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Common descent

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1599: 1521: 1652:, they might independently have acquired similar genetic sequences. Theobald's "formal test" was accordingly criticised by Takahiro Yonezawa and colleagues for not including consideration of convergence. They argued that Theobald's test was insufficient to distinguish between the competing hypotheses. Theobald has defended his method against this claim, arguing that his tests distinguish between phylogenetic structure and mere sequence similarity. Therefore, Theobald argued, his results show that "real universally 62: 4686: 617: 1689:. During the course of evolution, this RNA world was replaced by the evolutionary emergence of the DNA world. A world of independently self-replicating RNA genomes apparently no longer exists (RNA viruses are dependent on host cells with DNA genomes). Because the RNA world is apparently gone, it is not clear how scientific evidence could be brought to bear on the question of whether there was a single origin of life event from which all life descended. 4696: 1634:. This has led to questions about the single ancestry of life. However, biologists consider it very unlikely that completely unrelated proto-organisms could have exchanged genes, as their different coding mechanisms would have resulted only in garble rather than functioning systems. Later, however, many organisms all derived from a single ancestor could readily have shared genes that all worked in the same way, and it appears that they have. 1470:(DNA triplets) that code redundantly for the same amino acid. Since many species use the same codon at the same place to specify an amino acid that can be represented by more than one codon, that is evidence for their sharing a recent common ancestor. Had the amino acid sequences come from different ancestors, they would have been coded for by any of the redundant codons, and since the correct amino acids would already have been in place, 4722: 604: 862:"We do not know all the possible transitional gradations between the simplest and the most perfect organs; it cannot be pretended that we know all the varied means of Distribution during the long lapse of years, or that we know how imperfect the Geological Record is. Grave as these several difficulties are, in my judgment they do not overthrow the theory of descent from a few created forms with subsequent modification". 1681:, and that this implies a single origin for life. Although such a universal common ancestor may have existed, such a complex entity is unlikely to have arisen spontaneously from non-life and thus a cell with a DNA genome cannot reasonably be regarded as the origin of life. To understand the origin of life, it has been proposed that DNA based cellular life descended from relatively simple pre-cellular self-replicating 1629:
could have occurred during early evolution. Bacteria today remain capable of gene exchange between distantly-related lineages. This weakens the basic assumption of phylogenetic analysis, that similarity of genomes implies common ancestry, because sufficient gene exchange would allow lineages to share
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endued with animality, with the power of acquiring new parts attended with new propensities, directed by irritations, sensations, volitions, and associations; and thus possessing the faculty of continuing to improve by its own inherent activity, and of delivering down those improvements by generation
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There is grandeur in this view of life, with its several powers, having been originally breathed into a few forms or into one; and that, whilst this planet has gone cycling on according to the fixed law of gravity, from so simple a beginning endless forms most beautiful and most wonderful have been,
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throughout all living organisms, especially how the rotating elements are bound to the membrane. This supports the assumption of a LUCA as a cellular organism, although primordial membranes may have been semipermeable and evolved later to the membranes of modern bacteria, and on a second path to
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Another important piece of evidence is from detailed phylogenetic trees (i.e., "genealogic trees" of species) mapping out the proposed divisions and common ancestors of all living species. In 2010, Douglas L. Theobald published a statistical analysis of available genetic data, mapping them to
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No reliable observation has ever been found to contradict the general notion of common descent. It should come as no surprise, then, that the scientific community at large has accepted evolutionary descent as a historical reality since Darwin's time and considers it among the most reliably
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ould it be too bold to imagine, that in the great length of time, since the earth began to exist, perhaps millions of ages before the commencement of the history of mankind, would it be too bold to imagine, that all warm-blooded animals have arisen from one living filament, which
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could change the codons, but it would be extremely unlikely to make all the redundant codons in a whole sequence match exactly across multiple lineages. Similarly, shared nucleotide sequences, especially where these are apparently neutral such as the positioning of
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But he precedes that remark by, "Analogy would lead me one step further, namely, to the belief that all animals and plants have descended from some one prototype. But analogy may be a deceitful guide." And in the subsequent edition, he asserts rather,
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sequences and DNA sequences. Proteins with the same three-dimensional structure need not have identical amino acid sequences; any irrelevant similarity between the sequences is evidence for common descent. In certain cases, there are several
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applicable when one species is the ancestor of two or more species later in time. According to modern evolutionary biology, all living beings could be descendants of a unique ancestor commonly referred to as the
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The idea that all living things (including things considered non-living by science) are related is a recurring theme in many indigenous worldviews across the world. Later on, in the 1740s, the French
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Darwin, C. R. 1860. On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life. London: John Murray. 2nd edition, second issue, page 466
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Therefore I should infer from analogy that probably all the organic beings which have ever lived on this earth have descended from some one primordial form, into which life was first breathed.
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Biologists often point to the universality of many aspects of cellular life as supportive evidence to the more compelling evidence listed above. These similarities include the energy carrier
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has no influence over external morphology. That phylogenetic trees based on different types of information agree with each other is strong evidence of a real underlying common descent.
2865: 1507:- rather than through universal common descent - and therefore resulted in convergent evolution. In contrast, there is evidence for homology of the central subunits of 4418: 333: 2421: 2383:
Ohtomo, Yoko; Kakegawa, Takeshi; Ishida, Akizumi; et al. (January 2014). "Evidence for biogenic graphite in early Archaean Isua metasedimentary rocks".
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carrying out core functions like DNA replication. The fact that only one such set of enzymes exists is convincing evidence of a single ancestry. 6,331
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Weiss, Madeline C.; Sousa, Filipa L.; Mrnjavac, Natalia; Neukirchen, Sinje; Roettger, Mayo; Nelson-Sathi, Shijulal; Martin, William F. (2016-07-25).
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Knight, Robin; Freeland, Stephen J.; Landweber, Laura F. (January 2001). "Rewiring the keyboard: evolvability of the genetic code".
1537: 4671: 4126: 3237: 375: 1529: 1426:. The universality of this code is generally regarded by biologists as definitive evidence in favor of universal common descent. 3477: 1674:
The possibility is mentioned, above, that all living organisms may be descended from an original single-celled organism with a
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On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life
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All known forms of life are based on the same fundamental biochemical organization: genetic information encoded in
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commented that "practically no naturalists of position and recognized attainment doubt the theory of descent."
