 5902: 5889: 5884: 5879: 5874: 5725: 5655: 5561: 5392: 5375: 5210: 5163: 4879: 4874: 4857: 4847: 4795: 4504: 4404: 4344: 4332: 4177: 4013: 4002: 3996: 3984: 3724: 3640: 3520: 3515: 3480: 3405: 2408: 1891: 1848: 1840: 1471: 1382: 1348: 1158: 965: 784: 663: 544: 323: 203: 179: 94: 90: 51: 8768: 9105: 9089: 8236: 7146:. Réflexions sur la biologie et la phylogénie des protozoaires" 6654: 6652: 6073:. Fungal Conservation Committee, IUCN SSC. 2021. Archived from 5923: 5909: 5857: 5841: 5804: 5713: 5658:) are descended from the same ancestor as both the unicellular 5499: 5495: 5034: 5029: 5024: 5019: 5014: 5009: 5004: 4999: 4992: 4987: 4982: 4977: 4972: 4810: 4805: 4576: 4560: 4283: 2947: 2876: 2871: 2807: 2480: 2464: 2126: 2110: 1758: 1742: 1040: 912: 847: 731: 655: 458: 363: 311: 183: 106: 86: 78: 67: 7150:
Annales des Sciences Naturelles - Zoologie et Biologie Animale
6038:
Compared to the version Cavalier-Smith published in 2004, the
2207:
Cavalier-Smith does not accept the requirement for taxa to be
9307: 5916: 5819: 5800: 5773: 5682: 5076: 5051: 4965: 4960: 4955: 4950: 4945: 4940: 4935: 4528: 4137: 3047: 2543:        Infrakingdom 2432: 2244: 2094: 1887: 1726: 1246: 981: 838: 319: 237:) can be inserted between subkingdom and infrakingdom, e.g., 199: 175: 146: 140: 134: 82: 8104: 8007: 6649: 6382:"The Prokaryote-Eukaryote Dichotomy: Meanings and Mythology" 6253: 6251: 6249: 6247: 3743:        Superphylum 3675:        Superphylum 3609:        Superphylum 3599:        Superphylum 3549:        Superphylum 3514:        Superphylum 3108:        Infraphylum 3098:        Infraphylum 8764: 8762: 8659: 8598: 6046:
have been moved from Kingdom Protozoa to Kingdom Chromista.
5777: 5301: 5174: 5136: 4122: 3350:        Superclass 3340:        Superclass 1226: 306:, in 1735. He distinguished two kingdoms of living things: 8594:"The eocyte hypothesis and the origin of eukaryotic cells" 8287: 7768: 6749: 4181:        Subphylum 4156:        Subphylum 4136:        Subphylum 4131:        Subphylum 4126:        Subphylum 4098:        Subphylum 4083:        Subphylum 4057:        Subphylum 4052:        Subphylum 3459:        Subphylum 3454:        Subphylum 3433:        Subphylum 3428:        Subphylum 3423:        Subphylum 3034:        Subphylum 3029:        Subphylum 2933:        Subphylum 2928:        Subphylum 2918:        Subphylum 2913:        Subphylum 2875:        Subphylum 2870:        Subphylum 2855:        Subphylum 2850:        Subphylum 2845:        Subphylum 2835:        Subphylum 2830:        Subphylum 2825:        Subphylum 2748:        Subphylum 2733:        Subphylum 2718:        Subphylum 2703:        Subphylum 8338: 8062:
Harper, J. T.; Waanders, E. & Keeling, P. J. (2005).
7250:(January 1969). "New concepts of kingdoms of organisms". 7050:
Dagan, T.; Roettger, M.; Bryant & Martin, W. (2010).
7049: 6244: 5697: 3191:        Subclass 8884: 8759: 8497: 6623:
Palmer, E. Laurence; Fowler, Seymour H. (January 1975).
5944:); but this is beneath the top level classifications of 3196:        Subclas 639:) and organisms whose cells do have a distinct nucleus ( 8652: 8381: 7771:"A higher level classification of all living organisms" 7668:
Proceedings of the Royal Society B: Biological Sciences
6752:"A higher level classification of all living organisms" 5896: 4282:        Phylum 4257:        Phylum 4242:        Phylum 3955:        Phylum 3930:        Phylum 3882:        Phylum 3867:        Phylum 3816:        Phylum 3791:        Phylum 3662:        Phylum 2991:        Phylum 2966:        Phylum 2951:        Phylum 2683:        Phylum 2653:        Phylum 2638:        Phylum 2618:        Phylum 2603:        Phylum 2578:        Phylum 2558:        Phylum 2553:        Phylum 8841: 7830:"Evolution: Revisiting the Root of the Eukaryote Tree" 6745: 6743: 6741: 3320:        Class 3302:        Class 3287:        Class 3272:        Class 3252:        Class 3232:        Class 3227:        Class 3222:        Class 3088:        Class 3073:        Class 8175:"Phylogenomics reshuffles the eukaryotic supergroups" 8061: 6123:
International Union for Conservation of Nature (IUCN)
5704:
Tree of life (biology) § Developments since 1990
8835: 6929: 6927: 5986: 3974: 3704: 3437: 2231:. This means that all living eukaryotes are in fact 1586:
structure; this would later lead to the proposal of
1368: 470:, often called the "father of microscopy", sent the 8878: 7879: 7877: 7875: 7408: 7292: 6738: 6658: 6191: 6624: 6575: 6259: 5143:. All smaller branches can be considered kingdoms. 153: 66:. Kingdoms are divided into smaller groups called 8432: 7884:Simpson, Alastair G.B.; Roger, Andrew J. (2004). 7649: 6924: 6863: 6338: 6336: 6334: 6332: 6330: 6328: 9395: 8844:"Reasons to include viruses in the tree of life" 8747:. International Committee on Taxonomy of Viruses 8500:"Acytota - associated kingdom of neglected life" 7872: 7045: 7043: 6192:Woese, C.R.; Kandler, O.; Wheelis, M.L. (1990). 4237:      Superphylum 2986:      Superphylum 2946:      Superphylum 2678:      Superphylum 2598:      Superphylum 2573:      Superphylum 120:, noting that some traditional kingdoms are not 6535: 6430: 5100: 22: 7716: 7655: 7605: 7556: 7508: 7474:(1981). "Eukaryote kingdoms: Seven or nine?". 7470: 6985: 6933: 6810: 6325: 6257: 5938:International Committee on Taxonomy of Viruses 5915: 5732:(formerly known as eocytes or Crenarchaeota). 5066: 3335:      Subphylum 3315:      Subphylum 3247:      Subphylum 3217:      Subphylum 3093:      Subphylum 3068:      Subphylum 2799:      Subphylum 2794:      Subphylum 2784:      Subphylum 2779:      Subphylum 262:(384–322 BC) classified animal species in his 124:, meaning that they do not consist of all the 9008: 8816:"New views on the megaclassification of life" 8716:"New views on the megaclassification of life" 8426: 7979:"New views on the megaclassification of life" 7937:"Proposal for the recognition of super ranks" 7240: 7040: 6864:Poole, Anthony; Penny, David (21 June 2007). 6569: 6567: 6476: 6474: 6472: 5908: 5856: 878:), and Protoctista has a broader definition. 8547:Biological systematics: The state of the art 8232: 8230: 8168: 8166: 7883: 7827: 7710: 7212: 7174: 7094: 6996:Proceedings of the Royal Society of London B 6622: 6135: 5835: 298:(1707–1778) laid the foundations for modern 112:Some recent classifications based on modern 8807: 8332: 8151: 7512:(1992). "Origins of secondary metabolism". 7168:Titres et Travaux Scientifiques (1906–1937) 7130: 6342: 4310:      Phylum 4151:      Phylum 4121:      Phylum 4078:      Phylum 4047:      Phylum 4034:      Phylum 3472:      Phylum 3449:      Phylum 3418:      Phylum 3024:      Phylum 3019:      Phylum 2923:      Phylum 2908:      Phylum 2865:      Phylum 2860:      Phylum 2840:      Phylum 2820:      Phylum 2728:      Phylum 2698:      Phylum 494:also proposed a third kingdom of life, the 9015: 9001: 8577:: CS1 maint: location missing publisher ( 8003: 8001: 7970: 7828:Roger, A.J. & Simpson, A.G.B. (2009). 7340: 6564: 6469: 6387:Microbiology and Molecular Biology Reviews 6372: 6370: 6368: 6068:"IUCN SSC acceptance of Fauna Flora Funga" 5826: 5781: 5772: 5654:In this system the multicellular animals ( 4335:. (This was based on the consensus in the 3186:      Class 3181:      Class 2523:Cavalier-Smith's six kingdom system (1998) 1587: 8861: 8690: 8680: 8629: 8619: 8591: 8585: 8474: 8464: 8437:(2008). Redfield, Rosemary Jeanne (ed.). 8409: 8358: 8315: 8281: 8264: 8227: 8210: 8200: 8163: 8134: 8124: 8081: 8030: 7928: 7903: 7847: 7804: 7794: 7745: 7687: 7582: 7447: 7437: 7402: 7385: 7375: 7286: 7246: 7077: 7016: 6962: 6787: 6777: 6504: 6480: 6407: 6227: 6217: 5901: 5813: 5811: 5809: 5796: 5794: 5263: 4337:Taxonomic Outline of Bacteria and Archaea 846:recognized an additional kingdom for the 439: 245:in the classification of Cavalier-Smith. 144:(for animals), and, in the 21st century, 8733: 7559:"Kingdom protozoa and its 18 phyla" 7346: 7218: 7180: 7100: 5922: 5668: 5096: 5082: 4780: 854:; his Plantae were mostly multicellular 451: 379:Regnum Vegetabile ('vegetables'/plants) 105:(Animalia, Plantae, Fungi, Protista and 18: 8970:A Brief History of the Kingdoms of Life 8592:Archibald, John M. (23 December 2008). 