882:
31:
1222:
55:
936:
1289:
980:
1014:
1158:
342:
are small, stubby birds. Most have moderately long to long tails, while the stubtails and tesias have tiny tails that do not even emerge past their tail coverts. The group is typically clad in dull plumage, often with a line above the eye, but some are more colorful. Altogether the
Cettiidae are a
365:
The following cladogram showing the family relationships is based on a study by Carl
Oliveros and coworkers published in 2019. The number of species is taken from the bird list maintained by
287:
warblers, are mostly found in the forests of south and southeastern Asia, with one species of stubtail reaching as far north as Japan and
Siberia. Only one species,
1590:"Non-monophyly and intricate morphological evolution within the avian family Cettiidae revealed by multilocus analysis of a taxonomically densely sampled dataset"
2224:
2163:
343:
quite variable group containing many aberrant birds that hitherto had been uncomfortably placed with a wide range of unrelated families. The
2111:
2219:
2137:
1673:
374:
1440:
2214:
2142:
2229:
1701:
1404:; Ericson, P.G.P.; Olsson, U.; Sundberg, P. (2006). "Phylogeny and classification of the avian superfamily Sylvioidea".
1653:
Sefc, K.M.; Payne, R.B. & Sorenson, M.D. (2003): Phylogenetic relationships of
African sunbird-like warblers: Moho
881:
595:
The phylogenetic relationships between the eight genera were determined in a 2011 study by Per Alström and coworkers.
1647:
2168:
1637:
2150:
1846:
1243:
1456:
Irestedt, M.; Gelang, M.; Sangster, G.; Olsson, U.; Ericson, P.G.P.; Alström, P. (2011). "Neumann's
Warbler
2064:
2059:
1934:
1132:
1884:
1554:
1112:
366:
54:
1879:
1082:
347:, which contain birds which appear very similar to many cettiids, are far more uniform by contrast.
1990:
1957:
1359:
1233:
964:
332:
1914:
1904:
1856:
1694:
1263:
1052:
1032:
30:
2209:
2012:
1894:
1818:
1142:
920:
2181:
1929:
1899:
1874:
1851:
1679:
1253:
1122:
1102:
1042:
324:
350:
Most live in scrubland and frequently hunt food by clambering through thick tangled growth.
2085:
1985:
1813:
1369:
998:
900:
288:
8:
2050:
1952:
1808:
1221:
1206:
1176:
954:
910:
2204:
1924:
1919:
1909:
1841:
1687:
1616:
1589:
1563:"Cupwings, crombecs, bush warblers, Streaked Scrub Warbler, yellow flycatchers, hylias"
1528:
1501:
1273:
1186:
1072:
1062:
328:
49:
38:
2176:
2072:
1889:
1643:
1533:
1473:
1436:
1092:
244:
Some taxonomic authorities include this entire family, along with the related genera
220:
1641:(Volume 11: Old World Flycatchers to Old World Warblers). Lynx Edicions, Barcelona.
1980:
1611:
1601:
1558:
1523:
1513:
1469:
1413:
1327:
370:
205:
2077:
478:
2155:
1417:
1995:
1975:
1562:
1506:
Proceedings of the
National Academy of Sciences of the United States of America
1317:
1307:
409:
320:
316:
224:
1401:
359:
131:
2198:
2035:
1606:
501:
344:
257:
246:
209:
1518:
1944:
1836:
1778:
1537:
1460:(Sylvioidea): the sole African member of a Palaeotropic Miocene avifauna".
1349:
942:
768:
455:
228:
155:
935:
331:
are partly migratory over much of their range. A few species, such as the
2124:
2044:
1196:
1288:
2116:
1799:
1786:
1726:
888:
743:
283:
252:
149:
116:
2129:
1635:
del Hoyo, Josep; Elliott, Andrew & Sargatal, Jordi (eds.) (2006):
1588:
Alström, P.; Höhna, S.; Gelang, M.; Ericson, P.G.; Olsson, U. (2011).
