216:
75:
50:
31:
378:
Preliminary satellite analyses with Ladsat images show that from 1989 to 2002, 50 % of the primary forests in the species' range have been lost and the pasture areas have doubled. By 2002 only 32 % of the area still retained some vegetation cover (primary, secondary and flooded forest) in
387:
All 13 groups studied consisted of a single adult male, a single adult female and between one and four immature monkeys. Average group size, including adults and immatures, was 4.1 monkeys. This is consistent with other titi monkey species, which form small, cohesive groups led by a bonded pair of
392:
and pairs raise about one baby each year. Babies make purring noises similar to cats when they are content. Titi monkey diets typically consist primarily of fruit, with leaves the second most important food item, and with seeds forming only a small portion of the diet.
379:
which
Callicebus caquetensis can survive; however, the area corresponding to secondary forests (531 hectares) is severely fragmented and the primary and flooded forests are restricted to the banks of the Orteguaza and Caquetá rivers.
368:
in order to reach new nearby forest fragments. The species has a geographic range of just about 100 square kilometers (39 sq mi) and actually occupies only about 10 square kilometers (3.9 sq mi) within that range.
360:(IUCN). According to the authors' estimate, the population size of the species may be fewer than 250 adult animals. It lives in forests fragmented by agricultural activity, and dispersal is hazardous since it must cross open
592:
600:
IUCN SSC Primate
Specialist Group (PSG), International Primatological Society (IPS), Conservation International (CI), and Bristol Zoological Society, Arlington, VA
1807:
566:
1777:
320:
except that it does not have a white forehead bar, nor does it have the white hands and feet of the ornate titi. It also has a red beard. It has 22
357:
1723:
1578:
693:
622:
Norconk, M. (2007). "Sakis, Uakaris and Titi
Monkeys". In Campbell, C.; Fuentes, A.; MacKinnon, K.; Panger, M.; Bearder, S. (eds.).
1684:
1526:
1710:
1552:
312:
Its fur is generally brown, with a lighter tail and chestnut-red undersides, neck and cheeks. It is similar in appearance to the
523:
1802:
1715:
1557:
631:
344:
in the
Colombian Department of Caquetá at altitudes between 190 and 260 m, but the exact distribution area is still unknown.
1817:
1491:
1812:
1787:
1399:
1728:
1331:
1128:
570:
1650:
1092:
650:
1322:
1313:
884:
1792:
1281:
686:
54:
1797:
1263:
1209:
74:
325:
1437:
1164:
1110:
961:
943:
500:
1340:
1200:
1119:
934:
332:
of 2n=46. Its body is approximately 35 cm long and its tail 61 cm, it weighs between 800 and 1400 g.
1254:
1101:
679:
279:
1227:
1191:
1182:
1173:
1065:
925:
1609:
1782:
1453:
1408:
1155:
1570:
1741:
1604:
1596:
476:
Defler, T.R.; García-Gutiérrez, J.; Guzmán-Caro, D.C.; Palacios, E.; Stevenson, P.R. (2021).
475:
353:
352:
In the paper describing the species, the authors recommend that the species be classified as
301:
288:
189:
227:
1671:
1513:
1417:
1349:
1083:
1074:
8:
1381:
1218:
1146:
1002:
477:
389:
39:
830:
1272:
1236:
916:
671:
548:
458:
235:
69:
1736:
1658:
1591:
1500:
1390:
1137:
803:
785:
627:
317:
552:
521:
462:
1505:
952:
875:
857:
839:
794:
538:
495:
448:
341:
215:
1663:
1476:
776:
1754:
1583:
1011:
1636:
1020:
821:
543:
437:: A New and Critically Endangered Titi Monkey from Southern Caquetá, Colombia"
1771:
1565:
1048:
486:
297:
205:
166:
59:
899:
1697:
1539:
1485:
1038:
993:
759:
749:
703:
590:
453:
432:
365:
313:
271:
156:
300:, Marta Bueno and Javier Garcia in 2010. It is highly endangered due to
1689:
1531:
1364:
1296:
812:
739:
524:"Conservation of Colombian primates: an analysis of published research"
375:
has been listed among the World's 25 Most
Endangered Primates in 2017.
321:
136:
1702:
1544:
593:"Primates in Peril: The World's 25 Most Endangered Primates 2016–2018"
866:
848:
86:
1447:
1630:
1470:
733:
727:
721:
275:
106:
30:
1676:
1518:
522:
Stevenson, Pablo R.; Guzmán, Diana C.; Defler, Thomas R. (2010).
