33:
55:
1412:
Nevertheless, presumably, these males are able to give offspring, which leads to the appearance of tetraploid hybrids. Sometimes males emerge from unfertilized eggs laid by parthenogenetic females. Like males of hybrid origin, they have reproductive disorders, which may not interfere with their offspring. A small amount of parthenogenetic female males emerging from clutches is explained by frequently occurring mutations incompatible with life.
185:
sides of the body. In some species, blue or violet spots with white circles in the center and / or monotonous blue-violet spots at the junction of the abdominal scutes with trunk scales are located on the anterior third of the body. Most types of rock lizards are characterized by a diverse color of the abdominal side of the body, ranging from various shades of pink, red and orange to yellow and green.
1339:
hours in the middle of the day . After heating (basking), the body temperature of the lizard reaches about 30–34 °C (86–93 °F), and it begins a routine activity aimed at supporting the body. Late in the evening, when the heat subsides, the animals return to the basking places and stay there for some time, after which they go to their night shelters.
272:. For several decades, scientists from different countries studied the monophyletic group of rock lizards independently, highlighting new subspecies, simplifying some taxa and describing new species. It was on this group of lizards that the phenomenon of parthenogenesis in amniotic vertebrates was first discovered by the Russian zoologist
1335:
until the end of May, exit from wintering shelters occurs, and the active period is from 6–7 months (in the mountains) and up to 9–10 months (in the valleys and on the seashore). During this period, mature individuals mate, and females lay eggs. Hibernation takes place from late
September to mid November.
1272:
3) Rock lizards of dry and moderately dry landscapes (alpine steppes) of rocks and their feet on slopes with dry-loving shrubbery and grassy vegetation, road slopes. Such habitats have a large number of crevices and voids serving as shelters for lizards. Such species are found in such landscapes as:
300:
includes four groups (clades), combining species by origin and kinship: “raddei”, “rudis”, “saxicola” and “caucasica”. Subsequently, three more clades were identified: “praticola”, “chlorogaster”, and “defilippii”. In total, the genus includes 35 species, 7 of which breed parthenogenetically, and 22
1232:
Rock lizards are common in
Abkhazia, Azerbaijan, Armenia, Georgia, Iran, Nagorno-Karabakh, Russia (Adygea, Dagestan, Ingushetia, Kabardino-Balkaria, Karachay-Cherkessia, Krasnodar Territory, Republic of Crimea, North Ossetia-Alania, Stavropol Territory and Chechnya), in Turkey and South Ossetia. It
1407:
The appearance of parthenogenetic species is usually associated with the last, Wurm glaciation. Parthenogenetic rock lizards appeared about 10 thousand years ago when, due to the formation of mountain glaciers, the habitats of the parent bisexual species were disturbed, which led to the overlap of
1403:
always enter as maternal species. From the point of view of the classical concept of a species, homosexual taxa cannot be assigned a species status due to the lack of exchange of genetic material between individuals within the same population. However, on the basis of morphological and cytogenetic
1338:
The beginning and end of the daily activity of the lizard is determined by the lighting conditions within the individual area, and in some individuals it can start early in the morning, while individuals living on the slopes of the northern exposure or in deep forest valleys are active for several
1301:
Rock lizards feed on various invertebrates with a body size from a few millimeters to 4 cm (1.6 in): spiders, diptera, lepidoptera, hymenoptera, cockroaches, orthoptera, semi-rigid-winged, coleoptera, wood lice, worms, slugs, marine and freshwater amphipods, caddis flies, chironomids and
184:
The color of rock lizards varies from different tones of green to sand. Females are usually colored paler than males. On the dorsal side of the body, rock lizards have an occipital stripe composed of a set of black or brown spots and a wide line of the lizard's main color, and dark patterns on the
1415:
Despite the fact that as a result of parthenogenesis, individuals are born that receive hereditary material only from the mother's body, a small intraspecific genetic diversity has been revealed due to mutational processes, genetic instability and the appearance of same-sex species as a result of
1334:
The seasonal activity of rock lizards is determined by temperature indicators. Therefore, species living at different heights differ in terms of exit from wintering shelters, mating period, laying of eggs, hatching of young individuals and time of leaving for wintering. Around the end of
February
1411:
Hybridization occurs in places where the ranges of parthenogenetic and bisexual species overlap, resulting in the appearance of triploid sterile females and a small number of males, with changes in the reproductive system (malfunctions in the formation of reproductive products, hermaphroditism).
