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Kambara

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3948:, the upper forelimb may have been able to swing with much greater force, while also providing greater stability against gravity and thus improving the animals ability to perform the crocodilian high walk, the mode of locomotion during which crocodiles lift their body off the ground. Additionally, these factors may also increase stride-length and walking speed. The change to the deltapectoral crest could have advantages for swimming, allowing the limbs to better resist the drag during the recovery stroke. Furthermore, the shortening of the humeral shaft may have also come with advantages, although this is less clear due to the fact that no material of the lower forelimb has been described yet, making it ambiguous how the musculature of the lower and upper arm would connect. Regardless, it is possible that the shortening could have brought the musculature into a more proximal position, which would mean that the lower limbs would have exerted less force while walking or swimming, allowing the animal to swing the lower limb faster. 3964:, based on material collected from Tingamarra, were described by Stein and colleagues in 2017 and feature a robust ilium similar to what is seen in modern alligators and gharials, but with notably shallower peduncles (the parts of the ilium that connect to the other bones of the hip). The joint for the femur is shallowly concave and the postacetabular process, the section of the ilium behind this joint, lacks the constriction seen in modern crocodiles. Otherwise the ilium does not differ significantly for what is seen in other Eusuchians. This morphotype is designated "pelvic form one" by Stein and colleagues and differs in its morphology from fossils found in the Rundle Formation that may pertain to another species of 820: 3904:, shows several circular pits arranged in a linear fashion, which is in line with the bite marks left by a crocodilian. The amount of tooth marks present on the plastron indicates that the individual that tried to eat the turtle bit it several times, behaviour that would match the "juggling" performed by modern crocodiles. "Juggling" in this case describes the act of repeatedly biting prey like turtles in order to change its orientation, helping to align it either with the teeth or to make it easier to swallow. However, it is believed that in this instance the crocodile was overambitious, as the plastron's width and thickness would be too great for the average 4083:. Salisbury and Molnar suggest that, if not ecologically different, the two species may have differed in their habitat preferences and possibly wouldn't have crossed paths under normal circumstances. They point out that the fossil site near Murgon shows signs of having undergone both dry and wet periods, with the former greatly reducing the present bodies of water. A drought could have driven one of the species from their natural habitat and forced them to look for sanctuary in water that would otherwise be home to the other form. Somewhat similar circumstances may have led to the creation of the bonebed that preserves the fossils of 4120: 4064: 4091:
large, adult males prefer deeper open waters. This could explain why the Murgon site primarily preserves large individuals alongside some remains of eggs and hatchlings, with only few animals of intermediate size. Such subadults may have lived largely separate from adults in so called "stockyards", as they would pose a threat to small hatchlings while themselves being threatened by cannibalistic adult males. This could explain why the Murgon site primarily preserves large individuals alongside some remains of eggs and hatchlings, with only few animals of intermediate size.
6746: 6741: 5901: 3881: 157: 3973: 831: 5906: 1001:, which had been studied and compared to the equivalent bones in saltwater and freshwater crocodiles. In both of these modern forms, the deltopectoral crest is offset medially from the lateral margin of the shaft. The apex of the crest is also directed medially, which essentially places it just above the midline of the shaft. The deltapectoral crest sits at a right angle relative to the ventral face of the humeral shaft. However, in 131: 603: 3817: 4056:. Both species are known from a single fossil site near Murgon, with the remains being intermingled with one another, suggesting they occurred in the same environment at the time of their deaths. Morphologically, both species share a variety of similarities and differences. Both are semi-aquatic animals with platyrostral snouts and both are believed to have reached similar lengths. The two however differ in that 983: 938:. Around the seventh and eight dentary teeth, as well as from the twelfth to seventeenth, the teeth appear to have interlocked. However, the tenth to twelfth teeth of the dentary are confluent, meaning they are too closely spaced to interlock with those of the upper jaw, necessitating an overbite in this area. This clearly sets it apart from all other species of 1025:, the dorsal and ventral faces are compressed, which in turn lengthens the medial and lateral condyles, while also directing these surfaces towards the midline. The profile of the distal end is subsequently hexagonal. Profile and size of the medial and lateral condyles in saltwater and freshwater crocodiles are nearly identical, but very different in 4020:, is mentioned, but cannot be fully confirmed due to the poor preservation of that particular element. Simultaneously, Buchanan notes that the skeletal is not any more specialised in aquatic life than that of modern crocodilians either, suggesting a similarly generalized semi-aquatic mode of life. Finally, it is pointed out that the 903:
interlocking teeth in some areas and a slight overlap further back in the jaw. The enlarged fifth tooth of the maxilla does however occupy more space, with its pit being set much further laterally than any of the others. Regardless, this clearly sets not just the skulls but also the dentaries apart from those assigned to
559:, named in 2009 by Lucas A. Buchanan. Although the type description is limited to the skull material (two crania and a lower jaw), which is easily comparable to the previously established species, Buchanan takes note of a "considerable amount" of postcranial remains. These remains were partly described in a 4152:
attacked members of their own species. Such behavior is commonly seen in extant crocodilians and even recorded in a multitude of fossils. Crocodilians often attack limb bones, which would explain the fractures seen on the pathological humerus and metatarsals. The high number of individuals present at
3912:. A costal scute of what is thought to be the same individual has also been recovered, also showing signs of a crocodile bite, some healing and infection. While this suggests that turtles, which were a readily available food source, were fed on at least occasionally, it does not necessarily mean that 957:
All teeth are slightly compressed side to side and possess distinct carinae (cutting edges), but are not ziphodont like in some later mekosuchines. Both this compression and the development of the edges is strongest towards the front of the jaw and gradually grows weaker further back in the toothrow,
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differ significantly, as neither preserve fully overlapping dentition. Instead, distinct occlusal pits reveal that the teeth slid in between each other when the jaw was closed. This means that the teeth of these two species would interlock in a fashion much more similar to what is observed in species
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was a semi-aquatic generalist. One method to determine the details of such a lifestyle may be found in the way the teeth of these crocodilians occlude with another, although the specifics of this are still poorly understood. One hypothesis by Charles C. Mook proposed that an overbite in crocodilians
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bonebed lends itself well to this hypothesis as well, with the crowded nature of the area leading to increased aggression between its residents. In both cases, the injured animal survived and healed. The osteomyelitis of NMV P227802 meanwhile was caused by a bacterial infection causing necrosis and
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It is also possible that the animals were typically kept apart not by preferences unique to the two species but through preferences based on size, age and sex. In modern saltwater crocodiles, the ranges of nesting females and juveniles often overlap upriver or in areas of denser vegetation, whereas
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Another important factor that differentiates the ecology of the various species is the development of the retroarticular process. The retroarticular process serves as an attachment site for important jaw adductor and abductor muscles and is subsequently of great importance in capturing and subduing
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and subsequently formed a callus, a growth of cancellous bone, during healing. Another specimen represents a series of metatarsals, toe bones, which were fractured during the animal's life and then fused into a single element as it healed, with the individual elements rotating due to the continued
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seemingly displays a combination of adaptions for both semi-aquatic and somewhat terrestrial life, with the limbs suggesting a superior ability to walk on land relative to modern forms, whereas the shape and form of the skull are indicative of a semi-aquatic ambush hunting lifestyle not unlike the
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things are notably different. Neither the deltapectoral crest nor its apex are directed medially, with the crest as a whole being just above the lateral margin of the shaft and the apex being directed ventrally. The right angle formed by the crest and shaft however remains. This is associated with
442:
at Murgon. It is unclear how exactly these two species differed in their niche, but they did differ notably in their anatomy, with the latter being slightly more gracile and with interlocking teeth. It is also possible that the two species didn't coexist at all, but instead were driven together by
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is poorly understood. The general shape of the skull, which is flattened, and the environment the fossils were deposited in, all suggest that they were semi-aquatic ambush predators like modern crocodilians. However, subtle differences in the postcranial remains, especially the limb bones, suggest
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is an additional point of consideration. The olecranon fossa and the articular surface of the humerus both suggest a wider range of motion than in saltwater and freshwater crocodiles, but especially the former would require the description of the lower arm to verify this conclusion. The increased
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clearly interlock with each other based on the presence of the aforementioned occlusal pits between the individual tooth sockets up to the ninth tooth of the upper jaw. After this point, the pits are located further medially. This is also confirmed by the pits present in the lower jaw, indicating
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or possibly a different genus altogether. Assuming the former, the Rundle fossils ("pelvic form two") feature much deeper peduncles which are shared with younger mekosuchines from the Oligocene and Miocene, but lack the specific expansion of the pubic peduncle seen in later taxa. Additionally,
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with its intermediate form, various undetermined crocodilians of the time with interlocking teeth and phylogenetic analysis all seem to suggest that the genus actually began with interlocking dentition, acquiring the overbite later. Recent phylogenies consistently point towards
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differs significantly and is used to set them apart. Generally, three different morphologies can be observed, with the difference being primarily expressed through how the upper and lower toothrow interact with one another. As in many other crocodilians, the dental margin of
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The mid 1980s saw the discovery of mammal remains in the area, leading to increased study and revised dating. This new-found attention led to the discovery of several additional crocodilian fossils, including a largely complete skull (QM F21115) collected in 1991. The genus
4075:, Salisbury and Willis argue that the difference in head shape was of little value to their respective ecologies. They suggest that the differences are so minor that they are effectively still the same ecomorph, which they determine may have been similar to that of today's 532:
seemingly showed a high degree of variation among its fossils, in particular regarding the better preserved lower jaws. For this reason, it was suggested that the fossils could belong to multiple species of a single genus. This was confirmed when a second species,
3841:
of mammalian predators, being used to break and slice. This would then suggest that interlocking dentition may be better suited when having to restrain large, struggling prey, requiring greater force that would be more evenly spread out across the toothrow.
434:. While this could suggest that it was faster on land than living crocodilians, it does not necessarily mean that it was terrestrial like some later mekosuchines are thought to have been. Another question regarding its ecology is posed by the coexistence of 3928:
is typically regarded as such a semi-aquatic animal, which would be supported by the environment it was found in. While the skull displays no particular adaptations that would suggest any degree of terrestriality, as is the case in mekosuchines like
4127:
In addition to the many well preserved bones that show no signs of non-taphonomic distortion, there is plentiful material that shows clear signs of injury. This material, covered in Buchanan's thesis, includes a femur that has suffered a
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and other extinct Australian crocodilians were noted since the description of the Murgon mandible in 1982. By the 1993 description, the idea of an endemic Australian crocodilian radiation was better established, at the time including
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was described, it was recognised as a highly variable genus, retaining certain key traits but differing notably in others. As noted by Buchanan, the most prominent of these differentiating traits centre around the anatomy of the
4040:. However it is also possible that this was no adaption towards movement on land, but rather used for bottom-walking in shallow waters, a habit that could have led to more terrestrial mekosuchines later during the Cenozoic. 1006:
the torsion of the shaft. In the modern forms, the humerus is given a sigmoid shape by the fact that the articulating surfaces, the parts of the bone that connect to the joints, are at an angle to each other. The humerus of
2383:. In addition to the monophyletic Mekosuchinae recovered by the vast majority of their analysis, two more novel results were also found. These alternative topologies render Mekosuchinae paraphyletic, including the clade 4239:
may have been found across many of the inland waterways of Queensland, possibly even all of eastern Australia where temperatures allowed for the presence of crocodilians. If this was the case, then the distribution of
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stands out as having a long, high and wide process, offering a greater surface area for the musculature to attach to. By comparison, the retroarticular processes in older species are less well developed, with that of
4432:
Stein, M.; Salisbury, S. W.; Hand, S. J.; Archer, M.; Godthelp, H. (2012). "Humeral morphology of the early Eocene mekosuchine crocodylian Kambara from the Tingamarra Local Fauna southeastern Queensland, Australia".
4202:, although none of them were especially large, seemingly maxing out at 100 mm (3.9 in) in length. This would suggest that the waters of this locality were rather shallow, which could indicate that it was a 3992:
does indeed possess some traits associated with terrestrial locomotion, however, they are generally not as well developed as one would expect from a fully terrestrial animal. Among these is the development of the
898:, the first teeth of the lower jaw slide into very pronounced pits that pierce the bones of the upper jaw, emerging on its upper surface in front of the nostrils. The remaining premaxillary and maxillary teeth of 764:
appears typically crocodilian, being roughly triangular with a rostrum that begins relatively narrow and continues to widen towards the back. The precise degree to which the snout tapers differs between species.
4137:. Based on the advanced state of the infection, it is thought to have been chronic and to have been affecting the individual for years. The final pathological specimen noted by Buchanan is a highly deformed 640:
was not recognised as a distinct species until the discovery of nearly complete skulls in 1993, clearly showing major differences with the type species. Among these differences is the fact that the teeth of
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and their position in the lower jaw may have increased and focused the force of the bite in a smaller area, enhancing their ability to puncture and drawing further parallels to mammalian carnassials.