361:
329:
126:
1645:
976:
715:
591:
Defler, Thomas R.; Javier García: Diana C. Guzmán-Caro (2017).
146:
116:
96:
1749:
469:
701:
431:
Defler, Thomas R.; Bueno, Marta; Garcia, Javier (2010).
340:
The species occurs between the Río
Orteguaza and the
651:"Rare Redbearded Monkeys Found Living in Colombia"
501:10.2305/IUCN.UK.2021-1.RLTS.T14699281A192453101.en
430:
388:adults. Like other titi monkey species, they are
1769:
569:. ScienceNews. August 12, 2010. Archived from
358:International Union for Conservation of Nature
282:region. Taxonomically, it is a member of the "
687:
626:. Oxford University Press. pp. 123–138.
617:
615:
1808:Critically endangered biota of South America
1778:IUCN Red List critically endangered species
621:
335:
694:
680:
612:
584:
214:
48:
29:
542:
499:
452:
426:
1770:
424:
422:
420:
418:
416:
414:
412:
410:
408:
406:
1452:
1451:
675:
645:
643:
296:grouping. It was first described by
487:IUCN Red List of Threatened Species
403:
13:
640:
292:), following Shunsuke Kobayashi's
14:
1829:
73:
347:
559:
515:
307:
1:
1803:Critically endangered animals
1400:Colombian black-handed titi (
885:Vanzolini's bald-faced saki (
531:Tropical Conservation Science
396:
962:Reddish-brown bearded saki (
7:
1818:Taxa named by Thomas Defler
1332:Coastal black-handed titi (
935:Brown-backed bearded saki (
16:Species of New World monkey
10:
1834:
567:"New titi monkey, at last"
544:10.1177/194008291000300105
382:
1813:Mammals described in 2010
1788:Endemic fauna of Colombia
1620:
1492:Plecturocebus caquetensis
1462:Plecturocebus caquetensis
1460:
1434:
1362:
1294:
1046:
1037:
974:
944:Uta Hick's bearded saki (
926:Red-backed bearded saki (
897:
757:
748:
710:
702:Extant species of family
494:: e.T14699281A192453101.
480:Plecturocebus caquetensis
330:diploid chromosome number
259:Plecturocebus caquetensis
241:
234:
222:
213:
199:Plecturocebus caquetensis
195:
188:
70:Scientific classification
68:
46:
37:
28:
23:
1129:Prince Bernhard's titi (
336:Distribution and habitat
324:pairs (not counting the
304:and a small population.
1210:Chestnut-bellied titi (
1093:Olalla brothers' titi (
624:Primates in Perspective
1651:callicebus-caquetensis
1622:Callicebus caquetensis
1323:Coimbra Filho's titi (
1314:Barbara Brown's titi (
435:Callicebus caquetensis
244:Callicebus caquetensis
1605:Paleobiology Database
1282:Stephen Nash's titi (
1111:Urubamba brown titi (
1003:Black-headed uakari (
354:critically endangered
302:habitat fragmentation
289:Plecturocebus cupreus
280:Department of Caquetá
262:), also known as the
55:Critically Endangered
1341:Black-fronted titi (
1264:Hershkovitz's titi (
1201:Alta Floresta titi (
1120:Baptista Lake titi (
917:Black bearded saki (
454:10.1896/052.025.0101
441:Primate Conservation
1798:Mammals of Colombia
1255:White-tailed titi (
1102:White-coated titi (
804:Golden-faced saki (
786:White-footed saki (
254:Caquetá titi monkey
181:P. caquetensis
40:Conservation status
1793:Caquetá Department
1174:Red-bellied titi (
1165:Hoffmanns's titi (
1066:White-eared titi (
953:White-nosed saki (
876:White-faced saki (
831:Rio Tapajós saki (
573:on August 15, 2010
284:Callicebus cupreus
270:, is a species of
268:bushy-bearded titi
228:Caquetá department
1765:
1764:
1737:Open Tree of Life
1592:Open Tree of Life
1454:Taxon identifiers
1445:
1444:
1430:
1429:
1409:Red-headed titi (
1375:
1316:C. barbarabrownae
1307:
1156:Ashy black titi (
1059:
1033:
1032:
1005:C. melanocephalus
