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Nevertheless, presumably, these males are able to give offspring, which leads to the appearance of tetraploid hybrids. Sometimes males emerge from unfertilized eggs laid by parthenogenetic females. Like males of hybrid origin, they have reproductive disorders, which may not interfere with their offspring. A small amount of parthenogenetic female males emerging from clutches is explained by frequently occurring mutations incompatible with life.
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sides of the body. In some species, blue or violet spots with white circles in the center and / or monotonous blue-violet spots at the junction of the abdominal scutes with trunk scales are located on the anterior third of the body. Most types of rock lizards are characterized by a diverse color of the abdominal side of the body, ranging from various shades of pink, red and orange to yellow and green.
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hours in the middle of the day . After heating (basking), the body temperature of the lizard reaches about 30–34 Â°C (86–93 Â°F), and it begins a routine activity aimed at supporting the body. Late in the evening, when the heat subsides, the animals return to the basking places and stay there for some time, after which they go to their night shelters.
272:. For several decades, scientists from different countries studied the monophyletic group of rock lizards independently, highlighting new subspecies, simplifying some taxa and describing new species. It was on this group of lizards that the phenomenon of parthenogenesis in amniotic vertebrates was first discovered by the Russian zoologist 1335:
until the end of May, exit from wintering shelters occurs, and the active period is from 6–7 months (in the mountains) and up to 9–10 months (in the valleys and on the seashore). During this period, mature individuals mate, and females lay eggs. Hibernation takes place from late September to mid November.
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3) Rock lizards of dry and moderately dry landscapes (alpine steppes) of rocks and their feet on slopes with dry-loving shrubbery and grassy vegetation, road slopes. Such habitats have a large number of crevices and voids serving as shelters for lizards. Such species are found in such landscapes as:
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includes four groups (clades), combining species by origin and kinship: “raddei”, “rudis”, “saxicola” and “caucasica”. Subsequently, three more clades were identified: “praticola”, “chlorogaster”, and “defilippii”. In total, the genus includes 35 species, 7 of which breed parthenogenetically, and 22
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Rock lizards are common in Abkhazia, Azerbaijan, Armenia, Georgia, Iran, Nagorno-Karabakh, Russia (Adygea, Dagestan, Ingushetia, Kabardino-Balkaria, Karachay-Cherkessia, Krasnodar Territory, Republic of Crimea, North Ossetia-Alania, Stavropol Territory and Chechnya), in Turkey and South Ossetia. It
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The appearance of parthenogenetic species is usually associated with the last, Wurm glaciation. Parthenogenetic rock lizards appeared about 10 thousand years ago when, due to the formation of mountain glaciers, the habitats of the parent bisexual species were disturbed, which led to the overlap of
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always enter as maternal species. From the point of view of the classical concept of a species, homosexual taxa cannot be assigned a species status due to the lack of exchange of genetic material between individuals within the same population. However, on the basis of morphological and cytogenetic
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The beginning and end of the daily activity of the lizard is determined by the lighting conditions within the individual area, and in some individuals it can start early in the morning, while individuals living on the slopes of the northern exposure or in deep forest valleys are active for several
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Rock lizards feed on various invertebrates with a body size from a few millimeters to 4 cm (1.6 in): spiders, diptera, lepidoptera, hymenoptera, cockroaches, orthoptera, semi-rigid-winged, coleoptera, wood lice, worms, slugs, marine and freshwater amphipods, caddis flies, chironomids and
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The color of rock lizards varies from different tones of green to sand. Females are usually colored paler than males. On the dorsal side of the body, rock lizards have an occipital stripe composed of a set of black or brown spots and a wide line of the lizard's main color, and dark patterns on the
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Despite the fact that as a result of parthenogenesis, individuals are born that receive hereditary material only from the mother's body, a small intraspecific genetic diversity has been revealed due to mutational processes, genetic instability and the appearance of same-sex species as a result of
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The seasonal activity of rock lizards is determined by temperature indicators. Therefore, species living at different heights differ in terms of exit from wintering shelters, mating period, laying of eggs, hatching of young individuals and time of leaving for wintering. Around the end of February
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Hybridization occurs in places where the ranges of parthenogenetic and bisexual species overlap, resulting in the appearance of triploid sterile females and a small number of males, with changes in the reproductive system (malfunctions in the formation of reproductive products, hermaphroditism).