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arrived at the idea that all organisms had a common ancestor, and had diverged through random variation and
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and others as energy sources. Analysis of small sequence differences in widely shared substances such as
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phylogenetic trees, that gave "strong quantitative support, by a formal test, for the unity of life."
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further supports universal common descent. Some 23 proteins are found in all organisms, serving as
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Theobald, Douglas L. (13 May 2010). "A formal test of the theory of universal common ancestry".
819:) that the similarity of animal forms implies a common original type, and thus a common parent. 4656: 4305: 4174: 4136: 4037: 4008: 3981: 3976: 3574: 3268: 2897:"Primal Eukaryogenesis: On the Communal Nature of Precellular States, Ancestral to Modern Life" 2471: 1699: 1603: 1598: 1573: 1546: 1496: 927: 489: 444: 266: 161: 39: 24: 4162: 1545: 4597: 4471: 4390: 4385: 4370: 4355: 4345: 4261: 4236: 4071: 4027: 3991: 3959: 3900: 3878: 3859: 3722: 3675: 3626: 3621: 3579: 2955: 2574: 587: 469: 414: 380: 293: 2541: 1582:
Traditionally, these trees have been built using morphological methods, such as appearance,
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in 1859. Scientists accepted it because it explained so many facets of the living world."
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common to all living animals have been identified; these may have arisen from a single
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common to all living animals have been identified; these may have arisen from a single
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would not have driven any change in the codons, however much time was available.
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with water-repelling regions able to support the essential electron exchange (
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Evolution: Genesis and Revelations: With Readings from Empedocles to Wilson
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If early organisms had been driven by the same environmental conditions to
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seems to be strongly optimised. Richard Egel argues that in particular the
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Forterre, Patrick; Gribaldo, Simonetta; Brochier, CĂ©line (October 2005).
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29+ Evidences for Macroevolution: The Scientific Case for Common Descent
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29+ Evidences for Macroevolution: The Scientific Case for Common Descent
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Zoonomia; or The laws of organic life: in three parts (Volume 1) (1818)
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Similarities which have no adaptive relevance cannot be explained by
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established and fundamentally important facts in all of science.
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Glansdorff, Nicolas; Ying Xu; Labedan, Bernard (9 July 2008).
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Characteristic of a group of organisms with a common ancestor
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Yonezawa, Takahiro; Hasegawa, Masami (16 December 2010).
2532:"The Very First Animal Appeared Amid an Explosion of DNA" 2453: 1682: 1675: 911: 907: 723:. All currently living organisms on Earth share a common 3013:"Prediction 1.3: Consilience of independent phylogenies" 2329: 2082: 2818: 2753:
On the Origin of Species by Means of Natural Selection
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during early evolution has led to questions about the
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For use of the term in linguistics and philology, see
