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Eudicots

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are any modification of the wall of the pollen grain. These modifications include thinning, ridges and pores, they serve as an exit for the pollen contents and allow shrinking and swelling of the grain caused by changes in moisture content. The elongated apertures/ furrows in the pollen grain are
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evidence confirmed the genetic basis for the evolutionary relationships among flowering plants with tricolpate pollen grains and dicotyledonous traits. The term means "true dicotyledons", as it contains the majority of plants that have been considered dicots and have
614:, the monocots and the paleodicots) produce monosulcate pollen, with a single pore set in a differently oriented groove called the sulcus. The name "tricolpates" is preferred by some botanists to avoid confusion with the dicots, a nonmonophyletic group. 606:. Members of the group have tricolpate pollen, or forms derived from it. These pollens have three or more pores set in furrows called colpi. In contrast, most of the other 2173: 3053: 2997: 2205: 2253: 1837: 542:. These plants have a distinct trait in their pollen grains of exhibiting three colpi or grooves paralleling the polar axis. 717:
group. The core eudicots are a monophyletic group. A 2010 study suggested the core eudicots can be divided into two clades,
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Moore, Michael J.; Soltis, Pamela S.; Bell, Charles D.; Burleigh, J. Gordon & Soltis, Douglas E. (2010).
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or paleodicots, but these terms have not been widely or consistently adopted, as they do not refer to a
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Scores of familiar plants are eudicots, including many commonly cultivated and edible plants, numerous
502:). Most leafy, mid-latitude trees are also classified as eudicots, with notable exceptions being the 143: 2243: 1937:"Phylogenetic analysis of 83 plastid genes further resolves the early diversification of eudicots" 1702: 507: 17: 2937: 1997: 3015: 2984: 2156: 1948: 8: 2084:. Systematics Association Publications. Vol. 44. Clarendon Press. pp. 169–195. 410: 402: 1952: 2975: 2904: 2238: 1971: 1936: 1821: 1804: 1766: 1741: 1683: 1675: 1625: 424:, tropicals and ornamentals. Among the most well-known eudicot genera are those of the 194: 138: 2183: 1845: 701:), which, along with pores, are a chief criterion for identifying the pollen classes. 3038: 3010: 2898: 2713: 2122: 2085: 1976: 1915: 1899: 1771: 1722: 1456: 1027: 759: 689: 583: 1687: 2910: 2495: 2324: 2294: 2112: 2052: 1966: 1956: 1907: 1903: 1879: 1816: 1761: 1753: 1714: 1667: 682: 555: 551: 41: 1718: 1662:. Structural Botany in Systematics: A Symposium in Memory of William C. Dickison. 2798: 2656: 2306: 2269: 2215: 2177: 1406: 1270: 801: 670: 364: 282: 233: 1868:"Phylogeny of basal eudicots: insights from non-coding and rapidly evolving DNA" 2821: 2778: 2723: 2395: 2319: 1538: 1468: 1049: 763: 207: 1884: 1867: 3032: 2960: 2700: 2661: 2593: 2511: 2371: 2219: 1590: 1514: 1449: 1378: 992: 748: 626: 522: 445: 298: 254: 73: 2137: 1961: 1866:
Worberg, A; Quandt, D; Barniske, A-M; Löhne, C; Hilu, KW; Borsch, T (2007).
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of the dicots. One of the genetic traits which defines the eudicots is the
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by past authors. The current botanical terms were introduced in 1991, by
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The eudicots can be divided into two groups: the basal eudicots and the
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pollen in a cladistic context". In Blackmore, S.; Barnes, S.H. (eds.).