1305:
Despite the diversity of the food supply, rock lizards can develop preferences for feeding on invertebrates of a certain group (for example, flying forms of ants), which is caused by seasonal changes in the availability or abundance of this type of prey. Even after a significant reduction in the
1241:
Rock lizards are found in various high-altitude zones from 0 to 3,000 m (0 to 9,843 ft) above sea level and occupy a variety of landscapes: mountain-steppe, forest-steppe, mountain meadow, mountain forest, anthropogenic and coastal. By confinement to one or another habitat, they can be
180:
species are small lizards with a head and body length of 50–85 mm (2.0–3.3 in), and with a tail length about two times longer. The body is usually flattened, the head is pointed in shape and in most species flattened in a vertical plane, which allows lizards to hide in narrow crevices
1314:
Rock lizards are extremely rare on their own, usually forming settlements. The population density of parthenogenetic rock lizards can vary in a wider range than that of bisexual species: up to 200 individuals per 1 km of the route in unisexual species and up to 80 individuals in bisexual
1322:
The basis of the settlements of bisexual rock lizards are sedentary males and females with individual sites, often overlapping. In a number of species, some males possess territories protected from other males. The territories of males never overlap, but their centers of activity, primarily
1408:
their ranges and the hybridogenic formation of parthenogenetic individuals better adapted to the conditions of short summers and long winters. Due to the doubled reproduction rate and successful resettlement, same-sex populations later began to exist independently of the parent species.
1318:
Rock lizards are characterized by complex and diverse social systems, which, in particular, are characterized by stable long-term friendly relations between the male and the female and territorial or hierarchical relations between individuals of the same sex.
1292:
on the southern slope of the
Greater Caucasus Mountain Range (South Ossetia) is distributed at altitudes of 1,500–1,800 m (4,900–5,900 ft) above sea level, and on the northern slopes (Dagestan) at 50–2,100 m (160–6,890 ft) above sea level.
1347:
Rock lizards reach maturity in the second (females) and third (males) year of life with a total life expectancy of up to 13 years. Mating of some species of rock lizards occurs after the first molt, approximately 3–5 weeks after leaving wintering
1287:
Rock lizards are found at heights of 0 – 3000 m above sea level. Zonal and geographical distribution is determined by the amount of precipitation, average annual temperature, duration of the adverse season, and exposure of the slope. For example,
181:
between stones and rocks. Rock lizards have relatively long legs with special calluses on the inner surfaces of the paws and sharp claws, thanks to which they quickly move along the vertical rough surfaces of rocks and stones.
1256:
2) Inhabitants in areas of bedrock outcrops and clayey cliffs in alpine and subalpine meadows. As shelters, they often use rodent burrows, cavities between stones and cracks in the rocks. These habitats are adhered to by
1233:
is worth noting that the boundaries of the range of some species are not reliably known, but the expected areas of their encounters coincide with the already indicated distribution sites for the whole genus
1326:
The study of the social behavior and spatial structure of rock lizard populations has been the subject of a number of scientific papers published based on the results of many years of research
169:. Member species are native to the Caucasus, Iran and Turkey, living in forest and grassy habitats with numerous rock outcrops. Among rock lizards, seven parthenogenetic species are known.
1280:
4) Occupying anthropogenic habitats: an abandoned building, walls in cities, abandoned temples, monasteries, etc., where their number often exceeds that in natural habitats. For example,
1380:
penetrated the territory of the
Western Caucasus in the Middle Miocene or in the Middle Pliocene, when there was a land connection between Asia Minor and the Balkans.
1249:
and independent of them, able to live in habitats in which there are no rocks, using rodent burrows as shelters, deciduous litter, cavities in trees and bark (e.g.
276:(1924–2009), who also made a significant contribution to understanding the ecology, systematics, and morphology of rock lizards. In 1997, the Spanish scientist
32:
1376:
from the subfamily
Lacertinae occurred approximately 12-16 million years ago, in the middle of the Miocene. Presumably, the ancestral form of the genus
1737:
1763:
1531:
1306:
density of invertebrates of this group, lizards continue to hunt for them for some time in the presence of more accessible food objects.