4141:, which in some areas is twice as wide as it should be. The reason for the extreme deformation of this particular specimen is not entirely clear, but could have been caused by one of several possible 4032:
one seen in most extant crocodilians, while certain adaptations of the arm are also advantageous for swimming. Buchanan interprets this to mean that while not being a true terrestrial crocodilian,
4344:
Salisbury, S. W.; Willis, P. M. A. (1996). "A new crocodylian from the early Eocene of south-eastern Queensland and a preliminary investigation of the phylogenetic relationships of crocodyloids".
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show well defined attachment sites for musculature used to hold and crush prey, an activity the pterygoid would not be involved in. This collaboration between different muscle groups would allow
645:
interlock, rather than forming an overbite. It is this fact that the species name derives from. Additionally, the skull of this species was relatively more gracile, as are this species' teeth.
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use of the limb. NMV P227802 on the other hand represents a heavily altered humerus, characterised by being swollen and containing numerous sinuses filled with small bone fragments, signs of
4178:
were a part of the Tingamarra Fauna, which also included a variety of terrestrial and aquatic animals. Mammals are represented through various marsupials, while the reptile fauna includes
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There are also differences concerning the distal end, the end towards the toes rather than the hip. Here, modern Australian crocodiles have flat medial and lateral surfaces bordering the
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individual to manage. Subsequently, the turtle was let go or escaped the crocodile, allowing the bite to somewhat heal. Despite this however, the turtle later died from infection and
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stands out as having the most strongly developed processes, which correlates with larger pterygoid muscles and a stronger bite than that of its older relatives. This may suggest that
777:
was relatively flat, lacking the raised edges seen in today's crocodiles, but instead multiple specimens across species show the presence of well developed pits just before the
355:. It is generally thought to have been a semi-aquatic generalist, living a lifestyle similar to many of today's crocodiles. Four species are currently recognised, the sympatric 1091:, which may be a subadult, has been estimated to have been 2.5 m (8 ft 2 in), while a much larger mandible suggests lengths of up to 3.5 m (11 ft). 911:
the teeth clearly interlock in the premaxilla as well, however, while the first pair of dentary teeth also extend into the bone, they do not breach the dorsal surface as in
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is stated to derive from an Aboriginal word for crocodile, however, it is not specified which language in particular, only that it stems from the Queensland language group.
537:, was named three years later on the basis of new material collected from the same locality, with some of the previously recovered fossils now being assigned to this taxon. 487: 2403:. However, these results are a minority within the team's analysis and even then nodal support for paraphyletic Mekosuchinae is considered to be poor by the authors. 3940:
Most postcranial material remains unpublished, the exception being the humerus, which is known from an undetermined species found at the Murgon locality (possibly
3786:
was the basal most species of the genus, which would have rendered the overbite characteristic for this species the ancestral condition. However, the discovery of
790: 6680: 6454: 3969:"pelvic form two" displays a knob situated atop the postacetabular crest which appears to be convergently developed with the constriction of modern crocodylids. 6478: 713:. It was named for Murgon, the closest township to the type locality. It was a moderately robust animal with a skull much wider than that of the contemporary 958:
while also becoming more bulbous and peg-like. While the number of premaxillary teeth is consistent, the amount of teeth in the maxilla and dentary varies.
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species, but has enough differences to the other forms to be considered distinct. Most notably, the teeth appear to be intermediate between the overbite of
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in articulation. Ribs and chevrons are likewise known. Most of this material however have not seen formal publication and are only described in Buchanan's
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in 1982, who reported a lower jaw bone he believed to belong to a Miocene crocodylid. While noting similarities to other Australian crocodilians, notably
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bone death. While it is unclear how the animal was infected, it is possible that the area was exposed by an injury, possibly from the attack of another
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Some additional discussion of other postcranial material can be found in Buchanan's PhD thesis, which also contained the later published description of
1140:
in 1993, but it was still recognised as the basalmost member of Mekosuchinae. Subsequent papers largely supported this suggestion, typically recovering
773:, although still platyrostral, is notably slender compared to its contemporary relative. Similarly, the teeth are also more slender. The skull table of 4694:"Phylogenetic analysis of a new morphological dataset elucidates the evolutionary history of Crocodylia and resolves the long-standing gharial problem" 668: 372: 6857: 4024:
bonebed was likely the result of mass death caused by drought, something that would be less likely to affect an animal less dependent on water.
962:
has the lowest number of teeth in the maxillae (and thus throughout the upper jaw) with only 14 teeth on either side. The number is unknown for
4747:"Variation in the pelvic and pectoral girdles of Australian Oligo–Miocene mekosuchine crocodiles with implications for locomotion and habitus" 3869:
was capable of taking on larger, more terrestrial prey items, whereas older species fed on smaller animals. Furthermore, multiple species of
528:, Ralph Molnar and John D. Scanlon, who tentatively referred all Murgon crocodile fossils to this taxon. Even at this time it was noted that 4814: 4001:, the later of which only differing slightly from saltwater crocodiles and with only limited mobility of the ankle. The possibility that 1144:
as one of the earliest branching mekosuchines, although other taxa may be found as slightly more basal or on an even level as part of a
751:
people and simply means crocodile. This language was chosen as a proxy for that of the Bailai due to the links between the two cultures.
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With its appearance generally resembling today's crocodiles, in particular the flat, platyrostral snout, it is generally thought that
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was known for a while, with the type specimen found in 1991. However the material was not closely examined until the description of
6937: 6844: 4107:, but also at least two hatchlings and some fragmentary egg shells. This may suggest that the Murgon site could have been used by 3960:
would have been capable of high walking at a greater speed than the crocodiles found in Australia now. The pelvic bones of
2353:
was still recovered as a mekosuchine, however, the overall make up of the clade was changed significantly. The study found
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meanwhile is much straighter, with both surfaces being level to each other. As a consequence, the shaft of the humerus in
454:
analyses recover it as one of the earliest diverging mekosuchines, but recent studies have commonly found other taxa like
4568:
The systematics, palaeobiology and palaeoecology of Kambara taraina sp. nov. from the Eocene Rundle Formation, Queensland
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sp. nov. (Crocodylia, Crocodyloidea), a new Eocene mekosuchine from Queensland, Australia, and a revision of the genus".
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Ristevski, J.; Willis, P.M.A.; Yates, A.M.; White, M.A.; Hart, L.J.; Stein, M.D.; Price, G.J.; Salisbury, S.W. (2023).
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was slightly more gracile both in regards to its skull and teeth in addition to the different ways the teeth occluded.
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were not found to be mekosuchines in this analysis. Another, much more different placement was also shown in Ristevski
4836: 544:, although it stands out as the species with the least amount of available material. Like with the species before it, 426:
may have been better adapted at walking on land than the two crocodile species still inhabiting Australia today, the
383:
were medium-sized crocodilians, with mature specimens generally reaching lengths from 3–4 m (9.8–13.1 ft).
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and the dentition. Regarding the retroarticular process, an important attachment site for the abductor musculature,
7001: 4807: 4638:"Tip-dating and homoplasy: reconciling the shallow molecular divergences of modern gharials with their long fossil" 508:(named nine years later), Molnar refrained from naming the material due to the fragmentary nature of the dentary. 6996: 6740: 6073: 6745: 6119: 5988: 2852: 1777: 1136:
in this group and redefining Mekosuchidae as Mekosuchinae. No detailed phylogenetic analysis was conducted for
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interlock and thus set it apart from the other known species. The species name derives from the dialect of the
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in the Boat Mountain area near Murgon, Queensland, indicating that the two species coexisted. Because of this,
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for instance is described as being moderately broad-snouted with a robust build. By comparison, the skull of
735:, it stems from the Kerosene Creek Member of the Rundle Formation, which means that it did not coexist with 3528: 3178: 4398:"A new species of mekosuchine crocodilian from the middle Palaeogene Rundle Formation, central Queensland" 4103:
other than the fact that the Murgon fossil site preserves not just the remains of large, presumably adult
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As indicated by aspects of the skull shape, namely the resemblance to modern semi-aquatic ambush hunters,
1148:. For instance, a 2018 study by Lee and Yates, using genetic, morphological and stratigraphic data, found 5980: 5147: 4800: 4589:"Migrations, diversifications and extinctions: the evolutionary history of crocodyliforms in Australasia" 4148:
At least some of these pathologies are explainable through intraspecific combat, meaning that individual
3461: 2677: 2055: 872:, the teeth of the upper jaw overlap those of the lower jaw, giving it an overbite similar to that of an 6201: 6173: 4119: 3030: 2341:
Although Mekosuchinae as a monophyletic clade became a well established group in the years following
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is considered an important piece in understanding the evolutionary history and origin of this group.
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An example for modern sympatric crocodilians includes American alligators and American crocodiles.
687:. It was named in honour of Ralph Molnar for his consistent contributions to mekosuchine research. 6194: 4235:
may have been much more widespread than currently known. Holt, Salisbury and Willis propose that
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stemming from the younger deposits of the Kerosene Creek Member instead of the Brick Kiln Seam.
563:, but unlike the cranial material have not been formally published. An exception to this is the 6784: 6704: 6187: 4224:. Unlike with the species of the Tingamarra Fauna, the Rundle species were not sympatric, with 4063: 3584: 2780: 1940: 33: 6862: 1095:
is thought to have been of similar size, albeit with a more robust build. The same applies to
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was how two rather similar species of the same genus could coexist as closely as it did with
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s description, it is not without competing ideas. This in particular affects the position of
2028: 6924: 6831: 6180: 5869: 5862: 5812: 5610: 4600: 4516: 4442: 4353: 4245: 3294: 2612: 1974: 1526: 431: 4206:. Furthermore, it is known that the locality was subject to cyclical wet and dry seasons. 8: 5848: 5575: 4129: 3845:
Buchanan speculated on the potential implications for the intermediate dentition seen in
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range of motion is more certain based on the articular surface, and would suggest that
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Although so far only known from two sedimentary basins, each preserving two species,
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is known from a partial mandible and several isolated remains all collected from the
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has been found, most of said material remains undescribed with the exception of the
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of Australia, with the first remains found in rock units of what may be part of the
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are easiest differentiated by the different occlusal patterns of their teeth. While
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possess 16 teeth in either maxilla. The lower jaw contains 17 or 18 dentary teeth.
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which appears to have been attacked by a crocodile. The fossil, a nearly complete
498:. Although known for half a century, the material was first formally described by 6915: 6696: 6672: 6635: 6555: 6303: 6030: 5789: 5774: 5739: 5682: 5552: 5408: 5353: 5170: 5154: 4454: 4123:
Crocodiles will often attack their opponents' limbs during intraspecific combat.
6807: 6569: 6547: 6289: 6281: 6239: 6037: 5972: 5958: 5831: 5544: 5528: 5457: 5415: 5373: 5345: 5323: 5280: 5215: 5202: 5177: 5162: 5133: 5064: 5020: 5005: 2968: 2946: 1616: 1214: 1033: 504: 462: 4365: 3880: 1099:, which was described as reaching lengths from 3–4 m (9.8–13.1 ft). 470:
was no mekosuchine at all, but said results are not supported by all authors.
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may have been much more adapt at leaving the water than members of the genus
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Stein, Michael D.; Yates, Adam; Hand, Suzanne J.; Archer, Michael (2017).
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One particular question that was raised in the wake of the description of
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was a medium-sized crocodilian, smaller than the more recent mekosuchines
865:
appears wave-like, though it is more gentle than some other mekosuchines.
6816: 6540: 6533: 6331: 6310: 6274: 6246: 6225: 5767: 5760: 5732: 5725: 5711: 5652: 5505: 5441: 5401: 5380: 5308: 5287: 5229: 5103: 5071: 5028: 4528: 4187: 3561: 3154: 1345: 1166: 1049: 1041: 525: 456: 63: 4712: 930:, which shows a somewhat intermediate condition between the overbite of 414:, although incompletely known, seems to represent an intermediate form. 6942: 6849: 6494: 6358: 6163: 6087: 6080: 6066: 5645: 5262: 5042: 4942: 4854: 4824: 4792: 4763: 4191: 3998: 3352: 2878: 1589: 1132: 1061: 890: 495: 344: 234: 108: 73: 3972: 6444: 5782: 5565: 5049: 4915: 4221: 4203: 4080: 4013: 3937:, some elements of the postcranium suggest a more complex lifestyle. 3095: 2364: 2174: 1083: 873: 830: 576: 483: 407: 395: 352: 168: 113: 57: 6778: 4289: 6909: 6801: 6647: 6135: 5696: 5316: 5056: 4872: 4848: 4218: 4195: 4006: 3909: 3901: 3893: 3054: 2835: 2221: 1636: 1145: 1126: 1037: 748: 188: 103: 98: 83: 78: 68: 4290:"An early Eocene crocodilian from Murgon, southeastern Queensland" 1014:
is not lengthened as is the case in modern Australian crocodiles.