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This division of the eudicots is shown in the following cladogram:
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Doyle, J.A.; Hotton, C.L. (1991). "Diversification of early
1653:"Morphology and Angiosperm Systematics in the Molecular Era" 1672:
10.1663/0006-8101(2002)068[0545:maasit]2.0.co;2
421: 367:(angiosperms) which are mainly characterized by having two 2038: 1865: 1742:"Evolution of DELLA function and signaling in land plants" 1898: 1243:
The following is a more detailed breakdown according to
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The Pentapetalae can be then divided into three clades:
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Carol L. Hotton, to emphasize the later evolutionary
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The close relationships among flowering plants with
578:being the other. The remaining angiosperms include 574:(constituting over 70% of the angiosperm species), 1928: 1701:Furness, Carol A.; Rudall, Paula J. (March 2004). 3030: 1941:Proceedings of the National Academy of Sciences 538:was initially seen in morphological studies of 2099:Judd, Walter S.; Olmstead, Richard G. (2004). 2082:Pollen and Spores. Patterns of Diversification 727:, comprising all the remaining core eudicots. 2199: 1700: 570:to refer to one of the two largest clades of 2098: 2013: 1995: 1791: 1739: 617:The name "eudicots" (plural) is used in the 2075: 1644: 393:= seed leaf). Previously, they were called 2206: 2192: 1859: 713:. Basal eudicot is an informal name for a 114: 3054:Extant Early Cretaceous first appearances 2116: 2056: 1970: 1960: 1892: 1883: 1820: 1765: 2045:Botanical Journal of the Linnean Society 1787: 1785: 1247:, showing within each clade and orders: 417:from earlier, less specialized, dicots. 331: 1650: 14: 3031: 1914:. Sunderland, MA: Sinauer Associates. 1912:Phylogeny and Evolution of Angiosperms 1802: 582:and what are sometimes referred to as 2936: 2935: 2187: 2032: 1835: 1782: 598:The earlier name for the eudicots is 2039:Angiosperm Phylogeny Group (2016). 1872:Organisms Diversity & Evolution 1842:University of Arizona - Geosciences 751:includes the Vitales in the rosids) 24: 2136:in Stevens, P. F. (2001 onwards). 1822:10.1111/j.1469-8137.1972.tb04826.x 1740:Phokas, A.; Coates, J. C. (2021). 25: 3065: 2145: 2916: 2915: 2884: 142: 46: 2068: 1987: 704: 681:), and other groups having one 1829: 1796: 1733: 1694: 1618: 529: 526:, which is not an angiosperm. 13: 1: 1838:"Pollen Aperture Definitions" 1719:10.1016/j.tplants.2004.01.001 1612: 673:, with eudicots having three 2138:Angiosperm Phylogeny Website 2020:Angiosperm Phylogeny Website 2002:Angiosperm Phylogeny Website 754:superasterids consisting of 560:DELLA protein-encoding genes 7: 2014:Stevens, P.F. (2001–2016). 1996:Stevens, P.F. (2001–2014). 1803:Sporne, Kenneth R. (1972). 