979:
2098:
1967:
1770:
1738:
1344:
1295:
1013:
986:
793:
648:
630:
277:
191:
185:
161:
106:
66:
2006:
1157:
2103:
2029:
1866:
1754:
1020:
811:
432:
303:
216:
212:
167:
86:
2090:
1430:
339:
268:
1828:
1762:
1228:
700:
297:
272:
233:
179:
140:
76:
1709:
1746:
1164:
682:
312:
173:
1455:
1400:
315:. Most of the species in the family are sedentary, but the
308:
275:. The pseudo-tailorbirds, tesias and stubtails, as well as
264:
96:
1587:
1553:
2196:
1502:"Earth history and the passerine superradiation"
863:The family contains 31 species in eight genera.
1431:Winkler D.; Billerman, S.; Lovette I. (2015).
301:extends west to Western Europe, while that of
1695:
295:), occurs in Africa. The range of the genus
2225:Taxa named by Per Sundberg (ornithologist)
1702:
1688:
237:) and their relatives. As a common name,
29:
1615:
1605:
1527:
1517:
1396:
1394:
307:extends well into the Pacific, as far as
1499:
520:– bush warblers and allies (32 species)
1493:
358:The family Cettiidae was introduced by
134:, Ericson, Olsson, & Sundberg, 2006
2197:
1391:
375:International Ornithological Committee
227:" assemblage. It contains the typical
2011:
2010:
1683:
1569:. International Ornithologists' Union
1549:
1547:
1406:Molecular Phylogenetics and Evolution
1500:Oliveros, C.H.; et al. (2019).
1766:– typical bush warblers (4 species)
1435:. Lynx Editions. pp. 442–444.
373:and David Donsker on behalf of the
13:
1544:
481:– bristle flycatchers (3 species)
14:
2241:
1710:Genera and members of the family
1567:IOC World Bird List Version 12.2
1474:10.1111/j.1474-919X.2010.01084.x
1287:
1220:
1156:
1012:
978:
934:
880:
53:
2220:Taxa named by Per G.P. Ericson
1638:Handbook of Birds of the World
1581:
1480:
1449:
1424:
1:
1847:Chestnut-crowned bush warbler
1384:
1244:Chestnut-crowned bush warbler
815:– bush warblers (13 species)
412:– leaf warblers (81 species)
335:, are altitudinal migrants.
263:Its members occur mainly in
7:
1935:Yellow-bellied bush warbler
1418:10.1016/j.ympev.2005.05.015
1133:Yellow-bellied bush warbler
353:
219:"), formerly placed in the
10:
2246:
2230:Taxa named by Urban Olsson
1885:Brown-flanked bush warbler
1628:
1433:Bird Families of the World
1113:Brown-flanked bush warbler
772:– tailorbirds (2 species)
2215:Taxa named by Per Alström
2019:
1966:
1943:
1880:Bougainville bush warbler
1865:
1827:
1797:
1718:
1083:Bougainville bush warbler
969:Phyllergates heterolaemus
808:
790:
783:
765:
758:
740:
733:
697:
679:
672:
645:
627:
620:
613:
606:
515:
504:– streaked scrub warbler
499:
492:
476:
469:
453:
446:
430:
423:
407:
400:
393:
385:
146:
139:
50:Scientific classification
48:
37:
28:
23:
1991:Pale-footed bush warbler
1958:Rufous-headed tailorbird
1790:- streaked scrub warbler
1607:10.1186/1471-2148-11-352
1594:BMC Evolutionary Biology
1360:Pale-footed bush warbler
965:Rufous-headed tailorbird
925:Abroscopus superciliaris
652:– stubtails (3 species)
458:– bushtits (13 species)
333:pale-footed bush warbler
256:, in an enlarged family
1915:Philippine bush warbler
1905:Manchurian bush warbler
1857:Grey-sided bush warbler
1742:– stubtails (5 species)
1519:10.1073/pnas.1813206116
1264:Grey-sided bush warbler
1258:Cettia castaneocoronata
1238:– typical bush warblers
1053:Philippine bush warbler
1033:Manchurian bush warbler
959:Phyllergates cucullatus
797:– Broad-billed warbler
747:– warblers (3 species)
362:and coworkers in 2006.
