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Archer City Formation

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2644: 1183: 2672: 1925: 246: 209: 2052: 1159: 1611: 598:. Some publications have labelled the Nocona Formation as the "Upper Archer City Formation", thus considering the Archer City Formation to be a unit of the Wichita Group. This proposal is not widely agreed upon. The Cisco Group and its constituent formations have been restricted to marine strata further south and west. Of these strata, the Archer City Formation is roughly equivalent in time to the uppermost Pueblo Formation, the Moran Formation, the 1210: 1991: 2022: 2216: 1466: 1058: 1723: 1958: 1546: 2658: 191: 2181: 1279: 1094: 1312: 1414: 1822: 1895: 1576: 707:. These data points suggest soil temperatures of 31–35 Β°C (88–95 Β°F), significantly warmer than the cool and wet conditions of the Markley Formation. This fits with climate evidence from rapid sedimentary changes between the formations, as well as an increasingly patchy distribution of high-humidity flora. The Archer City Formation would have had a hot 813:
Four different types of paleosol (labelled types E through H) have been identified in the Archer City Formation. Three (F, G, and H) first appear in this formation, while type E paleosols are retained from the underlying Markley Formation. All four continue to persist through the Texas Lower Permian
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Based on correlations with coastal strata further west, the Carboniferous-Permian boundary is positioned near the top of the Markley Formation. This suggests that the Archer City Formation occupies most of the Asselian (299–293 Ma), the first global stage of the Permian Period. The plant fossils of
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The Kola Switch site is divided into three different layers in a 1.4 meters (4.6 feet) thick interval, each with a different set of fossils. The lower bed is pale upwards-fining siltstone, the middle bed is dark shale with carbonaceous impressions, and the upper bed is greenish claystone.
2837:"Morphology and distribution of fossil soils in the Permo-Pennsylvanian Wichita and Bowie Groups, north-central Texas, USA: implications for western equatorial Pangean palaeoclimate during icehouse-greenhouse transition: Permo-Carboniferous palaeoclimate from palaeosols" 852:. They develop in stable forested uplands with a low water table and a semi-arid to subhumid climate (~70 cm of annual precipitation). Type G paleosols are similar, with sandy vertical cracks and abundant calcareous nodules. They qualify as 797:
Some sandstone beds are isolated straight channel fills, without corresponding riverbank deposits. These straight beds may reach 30 meters in width (though most are much narrower) and several kilometers in length. Coarser beds preserve sinuous
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Plant fossils in the Archer City Formation are concentrated at two sites in Clay County: Kola Switch and Sanzenbacher Ranch. Both sites were discovered in 1940–1941 by Adolph H. Witte, a local geologist working for the
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The Sanzenbacher flora occupies a narrow layer of greyish claystone coarsening upwards to buff-colored siltstone. The most abundant plants are dry-soil species of seed ferns and conifers. Seed fern species include
3091:"Paleoecological and paleoenvironmental interpretation of three successive macrofloras and palynofloras from the Kola Switch locality, lower Permian (Archer City Formation, Bowie Group) of Clay County, Texas, USA" 3477:
Permo-Carboniferous Vertebrate Paleontology, Lithostratigraphy, and Depositional Environments of North-Central Texas. Field Trip Guidebook No. 2, 49th Annual Meeting of the Society of Vertebrate Paleontology
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ponds. Most of the pond bonebeds are too small to continuously support an ecosystem of large aquatic vertebrates, suggesting that the carcasses may have been washed in from larger bodies of water.
856:, inundated and dried at a seasonal frequency along the margins of a floodplain. Type H paleosols have an even greater proportion of calcareous minerals, forming blocky crusts interspersed among 802:
while finer channels develop small climbing ripples towards the top. Straight channels would have formed during high-energy erosion events, but the climate was probably too moist for frequent
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ferns in the Bursum Formation and a greater variety of swamp foliage in the Neal Ranch Formation. Many of the broader groups prevalent in these formations are equally abundant in the
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plants are less common but far from rare, giving Sanzenbacher a 'mixed' ecosystem character similar to, but drier than, the upper bed of Kola Switch. Marattialean ferns such as
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Hentz, Tucker F. (1988). "Lithostratigraphy and Paleoenvironments of Upper Paleozoic Continental Red Beds, North-central Texas: Bowie (new) and Wichita (revised) Groups".