1746:Evolution & Development 739:superrosids consisting of 593: 564:most recent common ancestor 10: 3070: 2167:, Tree of Life Web Project 2105:American Journal of Botany 1651:Endress, Peter K. (2002). 658:structure). The number of 2944: 2893: 2882: 2807: 2759: 2741: 2732: 2709: 2698: 2637: 2584: 2571: 2562: 2549: 2538: 2520: 2509: 2481: 2470: 2429: 2379: 2370: 2337: 2333: 2315: 2304: 2278: 2267: 2227: 1885:10.1016/j.ode.2006.08.001 1135: 1119: 1112: 1094: 1087: 1072: 1065: 1047: 1040: 1025: 1018: 1003: 996: 950: 934: 927: 909: 902: 884: 877: 867: 852: 845: 827: 820: 790: 782: 540:shared derived characters 261: 252: 139:Scientific classification 137: 122: 113: 34: 27:Clade of flowering plants 1792:Judd & Olmstead 2004 536:tricolpate pollen grains 375:. The term derives from 3049:Early Cretaceous plants 1962:10.1073/pnas.0907801107 1707:Trends in Plant Science 341:pollen has three colpi. 2140:. Version 7, May 2006. 2118:10.3732/ajb.91.10.1627 639:is a synonym for the " 516:)—which belong to the 342: 3044:Plant unranked clades 1906:; Endress, Peter K.; 792:(paraphyletic group: 403:evolutionary botanist 335: 2157:Encyclopedia of Life 1836:Davis, Owen (1999). 1632:. Basic Biology 2020 788:basal eudicots 399:non-magnoliid dicots 127:during germination ( 1953:2010PNAS..107.4623M 818:core eudicots 721:and a clade called 669:helps classify the 508:American tulip tree 405:James A. Doyle and 195:Polysporangiophytes 2176:2015-02-24 at the 1900:Soltis, Douglas E. 371:(cotyledons) upon 343: 3026: 3025: 3011:Open Tree of Life 2938:Taxon identifiers 2929: 2928: 2899:Basal angiosperms 2880: 2879: 2876: 2875: 2872: 2871: 2868: 2867: 2864: 2863: 2860: 2859: 2714:Berberidopsidales 2694: 2693: 2690: 2689: 2466: 2465: 2462: 2461: 2058:10.1111/boj.12385 1904:Soltis, Pamela S. 1758:10.1111/ede.12365 1457:Berberidopsidales 1239: 1238: 1230: 1229: 1221: 1220: 1212: 1211: 1203: 1202: 1194: 1193: 1185: 1184: 1176: 1175: 1167: 1166: 1158: 1157: 1149: 1148: 1028:Berberidopsidales 982: 981: 973: 972: 964: 963: 760:Berberidopsidales 647:group, the "true 584:basal angiosperms 381:(etymologically, 330: 329: 16:(Redirected from 3061: 3019: 3018: 3006: 3005: 2993: 2992: 2980: 2979: 2978: 2965: 2964: 2963: 2933: 2932: 2919: 2918: 2888: 2739: 2738: 2707: 2706: 2569: 2568: 2547: 2546: 2518: 2517: 2479: 2478: 2377: 2376: 2335: 2334: 2313: 2312: 2295:Austrobaileyales 2276: 2275: 2254:Acrogymnospermae 2216:flowering plants 2208: 2201: 2194: 2185: 2184: 2180:Plant Life Forms 2130: 2120: 2095: 2063: 2062: 2060: 2036: 2030: 2029: 2027: 2026: 2011: 2009: 2008: 1991: 1985: 1984: 1974: 1964: 1932: 1926: 1925: 1896: 1890: 1889: 1887: 1863: 1857: 1856: 1854: 1853: 1844:. Archived from 1833: 1827: 1826: 1824: 1800: 1794: 1789: 1780: 1779: 1769: 1737: 1731: 1730: 1698: 1692: 1691: 1660:Botanical Review 1657: 1648: 1642: 1641: 1639: 1637: 1622: 1115: 1114: 1090: 1089: 1068: 1067: 1043: 1042: 1021: 1020: 999: 