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(30 April 2018). 1452: 2206:"Origins of life: Common ancestry put to the test" 1922: 1831: 1728:The Rise of Great Britain and Russia, 1688–1715/25 4734: 3039: 2522: 2520: 1523: 866:Common descent was widely accepted amongst the 843:'s views about common descent, as expressed in 727:heritage, though the suggestion of substantial 2987:The Vital Question: Why Is Life The Way It Is? 2563: 2561: 2422:"Oldest fossil found: Meet your microbial mom" 1487:, provide strong evidence of common ancestry. 3431: 3231: 2663: 1718: 642: 2517: 765:in the concluding sentence of his 1859 book 3042:"Was the universal common ancestry proved?" 2567: 2558: 2447: 1562:genes implies a single origin for all life. 735:(single ancestry) of life. 6,331 groups of 3438: 3424: 3238: 3224: 2947: 2620: 2419: 2413: 2376: 2200: 2140: 2138: 2136: 2134: 2132: 2130: 2128: 1980:: CS1 maint: location missing publisher ( 1967:Der Begriff des Instinktes einst und jetzt 1802:The Complete Work of Charles Darwin Online 1615:Gene exchange clouds phylogenetic analysis 758:process was first proposed by the British 649: 635: 3173: 3163: 3146:Theobald, Douglas L. (24 November 2011). 3122: 3097:Theobald, Douglas L. (16 December 2010). 3065: 2930: 2920: 2814: 2812: 2800: 2603: 2500: 2359: 2349: 2260: 2229: 2100: 1630:much of their genome whether or not they 4672:Transgenerational epigenetic inheritance 3445: 3145: 3096: 2857: 2678: 2144: 1920: 1650:evolve similar biochemistry convergently 1597: 1519: 2782: 2196: 2194: 2125: 1963: 1637: 822:In 1794, Charles Darwin's grandfather, 4735: 2953: 2809: 2718: 2706: 2626: 2526: 1925:The Making of Modern Europe, 1648-1780 1829: 1807: 1768: 961: 901: 4435:Dialogues Concerning Natural Religion 3419: 3219: 2960:. Mittal Publications. pp. 32–. 2785:"Evolution as Fact, Theory, and Path" 2741:Biology: A Guide to the Natural World 2639:Staff, I. C. T. (13 September 2018). 2638: 2420:Borenstein, Seth (13 November 2013). 1814:Zoonomia; or the Laws of Organic Life 1610:, implying multiple possible origins. 1515: 1490: 870:after Darwin's publication. In 1907, 3010: 2980: 2894: 2888: 2863: 2690: 2191: 1872: 836:to its posterity, world without end? 754:Universal common descent through an 3302:Formamide-based prebiotic chemistry 2957:Continuity And Evolution Of Animals 2457:; Christian, Daniel; Wacey, David; 13: 3840:Evolutionary developmental biology 3245: 1840:State University of New York Press 14: 4764: 3307:Alternative abiogenesis scenarios 3194: 2789:Evolution: Education and Outreach 2743:. Pearson/Prentice Hall. p. 323. 2295:10.1038/scientificamerican0200-90 1449:) reactions for energy transfer. 4720: 4694: 4685: 4684: 1964:Ziegler, Heinrich Ernst (1904). 1732:The New Cambridge Modern History 1625:Theobald noted that substantial 1453:Selectively neutral similarities 616: 615: 602: 60: 4497:Extended evolutionary synthesis 3686:Gene-centered view of evolution 3139: 3090: 3033: 3004: 2974: 2776: 2758: 2733: 2724: 2712: 2700: 2684: 2672: 2657: 2632: 1997: 1711: 787:History of evolutionary thought 715:found in 3.48 billion-year-old 684:Common descent is an effect of 609:Evolutionary biology portal 4625:Hologenome theory of evolution 4492:History of molecular evolution 3718:Evolutionarily stable strategy 3607:Last universal common ancestor 3383:Last universal common ancestor 2773:. Henry Holt and Company. p. 3 2323: 2254: 2204:; Penny, David (13 May 2010). 2076: 2021: 1632:shared an ancestor (monophyly) 1512:those of modern archaea also. 671:last universal common ancestor 568:Creation–evolution controversy 322:History of evolutionary theory 1: 4419:Renaissance and Enlightenment 2014: 1593: 1505:laws of physics and chemistry 1433:(DNA triplets) are mapped to 4753:Most recent common ancestors 4630:Missing heritability problem 4257:Gamete differentiation/sexes 3211:The Tree of Life Web Project 2895:Egel, Richard (March 2012). 2266:"Uprooting the Tree of Life" 1663: 553:Evolution as fact and theory 7: 2046:10.1038/nmicrobiol.2016.116 1921:Treasure, Geoffrey (1985). 1800:The book is available from 1692: 890: 10: 4769: 4262:Life cycles/nuclear phases 3814:Trivers–Willard hypothesis 3021:The TalkOrigins Foundation 2596:10.1038/s41467-018-04136-5 2102:10.1051/medsci/20052110860 1888:Hackett Publishing Company 1736:Cambridge University Press 1685:molecules able to undergo 1667: 1641: 1618: 1571: 1565: 965: 897:Evidence of common descent 894: 784: 780: 588:Nature-nurture controversy 37: 18: 4680: 4580: 4505: 4409: 4336: 4292: 4147: 4051: 3868: 3827: 3760:Parent–offspring conflict 3696: 3565:Earliest known life forms 3486: 3453: 3392: 3378:Earliest known life forms 3370: 3315: 3284: 3253: 2864:Than, Ker (14 May 2010). 2802:10.1007/s12052-007-0001-z 2783:Gregory, T. Ryan (2008). 2664:Crombie & Hoskin 1970 1608:horizontal gene transfers 1374: 1344: 1334: 1326: 1318: 1312: 1286: 1278: 1248: 1240: 1231: 1223: 1217: 1193: 1163: 1155: 1147: 1138: 1087: 1063: 1055: 1047: 1037: 1031: 1026: 1023: 1020: 1017: 1012: 1007: 805:In 1790, the philosopher 475:Evolutionary neuroscience 450:Evolutionary epistemology 430:Evolutionary anthropology 410:Applications of evolution 4613:Cultural group selection 4477:The eclipse of Darwinism 4449:On the Origin of Species 4424:Transmutation of species 1990: 1830:Harris, C. Leon (1981). 