620: 603: 3688:"New Upper Pennsylvanian armored dissorophid records (Temnospondyli, Dissorophoidea) from the U.S. midcontinent and the stratigraphic distributions of dissorophids" 2280:
The floral diversity of the Archer City Formation strongly overlaps with other Asselian-Sakmarian formations in the southwestern United States. These include the
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counties. Many fossils are rare fragments of species which are better-preserved in younger parts of the red beds. Nevertheless, complete skeletons and important
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in the west and patches of steadily eroding mountains in the north and east. Some of the mountains are still standing in the present day, such as the
2089: 2517:). Otherwise, the floral diversity of the upper bed encompasses the same species as the other Kola Switch beds. The upper bed palynoflora is mostly 2419: 2318:. This is most likely a consequence of local environmental factors rather than a turnover of plant diversity at the Permo-Carboniferous boundary. 2624: 2574: 2469: 2095: 826:
in the type E variety. The Archer City Formation mostly lacks the high-humidity paleosol varieties of the Markley Formation (types A through D).
308:. The Archer City Formation was not named as a unique geological unit until the late 1980s. Older studies generally labelled its outcrops as the 3939:"Returning to the roots: resolution, reproducibility, and robusticity in the phylogenetic inference of Dissorophidae (Amphibia: Temnospondyli)" 3490: 2358: 353: 2836: 680:
northward and eastward. The major bonebed sites of the Archer City Formation were probably 40–60 km (25–37 mi) away from the coast.
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the Archer City Formation also resemble those of Asselian Europe. The upper part of the Archer City Formation may extend into the succeeding
583: 124: 2618: 1496: 4068:"Trimerorhachis (Amphibia: Temnospondyli) from the Lower Permian of Texas and New Mexico: cranial osteology, taxonomy and biostratigraphy" 1788:
A trimerorhachid dvinosaur. Reports from the Archer City Formation are unsubstantiated, though it is known from time-equivalent strata in
2545:. The combination of fossils from the lower and middle bed is peculiar considering their strongly different depositional environments. 2306: 347: 3805:"The Appendicular Skeleton of Eryops Megacephalus Cope, 1877 (Temnospondyli: Eryopoidea) from the Lower Permian of North America" 4301:"Possible origin of the xenacanth sharks Orthacanthus texensis and Orthacanthus platypternus in the Lower Permian of Texas, USA" 2873: 3313: 3332: 3294:"From wetlands to wet spots: Environmental tracking and the fate of Carboniferous elements in Early Permian tropical floras" 3804: 2490:
spp. (marattialean spores) are even more dominant. The middle bed represents foliage washed into a nearby stagnant pond.
2274: 4435: 4396: 4109: 3473:"Stratigraphic distribution of tetrapods in the Bowie and Wichita Groups, Permo-Carboniferous of north-central Texas" 460:
spp. make up a small component of the terrestrial fauna. Amphibians are common in every niche, with examples such as
3250: 845:, patterned with deep root casts and numerous horizons, including intermediate layers with calcareous nodules and 579:
in the west. It is most broadly exposed in Clay and Archer counties, including under its namesake of Archer City.
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of the early Sakmarian. The Asselian and Sakmarian global stages are equivalent to the early-middle parts of the
2310:) are common in both regions. Conversely, there is little similarity with swampy coal-bearing areas such as the 2685: 649: 289: 3820: 2632:) and spores are rarer. Still, the palynomorph diversity of Sanzenbacher is much greater than at Kola Switch. 4372:
DiMichele, William A.; Hook, Robert W.; Kerp, Hans; Hotton, Carol L.; Looy, Cindy V.; Chaney, Dan S. (2018),
3915: 2965:, Geological Society of America Special Papers, vol. 28, Geological Society of America, pp. 1–534, 2266: 711:
climate, with moist or swampy areas shrinking to the vicinity of riverbanks and ponds during the dry season.