8544: 8098: 7998: 7228:. Palo Alto: Pacific Books. p. 6. 7162: 7156: 7136: 7118: 7112: 6577:"Teaching Taxonomy: How Many Kingdoms?" 6424: 6365: 6187: 6185: 6183: 6152:McNeill, J.; et al., eds. (2006). 6151: 5979:, that possess typical cellular genes. 5888: 1874:. This superkingdom was opposed to the 581:  Non‑life   398:  Non‑life   253: 9396: 8944:Empire Biota: a comprehensive taxonomy 8813: 8713: 7976: 7611:"A revised six-kingdom system of life" 6308: 6261:"A revised six-kingdom system of life" 5951:There is ongoing debate as to whether 5698:Comparison of top level classification 2247:. Others were reclassified in kingdom 2174: 1031:Whittaker's five kingdom system (1969) 8996: 7934: 7225:The Classification of Lower Organisms 7184:(1938). "The kingdoms of organisms". 6536:Margulis L, Chapman MJ (2009-03-19). 5155:accurately introduced term dominion. 2723:    Infrakingdom 2693:    Infrakingdom 2593:    Infrakingdom 2196:. Unibacteria was divided into phyla 701:of Archaea, Bacteria, and Eukaryota. 9032: 8646: 7124:Generelle Morphologie der Organismen 6573: 6376: 6180: 5046:Kingdoms such as Protozoa represent 3922:       3859:       3783:       3667:       3654:       3506:       2520: 1028: 162:introduced the rank-based system of 9022: 7886:"The real 'kingdoms' of eukaryotes" 7412:; Kandler, O.; Wheelis, M. (1990). 4785: 4775: 2538:    Subkingdom 1851:. Moreover, only chromists contain 1591: 330:, placing them in a third kingdom, 116:have explicitly abandoned the term 13: 8936: 8018:Journal of Eukaryotic Microbiology 7627:10.1111/j.1469-185X.1998.tb00030.x 6280:10.1111/j.1469-185X.1998.tb00030.x 4762: 4757: 2255:which is now part of infrakingdom 666:. Ernst Haeckel, in his 1904 book 102: 14: 9420: 8963: 8498:Trifonov EN, Kejnovsky E (2016). 5866: 5861: 5851: 5846: 5741:Two superdomains (controversial) 5724:emerged from a phylum within the 4767: 4321: 2243:, were reclassified into kingdom 1799: 74: 9052: 9041: 8032:10.1111/j.1550-7408.2005.00053.x 7516:. Novartis Foundation Symposia. 5989: 5883: 832: 622: 9246: 9185: 8707: 8375: 8346:Molecular Biology and Evolution 8055: 7821: 7762: 7599: 7550: 7502: 7464: 6979: 6857: 6804: 6705: 6616: 6529: 6032: 5878: 5873: 4216:    Phylum 4001:    Phylum 3485:    Phylum 2789:    Phylum 2774:    Phylum 1573: 154:Definition and associated terms 9375: 9364: 9332: 9321: 9114: 9096: 8959:, New York: Worth Publishers. 7575:10.1128/mmbr.57.4.953-994.1993 7298:Journal of Molecular Evolution 6662:Journal of Molecular Evolution 6400:10.1128/MMBR.69.2.292-305.2005 6302: 6144: 6129: 6111: 6060: 5817: 2262:Because Cavalier-Smith allows 862:, and his Fungi multicellular 210:was introduced above kingdom. 1: 8516:10.1080/07391102.2015.1086959 8433:Kim, E.; Graham, L. E. & 7350:; Fox, G.E. (November 1977). 6053: 5662:and the fungi which form the 4358:  Life   2273:  Life   2180:recent research. The kingdom 1909:  Life   1609:  Life   889:  Life   858:, his Animalia multicellular 708:  Life   515:  Life   351:  Life   286:) wrote a parallel work, the 280: 273: 8545:Minelli, Alessandro (1993). 8466:10.1371/journal.pone.0002621 8389:Genome Biology and Evolution 8295:Genome Biology and Evolution 8202:10.1371/journal.pone.0000790 8126:10.1371/journal.pgen.0020220 7796:10.1371/journal.pone.0119248 7488:10.1016/0303-2647(81)90050-2 7272:10.1126/science.163.3863.150 7057:Genome Biology and Evolution 6779:10.1371/journal.pone.0119248 6627:Fieldbook of Natural History 6515:10.1126/science.163.3863.150 1293:Branch Opisthomastigomycota 7: 8942:Pelentier, B. (2007-2015). 8849:Nature Reviews Microbiology 8772:Nature Reviews Microbiology 7661:"Only six kingdoms of life" 7557:Cavalier-Smith, T. (1993). 7186:Quarterly Review of Biology 7170:. E. Sottano, Sète, France. 6988:"Only six kingdoms of life" 6986:Cavalier-Smith, T. (2004). 6313:. Oxford: Clarendon Press. 6309:Singer, Charles J. (1931). 6258:Cavalier-Smith, T. (1998). 5982: 5147:Based on such RNA studies, 5067:Beyond traditional kingdoms 5056:phylogenetic classification 1276:Subkingdom Dimastigomycota 592:Regnum Lapideum (minerals) 409:Regnum Lapideum (minerals) 10: 9425: 8988:Whittaker's classification 8955:and Helena Curtis (1970), 7520:: 64–80, discussion 80–7. 6574:Case, Emily (2008-10-01). 6346:International Microbiology 6100:animals, fungi, and plants 5931: 5701: 5678:six proposed supergroups. 5269: 5086: 4617: 2167:‡ No longer recognized by 445: 322:). Linnaeus also included 318:('vegetable kingdom', for 248: 9030: 8435:Redfield, Rosemary Jeanne 7905:10.1016/j.cub.2004.08.038 7849:10.1016/j.cub.2008.12.032 7526:10.1002/9780470514344.ch5 7514:Ciba Foundation Symposium 6866:"Engulfed by speculation" 5599: 5551: 5522: 5493: 5468: 5446: 5435: 5423: 5387: 5380: 5368: 5332: 5325: 5313: 5306: 5228: 5207: 5186: 5179: 5054:, and so are rejected by 4939: 4934: 4929: 4922: 4917: 4912: 4905: 4898: 4893: 4888: 4883: 4878: 4873: 4868: 4863: 4829: 4809: 4804: 4799: 4794: 4791: 4573: 4557: 4525: 4481: 4453: 4446: 4435: 4401: 4385: 4378: 4367: 4360: 4347:is any of the eukaryotic 2477: 2461: 2429: 2385: 2357: 2350: 2339: 2300: 2293: 2282: 2275: 2123: 2107: 2091: 2075: 2059: 2052: 2040: 2005: 1998: 1986: 1952: 1936: 1929: 1918: 1911: 1898:). This was known as the 1755: 1739: 1723: 1706: 1699: 1688: 1653: 1636: 1629: 1618: 1611: 994: 978: 962: 945: 938: 909: 902: 891: 797: 781: 764: 757: 728: 721: 710: 590: 583: 557: 541: 524: 517: 509: 407: 400: 377: 360: 353: 345: 213:Prefixes can be added so 36:biological classification 8976:The five kingdom concept 6583:American Biology Teacher 6434:Archiv fĂĽr Mikrobiologie 6025: 5716:" or cellular life. The 5708:Some authors have added 4302:     4229:     4113:     4070:     4039:     4026:     3851:     3498:     3464:     3441:     3410:     3307:     3209:     3173:     3060:     3011:     2938:     2900:     2812:     2310:as part of a subkingdom 2184:(sole kingdom of empire 837:The differences between 8910:10.1126/science.1101485 8814:Luketa, Stefan (2012). 8714:Luketa, Stefan (2012). 8682:10.1073/pnas.81.12.3786 8621:10.1073/pnas.0811118106 7563:Microbiological Reviews 7439:10.1073/pnas.87.12.4576 7377:10.1073/pnas.74.11.5088 6219:10.1073/pnas.87.12.4576 6104:fauna, flora, and funga 5963:and are not capable of 5576:(or cryptomonads), and 2688:  Subkingdom 1588:three "domains" of life 1253:Subkingdom Gymnomycota 468:Antonie van Leeuwenhoek 302:, now regulated by the 300:biological nomenclature 206:. In 1990, the rank of 8863:10.1038/nrmicro2108-c1 8257:10.1098/rsbl.2008.0224 7738:10.1098/rsbl.2009.0948 7680:10.1098/rspb.2004.2705 7234:10.5962/bhl.title.4474 7009:10.1098/rspb.2004.2705 6955:10.1098/rsbl.2009.0948 6935:Cavalier-Smith, Thomas 5738:Taxonomical root node 5674: 5264:Eukaryotic supergroups 5144: 1833:Gram-positive bacteria 1825:Gram-negative bacteria 1819:into two subkingdoms: 463: 448:Tree of life (biology) 440:Three kingdoms of life 58:is the second highest 47: 8360:10.1093/molbev/msm089 8083:10.1099/ijs.0.63216-0 6136:Linnaeus, C. (1735). 5892:including eukaryotes 5837:Prokaryota / Procarya 5702:Further information: 5672: 5139:data showing Woese's 5130: 5083:Three domains of life 4349:unicellular organisms 1845:endoplasmic reticulum 1805:Thomas Cavalier-Smith 1344:Subkingdom Agnotozoa 1061:Branch Mastigomonera 455: 446:Further information: 328:classification system 97:/Archaebacteria, and 33: 7935:Moore, R.T. (1974). 7144:Pansporella perplexa 6721:Rhode Island College 5970:nucleotide sequences 5531:, and various other 5272:Supergroup (biology) 1530:Phylum Brachiata or 1119:Plasmodiophoromycota 1113:Hyphochytridiomycota 1056:Phylum Myxobacteriae 254:Two kingdoms of life 9409:Biology terminology 9081:Infrakingdom/Branch 8972:at Earthling Nature 8902:2004Sci...306.1344R 8896:(5700): 1344–1350. 8785:10.1038/nrmicro2108 8745:talk.ictvonline.org 8673:1984PNAS...81.3786L 8612:2008PNAS..10520049A 8606:(51): 20049–20050. 8504:J Biomol Struct Dyn 8457:2008PLoSO...3.2621K 8193:2007PLoSO...2..790B 7977:Luketa, S. (2012). 7787:2015PLoSO..1019248R 7674:(1545): 1251–1262. 7430:1990PNAS...87.4576W 7368:1977PNAS...74.5088W 7310:1977JMolE...9..305B 7264:1969Sci...163..150W 7003:(1545): 1251–1262. 