1778:(1st ed.). London: 1627:horizontal gene transfer 1621:Horizontal gene transfer 914:, through the effect of 846:On the Origin of Species 768:On the Origin of Species 729:horizontal gene transfer 700:in 3.7 billion-year-old 465:Evolutionary linguistics 460:Evolutionary game theory 435:Evolutionary computation 4618:Dual inheritance theory 4457:History of paleontology 2822:Nature Reviews Genetics 1804:. Retrieved 2015-11-23. 1734:. Vol. 6. London: 811:Kritik der Urteilskraft 796:Pierre Louis Maupertuis 776:and are being, evolved. 578:Objections to evolution 485:Evolutionary psychology 480:Evolutionary physiology 425:Evolutionary aesthetics 404:Fields and applications 386:History of paleontology 4306:Punctuated equilibrium 3627:Non-adaptive radiation 3575:Evolutionary arms race 3297:Miller–Urey experiment 3269:Spontaneous generation 3165:10.1186/1745-6150-6-60 2954:Sharma, N. S. (2005). 2739:Krogh, David. (2005). 2351:10.1186/1745-6150-3-29 1611: 1574:Tree of life (biology) 1563: 1497:adenosine triphosphate 1003:Standard genetic code 888: 864: 855: 838: 778: 704:discovered in western 510:Speciation experiments 490:Experimental evolution 445:Evolutionary economics 267:Recent human evolution 125:Processes and outcomes 40:Lowest common ancestor 25:Historical linguistics 4598:Evolutionary medicine 4472:Mendelian inheritance 4180:Biological complexity 4168:Programmed cell death 3860:Phenotypic plasticity 3580:Evolutionary pressure 3570:Evidence of evolution 3468:Timeline of evolution 3011:Theobald, Douglas L. 2582:(1730 (2018)): 1730. 2575:Nature Communications 2493:10.1089/ast.2013.1030 1826:Retrieved 2015-11-23. 1642:Further information: 1619:Further information: 1601: 1553: 1509:transmembrane ATPases 966:Further information: 952:650 million years ago 895:Further information: 883: 860: 851: 833:the great First Cause 828: 773: 745:650 million years ago 702:metasedimentary rocks 690:3.9 billion years ago 470:Evolutionary medicine 415:Biosocial criminology 381:History of speciation 294:Evolutionary taxonomy 257:Timeline of evolution 4743:Evolutionary biology 4727:Evolutionary biology 4572:Teleology in biology 4467:Blending inheritance 3845:Genetic assimilation 3708:Artificial selection 3447:Evolutionary biology 2461:(16 December 2013). 1886:. Indianapolis, IN: 1879:Critique of Judgment 1644:Convergent evolution 1638:Convergent evolution 1459:convergent evolution 868:scientific community 816:Critique of Judgment 666:evolutionary biology 440:Evolutionary ecology 54:Evolutionary biology 4635:Molecular evolution 4593:Ecological genetics 4462:Transitional fossil 4252:Sexual reproduction 4092:endomembrane system 4021:pollinator-mediated 3977:dolphins and whales 3755:Parental investment 3340:Primordial sandwich 3285:Prebiotic synthesis 3255:History of research 3205:TalkOrigins Archive 3124:10.1038/nature09483 3115:2010Natur.468E..10T 3067:10.1038/nature09482 3058:2010Natur.468E...9Y 2922:10.3390/life2010170 2913:2012Life....2..170E 2870:National Geographic 2588:2018NatCo...9.1730P 2485:2013AsBio..13.1103N 2399:2014NatGe...7...25O 2287:2000SciAm.282b..90D 2274:Scientific American 2222:2010Natur.465..168S 2169:10.1038/nature09014 2161:2010Natur.465..219T 2034:Nature Microbiology 1700:The Ancestor's Tale 1004: 962:Common genetic code 910:, transcribed into 902:Common biochemistry 877:In 2008, biologist 542:Social implications 530:Universal Darwinism 520:Island biogeography 455:Evolutionary ethics 420:Ecological genetics 366:Molecular evolution 304:Transitional fossil 132:Population genetics 48:Part of a series on 4608:Cultural evolution 3723:Fisher's principle 3652:Handicap principle 3642:Parallel evolution 3506:Adaptive radiation 3371:Earliest organisms 3357:Quasispecies model 3325:GADV-protein world 2537:The New York Times 2262:Doolittle, W. Ford 1612: 1564: 1516:Phylogenetic trees 1491:Other similarities 1002: 573:Theistic evolution 505:Selective breeding 217:Parallel evolution 182:Adaptive radiation 21:Comparative method 4708: 4707: 4324:Uniformitarianism 4277:Sex-determination 3782:Sexual dimorphism 3777:Natural selection 3681:Unit of selection 3647:Signalling theory 3413: 3412: 3352:Peptide-RNA world 3335:Iron–sulfur world 3292:Pseudo-panspermia 2990:. Profile Books. 2967:978-81-8293-018-6 2765:Kellogg, Vernon L 2479:(12): 1103–1124. 2386:Nature Geoscience 2264:(February 2000). 2216:(7295): 168–169. 2155:(7295): 219–222. 2089:MĂ©decine/Sciences 1940:978-0-416-72370-0 1897:978-0-87220-025-8 1849:978-0-87395-487-7 1745:978-0-521-07524-4 1687:natural selection 1588:genetic variation 1568:Phylogenetic tree 1556:phylogenetic tree 1472:natural selection 1400: 1399: 1000: 999: 800:natural selection 721:Western Australia 659: 658: 350:Origin of Species 152:Natural selection 33:Textual criticism 4760: 4725: 4724: 4716: 4698: 4688: 4687: 4487:Modern synthesis 4247:Multicellularity 4242:Mosaic evolution 4127:auditory ossicle 3809:Social selection 3792:Flowering plants 3787:Sexual selection 3440: 3433: 3426: 3417: 3416: 3271:(4th century BC) 3265:(5th century BC) 3240: 3233: 3226: 3217: 3216: 3188: 3187: 3177: 3167: 3143: 3137: 3136: 3126: 3099:"Theobald reply" 3094: 3088: 3087: 3069: 3037: 3031: 3030: 3028: 3027: 3019:. Version 2.89. 