1711:
1827:
1750:
1315:
species, which is explained by the fact that parthenogenetic species are less aggressive and have a high population growth rate.
1755:
1513:
1167:
846:
1468:
1464:
223:(1794–1860) made an expedition to the North Caucasus. As a result of which he described two new species: a meadow lizard,
1094:
1448:
773:
723:
1245:
1) Lizards living in the forest, according to the occupied microreliefs, are divided into: adhering to rock outcrops (
1482:
1302:
sometimes parts of plants. Also, particular cases of cannibalism were recorded when adults ate juvenile individuals.
1690:
1008:
1768:
1703:
1004:
895:
719:
1323:
associated with basking, coincide with the centers of activity of females that live within their territories.
1794:
1098:
1646:
1651:
1360:
size is usually 2 to 7 eggs. The incubation period lasts approximately 55–65 days. Young animals with a
1192:
1037:
914:
807:
247:
220:
54:
1456:
1138:
1552:
767:
579:
277:
143:
1501:
1391:, resulting from the hybridization of males and females of different bisexual species. Moreover,
1023:
1017:
831:
435:
257:
1822:
1586:
1090:
1061:
938:
854:
825:
629:
571:
535:
417:
1742:
1781:
1472:
1361:
1217:
1176:
645:
1672:
1633:
1161:
1106:
1077:
874:
706:
677:
543:
409:
1555:(1999). "Phylogeny and relationships of the mountain lizards of Europe and the Near East (
8:
1624:
1385:
D. armeniaca, D. bendimahiensis, D. dahli, D. rostombekovi, D. sapphirina, D. unisexualis
1220:
in parentheses indicates that the species was originally described in a genus other than
1200:
1126:
1031:
964:
950:
889:
840:
815:
781:
741:
714:
599:
525:
451:
443:
427:
401:
393:
358:
348:
1474:
The
Reptiles of Northern Eurasia: Taxonomic Diversity, Distribution, Conservation Status
1452:
1817:
1497:
1150:
1116:
791:
756:
697:
621:
491:
469:
375:
340:
49:
237:. At that time, European scientists did not recognize the independence of the species
1776:
1659:
1509:
1478:
1186:
1085:
998:
986:
802:
747:
661:
591:
367:
253:
603:
Ahmadzadeh, Flecks, Carretero, Mozaffari, Bohme, Harris, Freitas & Rodder, 2013
595:
Ahmadzadeh, Flecks, Carretero, Mozaffari, Bohme, Harris, Freitas & Rodder, 2013
567:
Ahmadzadeh, Flecks, Carretero, Mozaffari, Bohme, Harris, Freitas & Rodder, 2013
1664:
1367:
1046:
956:
903:
880:
731:
483:
383:
324:
163:
41:
1638:
920:
Arribas, Candan, Kornilios, Ayaz, Kumlutas, Gul, Yilmaz, Caynak, & Ilgaz, 2022
1071:
976:
926:
864:
607:
563:
551:
461:
332:
1698:
1685:
1559:
1052:
575:
Ahmadzadeh, Flecks, Carretero, Mozaffari, Bohme, Harris, Freitas, Rodder, 2013
1811:
1609:
1477:, Faunastica, vol. 47, Sofia, Bulgaria: Pensoft Publishers, p. 79,
1460:
1426:
1357:
1356:. Egg laying begins in the second half of June and lasts until early August.
1309:
702:
273:
208:
260:(1858–1937) there was a lengthy discussion about the taxonomic position of
1724:
1618:
205:
1416:
repeated and independent crossing of parental species among themselves.
1716:
166:
126:
116:
1729:
1212:
66:
1580:
252:. But at the beginning of the XX century between the two zoologists
1603:
1536:
202:
106:
86:
1566:) and their relationships among the Eurasian lacertid radiation".
1364:(SVL) of about 25 mm (0.98 in) are born in late summer.
1275:
D. rudis, D. portschinski, D. daghestanica, D. raddei, D. saxicola
1789:
96:
1430:, another lizard genus containing several parthenogenic species.
1368:
The evolution and origin of parthenogenetic species of
Darevskia
76:
1677:
1447:
473:
Arribas, Candan, Jurnaz, KumlutaĹź, Caynak, & Ilgaz, 2022
194:
159:
1508:. Baltimore: Johns Hopkins University Press. xiii + 296 pp.