1819:Yellow-bellied warbler
1774:– broad-billed warbler
1561:, eds. (August 2022).
1137:Horornis acanthizoides
993:– broad-billed warbler
921:Yellow-bellied warbler
915:Abroscopus schisticeps
905:Abroscopus albogularis
1930:Tanimbar bush warbler
1900:Japanese bush warbler
1875:Aberrant bush warbler
1852:Chestnut-headed tesia
1312:Urosphena squameiceps
1254:Chestnut-headed tesia
1127:Horornis flavolivacea
1123:Aberrant bush warbler
1103:Tanimbar bush warbler
1043:Japanese bush warbler
686:– tesias (4 species)
435:– hylias (2 species)
325:Japanese bush warbler
204:is a newly validated
1814:Rufous-faced warbler
1750:– tesias (5 species)
1734:warblers (3 species)
1364:Hemitesia pallidipes
1332:Urosphena whiteheadi
1147:Horornis brunnescens
1097:Horornis ruficapilla
999:Broad-billed warbler
901:Rufous-faced warbler
1953:Mountain tailorbird
1895:Hume's bush warbler
1809:Black-faced warbler
1663:Pholidornis rushiae
1655:Hypergerus atriceps
1207:Russet-capped tesia
1201:Tesia superciliaris
1177:Slaty-bellied tesia
1143:Hume's bush warbler
1037:Horornis canturians
955:Mountain tailorbird
911:Black-faced warbler
1925:Sunda bush warbler
1920:Shade bush warbler
1910:Palau bush warbler
1557:; Donsker, David;
1458:Hemitesia neumanni
1374:Hemitesia neumanni
1322:Urosphena subulata
1268:Cettia brunnifrons
1187:Grey-bellied tesia
1107:Horornis carolinae
1073:Shade bush warbler
1063:Palau bush warbler
1003:Tickellia hodgsoni
293:Hemitesia neumanni
2192:
2191:
2177:Open Tree of Life
2013:Taxon identifiers
2004:
2003:
1986:Neumann's warbler
1890:Fiji bush warbler
1650:
1559:Rasmussen, Pamela
1512:(16): 7916–7925.
1442:978-84-941892-0-3
1382:
1381:
1370:Neumann's warbler
1191:Tesia cyaniventer
1117:Horornis fortipes
1093:Fiji bush warbler
1057:Horornis seebohmi
860:
859:
851:
850:
842:
841:
833:
832:
824:
823:
722:
721:
713:
712:
661:
660:
592:
591:
583:
582:
574:
573:
565:
564:
556:
555:
547:
546:
538:
537:
529:
528:
289:Neumann's warbler
241:is usually used.
221:Old World warbler
199:
198:
135:
2237:
2185:
2184:
2172:
2171:
2159:
2158:
2156:NHMSYS0021003930
2146:
2145:
2133:
2132:
2120:
2119:
2107:
2106:
2094:
2093:
2081:
2080:
2068:
2067:
2055:
2054:
2053:
2040:
2039:
2038:
2008:
2007:
1981:Bornean stubtail
1704:
1697:
1690:
1681:
1680:
1642:
1622:
1621:
1619:
1609:
1585:
1579:
1578:
1576:
1574:
1551:
1542:
1541:
1531:
1521:
1497:
1491:
1484:
1478:
1477:
1453:
1447:
1446:
1428:
1422:
1421:
1398:
1353:
1328:Bornean stubtail
1301:
1300:R. Swinhoe, 1877
1291:
1237:
1224:
1170:
1160:
1087:Horornis haddeni
1047:Horornis diphone
1026:
1016:
992:
982:
948:
938:
894:
884:
866:
865:
786:
785:
761:
760:
736:
735:
675:
674:
623:
622:
616:
615:
609:
608:
599:
598:
495:
494:
472:
471:
449:
448:
426:
425:
403:
402:
396:
395:
388:
387:
381:
380:
371:Pamela Rasmussen
239:cettiid warblers
130:
58:
57:
33:
21:
20:
2245:
2244:
2240:
2239:
2238:
2236:
2235:
2234:
2195:
2194:
2193:
2188:
2180:
2175:
2167:
2162:
2154:
2149:
2141:
2136:
2128:
2123:
2115:
2110:
2102:
2097:
2089:
2084:
2076:
2071:
2063:
2058:
2049:
2048:
2043:
2034:
2033:
2028:
2015:
2005:
2000:
1962:
1939:
1861:
1842:Cetti's warbler
1823:
1793:
1714:
1708:
1678:
1631:
1626:
1625:
1586:
1582:
1572:
1570:
1552:
1545:
1498:
1494:
1485:
1481:
1454:
1450:
1443:
1429:
1425:
1399:
1392:
1387:
1348:
1299:
1274:Cetti's warbler
1232:
1168:
1077:Horornis parens
1027:– bush warblers
1024:
990:
946:
892:
875:Living species
861:
852:
843:
834:
825:
723:
714:
662:
593:
584:
575:
566:
557:
548:
539:
530:
479:Erythrocercidae
356:
329:Cetti's warbler
267:, ranging into
189:
183:
177:
171:
165:
159:
153:
129:
52:
39:Cetti's warbler
17:
16:Family of birds
12:
11:
5:
2243:
2233:
2232:
2227:
2222:
2217:
2212:
2207:
2190:
2189:
2187:
2186:
2173:
2160:
2147:
2134:
2121:
2108:
2095:
2082:
2069:
2056:
2041:
2025:
2023:
2017:
2016:
2002:
2001:
1999:
1998:
1996:Timor stubtail
1993:
1988:
1983:
1978:
1976:Asian stubtail
1972:
1970:
1964:
1963:
1961:
1960:
1955:
1949:
1947:
1941:
1940:
1938:
1937:
1932:
1927:
1922:
1917:
1912:
1907:
1902:
1897:
1892:
1887:
1882:
1877:
1871:
1869:
1863:
1862:
1860:
1859:
1854:
1849:
1844:
1839:
1833:
1831:
1825:
1824:
1822:
1821:
1816:
1811:
1805:
1803:
1795:
1794:
1792:
1791:
1783:
1775:
1767:
1759:
1758:– (14 species)
1751:
1743:
1735:
1722:
1720:
1716:
1715:
1707:
1706:
1699:
1692:
1684:
1677:
1676:
1661:and Tit-hylia
1657:, Green Hylia
1651:
1632:
1630:
1627:
1624:
1623:
1580:
1543:
1492:
1479:
1448:
1441:
1423:
1412:(2): 381–397.
1389:
1388:
1386:
1383:
1380:
1379:
1378:
1377:
1367:
1355:
1341:
1338:
1337:
1336:
1335:
1325:
1318:Timor stubtail
1315:
1308:Asian stubtail
1303:
1292:
1284:
1283:
1282:
1281:
1271:
1261:
1251:
1239:
1225:
1217:
1216:
1215:
1214:
1211:Tesia everetti
1204:
1194:
1184:
1172:
1161:
1153:
1152:
1151:
1150:
1140:
1130:
1120:
1110:
1100:
1090:
1080:
1070:
1067:Horornis annae
1060:
1050:
1040:
1028:
1017:
1009:
1008:
1007:
1006:
994:
983:
975:
974:
973:
972:
962:
950:
939:
931:
930:
929:
928:
918:
908:
896:
885:
877:
876:
873:
870:
858:
857:
854:
853:
849:
848:
845:
844:
840:
839:
836:
835:
831:
830:
827:
826:
822:
821:
818:
817:
807:
804:
803:
800:
799:
789:
784:
782:
779:
778:
775:
774:
764:
759:
757:
754:
753:
750:
749:
739:
734:
732:
729:
728:
725:
724:
720:
719:
716:
715:
711:
710:
707:
706:
704:– (4 species)
696:
693:
692:
689:
688:
678:
673:
671:
668:
667:
664:
663:
659:
658:
655:
654:
644:
641:
640:
637:
636:
634:– (2 species)