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Despite the diversity of the food supply, rock lizards can develop preferences for feeding on invertebrates of a certain group (for example, flying forms of ants), which is caused by seasonal changes in the availability or abundance of this type of prey. Even after a significant reduction in the
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Rock lizards are found in various high-altitude zones from 0 to 3,000 m (0 to 9,843 ft) above sea level and occupy a variety of landscapes: mountain-steppe, forest-steppe, mountain meadow, mountain forest, anthropogenic and coastal. By confinement to one or another habitat, they can be
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species are small lizards with a head and body length of 50–85 mm (2.0–3.3 in), and with a tail length about two times longer. The body is usually flattened, the head is pointed in shape and in most species flattened in a vertical plane, which allows lizards to hide in narrow crevices
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Rock lizards are extremely rare on their own, usually forming settlements. The population density of parthenogenetic rock lizards can vary in a wider range than that of bisexual species: up to 200 individuals per 1 km of the route in unisexual species and up to 80 individuals in bisexual
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The basis of the settlements of bisexual rock lizards are sedentary males and females with individual sites, often overlapping. In a number of species, some males possess territories protected from other males. The territories of males never overlap, but their centers of activity, primarily
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their ranges and the hybridogenic formation of parthenogenetic individuals better adapted to the conditions of short summers and long winters. Due to the doubled reproduction rate and successful resettlement, same-sex populations later began to exist independently of the parent species.
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Rock lizards are characterized by complex and diverse social systems, which, in particular, are characterized by stable long-term friendly relations between the male and the female and territorial or hierarchical relations between individuals of the same sex.
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on the southern slope of the Greater Caucasus Mountain Range (South Ossetia) is distributed at altitudes of 1,500–1,800 m (4,900–5,900 ft) above sea level, and on the northern slopes (Dagestan) at 50–2,100 m (160–6,890 ft) above sea level.
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Rock lizards reach maturity in the second (females) and third (males) year of life with a total life expectancy of up to 13 years. Mating of some species of rock lizards occurs after the first molt, approximately 3–5 weeks after leaving wintering
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Rock lizards are found at heights of 0 – 3000 m above sea level. Zonal and geographical distribution is determined by the amount of precipitation, average annual temperature, duration of the adverse season, and exposure of the slope. For example,
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between stones and rocks. Rock lizards have relatively long legs with special calluses on the inner surfaces of the paws and sharp claws, thanks to which they quickly move along the vertical rough surfaces of rocks and stones.
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2) Inhabitants in areas of bedrock outcrops and clayey cliffs in alpine and subalpine meadows. As shelters, they often use rodent burrows, cavities between stones and cracks in the rocks. These habitats are adhered to by
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is worth noting that the boundaries of the range of some species are not reliably known, but the expected areas of their encounters coincide with the already indicated distribution sites for the whole genus
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The study of the social behavior and spatial structure of rock lizard populations has been the subject of a number of scientific papers published based on the results of many years of research
169:. Member species are native to the Caucasus, Iran and Turkey, living in forest and grassy habitats with numerous rock outcrops. Among rock lizards, seven parthenogenetic species are known. 1280:
4) Occupying anthropogenic habitats: an abandoned building, walls in cities, abandoned temples, monasteries, etc., where their number often exceeds that in natural habitats. For example,
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penetrated the territory of the Western Caucasus in the Middle Miocene or in the Middle Pliocene, when there was a land connection between Asia Minor and the Balkans.
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and independent of them, able to live in habitats in which there are no rocks, using rodent burrows as shelters, deciduous litter, cavities in trees and bark (e.g.
276:(1924–2009), who also made a significant contribution to understanding the ecology, systematics, and morphology of rock lizards. In 1997, the Spanish scientist 32: 1376:
from the subfamily Lacertinae occurred approximately 12-16 million years ago, in the middle of the Miocene. Presumably, the ancestral form of the genus
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density of invertebrates of this group, lizards continue to hunt for them for some time in the presence of more accessible food objects.