1352:, in others, immediately after leaving wintering shelter (
1310:
Population density and spatial structure of the population
1383:
The genus of rock lizards has 7 parthenogenetic species:
970:
Arribas, Ilgaz, Kumluras, Durmus, Avci, & Ăśzum, 2013
268:
and subspecies were further considered separately from
692:
The following species are recognized as being valid.
264:, which was resolved in favor of the former, and
1809:
1404:characters, they are assigned a species rank.
1146:) - Caucasian rock lizard, Valentin's lizard
1441:
1242:conditionally divided into several groups:
214:
1207:– Caucasian rock lizard, Valentin's lizard
681:(Schmidtler, Eiselt & Darevsky, 1994)
583:(Schmidtler, Eiselt & Darevsky, 1994)
219:In 1830, a professor at Kazan University,
31:
1111:(Schmidtler, Eiselt & Darevsky, 1994)
241:, considering it as part of the European
1144:Eiselt, Darevsky, & Schmidtler, 1992
296:According to Arribas (1997), the genus
1810:
1399:always enter as paternal species, and
1585:
1584:
761:(Tuniyev, Petrova & Lotiev, 2023)
1795:DBEC2AE0-D337-4660-8FAF-072A42C604D2
1704:89c08cb0-c039-4b1f-b67b-5ce5169719e3
1282:D. armeniaca, D. lindholmi, D. dahli
284:and designated the type species as
13:
1546:
1523:
431:(Darevsky & Vedmederja, 1977)
14:
1839:
1506:The Eponym Dictionary of Reptiles
1401:D. mixta, D. nairensis, D. raddei
665:(Darevsky & Danielyan, 1977)
280:named the genus of rock lizards
53:
1342:
1227:
487:(Darevsky & Tuniyev, 1997)
1828:Taxa named by Oscar J. Arribas
1568:Russian Journal of Herpetology
1491:
908:(Darevsky & Tuniyev, 1997)
776:, Eiselt & Darevsky, 1994)
495:(Darevsky & Eiselt, 1980)
172:
1:
1434:
1251:D. chlorogaster, D. armeniaca
291:
1296:
188:
7:
1504:; Grayson, Michael (2011).
1419:
1329:
10:
1844:
1236:
915:Darevskia josefschmidtleri
687:
447:(Lantz & Cyren, 1936)
371:(Lantz & Cyren, 1913)
221:Eduard Friedrich Eversmann
1593:
1539:www.reptile-database.org.
1139:Darevskia spitzenbergerae
274:Ilya Sergeyevich Darevsky
209:Ilya Sergeyevich Darevsky
50:Scientific classification
48:
39:
30:
23:
1372:Separation of the genus
768:Darevskia bendimahiensis
580:Darevskia bendimahiensis
519:Parthenogenetic species
215:Taxonomy and systematics
1247:D. raddei, D. brauneri)
1132:Ahmadzadeh et al., 2013
1018:Darevskia portschinskii
992:Lantz & Cyrén, 1936
944:Ahmadzadeh et al., 2013
932:Ahmadzadeh et al., 2013
798:– Brauner's rock lizard
436:Darevskia portschinskii
258:George Albert Boulenger
162:of wall lizards of the
1263:D. mixta, D. armeniaca
1062:Darevskia rostombekowi
939:Darevskia kopetdaghica
855:Darevskia daghestanica
826:Darevskia chlorogaster
763:– Arribas’ rock lizard
630:Darevskia rostombekowi
572:Darevskia kopetdaghica
536:Darevskia chlorogaster
418:Darevskia daghestanica
1177:Darevskia unisexualis
646:Darevskia unisexualis
1699:Fauna Europaea (new)
1537:The Reptile Database
1469:Barabanov, Andrei V.
1449:Ananjeva, Natalia B.