3008: 3002: 3001: 2978: 2972: 2971: 2951: 2945: 2944: 2934: 2924: 2892: 2886: 2885: 2883: 2881: 2872:. Archived from 2861: 2855: 2854: 2835:10.1038/35047500 2816: 2807: 2806: 2804: 2780: 2774: 2770:Darwinism To-Day 2762: 2756: 2737: 2731: 2728: 2722: 2716: 2710: 2704: 2698: 2688: 2682: 2676: 2670: 2661: 2655: 2654: 2652: 2651: 2636: 2630: 2624: 2618: 2617: 2607: 2565: 2556: 2555: 2553: 2551: 2545: 2540:. Archived from 2524: 2515: 2514: 2504: 2459:Hazen, Robert M. 2451: 2445: 2444: 2442: 2441: 2432:. Archived from 2430:Associated Press 2417: 2411: 2410: 2407:10.1038/ngeo2025 2380: 2374: 2373: 2363: 2353: 2327: 2321: 2320: 2318: 2317: 2311: 2305:. Archived from 2270: 2258: 2252: 2251: 2233: 2198: 2189: 2188: 2142: 2123: 2122: 2104: 2080: 2074: 2073: 2025: 2008: 2001: 1985: 1979: 1971: 1960: 1928: 1917: 1869: 1837: 1824:Internet Archive 1818: 1799: 1765: 1522: 1005: 1001: 972: 971: 834: 664:is a concept in 651: 644: 637: 624: 619: 618: 611: 607: 606: 583:Level of support 376:Current research 361:Modern synthesis 356:Before synthesis 309:Extinction event 67:Darwin's finches 64: 45: 44: 4768: 4767: 4763: 4762: 4761: 4759: 4758: 4757: 4733: 4732: 4731: 4719: 4711: 4709: 4704: 4676: 4603:Group selection 4576: 4501: 4405: 4332: 4294:Tempo and modes 4288: 4143: 4047: 3864: 3823: 3699: 3692: 3669:Species complex 3482: 3473:History of life 3449: 3444: 3414: 3409: 3388: 3366: 3330:Clay hypothesis 3311: 3280: 3275:Primordial soup 3249: 3244: 3197: 3192: 3191: 3144: 3140: 3095: 3091: 3038: 3034: 3025: 3023: 3009: 3005: 2998: 2979: 2975: 2968: 2952: 2948: 2893: 2889: 2879: 2877: 2876:on May 15, 2010 2862: 2858: 2817: 2810: 2781: 2777: 2763: 2759: 2738: 2734: 2729: 2725: 2717: 2713: 2705: 2701: 2689: 2685: 2677: 2673: 2662: 2658: 2649: 2647: 2637: 2633: 2625: 2621: 2566: 2559: 2549: 2547: 2525: 2518: 2452: 2448: 2439: 2437: 2418: 2414: 2381: 2377: 2328: 2324: 2315: 2313: 2309: 2268: 2259: 2255: 2231:10.1038/465168a 2199: 2192: 2143: 2126: 2095:(10): 860–865. 2081: 2077: 2026: 2022: 2017: 2012: 2011: 2002: 1998: 1993: 1988: 1973: 1972: 1941: 1898: 1850: 1809:Darwin, Erasmus 1770:Darwin, Charles 1746: 1724:Hoskin, Michael 1714: 1695: 1672: 1666: 1646: 1640: 1623: 1617: 1596: 1576: 1570: 1552: 1520: 1518: 1493: 1455: 1377: 1337: 1040: 1009: 970: 964: 948:common ancestor 904: 899: 893: 879:T. Ryan Gregory 832: 789: 783: 741:common ancestor 655: 614: 601: 600: 593: 592: 543: 535: 534: 405: 397: 396: 395: 323: 315: 314: 313: 262:Human evolution 252:History of life 236: 235:Natural history 228: 227: 226: 126: 118: 73: 43: 36: 17: 12: 11: 5: 4766: 4756: 4755: 4750: 4745: 4730: 4729: 4706: 4705: 4703: 4702: 4692: 4681: 4678: 4677: 4675: 4674: 4669: 4664: 4659: 4654: 4653: 4652: 4642: 4637: 4632: 4627: 4622: 4621: 4620: 4615: 4610: 4600: 4595: 4590: 4584: 4582: 4578: 4577: 4575: 4574: 4569: 4568: 4567: 4562: 4557: 4556: 4555: 4545: 4540: 4535: 4530: 4525: 4515: 4509: 4507: 4503: 4502: 4500: 4499: 4494: 4489: 4484: 4479: 4474: 4469: 4464: 4459: 4454: 4453: 4452: 4443:Charles Darwin 4440: 4439: 4438: 4426: 4421: 4415: 4413: 4407: 4406: 4404: 4403: 4398: 4393: 4388: 4383: 4381:Non-ecological 4378: 4373: 4368: 4363: 4358: 4353: 4348: 4342: 4340: 4334: 4333: 4331: 4330: 4321: 4312: 4298: 4296: 4290: 4289: 4287: 4286: 4281: 4280: 4279: 4274: 4269: 4264: 4259: 4249: 4244: 4239: 4234: 4229: 4224: 4219: 4214: 4209: 4204: 4199: 4198: 4197: 4187: 4182: 4177: 4172: 4171: 4170: 4165: 4154: 4152: 4145: 4144: 4142: 4141: 4140: 4139: 4134: 4132:nervous system 4129: 4124: 4119: 4111: 4110: 4109: 4104: 4099: 4094: 4089: 4084: 4074: 4069: 4064: 4058: 4056: 4049: 4048: 4046: 4045: 4040: 4035: 4030: 4025: 4024: 4023: 4013: 4012: 4011: 4006: 4005: 4004: 3999: 3989: 3984: 3979: 3974: 3969: 3968: 3967: 3962: 3952: 3942: 3937: 3936: 3935: 3925: 3920: 3915: 3910: 3909: 3908: 3898: 3893: 3892: 3891: 3881: 3875: 3873: 3866: 3865: 3863: 3862: 3857: 3852: 3847: 3842: 3837: 3831: 3829: 3825: 3824: 3822: 3821: 3816: 3811: 3806: 3805: 3804: 3799: 3794: 3784: 3779: 3774: 3769: 3764: 3763: 3762: 3757: 3747: 3742: 3737: 3736: 3735: 3725: 3720: 3715: 3710: 3704: 3702: 3694: 3693: 3691: 3690: 3689: 3688: 3678: 3673: 3672: 3671: 3666: 3656: 3655: 3654: 3644: 3639: 3634: 3632:Origin of life 3629: 3624: 3619: 3617:Microevolution 3614: 3612:Macroevolution 3609: 3604: 3599: 3598: 3597: 3587: 3582: 3577: 3572: 