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species, which is explained by the fact that parthenogenetic species are less aggressive and have a high population growth rate.
1755: 1513: 1167: 846: 1468: 1464: 223:(1794–1860) made an expedition to the North Caucasus. As a result of which he described two new species: a meadow lizard, 1094: 1448: 773: 723: 1245:
1) Lizards living in the forest, according to the occupied microreliefs, are divided into: adhering to rock outcrops (
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sometimes parts of plants. Also, particular cases of cannibalism were recorded when adults ate juvenile individuals.
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associated with basking, coincide with the centers of activity of females that live within their territories.
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size is usually 2 to 7 eggs. The incubation period lasts approximately 55–65 days. Young animals with a
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D. armeniaca, D. bendimahiensis, D. dahli, D. rostombekovi, D. sapphirina, D. unisexualis
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in parentheses indicates that the species was originally described in a genus other than
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The Reptiles of Northern Eurasia: Taxonomic Diversity, Distribution, Conservation Status
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Ahmadzadeh, Flecks, Carretero, Mozaffari, Bohme, Harris, Freitas & Rodder, 2013
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Ahmadzadeh, Flecks, Carretero, Mozaffari, Bohme, Harris, Freitas & Rodder, 2013
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Ahmadzadeh, Flecks, Carretero, Mozaffari, Bohme, Harris, Freitas & Rodder, 2013
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Arribas, Candan, Kornilios, Ayaz, Kumlutas, Gul, Yilmaz, Caynak, & Ilgaz, 2022
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Ahmadzadeh, Flecks, Carretero, Mozaffari, Bohme, Harris, Freitas, Rodder, 2013
1811: 1609: 1477:, Faunastica, vol. 47, Sofia, Bulgaria: Pensoft Publishers, p. 79, 1460: 1426: 1357: 1356:. Egg laying begins in the second half of June and lasts until early August. 1309: 702: 273: 208: 260:(1858–1937) there was a lengthy discussion about the taxonomic position of 1724: 1618: 205: 1416:
repeated and independent crossing of parental species among themselves.
1716: 166: 126: 116: 1729: 1212: 66: 1580: 252:. But at the beginning of the XX century between the two zoologists 1603: 1536: 202: 106: 86: 1566:) and their relationships among the Eurasian lacertid radiation". 1364:(SVL) of about 25 mm (0.98 in) are born in late summer. 1275:
D. rudis, D. portschinski, D. daghestanica, D. raddei, D. saxicola
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The evolution and origin of parthenogenetic species of Darevskia
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Arribas, Candan, Jurnaz, KumlutaĹź, Caynak, & Ilgaz, 2022
194: 159: 1508:. Baltimore: Johns Hopkins University Press. xiii + 296 pp. 1352:, in others, immediately after leaving wintering shelter ( 1310:
Population density and spatial structure of the population
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The genus of rock lizards has 7 parthenogenetic species:
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Arribas, Ilgaz, Kumluras, Durmus, Avci, & Ăśzum, 2013
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and subspecies were further considered separately from
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The following species are recognized as being valid.