1362:snout-to-vent length
1172:– Szczerbak's lizard
1162:Darevskia szczerbaki
1107:Darevskia sapphirina
1013:– red-bellied lizard
875:Darevskia defilippii
678:Darevskia sapphirina
544:Darevskia defilippii
230:, and a rock lizard
1201:Darevskia valentini
1127:Darevskia schaekeli
1057:– Azerbaijan lizard
1032:Darevskia praticola
965:Darevskia mirabilis
951:Darevskia lindholmi
890:Darevskia derjugini
841:Darevskia clarkorum
836:– greenbelly lizard
816:Darevskia caucasica
782:Darevskia bithynica
742:Darevskia armeniaca
715:Darevskia aghasyani
600:Darevskia schaekeli
552:Darevskia armeniaca
526:Darevskia praticola
506:
452:Darevskia derjugini
444:Darevskia lindholmi
428:Darevskia clarkorum
410:Darevskia sczerbaki
402:Darevskia valentini
394:Darevskia bithynica
359:Darevskia nairensis
349:Darevskia caucasica
305:
1457:Khalikov, Roman G.
1218:binomial authority
1157:– Steiner's lizard
1151:Darevskia steineri
1117:Darevskia saxicola
792:Darevskia brauneri
757:Darevskia arribasi
698:Darevskia adjarica
622:Darevskia steineri
539:(Boulenger, 1908)
530:(Eversmann, 1834)
505:
492:Darevskia adjarica
470:Darevskia tuniyevi
376:Darevskia brauneri
344:(Eversmann, 1834)
341:Darevskia saxicola
304:
1805:
1804:
1777:Open Tree of Life
1587:Taxon identifiers
1514:978-1-4214-0135-5
1465:Ryabov, Sergei A.
1461:Darevsky, Ilya S.
1453:Orlov, Nikolai L.
1206:
1196:
1187:Darevskia uzzelli
1182:
1171:
1156:
1122:
1121:(Eversmann, 1834)
1112:
1102:
1086:Darevskia salihae
1081:
1067:
1056:
1041:
1027:
1012:
999:Darevskia parvula
987:Darevskia obscura
982:
960:
910:– Charnali lizard
909:
899:
884:
870:
860:
850:
835:
821:
803:Darevskia caspica
797:
787:
777:
762:
752:– Armenian lizard
751:
737:
727:
710:
685:
684:
662:Darevskia uzzelli
649:(Darevsky, 1966)
633:(Darevsky, 1957)
611:(Darevsky, 1957)
592:Darevskia caspica
547:(Camerano, 1877)
503:
502:
455:(Nikolsky, 1898)
421:(Darevsky, 1967)
405:(Boettger, 1892)
387:(Darevsky, 1967)
368:Darevskia parvula
336:(Bedriaga, 1886)
328:(Boettger, 1892)
235:(Eversmann, 1834)
228:(Eversmann, 1834)
225:Lacerta praticola
201:, is in honor of
151:
150:
147:
1835:
1798:
1797:
1785:
1784:
1772:
1771:
1759:
1758:
1746:
1745:
1733:
1732:
1720:
1719:
1707:
1706:
1694:
1693:
1681:
1680:
1668:
1667:
1655:
1654:
1642:
1641:
1629:
1628:
1627:
1614:
1613:
1612:
1582:
1581:
1540:
1527:
1521:
1502:Watkins, Michael
1495:
1489:
1488:
1445:
1397:D. portschinskii
1205:(Boettger, 1892)
1204:
1191:(Darevsky &
1190:
1181:(Darevsky, 1966)
1180:
1165:
1154:
1145:
1133:
1120:
1110:
1089:
1075:
1066:(Darevsky, 1957)
1065:
1050:
1047:Darevskia raddei
1035:
1021:
1002:
993:
980:
971:
954:
945:
933:
921:
907:
904:Darevskia dryada
893:
878:
869:(Darevsky, 1957)
868:
859:(Darevsky, 1967)
858:
845:(Darevsky &
844:
829:
819:
811:
795:
785:
771:
760:
745:
736:(Darevsky, 1967)
735:
732:Darevskia alpina