575:
has been recovered from the Rundle Formation, however, it was not
130: 6929: 6836: 4199: 1018: 998: 602: 564: 334: 198: 118: 93: 3816: 3803:
is typically removed due to the limited nature of its material.
1064:. Their general proportions seem to resemble modern crocodiles. 5632: 4842: 4138: 4099:
Little has been published on the potential nesting behavior of
4076: 3944:). Based on the specific anatomy of the deltapectoral crest of 982: 850:
had interlocking teeth similar to that of a crocodile (bottom).
560: 491: 364: 348: 178: 50: 4635: 3892:, but does exist. Specifically, Buchanan's thesis describes a 1130:. Willis and colleagues took this a step further by including 1032:
Additional known elements of the postcranial skeleton include
5078: 4698: 4142: 1562: 337: 842:
had an overbite, much like modern alligators (top), whereas
6151: 5013: 4017: 3078: 2201: 1448: 1077: 3888:
Direct evidence for predator prey interaction is rare for
3877:
to attack large prey, before restraining and crushing it.
2399:
as an early diverging crocodyloid and immediate sister to
4692:
Rio, Jonathan P.; Mannion, Philip D. (6 September 2021).
4586: 4194:
environment. Fish are also known from this fauna, namely
32:
For an Indian poet in Kannada language by same name, see
4571:(PhD thesis). Melbourne: Monash University. p. 340. 4431: 466:
to be the oldest diverging. Other studies suggest that
4396:
Holt, T. R.; Salisbury, S. W.; Willis, P.M.A. (2005).
4288:
Willis, P. M. A.; Molnar, R.E.; Scanlon, J.D. (1993).
3849:. Buchanan argued that the confluent tooth sockets of 1021:, which makes the end look somewhat quadrilateral. In 406:
had interlocking dentition much more similar to true
4744: 4593:
Alcheringa: An Australasian Journal of Palaeontology
4435:
Alcheringa: An Australasian Journal of Palaeontology
2225: 2205: 2178: 2106: 2086: 2059: 2032: 2005: 1978: 1944: 1899: 1879: 1852: 1818: 1781: 1640: 1620: 1593: 1566: 1530: 1510: 1453: 1432: 1405: 1369: 1349: 1308: 1272: 1252: 1218: 314: 305: 296: 283: 4395: 4287: 4016:, which would allow for hinge-like movement of the 2349:. In a study dealing exclusively with morphology, 802:being described as long but narrow, while that of 4636:Michael S. Y. Lee; Adam M. Yates (27 June 2018). 934:and the more extensive interlocking dentition of 540:A third species was named in 2005 in the form of 6983: 2367:crocodyloid no longer believed to be related to 4343: 1056:, several dorsal and tail vertebrae as well as 571:and noted to differ greatly from modern forms. 446:As the oldest described and named mekosuchine, 856:The toothrow across all four known species of 5213: 4808: 4209:The Rundle Formation, which was home to both 1048:, hindlimbs as well as multiple parts of the 4217:, on the other hand is thought to represent 4887: 4079:. One possible explanation may be found in 3919: 942:and more closely resembles animals such as 709:and have been found in areas overlying the 555:The most recent addition to this genus was 4815: 4801: 4691: 1178:to be the basalmost forms. In both cases, 993:Although a lot of postcranial material of 701:The type species of the genus, remains of 417:Despite plentiful fossils, the ecology of 129: 4772: 4762: 4721: 4711: 4665: 4655: 4612: 4427: 4425: 4423: 4421: 4419: 4417: 4415: 4005:possessed a "fibular condyle" similar to 3828:. It broadly resembles modern crocodiles. 809: 4822: 4564: 4502: 4283: 4281: 4118: 4062: 3971: 3879: 3815: 981: 923:is characterised by being interlocking. 671:. It is the least well preserved of all 601: 4687: 4685: 4339: 4337: 4335: 4333: 4331: 4329: 4327: 4279: 4277: 4275: 4273: 4271: 4269: 4267: 4265: 4263: 4261: 3884:A modern Alligator "juggling" a turtle. 3776: 2335: 14: 6984: 4982: 4738: 4582: 4580: 4578: 4498: 4496: 4494: 4492: 4412: 4325: 4323: 4321: 4319: 4317: 4315: 4313: 4311: 4309: 4307: 3837:could serve a similar function as the 2929: 2408: 1750: 1187: 614:, scalebar = 10 cm (3.9 in). 6783: 6782: 5921: 5200: 4981: 4835: 4796: 4560: 4558: 4556: 4554: 4552: 4550: 4548: 4546: 4490: 4488: 4486: 4484: 4482: 4480: 4478: 4476: 4474: 4472: 4391: 4389: 4387: 4385: 4383: 4381: 4379: 4377: 4375: 3951:The range of motion for the limbs of 524:, was finally established in 1993 by 473: 4682: 4258: 4248:inhabit most of northern Australia. 1107:Similarities between a then unnamed 394:had an overbite similar to a modern 289:Willis, Molnar & Scanlon, 1993 ( 4629: 4575: 4304: 4165: 3811: 24: 5201: 4643:Proceedings of the Royal Society B 4543: 4509:Journal of Vertebrate Paleontology 4469: 4372: 3916:was specialised for hunting them. 2357:in a basal position and including 679:and the interlocking dentition of 379:, also from the Rundle Formation. 311:Holt, Salisbury & Willis, 2005 270:Willis, Molnar & Scanlon, 1993 25: 7018: 915:. Regardless, much like those of 784:Ever since the first material of 590:In the type description the name 7007:Prehistoric pseudosuchian genera 6744: 6739: 5922: 5904: 5899: 4402:Memoirs of the Queensland Museum 829: 818: 155: 61: 3806: 3782:Initially, it was thought that 2395:. Instead, these trees recover 1182:was found to be a mekosuchine. 482:are exclusively known from the 4244:may be similar to how today's 4114: 1052:, including nearly the entire 926:The third type is observed in 755: 632:were found alongside those of 375:and the youngest of the four, 13: 1: 4614:10.1080/03115518.2023.2201319 4251: 4071:In their 1996 description of 3820:The skull of an indetermined 3795:as the basal most species of 977: 567:, which was covered by Stein 4455:10.1080/03115518.2012.671697 3988:. Here, Buchanan notes that 3529:Brachyuranochampsa eversolei 3179:Orientalosuchus naduongensis 1102: 302:Salisbury & Willis, 1996 7: 4043: 2678:Mekosuchus whitehunterensis 2505:"Asiatosuchus" nanlingensis 2360:"Asiatosuchus" nanlingensis 2056:Mekosuchus whitehunterensis 1087:. The holotype specimen of 711:Oakdale Sandstone Formation 488:Oakdale Sandstone Formation 10: 7023: 4094: 3896:from the same locality as 3479:"Crocodylus" depressifrons 3203:Jiangxisuchus nankangensis 1040:, bones of the lower arm, 597: 31: 6899: 6791: 6757: 6737: 6645: 6506: 6493: 6442: 6395: 6382: 6367: 6356: 6234: 6223: 6161: 6114: 6105: 6056: 6025: 6014: 5945: 5934: 5930: 5917: 5897: 5829: 5800: 5640: 5631: 5563: 5485: 5476: 5429: 5363: 5275: 5271: 5260: 5224: 5209: 5196: 5088: 4995: 4991: 4977: 4941: 4923: 4914: 4896: 4883: 4831: 4366:10.1080/03115519608619189 3688: 3653: 3636: 3629: 3612: 3605: 3598: 3581: 3574: 3559: 3552: 3525: 3508: 3501: 3475: 3458: 3451: 3444: 3427: 3420: 3349: 3332: 3315: 3308: 3291: 3284: 3267: 3260: 3216: 3199: 3192: 3175: 3168: 3155:Krabisuchus siamogallicus 3151: 3144: 3136: 3092: 3075: 3068: 3051: 3044: 3031:Kalthifrons aurivellensis 3027: 3020: 3013: 2999:"Asiatosuchus" germanicus 2995: 2988: 2981: 2966: 2959: 2944: 2937: 2875: 2849: 2832: 2825: 2818: 2802: 2795: 2777: 2770: 2708: 2691: 2674: 2657: 2650: 2633: 2626: 2609: 2602: 2576: 2559: 2552: 2545: 2501: 2484: 2477: 2460: 2453: 2436: 2429: 2422: 2415: 2292: 2218: 2198: 2191: 2171: 2164: 2099: 2079: 2072: 2052: 2045: 2025: 2018: 1998: 1991: 1971: 1964: 1957: 1937: 1930: 1892: 1872: 1865: 1845: 1838: 1831: 1815:Kalthifrons aurivellensis 1811: 1804: 1794: 1774: 1767: 1757: 1725: 1633: 1613: 1606: 1586: 1579: 1559: 1552: 1523: 1507: 1500: 1493: 1445: 1425: 1418: 1398: 1391: 1362: 1342: 1335: 1328: 1321: 1301: 1294: 1265: 1245: 1238: 1231: 1211: 1204: 1194: 583:, which stems from older 280: 275: 152:Scientific classification 150: 137: 128: 43: 29:Extinct genus of reptiles 6074:"Crocodylus" gariepensis 4503:Buchanan, L.A. (2009). " 3980:performing the highwalk. 3920:Locomotion and lifestyle 731:The youngest species of 7002:Crocodiles of Australia 6347:"Tomistoma" lusitanicum 6120:"Crocodylus" megarhinus 5989:"Crocodylus" megarhinus 4565:Buchanan, L.A. (2008). 