3567: 3562: 3557: 3552: 3550:Common descent 3547: 3546: 3545: 3535: 3530: 3528:Baldwin effect 3525: 3524: 3523: 3518: 3508: 3503: 3498: 3492: 3490: 3484: 3483: 3481: 3480: 3475: 3470: 3465: 3460: 3454: 3451: 3450: 3443: 3442: 3435: 3428: 3420: 3411: 3410: 3408: 3407: 3402: 3396: 3394: 3390: 3389: 3387: 3386: 3380: 3374: 3372: 3368: 3367: 3365: 3364: 3359: 3354: 3349: 3344: 3343: 3342: 3332: 3327: 3321: 3319: 3313: 3312: 3310: 3309: 3304: 3299: 3294: 3288: 3286: 3282: 3281: 3279: 3278: 3277:(19th century) 3272: 3266: 3259: 3257: 3251: 3250: 3247:Origin of life 3243: 3242: 3235: 3228: 3220: 3214: 3213: 3208: 3196: 3195:External links 3193: 3190: 3189: 3152:Biology Direct 3138: 3089: 3032: 3003: 2997:978-1781250365 2996: 2973: 2966: 2946: 2907:(1): 170–212. 2887: 2856: 2808: 2775: 2757: 2749:978-0321946768 2732: 2723: 2711: 2709:, p. 397 2699: 2683: 2671: 2656: 2631: 2619: 2557: 2530:(4 May 2018). 2516: 2446: 2412: 2375: 2337:Biology Direct 2322: 2253: 2190: 2124: 2075: 2019: 2018: 2016: 2013: 2010: 2009: 1995: 1994: 1992: 1989: 1987: 1986: 1961: 1939: 1918: 1896: 1884:Mary J. Gregor 1874:Kant, Immanuel 1870: 1848: 1838:. Albany, NY: 1827: 1805: 1766: 1744: 1720:Crombie, A. C. 1715: 1713: 1710: 1709: 1708: 1703: 1694: 1691: 1668:Main article: 1665: 1662: 1639: 1636: 1616: 1613: 1595: 1592: 1566:Main article: 1517: 1514: 1492: 1489: 1454: 1451: 1398: 1397: 1394: 1391: 1388: 1384: 1383: 1380: 1373: 1370: 1367: 1363: 1362: 1359: 1356: 1353: 1349: 1348: 1343: 1340: 1333: 1330: 1325: 1322: 1317: 1314: 1310: 1309: 1306: 1303: 1300: 1295: 1291: 1290: 1285: 1282: 1277: 1274: 1271: 1267: 1266: 1263: 1260: 1257: 1253: 1252: 1247: 1244: 1239: 1236: 1230: 1227: 1222: 1219: 1215: 1214: 1211: 1208: 1205: 1201: 1200: 1197: 1192: 1189: 1186: 1182: 1181: 1178: 1175: 1172: 1168: 1167: 1162: 1159: 1154: 1151: 1146: 1143: 1140: 1136: 1135: 1129: 1126: 1121: 1118: 1115: 1111: 1110: 1105: 1102: 1097: 1094: 1091: 1086: 1082: 1081: 1078: 1075: 1072: 1068: 1067: 1062: 1059: 1054: 1051: 1046: 1043: 1036: 1033: 1029: 1028: 1025: 1022: 1019: 1015: 1014: 1011: 998: 997: 994: 992: 989: 986: 983: 980: 963: 960: 903: 900: 892: 889: 872:Vernon Kellogg 841:Charles Darwin 824:Erasmus Darwin 782: 779: 763:Charles Darwin 719:discovered in 673:(LUCA) of all 662:Common descent 657: 656: 654: 653: 646: 639: 631: 628: 627: 626: 625: 612: 595: 594: 591: 590: 585: 580: 575: 570: 565: 563:Social effects 560: 555: 550: 544: 541: 540: 537: 536: 533: 532: 527: 522: 517: 512: 507: 502: 497: 492: 487: 482: 477: 472: 467: 462: 457: 452: 447: 442: 437: 432: 427: 422: 417: 412: 406: 403: 402: 399: 398: 394: 393: 383: 378: 373: 368: 363: 358: 353: 346: 341: 336: 331: 325: 324: 321: 320: 317: 316: 312: 311: 306: 301: 296: 291: 289:Classification 286: 281: 276: 271: 270: 269: 259: 254: 249: 247:Common descent 244: 242:Origin of life 238: 237: 234: 233: 230: 229: 225: 224: 219: 214: 209: 204: 199: 194: 189: 184: 179: 174: 169: 164: 159: 154: 149: 144: 139: 134: 128: 127: 124: 123: 120: 119: 117: 116: 111: 106: 100: 99: 94: 89: 84: 78: 75: 74: 65: 57: 56: 50: 49: 29:Proto-language 15: 9: 6: 4: 3: 2: 4765: 4754: 4751: 4749: 4746: 4744: 4741: 4740: 4738: 4728: 4723: 4718: 4717: 4714: 4701: 4697: 4693: 4691: 4683: 4682: 4679: 4673: 4670: 4668: 4665: 4663: 4660: 4658: 4655: 4651: 4648: 4647: 4646: 4645:Phylogenetics 4643: 4641: 4638: 4636: 4633: 4631: 4628: 4626: 4623: 4619: 4616: 4614: 4611: 4609: 4606: 4605: 4604: 4601: 4599: 4596: 4594: 4591: 4589: 4586: 4585: 4583: 4579: 4573: 4570: 4566: 4563: 4561: 4558: 4554: 4551: 4550: 4549: 4548:Structuralism 4546: 4544: 4541: 4539: 4536: 4534: 4531: 4529: 4526: 4524: 4523:Catastrophism 4521: 4520: 4519: 4516: 4514: 4511: 4510: 4508: 4504: 4498: 4495: 4493: 4490: 4488: 4485: 4483: 4482:Neo-Darwinism 4480: 4478: 4475: 4473: 4470: 4468: 4465: 4463: 4460: 4458: 4455: 4451: 4450: 4446: 4445: 4444: 4441: 4437: 4436: 4432: 4431: 4430: 4427: 4425: 4422: 4420: 4417: 4416: 4414: 4412: 4408: 4402: 4399: 4397: 4396:Reinforcement 4394: 4392: 4389: 4387: 4384: 4382: 4379: 4377: 4374: 4372: 4369: 4367: 4364: 4362: 4359: 4357: 4354: 4352: 4349: 4347: 4344: 4343: 4341: 4339: 4335: 4329: 4328:Catastrophism 4325: 4322: 4320: 4319:Macromutation 4316: 4315:Micromutation 4313: 4311: 4307: 4303: 4300: 4299: 4297: 4295: 4291: 4285: 4282: 4278: 4275: 4273: 4270: 4268: 4265: 4263: 4260: 4258: 4255: 4254: 4253: 4250: 4248: 4245: 4243: 4240: 4238: 4235: 4233: 4230: 4228: 4225: 4223: 4222:Immune system 4220: 4218: 4215: 4213: 4210: 4208: 4205: 4203: 4200: 4196: 4193: 4192: 4191: 4188: 4186: 4183: 4181: 4178: 4176: 4173: 4169: 4166: 4164: 4161: 4160: 4159: 4156: 4155: 4153: 4151: 4146: 4138: 4135: 4133: 4130: 4128: 4125: 4123: 4120: 4118: 4115: 4114: 4112: 4108: 4105: 4103: 4100: 4098: 