594:
114:
1230:
575:. The first is the secondary regaining of the limbs from legless ancestors, while the second is the possibility of multiple episodes of leg loss in snakes. Unfortunately the current number of specimens from the family Pachyophiidae, and the complexity of snake
536:. The small overall size, underdevelopment of the tarsal ossifications, less extensive pachyostosis, and larger proportional size of the tibia and fibula are all possible traits of a juvenile. However the skull and tooth structures of
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of snakes. This is supported by the general characteristics of the skeleton. The presence of the hind limbs presents a problem with this placement however. There are two major ideas for the presence of hind limbs in
646:
748:
et al. 2003 "The
Anatomy and Relationships of Haasiophis terrasanctus, a Fossil Snake with Well-Developed Hind Limbs from the Mid-Cretaceous of the Middle East"
552:
are much taller in comparison and the rib shape is quite distinct. Therefore, while regarded as very closely related, the two are different genera.
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440:' ribs) are found beginning on the 155th vertebrae and continuing to the 160th. The ribs from the 45-48th and 105-108th vertebrae show signs of
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girdle. The pelvic girdle is partly obscured by overlaying bones reducing the detail which can be seen. Radiographs of the specimen reveal
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is 199, plus fragments of a 200th, with the rib cage extending from the 5th to the 154th vertebrae. Lymphapophyses (forked '
512:. The tarsal ossifications do not show signs that the bones were connected to each other during life. Traces of two
360:
as specimen HUJ-Pal. EJ 695., measures 88 cm, missing only the tip of the tail, and is considered a small snake.
296:, who first began work in the fossils of Ein Yabrud and started the description of the genus before he died, plus the
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and its sister genera are indicated as either a sister group to, or basal members of, the
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there is no preserved indication of a connection between the vertebral column and the
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the cranium is described as showing a mix of basal characters, like those found in
413:
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area of the skull, being small and slender, is similar to the preorbital area of
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bones of the skull are very similar to ones found in the sister genus
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The skull is well preserved though slightly compressed. In the
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300:"ophis", for snake . The specific names is from the
582:are not enough to favor one option over the other.
516:are discernible behind metatarsals two and three.
352:platform environment. The only known specimen of
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504:slightly. The foot is composed of three tarsal
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707:"The Cenomanian: stage of hindlimbed snakes"
315:is known from a single fossil discovered at
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304:"terrasanctus" meaning "of the holy land".
781:
767:
724:
692:et al. 2000. "A fossil snake with limbs".
281:snakes known to have possessed hindlimbs.
699:
528:may be interpreted as an indication that
289:The genus was named in honor of the late
1346:
524:Several of the skeletal characters of
1266:
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606:Other known fossil snakes with legs:
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356:, housed in the collections of the
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500:are preserved though the tibia is
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256:, consisting of the sole species
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412:snakes. The contact between the
348:which deposited in a low-energy
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376:, and advanced traits found in
277:. It is one of three genera of
444:, bony thickening and growth.
