68:
43:
263:
is a small megabat, weighing only 10–15 g (0.35–0.53 oz). It has a forearm length of 40–45 mm (1.6–1.8 in). Its fur is blackish brown on its back and somewhat paler on its underside.
2705:
295:
ferns, and, occasionally, caves. Its habitat includes lowland rainforests at elevations from sea level to 1,500 m (4,900 ft) above sea level.
3424:
3465:
390:
Diversification of Old World Bats in
Malaysia: An Evolutionary and Phylogeography Hypothesis tested through the Genetic Species Concept
3376:
3385:
3069:
3398:
3180:
3045:
3036:
510:
3485:
3442:
2962:
476:
3403:
466:
1374:
1365:
2894:
1419:
3470:
2980:
2930:
1464:
972:
608:
2912:
1383:
3171:
2971:
2876:
2867:
2400:
1392:
990:
644:
2577:
1774:
1473:
1437:
1410:
707:
662:
2903:
2568:
1455:
1446:
758:
2123:
1974:
1938:
1825:
1428:
1130:
981:
635:
3226:
2885:
2858:
1947:
1064:
936:
689:
680:
67:
3437:
3272:
1534:
1401:
1031:
749:
671:
653:
617:
2921:
2472:
2204:
2069:
1597:
1022:
725:
626:
503:
343:
3480:
3189:
2737:
1656:
405:"A checklist of the bats of Peninsular Malaysia and progress towards a DNA barcode reference library"
2526:
2481:
2276:
2186:
2150:
1965:
1956:
1866:
1497:
1139:
1121:
1088:
945:
873:
849:
217:
3263:
2373:
2105:
2033:
1914:
1890:
1816:
1706:
1615:
1606:
1564:
1252:
1219:
279:. Its range likely includes other countries in Southeast Asia, including Thailand and Indonesia (
3338:
2710:
2382:
2285:
2168:
2060:
1573:
1228:
904:
716:
3416:
3305:
2454:
2436:
2096:
1697:
1647:
1333:
1286:
954:
698:
496:
162:
3281:
2782:
2764:
2463:
2427:
2312:
2240:
2231:
2006:
1857:
416:
8:
2791:
2653:
2499:
2409:
2294:
2078:
1756:
276:
209:
32:
488:
420:
320:
2728:
2635:
2490:
2418:
2349:
2222:
2132:
2042:
2015:
1688:
1112:
439:
404:
62:
3475:
3139:
3130:
3004:
2719:
2508:
2364:
2339:
2321:
2267:
2258:
2249:
2213:
2114:
2087:
1783:
1765:
1311:
591:
472:
444:
366:
3106:
2746:
2692:
2544:
2535:
2517:
2445:
2391:
2355:
2303:
2177:
2141:
2024:
1792:
963:
816:
434:
424:
338:
284:
241:
3361:
2755:
2644:
2330:
2159:
1840:
1506:
1488:
1276:
1187:
1163:
1040:
825:
429:
3429:
3244:
2195:
2051:
556:
240:, Malaysia. Later genetic studies showed that there was a larger-than-expected
213:
178:
3459:
3411:
3296:
3121:
3097:
3019:
2819:
2777:
1055:
864:
740:
329:
139:
52:
47:
3254:
3087:
3060:
2995:
2831:
2559:
1734:
1671:
1588:
1547:
840:
786:
578:
448:
3370:
3154:
2841:
2605:
1202:
1154:
1005:
403:
Lim, Voon-Ching; Ramli, Rosli; Bhassu, Subha; Wilson, John-James (2017).
3390:
2618:
1929:
1807:
1630:
1267:
919:
562:
288:
3207:
2668:
1905:
1324:
1243:
1178:
1103:
799:
292:
79:
3332:
3355:
3217:
1989:
1881:
1747:
1348:
1079:
895:
550:
544:
538:
532:
225:
99:
2945:
520:
280:
229:
194:
129:
2687:
237:
109:
89:
252:, which was used to justify its elevation to a full species.
518:
119:
228:
had been collected by Eibert
Seimund, who became the
402:
344:10.2305/IUCN.UK.2021-1.RLTS.T84454980A84454984.en
3457:
392:(MSc thesis). Texas Tech University. p. 37.
1309:
504:
468:Field Guide to the Mammals of South-East Asia
381:
287:). They roost in small groups in palm trees,
1532:
2817:
1732:
784:
576:
511:
497:
41:
2603:
471:(2nd ed.). Bloomsbury. p. 220.
460:
458:
438:
428:
342:
236:. Seimmund had collected the holotype in
3181:East African little collared fruit bat (
388:Khan, Faisal Ali Anwarali (2008-04-23).