4095: 4093: 4090: 4088: 4085: 4083: 4082:symbiogenesis 4080: 4079: 4078: 4075: 4073: 4070: 4068: 4065: 4063: 4060: 4059: 4057: 4055: 4050: 4044: 4041: 4039: 4036: 4034: 4031: 4029: 4026: 4022: 4019: 4018: 4017: 4014: 4010: 4007: 4003: 4000: 3998: 3995: 3994: 3993: 3990: 3988: 3985: 3983: 3980: 3978: 3975: 3973: 3970: 3966: 3963: 3961: 3958: 3957: 3956: 3953: 3951: 3948: 3947: 3946: 3943: 3941: 3938: 3934: 3931: 3930: 3929: 3926: 3924: 3921: 3919: 3916: 3914: 3911: 3907: 3904: 3903: 3902: 3899: 3897: 3894: 3890: 3887: 3886: 3885: 3882: 3880: 3877: 3876: 3874: 3872: 3867: 3861: 3858: 3856: 3853: 3851: 3848: 3846: 3843: 3841: 3838: 3836: 3833: 3832: 3830: 3826: 3820: 3817: 3815: 3812: 3810: 3807: 3803: 3800: 3798: 3795: 3793: 3790: 3789: 3788: 3785: 3783: 3780: 3778: 3775: 3773: 3770: 3768: 3765: 3761: 3758: 3756: 3753: 3752: 3751: 3750:Kin selection 3748: 3746: 3745:Genetic drift 3743: 3741: 3738: 3734: 3731: 3730: 3729: 3726: 3724: 3721: 3719: 3716: 3714: 3711: 3709: 3706: 3705: 3703: 3701: 3695: 3687: 3684: 3683: 3682: 3679: 3677: 3674: 3670: 3667: 3665: 3662: 3661: 3660: 3657: 3653: 3650: 3649: 3648: 3645: 3643: 3640: 3638: 3635: 3633: 3630: 3628: 3625: 3623: 3620: 3618: 3615: 3613: 3610: 3608: 3605: 3603: 3600: 3596: 3593: 3592: 3591: 3588: 3586: 3583: 3581: 3578: 3576: 3573: 3571: 3568: 3566: 3563: 3561: 3558: 3556: 3553: 3551: 3548: 3544: 3541: 3540: 3539: 3536: 3534: 3531: 3529: 3526: 3522: 3519: 3517: 3514: 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2891: 2875: 2871: 2867: 2860: 2852: 2848: 2844: 2840: 2836: 2832: 2828: 2824: 2823: 2815: 2813: 2803: 2798: 2794: 2790: 2786: 2779: 2772: 2771: 2766: 2761: 2754: 2750: 2746: 2742: 2736: 2727: 2721:, p. 484 2720: 2715: 2708: 2703: 2696: 2692: 2687: 2681:, p. 142 2680: 2679:Treasure 1985 2675: 2669: 2665: 2660: 2646: 2642: 2635: 2629:, p. 490 2628: 2623: 2615: 2611: 2606: 2601: 2597: 2593: 2589: 2585: 2581: 2577: 2576: 2571: 2564: 2562: 2546:on 2022-01-01 2544: 2539: 2538: 2533: 2529: 2523: 2521: 2512: 2508: 2503: 2498: 2494: 2490: 2486: 2482: 2478: 2474: 2473: 2468: 2466: 2460: 2456: 2450: 2436:on 2019-11-10 2435: 2431: 2427: 2423: 2416: 2408: 2404: 2400: 2396: 2392: 2388: 2387: 2379: 2371: 2367: 2362: 2357: 2352: 2347: 2343: 2339: 2338: 2333: 2326: 2312:on 2006-09-07 2308: 2304: 2300: 2296: 2292: 2288: 2284: 2280: 2276: 2275: 2267: 2263: 2257: 2249: 2245: 2241: 2237: 2232: 2227: 2223: 2219: 2215: 2211: 2207: 2203: 2197: 2195: 2186: 2182: 2178: 2174: 2170: 2166: 2162: 2158: 2154: 2150: 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(1907). 2693:, p.  2650:2021-05-05 2440:2015-11-22 2426:Yahoo News 2316:2015-11-22 2015:References 1706:Urmetazoan 1658:homologous 1594:Objections 1584:embryology 1572:See also: 1463:amino acid 1298:Methionine 1242:Asparagine 1225:Isoleucine 1132:Tryptophan 918:- and RNA- 785:See also: 760:naturalist 686:speciation 299:Cladistics 222:Extinction 207:Divergence 177:Speciation 157:Adaptation 71:John Gould 4662:Protocell 4513:Darwinism 4401:Sympatric 4150:processes 4038:Tetrapods 3987:Kangaroos 3913:Dinosaurs 3850:Inversion 3819:Variation 3740:Gene flow 3733:Inclusive 3543:Mutualism 3488:Evolution 3362:RNA world 3347:PAH world 3203:from the 3158:(1): 60. 2795:: 46–52. 2691:Kant 1987 2248:205055573 2111:0767-0974 2054:2058-5276 1976:cite book 1876:(1987) . 1811:(1818) . 1796:741260650 1670:RNA world 1664:RNA world 1654:conserved 1558:based on 1233:Threonine 1195:Glutamine 1157:Histidine 1013:2nd base 982:nonpolar 924:ribosomes 809:wrote in 733:monophyly 717:sandstone 706:Greenland 558:Dysgenics 274:Phylogeny 172:Gene flow 142:Diversity 137:Variation 4690:Category 4565:Vitalism 4560:Theistic 4553:Spandrel 4237:Morality 4232:Monogamy 4107:plastids 4072:Flagella 4028:Reptiles 4009:sea cows 3992:primates 3901:Molluscs 3879:Bacteria 3767:Mutation 3700:genetics 3676:Taxonomy 3622:Mismatch 3602:Homology 3516:Cheating 3511:Altruism 3393:Research 3184:22114984 3076:21164432 2984:(2015). 2941:25382122 2851:12267003 2843:11253070 2645:Ict News 2614:29712911 2511:24205812 2370:18613974 2303:10710791 2240:20463725 2177:20463738 2119:16197904 2062:27562259 2005:homology 1957:11623262 1949:85000255 1914:13796153 1906:86014852 1858:81002555 1788:06017473 1772:(1859). 