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Type E paleosols are silty or fine sandy soils with persistent mottling, hematite nodules, and few distinct
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is based on more fragmentary fossils and may belong to an unrelated genus, perhaps a trematopid related to
2390:(horsetails) are very rare, despite the evidence for deposition in a slow stream channel. The most common 3364:"Apsisaurus witteri from the Lower Permian of Texas: yet another small varanopid synapsid, not a diapsid" 746:
beds and paleosols are also common. Fossils and other organic debris are concentrated into rare greyish
4340:"Revised generic diagnoses of the fossil fishes Megalichthys and Ectosteorhachis (family Osteolepidae)" 2269:. Though Witte's original collections have been lost, he sampled the sites further in 1961, along with 576: 782:, with channels only a few meters wide. Other common sandstone features include subdued, fine-grained 4339: 3661: 3575: 2984: 4028: 3505: 3363: 3209:"Late Paleozoic continental warming of a cold tropical basin and floristic change in western Pangea" 3162: 2960: 708: 328: 114: 4300: 3727:"Revision of the Early Permian dissorophid 'Dissorophus' angustus (Temnospondyli: Dissorophoidea)" 3687: 3293: 3994: 560: 285: 723:, the Archer City and Nocona formations are nearly indistinguishable, with equivalent sets of 4029:"A new trematopsid amphibian (Temnospondyli: Dissorophoidea) from the Lower Permian of Texas" 2695: 2649: 1117: 4411: 3995:"Reappraisal of the Early Permian amphibamid Tersomius texensis and some referred material" 3686:
May, William; Huttenlocker, Adam K.; Pardo, Jason D.; Benca, Jeff; Small, Bryan J. (2011).
2771:"Early Permian depositional environments and pond bonebeds in central Archer County, Texas" 2289: 677: 568: 529: 409: 336: 278: 676:
oil field. Rising sea levels through the Early Permian allowed the coastline to gradually
292:, introducing a tropical ecosystem which will persist in the area through the rest of the 8: 3333:"The osteology of a Lower Permian eosuchian from Texas and a review of diapsid phylogeny" 2677: 1182: 775: 572: 259: 4388: 4233: 4206: 3973: 3765: 3629: 3601: 3484: 3411: 3348: 3123: 3090: 3043:
The University of Texas at Austin, Bureau of Economic Geology, Report of Investigations
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of 0Β° to 5Β° N. Temperature estimates can be inferred for the Archer City Formation via
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deposits. The crevasse splays are often associated with plant fossils which have been
556:. The formation and its stratigraphic placement were formalized in text a year later. 4392: 4320: 4274: 4224:
Zidek, Jiri (1978). "New Chondrichthyan Spines from the Late Paleozoic of Oklahoma".
4198: 4159: 4087: 4067: 4048: 3960: 3893: 3824: 3785: 3746: 3726: 3707: 3621: 3527: 3472: 3383: 3309: 3251:"Age assignment of the Pennsylvanian-Early Permian succession of North Central Texas" 3231: 3182: 3128: 3110: 2924: 2855: 2790: 2786: 2770: 2315: 1863: 1320: 1302: 724: 661: 599: 456: 335:
and plant assemblages ranging from dry upland forests to moist riverside galleries. "
4083: 3874:"Osteology of the temnospondyl Neldasaurus and the evolution of basal dvinosaurians" 3014: 672:. The marine basin is formally known as the Midland Basin, a major component of the 4384: 4312: 4264: 4194: 4190: 4151: 4079: 4044: 4040: 4006: 3993:
Maddin, Hillary C.; FrΓΆbisch, Nadia B.; Evans, David C.; Milner, Andrew R. (2013).
3968: 3950: 3885: 3816: 3777: 3738: 3699: 3613: 3517: 3375: 3344: 3301: 3223: 3207:
Tabor, Neil J.; DiMichele, William A.; MontaΓ±ez, Isabel P.; Chaney, Dan S. (2013).
3174: 3118: 3102: 2966: 2916: 2851: 2782: 2281: 1601: 1051: 819: 787: 751: 739: 591: 541: 533: 313: 305: 134: 3742: 3178: 668:, while others have been fully eroded or buried, such as the Texan portion of the 4316: 3703: 3379: 3089:
DiMichele, William A.; Hotton, Carol L.; Looy, Cindy V.; Hook, Robert W. (2019).
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Conversely, the middle bed is almost entirely marattialean ferns. These include
837:, young soils corresponding to isolated, frequently disturbed areas with a high 4010: 3292:
DiMichele, William A.; Tabor, Neil J.; Chaney, Dan S.; Nelson, W. John (2006),
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The upper bed has the greatest portion of sphenophytes, not just sphenophylls (
2208: 2118: 1918: 1775: 1083: 783: 763: 700: 532:, starting in the late 1920s. At the time, these bonebeds were assigned to the 417: 4252: 3548:
Llewellyn Ivor Price (1937). "Two new cotylosaurs from the Permian of Texas".