3215:
3070:Veldkamp's dwarf epauletted fruit bat (
464:
318:
3458:
455:
314:
312:
310:
308:
3337:
3336:
3037:Hayman's dwarf epauletted fruit bat (
2895:Lesser Angolan epauletted fruit bat (
2816:
2602:
1731:
1531:
1308:
783:
575:
492:
364:
3046:Peters' dwarf epauletted fruit bat (
2963:Buettikofer's epauletted fruit bat (
1465:New Britain naked-backed fruit bat (
973:Nusatenggara short-nosed fruit bat (
609:Broad-striped tube-nosed fruit bat (
387:
267:
3466:IUCN Red List least concern species
2913:East African epauletted fruit bat (
1384:Philippine naked-backed fruit bat (
1375:Beaufort's naked-backed fruit bat (
1366:Andersen's naked-backed fruit bat (
330:IUCN Red List of Threatened Species
305:
13:
2401:Little golden-mantled flying fox (
1420:Solomon's naked-backed fruit bat (
1393:Halmahera naked-backed fruit bat (
991:Indonesian short-nosed fruit bat (
645:Round-eared tube-nosed fruit bat (
367:"XII: Seven New Malaysian Mammals"
14:
3497:
2981:Franquet's epauletted fruit bat (
2931:Wahlberg's epauletted fruit bat (
2578:Sulawesi stripe-faced fruit bat (
1775:Giant golden-crowned flying fox (
1474:Greenish naked-backed fruit bat (
1438:Moluccan naked-backed fruit bat (
1411:Sulawesi naked-backed fruit bat (
708:Philippine tube-nosed fruit bat (
216:. He noted it as a subspecies of
2904:Ethiopian epauletted fruit bat (
2569:Mindoro stripe-faced fruit bat (
1456:Western naked-backed fruit bat (
1447:Panniet naked-backed fruit bat (
759:Steadfast tube-nosed fruit bat (
66:
3377:Balionycteris maculata seimundi
2877:Peters's epauletted fruit bat (
2868:Ansell's epauletted fruit bat (
2124:New Ireland masked flying fox (
1975:New Georgian monkey-faced Bat (
1939:Bougainville monkey-faced Bat (
1826:Small white-winged flying fox (
1429:Lesser naked-backed fruit bat (
982:Greater short-nosed fruit bat (
636:Mountain tube-nosed fruit bat (
371:Journal of the F. M. S. Museums
222:Balionycteris maculata seimundi
2886:Gambian epauletted fruit bat (
2859:Angolan epauletted fruit bat (
1948:Guadalcanal monkey-faced Bat (
937:Lesser short-nosed fruit bat (
690:Demonic tube-nosed fruit bat (
681:Malaita tube-nosed fruit bat (
663:Keast's tube-nosed fruit bat (
396:
358:
255:
1:
3172:São Tomé collared fruit bat (
1402:Biak naked-backed fruit bat (
750:Lesser tube-nosed fruit bat (
672:Island tube-nosed fruit bat (
654:Dragon tube-nosed fruit bat (
618:Common tube-nosed fruit bat (
298:
3486:Taxa named by C. Boden Kloss
3162:(Little collared fruit bats)
2922:Minor epauletted fruit bat (
2473:Philippine gray flying fox (
2070:Bismarck masked flying fox (
1131:Ratanaworabhan's fruit bat (
726:Umboi tube-nosed fruit bat (
430:10.1371/journal.pone.0179555
220:, with a scientific name of
7:
3190:Little collared fruit bat (
1848:(Straw-coloured fruit bats)
1657:New Caledonia blossom bat (
1065:Fischer's pygmy fruit bat (
200:
10:
3502:
2527:New Caledonia flying fox (
2277:Black-bearded flying fox (
1966:Montane monkey-faced Bat (
1957:Greater monkey-faced Bat (
1867:Straw-coloured fruit bat (
1498:Sulawesi harpy fruit bat (
1140:White-collared fruit bat (
1122:Javan tailless fruit bat (
1032:Rickart's dyak fruit bat (
874:Spotted-winged fruit bat (
850:Mindanao pygmy fruit bat (
3471:Mammals described in 1921
3345:
3294:
3264:Short-palated fruit bat (
3252:
3243:
3206:
3152:
3119:
3095:
3086:
3058:
3017:
2993:
2943:
2839:
2830:
2826:
2812:
2666:
2616:
2612:
2598:
2557:
2374:Geelvink Bay flying fox (
2205:Livingstone's fruit bat (
2106:Large Samoan flying fox (
2034:Small Samoan flying fox (
1987:
1927:
1915:Small-toothed fruit bat (
1903:
1891:Fijian monkey-faced Bat (
1879:
1838:
1817:White-winged flying fox (
1805:
1745:
1741:
1727:
1707:Moss-forest blossom bat (
1669:
1628:
1607:Black-bellied fruit bat (
1598:Fardoulis's blossom bat (
1586:
1565:Long-tongued nectar bat (
1555:(Long-tongued fruit bats)
1545:
1541:
1527:
1486:
1346:
1322:
1318:
1304:
1265:
1241:
1220:Greater musky fruit bat (
1200:
1176:
1152:
1101:
1077:
1053:
1023:Brooks's dyak fruit bat (
1003:
917:
893:
862:
838:
797:
793:
779:
738:
627:Pallas's tube-nosed bat (
589:
585:
571:
527:
519:Extant species of family
465:Francis, Charles (2019).
275:is widely distributed in
168:
161:
63:Scientific classification
61:
39:
30:
23:
3227:D'Anchieta's fruit bat (
2738:Leschenault's rousette (
2383:Grey-headed flying fox (
2286:Black-eared flying fox (
2169:Ontong Java flying fox (
2061:Ashy-headed flying fox (
1638:(Long-tailed fruit bats)
1574:Long-tongued fruit bat (
1356:(Bare-backed fruit bats)
1229:Lesser musky fruit bat (
905:Black-capped fruit bat (
717:Eastern tube-nosed Bat (
337:: e.T84454980A84454984.