4296:(3): 27–33 – via 3336:Trilophosuchus rackhami 2853:"Crocodylus" megarhinus 2712:Mekosuchus inexpectatus 2637:Trilophosuchus rackhami 2103:Mekosuchus inexpectatus 2002:Trilophosuchus rackhami 1778:"Crocodylus" megarhinus 1067: 779:supratemporal fenestrae 367:, the poorly preserved 6997:Eocene crocodylomorphs 4657:10.1098/rspb.2018.1071 4124: 4068: 3981: 3885: 3829: 3585:Australosuchus clarkae 2781:Australosuchus clarkae 1941:Australosuchus clarkae 990: 810:Toothrow and dentition 791:retroarticular process 615: 347:that lived during the 34:Chandrashekhar Kambara 6964:Paleobiology Database 6884:Paleobiology Database 4246:freshwater crocodiles 4122: 4066: 3975: 3883: 3819: 3431:Prodiplocynodon langi 3319:Volia athollandersoni 2371:. Additionally, both 2029:Volia athollandersoni 1152:to be second only to 985: 659:The third species of 605: 490:near the township of 142:and the lower jaw of 6622:"Tomistoma" coppensi 6614:"Tomistoma" cairense 6340:"Tomistoma" cairense 5981:"Crocodylus" affinis 5148:"Crocodylus" affinis 4529:10.1671/039.029.0220 4190:, which inhabited a 3462:"Crocodylus" affinis 1160:alternatively found 1156:, whereas Ristevski 432:freshwater crocodile 363:from sediments near 6901:Kambara murgonensis 6629:"Tomistoma" dowsoni 4713:10.7717/peerj.12094 4605:2023Alch...47..370R 4521:2009JVPal..29..473B 4447:2012Alch...36..473S 4358:1996Alch...20..179S 4143:tumorous conditions 4130:comminuted fracture 4054:Kambara murgonensis 4050:Kambara implexidens 3784:Kambara murgonensis 3657:Kambara murgonensis 3616:Kambara implexidens 3271:Dongnanosuchus hsui 3220:Eoalligator chunyii 2661:Ultrastenos willisi 2488:Kambara murgonensis 2440:Kambara implexidens 1896:Kambara murgonensis 1849:Kambara implexidens 1509:Bullock Creek taxon 1305:Kambara murgonensis 1269:Kambara implexidens 1138:Kambara murgonensis 988:Kambara implexidens 986:Size comparison of 919:, the dentition of 806:was short and low. 612:Kambara murgonensis 608:Kambara implexidens 606:Comparison between 535:Kambara implexidens 361:Kambara implexidens 357:Kambara murgonensis 140:Kambara implexidens 6202:C. thorbjarnarsoni 5251:Listrognathosuchus 5127:Brachyuranochampsa 4764:10.7717/peerj.3501 4184:meiolaniid turtles 4125: 4069: 3982: 3978:American crocodile 3886: 3830: 2695:Mekosuchus sanderi 2083:Mekosuchus sanderi 991: 616: 474:History and naming 6979: 6978: 6951:Open Tree of Life 6871:Open Tree of Life 6785:Taxon identifiers 6776: 6775: 6753: 6752: 6735: 6734: 6731: 6730: 6727: 6726: 6723: 6722: 6489: 6488: 6219: 6218: 6215: 6214: 6211: 6210: 6174:C. anthropophagus 5913: 5912: 5895: 5894: 5891: 5890: 5887: 5886: 5883: 5882: 5879: 5878: 5627: 5626: 5425: 5424: 5388:Eurycephalosuchus 5192: 5191: 5188: 5187: 5141:"Crocodylus" acer 4973: 4972: 4969: 4968: 4965: 4964: 4961: 4960: 4180:softshell turtles 3780: 3779: 3773: 3772: 3764: 3763: 3755: 3754: 3746: 3745: 3737: 3736: 3728: 3727: 3719: 3718: 3710: 3709: 3701: 3700: 3677: 3676: 3668: 3667: 3540: 3539: 3512:"Crocodylus" acer 3490: 3489: 3409: 3408: 3400: 3399: 3391: 3390: 3382: 3381: 3373: 3372: 3364: 3363: 3249: 3248: 3240: 3239: 3231: 3230: 3142:Orientalosuchina 3125: 3124: 3116: 3115: 3107: 3106: 2926: 2925: 2917: 2916: 2908: 2907: 2899: 2898: 2890: 2889: 2864: 2863: 2759: 2758: 2750: 2749: 2741: 2740: 2732: 2731: 2723: 2722: 2591: 2590: 2534: 2533: 2525: 2524: 2516: 2515: 2339: 2338: 2332: 2331: 2323: 2322: 2314: 2313: 2305: 2304: 2281: 2280: 2272: 2271: 2263: 2262: 2254: 2253: 2245: 2244: 2236: 2235: 2153: 2152: 2144: 2143: 2135: 2134: 2126: 2125: 2117: 2116: 1919: 1918: 1910: 1909: 1747: 1746: 1738: 1737: 1714: 1713: 1705: 1704: 1696: 1695: 1687: 1686: 1678: 1677: 1669: 1668: 1660: 1659: 1651: 1650: 1541: 1540: 1482: 1481: 1473: 1472: 1464: 1463: 1380: 1379: 1283: 1282: 705:were part of the 663:to be described, 326: 325: 271: 16:(Redirected from 7014: 6972: 6971: 6959: 6958: 6946: 6945: 6933: 6932: 6920: 6919: 6918: 6892: 6891: 6879: 6878: 6866: 6865: 6853: 6852: 6840: 6839: 6827: 6826: 6825: 6812: 6811: 6810: 6780: 6779: 6748: 6743: 6705:G. pachyrhynchus 6607:Siquisiquesuchus 6504: 6503: 6393: 6392: 6365: 6364: 6232: 6231: 6112: 6111: 6023: 6022: 5943: 5942: 5932: 5931: 5919: 5918: 5908: 5903: 5870:C. wannlangstoni 5863:C. venezuelensis 5705:Globidentosuchus 5668:Centenariosuchus 5638: 5637: 5499:Allognathosuchus 5483: 5482: 5474: 5473: 5365:Orientalosuchina 5273: 5272: 5269: 5268: 5222: 5221: 5211: 5210: 5198: 5197: 4993: 4992: 4979: 4978: 4921: 4920: 4894: 4893: 4885: 4884: 4878: 4877: 4833: 4832: 4817: 4810: 4803: 4794: 4793: 4787: 4786: 4776: 4766: 4742: 4736: 4735: 4725: 4715: 4689: 4680: 4679: 4669: 4659: 4633: 4627: 4626: 4616: 4584: 4573: 4572: 4562: 4541: 4540: 4500: 4467: 4466: 4429: 4410: 4409: 4393: 4370: 4369: 4341: 4302: 4301: 4285: 4188:madtsoiid snakes 4166:Paleoenvironment 3839:carnassial teeth 3826:Melbourne Museum 3812:Ecology and diet 3632: 3631: 3608: 3607: 3601: 3600: 3577: 3576: 3555: 3554: 3504: 3503: 3454: 3453: 3447: 3446: 3423: 3422: 3311: 3310: 3287: 3286: 3263: 3262: 3195: 3194: 3171: 3170: 3147: 3146: 3139: 3138: 3071: 3070: 3047: 3046: 3023: 3022: 3016: 3015: 2991: 2990: 2984: 2983: 2962: 2961: 2940: 2939: 2932: 2931: 2828: 2827: 2821: 2820: 2798: 2797: 2773: 2772: 2653: 2652: 2629: 2628: 2605: 2604: 2555: 2554: 2548: 2547: 2480: 2479: 2456: 2455: 2432: 2431: 2425: 2424: 2418: 2417: 2411: 2410: 2406: 2405: 2387:, but excluding 2385:Orientalosuchina 2227: 2207: 2194: 2193: 2180: 2167: 2166: 2108: 2088: 2075: 2074: 2061: 2048: 2047: 2034: 2021: 2020: 2007: 1994: 1993: 1980: 1967: 1966: 1960: 1959: 1946: 1933: 1932: 1901: 1881: 1868: 1867: 1854: 1841: 1840: 1834: 1833: 1820: 1807: 1806: 1797: 1796: 1783: 1770: 1769: 1760: 1759: 1753: 1752: 1642: 1622: 1609: 1608: 1595: 1582: 1581: 1568: 1555: 1554: 1532: 1512: 1503: 1502: 1496: 1495: 1455: 1434: 1421: 1420: 1407: 1394: 1393: 1371: 1351: 1338: 1337: 1331: 1330: 1324: 1323: 1310: 1297: 1296: 1274: 1254: 1241: 1240: 1234: 1233: 1220: 1207: 1206: 1197: 1196: 1190: 1189: 1185: 1184: 1058:lumbar vertebrae 833: 822: 707:Tingamarra Fauna 669:Rundle Formation 373:Rundle Formation 316: 307: 298: 285: 269: 262: 249: 225:Archosauriformes 212:Archosauromorpha 160: 159: 133: 123: 60: 49:Temporal range: 41: 40: 21: 7022: 7021: 7017: 7016: 7015: 7013: 7012: 7011: 6982: 6981: 6980: 6975: 6967: 6962: 6954: 6949: 6941: 6936: 6928: 6923: 6914: 6913: 6908: 6895: 6887: 6882: 6874: 6869: 6861: 6856: 6848: 6843: 6835: 6830: 6821: 6820: 6815: 6806: 6805: 6800: 6787: 6777: 6772: 6749: 6719: 6681:G. curvirostris 6658:G. bengawanicus 6641: 6636:Toyotamaphimeia 6556:Hesperogavialis 6497: 6485: 6438: 6386: 6378: 6361: 6352: 6304:Megadontosuchus 6228: 6207: 6195:C. palaeindicus 6157: 6101: 6052: 6031:Aldabrachampsus 6019: 6010: 5939: 5926: 5909: 5875: 5849:C. brevirostris 5825: 5796: 5790:Wannaganosuchus 5775:Stangerochampsa 5740:Orthogenysuchus 5683:Chinatichampsus 5623: 5559: 5553:Wannaganosuchus 5472: 5434: 5421: 5409:Orientalosuchus 5359: 5354:Stangerochampsa 5265: 5256: 5218: 5205: 5184: 5171:Prodiplocynodon 5155:Portugalosuchus 5084: 4987: 4957: 4937: 4910: 4879: 4838: 4837: 4827: 4821: 4791: 4790: 4743: 4739: 4690: 4683: 4634: 4630: 4585: 4576: 4563: 4544: 4505:Kambara taraina 4501: 4470: 4430: 4413: 4394: 4373: 4342: 4305: 4286: 4259: 4254: 4168: 4155:Kambara taraina 4117: 4097: 4046: 4022:Kambara taraina 3986:Kambara taraina 3922: 3898:Kambara taraina 3847:Kambara molnari 3824:species at the 3814: 3809: 3788:Kambara molnari 3774: 3765: 3756: 3747: 3738: 3729: 3720: 3711: 3702: 3678: 3669: 3640:Kambara taraina 3541: 3491: 3410: 3401: 3392: 3383: 3374: 3365: 3353:Mekosuchus spp. 3250: 3241: 3232: 3126: 3117: 3108: 2927: 2918: 2909: 2900: 2891: 2865: 2760: 2751: 2742: 2733: 2724: 2592: 2535: 2526: 2517: 2464:Kambara taraina 2333: 2324: 2315: 2306: 2282: 2273: 2264: 2255: 2246: 2237: 2154: 2145: 2136: 2127: 2118: 1920: 1911: 1876:Kambara taraina 1748: 1739: 1715: 1706: 1697: 1688: 1679: 1670: 1661: 1652: 1542: 1483: 1474: 1465: 1381: 1284: 1249:Kambara taraina 1105: 1070: 1054:cervical series 1034:shoulder blades 980: 854: 853: 852: 851: 836: 835: 834: 825: 824: 823: 812: 758: 743:, the teeth of 628:The fossils of 600: 476: 377:Kambara taraina 369:Kambara molnari 268: 260: 247: 154: 144:Kambara molnari 124: 122: 121: 116: 111: 106: 101: 96: 91: 86: 81: 76: 71: 66: 55: 54: 47: 37: 30: 23: 22: 15: 12: 11: 5: 7020: 7010: 7009: 7004: 6999: 6994: 6977: 6976: 6974: 6973: 6960: 6947: 6934: 6921: 6905: 6903: 6897: 6896: 6894: 6893: 6880: 6867: 6854: 6841: 6828: 6813: 6797: 6795: 6789: 6788: 6774: 6773: 6771: 6770: 6765: 6758: 6755: 6754: 6751: 6750: 6738: 6736: 6733: 6732: 6729: 6728: 6725: 6724: 6721: 6720: 6718: 6717: 6709: 6701: 6693: 6685: 6677: 6669: 6661: 6653: 6651: 6643: 6642: 6640: 6639: 6632: 6625: 6618: 6610: 6603: 6596: 6589: 6582: 6574: 6570:Maomingosuchus 6566: 6559: 6552: 6548:Harpacochampsa 6544: 6537: 6530: 6522: 6515: 6507: 6501: 6491: 6490: 6487: 6486: 6484: 6483: 6475: 6471:T. lusitanicum 6467: 6459: 6455:T. calaritanum 6450: 6448: 6440: 6439: 6437: 6436: 6428: 6420: 6413: 6405: 6396: 6390: 6380: 6379: 6377: 6376: 6368: 6362: 6357: 6354: 6353: 6351: 6350: 6343: 6336: 6328: 6321: 6314: 6307: 