3305: 4429: 4409: 4324: 4278: 4202: 4163: 4091: 4052: 3964: 3897: 3828: 3789: 3750: 3711: 3625: 3531: 3387: 3235: 3186: 3114: 2928: 2794: 2589: 2456: 2311: 2285: 2101: 2066: 2014: 1948: 1688: 1437: 1365: 1335: 1287: 815: 653: 625: 595: 450: 381: 297: 214: 71: 31: 27: 245: 3132: 2663: 2335: 2204: 2189: 2083: 1815: 1731: 1642: 1422: 1407: 1272: 1269: 1068: 830: 633: 549: 525: 495: 490: 432: 341: 4269: 3504:
Lucas, Spencer G.; Rinehart, Larry F.; Celeskey, Matthew D. (2018-11-14).
2920: 2502: 2450: 2387: 2379: 2297: 2113: 2071: 1999: 1984: 1933: 1924: 1878: 1701: 1604: 1584: 1512: 1474: 1447: 1158: 1102: 1021: 940: 873: 838: 803: 637: 553: 545: 466: 462: 375: 364: 46: 3955: 3633: 822:. The paleosols are generally dark red or brown, with only intermittent 4237: 4210: 2433: 2391: 2301: 2293: 2051: 1981: 1881: 1789: 1678: 1671: 1650: 1610: 1305: 1254: 1128: 1036: 1006: 926: 869: 799: 736: 728: 657: 444: 438: 426: 401: 369: 320: 293: 91: 56: 2874:"Vertebrate faunal horizons in the Texas Permo-Carboniferous red beds" 2468:
continue to make up a significant portion of the palynomorphs, though
2021: 1990: 1209: 2970: 2584: 2513: 2507: 2331: 2234: 2215: 2122: 2030: 1951: 1903: 1800: 1753: 1659: 1569: 1539: 1465: 1057: 767: 747: 743: 720: 616: 393: 387: 275: 173: 169: 96: 40: 4155: 2125:
have also been reported from bonebeds of the Archer City Formation.
1957: 1722: 316:
formations, which are age-equivalent marine units to the southwest.
4374:"Lower Permian Flora of the Sanzenbacher Ranch, Clay County, Texas" 4344:
Bulletin of the Museum of Comparative Zoology at Harvard University
3938: 3781: 3766:"Edops, a Primitive Rhachitomous Amphibian from the Texas Red Beds" 3617: 2444: 2249: 1966: 1886: 1637: 1620: 1545: 1392: 979: 865: 857: 853: 846: 732: 704: 696: 688: 630: 517: 484: 421: 405: 332: 271: 159: 86: 81: 66: 61: 51: 36: 2180: 731:
sediments. The vast majority of each formation is homogeneous red
4110:"The Appendicular Skeleton of the Permian Embolomerous Amphibian 3851:, a new rhachitomous labyrinthodont from the Texas Lower Permian" 3522: 2700: 2344: 2340: 2273:
geologists Sergius H. Mamay and Arthur D. Watt. On behalf of the
2107: 2077: 1278: 1227: 1093: 1025: 933: 842: 834: 823: 779: 684: 359: 324: 319:
The sediments of the Archer City Formation reconstruct a coastal
282: 267: 101: 76: 3021:, Austin: The University of Texas and Bureau of Economic Geology 1413: 1311: 692: 2414: 1821: 1554: 791: 478: 1894: 619:
stage (293–290 Ma). The Santa Anna Branch Shale and overlying
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The Archer City Formation preserves a diverse fauna of fossil
2404: 1524: 904: 472: 304:, and succeeded by the equally fossiliferous red beds of the 263: 196: 4140:"The postcranial skeleton of the Permian microsaur Pantylus" 4072:
Neues Jahrbuch fΓΌr Geologie und PalΓ€ontologie - Abhandlungen
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Neues Jahrbuch fΓΌr Geologie und PalΓ€ontologie - Abhandlungen
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Reisz, Robert R.; Laurin, Michel; Marjanović, David (2010).