358:Hebrew University of Jerusalem
1:
1369:Fossil taxa described in 2000
632:
480:similar to that possessed by
464:rib or vertebrae. Also like
447:
840:Genera with hindlimbs and a
559:analysis of skull features,
544:are markedly different. The
452:Along with its sister genus
404:bones host 73-75 teeth. The
380:. The cranium shows a small
284:
7:
647:"The Paleobiology Database
585:
10:
1385:
1251:Category:Cretaceous snakes
1246:List of Serpentes families
532:is a juvenile specimen of
460:does not possess either a
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1173:Vipera latastei ebusitana
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109:Scientific classification
107:
23:
363:
1276:Haasiophis terrasanctus
750:Journal of Paleontology
259:Haasiophis terrasanctus
238:Haasiophis terrasanctus
16:Extinct genus of snakes
327:, 20 km north of
1326:Paleobiology Database
244:Tchernov et al., 2000
1364:Transitional fossils
1354:Fossils of Palestine
476:possessing a simple
217:H. terrasanctus
1213:Psammophis odysseus
600:Paleontology portal
1105:Liasis dubudingala
1015:Boa blanchardensis
520:Taxonomic position
426:and the unrelated
341:. This site is a
1359:Cretaceous snakes
1341:
1340:
1313:Open Tree of Life
1268:Taxon identifiers
1259:
1258:
1235:Snakes portal
331:, in the central
308:Specimen material
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709:. Archived from
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29:Temporal range:
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313:H. terrasanctus
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101:Late Cretaceous
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1241:List of snakes
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546:neural spines
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506:ossifications
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1071:Rieppelophis
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1057:Phosphoroboa
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1050:Palaeopython
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986:Dinilysiidae
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892:Pachyrhachis
890:
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849:
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715:. Retrieved
711:the original
701:
693:
659:. Retrieved
655:the original
648:
641:
626:Pachyrhachis
624:
617:
610:
605:
577:evolutionary
560:
554:
550:Pachyrhachis
549:
541:
538:Pachyrhachis
537:
534:Pachyrhachis
533:
529:
525:
523:
482:Pachyrhachis
481:
473:
466:Pachyrhachis
465:
457:
454:Pachyrhachis
453:
451:
442:pachyostosis
427:
423:Pachyrhachis
421:
388:, while the
367:
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338:Pachyrhachis
336:
321:Judean hills
312:
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288:
258:
257:
252:
251:
250:
237:
232:
216:
215:
202:
201:
24:
18:
1193:Naja romani
1132:Archaeophis
1043:Messelophis
1029:Chubutophis
947:Powellophis
919:Gigantophis
912:Alamitophis
903:Madtsoiidae
510:metatarsals
486:zeugopodium
374:pipe snakes
1348:Categories
1139:Palaeophis
1096:Pythonidae
1085:Waincophis
975:Yurlunggur
885:Pachyophis
878:Haasiophis
871:Eupodophis
822:Sauropsida
717:2008-04-10
661:2009-02-06
649:Haasiophis
633:References
612:Eupodophis
561:Haasiophis
542:Haasiophis
530:Haasiophis
526:Haasiophis
478:triradiate
458:Haasiophis
448:Hind limbs
414:prefrontal
406:preorbital
382:premaxilla
317:Ein Yabrud
294:Georg Haas
279:Cenomanian
275:hind limbs
253:Haasiophis
203:Haasiophis
172:Suborder:
25:Haasiophis
1157:Viperidae
1078:Titanoboa
995:Dinilysia
803:Kingdom:
557:cladistic
555:Based on
514:phalanges
508:and five
434:vertebrae
429:Dinilysia
410:pythonine
398:pterygoid
346:carbonate
343:limestone
333:West Bank
329:Jerusalem
285:Etymology
210:Species:
176:Serpentes
132:Kingdom:
126:Eukaryota
1291:Q3782313
1285:Wikidata
1184:Elapidae
940:Nidophis
933:Menarana
926:Madtsoia
830:Squamata
814:Chordata
811:Phylum:
806:Animalia
586:See also
580:taxonomy
394:palatine
325:Ramallah
262:, is an
182:Family:
166:Squamata
156:Reptilia
146:Chordata
142:Phylum:
136:Animalia
122:Domain:
1318:4125764
1305:4968205
1166:Laophis
1064:Rageryx
968:Wonambi
954:Sanajeh
827:Order:
819:Class:
789:Fossil
694:Science
438:cloacal
418:frontal
390:maxilla
323:, near
319:in the
264:extinct
235:†
214:†
195:Genus:
162:Order:
152:Class:
1022:Boavus
1006:Boidae
961:Vasuki
851:Najash
842:sacrum
792:snakes
651:entry"
619:Najash
498:fibula
496:, and
490:tarsus
470:pelvic
462:sacral
402:dentar
400:, and
384:which
350:marine
271:snakes
1331:65326
568:clade
494:tibia
364:Skull
302:Latin
298:Greek
273:with
267:genus
1300:GBIF
540:and
416:and
40:Preκ
548:of
269:of
1350::
1328::
1315::
1302::
1287::
726:^
670:^
492:,
488:,
456:,
396:,
392:,
90:Pg
34:Ma
782:e
775:t
768:v
720:.
664:.
199:β
186:β
95:N
85:K
80:J
75:T
70:P
65:C
60:D
55:S
50:O
45:κ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