6827:1987Natur.326..332C 6770:2015PLoSO..1019248R 6675:1977JMolE...9..305B 6497:1969Sci...163..150W 6210:1990PNAS...87.4576W 5898:Eukaryota / Eukarya 5141:three-domain system 5089:Three-domain system 3299:Loculoascomycetidae 2525: 2175:Six kingdoms (1998) 1900:Archezoa hypothesis 1355:Subkingdom Parazoa 1033: 699:three-domain system 668:The Wonders of Life 652:Herbert F. Copeland 627:The development of 70:(singular phylum). 9404:Kingdoms (biology) 9353: 9310: 9289: 9263: 9221: 9155: 9120: 9108: 9072: 9063: 8981:2021-11-07 at the 8402:10.1093/gbe/evp011 8308:10.1093/gbe/evp022 7718:Cavalier-Smith, T. 7657:Cavalier-Smith, T. 7615:Biological Reviews 7607:Cavalier-Smith, T. 7510:Cavalier-Smith, T. 7472:Cavalier-Smith, T. 7318:10.1007/BF01796092 7070:10.1093/gbe/evq025 6713:"The Six Kingdoms" 6683:10.1007/BF01796092 6447:10.1007/BF00425185 6267:Biological Reviews 6092:animals and plants 5972:from their hosts. 5720:proposes that the 5675: 5145: 4441:Superkingdom  4373:Superkingdom  3377:Homobasidiomycetes 3342:Hemibasidiomycetia 2521: 2239:, like the phylum 2046:Superkingdom  1992:Superkingdom  1924:Superkingdom  1847:instead of in the 1270:Labyrinthulomycota 1064:Phylum Eubacteriae 1047:Branch Myxomonera 1029: 464: 304:Nomenclature Codes 289:Historia Plantarum 268:, while his pupil 265:History of Animals 184:phylum or division 103:five kingdoms only 48: 9391: 9390: 9386: 9385: 9349: 9306: 9285: 9259: 9217: 9151: 9110: 9104: 9068: 9061: 8957:Biology of Plants 8667:(12): 3786–3790. 8158:Burki et al. 2007 8076:(Pt 1): 487–496. 7898:(17): R693–R696. 7424:(12): 4576–4579. 7362:(11): 5088–5090. 7258:(3863): 150–160. 7126:. Reimer, Berlin. 6821:(6111): 332–333. 6642:978-0-070-48425-2 6162:on 6 October 2012 6098:be replaced with 5929: 5928: 5791:non-cellular life 5767:Eocyte hypothesis 5718:eocyte hypothesis 5710:non-cellular life 5660:choanoflagellates 5651: 5650: 5642: 5641: 5633: 5632: 5624: 5623: 5504:choanoflagellates 5412: 5411: 5403: 5402: 5357: 5356: 5348: 5347: 5256: 5255: 5247: 5246: 5133:phylogenetic tree 5040: 5039: 4614: 4613: 4605: 4604: 4596: 4595: 4587: 4586: 4424: 4423: 4415: 4414: 4341:Catalogue of Life 4319: 4318: 4059:Macrorhodophytina 3362:Tremellomycetidae 3327:Sporidiomycetidae 3244:Saccharomycetidae 2518: 2517: 2509: 2508: 2500: 2499: 2491: 2490: 2328: 2327: 2319: 2318: 2164: 2163: 2155: 2154: 2146: 2145: 2137: 2136: 2029: 2028: 2020: 2019: 1975: 1974: 1966: 1965: 1796: 1795: 1787: 1786: 1778: 1777: 1769: 1768: 1677: 1676: 1668: 1667: 1571: 1570: 1026: 1025: 1017: 1016: 1008: 1007: 923: 922: 829: 828: 820: 819: 811: 810: 742: 741: 695:two-empire system 619: 618: 610: 609: 601: 600: 571: 570: 436: 435: 427: 426: 418: 417: 388: 387: 316:Regnum Vegetabile 34:The hierarchy of 9416: 9377: 9366: 9334: 9323: 9248: 9187: 9116: 9098: 9054: 9043: 9033: 9017: 9010: 9003: 8994: 8993: 8930: 8929: 8882: 8876: 8875: 8865: 8839: 8833: 8832: 8820: 8811: 8805: 8804: 8766: 8757: 8756: 8754: 8752: 8737: 8731: 8730: 8720: 8711: 8705: 8704: 8694: 8684: 8650: 8644: 8643: 8633: 8623: 8589: 8583: 8582: 8576: 8568: 8542: 8536: 8535: 8495: 8489: 8488: 8478: 8468: 8430: 8424: 8423: 8413: 8379: 8373: 8372: 8362: 8353:(8): 1702–1713. 8336: 8330: 8329: 8319: 8285: 8279: 8278: 8268: 8234: 8225: 8224: 8214: 8204: 8170: 8161: 8155: 8149: 8148: 8138: 8128: 8102: 8096: 8095: 8085: 8059: 8053: 8052: 8034: 8014: 8005: 7996: 7995: 7983: 7974: 7968: 7967: 7941: 7932: 7926: 7925: 7907: 7881: 7870: 7869: 7851: 7825: 7819: 7818: 7808: 7798: 7766: 7760: 7759: 7749: 7714: 7708: 7707: 7705: 7704: 7691: 7665: 7653: 7647: 7646: 7603: 7597: 7596: 7586: 7554: 7548: 7547: 7506: 7500: 7499: 7482:(3–4): 461–481. 7468: 7462: 7461: 7451: 7441: 7406: 7400: 7399: 7389: 7379: 7344: 7338: 7337: 7290: 7284: 7283: 7244: 7238: 7237: 7216: 7210: 7209: 7178: 7172: 7171: 7160: 7154: 7153: 7152:. 10-VIII: 5–84. 7134: 7128: 7127: 7116: 7110: 7109: 7098: 7092: 7091: 7081: 7047: 7038: 7037: 7035: 7033: 7020: 6992: 6983: 6977: 6976: 6966: 6931: 6922: 6921: 6919: 6917: 6911: 6905:. Archived from 6870: 6861: 6855: 6854: 6835:10.1038/326332a0 6808: 6802: 6801: 6791: 6781: 6747: 6736: 6735: 6733: 6732: 6723:. Archived from 6709: 6703: 6702: 6656: 6647: 6646: 6631:(2nd ed.). 6630: 6620: 6614: 6613: 6611: 6610: 6596:10.2307/30163328 6579: 6571: 6562: 6561: 6533: 6527: 6526: 6508: 6491:(3863): 150–60. 6478: 6467: 6466: 6428: 6422: 6421: 6411: 6374: 6363: 6362: 6340: 6323: 6322: 6306: 6300: 6299: 6263: 6255: 6242: 6241: 6231: 6221: 6189: 6178: 6177: 6170: 6168: 6167: 6148: 6142: 6141: 6133: 6127: 6126: 6125:. 3 August 2021. 6115: 6109: 6108: 6086: 6085: 6079: 6072: 6064: 6047: 6036: 5999: 5994: 5993: 5735: 5734: 5438: 5437: 5426: 5425: 5383: 5382: 5371: 5370: 5328: 5327: 5316: 5315: 5309: 5308: 5297: 5296: 5288: 5182: 5181: 5170: 5169: 5099: 5093:Domain (biology) 4733:3 superkingdoms 4622: 4621: 4449: 4448: 4438: 4437: 4381: 4380: 4370: 4369: 4363: 4362: 4355: 4354: 4314: 4195: 4005: 3976: 3706: 3439: 3389: 3279:Verrucomycetidae 3219:Hemiascomycotina 3075:Chytridiomycetes 3039: 2753: 2670:Chromatibacteria 2610:Chlorobibacteria 2526: 2524: 2353: 2352: 2342: 2341: 2296: 2295: 2285: 2284: 2278: 2277: 2270: 2269: 2055: 2054: 2043: 2042: 2001: 2000: 1989: 1988: 1932: 1931: 1921: 1920: 1914: 1913: 1906: 1905: 1702: 1701: 1691: 1690: 1632: 1631: 1621: 1620: 1614: 1613: 1606: 1605: 1370: 1201:Chlorophycophyta 1034: 1032: 941: 940: 905: 904: 894: 893: 886: 885: 844:Robert Whittaker 760: 759: 724: 723: 713: 712: 705: 704: 586: 585: 520: 519: 512: 511: 504: 503: 403: 402: 356: 355: 348: 347: 340: 339: 285: 282: 278: 275: 9424: 9423: 9419: 9418: 9417: 9415: 9414: 9413: 9394: 9393: 9392: 9387: 9345:Species complex 9095:Superdivision ( 9026: 9024:Taxonomic ranks 9021: 8983:Wayback Machine 8966: 8939: 8937:Further reading 8934: 8933: 8883: 8879: 8840: 8836: 8818: 8812: 8808: 8767: 8760: 8750: 8748: 8739: 8738: 8734: 8718: 8712: 8708: 8651: 8647: 8590: 8586: 8570: 8569: 8557: 8543: 8539: 8496: 8492: 8431: 8427: 8380: 8376: 8337: 8333: 8286: 8282: 8244:Biology Letters 8235: 8228: 8171: 8164: 8156: 8152: 8103: 8099: 8060: 8056: 8012: 8006: 7999: 7981: 7975: 7971: 7956:10.2307/1218807 7939: 7933: 7929: 7891:Current Biology 7882: 7873: 7835:Current Biology 7826: 7822: 7781:(4): e0119248. 7767: 7763: 7715: 7711: 7702: 7700: 7663: 7654: 7650: 7604: 7600: 7555: 7551: 7536: 7507: 7503: 7469: 7465: 7407: 7403: 7345: 7341: 7291: 7287: 7248:Whittaker, R.H. 7245: 7241: 7217: 7213: 7179: 7175: 7161: 7157: 7135: 7131: 7117: 7113: 7099: 7095: 7048: 7041: 7031: 7029: 6990: 6984: 6980: 6943:Biology Letters 6932: 6925: 6915: 6913: 6909: 6887:10.1038/447913a 6868: 6862: 6858: 6809: 6805: 6764:(4): e0119248. 6748: 6739: 6730: 6728: 6711: 6710: 6706: 6657: 6650: 6643: 6621: 6617: 6608: 6606: 6572: 6565: 6554: 6534: 6530: 6506:10.1.1.403.5430 6479: 6470: 6429: 6425: 6375: 6366: 6341: 6326: 6307: 6303: 6256: 6245: 6190: 6181: 6172: 6165: 6163: 6149: 6145: 6134: 6130: 6117: 6116: 6112: 6096:fauna and flora 6083: 6081: 6077: 6070: 6066: 6065: 6061: 6056: 6051: 6050: 6037: 6033: 6028: 5995: 5988: 5985: 5934: 5839: 5830: 5789: 5754:Five Dominiums 5706: 5700: 5652: 5643: 5634: 5625: 5593: (or  5413: 5404: 5358: 5349: 5286: 5274: 5266: 5257: 5248: 5129: 5097: 5095: 5087:Main articles: 5085: 5069: 4742:2 superkingdoms 4696: 4687: 4682: 4675: 4667: 4656: 4649: 4642: 4635: 4628: 4620: 4615: 4606: 4597: 4588: 4425: 4416: 4324: 4309: 4301: 4296: 4291: 4286: 4281: 4276: 4271: 4266: 4261: 4256: 4251: 4246: 4241: 4236: 4228: 4220: 4215: 4207: 4200: 4190: 4185: 4183:Spermatophytina 4180: 4175: 4170: 4165: 4160: 4155: 4150: 4145: 4140: 4135: 4130: 4125: 4120: 4112: 4107: 4102: 4097: 4092: 4087: 4082: 4077: 4069: 4061: 4056: 4054:Rhodellophytina 4051: 4046: 4038: 4033: 4025: 4017: 4010: 4000: 3992: 3987: 3982: 3977: 3969: 3967:Cephalochordata 3964: 3959: 3954: 3949: 3944: 3939: 3934: 3929: 3921: 3916: 3911: 3906: 3901: 3896: 3891: 3886: 3881: 3876: 3871: 3866: 3858: 3850: 3848:Cercomeromorpha 3845: 3840: 3835: 3830: 3825: 3820: 3818:Platyhelminthes 3815: 3810: 3805: 3800: 3795: 3790: 3782: 3777: 3772: 3767: 3762: 3757: 3752: 3750:Nemathelminthes 3747: 3745:Nemathelminthes 3742: 3737: 3732: 3727: 3722: 3717: 3712: 3707: 3702:Trilobitomorpha 3699: 3694: 3689: 3687:Cheliceromorpha 3684: 3679: 3674: 3666: 3661: 3653: 3648: 3643: 3638: 3633: 3628: 3623: 3618: 3613: 3608: 3603: 3598: 3593: 3588: 