6300: 6293: 6290:Maomingosuchus 6286: 6282:Leptorrhamphus 6278: 6271: 6264: 6257: 6250: 6243: 6240:Dollosuchoides 6235: 6229: 6224: 6221: 6220: 6217: 6216: 6213: 6212: 6209: 6208: 6206: 6205: 6198: 6191: 6188:C. falconensis 6184: 6177: 6169: 6167: 6159: 6158: 6156: 6155: 6148: 6140: 6132: 6124: 6115: 6109: 6103: 6102: 6100: 6099: 6091: 6084: 6077: 6070: 6062: 6060: 6054: 6053: 6051: 6050: 6042: 6038:Dzungarisuchus 6034: 6026: 6020: 6015: 6012: 6011: 6009: 6008: 6000: 5992: 5985: 5977: 5973:Australosuchus 5969: 5962: 5959:Antecrocodylus 5955: 5946: 5940: 5935: 5928: 5927: 5915: 5914: 5911: 5910: 5898: 5896: 5893: 5892: 5889: 5888: 5885: 5884: 5881: 5880: 5877: 5876: 5874: 5873: 5866: 5859: 5852: 5845: 5837: 5835: 5827: 5826: 5824: 5823: 5820:M. latrubessei 5816: 5808: 5806: 5798: 5797: 5795: 5794: 5786: 5779: 5771: 5764: 5757: 5750: 5743: 5736: 5729: 5722: 5715: 5708: 5701: 5693: 5686: 5679: 5671: 5664: 5656: 5649: 5641: 5635: 5629: 5628: 5625: 5624: 5622: 5621: 5614: 5607: 5600: 5593: 5586: 5579: 5571: 5569: 5561: 5560: 5558: 5557: 5549: 5545:Procaimanoidea 5541: 5533: 5529:Hassiacosuchus 5525: 5518: 5510: 5502: 5495: 5486: 5480: 5471: 5470: 5462: 5458:Menatalligator 5454: 5446: 5437: 5435: 5430: 5427: 5426: 5423: 5422: 5420: 5419: 5416:Protoalligator 5412: 5405: 5398: 5391: 5384: 5377: 5374:Dongnanosuchus 5369: 5367: 5361: 5360: 5358: 5357: 5350: 5346:Procaimanoidea 5342: 5335: 5327: 5324:Hassiacosuchus 5320: 5313: 5305: 5298: 5291: 5284: 5281:Albertochampsa 5276: 5266: 5261: 5258: 5257: 5255: 5254: 5247: 5240: 5233: 5225: 5219: 5216:Alligatoroidea 5214: 5207: 5206: 5203:Alligatoroidea 5194: 5193: 5190: 5189: 5186: 5185: 5183: 5182: 5178:Planocraniidae 5174: 5167: 5163:Pristichampsus 5159: 5151: 5144: 5137: 5134:Charactosuchus 5130: 5123: 5115: 5108: 5100: 5092: 5090: 5086: 5085: 5083: 5082: 5075: 5068: 5065:Trilophosuchus 5061: 5053: 5046: 5039: 5032: 5025: 5021:Harpacochampsa 5017: 5010: 5006:Australosuchus 5001: 4999: 4989: 4988: 4975: 4974: 4971: 4970: 4967: 4966: 4963: 4962: 4959: 4958: 4956: 4955: 4954: 4953: 4947: 4945: 4939: 4938: 4936: 4935: 4934: 4933: 4924: 4918: 4912: 4911: 4909: 4908: 4907: 4906: 4897: 4891: 4881: 4880: 4876: 4875: 4866: 4857: 4851: 4845: 4829: 4828: 4820: 4819: 4812: 4805: 4797: 4789: 4788: 4737: 4681: 4628: 4599:(4): 370–415. 4574: 4542: 4515:(2): 473–486. 4468: 4441:(4): 473–486. 4411: 4371: 4352:(3): 179–226. 4303: 4256: 4255: 4253: 4250: 4176:K. implexidens 4172:K. murgonensis 4167: 4164: 4116: 4113: 4096: 4093: 4073:K. implexidens 4058:K. implexidens 4045: 4042: 3942:K. implexidens 3935:Trilophosuchus 3921: 3918: 3813: 3810: 3808: 3805: 3793:K. implexidens 3778: 3777: 3775: 3771: 3770: 3767: 3766: 3762: 3761: 3758: 3757: 3753: 3752: 3749: 3748: 3744: 3743: 3740: 3739: 3735: 3734: 3731: 3730: 3726: 3725: 3722: 3721: 3717: 3716: 3713: 3712: 3708: 3707: 3704: 3703: 3699: 3698: 3695: 3694: 3687: 3684: 3683: 3680: 3679: 3675: 3674: 3671: 3670: 3666: 3665: 3662: 3661: 3652: 3649: 3648: 3645: 3644: 3635: 3630: 3628: 3625: 3624: 3621: 3620: 3611: 3606: 3604: 3599: 3597: 3594: 3593: 3590: 3589: 3580: 3575: 3573: 3570: 3569: 3566: 3565: 3558: 3553: 3551: 3547: 3546: 3543: 3542: 3538: 3537: 3534: 3533: 3524: 3521: 3520: 3517: 3516: 3507: 3502: 3500: 3497: 3496: 3493: 3492: 3488: 3487: 3484: 3483: 3474: 3471: 3470: 3467: 3466: 3457: 3452: 3450: 3445: 3443: 3440: 3439: 3436: 3435: 3426: 3421: 3419: 3416: 3415: 3412: 3411: 3407: 3406: 3403: 3402: 3398: 3397: 3394: 3393: 3389: 3388: 3385: 3384: 3380: 3379: 3376: 3375: 3371: 3370: 3367: 3366: 3362: 3361: 3358: 3357: 3348: 3345: 3344: 3341: 3340: 3331: 3328: 3327: 3324: 3323: 3314: 3309: 3307: 3304: 3303: 3300: 3299: 3290: 3285: 3283: 3280: 3279: 3276: 3275: 3266: 3261: 3259: 3256: 3255: 3252: 3251: 3247: 3246: 3243: 3242: 3238: 3237: 3234: 3233: 3229: 3228: 3225: 3224: 3215: 3212: 3211: 3208: 3207: 3198: 3193: 3191: 3188: 3187: 3184: 3183: 3174: 3169: 3167: 3164: 3163: 3160: 3159: 3150: 3145: 3143: 3137: 3135: 3132: 3131: 3128: 3127: 3123: 3122: 3119: 3118: 3114: 3113: 3110: 3109: 3105: 3104: 3101: 3100: 3096:Paludirex spp. 3091: 3088: 3087: 3084: 3083: 3074: 3069: 3067: 3064: 3063: 3060: 3059: 3050: 3045: 3043: 3040: 3039: 3036: 3035: 3026: 3021: 3019: 3014: 3012: 3008: 3007: 3004: 3003: 2994: 2989: 2987: 2982: 2980: 2977: 2976: 2973: 2972: 2969:Planocraniidae 2965: 2960: 2958: 2955: 2954: 2951: 2950: 2947:Alligatoroidea 2943: 2938: 2936: 2930: 2928: 2924: 2923: 2920: 2919: 2915: 2914: 2911: 2910: 2906: 2905: 2902: 2901: 2897: 2896: 2893: 2892: 2888: 2887: 2884: 2883: 2874: 2871: 2870: 2867: 2866: 2862: 2861: 2858: 2857: 2848: 2845: 2844: 2841: 2840: 2831: 2826: 2824: 2819: 2817: 2813: 2812: 2809: 2808: 2801: 2796: 2794: 2790: 2789: 2786: 2785: 2776: 2771: 2769: 2766: 2765: 2762: 2761: 2757: 2756: 2753: 2752: 2748: 2747: 2744: 2743: 2739: 2738: 2735: 2734: 2730: 2729: 2726: 2725: 2721: 2720: 2717: 2716: 2707: 2704: 2703: 2700: 2699: 2690: 2687: 2686: 2683: 2682: 2673: 2670: 2669: 2666: 2665: 2656: 2651: 2649: 2646: 2645: 2642: 2641: 2632: 2627: 2625: 2622: 2621: 2618: 2617: 2608: 2603: 2601: 2598: 2597: 2594: 2593: 2589: 2588: 2585: 2584: 2575: 2572: 2571: 2568: 2567: 2558: 2553: 2551: 2546: 2544: 2541: 2540: 2537: 2536: 2532: 2531: 2528: 2527: 2523: 2522: 2519: 2518: 2514: 2513: 2510: 2509: 2500: 2497: 2496: 2493: 2492: 2483: 2478: 2476: 2473: 2472: 2469: 2468: 2459: 2454: 2452: 2449: 2448: 2445: 2444: 2435: 2430: 2428: 2423: 2421: 2416: 2414: 2409: 2389:Australosuchus 2377:Australosuchus 2337: 2336: 2334: 2330: 2329: 2326: 2325: 2321: 2320: 2317: 2316: 2312: 2311: 2308: 2307: 2303: 2302: 2299: 2298: 2291: 2288: 2287: 2284: 2283: 2279: 2278: 2275: 2274: 2270: 2269: 2266: 2265: 2261: 2260: 2257: 2256: 2252: 2251: 2248: 2247: 2243: 2242: 2239: 2238: 2234: 2233: 2230: 2229: 2217: 2214: 2213: 2210: 2209: 2197: 2192: 2190: 2187: 2186: 2183: 2182: 2175:Paludirex spp. 2170: 2165: 2163: 2160: 2159: 2156: 2155: 2151: 2150: 2147: 2146: 2142: 2141: 2138: 2137: 2133: 2132: 2129: 2128: 2124: 2123: 2120: 2119: 2115: 2114: 2111: 2110: 2098: 2095: 2094: 2091: 2090: 2078: 2073: 2071: 2068: 2067: 2064: 2063: 2051: 2046: 2044: 2041: 2040: 2037: 2036: 2024: 2019: 2017: 2014: 2013: 2010: 2009: 1997: 1992: 1990: 1987: 1986: 1983: 1982: 1970: 1965: 1963: 1958: 1956: 1953: 1952: 1949: 1948: 1936: 1931: 1929: 1926: 1925: 1922: 1921: 1917: 1916: 1913: 1912: 1908: 1907: 1904: 1903: 1891: 1888: 1887: 1884: 1883: 1871: 1866: 1864: 1861: 1860: 1857: 1856: 1844: 1839: 1837: 1832: 1830: 1827: 1826: 1823: 1822: 1810: 1805: 1803: 1795: 1793: 1790: 1789: 1786: 1785: 1773: 1768: 1766: 1758: 1756: 1751: 1749: 1745: 1744: 1741: 1740: 1736: 1735: 1732: 1731: 1724: 1721: 1720: 1717: 1716: 1712: 1711: 1708: 1707: 1703: 1702: 1699: 1698: 1694: 1693: 1690: 1689: 1685: 1684: 1681: 1680: 1676: 1675: 1672: 1671: 1667: 1666: 1663: 1662: 1658: 1657: 1654: 1653: 1649: 1648: 1645: 1644: 1632: 1629: 1628: 1625: 1624: 1617:Trilophosuchus 1612: 1607: 1605: 1602: 1601: 1598: 1597: 1585: 1580: 1578: 1575: 1574: 1571: 1570: 1558: 1553: 1551: 1548: 1547: 1544: 1543: 1539: 1538: 1535: 1534: 1522: 1519: 1518: 1515: 1514: 1506: 1501: 1499: 1494: 1492: 1489: 1488: 1485: 1484: 1480: 1479: 1476: 1475: 1471: 1470: 1467: 1466: 1462: 1461: 1458: 1457: 1444: 1441: 1440: 1437: 1436: 1424: 1419: 1417: 1414: 1413: 1410: 1409: 1397: 1392: 1390: 1387: 1386: 1383: 1382: 1378: 1377: 1374: 1373: 1361: 1358: 1357: 1354: 1353: 1341: 1336: 1334: 1329: 1327: 1322: 1320: 1317: 1316: 1313: 1312: 1300: 1295: 1293: 1290: 1289: 1286: 1285: 1281: 1280: 1277: 1276: 1264: 1261: 1260: 1257: 1256: 1244: 1239: 1237: 1232: 1230: 1227: 1226: 1223: 1222: 1215:Australosuchus 1210: 1205: 1203: 1195: 1193: 1188: 1162:Australosuchus 1154:Australosuchus 1114:Australosuchus 1104: 1101: 1093:K. murgonensis 1089:K. implexidens 1069: 1066: 979: 976: 972:K. murgonensis 968:K. implexidens 936:K. implexidens 932:K. murgonensis 917:K. implexidens 913:K. implexidens 905:K. murgonensis 900:K. implexidens 896:K. implexidens 881:K. implexidens 879:However, both 870:K. murgonensis 844:K. implexidens 840:K. murgonensis 838: 837: 828: 827: 826: 817: 816: 815: 814: 813: 811: 808: 804:K. implexidens 771:K. implexidens 767:K. murgonensis 757: 754: 753: 752: 741:K. implexidens 728: 727: 719: 718: 715:K. implexidens 703:K. murgonensis 698: 697: 694:K. murgonensis 689: 688: 681:K. implexidens 677:K. murgonensis 656: 655: 647: 646: 643:K. implexidens 638:K. implexidens 634:K. murgonensis 630:K. implexidens 625: 624: 621:K. implexidens 599: 596: 550:K. implexidens 