718:
701:
507:
504:
484:Darevskia dryada
439:(Kessler, 1878)
384:Darevskia alpina
363:Darevsky, 1967)
325:Darevskia raddei
306:
303:
278:Oscar J. Arribas
256:(1862–1953) and
251:
236:
232:Lacerta saxicola
229:
142:
58:
57:
42:Darevskia raddei
35:
21:
20:
16:Genus of lizards
1843:
1842:
1838:
1837:
1836:
1834:
1833:
1832:
1808:
1807:
1806:
1801:
1793:
1788:
1780:
1775:
1767:
1762:
1754:
1749:
1741:
1736:
1728:
1723:
1715:
1710:
1702:
1697:
1689:
1684:
1676:
1671:
1663:
1658:
1650:
1645:
1637:
1632:
1623:
1622:
1617:
1608:
1607:
1602:
1589:
1549:
1547:Further reading
1544:
1543:
1528:
1524:
1496:
1492:
1485:
1446:
1442:
1437:
1422:
1370:
1345:
1332:
1312:
1299:
1290:D. daghestanica
1239:
1230:
1143:
1131:
1072:Darevskia rudis
1042:– meadow lizard
991:
977:Darevskia mixta
969:
943:
931:
927:Darevskia kamii
919:
885:– Elburs lizard
865:Darevskia dahli
806:
690:
625:(Eiselt, 1995)
608:Darevskia dahli
564:Darevskia kamii
555:(Mehely, 1909)
465:(Mehely, 1909)
462:Darevskia mixta
413:(Lukina, 1963)
397:(Mehely, 1909)
379:(Mehely, 1909)
352:(Mehely, 1909)
333:Darevskia rudis
294:
245:
234:
227:
217:
191:
175:
141:
52:
17:
12:
11:
5:
1841:
1831:
1830:
1825:
1820:
1803:
1802:
1800:
1799:
1786:
1773:
1760:
1747:
1734:
1721:
1708:
1695:
1686:Fauna Europaea
1682:
1669:
1656:
1643:
1630:
1615:
1599:
1597:
1591:
1590:
1579:
1578:
1553:Arribas, Oscar
1548:
1545:
1542:
1541:
1522:
1490:
1483:
1439:
1438:
1436:
1433:
1432:
1431:
1421:
1418:
1369:
1366:
1344:
1341:
1331:
1328:
1311:
1308:
1298:
1295:
1238:
1235:
1229:
1226:
1209:
1208:
1197:
1183:
1173:
1158:
1155:(Eiselt, 1995)
1147:
1135:
1123:
1113:
1103:
1082:
1068:
1058:
1043:
1028:
1014:
995:
983:
981:(MĂ©helĂż, 1909)
973:
961:
947:
935:
923:
911:
900:
886:
871:
861:
851:
837:
822:
820:(MĂ©helĂż, 1909)
812:
799:
796:(MĂ©helĂż, 1909)
788:
786:(MĂ©helĂż, 1909)
778:
764:
753:
738:
728:
711:
689:
686:
683:
682:
674:
672:
670:
667:
666:
658:
656:
654:
651:
650:
642:
640:
638:
635:
634:
626:
618:
616:
613:
612:
604:
596:
588:
585:
584:
576:
568:
560:
557:
556:
548:
540:
532:
521:
520:
517:
514:
511:
501:
500:
498:
496:
488:
479:
478:
476:
474:
466:
457:
456:
448:
440:
432:
423:
422:
414:
406:
398:
389:
388:
380:
372:
364:
354:
353:
345:
337:
329:
320:
319:
316:
313:
310:
293:
290:
216:
213:
190:
187:
174:
171:
149:
148:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
74:
70:
69:
64:
60:
59:
46:
45:
37:
36:
28:
27:
15:
9:
6:
4:
3:
2:
1840:
1829:
1826:
1824:
1823:Lizard genera
1821:
1819:
1816:
1815:
1813:
1796:
1791:
1787:
1783:
1778:
1774:
1770:
1765:
1761:
1757:
1752:
1748:
1744:
1739:
1735:
1731:
1726:
1722:
1718:
1713:
1709:
1705:
1700:
1696:
1692:
1687:
1683:
1679:
1674:
1670:
1666:
1661:
1657:
1653:
1648:
1644:
1640:
1635:
1631:
1626:
1620:
1616:
1611:
1605:
1601:
1600:
1598:
1596:
1592:
1588:
1583:
1577:, new genus).