987:
910:
777:Equatorial saki (
770:
657:. August 13, 2010
633:978-0-19-517133-4
318:white-tailed titi
250:
249:
63:
1825:
1758:
1757:
1745:
1744:
1732:
1731:
1719:
1718:
1706:
1705:
1693:
1692:
1680:
1679:
1667:
1666:
1654:
1653:
1641:
1640:
1639:
1613:
1612:
1600:
1599:
1587:
1586:
1574:
1573:
1561:
1560:
1548:
1547:
1535:
1534:
1522:
1521:
1509:
1508:
1496:
1495:
1494:
1481:
1480:
1479:
1449:
1448:
1374:
1373:
1369:
1306:
1305:
1301:
1058:
1057:
1053:
1044:
1043:
1021:Neblina uakari (
986:
985:
981:
909:
908:
904:
822:Burnished saki (
779:P. aequatorialis
769:
768:
764:
755:
754:
696:
689:
682:
673:
672:
666:
665:
663:
662:
647:
638:
637:
619:
610:
609:
607:
606:
597:
588:
582:
581:
579:
578:
563:
557:
556:
546:
528:
519:
513:
512:
510:
508:
503:
473:
467:
466:
456:
428:
264:red-bearded titi
218:
201:
78:
77:
57:
52:
51:
33:
21:
20:
1833:
1832:
1828:
1827:
1826:
1824:
1823:
1822:
1768:
1767:
1766:
1761:
1753:
1748:
1740:
1735:
1727:
1722:
1714:
1709:
1701:
1696:
1688:
1683:
1675:
1670:
1662:
1657:
1649:
1644:
1635:
1634:
1629:
1616:
1608:
1603:
1595:
1590:
1582:
1577:
1569:
1564:
1556:
1551:
1543:
1538:
1530:
1525:
1517:
1512:
1504:
1499:
1490:
1489:
1484:
1475:
1474:
1469:
1456:
1446:
1441:
1426:
1418:Collared titi (
1371:
1370:
1368:
1358:
1350:Atlantic titi (
1303:
1302:
1300:
1290:
1284:P. stephennashi
1239:P. aureipalatii
1228:Toppin's titi (
1192:Milton's titi (
1183:Vieira's titi (
1113:P. urubambensis
1084:Rio Mayo titi (
1075:Rio Beni titi (
1068:P. donacophilus
1055:
1054:
1052:
1029:
983:
982:
980:
970:
907:(Bearded sakis)
906:
905:
903:
893:
858:Miller's saki (
840:Isabel's saki (
806:P.chrysocephala
795:Cazuza's saki (
766:
765:
763:
744:
706:
700:
670:
669:
660:
658:
649:
648:
641:
634:
620:
613:
604:
602:
595:
589:
585:
576:
574:
565:
564:
560:
526:
520:
516:
506:
504:
474:
470:
429:
404:
399:
385:
350:
338:
326:sex chromosomes
310:
209:
203:
197:
184:
72:
64:
53:
49:
42:
17:
12:
11:
5:
1831:
1821:
1820:
1815:
1810:
1805:
1800:
1795:
1790:
1785:
1780:
1763:
1762:
1760:
1759:
1746:
1733:
1720:
1707:
1694:
1681:
1668:
1655:
1642:
1626:
1624:
1618:
1617:
1615:
1614:
1601:
1588:
1575:
1562:
1549:
1536:
1523:
1510:
1497:
1482:
1466:
1464:
1458:
1457:
1443:
1442:
1435:
1432:
1431:
1428:
1427:
1425:
1424:
1415:
1406:
1397:
1388:
1382:Lucifer titi (
1378:
1376:
1360:
1359:
1357:
1356:
1347:
1338:
1329:
1320:
1310:
1308:
1292:
1291:
1289:
1288:
1279:
1270:
1261:
1252:
1248:P. caquetensis
1246:Caquetá titi (
1243:
1234:
1225:
1219:Coppery titi (
1216:
1207:
1198:
1189:
1180:
1171:
1162:
1158:P. cinerascens
1153:
1147:Parecis titi (
1144:
1135:
1126:
1117:
1108:
1099:
1090:
1081:
1072:
1062:
1060:
1041:
1035:
1034:
1031:
1030:
1028:
1027:
1018:
1012:Aracá uakari (
1009:
1000:
990:
988:
972:
971:
969:
968:
959:
950:
941:
932:
923:
913:
911:
895:
894:
892:
891:
882:
873:
864:
855:
846:
837:
828:
819:
810:
801:
792:
783:
773:
771:
767:(Saki monkeys)
752:
746:
745:
743:
742:
736:
730:
724:
718:
711:
708:
707:
699:
698:
691:
684:
676:
668:
667:
639:
632:
611:
583:
558:
514:
468:
401:
400:
398:
395:
384:
381:
373:P. caquetensis
349:
346:
337:
334:
309:
306:
248:
247:
239:
238:
232:
231:
224:P. caquetensis
220:
219:
211:
210:
208:, et al., 2010
204:
193:
192:
186:
185:
178:
176:
172:
171:
164:
160:
159:
154:
150:
149:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
66:
65:
47:
44:
43:
38:
35:
34:
26:
25:
15:
9:
6:
4:
3:
2:
1830:
1819:
1816:
1814:
1811:
1809:
1806:
1804:
1801:
1799:
1796:
1794:
1791:
1789:
1786:
1784:
1783:Plecturocebus
1781:
1779:
1776:
1775:
1773:
1756:
1751:
1747:
1743:
1738:
1734:
1730:
1725:
1721:
1717:
1712:
1708:
1704:
1699:
1695:
1691:
1686:
1682:
1678:
1673:
1669:
1665:
1660:
1656:
1652:
1647:
1643:
1638:
1632:
1628:
1627:
1625:
1623:
1619:
1611:
1606:
1602:
1598:
1593:
1589:
1585:
1580:
1576:
1572:
1567:
1563:
1559:
1554:
1550:
1546:
1541:
1537:
1533:
1528:
1524:
1520:
1515:
1511:
1507:
1502:
1498:
1493:
1487:
1483:
1478:
1472:
1468:
1467:
1465:
1463:
1459:
1455:
1450:
1440:
1439:
1433:
1423:
1421:
1416:
1414:
1412:
1407:
1405:
1403:
1398:
1396:
1394:
1389:
1387:
1385:
1380:
1379:
1377:
1367:
1366:
1361:
1355:
1353:
1352:C. personatus
1348:
1346:
1344:
1343:C. nigrifrons
1339:
1337:
1335:
1334:C. melanochir
1330:
1328:
1326:
1321:
1319:
1317:
1312:
1311:
1309:
1299:
1298:
1293:
1287:
1285:
1280:
1278:
1276:
1273:Ornate titi (
1271:
1269:
1267:
1262:
1260:
1258:
1253:
1251:
1249:
1244:
1242:
1240:
1237:Madidi titi (
1235:
1233:
1231:
1226:
1224:
1222:
1217:
1215:
1213:
1208:
1206:
1204:
1199:
1197:
1195:
1190:
1188:
1186:
1181:
1179:
1177:
1172:
1170:
1168:
1167:P. hoffmannsi
1163:
1161:
1159:
1154:
1152:
1150:
1145:
1143:
1141:
1136:
1134:
1132:
1127:
1125:
1123:
1118:
1116:
1114:
1109:
1107:
1105:
1104:P. pallescens
1100:
1098:
1096:
1091:
1089:
1087:
1082:
1080:
1078:
1073:
1071:
1069:
1064:
1063:
1061:
1051:
1050:
1049:Plecturocebus
1045:
1042:
1040:
1036:
1026:
1024:
1019:
1017:
1015:
1010:
1008:
1006:
1001:
999:
997:
994:Bald uakari (
992:
991:
989:
979:
978:
973:
967:
965:
960:
958:
956:
951:
949:
947:
942:
940:
938:
933:
931:
929:
928:C. chiropotes
924:
922:
920:
915:
914:
912:
902:
901:
896:
890:
888:
887:P. vanzolinii
883:
881:
879:
874:
872:
870:
865:
863:
861:
856:
854:
852:
847:
845:
843:
838:
836:
834:
829:
827:
825:
820:
818:
816:
811:
809:
807:
802:
800:
798:
793:
791:
789:
784:
782:
780:
775:
774:
772:
762:
761:
756:
753:
751:
747:
741:
737:
735:
731:
729:
725:
723:
719:
717:
713:
712:
709:
705:
697:
692:
690:
685:
683:
678:
677:
674:
656:
652:
646:
644:
635:
629:
625:
618:
616:
601:
594:
587:
572:
568:
562:
554:
550:
545:
540:
536:
532:
525:
518:
502:
497:
493:
489:
488:
483:
481:
472:
464:
460:
455:
450:
446:
442:
438:
436:
427:
425:
423:
421:
419:
417:
415:
413:
411:
409:
407:
402:
394:
391:
380:
376:
374:
370:
367:
363:
359:
355:
345:
343:
333:
331:
327:
323:
319:
315:
305:
303:
299:
298:Thomas Defler
295:
291:
290:
285:
281:
278:found in the
277:
273:
269:
265:
261:
260:
255:
246:
245:
240:
237:
233:
229:
225:
221:
217:
212:
207:
202:
200:
194:
191:
190:Binomial name
187:
183:
182:
177:
174:
173:
170:
169:
168:Plecturocebus
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
76:
71:
67:
61:
56:
45:
41:
36:
32:
27:
24:Caquetá titi
22:
19:
1621:
1461:
1436:
1420:C. torquatus
1419:
1410:
1401:
1392:
1391:Black titi (
1383:
1363:
1351:
1342:
1333:
1324:
1315:
1295:
1283:
1274:
1265:
1256:
1247:
1245:
1238:
1229:
1220:
1212:P. caligatus
1211:
1202:
1193:
1184:
1175:
1166:
1157:
1148:
1139:
1138:Brown titi (
1131:P. bernhardi
1130:
1121:
1112:
1103:
1094:
1085:
1076:
1067:
1047:
1039:Callicebinae
1022:
1013:
1004:
995:
975:
964:C. sagulatus
963:
955:C. albinasus
954:
946:C. utahickae
945:
937:C. israelita
936:
927:
918:
898:
886:
877:
868:
859:
850:
841:
832:
823:
814:
813:Hairy saki (
805:
796:
787:
778:
758:
659:. Retrieved
654:
623:
603:. Retrieved
599:
586:
575:. Retrieved
571:the original
561:
537:(1): 45–62.