518:K. murgonensis 505:Australosuchus 475: 472: 463:Australosuchus 440:K. implexidens 436:K. murgonensis 400:K. implexidens 392:K. murgonensis 324: 323: 322: 321: 320:Buchanan, 2009 312: 303: 300:K. implexidens 294: 287:K. murgonensis 278: 277: 273: 272: 258: 254: 253: 245: 238: 237: 232: 228: 227: 222: 215: 214: 209: 202: 201: 196: 192: 191: 186: 182: 181: 176: 172: 171: 166: 162: 161: 148: 147: 135: 134: 126: 125: 117: 112: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 48: 28: 9: 6: 4: 3: 2: 7019: 7008: 7005: 7003: 7000: 6998: 6995: 6993: 6990: 6989: 6987: 6970: 6965: 6961: 6957: 6952: 6948: 6944: 6939: 6935: 6931: 6926: 6922: 6917: 6911: 6907: 6906: 6904: 6902: 6898: 6890: 6885: 6881: 6877: 6872: 6868: 6864: 6859: 6855: 6851: 6846: 6842: 6838: 6833: 6829: 6824: 6818: 6814: 6809: 6803: 6799: 6798: 6796: 6794: 6790: 6786: 6781: 6769: 6768:Gryposuchinae 6766: 6764: 6760: 6759: 6756: 6747: 6742: 6715: 6714: 6710: 6707: 6706: 6702: 6699: 6698: 6694: 6691: 6690: 6686: 6683: 6682: 6678: 6675: 6674: 6670: 6667: 6666: 6662: 6660: 6659: 6655: 6654: 6652: 6650: 6649: 6644: 6638: 6637: 6633: 6631: 6630: 6626: 6624: 6623: 6619: 6616: 6615: 6611: 6609: 6608: 6604: 6602: 6601: 6600:Rhamphosuchus 6597: 6595: 6594: 6593:Piscogavialis 6590: 6588: 6587: 6583: 6580: 6579: 6578:Paratomistoma 6575: 6572: 6571: 6567: 6565: 6564: 6563:Ikanogavialis 6560: 6558: 6557: 6553: 6550: 6549: 6545: 6543: 6542: 6538: 6536: 6535: 6531: 6528: 6527: 6526:Gavialosuchus 6523: 6521: 6520: 6516: 6514: 6513: 6512:Aktiogavialis 6509: 6508: 6505: 6502: 6500: 6496: 6492: 6481: 6480: 6479:T. taiwanicum 6476: 6473: 6472: 6468: 6465: 6464: 6460: 6457: 6456: 6452: 6451: 6449: 6447: 6446: 6441: 6434: 6433: 6429: 6426: 6425: 6424:Paratomistoma 6421: 6419: 6418: 6414: 6411: 6410: 6409:Gavialosuchus 6406: 6403: 6402: 6401:Brasilosuchus 6398: 6397: 6394: 6391: 6389: 6388:sensu stricto 6385: 6381: 6375: 6374: 6370: 6369: 6366: 6363: 6360: 6355: 6349: 6348: 6344: 6342: 6341: 6337: 6334: 6333: 6329: 6327: 6326: 6322: 6320: 6319: 6318:Paratomistoma 6315: 6313: 6312: 6308: 6306: 6305: 6301: 6299: 6298: 6297:Maroccosuchus 6294: 6292: 6291: 6287: 6284: 6283: 6279: 6277: 6276: 6272: 6270: 6269: 6268:Gunggamarandu 6265: 6263: 6262: 6261:Gavialosuchus 6258: 6256: 6255: 6254:Ferganosuchus 6251: 6249: 6248: 6244: 6242: 6241: 6237: 6236: 6233: 6230: 6227: 6222: 6204: 6203: 6199: 6197: 6196: 6192: 6190: 6189: 6185: 6183: 6182: 6178: 6176: 6175: 6171: 6170: 6168: 6166: 6165: 6160: 6154: 6153: 6149: 6146: 6145: 6144:Tzaganosuchus 6141: 6138: 6137: 6133: 6130: 6129: 6125: 6122: 6121: 6117: 6116: 6113: 6110: 6108: 6104: 6097: 6096: 6092: 6090: 6089: 6085: 6083: 6082: 6078: 6076: 6075: 6071: 6069: 6068: 6064: 6063: 6061: 6059: 6058:Osteolaeminae 6055: 6048: 6047: 6046:Oxysdonsaurus 6043: 6040: 6039: 6035: 6033: 6032: 6028: 6027: 6024: 6021: 6018: 6013: 6006: 6005: 6001: 5998: 5997: 5996:Jiangxisuchus 5993: 5991: 5990: 5986: 5983: 5982: 5978: 5975: 5974: 5970: 5968: 5967: 5966:Astorgosuchus 5963: 5961: 5960: 5956: 5953: 5952: 5951:Albertosuchus 5948: 5947: 5944: 5941: 5938: 5937:Crocodyloidea 5933: 5929: 5925: 5920: 5916: 5907: 5902: 5872: 5871: 5867: 5865: 5864: 5860: 5858: 5857: 5856:C. gasparinae 5853: 5851: 5850: 5846: 5844: 5843: 5839: 5838: 5836: 5834: 5833: 5828: 5822: 5821: 5817: 5815: 5814: 5810: 5809: 5807: 5805: 5804: 5799: 5792: 5791: 5787: 5785: 5784: 5780: 5777: 5776: 5772: 5770: 5769: 5765: 5763: 5762: 5758: 5756: 5755: 5751: 5749: 5748: 5744: 5742: 5741: 5737: 5735: 5734: 5730: 5728: 5727: 5723: 5721: 5720: 5719:Kuttanacaiman 5716: 5714: 5713: 5709: 5707: 5706: 5702: 5699: 5698: 5694: 5692: 5691: 5690:Culebrasuchus 5687: 5685: 5684: 5680: 5677: 5676: 5672: 5670: 5669: 5665: 5662: 5661: 5660:Brachychampsa 5657: 5655: 5654: 5650: 5648: 5647: 5643: 5642: 5639: 5636: 5634: 5630: 5620: 5619: 5615: 5613: 5612: 5611:A. prenasalis 5608: 5606: 5605: 5601: 5599: 5598: 5594: 5592: 5591: 5587: 5585: 5584: 5580: 5578: 5577: 5573: 5572: 5570: 5568: 5567: 5562: 5555: 5554: 5550: 5547: 5546: 5542: 5539: 5538: 5534: 5531: 5530: 5526: 5524: 5523: 5522:Chrysochampsa 5519: 5516: 5515: 5511: 5508: 5507: 5503: 5501: 5500: 5496: 5493: 5492: 5491:Akanthosuchus 5488: 5487: 5484: 5481: 5479: 5478:Alligatorinae 5475: 5468: 5467: 5466:Sajkanosuchus 5463: 5460: 5459: 5455: 5452: 5451: 5450:Lianghusuchus 5447: 5444: 5443: 5439: 5438: 5436: 5433: 5432:Alligatoridae 5428: 5418: 5417: 5413: 5411: 5410: 5406: 5404: 5403: 5399: 5397: 5396: 5395:Jiangxisuchus 5392: 5390: 5389: 5385: 5383: 5382: 5378: 5376: 5375: 5371: 5370: 5368: 5366: 5362: 5356: 5355: 5351: 5348: 5347: 5343: 5341: 5340: 5336: 5333: 5332: 5328: 5326: 5325: 5321: 5319: 5318: 5314: 5311: 5310: 5306: 5304: 5303: 5299: 5297: 5296: 5295:Brachychampsa 5292: 5290: 5289: 5285: 5283: 5282: 5278: 5277: 5274: 5270: 5267: 5264: 5259: 5253: 5252: 5248: 5246: 5245: 5241: 5239: 5238: 5234: 5232: 5231: 5227: 5226: 5223: 5220: 5217: 5212: 5208: 5204: 5199: 5195: 5180: 5179: 5175: 5173: 5172: 5168: 5165: 5164: 5160: 5157: 5156: 5152: 5150: 5149: 5145: 5143: 5142: 5138: 5136: 5135: 5131: 5129: 5128: 5124: 5121: 5120: 5119:Borealosuchus 5116: 5114: 5113: 5109: 5106: 5105: 5101: 5099: 5098: 5097:Albertosuchus 5094: 5093: 5091: 5087: 5081: 5080: 5076: 5074: 5073: 5069: 5067: 5066: 5062: 5059: 5058: 5054: 5052: 5051: 5047: 5045: 5044: 5040: 5038: 5037: 5033: 5031: 5030: 5026: 5023: 5022: 5018: 5016: 5015: 5011: 5008: 5007: 5003: 5002: 5000: 4998: 4994: 4990: 4985: 4980: 4976: 4951: 4950: 4949: 4948: 4946: 4944: 4940: 4932: 4928: 4927: 4926: 4925: 4922: 4919: 4917: 4913: 4905: 4901: 4900: 4899: 4898: 4895: 4892: 4890: 4886: 4882: 4874: 4870: 4867: 4865: 4861: 4858: 4856: 4852: 4850: 4846: 4844: 4840: 4839: 4834: 4830: 4826: 4818: 4813: 4811: 4806: 4804: 4799: 4798: 4795: 4784: 4780: 4775: 4770: 4765: 4760: 4756: 4752: 4748: 4741: 4733: 4729: 4724: 4719: 4714: 4709: 4705: 4701: 4700: 4695: 4688: 4686: 4677: 4673: 4668: 4663: 4658: 4653: 4649: 4645: 4644: 4639: 4632: 4624: 4620: 4615: 4610: 4606: 4602: 4598: 4594: 4590: 4583: 4581: 4579: 4570: 4569: 4561: 4559: 4557: 4555: 4553: 4551: 4549: 4547: 4538: 4534: 4530: 4526: 4522: 4518: 4514: 4510: 4506: 4499: 4497: 4495: 4493: 4491: 4489: 4487: 4485: 4483: 4481: 4479: 4477: 4475: 4473: 4464: 4460: 4456: 4452: 4448: 4444: 4440: 4436: 4428: 4426: 4424: 4422: 4420: 4418: 4416: 4407: 4403: 4399: 4392: 4390: 4388: 4386: 4384: 4382: 4380: 4378: 4376: 4367: 4363: 4359: 4355: 4351: 4347: 4340: 4338: 4336: 4334: 4332: 4330: 4328: 4326: 4324: 4322: 4320: 4318: 4316: 4314: 4312: 4310: 4308: 4299: 4295: 4291: 4284: 4282: 4280: 4278: 4276: 4274: 4272: 4270: 4268: 4266: 4264: 4262: 4257: 4249: 4247: 4243: 4238: 4234: 4229: 4227: 4223: 4220: 4216: 4212: 4207: 4205: 4201: 4197: 4193: 4189: 4185: 4181: 4177: 4173: 4163: 4161: 4156: 4151: 4146: 4144: 4140: 4136: 4135:osteomyelitis 4131: 4121: 4112: 4110: 4106: 4102: 4092: 4088: 4086: 4082: 4078: 4074: 4065: 4061: 4059: 4055: 4051: 4041: 4039: 4035: 4030: 4025: 4023: 4019: 4015: 4011: 4008: 4004: 4000: 3996: 3991: 3987: 3979: 3974: 3970: 3967: 3963: 3959: 3954: 3949: 3947: 3943: 3938: 3936: 3932: 3927: 3917: 3915: 3911: 3907: 3903: 3899: 3895: 3894:chelid turtle 3891: 3882: 3878: 3876: 3872: 3868: 3864: 3860: 3854: 3852: 3848: 3843: 3840: 3835: 3827: 3823: 3818: 3804: 3802: 3798: 3794: 3789: 3785: 3769: 3768: 3760: 3759: 3751: 3750: 3742: 3741: 3733: 3732: 3724: 3723: 3715: 3714: 3706: 3705: 3697: 3696: 3693: 3692: 3686: 3685: 3682: 3681: 3673: 3672: 3664: 3663: 3660: 3659: 3658: 3651: 3650: 3647: 3646: 3643: 3642: 3641: 3634: 3633: 3627: 3626: 3623: 3622: 3619: 3618: 3617: 3610: 3609: 3603: 3602: 3596: 3595: 3592: 3591: 3588: 3587: 3586: 3579: 3578: 3572: 3571: 3568: 3567: 3564: 3563: 3557: 3556: 3550:Longirostres 3549: 3548: 3545: 3544: 3536: 3535: 3532: 3531: 3530: 3523: 3522: 3519: 3518: 3515: 3514: 3513: 3506: 3505: 3499: 3498: 3495: 3494: 3486: 3485: 3482: 3481: 3480: 3473: 3472: 3469: 3468: 3465: 3464: 3463: 3456: 3455: 3449: 3448: 3442: 3441: 3438: 3437: 3434: 3433: 3432: 3425: 3424: 3418: 3417: 3414: 3413: 3405: 3404: 3396: 3395: 3387: 3386: 3378: 3377: 3369: 3368: 3360: 3359: 3356: 3355: 3354: 3347: 3346: 3343: 3342: 3339: 3338: 3337: 3330: 3329: 3326: 3325: 3322: 3321: 3320: 3313: 3312: 3306: 3305: 3302: 3301: 3298: 3297: 3296: 3295:"Baru" huberi 3289: 3288: 3282: 3281: 3278: 3277: 3274: 3273: 3272: 3265: 3264: 3258: 3257: 3254: 3253: 3245: 3244: 3236: 3235: 3227: 3226: 3223: 3222: 3221: 3214: 3213: 3210: 3209: 3206: 3205: 3204: 3197: 3196: 3190: 3189: 3186: 3185: 3182: 