1754:57014935 1693:See also 1443:peptides 1412:bacteria 1376:Glutamic 1336:Aspartic 1288:Arginine 1165:Arginine 1065:Cysteine 1057:Tyrosine 891:Evidence 698:biogenic 694:graphite 622:Category 548:Eugenics 390:timeline 371:Evo-devo 329:Overview 147:Mutation 109:Evidence 104:Glossary 4748:Descent 4581:Related 4411:History 4272:Meiosis 4207:Empathy 4202:Emotion 4102:nucleus 4043:Viruses 4033:Spiders 3945:Mammals 3928:Insects 3728:Fitness 3664:Species 3463:Outline 3175:3314578 3133:4317014 3111:Bibcode 3084:4318346 3054:Bibcode 2932:4187143 2909:Bibcode 2605:5928047 2584:Bibcode 2502:3870916 2481:Bibcode 2395:Bibcode 2361:2478661 2283:Bibcode 2218:Bibcode 2185:4422345 2157:Bibcode 2070:2997255 1970:. Jena. 1931:Methuen 1866:7278190 1762:7588392 1481:introns 1420:animals 1416:archaea 1346:Glycine 1328:Alanine 1235:  1149:Proline 1134:  1089:Leucine 1041:alanine 1039:Phenyl- 991:acidic 954:in the 940:enzymes 926:, with 920:enzymes 916:protein 826:asked: 781:History 747:in the 725:genetic 713:fossils 114:History 97:Outline 4713:Portal 4700:Portal 4376:Hybrid 4212:Ethics 4054:organs 4016:Plants 4002:lemurs 3997:humans 3982:horses 3972:hyenas 3960:wolves 3955:canids 3889:origin 3385:(LUCA) 3182:  3172:  3131:  3103:Nature 3082:  3074:  3046:Nature 2994:  2964:  2939:  2929:  2849:  2841:  2747:  2612:  2602:  2509:  2499:  2368:  2358:  2344:: 29. 2301:  2246:  2238:  2210:Nature 2183:  2175:  2148:Nature 2117:  2109:  2068:  2060:  2052:  1955:  1947:  1937:  1912:  1904:  1894:  1864:  1856:  1846:  1794:  1786:  1760:  1752:  1742:  1679:genome 1468:codons 1431:codons 1424:plants 1320:Valine 1280:Lysine 1250:Serine 1049:Serine 988:basic 985:polar 620:  344:Darwin 31:, and 4163:Death 4158:Aging 4137:brain 3923:Fungi 3884:Birds 3797:Fungi 3595:Event 3478:Index 3129:S2CID 3080:S2CID 2847:S2CID 2668:62–63 2550:4 May 2310:(PDF) 2269:(PDF) 2244:S2CID 2181:S2CID 2066:S2CID 1991:Notes 1602:2005 1447:redox 1010:base 944:genes 932:NADPH 737:genes 679:Earth 82:Index 4650:Tree 4122:hair 4062:Cell 3965:dogs 3950:cats 3940:Life 3918:Fish 3871:taxa 3180:PMID 3072:PMID 2992:ISBN 2962:ISBN 2937:PMID 2901:Life 2882:2017 2839:PMID 2745:ISBN 2610:PMID 2552:2018 2507:PMID 2366:PMID 2299:PMID 2236:PMID 2173:PMID 2115:PMID 2107:ISSN 2058:PMID 2050:ISSN 1982:link 1953:OCLC 1945:LCCN 1935:ISBN 1910:OCLC 1902:LCCN 1892:ISBN 1862:OCLC 1854:LCCN 1844:ISBN 1792:OCLC 1784:LCCN 1758:OCLC 1750:LCCN 1740:ISBN 1483:and 1422:and 1414:and 1402:The 1396:GGG 1393:GAG 1390:GCG 1387:GTG 1382:GGA 1378:acid 1372:GAA 1369:GCA 1366:GTA 1361:GGC 1358:GAC 1355:GCC 1352:GTC 1342:GGT 1338:acid 1332:GAT 1324:GCT 1316:GTT 1308:AGG 1305:AAG 1302:ACG 1294:ATG 1284:AGA 1276:AAA 1273:ACA 1270:ATA 1265:AGC 1262:AAC 1259:ACC 1256:ATC 1246:AGT 1238:AAT 1229:ACT 1221:ATT 1213:CGG 1210:CAG 1207:CCG 1204:CTG 1199:CGA 1191:CAA 1188:CCA 1185:CTA 1180:CGC 1177:CAC 1174:CCC 1171:CTC 1161:CGT 1153:CAT 1145:CCT 1142:CTT 1128:TGG 1124:Stop 1120:TAG 1117:TCG 1114:TTG 1108:Stop 1104:TGA 1100:Stop 1096:TAA 1093:TCA 1085:TTA 1080:TGC 1077:TAC 1074:TCC 1071:TTC 1061:TGT 1053:TAT 1045:TCT 1035:TTT 708:and 675:life 92:Main 4148:Of 4117:eye 4067:DNA 4052:Of 3869:Of 3170:PMC 3160:doi 3119:doi 3107:468 3062:doi 3050:468 2927:PMC 2917:doi 2831:doi 2797:doi 2695:304 2600:PMC 2592:doi 2497:PMC 2489:doi 2403:doi 2356:PMC 2346:doi 2291:doi 2279:282 2226:doi 2214:465 2165:doi 2153:465 2097:doi 2042:doi 1683:RNA 1676:DNA 1660:." 1418:to 1008:1st 928:ATP 912:RNA 908:DNA 677:on 69:by 4739:: 3178:. 3168:. 3154:. 3150:. 3127:. 3117:. 3105:. 3101:. 3078:. 3070:. 3060:. 3048:. 3044:. 3015:. 2935:. 2925:. 2915:. 2903:. 2899:. 2868:. 2845:. 2837:. 2825:. 2811:^ 2791:. 2787:. 2643:. 2608:. 2598:. 2590:. 2578:. 2572:. 2560:^ 2534:. 2519:^ 2505:. 2495:. 2487:. 2477:13 2475:. 2469:. 2465:ca 2428:. 2424:. 2401:. 2389:. 2364:. 2354:. 2340:. 2334:. 2297:. 2289:. 2277:. 2271:. 2242:. 2234:. 2224:. 2212:. 2208:. 2193:^ 2179:. 2171:. 2163:. 2151:. 2127:^ 2113:. 2105:. 2093:21 2091:. 2087:. 2064:. 2056:. 2048:. 2036:. 2032:. 1978:}} 1974:{{ 1951:. 1943:. 1933:. 1908:. 1900:. 1890:. 1860:. 1852:. 1842:. 1790:. 1782:. 1756:. 1748:. 1738:. 1730:. 1722:; 1554:A 1313:G 1218:A 1139:C 1032:T 1027:G 1024:A 1021:C 1018:T 958:. 930:, 802:. 771:: 751:. 681:. 27:, 23:, 4715:: 4326:/ 4317:/ 4308:/ 4304:/ 3439:e 3432:t 3425:v 3239:e 3232:t 3225:v 3207:. 3186:. 3162:: 3156:6 3135:. 3121:: 3113:: 3086:. 3064:: 3056:: 3029:. 3000:. 2970:. 2943:. 2919:: 2911:: 2905:2 2884:. 2853:. 2833:: 2827:2 2805:. 2799:: 2793:1 2653:. 2616:. 2594:: 2586:: 2580:9 2554:. 2513:. 2491:: 2483:: 2443:. 2409:. 2405:: 2397:: 2391:7 2372:. 2348:: 2342:3 2319:. 2293:: 2285:: 2250:. 2228:: 2220:: 2187:. 2167:: 2159:: 2121:. 2099:: 2072:. 2044:: 2038:1 2007:. 1984:) 1959:. 1916:. 1868:. 1798:. 1764:. 813:( 650:e 643:t 636:v 392:) 388:( 42:. 35:.

Index

Comparative method
Historical linguistics
Proto-language
Textual criticism
Lowest common ancestor
Evolutionary biology

Darwin's finches
John Gould
Index
Introduction
Main
Outline
Glossary
Evidence
History
Population genetics
Variation
Diversity
Mutation
Natural selection
Adaptation
Polymorphism
Genetic drift
Gene flow
Speciation
Adaptive radiation
Co-operation
Coevolution
Coextinction

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