3583: 3578: 3573: 3568: 3563: 3558: 3553: 3548: 3543: 3538: 3533: 3528: 3523: 3518: 3513: 3505: 3497: 3489: 3484: 3476: 3471: 3463: 3458: 3453: 3448: 3440: 3432: 3427: 3422: 3417: 3409: 3401: 3394: 3384: 3379: 3374: 3369: 3364: 3359: 3354: 3349: 3344: 3339: 3334: 3329: 3324: 3319: 3314: 3306: 3301: 3296: 3294:Dendromycetidae 3291: 3286: 3281: 3276: 3271: 3266: 3261: 3259:Calycomycetidae 3256: 3251: 3246: 3241: 3236: 3231: 3226: 3224:Taphrinomycetes 3221: 3216: 3208: 3200: 3195: 3190: 3185: 3180: 3172: 3167: 3165:Trichomycetalia 3162: 3157: 3152: 3147: 3142: 3140:Mucoromycetidae 3137: 3132: 3127: 3122: 3117: 3112: 3107: 3102: 3097: 3092: 3087: 3082: 3077: 3072: 3067: 3059: 3051: 3044: 3038: 3033: 3028: 3023: 3018: 3010: 3005: 3000: 2998:Tubulicorticata 2995: 2990: 2985: 2980: 2975: 2970: 2965: 2960: 2955: 2950: 2945: 2937: 2932: 2927: 2922: 2917: 2912: 2907: 2899: 2894: 2889: 2884: 2879: 2874: 2869: 2864: 2859: 2854: 2849: 2844: 2839: 2834: 2829: 2824: 2819: 2811: 2803: 2798: 2793: 2788: 2783: 2778: 2773: 2765: 2758: 2752: 2747: 2742: 2737: 2732: 2727: 2722: 2717: 2712: 2707: 2702: 2697: 2692: 2687: 2685:Planctobacteria 2682: 2680:Planctobacteria 2677: 2672: 2667: 2662: 2657: 2652: 2647: 2642: 2637: 2632: 2627: 2622: 2617: 2612: 2607: 2605:Sphingobacteria 2602: 2597: 2592: 2587: 2582: 2577: 2572: 2567: 2562: 2557: 2552: 2547: 2542: 2537: 2530: 2522: 2519: 2510: 2501: 2492: 2329: 2320: 2177: 2165: 2156: 2147: 2138: 2030: 2021: 1976: 1967: 1864:proteobacterium 1802: 1797: 1788: 1779: 1770: 1712:or Protoctista 1678: 1669: 1576: 1430:Pseudocoelomata 1423:or Rhynchocoela 1415:Platyhelminthes 1298:Chytridiomycota 1181:Phaeophycophyta 1167:Rhodophycophyta 1030: 1027: 1018: 1009: 951:or Protoctista 924: 835: 830: 821: 812: 770:or Protoctista 743: 645:Édouard Chatton 625: 620: 611: 602: 572: 530:or Protoctista 450: 442: 437: 428: 419: 389: 333:Regnum Lapideum 283: 276: 256: 251: 225:(also known as 156: 40:taxonomic ranks 38:'s eight major 32: 17: 12: 11: 5: 9422: 9412: 9411: 9406: 9389: 9388: 9384: 9383: 9382: 9381: 9370: 9359: 9354: 9347: 9340: 9339: 9338: 9327: 9316: 9311: 9302: 9301: 9300: 9295: 9290: 9283: 9276: 9275: 9274: 9269: 9264: 9257: 9252: 9239: 9238: 9237: 9232: 9227: 9222: 9215: 9210: 9203: 9202: 9201: 9196: 9191: 9178: 9177: 9176: 9171: 9166: 9161: 9156: 9149: 9142: 9141: 9140: 9135: 9130: 9121: 9102: 9085: 9084: 9083: 9078: 9073: 9066: 9058: 9047: 9031: 9028: 9027: 9020: 9019: 9012: 9005: 8997: 8991: 8990: 8985: 8973: 8965: 8964:External links 8962: 8961: 8960: 8953:Peter H. Raven 8950: 8938: 8935: 8932: 8931: 8877: 8834: 8806: 8779:(4): 306–311. 8758: 8732: 8706: 8645: 8584: 8555: 8537: 8490: 8425: 8374: 8331: 8280: 8251:(4): 366–369. 8226: 8162: 8150: 8097: 8054: 8025:(5): 399–451. 7997: 7969: 7950:(4): 650–652. 7927: 7871: 7820: 7761: 7732:(3): 342–345. 7709: 7648: 7621:(3): 203–266. 7598: 7569:(4): 953–994. 7549: 7534: 7501: 7463: 7401: 7339: 7304:(4): 305–311. 7285: 7239: 7220:Copeland, H.F. 7211: 7198:10.1086/394568 7192:(4): 383–420. 7182:Copeland, H.F. 7173: 7155: 7129: 7111: 7093: 7039: 6978: 6949:(3): 342–345. 6923: 6912:on 6 July 2011 6856: 6803: 6737: 6704: 6669:(4): 305–311. 6648: 6641: 6615: 6590:(8): 472–477. 6563: 6552: 6544:Academic Press 6528: 6468: 6423: 6394:(2): 292–305. 6364: 6324: 6301: 6243: 6204:(12): 4576–9. 6179: 6143: 6128: 6110: 6058: 6057: 6055: 6052: 6049: 6048: 6030: 6029: 6027: 6024: 6023: 6022: 6017: 6012: 6007: 6001: 6000: 5997:Biology portal 5984: 5981: 5948:and subrealm. 5933: 5930: 5927: 5926: 5920: 5919: 5913: 5912: 5906: 5905: 5900: 5894: 5893: 5887: 5885:Archaebacteria 5882: 5877: 5871: 5870: 5865: 5860: 5855: 5850: 5845: 5834: 5824: 5823: 5815: 5814: 5812: 5810: 5808: 5797: 5795: 5793: 5780: 5770: 5769: 5764: 5761: 5755: 5752: 5747: 5742: 5739: 5730:Thermoproteota 5699: 5696: 5687:SAR supergroup 5649: 5648: 5645: 5644: 5640: 5639: 5636: 5635: 5631: 5630: 5627: 5626: 5622: 5621: 5618: 5617: 5598: 5591:Archaeplastida 5586: 5585: 5582: 5581: 5550: 5547:Chromalveolata 5542: 5541: 5538: 5537: 5521: 5513: 5512: 5509: 5508: 5492: 5484: 5483: 5480: 5479: 5467: 5459: 5458: 5455: 5454: 5445: 5436: 5434: 5424: 5422: 5419: 5418: 5415: 5414: 5410: 5409: 5406: 5405: 5401: 5400: 5397: 5396: 5386: 5381: 5379: 5369: 5367: 5364: 5363: 5360: 5359: 5355: 5354: 5351: 5350: 5346: 5345: 5342: 5341: 5331: 5326: 5324: 5314: 5312: 5307: 5305: 5295: 5270:Main article: 5265: 5262: 5254: 5253: 5250: 5249: 5245: 5244: 5241: 5240: 5227: 5224: 5223: 5220: 5219: 5215:Archaebacteria 5206: 5203: 5202: 5199: 5198: 5185: 5180: 5178: 5168: 5164:Archaebacteria 5084: 5081: 5077:natural groups 5068: 5065: 5064: 5063: 5059: 5038: 5037: 5032: 5027: 5022: 5017: 5012: 5007: 5002: 4996: 4995: 4990: 4985: 4980: 4975: 4969: 4968: 4963: 4958: 4953: 4948: 4943: 4938: 4933: 4927: 4926: 4921: 4916: 4910: 4909: 4903: 4902: 4897: 4892: 4887: 4882: 4877: 4872: 4867: 4861: 4860: 4855: 4853:Archaebacteria 4850: 4845: 4843:Archaebacteria 4839: 4838: 4833: 4828: 4823: 4818: 4813: 4808: 4803: 4798: 4793: 4789: 4788: 4783: 4778: 4773: 4770: 4765: 4760: 4755: 4752: 4749: 4745: 4744: 4739: 4734: 4731: 4726: 4721: 4716: 4711: 4706: 4703: 4699: 4698: 4689: 4686:Cavalier-Smith 4684: 4680:Cavalier-Smith 4677: 4669: 4658: 4651: 4644: 4637: 4630: 4619: 4616: 4612: 4611: 4608: 4607: 4603: 4602: 4599: 4598: 4594: 4593: 4590: 4589: 4585: 4584: 4581: 4580: 4572: 4569: 4568: 4565: 4564: 4556: 4553: 4552: 4549: 4548: 4524: 4521: 4520: 4517: 4516: 4480: 4477: 4476: 4473: 4472: 4452: 4447: 4445: 4436: 4434: 4431: 4430: 4427: 4426: 4422: 4421: 4418: 4417: 4413: 4412: 4409: 4408: 4400: 4397: 4396: 4393: 4392: 4384: 4379: 4377: 4368: 4366: 4361: 4359: 4353: 4323: 4322:Seven kingdoms 4320: 4317: 4316: 4197: 4158:Pteridophytina 4100:Phragmophytina 4007: 3952:Appendicularia 3808:Acanthocephala 3755:Scalidorhyncha 3391: 3387:Pileomycetidae 3382:Clavomycetidae 3367:Dacrymycetidae 3332:Uredomycetidae 3284:Ostiomycetidae 3249:Euascomycotina 3130:Neozygomycetia 3095:Melanomycotina 3085:Spizomycetidae 3080:Rumpomycetidae 3070:Dictyomycotina 3041: 2978:Coccidiomorpha 2963:Dinoflagellata 2852:Reticulofilosa 2816:Sarcomastigota 2755: 2740:Halomebacteria 2725:Archaebacteria 2715:Actinobacteria 2705:Teichobacteria 2655:Proteobacteria 2625:Sclenobacteria 2600:Pimelobacteria 2585:Euspirochaetae 2575:Endoflagellata 2565:Chlorobacteria 2516: 2515: 2512: 2511: 2507: 2506: 2503: 2502: 2498: 2497: 2494: 2493: 2489: 2488: 2485: 2484: 2476: 2473: 2472: 2469: 2468: 2460: 2457: 2456: 2453: 2452: 2428: 2425: 2424: 2421: 2420: 2384: 2381: 2380: 2377: 2376: 2356: 2351: 2349: 2340: 2338: 2335: 2334: 2331: 2330: 2326: 2325: 2322: 2321: 2317: 2316: 2313: 2312: 2308:Archaebacteria 2299: 2294: 2292: 2283: 2281: 2276: 2274: 2268: 2229:Hydrogenosomes 2198:Archaebacteria 2176: 2173: 2162: 2161: 2158: 2157: 2153: 2152: 2149: 2148: 2144: 2143: 2140: 2139: 2135: 2134: 2131: 2130: 2122: 2119: 2118: 2115: 2114: 2106: 2103: 2102: 2099: 2098: 2090: 2087: 2086: 2083: 2082: 2074: 2071: 2070: 2067: 2066: 2058: 2053: 2051: 2041: 2039: 2036: 2035: 2032: 2031: 2027: 2026: 2023: 2022: 2018: 2017: 2014: 2013: 2004: 1999: 1997: 1987: 1985: 1982: 1981: 1978: 1977: 1973: 1972: 1969: 1968: 1964: 1963: 1960: 1959: 1956:Archaebacteria 1951: 1948: 1947: 1944: 1943: 1935: 1930: 1928: 1919: 1917: 1912: 1910: 1904: 1868:eukaryogenesis 1813:Archaebacteria 1801: 1800:Eight kingdoms 1798: 1794: 1793: 1790: 1789: 1785: 1784: 1781: 1780: 1776: 1775: 1772: 1771: 1767: 1766: 1763: 1762: 1754: 1751: 1750: 1747: 1746: 1738: 1735: 1734: 1731: 1730: 1722: 1719: 1718: 1715: 1714: 1705: 1700: 1698: 1689: 1687: 1684: 1683: 1680: 1679: 1675: 1674: 1671: 1670: 1666: 1665: 1662: 1661: 1657:Archaebacteria 1652: 1649: 1648: 1645: 1644: 1635: 1630: 1628: 1619: 1617: 1612: 1610: 1604: 1592:seven kingdoms 1575: 1572: 1569: 1568: 1567: 1566: 1565: 1564: 1563: 1562: 1561: 1560: 1559: 1558: 1552: 1546: 1540: 1534: 1522: 1521: 1520: 1514: 1508: 1502: 1496: 1490: 1484: 1478: 1456: 1455: 1454: 1444: 1438: 1436:Acanthocephala 1426: 1425: 1424: 1417: 1399: 1398: 1397: 1391: 1373: 1372: 1371: 1362: 1353: 1352: 1351: 1332: 1331: 1330: 1329: 1328: 1327: 1326: 1320: 1314: 1306:Amastigomycota 1302: 1301: 1300: 1285: 1284: 1283: 1274: 1273: 1272: 1266: 1260: 1241: 1240: 1239: 1238: 1237: 1236: 