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At least seven skulls with associated postcranial fragments
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A medium-sized ophiacodontid, also described under the name
3163:"Global Permian tetrapod biostratigraphy and biochronology" 2454:
sp, and several more unnamed species. Sphenophytes such as
2383: 872:, deposited in dry upland areas free from the influence of 861: 849: 3291: 2985:"The stratigraphy of the Permian Wichita redbeds of Texas" 3992: 3821:
10.1666/0022-3360(2006)80[561:TASOEM]2.0.CO;2
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Isolated neural spines and at least one partial skeleton
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is the most abundant plant fossil in the lower bed. The
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of Europe, even if only a few specific species (such as
2296:. There are some notable differences, such as a lack of 1839:
Non-temnospondyl amphibians of the Archer City Formation
4371: 3088: 548:. The Archer City Formation was first named in a 1987 3573: 3503: 3361: 2612:. The most common palynomorphs are seed fern pollen ( 1977:
Multiple skeletons in varying states of completeness
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Tabor, Neil John; MontaΓ±ez, Isabel P. (2004-06-24).
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Teeth, fin spines, cartilage fragments, and a skull
296:. The Archer City Formation is preceded by the cool 4412:"Fossilworks: Gateway to the Paleobiology Database" 4410:Various Contributors to the Paleobiology Database. 4066:Milner, Andrew R.; Schoch, Rainer R. (2013-10-01). 3412:"New genera and species of pelycosaurian reptiles" 2691:List of fossiliferous stratigraphic units in Texas 2378:). The plants tend to be adapted to low humidity; 1884:, often conflated with the more fragmentary genus 3916:"The relationships of the rhachitomous amphibian 2775:Palaeogeography, Palaeoclimatology, Palaeoecology 2600:are occasionally prevalent. Sphenophytes include 1298:Five skulls with associated postcranial material 762:By volume, the most prevalent sandstone beds are 4427: 3167:Geological Society, London, Special Publications 1510:A dissorophid, formerly considered a species of 778:fill deposits. These are all hallmarks of small 488:, just to name a few. The rivers were roamed by 420:are also known from the formation. Some notable 3662:"Early evolution of the dissorophid amphibians" 1715:A trematopid, formerly considered a species of 1171:A partial skeleton with a well-preserved skull 652:, the Archer City Formation was deposited on a 606:(= Putnam Formation), from oldest to youngest. 3550:Proceedings of the New England ZoΓΆlogical Club 3419:Proceedings of the New England Zoological Club 4121:Contributions from the Museum of Paleontology 3855:Bulletin of the Museum of Comparative Zoology 3666:Bulletin of the Museum of Comparative Zoology 3580:Bulletin of the Museum of Comparative Zoology 2834: 2768: 2356:) is also common, followed by the seed ferns 2277:, Mamay led another expedition in 1990–1991. 4253:"Notes on certain American Paleozoic fishes" 4065: 3802: 3764:Romer, Alfred S.; Witter, Robert V. (1942). 2905:"Early history of Texas redbeds vertebrates" 899: 397:, etc.) were prominent in more humid areas. 327:and isolated ponds. The climate was hot and 3763: 3569: 3567: 3565: 3563: 2983:Romer, Alfred Sherwood (27 November 1974). 841:, such as riverbanks. Type F paleosols are 582:The Archer City Formation is a unit of the 511: 4181:, an Early Permian nectridian amphibian". 4108:Romer, Alfred Sherwood (11 January 1957). 