626:
621:
619:
614:
612:
607:
605:
597:
590:
589:
586:
585:
581:
580:
577:
576:
572:
571:
568:
567:
563:
562:
559:
558:
554:
553:
550:
549:
545:
544:
541:
540:
536:
535:
532:
531:
527:
526:
523:
522:
514:
511:
510:
507:
506:
498:
493:
491:
488:
487:
484:
483:
475:
470:
468:
465:
464:
461:
460:
452:
447:
445:
442:
441:
438:
437:
429:
424:
422:
419:
418:
415:
414:
410:Phylloscopidae
406:
401:
399:
394:
392:
391:Aegithaloidea
386:
384:
379:
355:
352:
317:Asian stubtail
197:
196:
144:
143:
137:
136:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
74:
70:
69:
64:
60:
59:
46:
45:
35:
34:
26:
25:
15:
9:
6:
4:
3:
2:
2242:
2231:
2228:
2226:
2223:
2221:
2218:
2216:
2213:
2211:
2210:Bird families
2208:
2206:
2203:
2202:
2200:
2183:
2178:
2174:
2170:
2165:
2161:
2157:
2152:
2148:
2144:
2139:
2135:
2131:
2126:
2122:
2118:
2113:
2109:
2105:
2100:
2096:
2092:
2087:
2083:
2079:
2074:
2070:
2066:
2061:
2057:
2052:
2046:
2042:
2037:
2031:
2027:
2026:
2024:
2022:
2018:
2014:
2009:
1997:
1994:
1992:
1989:
1987:
1984:
1982:
1979:
1977:
1974:
1973:
1971:
1969:
1965:
1959:
1956:
1954:
1951:
1950:
1948:
1946:
1942:
1936:
1933:
1931:
1928:
1926:
1923:
1921:
1918:
1916:
1913:
1911:
1908:
1906:
1903:
1901:
1898:
1896:
1893:
1891:
1888:
1886:
1883:
1881:
1878:
1876:
1873:
1872:
1870:
1868:
1864:
1858:
1855:
1853:
1850:
1848:
1845:
1843:
1840:
1838:
1835:
1834:
1832:
1830:
1826:
1820:
1817:
1815:
1812:
1810:
1807:
1806:
1804:
1802:
1801:
1796:
1789:
1788:
1784:
1782:- (2 species)
1781:
1780:
1776:
1773:
1772:
1768:
1765:
1764:
1760:
1757:
1756:
1752:
1749:
1748:
1744:
1741:
1740:
1736:
1733:
1729:
1728:
1724:
1723:
1721:
1717:
1713:
1705:
1700:
1698:
1693:
1691:
1686:
1685:
1682:
1675:
1672:(1-2): 8-17.
1671:
1668:
1664:
1660:
1659:Hylia prasina
1656:
1652:
1649:
1648:84-96553-06-X
1645:
1640:
1639:
1634:
1633:
1618:
1613:
1608:
1603:
1599:
1595:
1591:
1584:
1568:
1564:
1560:
1556:
1550:
1548:
1539:
1535:
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502:Scotocercidae
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345:Locustellidae
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258:Scotocercidae
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247:Erythrocercus
242:
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229:bush warblers
226:
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27:
22:
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2020:
1945:Phyllergates
1837:Bush-warbler
1798:
1785:
1779:Phyllergates
1777:
1769:
1761:
1753:
1745:
1737:
1731:
1725:
1711:
1674:PDF fulltext
1669:
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1636:
1597:
1593:
1583:
1571:. Retrieved
1566:
1509:
1505:
1495:
1487:
1482:
1468:(1): 78–86.