218:spotted-winged fruit bat
3273:Campo-Ma’an fruit bat (
2482:Temminck's flying fox (
2455:Little red flying fox (
2437:Madagascan flying fox (
2187:Andersen's flying fox (
2151:Gilliard's flying fox (
2097:Spectacled flying fox (
1698:Halmahera blossom bat (
1648:Long-tailed fruit bat (
1089:Salim Ali's fruit bat (
946:Horsfield's fruit bat (
699:Lesser tube-nosed bat (
599:(tube-nosed fruit bats)
3347:Balionycteris seimundi
3027:(Dwarf epauleted bats)
2849:(Epauleted fruit bats)
2783:Long-haired rousette (
2765:Bare-backed rousette (
2464:Seychelles fruit bat (
2428:Rodrigues flying fox (
2313:Mauritian flying fox (
2241:Big-eared flying fox (
2007:Admiralty flying fox (
1858:Madagascan fruit bat (
1616:Woodford's fruit bat (
1253:Blanford's fruit bat (
927:(Dog-faced fruit bats)
884:Balionycteris seimundi
323:Balionycteris seimundi
273:Balionycteris seimundi
261:Balionycteris seimundi
206:Balionycteris seimundi
190:Balionycteris seimundi
172:Balionycteris seimundi
25:Balionycteris seimundi
2792:Madagascan rousette (
2711:Geoffroy's rousette (
2676:(rousette fruit bats)
2654:Philippine dawn bat (
2500:Vanikoro flying fox (
2410:Solomons flying fox (
2295:Caroline flying fox (
2079:Moluccan flying fox (
1757:Sulawesi flying fox (
365:Kloss, C. B. (1921).
3306:Zenker's fruit bat (
2972:Dobson's fruit bat (
2636:Greater nectar bat (
2491:Insular flying fox (
2419:Rennell flying fox (
2223:Okinawa flying fox (
2133:Nicobar flying fox (
2043:Vanuatu flying fox (
2016:Aldabra flying fox (
1689:Common blossom bat (
1334:Bulmer's fruit bat (
1113:Tailless fruit bat (
955:Peters's fruit bat (
3282:Pohle's fruit bat (
3131:Azagnyi fruit bat (
3005:Hammer-headed Bat (
2794:R. madagascariensis
2729:Egyptian rousette (
2720:Sulawesi rousette (
2509:Kosrae flying fox (
2365:Masked flying fox (
2340:Ornate flying fox (
2322:Temotu flying fox (
2268:Indian flying fox (
2259:Mariana fruit bat (
2250:Lesser flying fox (
2232:Lyle's flying fox (
2214:Lombok flying fox (
2115:Ryukyu flying fox (
2088:Makira flying fox (
1828:D. microleucopterus
1784:Palawan fruit bat (
1766:Talaud flying fox (
421:2017PLoSO..1279555L
277:Peninsular Malaysia
197:found in Malaysia.
33:Conservation status
3107:Angolan rousette (
2747:Linduan Rousette (
2713:R. amplexicaudatus
2693:Manado fruit bat (
2545:Dwarf flying fox (
2536:Pemba flying fox (
2518:Large flying fox (
2446:Samoa flying fox (
2392:Bonin flying fox (
2356:Pelew flying fox (
2304:Great flying fox (
2178:Small flying fox (
2142:Banks flying fox (
2025:Black flying fox (
1793:Sunda flying fox (
1210:(Musky fruit bats)
1013:(Dayak fruit bats)
964:Minute fruit bat (
817:Borneo fruit bat (
807:(Pygmy fruit bats)
3453:
3452:
3339:Taxon identifiers
3330:
3329:
3326:
3325:
3322:
3321:
3318:
3317:
3275:C. campomaanensis
3239:
3238:
3202:
3201:
3165:
3082:
3081:
3030:
2956:
2852:
2808:
2807:
2804:
2803:
2756:Comoro rousette (
2679:
2645:Cave nectar bat (
2629:
2626:(Dawn fruit bats)
2594:
2593:
2590:
2589:
2331:Ceram fruit bat (
2160:Gray flying fox (
2099:P. conspicillatus
2000:
1851:
1723:
1722:
1719:
1718:
1682:
1641:
1558:
1523:
1522:
1519:
1518:
1507:Harpy fruit bat (
1359:
1312:Harpyionycterinae
1300:
1299:
1296:
1295:
1277:Swift fruit bat (
1213:
1188:Dusky fruit bat (
1166:O. cartilagonodus
1164:Luzon fruit bat (
1041:Dayak fruit bat (
1016:
993:C. titthaecheilus
930:
907:C. melanocephalus
826:Pygmy fruit bat (
810:
775:
774:
771:
770:
602:
478:978-1-4729-3499-4
268:Range and habitat
186:
185:
56:
3493:
3481:Bats of Malaysia
3446:
3445:
3433:
3432:
3420:
3419:
3407:
3406:
3394:
3393:
3381:
3380:
3379:
3366:
3365:
3364:
3334:
3333:
3250:
3249:
3213:
3212:
3174:M. brachycephala
3164:
3163:
3159:
3140:Woermann's bat (
3093:
3092:
3029:
3028:
3024:
2955:
2954:
2953:(Epauleted bats)
2950:
2851:
2850:
2846:
2837:
2836:
2828:
2827:
2814:
2813:
2678:
2677:
2673:
2628:
2627:
2623:
2614:
2613:
2600:
2599:
2466:P. seychellensis
2385:P. poliocephalus
2306:P. neohibernicus
2196:Kei flying fox (
2081:P. chrysoproctus
2052:Aru flying fox (
2009:P. admiralitatum
1999:
1998:
1994:
1850:
1849:
1845:
1743:
1742:
1729:
1728:
1681:
1680:
1676:
1659:N. neocaledonica
1640:
1639:
1635:
1557:
1556:
1552:
1543:
1542:
1529:
1528:
1358:
1357:
1353:
1320:
1319:
1306:
1305:
1212:
1211:
1207:
1015:
1014:
1010:
929:
928:
924:
809:
808:
804:
795:
794:
781:
780:
601:
600:
596:
587:
586:
573:
572:
513:
506:
499:
490:
489:
483:
482:
462:
453:
452:
442:
432:
400:
394:
393:
385:
379:
378:
362:
356:
355:
353:
351:
346:
319:Tan, P. (2021).