534:
530:
517:
505:. Retrieved
491:
485:
479:
471:
444:
440:
434:
386:
377:
372:
371:
351:
348:Conservation
339:
311:
293:
287:
286:group" (now
283:
267:
263:
258:
257:
253:
251:
243:
242:
223:
198:
196:
180:
179:
167:
143:Infraorder:
18:
1698:iNaturalist
1540:iNaturalist
1486:Wikispecies
1325:C. coimbrai
1257:P. discolor
1140:P. brunneus
1122:P. baptista
1086:P. oenanthe
1077:P. modestus
878:P. pithecia
869:P. napensis
867:Napo saki (
851:P. monachus
849:Monk saki (
833:P. irrorata
788:P. albicans
750:Pitheciinae
704:Pitheciidae
507:12 November
366:barbed wire
342:Río Caquetá
314:ornate titi
308:Description
274:endemic to
272:titi monkey
230:, Colombia
157:Pitheciidae
147:Simiiformes
1772:Categories
1411:C. regulus
1384:C. lucifer
1365:Cheracebus
1297:Callicebus
1275:P. ornatus
1230:P. toppini
1221:P. cupreus
1203:P. grovesi
1194:P. miltoni
1185:P. vieirai
1149:P. parecis
1095:P. olallae
1014:C. ayresii
919:C. satanas
900:Chiropotes
860:P. milleri
842:P. isabela
815:P. hirsuta
797:P. cazuzai
740:Haplorhini
738:Suborder:
661:2010-08-13
605:2022-10-20
577:2010-08-13
397:References
390:monogamous
328:), with a
322:chromosome
294:Callicebus
226:on map of
137:Haplorhini
133:Suborder:
1477:Q56116516
1402:C. medemi
1393:C. lugens
1266:P. dubius
1176:P. moloch
1023:C. hosomi
996:C. calvus
984:(Uakaris)
824:P. inusta
714:Kingdom:
465:. 136211.
175:Species:
93:Kingdom:
87:Eukaryota
1750:Species+
1677:17711226
1637:Q1035244
1631:Wikidata
1571:14699281
1519:51906298
1471:Wikidata
1438:Category
760:Pithecia
734:Primates
728:Mammalia
722:Chordata
720:Phylum:
716:Animalia
655:CBS News
553:87418117
463:83583912
276:Colombia
236:Synonyms
153:Family:
127:Primates
117:Mammalia
107:Chordata
103:Phylum:
97:Animalia
83:Domain:
60:IUCN 3.1
1742:5925586
1729:1861701
1716:1025147
1690:9163862
1597:5925586
1584:1000873
1558:1025130
1532:9438354
1372:(Titis)
1304:(Titis)
1056:(Titis)
977:Cacajao
732:Order:
726:Class:
447:: 1–9.
383:Biology
362:savanna
356:by the
266:or the
163:Genus:
123:Order:
113:Class:
58: (
1703:144088
1646:ARKive
1610:390299
1545:738201
630:
551:
461:
206:Defler
1506:4K5Y2
596:(PDF)
549:S2CID
527:(PDF)
459:S2CID
1755:7400
1724:NCBI
1711:ITIS
1685:GBIF
1664:PML6
1566:IUCN
1553:ITIS
1527:GBIF
628:ISBN
509:2021
492:2021
316:and
252:The
1672:EoL
1659:CoL
1579:MDD
1514:EoL
1501:CoL
539:doi
496:doi
449:doi
364:or
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642:^
614:^
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256:(
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