998: 930: 929: 905: 904: 880: 879: 870: 869: 848: 847: 823: 822: 785: 784: 777: 776: 671:flowering plants 365:flowering plants 264:Basal eudicots: 147: 146: 118: 108: 45: 42:Early Cretaceous 38:Temporal range: 32: 31: 21: 3069: 3068: 3064: 3063: 3062: 3060: 3059: 3058: 3029: 3028: 3027: 3022: 3014: 3009: 3001: 2996: 2988: 2983: 2974: 2973: 2968: 2959: 2958: 2953: 2940: 2930: 2925: 2889: 2856: 2803: 2755: 2728: 2703: 2686: 2657:Crossosomatales 2633: 2580: 2558: 2543: 2534: 2514: 2505: 2496:Trochodendrales 2475: 2458: 2425: 2366: 2329: 2325:Ceratophyllales 2309: 2300: 2272: 2263: 2244:Marchantiophyta 2223: 2212: 2178:Wayback Machine 2148: 2143: 2111:(10): 1627–44. 2092: 2071: 2066: 2037: 2033: 2024: 2022: 2012: 2006: 2004: 1994: 1992: 1988: 1933: 1929: 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2497: 2494: 2492: 2489: 2487: 2484: 2483: 2480: 2477: 2474: 2469: 2455: 2452: 2450: 2447: 2445: 2442: 2440: 2437: 2436: 2434: 2432: 2428: 2421: 2418: 2415: 2412: 2409: 2406: 2403: 2400: 2397: 2396:Petrosaviales 2394: 2391: 2388: 2385: 2382: 2381: 2378: 2375: 2373: 2369: 2363: 2360: 2358: 2355: 2353: 2350: 2348: 2345: 2344: 2342: 2340: 2336: 2332: 2326: 2323: 2321: 2320:Chloranthales 2318: 2317: 2314: 2311: 2308: 2303: 2296: 2293: 2290: 2287: 2284: 2281: 2280: 2277: 2274: 2271: 2266: 2260: 2257: 2255: 2252: 2250: 2247: 2245: 2242: 2240: 2237: 2234: 2230: 2229: 2226: 2221: 2217: 2209: 2204: 2202: 2197: 2195: 2190: 2189: 2186: 2179: 2175: 2172: 2169: 2166: 2165: 2161: 2159: 2158: 2153: 2150: 2149: 2139: 2135: 2132: 2128: 2124: 2119: 2114: 2110: 2106: 2102: 2097: 2093: 2091:9780198577461 2087: 2083: 2079: 2074: 2073: 2059: 2054: 2050: 2046: 2042: 2035: 2021: 2017: 2003: 1999: 1990: 1982: 1978: 1973: 1968: 1963: 1958: 1954: 1950: 1946: 1942: 1938: 1931: 1923: 1921:9780878938179 1917: 1913: 1909: 1905: 1901: 1895: 1886: 1881: 1877: 1873: 1869: 1862: 1848:on 2009-02-03 1847: 1843: 1839: 1832: 1823: 1818: 1814: 1810: 1806: 1799: 1793: 1788: 1786: 1777: 1773: 1768: 1763: 1759: 1755: 1751: 1747: 1743: 1736: 1728: 1724: 1720: 1716: 1712: 1708: 1704: 1697: 1689: 1685: 1681: 1677: 1673: 1669: 1665: 1661: 1654: 1647: 1631: 1630:Basic Biology 1627: 1621: 1617: 1598: 1594: 1592: 1591:Metteniusales 1588: 1586: 1582: 1580: 1576: 1574: 1570: 1568: 1564: 1562: 1558: 1556: 1552: 1551: 1550: 1549: 1544: 1540: 1536: 1534: 1530: 1528: 1524: 1522: 1518: 1516: 1515:Escalloniales 1512: 1510: 1506: 1504: 1500: 1499: 1498: 1497: 1492: 1490: 1486: 1484: 1480: 1479: 1478: 1477: 1472: 1470: 1466: 1464: 1460: 1458: 1454: 1453: 1452: 1451: 1450:Superasterids 1446: 1438: 1434: 1432: 1428: 1426: 1422: 1420: 1416: 1414: 1410: 1408: 1404: 1402: 1398: 1396: 1392: 1391: 1390: 1389: 1384: 1380: 1379:Zygophyllales 1376: 1374: 1370: 1368: 1364: 1362: 1358: 1356: 1352: 1350: 1346: 1344: 1340: 1338: 1334: 1333: 1332: 1331: 1326: 1324: 1320: 1319: 1318: 1317: 1312: 1310: 1306: 1305: 1304: 1303: 