1576:
1572:
1569:
1565:
1561:
1558:
1557:Archeolacerta
1554:
1551:
1550:
1538:
1534:
1533:
1526:
1519:
1515:
1511:
1507:
1503:
1499:
1494:
1486:
1484:954-642-269-X
1480:
1476:
1475:
1470:
1466:
1462:
1458:
1454:
1450:
1444:
1440:
1429:
1428:
1427:Cnemidophorus
1424:
1423:
1417:
1413:
1409:
1405:
1402:
1398:
1394:
1390:
1386:
1381:
1379:
1375:
1365:
1363:
1359:
1355:
1354:D. valentini)
1351:
1340:
1336:
1327:
1324:
1320:
1316:
1307:
1303:
1294:
1291:
1285:
1283:
1278:
1276:
1270:
1268:
1264:
1260:
1254:
1252:
1248:
1243:
1234:
1225:
1223:
1219:
1215:
1214:
1203:
1202:
1198:
1194:
1189:
1188:
1184:
1179:
1178:
1174:
1169:
1164:
1163:
1159:
1153:
1152:
1148:
1141:
1140:
1136:
1129:
1128:
1124:
1119:
1118:
1114:
1109:
1108:
1104:
1100:
1096:
1092:
1088:
1087:
1083:
1079:
1074:
1073:
1069:
1064:
1063:
1059:
1054:
1049:
1048:
1044:
1039:
1034:
1033:
1029:
1025:
1020:
1019:
1015:
1010:
1006:
1001:
1000:
996:
989:
988:
984:
979:
978:
974:
967:
966:
962:
958:
953:
952:
948:
941:
940:
936:
929:
928:
924:
917:
916:
912:
906:
905:
901:
897:
892:
891:
887:
882:
877:
876:
872:
867:
866:
862:
857:
856:
852:
848:
843:
842:
838:
833:
828:
827:
823:
818:
817:
813:
809:
805:
804:
800:
794:
793:
789:
784:
783:
779:
775:
770:
769:
765:
759:
758:
754:
749:
744:
743:
739:
734:
733:
729:
725:
721:
717:
716:
712:
708:
704:
700:
699:
695:
694:
693:
680:
679:
675:
673:
671:
669:
668:
664:
663:
659:
657:
655:
653:
652:
648:
647:
643:
641:
639:
637:
636:
632:
631:
627:
624:
623:
619:
617:
615:
614:
610:
609:
605:
602:
601:
597:
594:
593:
589:
587:
586:
582:
581:
577:
574:
573:
569:
566:
565:
561:
559:
558:
554:
553:
549:
546:
545:
541:
538:
537:
533:
531:
528:
527:
523:
522:
518:
515:
513:Chlorogaster
512:
509:
508:
499:
497:
494:
493:
489:
486:
485:
481:
480:
477:
475:
472:
471:
467:
464:
463:
459:
458:
454:
453:
449:
446:
445:
441:
438:
437:
433:
430:
429:
425:
424:
420:
419:
415:
412:
411:
407:
404:
403:
399:
396:
395:
391:
390:
386:
385:
381:
378:
377:
373:
370:
369:
365:
362:
360:
356:
355:
351:
350:
346:
343:
342:
338:
335:
334:
330:
327:
326:
322:
321:
317:
314:
311:
308:
307:
302:
299:
289:
287:
283:
279:
275:
271:
267:
263:
259:
255:
249:
244:
240:
233:
226:
222:
212:
210:
207:
206:herpetologist
204:
200:
196:
186:
182:
179:
170:
168:
165:
161:
157:
156:
145:
140:
139:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
78:
75:
72:
71:
68:
65:
62:
61:
56:
51:
47:
44:
43:
38:
34:
29:
26:
22:
19:
1594:
1574:
1573:(1): 1-22. (
1570:
1567:
1563:
1556:
1530:
1525:
1517:
1505:
1493:
1473:
1443:
1425:
1414:
1410:
1406:
1400:
1396:
1393:D. valentini
1392:
1388:
1384:
1382:
1377:
1373:
1371:
1353:
1350:D. brauneri)
1349:
1346:
1343:Reproduction
1337:
1333:
1325:
1321:
1317:
1313:
1304:
1300:
1289:
1286:
1281:
1279:
1274:
1271:
1267:D. valentini
1266:
1262:
1258:
1255:
1250:
1246:
1244:
1240:
1231:
1228:Distribution
1221:
1211:
1210:
1199:
1185:
1175:
1160:
1149:
1137:
1125:
1115:
1105:
1084:
1070:
1060:
1045:
1030:
1016:
997:
985:
975:
963:
949:
937:
925:
913:
902:
888:
873:
863:
853:
839:
824:
814:
810:et al., 2013
801:
790:
780:
766:
755:
740:
730:
713:
696:
691:
676:
660:
644:
628:
620:
606:
598:
590:
578:
570:
562:
550:
542:
534:
529:
524:
490:
482:
468:
460:
450:
442:
434:
426:
416:
408:
400:
392:
382:
374:
366:
357:
347:
339:
331:
323:
301:subspecies.