264:, which was resolved in favor of the former, and 1809: 1404:characters, they are assigned a species rank. 1146:) - Caucasian rock lizard, Valentin's lizard 1441: 1242:conditionally divided into several groups: 214: 1207:– Caucasian rock lizard, Valentin's lizard 681:(Schmidtler, Eiselt & Darevsky, 1994) 583:(Schmidtler, Eiselt & Darevsky, 1994) 219:In 1830, a professor at Kazan University, 31: 1111:(Schmidtler, Eiselt & Darevsky, 1994) 241:, considering it as part of the European 1144:Eiselt, Darevsky, & Schmidtler, 1992 296:According to Arribas (1997), the genus 1810: 1399:always enter as paternal species, and 1585: 1584: 761:(Tuniyev, Petrova & Lotiev, 2023) 1795:DBEC2AE0-D337-4660-8FAF-072A42C604D2 1704:89c08cb0-c039-4b1f-b67b-5ce5169719e3 1282:D. armeniaca, D. lindholmi, D. dahli 284:and designated the type species as 13: 1546: 1523: 431:(Darevsky & Vedmederja, 1977) 14: 1839: 1506:The Eponym Dictionary of Reptiles 1401:D. mixta, D. nairensis, D. raddei 665:(Darevsky & Danielyan, 1977) 280:named the genus of rock lizards 53: 1342: 1227: 487:(Darevsky & Tuniyev, 1997) 1828:Taxa named by Oscar J. Arribas 1568:Russian Journal of Herpetology 1491: 908:(Darevsky & Tuniyev, 1997) 776:, Eiselt & Darevsky, 1994) 495:(Darevsky & Eiselt, 1980) 172: 1: 1434: 1251:D. chlorogaster, D. armeniaca 291: 1296: 188: 7: 1504:; Grayson, Michael (2011). 1419: 1329: 10: 1844: 1236: 915:Darevskia josefschmidtleri 687: 447:(Lantz & Cyren, 1936) 371:(Lantz & Cyren, 1913) 221:Eduard Friedrich Eversmann 1593: 1539:www.reptile-database.org. 1139:Darevskia spitzenbergerae 274:Ilya Sergeyevich Darevsky 209:Ilya Sergeyevich Darevsky 50:Scientific classification 48: 39: 30: 23: 1372:Separation of the genus 768:Darevskia bendimahiensis 580:Darevskia bendimahiensis 519:Parthenogenetic species 215:Taxonomy and systematics 1247:D. raddei, D. brauneri) 1132:Ahmadzadeh et al., 2013 1018:Darevskia portschinskii 992:Lantz & CyrĂ©n, 1936 944:Ahmadzadeh et al., 2013 932:Ahmadzadeh et al., 2013 798:– Brauner's rock lizard 436:Darevskia portschinskii 258:George Albert Boulenger 162:of wall lizards of the 1263:D. mixta, D. armeniaca 1062:Darevskia rostombekowi 939:Darevskia kopetdaghica 855:Darevskia daghestanica 826:Darevskia chlorogaster 763:– Arribas’ rock lizard 630:Darevskia rostombekowi 572:Darevskia kopetdaghica 536:Darevskia chlorogaster 418:Darevskia daghestanica 1177:Darevskia unisexualis 646:Darevskia unisexualis 1699:Fauna Europaea (new) 1537:The Reptile Database 1469:Barabanov, Andrei V. 1449:Ananjeva, Natalia B. 1362:snout-to-vent length 1172:– Szczerbak's lizard 1162:Darevskia szczerbaki 1107:Darevskia sapphirina 1013:– red-bellied lizard 875:Darevskia defilippii 678:Darevskia sapphirina 544:Darevskia defilippii 230:, and a rock lizard 1201:Darevskia valentini 1127:Darevskia schaekeli 1057:– Azerbaijan lizard 1032:Darevskia praticola 965:Darevskia mirabilis 951:Darevskia