1298: 1296: 1292: 1290: 1286: 1285: 1284: 1283:Core eudicots 1280: 1278: 1274: 1272: 1268: 1266: 1262: 1260: 1256: 1255: 1254: 1250: 1249: 1248: 1246: 1235: 1234: 1226: 1225: 1217: 1216: 1208: 1207: 1199: 1198: 1190: 1189: 1181: 1180: 1172: 1171: 1163: 1162: 1154: 1153: 1145: 1144: 1141: 1139: 1133: 1132: 1129: 1128: 1125: 1123: 1117: 1116: 1110: 1107: 1106: 1103: 1102: 1099: 1098: 1092: 1091: 1085: 1084: 1081: 1080: 1077: 1076: 1070: 1069: 1063: 1060: 1059: 1056: 1055: 1052: 1051: 1045: 1044: 1038: 1037: 1034: 1033: 1030: 1029: 1023: 1022: 1016: 1015: 1012: 1011: 1008: 1007: 1001: 1000: 994: 993:superasterids 991: 990: 987: 986: 978: 977: 969: 968: 960: 959: 956: 954: 948: 947: 944: 943: 940: 938: 932: 931: 925: 922: 921: 918: 917: 914: 913: 907: 906: 900: 897: 896: 893: 892: 889: 888: 882: 881: 875: 872: 871: 865: 864: 861: 860: 857: 856: 850: 849: 843: 840: 839: 836: 835: 832: 831: 825: 824: 817: 816: 813: 812: 809: 807: 803: 799: 795: 787: 786: 779: 778: 774: 769: 765: 761: 757: 753: 750: 749:APG IV system 746: 742: 738: 736: 733: 732: 731: 728: 726: 725: 720: 716: 712: 711:core eudicots 702: 700: 696: 691: 686: 684: 680: 676: 672: 668: 664: 661: 657: 654: 650: 646: 642: 638: 634: 632: 628: 627:APG IV system 624: 620: 615: 613: 610:(that is the 609: 605: 601: 591: 589: 585: 581: 577: 573: 569: 565: 561: 557: 553: 548: 543: 541: 537: 527: 525: 524: 523:Ginkgo biloba 519: 515: 514: 509: 505: 501: 497: 493: 492: 487: 483: 482: 477: 473: 472: 467: 463: 462: 457: 453: 452: 447: 446:forget-me-not 443: 442: 437: 433: 432: 427: 423: 418: 416: 412: 408: 407:paleobotanist 404: 400: 396: 392: 388: 384: 380: 379: 374: 370: 366: 362: 358: 357: 352: 348: 340: 339: 334: 320: 317: 315: 312: 311: 309: 305: 302: 300: 299:Superasterids 297: 296: 294: 293: 291: 290:Core eudicots 288: 284: 281: 279: 276: 274: 271: 269: 266: 265: 263: 262: 260: 256: 251: 248: 245: 242: 239: 238: 235: 232: 229: 226: 225: 222: 219: 216: 213: 212: 209: 208:Tracheophytes 206: 203: 200: 199: 196: 193: 190: 187: 186: 183: 180: 177: 174: 173: 170: 167: 164: 161: 160: 157: 154: 151: 150: 145: 140: 136: 132: 131: 126: 121: 117: 112: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 43: 33: 30: 19: 2945: 2769:Aquifoliales 2662:Picramniales 2624:Cucurbitales 2604:Malpighiales 2554:Saxifragales 2486:Ranunculales 2472: 2454:Zingiberales 2449:Commelinales 2402:Dioscoreales 2283:Amborellales 2259:Angiospermae 2162: 2155: 2108: 2104: 2081: 2069:Bibliography 2048: 2044: 2034: 2023:. 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347:eudicots 253:Clades ( 247:Eudicots 130:Brassica 44:- recent 35:Eudicots 2961:Q165468 2809:Lamiids 2789:Apiales 2639:Malvids 2629:Fagales 2619:Rosales 2614:Fabales 2576:Vitales 2501:Buxales 2154:at the 1998:"Trees" 1972:2842043 1949:Bibcode 1767:9285615 1680:4354438 1548:Lamiids 1527:Apiales 1388:Malvids 1349:Fagales 1343:Rosales 1337:Fabales 1323:Vitales 1277:Buxales 1138:lamiids 953:malvids 912:Vitales 806:Buxales 693:called 688:Pollen 663:furrows 641:Eudicot 590:group. 