1235: 1229: 1217: 1216: 1215: 1209: 1191: 1190: 1189: 1177: 1176: 1175: 1153: 1152: 1151: 1145: 1139: 1133: 1127: 1121: 1115: 1109: 1103: 1097: 1081: 1080: 1079: 1078: 1077: 1071: 1065: 1059: 1058: 1057: 1054: 1024: 1023: 1020: 1019: 1015: 1014: 1011: 1010: 1006: 1005: 1002: 1001: 993: 990: 989: 986: 985: 977: 974: 973: 970: 969: 961: 958: 957: 954: 953: 944: 939: 937: 930: 929: 926: 925: 921: 920: 917: 916: 908: 903: 901: 892: 890: 884: 834: 831: 827: 826: 823: 822: 818: 817: 814: 813: 809: 808: 805: 804: 796: 793: 792: 789: 788: 780: 777: 776: 773: 772: 763: 758: 756: 749: 748: 745: 744: 740: 739: 736: 735: 727: 722: 720: 711: 709: 703: 683:C. B. van Niel 677:In the 1960s, 624: 621: 617: 616: 613: 612: 608: 607: 604: 603: 599: 598: 595: 594: 589: 584: 582: 578: 577: 574: 573: 569: 568: 565: 564: 556: 553: 552: 549: 548: 540: 537: 536: 533: 532: 523: 518: 516: 510: 508: 502: 441: 438: 434: 433: 430: 429: 425: 424: 421: 420: 416: 415: 412: 411: 406: 401: 399: 395: 394: 391: 390: 386: 385: 382: 381: 376: 373: 372: 369: 368: 359: 354: 352: 346: 344: 338: 314:kingdom') and 308:Regnum Animale 255: 252: 250: 247: 155: 152: 138:(for plants), 60:taxonomic rank 16:Taxonomic rank 15: 9: 6: 4: 3: 2: 9421: 9410: 9407: 9405: 9402: 9401: 9399: 9380: 9379: 9371: 9369: 9368: 9360: 9358: 9355: 9352: 9348: 9346: 9343: 9342: 9341: 9337: 9336: 9328: 9326: 9325: 9317: 9315: 9312: 9309: 9305: 9304: 9303: 9299: 9296: 9294: 9291: 9288: 9284: 9282: 9279: 9278: 9277: 9273: 9270: 9268: 9265: 9262: 9258: 9256: 9253: 9251: 9250: 9242: 9241: 9240: 9236: 9233: 9231: 9228: 9226: 9223: 9220: 9216: 9214: 9211: 9209: 9206: 9205: 9204: 9200: 9197: 9195: 9192: 9190: 9189: 9181: 9180: 9179: 9175: 9172: 9170: 9167: 9165: 9162: 9160: 9157: 9154: 9150: 9148: 9145: 9144: 9143: 9139: 9136: 9134: 9131: 9129: 9125: 9122: 9119: 9118: 9107: 9103: 9101: 9100: 9091: 9088: 9087: 9086: 9082: 9079: 9077: 9074: 9071: 9067: 9065: 9064:/Superkingdom 9059: 9057: 9056: 9048: 9046: 9045: 9037: 9036: 9035: 9034: 9029: 9025: 9018: 9013: 9011: 9006: 9004: 8999: 8998: 8995: 8989: 8986: 8984: 8980: 8977: 8974: 8971: 8968: 8967: 8958: 8954: 8951: 8948: 8945: 8941: 8940: 8927: 8923: 8919: 8915: 8911: 8907: 8903: 8899: 8895: 8891: 8890: 8881: 8873: 8869: 8864: 8859: 8855: 8851: 8850: 8845: 8838: 8831:(4): 218–237. 8830: 8826: 8825: 8817: 8810: 8802: 8798: 8794: 8790: 8786: 8782: 8778: 8774: 8773: 8765: 8763: 8746: 8742: 8736: 8729:(4): 218–237. 8728: 8724: 8717: 8710: 8702: 8698: 8693: 8688: 8683: 8678: 8674: 8670: 8666: 8662: 8661: 8656: 8649: 8641: 8637: 8632: 8627: 8622: 8617: 8613: 8609: 8605: 8601: 8600: 8595: 8588: 8580: 8574: 8566: 8562: 8558: 8556:0-412-36440-9 8552: 8548: 8541: 8533: 8529: 8525: 8521: 8517: 8513: 8510:(8): 1641–8. 8509: 8505: 8501: 8494: 8486: 8482: 8477: 8472: 8467: 8462: 8458: 8454: 8450: 8446: 8445: 8440: 8436: 8429: 8421: 8417: 8412: 8407: 8403: 8399: 8395: 8391: 8390: 8385: 8378: 8370: 8366: 8361: 8356: 8352: 8348: 8347: 8342: 8335: 8327: 8323: 8318: 8313: 8309: 8305: 8301: 8297: 8296: 8291: 8284: 8276: 8272: 8267: 8262: 8258: 8254: 8250: 8246: 8245: 8240: 8233: 8231: 8222: 8218: 8213: 8208: 8203: 8198: 8194: 8190: 8186: 8182: 8181: 8176: 8169: 8167: 8159: 8154: 8146: 8142: 8137: 8132: 8127: 8122: 8118: 8114: 8113: 8112:PLOS Genetics 8108: 8101: 8093: 8089: 8084: 8079: 8075: 8071: 8070: 8065: 8058: 8050: 8046: 8042: 8038: 8033: 8028: 8024: 8020: 8019: 8011: 8004: 8002: 7994:(4): 218–237. 7993: 7989: 7988: 7980: 7973: 7965: 7961: 7957: 7953: 7949: 7945: 7938: 7931: 7923: 7919: 7915: 7911: 7906: 7901: 7897: 7893: 7892: 7887: 7880: 7878: 7876: 7867: 7863: 7859: 7855: 7850: 7845: 7842:(4): R165–7. 7841: 7837: 7836: 7831: 7824: 7816: 7812: 7807: 7802: 7797: 7792: 7788: 7784: 7780: 7776: 7772: 7765: 7757: 7753: 7748: 7743: 7739: 7735: 7731: 7727: 7723: 7720:(June 2010). 7719: 7713: 7699: 7695: 7690: 7685: 7681: 7677: 7673: 7669: 7662: 7658: 7652: 7644: 7640: 7636: 7632: 7628: 7624: 7620: 7616: 7612: 7608: 7602: 7594: 7590: 7585: 7580: 7576: 7572: 7568: 7564: 7560: 7553: 7545: 7541: 7537: 7535:9780470514344 7531: 7527: 7523: 7519: 7515: 7511: 7505: 7497: 7493: 7489: 7485: 7481: 7477: 7473: 7467: 7459: 7455: 7450: 7445: 7440: 7435: 7431: 7427: 7423: 7419: 7415: 7411: 7405: 7397: 7393: 7388: 7383: 7378: 7373: 7369: 7365: 7361: 7357: 7353: 7349: 7343: 7335: 7331: 7327: 7323: 7319: 7315: 7311: 7307: 7303: 7299: 7295: 7289: 7281: 7277: 7273: 7269: 7265: 7261: 7257: 7253: 7249: 7243: 7235: 7231: 7227: 7226: 7221: 7215: 7207: 7203: 7199: 7195: 7191: 7187: 7183: 7177: 7169: 7165: 7159: 7151: 7147: 7145: 7139: 7133: 7125: 7121: 7115: 7107: 7103: 7097: 7089: 7085: 7080: 7075: 7071: 7067: 7063: 7059: 7058: 7053: 7046: 7044: 7028: 7024: 7019: 7014: 7010: 7006: 7002: 6998: 6997: 6989: 6982: 6974: 6970: 6965: 6960: 6956: 6952: 6948: 6944: 6940: 6936: 6930: 6928: 6908: 6904: 6900: 6896: 6892: 6888: 6884: 6881:(7147): 913. 6880: 6876: 6875: 6867: 6860: 6852: 6848: 6844: 6840: 6836: 6832: 6828: 6824: 6820: 6816: 6815: 6807: 6799: 6795: 6790: 6785: 6780: 6775: 6771: 6767: 6763: 6759: 6758: 6753: 6746: 6744: 6742: 6727:on 2021-05-10 6726: 6722: 6718: 6714: 6708: 6700: 6696: 6692: 6688: 6684: 6680: 6676: 6672: 6668: 6664: 6663: 6655: 6653: 6644: 6638: 6634: 6629: 6628: 6619: 6605: 6601: 6597: 6593: 6589: 6585: 6584: 6578: 6570: 6568: 6559: 6555: 6553:9780080920146 6549: 6545: 6541: 6540: 6532: 6524: 6520: 6516: 6512: 6507: 6502: 6498: 6494: 6490: 6486: 6485: 6477: 6475: 6473: 6464: 6460: 6456: 6452: 6448: 6444: 6440: 6436: 6435: 6427: 6419: 6415: 6410: 6405: 6401: 6397: 6393: 6389: 6388: 6383: 6379: 6373: 6371: 6369: 6360: 6356: 6353:(4): 207–16. 6352: 6348: 6347: 6339: 6337: 6335: 6333: 6331: 6329: 6320: 6316: 6312: 6305: 6297: 6293: 6289: 6285: 6281: 6277: 6274:(3): 203–66. 6273: 6269: 6268: 6262: 6254: 6252: 6250: 6248: 6239: 6235: 6230: 6225: 6220: 6215: 6211: 6207: 6203: 6199: 6195: 6188: 6186: 6184: 6175: 6174:"article 3.1" 6161: 6157: 6156: 6147: 6139: 6132: 6124: 6120: 6114: 6107: 6105: 6101: 6097: 6093: 6080:on 2021-11-11 6076: 6069: 6063: 6059: 6045: 6041: 6035: 6031: 6021: 6018: 6016: 6013: 6011: 6010:Phylogenetics 6008: 6006: 6003: 6002: 5998: 5992: 5987: 5980: 5978: 5973: 5971: 5966: 5962: 5958: 5954: 5949: 5947: 5943: 5939: 5925: 5921: 5918: 5914: 5911: 5907: 5904: 5899: 5895: 5891: 5886: 5881: 5876: 5872: 5869: 5864: 5859: 5854: 5849: 5843: 5838: 5833: 5832:cellular life 5829: 5825: 5821: 5818:Prionobiota ( 5816: 5806: 5802: 5799:Virusobiota ( 5798: 5792: 5788: 5784: 5779: 5775: 5771: 5768: 5765: 5763:Six kingdoms 5762: 5760: 5756: 5753: 5751: 5750:Three domains 5748: 5746: 5743: 5740: 5737: 5736: 5733: 5731: 5727: 5723: 5719: 5715: 5711: 5705: 5695: 5692: 5688: 5684: 5679: 5671: 5667: 5665: 5661: 5657: 5647: 5646: 5638: 5637: 5629: 5628: 5620: 5619: 5616: 5615: 5611: 5607: 5603: 5596: 5592: 5588: 5587: 5584: 5583: 5580: 5579: 5575: 5571: 5567: 5563: 5559: 5555: 5554:Stramenopiles 5549:   5548: 5544: 5543: 5540: 5539: 5536: 5534: 5530: 5526: 5520:   5519: 5515: 5514: 5511: 5510: 5507: 5505: 5501: 5497: 5491:   5490: 5486: 5485: 5482: 5481: 5478: 5477: 5473: 5466:   5465: 5461: 5460: 5457: 5456: 5453: 5451: 5444:   5443: 5440: 5439: 5433:   5432: 5428: 5427: 5421: 5420: 5417: 5416: 5408: 5407: 5399: 5398: 5395: 5394: 5391: 5385: 5384: 5378:   5377: 5373: 5372: 5366: 5365: 5362: 5361: 5353: 5352: 5344: 5343: 5340: 5339: 5336: 5330: 5329: 5323:   5322: 5318: 5317: 5311: 5310: 5304:   5303: 5299: 5298: 5294: 5290: 5283: 5279: 5273: 5268: 5261: 5252: 5251: 5243: 5242: 5239: 5237: 5233: 5226: 5225: 5222: 5221: 5218: 5216: 5212: 5205: 5204: 5201: 5200: 5197: 5195: 5191: 5184: 5183: 5177:   5176: 5172: 5171: 5167: 5165: 5161: 5156: 5152: 5150: 5142: 5138: 5134: 5094: 5090: 5080: 5078: 5074: 5060: 5057: 5053: 5049: 5045: 5044: 5043: 5036: 5033: 5031: 5028: 5026: 5023: 5021: 5018: 5016: 5013: 5011: 5008: 5006: 5003: 5001: 4998: 4997: 4994: 4991: 4989: 4986: 4984: 4981: 4979: 4976: 4974: 4971: 4970: 4967: 4964: 4962: 4959: 4957: 4954: 4952: 4949: 4947: 4944: 4942: 4937: 4932: 4928: 4925: 4920: 4915: 4911: 4908: 4904: 4901: 4896: 4891: 4886: 4881: 4876: 4871: 4866: 4862: 4859: 4856: 4854: 4851: 4849: 4846: 4844: 4841: 4840: 4837: 4834: 4832: 4827: 4824: 4822: 4819: 4817: 4814: 4812: 4807: 4802: 4797: 4790: 4787: 4784: 