lindholmi 890:Darevskia derjugini 841:Darevskia clarkorum 836:– greenbelly lizard 816:Darevskia caucasica 782:Darevskia bithynica 742:Darevskia armeniaca 715:Darevskia aghasyani 600:Darevskia schaekeli 552:Darevskia armeniaca 526:Darevskia praticola 506: 452:Darevskia derjugini 444:Darevskia lindholmi 428:Darevskia clarkorum 410:Darevskia sczerbaki 402:Darevskia valentini 394:Darevskia bithynica 359:Darevskia nairensis 349:Darevskia caucasica 305: 1457:Khalikov, Roman G. 1218:binomial authority 1157:– Steiner's lizard 1151:Darevskia steineri 1117:Darevskia saxicola 792:Darevskia brauneri 757:Darevskia arribasi 698:Darevskia adjarica 622:Darevskia steineri 539:(Boulenger, 1908) 530:(Eversmann, 1834) 505: 492:Darevskia adjarica 470:Darevskia tuniyevi 376:Darevskia brauneri 344:(Eversmann, 1834) 341:Darevskia saxicola 304: 1805: 1804: 1777:Open Tree of Life 1587:Taxon identifiers 1514:978-1-4214-0135-5 1465:Ryabov, Sergei A. 1461:Darevsky, Ilya S. 1453:Orlov, Nikolai L. 1206: 1196: 1187:Darevskia uzzelli 1182: 1171: 1156: 1122: 1121:(Eversmann, 1834) 1112: 1102: 1086:Darevskia salihae 1081: 1067: 1056: 1041: 1027: 1012: 999:Darevskia parvula 987:Darevskia obscura 982: 960: 910:– Charnali lizard 909: 899: 884: 870: 860: 850: 835: 821: 803:Darevskia caspica 797: 787: 777: 762: 752:– Armenian lizard 751: 737: 727: 710: 685: 684: 662:Darevskia uzzelli 649:(Darevsky, 1966) 633:(Darevsky, 1957) 611:(Darevsky, 1957) 592:Darevskia caspica 547:(Camerano, 1877) 503: 502: 455:(Nikolsky, 1898) 421:(Darevsky, 1967) 405:(Boettger, 1892) 387:(Darevsky, 1967) 368:Darevskia parvula 336:(Bedriaga, 1886) 328:(Boettger, 1892) 235:(Eversmann, 1834) 228:(Eversmann, 1834) 225:Lacerta praticola 201:, is in honor of 151: 150: 147: 1835: 1798: 1797: 1785: 1784: 1772: 1771: 1759: 1758: 1746: 1745: 1733: 1732: 1720: 1719: 1707: 1706: 1694: 1693: 1681: 1680: 1668: 1667: 1655: 1654: 1642: 1641: 1629: 1628: 1627: 1614: 1613: 1612: 1582: 1581: 1540: 1527: 1521: 1502:Watkins, Michael 1495: 1489: 1488: 1445: 1397:D. portschinskii 1205:(Boettger, 1892) 1204: 1191:(Darevsky & 1190: 1181:(Darevsky, 1966) 1180: 1165: 1154: 1145: 1133: 1120: 1110: 1089: 1075: 1066:(Darevsky, 1957) 1065: 1050: 1047:Darevskia raddei 1035: 1021: 1002: 993: 980: 971: 954: 945: 933: 921: 907: 904:Darevskia dryada 893: 878: 869:(Darevsky, 1957) 868: 859:(Darevsky, 1967) 858: 845:(Darevsky & 844: 829: 819: 811: 795: 785: 771: 760: 745: 736:(Darevsky, 1967) 735: 732:Darevskia alpina 718: 701: 507: 504: 484:Darevskia dryada 439:(Kessler, 1878) 384:Darevskia alpina 363:Darevsky, 1967) 325:Darevskia raddei 306: 303: 278:Oscar J. Arribas 256:(1862–1953) and 251: 236: 232:Lacerta saxicola 229: 142: 58: 57: 42:Darevskia raddei 35: 21: 20: 16:Genus of lizards 1843: 1842: 1838: 1837: 1836: 1834: 1833: 1832: 1808: 1807: 1806: 1801: 1793: 1788: 1780: 1775: 1767: 1762: 1754: 1749: 1741: 1736: 1728: 1723: 1715: 1710: 1702: 