3181: 3180: 3173: 3172: 3166: 3165: 3162: 3161: 3158: 3157: 3156: 3149: 3148: 3141: 3140: 3134: 3133: 3130: 3129: 3121: 3120: 3112: 3111: 3103: 3102: 3099: 3098: 3097: 3090: 3089: 3086: 3085: 3082: 3081: 3080: 3073: 3072: 3066: 3065: 3062: 3061: 3058: 3057: 3056: 3055:Quinkana spp. 3049: 3048: 3042: 3041: 3038: 3037: 3034: 3033: 3032: 3025: 3024: 3018: 3017: 3011:Mekosuchinae 3010: 3009: 3006: 3005: 3002: 3001: 3000: 2993: 2992: 2986: 2985: 2979: 2978: 2975: 2974: 2971: 2970: 2964: 2963: 2957: 2956: 2953: 2952: 2949: 2948: 2942: 2941: 2934: 2933: 2922: 2921: 2913: 2912: 2904: 2903: 2895: 2894: 2886: 2885: 2882: 2881: 2880: 2873: 2872: 2869: 2868: 2860: 2859: 2856: 2855: 2854: 2847: 2846: 2843: 2842: 2839: 2838: 2837: 2830: 2829: 2823: 2822: 2816:Crocodylinae 2815: 2814: 2811: 2810: 2807: 2806: 2805:Osteolaeminae 2800: 2799: 2793:Crocodylidae 2792: 2791: 2788: 2787: 2784: 2783: 2782: 2775: 2774: 2768: 2767: 2764: 2763: 2755: 2754: 2746: 2745: 2737: 2736: 2728: 2727: 2719: 2718: 2715: 2714: 2713: 2706: 2705: 2702: 2701: 2698: 2697: 2696: 2689: 2688: 2685: 2684: 2681: 2680: 2679: 2672: 2671: 2668: 2667: 2664: 2663: 2662: 2655: 2654: 2648: 2647: 2644: 2643: 2640: 2639: 2638: 2631: 2630: 2624: 2623: 2620: 2619: 2616: 2615: 2614: 2613:"Baru" huberi 2607: 2606: 2600: 2599: 2596: 2595: 2587: 2586: 2583: 2582: 2581: 2574: 2573: 2570: 2569: 2566: 2565: 2564: 2557: 2556: 2550: 2549: 2543: 2542: 2539: 2538: 2530: 2529: 2521: 2520: 2512: 2511: 2508: 2507: 2506: 2499: 2498: 2495: 2494: 2491: 2490: 2489: 2482: 2481: 2475: 2474: 2471: 2470: 2467: 2466: 2465: 2458: 2457: 2451: 2450: 2447: 2446: 2443: 2442: 2441: 2434: 2433: 2427: 2426: 2420: 2419: 2413: 2412: 2407: 2404: 2402: 2398: 2394: 2390: 2386: 2382: 2378: 2374: 2370: 2366: 2362: 2361: 2356: 2352: 2348: 2344: 2328: 2327: 2319: 2318: 2310: 2309: 2301: 2300: 2297: 2296: 2290: 2289: 2286: 2285: 2277: 2276: 2268: 2267: 2259: 2258: 2250: 2249: 2241: 2240: 2232: 2231: 2228: 2224: 2223: 2222:Quinkana spp. 2216: 2215: 2212: 2211: 2208: 2204: 2203: 2196: 2195: 2189: 2188: 2185: 2184: 2181: 2177: 2176: 2169: 2168: 2162: 2161: 2158: 2157: 2149: 2148: 2140: 2139: 2131: 2130: 2122: 2121: 2113: 2112: 2109: 2105: 2104: 2097: 2096: 2093: 2092: 2089: 2085: 2084: 2077: 2076: 2070: 2069: 2066: 2065: 2062: 2058: 2057: 2050: 2049: 2043: 2042: 2039: 2038: 2035: 2031: 2030: 2023: 2022: 2016: 2015: 2012: 2011: 2008: 2004: 2003: 1996: 1995: 1989: 1988: 1985: 1984: 1981: 1977: 1976: 1975:"Baru" huberi 1969: 1968: 1962: 1961: 1955: 1954: 1951: 1950: 1947: 1943: 1942: 1935: 1934: 1928: 1927: 1924: 1923: 1915: 1914: 1906: 1905: 1902: 1898: 1897: 1890: 1889: 1886: 1885: 1882: 1878: 1877: 1870: 1869: 1863: 1862: 1859: 1858: 1855: 1851: 1850: 1843: 1842: 1836: 1835: 1829: 1828: 1825: 1824: 1821: 1817: 1816: 1809: 1808: 1802: 1799: 1798: 1792: 1791: 1788: 1787: 1784: 1780: 1779: 1772: 1771: 1765: 1764:Crocodyloidea 1762: 1761: 1755: 1754: 1743: 1742: 1734: 1733: 1730: 1729: 1723: 1722: 1719: 1718: 1710: 1709: 1701: 1700: 1692: 1691: 1683: 1682: 1674: 1673: 1665: 1664: 1656: 1655: 1647: 1646: 1643: 1639: 1638: 1631: 1630: 1627: 1626: 1623: 1619: 1618: 1611: 1610: 1604: 1603: 1600: 1599: 1596: 1592: 1591: 1584: 1583: 1577: 1576: 1573: 1572: 1569: 1565: 1564: 1557: 1556: 1550: 1549: 1546: 1545: 1537: 1536: 1533: 1529: 1528: 1527:"Baru" huberi 1521: 1520: 1517: 1516: 1513: 1505: 1504: 1498: 1497: 1491: 1490: 1487: 1486: 1478: 1477: 1469: 1468: 1460: 1459: 1456: 1451: 1450: 1443: 1442: 1439: 1438: 1435: 1431: 1430: 1423: 1422: 1416: 1415: 1412: 1411: 1408: 1404: 1403: 1396: 1395: 1389: 1388: 1385: 1384: 1376: 1375: 1372: 1368: 1367: 1366:Pallimnarchus 1360: 1359: 1356: 1355: 1352: 1348: 1347: 1340: 1339: 1333: 1332: 1326: 1325: 1319: 1318: 1315: 1314: 1311: 1307: 1306: 1299: 1298: 1292: 1291: 1288: 1287: 1279: 1278: 1275: 1271: 1270: 1263: 1262: 1259: 1258: 1255: 1251: 1250: 1243: 1242: 1236: 1235: 1229: 1228: 1225: 1224: 1221: 1217: 1216: 1209: 1208: 1202: 1199: 1198: 1192: 1191: 1186: 1183: 1181: 1177: 1173: 1169: 1168: 1163: 1159: 1155: 1151: 1147: 1143: 1139: 1135: 1134: 1129: 1128: 1123: 1119: 1118:Pallimnarchus 1115: 1110: 1100: 1098: 1094: 1090: 1086: 1085: 1080: 1079: 1074: 1065: 1063: 1059: 1055: 1051: 1047: 1043: 1039: 1035: 1030: 1028: 1024: 1020: 1015: 1013: 1009: 1004: 1000: 996: 989: 984: 975: 973: 969: 966:, while both 965: 961: 955: 953: 952: 947: 946: 945:Borealosuchus 941: 937: 933: 929: 924: 922: 918: 914: 910: 906: 901: 897: 893: 892: 886: 882: 877: 875: 871: 866: 864: 859: 849: 845: 841: 832: 821: 807: 805: 801: 796: 792: 787: 782: 780: 776: 772: 768: 763: 760:The skull of 750: 746: 742: 738: 734: 730: 729: 726: 725: 721: 720: 716: 712: 708: 704: 700: 699: 696: 695: 691: 690: 686: 682: 678: 674: 670: 666: 662: 658: 657: 654: 653: 649: 648: 644: 639: 635: 631: 627: 626: 623: 622: 618: 617: 613: 609: 604: 595: 593: 588: 586: 585:stratigraphic 582: 578: 574: 570: 566: 562: 558: 553: 551: 547: 543: 538: 536: 531: 527: 523: 519: 515: 509: 507: 506: 501: 497: 493: 489: 485: 481: 471: 469: 465: 464: 459: 458: 453: 449: 444: 441: 437: 433: 429: 425: 420: 415: 413: 409: 405: 401: 397: 393: 389: 384: 382: 378: 374: 370: 366: 362: 358: 354: 350: 346: 343: 339: 336: 332: 331: 319: 313: 310: 304: 301: 295: 292: 288: 282: 281: 279: 274: 267: 266: 259: 256: 255: 252: 246: 243: 240: 239: 236: 233: 230: 229: 226: 223: 220: 217: 216: 213: 210: 207: 204: 203: 200: 197: 194: 193: 190: 187: 184: 183: 180: 177: 174: 173: 170: 167: 164: 163: 158: 153: 149: 145: 141: 138:The skull of 136: 132: 127: 120: 115: 110: 105: 100: 95: 90: 85: 80: 75: 70: 65: 59: 56:54.6–29  52: 46: 42: 39: 35: 27: 19: 6992:Mekosuchinae 6900: 6792: 6763:Brevirostres 6713:G. papuensis 6711: 6703: 6695: 6687: 6679: 6671: 6665:G. breviceps 6663: 6656: 6646: 6634: 6627: 6620: 6612: 6605: 6598: 6591: 6586:Penghusuchus 6584: 6576: 6568: 6561: 6554: 6546: 6539: 6532: 6524: 6519:Dadagavialis 6517: 6510: 6498: 6477: 6469: 6461: 6453: 6443: 6432:Thecachampsa 6430: 6422: 6417:Melitosaurus 6415: 6407: 6399: 6387: 6384:Tomistominae 6373:Sacacosuchus 6371: 6345: 6338: 6330: 6325:Thecachampsa 6323: 6316: 6309: 6302: 6295: 6288: 6280: 6273: 6266: 6259: 6252: 6245: 6238: 6200: 6193: 6186: 6181:C. checchiai 6179: 6172: 6162: 6150: 6142: 6134: 6126: 6118: 6107:Crocodylinae 6093: 6086: 6079: 6072: 6065: 6044: 6036: 6029: 6017:Crocodylidae 6004:Mekosuchinae 6002: 5994: 5987: 5979: 5971: 5964: 5957: 5949: 5924:Longirostres 5868: 5861: 5854: 5847: 5842:C. australis 5840: 5830: 5818: 5811: 5803:Melanosuchus 5801: 5788: 5781: 5773: 5766: 5759: 5754:Paranasuchus 5752: 5747:Paranacaiman 5745: 5738: 5731: 5724: 5717: 5710: 5703: 5695: 5688: 5681: 5675:Ceratosuchus 5673: 5666: 5658: 5651: 5644: 5616: 5609: 5602: 5595: 5588: 5581: 5576:A. hailensis 5574: 5564: 5551: 5543: 5537:Navajosuchus 5535: 5527: 5520: 5514:Ceratosuchus 5512: 5504: 5497: 5489: 5464: 5456: 5448: 5440: 5414: 5407: 5400: 5393: 5386: 5379: 5372: 5352: 5344: 5339:Navajosuchus 5337: 5331:Leidyosuchus 5329: 5322: 5315: 5307: 5302:Ceratosuchus 5300: 5293: 5286: 5279: 5249: 5244:Leidyosuchus 5242: 5237:Diplocynodon 5235: 5228: 5176: 5169: 5161: 5153: 5146: 5139: 5132: 5125: 5117: 5112:Asiatosuchus 5110: 5102: 5095: 5077: 5070: 5063: 5055: 5048: 5041: 5035: 5034: 5027: 5019: 5012: 5004: 4997:Mekosuchinae 4986:crocodilians 4904:Pseudosuchia 4889:Pseudosuchia 4868: 4864:Pseudosuchia 4859: 4825:crocodilians 4754: 4750: 4740: 4703: 4697: 4647: 4641: 4631: 4596: 4592: 4567: 4512: 4508: 4504: 4438: 4434: 4405: 4401: 4349: 4345: 4298:Academia.edu 4293: 4241: 4236: 4232: 4230: 4225: 4214: 4210: 4208: 4175: 4171: 4169: 4159: 4154: 4149: 4147: 4126: 4108: 4104: 4100: 4098: 4089: 4084: 4072: 4070: 4057: 4053: 4049: 4047: 4037: 4033: 4028: 4026: 4021: 4010:rauisuchians 4002: 3989: 3985: 3983: 3965: 3961: 3957: 3952: 3950: 3945: 3941: 3939: 3934: 3930: 3925: 3923: 3913: 3905: 3897: 3889: 3887: 3874: 3870: 3866: 3862: 3858: 3857:prey. Among 3855: 3850: 3846: 3844: 3833: 3831: 3821: 3807:Paleobiology 3800: 3796: 3792: 3787: 3783: 3781: 3691:Crocodylidae 3689: 3656: 3655: 3654: 3639: 3638: 3637: 3615: 3614: 3613: 3583: 3582: 3560: 3527: 3526: 3510: 3509: 3477: 3476: 3460: 3459: 3429: 3428: 3351: 3350: 3334: 3333: 3317: 3316: 3293: 3292: 3269: 3268: 3218: 3217: 3201: 3200: 3177: 3176: 3153: 3152: 3094: 3093: 3077: 3076: 3053: 3052: 3029: 3028: 2997: 2996: 2967: 2945: 2877: 2876: 2851: 2850: 2834: 2833: 2803: 2779: 2778: 2710: 2709: 2693: 2692: 2676: 2675: 2659: 2658: 2635: 2634: 2611: 2610: 2580:Baru wickeni 2578: 2577: 2563:Baru