4782: 4779: 4777: 4774: 4771: 4769: 4766: 4764: 4761: 4759: 4756: 4753: 4750: 4747: 4746: 4743: 4740: 4738: 4735: 4732: 4730: 4727: 4725: 4722: 4720: 4717: 4715: 4712: 4710: 4707: 4704: 4701: 4700: 4694: 4690: 4685: 4681: 4678: 4674: 4670: 4665: 4662: 4659: 4655: 4652: 4648: 4645: 4641: 4638: 4634: 4631: 4627: 4624: 4623: 4610: 4609: 4601: 4600: 4592: 4591: 4583: 4582: 4579: 4578: 4571: 4570: 4567: 4566: 4563: 4562: 4555: 4554: 4551: 4550: 4547: 4546: 4542: 4538: 4534: 4530: 4523: 4522: 4519: 4518: 4515: 4514: 4510: 4506: 4502: 4498: 4494: 4490: 4486: 4479: 4478: 4475: 4474: 4471: 4470: 4466: 4462: 4458: 4451: 4450: 4444: 4440: 4439: 4433: 4432: 4429: 4428: 4420: 4419: 4411: 4410: 4407: 4406: 4399: 4398: 4395: 4394: 4391: 4390: 4383: 4382: 4376: 4372: 4371: 4365: 4364: 4357: 4356: 4352: 4350: 4346: 4342: 4338: 4334: 4330: 4315: 4313: 4308: 4307: 4304:Infrakingdom 4300: 4295: 4290: 4285: 4280: 4275: 4270: 4265: 4260: 4255: 4254:Labyrinthista 4250: 4245: 4240: 4235: 4234: 4231:Infrakingdom 4227: 4226: 4219: 4214: 4213: 4206: 4205: 4198: 4196: 4194: 4189: 4184: 4179: 4174: 4169: 4164: 4159: 4154: 4149: 4144: 4139: 4134: 4129: 4124: 4119: 4118: 4115:Infrakingdom 4111: 4106: 4101: 4096: 4091: 4090:Prasinophytae 4086: 4085:Chlorophytina 4081: 4076: 4075: 4072:Infrakingdom 4068: 4067: 4066:Viridiplantae 4060: 4055: 4050: 4045: 4044: 4041:Infrakingdom 4037: 4032: 4031: 4028:Infrakingdom 4024: 4023: 4016: 4015: 4008: 4006: 4004: 3999: 3998: 3991: 3990:Gnathostomata 3986: 3981: 3973: 3968: 3963: 3958: 3953: 3948: 3943: 3938: 3933: 3928: 3927: 3924:Infrakingdom 3920: 3915: 3910: 3905: 3900: 3895: 3890: 3885: 3884:Echinodermata 3880: 3879:Enteropneusta 3875: 3874:Pterobranchia 3870: 3865: 3864: 3861:Infrakingdom 3857: 3856: 3855:Deuterostomia 3849: 3844: 3839: 3834: 3833:Rhabditophora 3829: 3824: 3819: 3814: 3809: 3804: 3799: 3794: 3793:Acanthognatha 3789: 3788: 3785:Infrakingdom 3781: 3776: 3771: 3766: 3761: 3756: 3751: 3746: 3741: 3736: 3731: 3726: 3721: 3716: 3711: 3703: 3698: 3693: 3688: 3683: 3678: 3673: 3672: 3669:Infrakingdom 3665: 3660: 3659: 3656:Infrakingdom 3652: 3647: 3642: 3637: 3632: 3627: 3622: 3617: 3612: 3607: 3602: 3597: 3592: 3587: 3582: 3577: 3572: 3567: 3562: 3557: 3552: 3547: 3542: 3537: 3532: 3527: 3522: 3517: 3512: 3511: 3508:Infrakingdom 3504: 3503: 3496: 3495: 3488: 3483: 3482: 3475: 3470: 3469: 3466:Infrakingdom 3462: 3457: 3452: 3447: 3446: 3443:Infrakingdom 3436: 3435:Archaeocyatha 3431: 3430:Calcispongiae 3426: 3425:Hyalospongiae 3421: 3416: 3415: 3412:Infrakingdom 3408: 3407: 3400: 3399: 3392: 3390: 3388: 3383: 3378: 3373: 3372:Auromycetidae 3368: 3363: 3358: 3353: 3352:Hymenomycetia 3348: 3343: 3338: 3337:Orthomycotina 3333: 3328: 3323: 3318: 3317:Septomycotina 3313: 3312: 3311:Basidiomycota 3305: 3304:Plectomycetes 3300: 3295: 3290: 3289:Loculomycetes 3285: 3280: 3275: 3274:Pyrenomycetes 3270: 3269:Pezomycetidae 3265: 3264:Lecomycetidae 3260: 3255: 3250: 3245: 3240: 3239:Dipomycetidae 3235: 3230: 3225: 3220: 3215: 3214: 3207: 3206: 3199: 3194: 3193:Pleistophorea 3189: 3184: 3179: 3178: 3177:Microsporidia 3171: 3170:Pyxomycetalia 3166: 3161: 3160:Pedomycetidae 3156: 3155:Entomycetidae 3151: 3146: 3145:Meromycetidae 3141: 3136: 3131: 3126: 3121: 3116: 3115:Eozygomycetia 3111: 3106: 3101: 3096: 3091: 3090:Enteromycetes 3086: 3081: 3076: 3071: 3066: 3065: 3058: 3057: 3050: 3049: 3042: 3040: 3037: 3032: 3027: 3022: 3017: 3016: 3013:Infrakingdom 3009: 3008:Filocorticata 3004: 2999: 2994: 2989: 2988:Heterokaryota 2984: 2979: 2974: 2969: 2964: 2959: 2958:Protalveolata 2954: 2949: 2944: 2943: 2940:Infrakingdom 2936: 2931: 2926: 2921: 2920:Pseudociliata 2916: 2911: 2906: 2905: 2904:Discicristata 2902:Infrakingdom 2898: 2893: 2888: 2883: 2878: 2873: 2868: 2863: 2858: 2853: 2848: 2843: 2838: 2833: 2828: 2823: 2818: 2817: 2814:Infrakingdom 2810: 2809: 2802: 2797: 2792: 2787: 2782: 2777: 2772: 2771: 2764: 2763: 2756: 2754: 2751: 2750:Sulfobacteria 2746: 2741: 2736: 2731: 2726: 2721: 2716: 2711: 2706: 2701: 2696: 2691: 2686: 2681: 2676: 2671: 2666: 2665:Alphabacteria 2661: 2660:Rhodobacteria 2656: 2651: 2650:Phycobacteria 2646: 2645:Gloeobacteria 2641: 2640:Cyanobacteria 2636: 2635:Fibrobacteria 2631: 2626: 2621: 2616: 2615:Flavobacteria 2611: 2606: 2601: 2596: 2595:Glycobacteria 2591: 2586: 2581: 2576: 2571: 2570:Deinobacteria 2566: 2561: 2556: 2555:Heliobacteria 2551: 2546: 2541: 2536: 2535: 2528: 2527: 2514: 2513: 2505: 2504: 2496: 2495: 2487: 2486: 2483: 2482: 2475: 2474: 2471: 2470: 2467: 2466: 2459: 2458: 2455: 2454: 2451: 2450: 2446: 2442: 2438: 2434: 2427: 2426: 2423: 2422: 2419: 2418: 2414: 2410: 2406: 2402: 2398: 2394: 2390: 2383: 2382: 2379: 2378: 2375: 2374: 2370: 2366: 2362: 2355: 2354: 2348: 2344: 2343: 2337: 2336: 2333: 2332: 2324: 2323: 2315: 2314: 2311: 2309: 2305: 2298: 2297: 2291: 2287: 2286: 2280: 2279: 2272: 2271: 2267: 2265: 2260: 2258: 2254: 2250: 2246: 2242: 2241:Microsporidia 2238: 2234: 2230: 2226: 2220: 2218: 2214: 2210: 2205: 2203: 2199: 2195: 2191: 2187: 2183: 2172: 2170: 2160: 2159: 2151: 2150: 2142: 2141: 2133: 2132: 2129: 2128: 2121: 2120: 2117: 2116: 2113: 2112: 2105: 2104: 2101: 2100: 2097: 2096: 2089: 2088: 2085: 2084: 2081: 2080: 2073: 2072: 2069: 2068: 2065: 2064: 2057: 2056: 2049: 2045: 2044: 2038: 2037: 2034: 2033: 2025: 2024: 2016: 2015: 2012: 2010: 2003: 2002: 1995: 1991: 1990: 1984: 1983: 1980: 1979: 1971: 1970: 1962: 1961: 1958: 1957: 1950: 1949: 1946: 1945: 1942: 1941: 1934: 1933: 1927: 1923: 1922: 1916: 1915: 1908: 1907: 1903: 1901: 1897: 1893: 1889: 1885: 1881: 1877: 1873: 1869: 1865: 1861: 1860:endosymbiosis 1856: 1854: 1853:chlorophyll c 1850: 1846: 1842: 1838: 1834: 1830: 1826: 1822: 1818: 1814: 1810: 1806: 1792: 1791: 1783: 1782: 1774: 1773: 1765: 1764: 1761: 1760: 1753: 1752: 1749: 1748: 1745: 1744: 1737: 1736: 1733: 1732: 1729: 1728: 1721: 1720: 1717: 1716: 1713: 1711: 1704: 1703: 1697: 1693: 1692: 1686: 1685: 1682: 1681: 1673: 1672: 1664: 1663: 1660: 1658: 1651: 1650: 1647: 1646: 1643: 1641: 1634: 1633: 1627: 1623: 1622: 1616: 1615: 1608: 1607: 1603: 1601: 1597: 1593: 1589: 1585: 1584:ribosomal RNA 1581: 1557: 1553: 1551: 1547: 1545: 1544:Echinodermata 1541: 1539: 1535: 1533: 1529: 1528: 1527: 1523: 1519: 1515: 1513: 1509: 1507: 1503: 1501: 1500:Sipunculoidea 1497: 1495: 1491: 1489: 1485: 1483: 1479: 1477: 1473: 1469: 1468: 1467: 1463: 1462: 1461: 1457: 1453: 1449: 1445: 1443: 1442:Aschelminthes 1439: 1437: 1433: 1432: 1431: 1427: 1422: 1418: 1416: 1412: 1411: 1410: 1406: 1405: 1404: 1400: 1396: 1392: 1390: 1386: 1385: 1384: 1380: 1379: 1378: 1374: 1367: 1366:Archaeocyatha 1363: 1361: 1357: 1356: 1354: 1350: 1346: 1345: 1343: 1342: 1341: 1340: 1339: 1333: 1325: 1324:Basidiomycota 1321: 1319: 1315: 1313: 1309: 1308: 1307: 1303: 1299: 1295: 1294: 1292: 1291: 1290: 1286: 1282: 1278: 1277: 1275: 1271: 1267: 1265: 1264:Acrasiomycota 1261: 1259: 1255: 1254: 1252: 1251: 1250: 1249: 1248: 1242: 1234: 1230: 1228: 1224: 1223: 1222: 1218: 1214: 1210: 1208: 1204: 1203: 1202: 1198: 1197: 1196: 1195:Euchlorophyta 1192: 1188: 1184: 1183: 1182: 1178: 1174: 1170: 1169: 1168: 1164: 1163: 1162: 1161: 1160: 1154: 1150: 1146: 1144: 1140: 1138: 1134: 1132: 1131:Cnidosporidia 1128: 1126: 1122: 1120: 1116: 1114: 1110: 1108: 1104: 1102: 1098: 1096: 1092: 1091: 1090: 1089: 1088: 1082: 1076: 1072: 1070: 1066: 1063: 1062: 1060: 1055: 1053: 1049: 1048: 1046: 1045: 1044: 1043: 1042: 1036: 1035: 1022: 1021: 1013: 1012: 1004: 1003: 1000: 999: 992: 991: 988: 987: 984: 983: 976: 975: 972: 971: 968: 967: 960: 959: 956: 955: 952: 950: 943: 942: 936: 932: 931: 928: 927: 919: 918: 915: 914: 907: 906: 900: 896: 895: 888: 887: 883: 879: 877: 873: 872:Lynn Margulis 867: 865: 861: 857: 853: 849: 845: 840: 833:Five kingdoms 825: 824: 816: 815: 807: 806: 803: 802: 795: 794: 791: 790: 787: 786: 779: 778: 775: 774: 771: 769: 762: 761: 755: 751: 750: 747: 746: 738: 737: 734: 733: 726: 725: 719: 715: 714: 707: 706: 702: 700: 696: 692: 688: 684: 680: 679:Roger Stanier 675: 673: 672:Cyanobacteria 669: 665: 661: 657: 653: 648: 646: 642: 638: 634: 630: 623:Four kingdoms 615: 614: 606: 605: 597: 596: 593: 588: 587: 580: 579: 576: 575: 567: 566: 563: 562: 555: 554: 551: 550: 547: 546: 539: 538: 535: 534: 531: 529: 522: 521: 514: 513: 506: 505: 501: 499: 498: 493: 492:Ernst Haeckel 489: 486:proposed the 485: 480: 476: 473: 472:Royal Society 469: 461: 460: 454: 449: 444: 432: 431: 423: 422: 414: 413: 410: 405: 404: 397: 396: 393: 392: 384: 383: 380: 375: 374: 371: 370: 367: 365: 358: 357: 350: 349: 342: 341: 337: 335: 334: 329: 325: 321: 317: 313: 309: 305: 301: 297: 296:Carl Linnaeus 293: 292:, on plants. 