1697: 1689: 1684: 1676: 1671: 1663: 1658: 1650: 1645: 1637: 1632: 1623: 1622: 1617: 1608: 1607: 1602: 1589: 1549: 1547:Further reading 1544: 1543: 1528: 1524: 1496: 1492: 1485: 1446: 1442: 1437: 1422: 1370: 1345: 1332: 1312: 1299: 1290:D. daghestanica 1239: 1230: 1143: 1131: 1072:Darevskia rudis 1042:– meadow lizard 991: 977:Darevskia mixta 969: 943: 931: 927:Darevskia kamii 919: 885:– Elburs lizard 865:Darevskia dahli 806: 690: 625:(Eiselt, 1995) 608:Darevskia dahli 564:Darevskia kamii 555:(Mehely, 1909) 465:(Mehely, 1909) 462:Darevskia mixta 413:(Lukina, 1963) 397:(Mehely, 1909) 379:(Mehely, 1909) 352:(Mehely, 1909) 333:Darevskia rudis 294: 245: 234: 227: 217: 191: 175: 141: 52: 17: 12: 11: 5: 1841: 1831: 1830: 1825: 1820: 1803: 1802: 1800: 1799: 1786: 1773: 1760: 1747: 1734: 1721: 1708: 1695: 1686:Fauna Europaea 1682: 1669: 1656: 1643: 1630: 1615: 1599: 1597: 1591: 1590: 1579: 1578: 1553:Arribas, Oscar 1548: 1545: 1542: 1541: 1522: 1490: 1483: 1439: 1438: 1436: 1433: 1432: 1431: 1421: 1418: 1369: 1366: 1344: 1341: 1331: 1328: 1311: 1308: 1298: 1295: 1238: 1235: 1229: 1226: 1209: 1208: 1197: 1183: 1173: 1158: 1155:(Eiselt, 1995) 1147: 1135: 1123: 1113: 1103: 1082: 1068: 1058: 1043: 1028: 1014: 995: 983: 981:(MĂ©helĂż, 1909) 973: 961: 947: 935: 923: 911: 900: 886: 871: 861: 851: 837: 822: 820:(MĂ©helĂż, 1909) 812: 799: 796:(MĂ©helĂż, 1909) 788: 786:(MĂ©helĂż, 1909) 778: 764: 753: 738: 728: 711: 689: 686: 683: 682: 674: 672: 670: 667: 666: 658: 656: 654: 651: 650: 642: 640: 638: 635: 634: 626: 618: 616: 613: 612: 604: 596: 588: 585: 584: 576: 568: 560: 557: 556: 548: 540: 532: 521: 520: 517: 514: 511: 501: 500: 498: 496: 488: 479: 478: 476: 474: 466: 457: 456: 448: 440: 432: 423: 422: 414: 406: 398: 389: 388: 380: 372: 364: 354: 353: 345: 337: 329: 320: 319: 316: 313: 310: 293: 290: 216: 213: 190: 187: 174: 171: 149: 148: 134: 130: 129: 124: 120: 119: 114: 110: 109: 104: 100: 99: 94: 90: 89: 84: 80: 79: 74: 70: 69: 64: 60: 59: 46: 45: 37: 36: 28: 27: 15: 9: 6: 4: 3: 2: 1840: 1829: 1826: 1824: 1823:Lizard genera 1821: 1819: 1816: 1815: 1813: 1796: 1791: 1787: 1783: 1778: 1774: 1770: 1765: 1761: 1757: 1752: 1748: 1744: 1739: 1735: 1731: 1726: 1722: 1718: 1713: 1709: 1705: 1700: 1696: 1692: 1687: 1683: 1679: 1674: 1670: 1666: 1661: 1657: 1653: 1648: 1644: 1640: 1635: 1631: 1626: 1620: 1616: 1611: 1605: 1601: 1600: 1598: 1596: 1592: 1588: 1583: 1577:, new genus). 1576: 1572: 1569: 1565: 1561: 1558: 1557:Archeolacerta 1554: 1551: 1550: 1538: 1534: 1533: 1526: 1519: 1515: 1511: 1507: 1503: 1499: 1494: 1486: 1484:954-642-269-X 1480: 1476: 1475: 1470: 1466: 1462: 1458: 1454: 1450: 1444: 1440: 1429: 1428: 1427:Cnemidophorus 1424: 1423: 1417: 1413: 1409: 1405: 1402: 1398: 1394: 1390: 1386: 1381: 1379: 1375: 1365: 1363: 1359: 1355: 1354:D. valentini) 1351: 1340: 1336: 1327: 1324: 1320: 1316: 1307: 1303: 1294: 1291: 1285: 1283: 1278: 1276: 1270: 1268: 1264: 