297:
295:
285:
281:
269:
265:
261:
254:Lajos MĂ©helĂż
242:
238:
231:
224:
218:
198:
195:generic name
192:
183:
177:
176:
154:
153:
152:
137:
136:
40:
24:
18:
1725:iNaturalist
1619:Wikispecies
1498:Beolens, Bo
516:Defilippii
286:D. saxicola
266:L. saxicola
262:L. saxicola
239:L. saxicola
173:Description
123:Subfamily:
1812:Categories
1564:sensu lato
1435:References
1389:D. uzzelli
847:Vedmederja
808:Ahmadzadeh
774:Schmidtler
510:Praticola
318:Caucasica
292:Cladistics
270:L. muralis
243:L. muralis
167:Lacertidae
127:Lacertinae
117:Lacertidae
1818:Darevskia
1639:Darevskia
1625:Darevskia
1595:Darevskia
1575:Darevskia
1532:Darevskia
1520:, p. 65).
1518:Darevskia
1378:Darevskia
1374:Darevskia
1297:Nutrition
1259:D. alpina
1222:Darevskia
1213:Nota bene
1193:Danielyan
1038:Eversmann
957:Szczerbak
832:Boulenger
703:(Darevsky
315:Saxicola
298:Darevskia
282:Darevskia
199:Darevskia
189:Etymology
178:Darevskia
155:Darevskia
138:Darevskia
73:Kingdom:
67:Eukaryota
25:Darevskia
1604:Wikidata
1562:, 1921,
1471:(2006),
1420:See also
1330:Activity
1078:Bedriaga
1053:Boettger
896:Nikolsky
881:Camerano
720:(Tuniyev
248:Laurenti
113:Family:
107:Squamata
97:Reptilia
87:Chordata
83:Phylum:
77:Animalia
63:Domain:
1790:ZooBank
1782:1071242
1756:1155158
1743:1062515
1717:2468734
1610:Q902055
1560:Mertens
1237:Habitat
1195:, 1977)
1170:, 1963)
1101:, 2022)
1091:(Kurnaz
1080:, 1886)
1055:, 1892)
1040:, 1834)
1026:, 1878)
1024:Kessler
1011:, 1913)
959:, 1962)
898:, 1898)
883:, 1877)
849:, 1977)
834:, 1908)
750:, 1909)
726:, 2019)
724:Petrova
709:, 1980)
688:Species
309:Raddei
250:, 1768)
203:Russian
144:Arribas
133:Genus:
103:Order:
93:Class:
1769:122330
1691:214623
1529:Genus
1512:
1481:
1358:Clutch
1168:Lukina
1099:EroÄźlu
1097:&
1007:&
748:MĂ©helĂż
722:&
707:Eiselt
705:&
312:Rudis
164:family
146:, 1999
1738:IRMNG
1730:35361
1678:17152
1665:6327R
1652:88012
1095:Ĺžahin
1009:Cyrén
1005:Lantz
160:genus
158:is a
1764:NCBI
1751:ITIS
1712:GBIF
1647:BOLD
1510:ISBN
1479:ISBN
1395:and
1387:and
1265:and
1261:and
1216:: A
193:The
1673:EoL
1660:CoL
1634:ADW
1535:at
1516:. (
1253:).
1814::
1792::
1779::
1766::
1753::
1740::
1727::
1714::
1701::
1688::
1675::
1662::
1649::
1636::
1621::
1606::
1500:;
1467:;
1463:;
1459:;
1455:;
1451:;
1284:.
1277:.
1269:.
1224:.
1093:,
288:.
211:.
197:,
1571:6
1487:.
1348:(
1166:(
1142:(
1134:)
1130:(
1076:(
1051:(
1036:(
1022:(
1003:(
994:)
990:(
972:)
968:(
955:(
946:)
942:(
934:)
930:(
922:)
918:(
894:(
879:(
830:(
772:(
746:(
361:(
246:(
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