291: 290: 284: 287 BC 271: 267: 266: 261: 246: 244: 243:Deuterostomia 240: 236: 232: 228: 224: 220: 216: 211: 209: 205: 201: 197: 193: 189: 185: 181: 177: 173: 169: 165: 161: 160:Carl Linnaeus 151: 149: 148: 143: 142: 137: 136: 131: 127: 123: 119: 115: 110: 108: 104: 100: 96: 92: 88: 84: 80: 76: 71: 69: 65: 62:, just below 61: 57: 53: 45: 41: 37: 21: 9372: 9361: 9329: 9318: 9243: 9182: 9111: 9093: 9080: 9075: 9069: 9049: 9038: 8956: 8943: 8893: 8887: 8880: 8853: 8847: 8837: 8828: 8824:Protistology 8822: 8809: 8776: 8770: 8749:. Retrieved 8744: 8735: 8726: 8723:Protistology 8722: 8709: 8664: 8658: 8648: 8603: 8597: 8587: 8546: 8540: 8507: 8503: 8493: 8451:(7): e2621. 8448: 8442: 8428: 8393: 8387: 8377: 8350: 8344: 8334: 8299: 8293: 8283: 8248: 8242: 8184: 8178: 8153: 8119:(12): e220. 8116: 8110: 8100: 8073: 8067: 8057: 8022: 8016: 7991: 7987:Protistology 7985: 7972: 7947: 7943: 7930: 7895: 7889: 7839: 7833: 7823: 7778: 7774: 7764: 7729: 7725: 7712: 7701:. Retrieved 7671: 7667: 7651: 7618: 7614: 7601: 7566: 7562: 7552: 7517: 7513: 7504: 7479: 7475: 7466: 7421: 7417: 7404: 7359: 7355: 7342: 7301: 7297: 7288: 7255: 7251: 7242: 7224: 7214: 7189: 7185: 7176: 7167: 7158: 7149: 7143: 7132: 7123: 7114: 7105: 7102:Linnaeus, C. 7096: 7061: 7055: 7030:. Retrieved 7000: 6994: 6981: 6946: 6942: 6914:. Retrieved 6907:the original 6878: 6872: 6859: 6818: 6812: 6806: 6761: 6755: 6729:. Retrieved 6725:the original 6716: 6707: 6666: 6660: 6626: 6618: 6607:. Retrieved 6587: 6581: 6558:Google Books 6556:– via 6538: 6531: 6488: 6482: 6441:(1): 17–35. 6438: 6432: 6426: 6391: 6385: 6350: 6344: 6310: 6304: 6271: 6265: 6201: 6197: 6164:. Retrieved 6160:the original 6154: 6146: 6137: 6131: 6122: 6113: 6103: 6099: 6095: 6091: 6088: 6082:. Retrieved 6075:the original 6062: 6034: 5974: 5950: 5941: 5935: 5827: 5787:Aphanobionta 5786: 5782: 5758: 5707: 5690: 5680: 5676: 5664:Opisthokonta 5653: 5614:glaucophytes 5600: 5595:Primoplantae 5589:   5552: 5545:   5525:Foraminifera 5523: 5516:   5494: 5489:Opisthokonta 5487:   5476:slime moulds 5470:most lobose 5469: 5462:   5447: 5429:Domain  5388: 5374:Domain  5333: 5319:Domain  5300:   5291: 5275: 5267: 5258: 5229: 5208: 5187: 5173:   5157: 5153: 5146: 5070: 5050:rather than 5041: 4692: 4672: 4663: 4574: 4558: 4533:glaucophytes 4526: 4493:cryptophytes 4482: 4454: 4402: 4386: 4325: 4303: 4289:Bigyromonada 4230: 4222: 4221:   4209: 4208:   4201: 4199: 4193:Angiospermae 4188:Gymnospermae 4173:Sphenophytae 4153:Tracheophyta 4133:Anthocerotae 4114: 4071: 4063: 4062:   4040: 4027: 4019: 4018:   4011: 4009: 3994: 3993:   3923: 3909:Eleutherozoa 3869:Hemichordata 3860: 3852: 3784: 3780:Nematomorpha 3668: 3664:Chaetognatha 3658:Chaetognathi 3655: 3526:Gymnolaemata 3507: 3499: 3491: 3490:   3487:Myxosporidia 3478: 3477:   3465: 3445:Coelenterata 3442: 3411: 3403: 3402:   3395: 3393: 3322:Septomycetes 3308: 3254:Discomycetes 3210: 3202: 3201:   3174: 3125:Glomomycetes 3110:Zygomycotina 3100:Allomycotina 3061: 3053: 3052:   3045: 3043: 3012: 2983:Manubrispora 2939: 2901: 2897:Dictyostelia 2862:Foraminifera 2857:Monadofilosa 2813: 2805: 2804:   2796:Anaeromonada 2767: 2766:   2759: 2757: 2735:Euryarcheota 2730:Mendosicutes 2720:Togobacteria 2710:Endobacteria 2700:Posibacteria 2695:Posibacteria 2675:Thiobacteria 2630:Fusobacteria 2620:Eurybacteria 2580:Spirochaetae 2560:Hadobacteria 2545:Lipobacteria 2540:Negibacteria 2531: 2529: 2478: 2462: 2437:glaucophytes 2430: 2397:cryptophytes 2386: 2358: 2345:Empire  2301: 2288:Empire  2261: 2233:metakaryotes 2221: 2209:monophyletic 2206: 2202:Posibacteria 2194:Negibacteria 2178: 2166: 2124: 2108: 2092: 2076: 2060: 2006: 1953: 1937: 1857: 1829:Posibacteria 1821:Negibacteria 1803: 1756: 1740: 1724: 1707: 1694:Empire  1654: 1637: 1624:Empire  1577: 1574:Six kingdoms 1550:Hemichordata 1538:Chaetognatha 1335: 1334: 1244: 1243: 1233:Tracheophyta 1156: 1155: 1137:Zoomastigina 1095:Euglenophyta 1084: 1083: 1075:Spirochaetae 1069:Actinomycota 1038: 1037: 995: 979: 963: 946: 933:Empire  910: 897:Empire  880: 868: 860:heterotrophs 836: 798: 782: 765: 752:Empire  729: 716:Empire  690: 687:superkingdom 686: 676: 667: 649: 626: 591: 558: 542: 525: 495: 487: 481: 477: 465: 462:with plants. 457: 443: 408: 378: 361: 331: 315: 307: 294: 287: 270:Theophrastus 263: 257: 234: 230: 226: 223:infrakingdom 222: 218: 214: 212: 164:nomenclature 157: 145: 139: 133: 132:. The terms 128:of a common 122:monophyletic 117: 111: 75:six kingdoms 72: 55: 49: 9272:Infrafamily 9255:Superfamily 9169:Subterclass 9138:Microphylum 9133:Infraphylum 9128:Subdivision 9090:Superphylum 8741:"ICTV Code" 8302:: 231–238. 8187:(8): e790. 8160:, p. 4 7476:Bio Systems 7348:Woese, C.R. 7294:Woese, C.R. 7164:Chatton, É. 7138:Chatton, É. 7120:Haeckel, E. 6717:www.ric.edu 6633:McGraw-Hill 6015:Systematics 5965:replication 5745:Two empires 5728:called the 5685:dubbed the 5606:green algae 5602:Land plants 5574:Cryptophyta 5558:Brown Algae 5390:prokaryotic 5335:prokaryotic 4931:Vegetabilia 4751:3 kingdoms 4748:2 kingdoms 4545:land plants 4541:green algae 4501:Brown Algae 4497:Heterokonta 4294:Pseudofungi 4269:Hypogyrista 4233:Heterokonta 4225:Chromobiota 4223:Subkingdom 4218:Cryptophyta 4210:Subkingdom 4163:Psilophytae 4105:Charophytae 4095:Tetraphytae 4080:Chlorophyta 4074:Chlorophyta 4064:Subkingdom 4036:Glaucophyta 4030:Glaucophyta 4020:Subkingdom 3995:Subkingdom 3863:Coelomopora 3823:Turbellaria 3765:Kinorhyncha 3735:Onychophora 3710:Mandibulata 3697:Chelicerata 3692:Pycnogonida 3646:Pogonophora 3591:Brachiopoda 3581:Cephalopoda 3566:Glossophora 3546:Cycliophora 3502:Protostomia 3492:Subkingdom 3479:Subkingdom 3404:Subkingdom 3357:Gelimycetes 3347:Ustomycetes 3234:Endomycetes 3203:Subkingdom 3188:Microsporea 3135:Zygomycetes 3120:Bolomycetes 3105:Allomycetes 3064:Archemycota 3054:Subkingdom 3003:Epiplasmata 2882:Archamoebae 2806:Subkingdom 2786:Axostylaria 2781:Eopharyngia 2768:Subkingdom 2690:Unibacteria 2590:Leptospirae 2449:land plants 2445:green algae 2405:Brown Algae 2401:Heterokonta 2306:— includes 2190:Unibacteria 2169:taxonomists 2048:Metakaryota 1876:Metakaryota 1642:(Bacteria) 1532:Pogonophora 1526:Enterocoela 1506:Echiuroidea 1482:Brachiopoda 1466:Schizocoela 1375:Subkingdom 1287:Subkingdom 1207:Chlorophyta 1193:Subkingdom 1179:Subkingdom 1165:Subkingdom 1101:Chrysophyta 876:Embryophyta 864:saprotrophs 643:). In 1937 637:prokaryotes 488:Protoctista 239:Protostomia 227:infraregnum 126:descendants 9398:Categories 9357:Subspecies 9298:Infratribe 9281:Supertribe 9230:Infraorder 9213:Superorder 9184:Division ( 9164:Infraclass 9147:Superclass 9113:Division ( 9076:Subkingdom 9051:Subrealm ( 8856:(8): 615. 8549:. London. 8396:: 99–113. 7726:Biol. Lett 7703:2010-04-29 7064:: 379–92. 