1260: 1254: 1252: 1248: 1243: 1234: 1225: 1223: 1219: 1215: 1214: 1203: 1202: 1198: 1194: 1189: 1188: 1184: 1179: 1178: 1174: 1169: 1164: 1163: 1159: 1153: 1152: 1148: 1141: 1140: 1136: 1129: 1128: 1124: 1119: 1118: 1114: 1109: 1108: 1104: 1100: 1096: 1092: 1088: 1087: 1083: 1079: 1074: 1073: 1069: 1064: 1063: 1059: 1054: 1049: 1048: 1044: 1039: 1034: 1033: 1029: 1025: 1020: 1019: 1015: 1010: 1006: 1001: 1000: 996: 989: 988: 984: 979: 978: 974: 967: 966: 962: 958: 953: 952: 948: 941: 940: 936: 929: 928: 924: 917: 916: 912: 906: 905: 901: 897: 892: 891: 887: 882: 877: 876: 872: 867: 866: 862: 857: 856: 852: 848: 843: 842: 838: 833: 828: 827: 823: 818: 817: 813: 809: 805: 804: 800: 794: 793: 789: 784: 783: 779: 775: 770: 769: 765: 759: 758: 754: 749: 744: 743: 739: 734: 733: 729: 725: 721: 717: 716: 712: 708: 704: 700: 699: 695: 694: 693: 680: 679: 675: 673: 671: 669: 668: 664: 663: 659: 657: 655: 653: 652: 648: 647: 643: 641: 639: 637: 636: 632: 631: 627: 624: 623: 619: 617: 615: 614: 610: 609: 605: 602: 601: 597: 594: 593: 589: 587: 586: 582: 581: 577: 574: 573: 569: 566: 565: 561: 559: 558: 554: 553: 549: 546: 545: 541: 538: 537: 533: 531: 528: 527: 523: 522: 518: 515: 513:Chlorogaster 512: 509: 508: 499: 497: 494: 493: 489: 486: 485: 481: 480: 477: 475: 472: 471: 467: 464: 463: 459: 458: 454: 453: 449: 446: 445: 441: 438: 437: 433: 430: 429: 425: 424: 420: 419: 415: 412: 411: 407: 404: 403: 399: 396: 395: 391: 390: 386: 385: 381: 378: 377: 373: 370: 369: 365: 362: 360: 356: 355: 351: 350: 346: 343: 342: 338: 335: 334: 330: 327: 326: 322: 321: 317: 314: 311: 308: 307: 302: 299: 289: 287: 283: 279: 275: 271: 267: 263: 259: 255: 249: 244: 240: 233: 226: 222: 212: 210: 207: 206:herpetologist 204: 200: 196: 186: 182: 179: 170: 168: 165: 161: 157: 156: 145: 140: 139: 135: 132: 131: 128: 125: 122: 121: 118: 115: 112: 111: 108: 105: 102: 101: 98: 95: 92: 91: 88: 85: 82: 81: 78: 75: 72: 71: 68: 65: 62: 61: 56: 51: 47: 44: 43: 38: 34: 29: 26: 22: 19: 1594: 1574: 1573:(1): 1-22. ( 1570: 1567: 1563: 1556: 1530: 1525: 1517: 1505: 1493: 1473: 1443: 1425: 1414: 1410: 1406: 1400: 1396: 1393:D. valentini 1392: 1388: 1384: 1382: 1377: 1373: 1371: 1353: 1350:D. brauneri) 1349: 1346: 1343:Reproduction 1337: 1333: 1325: 1321: 1317: 1313: 1304: 1300: 1289: 1286: 1281: 1279: 1274: 1271: 1267:D. valentini 1266: 1262: 1258: 1255: 1250: 1246: 1244: 1240: 1231: 1228:Distribution 1221: 1211: 1210: 1199: 1185: 1175: 1160: 1149: 1137: 1125: 1115: 1105: 1084: 1070: 1060: 1045: 1030: 1016: 997: 985: 975: 963: 949: 937: 925: 913: 902: 888: 873: 863: 853: 839: 824: 814: 810:et al., 2013 801: 790: 780: 766: 755: 740: 730: 713: 696: 691: 676: 660: 644: 628: 620: 606: 598: 590: 578: 570: 562: 550: 542: 534: 529: 524: 490: 482: 468: 460: 450: 442: 434: 426: 416: 408: 400: 392: 382: 374: 366: 357: 347: 339: 331: 323: 301:subspecies. 