1465:
1461:
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1451:
1432:
1426:
1409:
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1373:
1363:
1343:
1331:
1321:
1311:
1294:
1278:Cettia cetti
1277:
1267:
1257:
1248:Cettia major
1247:
1227:
1210:
1200:
1190:
1181:Tesia olivea
1180:
1163:
1146:
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1126:
1116:
1106:
1096:
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1076:
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985:
968:
958:
947:Sharpe, 1883
943:Phyllergates
941:
924:
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810:
809:
792:
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769:Phyllergates
767:
766:
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681:
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647:
646:
629:
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602:
594:
517:
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477:
456:Aegithalidae
454:
431:
408:
364:
357:
349:
337:
302:
296:
292:
282:
276:
262:
251:
245:
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201:
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190:
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178:
172:
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156:Phyllergates
154:
148:
147:
126:
43:Cettia cetti
42:
18:
2125:iNaturalist
2045:Wikispecies
1555:Gill, Frank
1402:Alström, P.
1302:– stubtails
1197:Javan tesia
991:Blyth, 1861
893:Baker, 1930
360:Per Alström
2199:Categories
1800:Abroscopus
1787:Scotocerca
1732:Abroscopus
1727:Abroscopus
1600:(1): 352.
1385:References
1354:– warblers
895:– warblers
889:Abroscopus
744:Abroscopus
367:Frank Gill
319:is wholly
284:Abroscopus
253:Scotocerca
150:Abroscopus
117:Sylvioidea
24:Cettiidae
2205:Cettiidae
2051:Cettiidae
2021:Cettiidae
1968:Urosphena
1771:Tickellia
1739:Urosphena
1712:Cettiidae
1573:26 August
1486:del Hoyo
1345:Hemitesia
1296:Urosphena
1234:Bonaparte
987:Tickellia
794:Tickellia
649:Urosphena
631:Hemitesia
603:Cettiidae
518:Cettiidae
321:migratory
278:Tickellia
213:songbirds
208:of small
202:Cettiidae
192:Hemitesia
186:Urosphena
162:Tickellia
127:Cettiidae
73:Kingdom:
67:Eukaryota
2091:10556136
2030:Wikidata
1867:Horornis
1755:Horornis
1538:30936315
1171:– tesias
1021:Horornis
812:Horornis
433:Hyliidae
354:Taxonomy
323:and the
304:Horornis
225:wastebin
217:warblers
168:Horornis
123:Family:
87:Chordata
83:Phylum:
77:Animalia
63:Domain:
2182:5846361
2169:1592216
2117:6100900
2036:Q222432
1667:Ostrich
1629:Sources
1617:3261208
1529:6475423
377:(IOC).
340:species
269:Oceania
132:Alström
103:Order:
93:Class:
2143:915695
2130:144308
2104:1CETTF
2065:743402
1829:Cettia
1763:Cettia
1719:Genera
1646:
1614:
1536:
1526:
1490:(2006)
1488:et al.
1439:
1352:, 1948
1350:Chapin
1236:, 1834
1229:Cettia
701:Cettia
298:Cettia
273:Europe
234:Cettia
206:family
180:Cettia
141:Genera
1747:Tesia
1165:Tesia
872:Genus
869:Image
683:Tesia
313:Palau
174:Tesia
2164:NCBI
2138:ITIS
2112:GBIF
2099:EPPO
2060:BOLD
1644:ISBN
1575:2022
1534:PMID
1462:Ibis
1437:ISBN
338:The
327:and
311:and
309:Fiji
281:and
271:and
265:Asia
250:and
97:Aves
2151:NBN
2086:EoL
2078:7Y5
2073:CoL
1612:PMC
1602:doi
1524:PMC
1514:doi
1510:116
1470:doi
1466:153
1414:doi
2201::
2179::
2166::
2153::
2140::
2127::
2114::
2101::
2088::
2075::
2062::
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2032::
1730:–
1670:74
1665:.
1610:.
1598:11
1596:.
1592:.
1565:.
1546:^
1532:.
1522:.
1508:.
1504:.
1464:.
1410:38
1408:.
1393:^
369:,
260:.
215:("
41:,
1703:e
1696:t
1689:v
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1189:(
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1145:(
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1125:(
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1005:)
1001:(
971:)
967:(
961:)
957:(
927:)
923:(
917:)
913:(
907:)
903:(
291:(
231:(
223:"
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