316:
285:Riau Archipelago
242:genetic distance
193:is a species of
174:
154:B. seimundi
71:
70:
50:
45:
44:
21:
20:
3501:
3500:
3496:
3495:
3494:
3492:
3491:
3490:
3456:
3455:
3454:
3449:
3441:
3436:
3428:
3423:
3415:
3410:
3402:
3397:
3389:
3384:
3375:
3374:
3369:
3360:
3359:
3354:
3341:
3331:
3314:
3290:
3235:
3198:
3161:
3160:
3158:
3148:
3115:
3078:
3054:
3026:
3025:
3023:
3013:
2989:
2965:E. buettikoferi
2952:
2951:
2949:
2939:
2848:
2847:
2845:
2822:
2800:
2740:R. leschenaulti
2675:
2674:
2672:
2662:
2625:
2624:
2622:
2608:
2586:
2553:
2502:P. tuberculatus
2324:P. nitendiensis
2207:P. livingstonii
2153:P. gilliardorum
1996:
1995:
1993:
1983:
1923:
1899:
1875:
1847:
1846:
1844:
1834:
1801:
1737:
1715:
1678:
1677:
1675:
1665:
1637:
1636:
1634:
1624:
1582:
1554:
1553:
1551:
1537:
1515:
1489:Harpyionycteris
1482:
1449:D. pannietensis
1355:
1354:
1352:
1342:
1314:
1292:
1261:
1237:
1209:
1208:
1206:
1196:
1172:
1148:
1097:
1073:
1049:
1012:
1011:
1009:
999:
975:C. nusatenggara
926:
925:
923:
913:
889:
858:
852:A. paucidentata
834:
806:
805:
803:
789:
767:
734:
598:
597:
595:
581:
567:
523:
517:
487:
486:
479:
463:
456:
415:(7): e0179555.
401:
397:
386:
382:
363:
359:
349:
347:
317:
306:
301:
270:
258:
244:of 12% between
203:
182:
176:
170:
157:
65:
57:
46:
42:
35:
17:
12:
11:
5:
3499:
3489:
3488:
3483:
3478:
3473:
3468:
3451:
3450:
3448:
3447:
3434:
3421:
3408:
3395:
3382:
3367:
3351:
3349:
3343:
3342:
3328:
3327:
3324:
3323:
3320:
3319:
3316:
3315:
3313:
3312:
3302:
3300:
3292:
3291:
3289:
3288:
3279:
3270:
3260:
3258:
3247:
3245:Scotonycterini
3241:
3240:
3237:
3236:
3234:
3233:
3223:
3221:
3210:
3204:
3203:
3200:
3199:
3197:
3196:
3187:
3178:
3168:
3166:
3150:
3149:
3147:
3146:
3137:
3127:
3125:
3117:
3116:
3114:
3113:
3103:
3101:
3090:
3084:
3083:
3080:
3079:
3077:
3076:
3066:
3064:
3056:
3055:
3053:
3052:
3043:
3039:M. intermedius
3033:
3031:
3015:
3014:
3012:
3011:
3001:
2999:
2991:
2990:
2988:
2987:
2978:
2969:
2959:
2957:
2941:
2940:
2938:
2937:
2928:
2919:
2910:
2901:
2892:
2883:
2874:
2865:
2855:
2853:
2834:
2824:
2823:
2810:
2809:
2806:
2805:
2802:
2801:
2799:
2798:
2789:
2772:
2771:
2762:
2753:
2744:
2735:
2731:R. aegyptiacus
2726:
2717:
2700:
2699:
2682:
2680:
2664:
2663:
2661:
2660:
2651:
2642:
2632:
2630:
2610:
2609:
2596:
2595:
2592:
2591:
2588:
2587:
2585:
2584:
2575:
2571:S. mindorensis
2565:
2563:
2555:
2554:
2552:
2551:
2542:
2533:
2524:
2515:
2506:
2497:
2488:
2479:
2470:
2461:
2452:
2443:
2434:
2430:P. rodricensis
2425:
2416:
2407:
2398:
2389:
2380:
2371:
2362:
2353:
2346:
2337:
2328:
2319:
2310:
2301:
2292:
2283:
2279:P. melanopogon
2274:
2265:
2256:
2247:
2238:
2229:
2225:P. loochoensis
2220:
2216:P. lombocensis
2211:
2202:
2193:
2189:P. intermedius
2184:
2180:P. hypomelanus
2175:
2166:
2157:
2148:
2139:
2130:
2121:
2112:
2103:
2094:
2085:
2076:
2072:P. capistratus
2067:
2058:
2049:
2040:
2031:
2022:
2018:P. aldabrensis
2013:
2003:
2001:
1997:(flying foxes)
1985:
1984:
1982:
1981:
1972:
1963:
1954:
1945:
1935:
1933:
1925:
1924:
1922:
1921:
1911:
1909:
1901:
1900:
1898:
1897:
1887:
1885:
1877:
1876:
1874:
1873:
1864:
1854:
1852:
1836:
1835:
1833:
1832:
1823:
1819:D. leucopterus
1813:
1811:
1803:
1802:
1800:
1799:
1790:
1781:
1772:
1763:
1753:
1751:
1739:
1738:
1725:
1724:
1721:
1720:
1717:
1716:
1714:
1713:
1704:
1695:
1685:
1683:
1679:(blossom bats)
1667:
1666:
1664:
1663:
1654:
1644:
1642:
1626:
1625:
1623:
1622:
1613:
1604:
1594:
1592:
1584:
1583:
1581:
1580:
1571:
1561:
1559:
1539:
1538:
1535:Macroglossinae
1525:
1524:
1521:
1520:
1517:
1516:
1514:
1513:
1504:
1494:
1492:
1484:
1483:
1481:
1480:
1471:
1462:
1453:
1444:
1440:D. moluccensis
1435:
1426:
1417:
1408:
1399:
1390:
1381:
1372:
1362:
1360:
1344:
1343:
1341:
1340:
1330:
1328:
1316:
1315:
1302:
1301:
1298:
1297:
1294:
1293:
1291:
1290:
1283:
1273:
1271:
1263:
1262:
1260:
1259:
1249:
1247:
1239:
1238:
1236:
1235:
1226:
1216:
1214:
1198:
1197:
1195:
1194:
1184:
1182:
1174:
1173:
1171:
1170:
1160:
1158:
1150:
1149:
1147:
1146:
1137:
1128:
1119:
1109:
1107:
1099:
1098:
1096:
1095:
1085:
1083:
1075:
1074:
1072:
1071:
1061:
1059:
1051:
1050:
1048:
1047:
1038:
1029:
1019:
1017:
1001:
1000:
998:
997:
988:
979:
970:
961:
957:C. luzoniensis
952:
948:C. horsfieldii
943:
933:
931:
915:
914:
912:
911:
901:
899:
891:
890:
888:
887:
880:
870:
868:
860:
859:
857:
856:
846:
844:
836:
835:
833:
832:
823:
813:
811:
791:
790:
777:
776:
773:
772:
769:
768:
766:
765:
756:
746:
744:
736:
735:
733:
732:
723:
714:
705:
696:
687:
683:N. malaitensis
678:
669:
660:
651:
642:
633:
624:
615:
605:
603:
583:
582:
569:
568:
566:
565:
559:
557:Laurasiatheria
553:
547:
541:
535:
528:
525:
524:
516:
515:
508:
501:
493:
485:
484:
477:
454:
395:
380:
357:
303:
302:
300:
297:
269:
266:
257:
254:
250:B. m. maculata
246:B. m. seimundi
214:C. Boden Kloss
202:
199:
184:
183:
177:
166:
165:
159:
158:
151:
149:
145:
144:
137:
133:
132:
127:
123:
122:
117:
113:
112:
107:
103:
102:
97:
93:
92:
87:
83:
82:
77:
73:
72:
59:
58:
40:
37:
36:
31:
28:
27:
16:Species of bat
15:
9:
6:
4:
3:
2:
3498:
3487:
3484:
3482:
3479:
3477:
3474:
3472:
3469:
3467:
3464:
3463:
3461:
3444:
3439:
3435:
3431:
3426:
3422:
3418:
3413:
3409:
3405:
3400:
3396:
3392:
3387:
3383:
3378:
3372:
3368:
3363:
3357:
3353:
3352:
3350:
3348:
3344:
3340:
3335:
3311:
3309:
3304:
3303:
3301:
3299:
3298:
3297:Scotonycteris
3293:
3287:
3285:
3280:
3278:
3276:
3271:
3269:
3267:
3262:
3261:
3259:
3257:
3256:
3251:
3248:
3246:
3242:
3232:
3230:
3225:
3224:
3222:
3220:
3219:
3214:
3211:
3209:
3205:
3195:
3193:
3188:
3186:
3184:
3179:
3177:
3175:
3170:
3169:
3167:
3157:
3156:
3151:
3145:
3143:
3138:
3136:
3134:
3129:
3128:
3126:
3124:
3123:
3122:Megaloglossus
3118:
3112:
3110:
3109:L. angolensis
3105:
3104:
3102:
3100:
3099:
3098:Lissonycteris
3094:
3091:
3089:
3085:
3075:
3073:
3072:N. veldkampii
3068:
3067:
3065:
3063:
3062:
3057:
3051:
3049:
3044:
3042:
3040:
3035:
3034:
3032:
3022:
3021:
3020:Micropteropus
3016:
3010:
3008:
3007:H. monstrosus
3003:
3002:
3000:
2998:
2997:
2992:
2986:
2984:
2979:
2977:
2975:
2970:
2968:
2966:
2961:
2960:
2958:
2948:
2947:
2942:
2936:
2934:
2929:
2927:
2925:
2920:
2918:
2916:
2911:
2909:
2907:
2902:
2900:
2898:
2893:
2891:
2889:
2884:
2882:
2880:
2875:
2873:
2871:
2866:
2864:
2862:
2861:E. angolensis
2857:
2856:
2854:
2844:
2843:
2838:
2835:
2833:
2829:
2825:
2821:
2820:Epomophorinae
2815:
2811:
2797:
2795:
2790:
2788:
2786:
2780:
2779:
2778:Stenonycteris
2774:
2773:
2770:
2768:
2767:R. spinalatus
2763:
2761:
2759:
2758:R. obliviosus
2754:
2752:
2750:
2749:R. linduensis
2745:
2743:
2741:
2736:
2734:
2732:
2727:
2725:
2723:
2722:R. celebensis
2718:
2716:
2714:
2708:
2707:
2702:
2701:
2698:
2696:
2690:
2689:
2684:
2683:
2681:
2671:
2670:
2665:
2659:
2657:
2652:
2650:
2648:
2643:
2641:
2639:
2634:
2633:
2631:
2621:
2620:
2615:
2611:
2607:
2601:
2597:
2583:
2581:
2576:
2574:
2572:
2567:
2566:
2564:
2562:
2561:
2556:
2550:
2548:
2543:
2541:
2539:
2538:P. voeltzkowi
2534:
2532:
2530:
2525:
2523:
2521:
2516:
2514:
2512:
2507:
2505:
2503:
2498:
2496:
2494:
2489:
2487:
2485:
2484:P. temminckii
2480:
2478:
2476:
2471:
2469:
2467:
2462:
2460:
2458:
2457:P. scapulatus
2453:
2451:
2449:
2444:
2442:
2440:
2435:
2433:
2431:
2426:
2424:
2422:
2417:
2415:
2413:
2408:
2406:
2404:
2399:
2397:
2395:
2390:
2388:
2386:
2381:
2379:
2377:
2372:
2370:
2368:
2367:P. personatus
2363:
2361:
2359:
2358:P. pelewensis
2354:
2352:
2351:
2347:
2345:
2343:
2338:
2336:
2334:
2329:
2327:
2325:
2320:
2318:
2316:
2311:
2309:
2307:
2302:
2300:
2298:
2297:P. molossinus
2293:
2291:
2289:
2284:
2282:
2280:
2275:
2273:
2271:
2266:
2264:
2262:
2257:
2255:
2253:
2248:
2246:
2244:
2239:
2237:
2235:
2230:
2228:
2226:
2221:
2219:
2217:
2212:
2210:
2208:
2203:
2201:
2199:
2194:
2192:
2190:
2185:
2183:
2181:
2176:
2174:
2172:
2167:
2165:
2163:
2158:
2156:
2154:
2149:
2147:
2145:
2140:
2138:
2136:
2131:
2129:
2127:
2122:
2120:
2118:
2117:P. dasymallus
2113:
2111:
2109:
2104:
2102:
2100:
2095:
2093:
2091:
2086:
2084:
2082:
2077:
2075:
2073:
2068:
2066:
2064:
2059:
2057:
2055:
2050:
2048:
2046:
2041:
2039:
2037:
2032:
2030:
2028:
2023:
2021:
2019:
2014:
2012:
2010:
2005:
2004:
2002:
1992:
1991:
1986:
1980:
1978:
1973:
1971:
1969:
1964:
1962:
1960:
1955:
1953:
1951:
1946:
1944:
1942:
1937:
1936:
1934:
1932:
1931:
1926:
1920:
1918:
1913:
1912:
1910:
1908:
1907:
1902:
1896:
1894:
1889:
1888:
1886:
1884:
1883:
1878:
1872:
1870:
1865:
1863:
1861:
1856:
1855:
1853:
1843:
1842:
1837:
1831:
1829:
1824:
1822:
1820:
1815:
1814:
1812:
1810:
1809:
1804:
1798:
1796:
1791:
1789:
1787:
1782:
1780:
1778:
1773:
1771:
1769:
1764:
1762:
1760:
1759:A. celebensis
1755:
1754:
1752:
1750:
1749:
1744:
1740:
1736:
1730:
1726:
1712:
1710:
1705:
1703:
1701:
1696:
1694:
1692:
1687:
1686:
1684:
1674:
1673:
1668:
1662:
1660:
1655:
1653:
1651:
1650:N. macdonaldi
1646:
1645:
1643:
1633:
1632:
1627:
1621:
1619:
1614:
1612:
1610:
1605:
1603:
1601:
1600:M. fardoulisi
1596:
1595:
1593:
1591:
1590:
1585:
1579:
1577:
1572:
1570:
1568:
1563:
1562:
1560:
1550:
1549:
1544:
1540:
1536:
1530:
1526:
1512:
1510:
1509:H. whiteheadi
1505:
1503:
1501:
1500:H. celebensis
1496:
1495:
1493:
1491:
1490:
1485:
1479:
1477:
1472:
1470:
1468:
1467:D. praedatrix
1463:
1461:
1459:
1454:
1452:
1450:
1445:
1443:
1441:
1436:
1434:
1432:
1427:
1425:
1423:
1418:
1416:
1414:
1409:
1407:
1405:
1400:
1398:
1396:
1391:
1389:
1387:
1382:
1380:
1378:
1373:
1371:
1369:
1364:
1363:
1361:
1351:
1350:
1345:
1339:
1337:
1332:
1331:
1329:
1327:
1326:
1321:
1317:
1313:
1307:
1303:
1289:
1288:
1287:T. suhaniahae
1284:
1282:
1280:
1279:T. nigrescens
1275:
1274:
1272:
1270:
1269:
1264:
1258:
1256:
1251:
1250:
1248:
1246:
1245:
1240:
1234:
1232:
1227:
1225:
1223:
1218:
1217:
1215:
1205:
1204:
1199:
1193:
1191:
1186:
1185:
1183:
1181:
1180:
1175:
1169:
1167:
1162:
1161:
1159:
1157:
1156:
1151:
1145:
1143:
1138:
1136:
1134:
1129:
1127:
1125:
1120:
1118:
1116:
1111:
1110:
1108:
1106:
1105:
1100:
1094:
1092:
1087:
1086:
1084:
1082:
1081:
1076:
1070:
1068:
1063:
1062:
1060:
1058:
1057:
1056:Haplonycteris
1052:
1046:
1044:
1039:
1037:
1035:
1030:
1028:
1026:
1021:
1020:
1018:
1008:
1007:
1002:
996:
994:
989:
987:
985:
980:
978:
976:
971:
969:
967:
962:
960:
958:
953:
951:
949:
944:
942:
940:
939:C. brachyotis
935:
934:
932:
922:
921:
916:
910:
908:
903:
902:
900:
898:
897:
892:
886:
885:
881:
879:
877:
872:
871:
869:
867:
866:
865:Balionycteris
861:
855:
853:
848:
847:
845:
843:
842:
837:
831:
829:
824:
822:
820:
815:
814:
812:
802:
801:
796:
792:
788:
782:
778:
764:
762:
757:
755:
753:
748:
747:
745:
743:
742:
741:Paranyctimene
737:
731:
729:
724:
722:
720:
715:
713:
711:
706:
704:
702:
697:
695:
693:
688:
686:
684:
679:
677:
675:
670:
668:
666:
661:
659:
657:
656:N. draconilla
652:
650:
648:
643:
641:
639:
634:
632:
630:
629:N. cephalotes
625:
623:
621:
620:N. albiventer
616:
614:
612:
607:
606:
604:
594:
593:
588:
584:
580:
574:
570:
564:
560:
558:
554:
552:
548:
546:
542:
540:
536:
534:
530:
529:
526:
522:
514:
509:
507:
502:
500:
495:
494:
491:
480:
474:
470:
469:
461:
459:
450:
446:
441:
436:
431:
426:
422:
418:
414:
410:
406:
399:
391:
384:
376:
372:
368:
361:
345:
340:
336:
332:
331:
326:
324:
315:
313:
311:
309:
304:
296:
294:
290:
289:termite nests
286:
282:
278:
274:
265:
262:
253:
251:
247:
243:
239:
235:
232:for the name
231:
227:
223:
219:
215:
211:
207:
198:
196:
192:
191:
180:
175:
173:
167:
164:
163:Binomial name
160:
156:
155:
150:
147:
146:
143:
142:
141:Balionycteris
138:
135:
134:
131:
128:
125:
124:
121:
118:
115:
114:
111:
108:
105:
104:
101:
98:
95:
94:
91:
88:
85:
84:
81:
78:
75:
74:
69:
64:
60:
54:
49:
48:Least Concern
38:
34:
29:
26:
22:
19:
3346:
3307:
3295:
3283:
3274:
3265:
3255:Casinycteris
3253:
3229:P. anchietae
3228:
3216:
3191:
3182:
3173:
3153:
3142:M. woermanni
3141:
3132:
3120:
3108:
3096:
3088:Myonycterini
3071:
3061:Nanonycteris
3059:
3047:
3038:
3018:
3006:
2996:Hypsignathus
2994:
2983:E. franqueti
2982:
2973:
2964:
2944:
2933:E. wahlbergi
2932:
2923:
2914:
2905:
2896:
2888:E. gambianus
2887:
2879:E. crypturus
2878:
2869:
2860:
2840:
2832:Epomophorini
2793:
2784:
2775:
2766:
2757:
2748:
2739:
2730:
2721:
2712:
2703:
2694:
2685:
2667:
2655:
2646:
2637:
2617:
2579:
2570:
2560:Styloctenium
2558:
2547:P. woodfordi
2546:
2537:
2528:
2519:
2510:
2501:
2492:
2483:
2475:P. speciosus
2474:
2465:
2456:
2448:P. samoensis
2447:
2438:
2429:
2420:
2411:
2402:
2394:P. pselaphon
2393:
2384:
2375:
2366:
2357:
2350:P. pelagicus
2348:
2341:
2332:
2323:
2314:
2305:
2296:
2288:P. melanotus
2287:
2278:
2269:
2261:P. mariannus
2260:
2252:P. mahaganus
2251:
2242:
2233:
2224:
2215:
2206:
2197:
2188:
2179:
2170:
2161:
2152:
2143:
2134:
2125:
2116:
2107:
2098:
2089:
2080:
2071:
2062:
2053:
2045:P. anetianus
2044:
2036:P. allenorum
2035:
2026:
2017:
2008:
1988:
1976:
1967:
1959:P. flanneryi
1958:
1949:
1940:
1928:
1916:
1904:
1893:M. acrodonta
1892:
1880:
1868:
1860:E. dupreanum
1859:
1839:
1827:
1818:
1806:
1794:
1785:
1776:
1767:
1758:
1746:
1735:Pteropodinae
1708:
1700:S. carolinae
1699:
1691:S. australis
1690:
1672:Syconycteris
1670:
1658:
1649:
1629:
1618:M. woodfordi
1617:
1608:
1599:
1589:Melonycteris
1587:
1575:
1566:
1548:Macroglossus
1546:
1508:
1499:
1487:
1475:
1466:
1457:
1448:
1439:
1430:
1421:
1412:
1403:
1395:D. crenulata
1394:
1385:
1377:D. beauforti
1376:
1368:D. anderseni
1367:
1347:
1335:
1323:
1285:
1278:
1266:
1255:S. blanfordi
1254:
1242:
1230:
1221:
1201:
1189:
1177:
1165:
1153:
1141:
1132:
1123:
1115:M. ecaudatus
1114:
1102:
1091:L. salimalii
1090:
1078:
1066:
1054:
1043:D. spadiceus
1042:
1033:
1024:
1004:
992:
983:
974:
965:
956:
947:
938:
918:
906:
894:
883:
882:
875:
863:
851:
841:Alionycteris
839:
827:
818:
798:
787:Cynopterinae
760:
751:
739:
728:N. vizcaccia
727:
719:N. robinsoni
718:
709:
700:
691:
682:
673:
664:
655:
646:
637:
628:
619:
610:
590:
579:Nyctimeninae
555:Superorder:
549:Infraclass:
521:Pteropodidae
467:
412:
408:
398:
389:
383:
374:
370:
360:
348:. Retrieved
334:
328:
322:
272:
271:
260:
259:
249:
245:
233:
221:
205:
204:
189:
188:
187:
171:
169:
153:
152:
140:
130:Pteropodidae
24:
18:
3371:Wikispecies
3284:C. ophiodon
3266:C. argynnis
3192:M. torquata
3155:Myonycteris
3048:M. pusillus
2906:E. labiatus
2842:Epomophorus
2606:Rousettinae
2580:S. wallacei
2520:P. vampyrus
2493:P. tonganus
2421:P. rennelli
2333:P. ocularis
2243:P. macrotis
2198:P. keyensis
2171:P. howensis
2144:P. fundatus
2135:P. faunulus
2090:P. cognatus
2063:P. caniceps
2054:P. aruensis
1808:Desmalopex
1795:A. mackloti
1786:A. leucotis
1609:M. melanops
1576:M. sobrinus
1386:D. chapmani
1336:A. bulmerae
1203:Ptenochirus
1155:Otopteropus
1142:M. wetmorei
1133:M. niphanae
1124:M. kusnotoi
1067:H. fischeri
1034:D. rickarti
1006:Dyacopterus
876:B. maculata
819:A. aequalis
647:N. cyclotis
350:15 December
256:Description
212:in 1921 by
3460:Categories
3308:S. zenkeri
3183:M. relicta
3133:M. azagnyi
2974:E. dobsoni
2915:E. minimus
2897:E. grandis
2870:E. anselli
2818:Subfamily
2785:R. lanosus
2656:E. robusta
2647:E. spelaea
2619:Eonycteris
2604:Subfamily
2529:P. vetulus
2511:P. ualanus
2412:P. rayneri
2403:P. pumilus
2342:P. ornatus
2162:P. griseus
2126:P. ennisae
1968:P. pulchra
1930:Pteralopex
1777:A. jubatus
1768:A. humilis
1733:Subfamily
1631:Notopteris
1567:M. minimus
1533:Subfamily
1476:D. viridis
1458:D. peronii
1422:D. inermis
1413:D. exoleta
1310:Subfamily
1268:Thoopterus
1025:D. brooksi
966:C. minutus
920:Cynopterus
785:Subfamily
701:N. minutus
692:N. masalai
638:N. certans
577:Subfamily
563:Chiroptera
299:References
120:Chiroptera
3362:Q20905623
3208:Plerotini
2776:Subgenus
2706:Rousettus
2704:Subgenus
2695:R. bidens
2686:Subgenus
2669:Rousettus
2376:P. pohlei
2270:P. medius
2027:P. alecto
1950:P. atrata
1941:P. anceps
1917:N. frosti
1906:Neopteryx
1869:E. helvum
1709:S. hobbit
1404:D. emersa
1325:Aproteles
1244:Sphaerias
1222:P. jagori
1190:P. lucasi
1179:Penthetor
1104:Megaerops
984:C. sphinx
828:A. alecto
800:Aethalops
752:P. raptor
710:N. rabori
665:N. keasti
592:Nyctimene
531:Kingdom:
293:epiphytic
210:described
148:Species:
86:Kingdom:
80:Eukaryota
3476:Megabats
3443:13800032
3417:84454980
3356:Wikidata
3218:Plerotes
2924:E. minor
2638:E. major
2439:P. rufus
2315:P. niger
2234:P. lylei
1990:Pteropus
1882:Mirimiri
1748:Acerodon
1431:D. minor
1349:Dobsonia
1231:P. minor
1080:Latidens
896:Chironax
761:P. tenax
674:N. major
611:N. aello
551:Eutheria
545:Mammalia
539:Chordata
537:Phylum:
533:Animalia
449:28742835
409:PLOS ONE
283:and the
234:seimundi
226:holotype
201:Taxonomy
126:Family:
110:Mammalia
100:Chordata
96:Phylum:
90:Animalia
76:Domain:
53:IUCN 3.1
3430:1004361
3391:4265623
2946:Epomops
2108:P. coxi
1977:P. taki
1841:Eidolon
561:Order:
543:Class:
440:5526618
417:Bibcode
281:Sumatra
230:epithet
195:megabat
136:Genus:
116:Order:
106:Class:
51: (
3404:947765
2688:Boneia
475:
447:
437:
377:: 229.
238:Pahang
224:. The
181:, 1921
179:Kloss
3412:IUCN
3399:ITIS
3386:GBIF
473:ISBN
445:PMID
352:2023
335:2021
248:and
208:was
3438:MSW
3425:MDD
435:PMC
425:doi
339:doi
3462::
3440::
3427::
3414::
3401::
3388::
3373::
3358::
2781::
2709::
2691::
457:^
443:.
433:.
423:.
413:12
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407:.
375:10
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307:^
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512:e
505:t
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55:)
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