297: 295: 285: 281: 269: 265: 261: 254:Lajos MĂ©helĂż 242: 238: 231: 224: 218: 198: 195:generic name 192: 183: 177: 176: 154: 153: 152: 137: 136: 40: 24: 18: 1725:iNaturalist 1619:Wikispecies 1498:Beolens, Bo 516:Defilippii 286:D. saxicola 266:L. saxicola 262:L. saxicola 239:L. saxicola 173:Description 123:Subfamily: 1812:Categories 1564:sensu lato 1435:References 1389:D. uzzelli 847:Vedmederja 808:Ahmadzadeh 774:Schmidtler 510:Praticola 318:Caucasica 292:Cladistics 270:L. muralis 243:L. muralis 167:Lacertidae 127:Lacertinae 117:Lacertidae 1818:Darevskia 1639:Darevskia 1625:Darevskia 1595:Darevskia 1575:Darevskia 1532:Darevskia 1520:, p. 65). 1518:Darevskia 1378:Darevskia 1374:Darevskia 1297:Nutrition 1259:D. alpina 1222:Darevskia 1213:Nota bene 1193:Danielyan 1038:Eversmann 957:Szczerbak 832:Boulenger 703:(Darevsky 315:Saxicola 298:Darevskia 282:Darevskia 199:Darevskia 189:Etymology 178:Darevskia 155:Darevskia 138:Darevskia 73:Kingdom: 67:Eukaryota 25:Darevskia 1604:Wikidata 1562:, 1921, 1471:(2006), 1420:See also 1330:Activity 1078:Bedriaga 1053:Boettger 896:Nikolsky 881:Camerano 720:(Tuniyev 248:Laurenti 113:Family: 107:Squamata 97:Reptilia 87:Chordata 83:Phylum: 77:Animalia 63:Domain: 1790:ZooBank 1782:1071242 1756:1155158 1743:1062515 1717:2468734 1610:Q902055 1560:Mertens 1237:Habitat 1195:, 1977) 1170:, 1963) 1101:, 2022) 1091:(Kurnaz 1080:, 1886) 1055:, 1892) 1040:, 1834) 1026:, 1878) 1024:Kessler 1011:, 1913) 959:, 1962) 898:, 1898) 883:, 1877) 849:, 1977) 834:, 1908) 750:, 1909) 726:, 2019) 724:Petrova 709:, 1980) 688:Species 309:Raddei 250:, 1768) 203:Russian 144:Arribas 133:Genus: 103:Order: 93:Class: 1769:122330 1691:214623 1529:Genus 1512:  1481:  1358:Clutch 1168:Lukina 1099:EroÄźlu 1097:& 1007:& 748:MĂ©helĂż 722:& 707:Eiselt 705:& 312:Rudis 164:family 146:, 1999 1738:IRMNG 1730:35361 1678:17152 1665:6327R 1652:88012 1095:Ĺžahin 1009:CyrĂ©n 1005:Lantz 160:genus 158:is a 1764:NCBI 1751:ITIS 1712:GBIF 1647:BOLD 1510:ISBN 1479:ISBN 1395:and 1387:and 1265:and 1261:and 1216:: A 193:The 1673:EoL 1660:CoL 1634:ADW 1535:at 1516:. ( 1253:). 1814:: 1792:: 1779:: 1766:: 1753:: 1740:: 1727:: 1714:: 1701:: 1688:: 1675:: 1662:: 1649:: 1636:: 1621:: 1606:: 1500:; 1467:; 1463:; 1459:; 1455:; 1451:; 1284:. 1277:. 1269:. 1224:. 1093:, 288:. 211:. 197:, 1571:6 1487:. 1348:( 1166:( 1142:( 1134:) 1130:( 1076:( 1051:( 1036:( 1022:( 1003:( 994:) 990:( 972:) 968:( 955:( 946:) 942:( 934:) 930:( 922:) 918:( 894:( 879:( 830:( 772:( 746:( 361:( 246:(

Index


Darevskia raddei
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Reptilia
Squamata
Lacertidae
Lacertinae
Darevskia
Arribas
genus
family
Lacertidae
generic name
Russian
herpetologist
Ilya Sergeyevich Darevsky
Eduard Friedrich Eversmann
Laurenti
Lajos MĂ©helĂż
George Albert Boulenger
Ilya Sergeyevich Darevsky
Oscar J. Arribas
Darevskia raddei
Darevskia rudis
Darevskia saxicola
Darevskia caucasica

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