darrowi 2561: 2560: 2503: 2502: 2487: 2486: 2485: 2463: 2462: 2461: 2439: 2438: 2437: 2401:Crocodylidae 2396: 2392: 2388: 2380: 2376: 2372: 2369:Asiatosuchus 2368: 2358: 2354: 2350: 2346: 2342: 2340: 2295:Crocodylidae 2293: 2220: 2219: 2200: 2199: 2173: 2172: 2101: 2100: 2081: 2080: 2054: 2053: 2027: 2026: 2000: 1999: 1973: 1972: 1939: 1938: 1895: 1894: 1893: 1875: 1874: 1873: 1848: 1847: 1846: 1813: 1812: 1801:Mekosuchinae 1776: 1775: 1728:Longirostres 1726: 1635: 1634: 1615: 1614: 1588: 1587: 1561: 1560: 1525: 1524: 1508: 1447: 1446: 1429:Baru darrowi 1427: 1426: 1402:Baru wickeni 1400: 1399: 1364: 1363: 1344: 1343: 1304: 1303: 1302: 1268: 1267: 1266: 1248: 1247: 1246: 1213: 1212: 1201:Mekosuchinae 1179: 1175: 1171: 1165: 1161: 1157: 1153: 1149: 1141: 1137: 1131: 1125: 1121: 1117: 1113: 1108: 1106: 1096: 1092: 1088: 1082: 1076: 1072: 1071: 1046:pelvic bones 1031: 1026: 1022: 1016: 1011: 1007: 1002: 994: 992: 987: 971: 967: 963: 959: 956: 951:Diplocynodon 949: 943: 939: 935: 931: 927: 925: 920: 916: 912: 908: 904: 899: 895: 889: 884: 880: 878: 869: 867: 862: 857: 855: 847: 843: 839: 803: 799: 794: 785: 783: 774: 770: 766: 761: 759: 744: 740: 739:. Like with 736: 732: 723: 722: 714: 702: 693: 692: 684: 680: 676: 672: 664: 660: 651: 650: 642: 637: 633: 629: 620: 619: 611: 607: 591: 589: 580: 572: 568: 556: 554: 549: 545: 541: 539: 534: 529: 522:type species 517: 513: 510: 503: 500:Ralph Molnar 479: 477: 467: 461: 455: 452:Phylogenetic 447: 445: 439: 435: 423: 418: 416: 411: 403: 399: 391: 387: 385: 380: 376: 368: 360: 356: 329: 328: 327: 317: 308: 299: 286: 264: 263: 251:Mekosuchinae 241: 218: 205: 143: 139: 44: 38: 26: 6817:Wikispecies 6689:G. leptodus 6541:Hanyusuchus 6534:Gryposuchus 6463:T. gaudense 6332:Tienosuchus 6311:Ocepesuchus 6275:Kentisuchus 6247:Dollosuchus 6226:Gavialoidea 5768:Purussaurus 5761:Protocaiman 5733:Necrosuchus 5726:Mourasuchus 5712:Gnatusuchus 5653:Bottosaurus 5618:A. thomsoni 5597:A. munensis 5590:A. mefferdi 5506:Arambourgia 5442:Balanerodus 5402:Krabisuchus 5381:Eoalligator 5309:Deinosuchus 5288:Arambourgia 5230:Deinosuchus 5104:Arenysuchus 5072:Ultrastenos 5029:Kalthifrons 4115:Pathologies 3863:K. taraina 3799:, although 3562:Gavialoidea 2935:Crocodilia 1346:Kalthifrons 1172:Kalthifrons 1167:Kalthifrons 1050:spinal cord 1042:finger bone 756:Description 526:Paul Willis 478:Remains of 457:Kalthifrons 386:Species of 345:crocodylian 342:mekosuchine 6986:Categories 6761:See also: 6499:sensu lato 6495:Gavialinae 6359:Gavialidae 6164:Crocodylus 6088:Rimasuchus 6081:Euthecodon 6067:Brochuchus 5813:M. fisheri 5646:Acresuchus 5633:Caimaninae 5583:A. mcgrewi 5263:Globidonta 5043:Mekosuchus 4952:see below↓ 4943:Crocodilia 4855:Sauropsida 4706:: e12094. 4408:: 207–218. 4346:Alcheringa 4252:References 4226:K. taraina 4215:K. taraina 4211:K. molnari 4192:lacustrine 4085:K. taraina 4038:Crocodylus 3999:astragalus 3875:K. taraina 3867:K. taraina 3851:K. molnari 3801:K. molnari 2879:Crocodylus 1590:Mekosuchus 1133:Mekosuchus 1097:K. molnari 1062:PhD thesis 978:Postcrania 964:K. molnari 960:K. taraina 928:K. molnari 921:K. taraina 909:K. taraina 891:Crocodylus 885:K. taraina 848:K. taraina 800:K. molnari 795:K. taraina 745:K. taraina 737:K. molnari 724:K. taraina 685:K. taraina 665:K. molnari 652:K. molnari 581:K. molnari 573:K. taraina 557:K. taraina 546:K. molnari 542:K. molnari 496:Queensland 412:K. molnari 408:crocodiles 404:K. taraina 318:K. taraina 309:K. molnari 235:Crocodilia 6916:Q20718518 6697:G. lewisi 6673:G. browni 6445:Tomistoma 5783:Tsoabichi 5604:A. olseni 5566:Alligator 5050:Paludirex 4931:Neosuchia 4916:Neosuchia 4841:Kingdom: 4757:: e3501. 4623:258878554 4463:129195186 4222:mud flats 4204:billabong 4111:to nest. 4081:taphonomy 4027:Overall, 4014:aetosaurs 3861:species, 3079:Baru spp. 2365:Paleocene 2202:Baru spp. 1103:Phylogeny 1084:Paludirex 1038:coracoids 874:alligator 577:sympatric 484:Paleogene 443:drought. 428:saltwater 396:alligator 371:from the 353:Australia 351:epoch in 175:Kingdom: 169:Eukaryota 6930:42335027 6910:Wikidata 6837:10588969 6808:Q5230226 6802:Wikidata 6648:Gavialis 6136:Quinkana 5697:Eocaiman 5317:Eocaiman 5057:Quinkana 4873:Eusuchia 4849:Chordata 4847:Phylum: 4843:Animalia 4823:Extinct 4783:28674657 4732:34567843 4676:30051855 4650:(1881). 4537:86254159 4219:lagoonal 4200:teleosts 4196:lungfish 4044:Sympatry 4007:Triassic 3997:and the 3931:Quinkana 3910:necrosis 3902:plastron 2836:Quinkana 2373:Quinkana 2343:Kambara' 1637:Quinkana 1146:polytomy 1127:Quinkana 1019:condyles 749:Darumbal 587:layers. 276:Species 199:Reptilia 189:Chordata 185:Phylum: 179:Animalia 165:Domain: 6956:6149935 6943:8610649 6876:3605941 6863:1201546 6850:4822295 6823:Kambara 6793:Kambara 6128:Kinyang 6095:Kinyang 5036:Kambara 4853:Class: 4774:5494174 4723:8428266 4667:6030529 4601:Bibcode 4517:Bibcode 4443:Bibcode 4354:Bibcode 4242:Kambara 4237:Kambara 4233:Kambara 4160:Kambara 4150:Kambara 4109:Kambara 4105:Kambara 4101:Kambara 4095:Nesting 4077:caimans 4034:Kambara 4029:Kambara 4003:Kambara 3990:Kambara 3966:Kambara 3962:Kambara 3958:Kambara 3953:Kambara 3946:Kambara 3926:Kambara 3914:Kambara 3906:Kambara 3890:Kambara 3871:Kambara 3859:Kambara 3834:Kambara 3822:Kambara 3797:Kambara 2397:Kambara 2393:Kambara 2355:Kambara 2351:Kambara 2347:Kambara 1452:Alcoota 1180:Kambara 1176:Kambara 1150:Kambara 1142:Kambara 1109:Kambara 1073:Kambara 1027:Kambara 1023:Kambara 1012:Kambara 1008:Kambara 1003:Kambara 999:humerus 995:Kambara 940:Kambara 863:Kambara 858:Kambara 786:Kambara 775:Kambara 762:Kambara 733:Kambara 673:Kambara 661:Kambara 598:Species 592:Kambara 565:humerus 530:Kambara 520:as the 516:, with 514:Kambara 480:Kambara 468:Kambara 448:Kambara 424:Kambara 419:Kambara 398:, both 388:Kambara 381:Kambara 335:extinct 330:Kambara 265:Kambara 257:Genus: 231:Order: 195:Class: 45:Kambara 18:Kembara 6969:265611 6889:157974 5832:Caiman 5089:Others 4781:  4771:  4730:  4720:  4674:  4664:  4621:  4535:  4461:  4294:Kaupia 4139:fibula 1158:et al. 569:et al. 561:thesis 492:Murgon 365:Murgon 349:Eocene 333:is an 51:Eocene 6858:IRMNG 5079:Volia 4984:Basal 4869:Clade 4860:Clade 4751:PeerJ 4699:PeerJ 4619:S2CID 4533:S2CID 4459:S2CID 4170:Both 3995:ilium 2381:et al 1563:Volia 907:. In 894:. In 579:with 422:that 338:genus 242:Clade 219:Clade 206:Clade 6938:GBIF 6845:GBIF 6152:Voay 5014:Baru 4929:see 4902:see 4779:PMID 4728:PMID 4672:PMID 4213:and 4198:and 4186:and 4174:and 4153:the 4018:knee 4012:and 3933:and 2391:and 2375:and 2363:, a 1449:Baru 1174:and 1124:and 1122:Baru 1081:and 1078:Baru 1068:Size 970:and 948:and 883:and 846:and 683:and 610:and 438:and 430:and 402:and 359:and 291:type 64:PreꞒ 6925:EoL 6832:EoL 4769:PMC 4759:doi 4718:PMC 4708:doi 4662:PMC 4652:doi 4648:285 4609:doi 4525:doi 4451:doi 4362:doi 3976:An 1170:or 888:of 876:. 868:In 460:or 340:of 6988:: 6966:: 6953:: 6940:: 6927:: 6912:: 6886:: 6873:: 6860:: 6847:: 6834:: 6819:: 6804:: 4871:: 4862:: 4777:. 4767:. 4753:. 4749:. 4726:. 4716:. 4702:. 4696:. 4684:^ 4670:. 4660:. 4646:. 4640:. 4617:. 4607:. 4597:47 4595:. 4591:. 4577:^ 4545:^ 4531:. 4523:. 4513:29 4511:. 4471:^ 4457:. 4449:. 4439:36 4437:. 4414:^ 4406:50 4404:. 4400:. 4374:^ 4360:. 4350:20 4348:. 4306:^ 4292:. 4260:^ 4182:, 4162:. 4145:. 4087:. 1164:, 1120:, 1116:, 1044:, 1036:, 1029:. 954:. 781:. 552:. 494:, 410:. 244:: 221:: 208:: 146:. 114:Pg 58:Ma 53:, 6716:? 6708:? 6700:? 6692:? 6684:? 6676:? 6668:? 6617:? 6581:? 6573:? 6551:? 6529:? 6482:? 6474:? 6466:? 6458:? 6435:? 6427:? 6412:? 6404:? 6335:? 6285:? 6147:? 6139:? 6131:? 6123:? 6098:? 6049:? 6041:? 6007:? 5999:? 5984:? 5976:? 5954:? 5793:? 5778:? 5700:? 5678:? 5663:? 5556:? 5548:? 5540:? 5532:? 5517:? 5509:? 5494:? 5469:? 5461:? 5453:? 5445:? 5349:? 5334:? 5312:? 5181:? 5166:? 5158:? 5122:? 5107:? 5060:? 5024:? 5009:? 4816:e 4809:t 4802:v 4785:. 4761:: 4755:5 4734:. 4710:: 4704:9 4678:. 4654:: 4625:. 4611:: 4603:: 4539:. 4527:: 4519:: 4465:. 4453:: 4445:: 4368:. 4364:: 4356:: 4300:. 2226:† 2206:† 2179:† 2107:† 2087:† 2060:† 2033:† 2006:† 1979:† 1945:† 1900:† 1880:† 1853:† 1819:† 1782:† 1641:† 1621:† 1594:† 1567:† 1531:† 1511:† 1454:† 1433:† 1406:† 1370:† 1350:† 1309:† 1273:† 1253:† 1219:† 717:. 315:† 306:† 297:† 293:) 284:† 261:† 248:† 119:N 109:K 104:J 99:T 94:P 89:C 84:D 79:S 74:O 69:Ꞓ 36:. 20:)

Index

Kembara
Chandrashekhar Kambara
Eocene
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Archosauromorpha
Archosauriformes
Crocodilia
Mekosuchinae
Kambara
type
extinct

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