6731:2020-07-25 6609:2020-07-28 6166:2011-02-20 6084:2022-03-04 6054:References 6044:rhizarians 6040:alveolates 6005:Cladistics 5961:metabolism 5959:that lack 5863:Eubacteria 5776:/ Vitae / 5722:eukaryotes 5570:Haptophyta 5529:Radiolaria 5450:flagellate 5282:eukaryotes 5194:Eubacteria 5160:Eubacteria 5149:Carl Woese 4826:Eubacteria 4816:Eubacteria 4801:Prokaryota 4786:7 kingdoms 4781:6 kingdoms 4776:8 kingdoms 4768:6 kingdoms 4763:5 kingdoms 4758:4 kingdoms 4509:Haptophyta 4375:Prokaryota 4339:, and the 4312:Haptophyta 4306:Haptophyta 4259:Ochrophyta 4168:Lycophytae 4117:Cormophyta 4110:Rudophytae 4049:Rhodophyta 4043:Rhodophyta 3980:Vertebrata 3894:Pelmatozoa 3838:Neodermata 3828:Mucorhabda 3740:Tardigrada 3682:Arthropoda 3636:Clitellata 3631:Pharyngata 3626:Operculata 3621:Polychaeta 3586:Brachiozoa 3541:Entoprocta 3414:Spongiaria 3229:Geomycetes 3213:Ascomycota 3183:Minisporea 3150:Zoomycetes 3036:Radiolaria 3015:Actinopoda 2993:Ciliophora 2973:Gregarinae 2935:Saccostoma 2930:Plicostoma 2925:Euglenozoa 2915:Tetramitia 2910:Percolozoa 2801:Parabasala 2776:Metamonada 2745:Eurytherma 2550:Eobacteria 2413:Haptophyta 2290:Prokaryota 2253:Metamonada 2225:archezoans 2186:Prokaryota 1940:Eubacteria 1926:Prokaryota 1817:Eubacteria 1809:Eubacteria 1659:(Archaea) 1640:Eubacteria 1626:Prokaryota 1580:Carl Woese 1518:Arthropoda 1476:Ectoprocta 1448:Entoprocta 1409:Acoelomata 1395:Ctenophora 1318:Ascomycota 1312:Zygomycota 1258:Myxomycota 1213:Charophyta 1187:Phaeophyta 1173:Rhodophyta 1149:Ciliophora 1107:Pyrrophyta 1052:Cyanophyta 899:Prokaryota 856:autotrophs 718:Prokaryota 693:—with the 641:eukaryotes 629:microscopy 366:(animals) 277: 371 215:subkingdom 114:cladistics 9363:Variety ( 9320:Section ( 9267:Subfamily 9245:Section ( 9235:Parvorder 9208:Magnorder 9174:Parvclass 9124:Subphylum 8573:cite book 7922:207051421 7410:Woese, C. 6501:CiteSeerX 6150:See e.g. 5977:Mimivirus 5957:parasites 5610:red algae 5578:Alveolata 5535:protozoa 5472:amoeboids 5464:Amoebozoa 5452:protozoa 5431:Eukaryota 5393:Archaeans 5285:'kingdoms 5236:Eukaryota 5135:based on 5073:cladistic 4924:Chromista 4919:Chromista 4914:Chromista 4865:Eukaryota 4737:2 empires 4729:3 domains 4724:2 empires 4719:2 empires 4714:2 empires 4709:2 empires 4691:Ruggiero 4654:Whittaker 4489:Alveolata 4485:Chromista 4465:Choanozoa 4461:Amoebozoa 4443:Eukaryota 4299:Opalinata 4279:Diatomeae 4274:Chrysista 4244:Sagenista 4239:Sagenista 4212:Cryptista 4204:Chromista 4143:Sphagneae 4128:Hepaticae 4123:Bryophyta 4022:Biliphyta 3972:Conodonta 3962:Acraniata 3932:Urochorda 3926:Chordonia 3919:Echinozoa 3914:Asterozoa 3899:Blastozoa 3889:Homalozoa 3843:Trematoda 3813:Monokonta 3770:Nematoida 3760:Priapozoa 3720:Myriapoda 3715:Crustacea 3677:Haemopoda 3671:Ecdysozoa 3651:Nemertina 3606:Sipuncula 3601:Sipuncula 3596:Phoronida 3576:Spiculata 3571:Univalvia 3551:Conchozoa 3536:Kamptozoa 3531:Lophopoda 3494:Bilateria 3461:Medusozoa 3205:Neomycota 3031:Spasmaria 2942:Alveolata 2887:Mycetozoa 2867:Amoebozoa 2847:Phytomyxa 2837:Choanozoa 2822:Neomonada 2791:Trichozoa 2393:Alveolata 2389:Chromista 2369:Choanozoa 2365:Amoebozoa 2347:Eukaryota 2264:paraphyly 2213:paraphyla 2079:Chromista 1896:Chromista 1839:from the 1837:Chromista 1696:Eukaryota 1600:Chromista 1578:In 1977, 1524:Subgrade 1488:Phoronida 1464:Subgrade 1460:Coelomata 1452:Kamptozoa 1403:Bilateria 1377:Eumetazoa 1227:Bryophyta 1221:Metaphyta 1143:Sarcodina 935:Eukaryota 852:nutrition 754:Eukaryota 650:In 1938, 484:John Hogg 466:In 1674, 260:Aristotle 219:subregnum 9331:Series ( 9314:Subgenus 9293:Subtribe 9225:Suborder 9159:Subclass 8979:Archived 8926:84298461 8918:15486256 8872:19561628 8793:19270719 8751:26 April 8640:19091952 8565:27895507 8532:38178747 8524:26305806 8485:18612431 8444:PLOS ONE 8420:20333181 8369:17488740 8326:20333193 8275:18522922 8221:17726520 8180:PLOS ONE 8145:17194223 8092:15653923 8041:16248873 7914:15341755 7866:13172971 7858:19243692 7815:25923521 7775:PLOS ONE 7756:20031978 7698:15306349 7659:(2004). 7609:(1998). 7334:27788891 7222:(1956). 7206:84634277 7166:(1937). 7140:(1925). 7122:(1866). 7104:(1735). 7088:20624742 7032:29 April 7027:15306349 6973:20031978 6937:(2009). 6916:15 March 6895:17581566 6798:25923521 6757:PLOS ONE 6699:27788891 6604:30163328 6463:29859498 6455:13916221 6418:15944457 6380:(2005). 6378:Sapp, J. 6359:10943416 6042:and the 6020:Taxonomy 5983:See also 5924:Animalia 5903:Protista 5868:Bacteria 5853:Bacteria 5848:Bacteria 5759:kingdoms 5597:)  5533:amoeboid 5518:Rhizaria 5448:various 5442:Excavata 5338:Bacteria 5321:Bacteria 5280:for all 5278:grab-bag 5190:Bacteria 5058:systems. 5035:Animalia 5030:Animalia 5025:Animalia 5020:Animalia 5015:Animalia 5010:Animalia 5005:Animalia 5000:Animalia 4907:Protozoa 4900:Protozoa 4895:Protozoa 4890:Archezoa 4880:Protista 4875:Protista 4870:Protista 4836:Bacteria 4831:Bacteria 4821:Bacteria 4796:Protista 4647:Copeland 4626:Linnaeus 4577:Animalia 4575:Kingdom 4559:Kingdom 4527:Kingdom 4513:Rhizaria 4483:Kingdom 4469:Excavata 4457:Protozoa 4455:Kingdom 4403:Kingdom 4389:Bacteria 4387:Kingdom 4329:Bacteria 4264:Phaeista 4249:Bicoecia 4202:Kingdom 4012:Kingdom 3957:Chordata 3942:Ascidiae 3937:Tunicata 3904:Crinozoa 3803:Rotifera 3798:Trochata 3787:Platyzoa 3775:Nematoda 3730:Lobopoda 3616:Annelida 3611:Vermizoa 3561:Bivalvia 3556:Mollusca 3510:Lophozoa 3474:Placozoa 3468:Placozoa 3456:Anthozoa 3451:Cnidaria 3420:Porifera 3398:Animalia 3396:Kingdom 3198:Disporea 3056:Eomycota 3046:Kingdom 3026:Radiozoa 3021:Heliozoa 2968:Sporozoa 2842:Cercozoa 2827:Apusozoa 2770:Archezoa 2762:Protozoa 2760:Kingdom 2534:Bacteria 2532:Kingdom 2481:Animalia 2479:Kingdom 2463:Kingdom 2431:Kingdom 2417:Rhizaria 2387:Kingdom 2373:Excavata 2361:Protozoa 2359:Kingdom 2304:Bacteria 2302:Kingdom 2257:Excavata 2249:Protozoa 2237:Archezoa 2182:Bacteria 2127:Animalia 2125:Kingdom 2109:Kingdom 2093:Kingdom 2077:Kingdom 2063:Protozoa 2061:Kingdom 2009:Archezoa 2007:Kingdom 1994:Archezoa 1954:Kingdom 1938:Kingdom 1884:Protozoa 1880:Animalia 1872:Archezoa 1759:Animalia 1757:Kingdom 1741:Kingdom 1725:Kingdom 1710:Protista 1708:Kingdom 1655:Kingdom 1638:Kingdom 1596:Protozoa 1556:Chordata 1512:Annelida 1494:Mollusca 1421:Nemertea 1389:Cnidaria 1360:Porifera 1338:Animalia 1336:Kingdom 1289:Eumycota 1281:Oomycota 1245:Kingdom 1157:Kingdom 1125:Sporozoa 1087:Protista 1085:Kingdom 1039:Kingdom 998:Animalia 996:Kingdom 980:Kingdom 964:Kingdom 949:Protista 947:Kingdom 911:Kingdom 801:Animalia 799:Kingdom 783:Kingdom 768:Protista 766:Kingdom 730:Kingdom 660:Bacteria 561:Animalia 559:Kingdom 543:Kingdom 528:Protista 526:Kingdom 507:  497:Protista 482:In 1860 343:  324:minerals 233:(Latin: 130:ancestor 99:Bacteria 91:Protista 79:Animalia 9351:Species 9070:Kingdom 9040:Realm ( 8898:Bibcode 8889:Science 8801:3907750 8701:6587394 8669:Bibcode 8631:2629348 8608:Bibcode 8476:2440802 8453:Bibcode 8411:2817406 8317:2817417 8266:2610160 8212:1949142 8189:Bibcode 8136:1713255 8049:8060916 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4333:Archaea 4178:Filices 4148:Bryatae 4014:Plantae 4003:Mesozoa 3997:Mesozoa 3985:Agnatha 3947:Thaliae 3853:Branch 3725:Insecta 3641:Echiura 3521:Bryozoa 3516:Polyzoa 3500:Branch 3481:Myxozoa 3406:Radiata 3309:Phylum 3211:Phylum 3175:Phylum 3062:Phylum 2953:Dinozoa 2832:Isomita 2435:— e.g. 2433:Plantae 2411:etc.), 2409:Diatoms 2391:— e.g. 2363:— e.g. 2251:, like 2095:Plantae 1892:Plantae 1849:cytosol 1841:Plantae 1727:Plantae 1554:Phylum 1548:Phylum 1542:Phylum 1536:Phylum 1516:Phylum 1510:Phylum 1504:Phylum 1498:Phylum 1492:Phylum 1486:Phylum 1480:Phylum 1472:Bryozoa 1470:Phylum 1446:Phylum 1440:Phylum 1434:Phylum 1419:Phylum 1413:Phylum 1401:Branch 1393:Phylum 1387:Phylum 1383:Radiata 1381:Branch 1364:Phylum 1358:Phylum 1349:Mesozoa 1347:Phylum 1322:Phylum 1316:Phylum 1310:Phylum 1304:Branch 1296:Phylum 1279:Phylum 1268:Phylum 1262:Phylum 1256:Phylum 1231:Phylum 1225:Phylum 1219:Branch 1211:Phylum 1205:Phylum 1199:Branch 1185:Phylum 1171:Phylum 1159:Plantae 1147:Phylum 1141:Phylum 1135:Phylum 1129:Phylum 1123:Phylum 1117:Phylum 1111:Phylum 1105:Phylum 1099:Phylum 1093:Phylum 1073:Phylum 1067:Phylum 1050:Phylum 966:Plantae 785:Plantae 664:Archaea 633:nucleus 545:Plantae 364:Animale 362:Regnum 326:in his 249:History 204:species 180:species 118:kingdom 95:Archaea 83:Plantae 56:kingdom 52:biology 9374:Form ( 9261:Family 9199:Cohort 9194:Legion 9106:Phylum 9062:Domain 8924:  8916:  8870:  8799:  8791:  8699:  8692:345305 8689:  8638:  8628:  8563:  8553:  8530:  8522:  8483:  8473:  8418:  8408:  8367:  8324:  8314:  8273:  8263:  8219:  8209:  8143:  8133:  8090:  8047:  8039:  7962:  7920:  7912:  7864:  7856:  7813:  7803:  7754:  7744:  7696:  7686:  7641:  7633:  7591:  7584:372943 7581:  7542:  7532:  7494:  7456:  7446:  7396:270744 7394:  7387:432104 7384:  7332:  7326:408502 7324:  7278:  7204:  7086:  7076:  7025:  7015:  6971:  6961:  6901:  6893:  6874:Nature 6849:  6841:  6814:Nature 6796:  6786:  6697:  6691:408502 6689:  6639:  6602:  6550:  6521:  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Index


biological classification
taxonomic ranks
domain
biology
taxonomic rank
domain
phyla
six kingdoms
Animalia
Plantae
Fungi
Protista
Archaea
Bacteria
five kingdoms only
Monera
cladistics
monophyletic
descendants
ancestor
flora
fauna
funga
Carl Linnaeus
nomenclature
class
order
genus
species

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