456:cabbage 389:= two; 310:Orders 295:Clades 268:Buxales 156:Plantae 18:Eudicot 3016:431495 2586:Fabids 2564:Rosids 2444:Poales 2220:APG IV 2171:Dicots 2125:  2088:  1979:  1969:  1918:  1774:  1764:  1725:  1686:  1678:  1636:6 July 1595:order 1589:order 1583:order 1577:order 1571:order 1565:order 1559:order 1553:order 1545:clade 1537:order 1531:order 1525:order 1519:order 1513:order 1507:order 1501:order 1493:clade 1487:order 1481:order 1473:clade 1467:order 1461:order 1455:order 1447:clade 1435:order 1429:order 1423:order 1417:order 1411:order 1405:order 1399:order 1393:order 1385:clade 1377:order 1371:order 1365:order 1359:order 1353:order 1347:order 1341:order 1335:order 1330:Fabids 1327:clade 1321:order 1316:Rosids 1313:clade 1307:order 1299:clade 1293:order 1287:order 1281:clade 1275:order 1269:order 1263:order 1257:order 1251:clade 1245:APG IV 937:fabids 899:rosids 745:rosids 699:colpus 683:sulcus 656:pollen 604:pollen 568:botany 545:Later 494:) and 415:dicots 359:are a 338:Arabis 255:APG IV 3003:71240 2233:Plant 1684:S2CID 1676:JSTOR 1656:(PDF) 747:(the 695:colpi 675:colpi 667:pores 631:clade 520:—and 486:maple 471:Malus 466:apple 422:trees 361:clade 353:, or 241:Clade 228:Clade 215:Clade 202:Clade 189:Clade 176:Clade 163:Clade 133:sp.) 2998:NCBI 2123:PMID 2086:ISBN 1977:PMID 1916:ISBN 1772:PMID 1723:PMID 1638:2020 766:and 743:and 506:and 491:Acer 345:The 123:Two 49:PreꞒ 2990:283 2985:EoL 2113:doi 2053:doi 2049:181 1967:PMC 1957:doi 1945:107 1880:doi 1817:doi 1762:PMC 1754:doi 1715:doi 1668:doi 665:or 619:APG 558:of 484:), 474:), 464:), 454:), 444:), 434:), 397:or 363:of 3035:: 3013:: 3000:: 2987:: 2972:: 2957:: 2121:. 2109:91 2107:. 2103:. 2047:. 2043:. 2018:. 2000:. 1975:. 1965:. 1955:. 1943:. 1939:. 1902:; 1874:. 1870:. 1840:. 1813:71 1811:. 1807:. 1784:^ 1770:. 1760:. 1750:23 1748:. 1744:. 1721:. 1709:. 1705:. 1682:. 1674:. 1664:68 1658:. 1628:. 808:) 804:, 800:, 796:, 762:, 758:, 685:. 643:" 387:di 383:eu 349:, 292:: 257:) 243:: 230:: 217:: 204:: 191:: 178:: 165:: 99:Pg 2909:³ 2903:² 2897:¹ 2422:³ 2416:³ 2410:³ 2404:³ 2398:³ 2392:² 2386:² 2297:¹ 2291:¹ 2285:¹ 2235:: 2222:) 2218:( 2207:e 2200:t 2193:v 2129:. 2115:: 2094:. 2061:. 2055:: 2028:. 2010:. 1983:. 1959:: 1951:: 1924:. 1888:. 1882:: 1876:7 1855:. 1825:. 1819:: 1778:. 1756:: 1729:. 1717:: 1711:9 1690:. 1670:: 1640:. 677:( 510:( 498:( 488:( 478:( 468:( 458:( 448:( 438:( 428:( 104:N 94:K 89:J 84:T 79:P 74:C 69:D 64:S 59:O 54:Ꞓ 20:)

Index

Eudicot
Early Cretaceous
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

seed leaves
Brassica
Scientific classification
Edit this classification
Plantae
Streptophyta
Embryophytes
Polysporangiophytes
Tracheophytes
Spermatophytes
Angiosperms
Eudicots
APG IV
Buxales
Proteales

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