3576:"Romeriid Reptiles from the Lower Permian" 3543: 3541: 3489:: CS1 maint: location missing publisher ( 3012: 2958: 683:During the Permian, the area was near the 244: 16:Geologic formation in Texas, United States 4268: 3972: 3954: 3521: 3337:Zoological Journal of the Linnean Society 3122: 3013:Hentz, Tucker F.; Brown, L.F. Jr (1987), 3840: 3838: 3560: 3405: 3403: 3401: 3399: 3397: 2582:is the most common conifer, followed by 2130: 1837: 1635:A temnospondyl of uncertain affinities. 1458:Two skulls, one with a partial skeleton 1363: 1225: 977: 4337: 4298: 4176: 4137: 3914:Carroll, Robert L. (21 February 1964). 3913: 3659: 3538: 3466: 3464: 3462: 3460: 3458: 3456: 3454: 3452: 3248: 4428: 4367: 4365: 4363: 4361: 4359: 4357: 4294: 4292: 4290: 4288: 4103: 4101: 4026: 4022: 4020: 3988: 3986: 3984: 3909: 3907: 3871: 3724: 3681: 3679: 3655: 3653: 3651: 3649: 3647: 3645: 3643: 3595: 3593: 3450: 3448: 3446: 3444: 3442: 3440: 3438: 3436: 3434: 3432: 3330: 2909:Geological Society of America Bulletin 2364:O. subcrenulata, O. osmundaeformis, O. 1649:, has variably been interpreted as an 559:The formation outcrops in an arc from 552:seeking to update the stratigraphy of 288:. It is the earliest component of the 4250: 4223: 4107: 3844: 3835: 3599: 3409: 3394: 3326: 3324: 3287: 3285: 3283: 3281: 3279: 3277: 3275: 3273: 3271: 3216:International Journal of Coal Geology 3202: 3200: 3198: 3196: 3160: 3156: 3154: 3152: 3150: 3148: 3146: 3144: 3142: 3084: 3082: 3080: 3078: 3076: 3040: 2982: 2902: 2898: 2896: 2894: 2871: 2867: 2865: 2830: 2828: 2826: 2824: 2764: 2762: 2760: 2758: 2756: 2754: 2752: 2750: 2748: 2746: 2744: 2742: 2740: 2738: 2736: 2548: 1350:A partial skull and partial skeleton 524:were first investigated in detail by 3470: 3074: 3072: 3070: 3068: 3066: 3064: 3062: 3060: 3058: 3056: 3036: 3034: 3032: 3030: 3028: 3008: 3006: 3004: 3002: 2954: 2952: 2950: 2948: 2946: 2944: 2942: 2940: 2938: 2822: 2820: 2818: 2816: 2814: 2812: 2810: 2808: 2806: 2804: 2734: 2732: 2730: 2728: 2726: 2724: 2722: 2720: 2718: 2716: 2608:, and to a lesser extent species of 363:, etc.) occupied drier soils, while 4354: 4285: 4098: 4017: 3981: 3936: 3904: 3676: 3640: 3590: 3429: 2959:Romer, A. S.; Price, L. W. (1940), 1830: 956:Uncertain or tentative taxa are in 643: 13: 4389:10.1016/B978-0-12-813012-4.00006-1 4183:Journal of Vertebrate Paleontology 4033:Journal of Vertebrate Paleontology 3803:Pawley, Kat; Warren, Anne (2006). 3731:Journal of Vertebrate Paleontology 3692:Journal of Vertebrate Paleontology 3368:Journal of Vertebrate Paleontology 3349:10.1111/j.1096-3642.1991.tb00886.x 3321: 3268: 3193: 3139: 2891: 2862: 2382:of moisture-loving plants such as 1056: 14: 4447: 4138:Carroll, Robert L. (1968-11-01). 3300:, Geological Society of America, 3053: 3025: 2999: 2935: 2801: 2713: 2288:formations of New Mexico and the 2132:Fish of the Archer City Formation 1595:Four partial skulls and postcania 1146:, also described under the names 1113:Limb and hip bones and vertebrae 916:Taxon falsely reported as present 864:-rich clay. They could be termed 757: 3872:Schoch, Rainer R. (2018-01-01). 3574:J. Clark; R. L. Carroll (1973). 2856:10.1111/j.1365-3091.2004.00655.x 2670: 2656: 2642: 2484:spp. (marattialean spores), and 2214: 2179: 2050: 2020: 1989: 1956: 1923: 1893: 1820: 1721: 1609: 1574: 1544: 1464: 1412: 1356: 1310: 1277: 1208: 1181: 1157: 1092: 207: 189: 44: 4338:Thomson, Keith Stewart (1964). 4331: 4244: 4217: 4170: 4131: 4059: 4027:Dilkes, David W. (1990-06-21). 3930: 3865: 3796: 3757: 3725:Dilkes, David W. (2020-07-03). 3718: 3497: 3355: 3242: 2127: 1834: 1360: 1222: 974: 921:Dubious taxon or junior synonym 4195:10.1080/02724634.1999.10011176 4045:10.1080/02724634.1990.10011809 3600:Olson, Everett Claire (1941). 2976: 2686:Red Beds of Texas and Oklahoma 2321: 1535:Several skulls and postcrania 1202:Skull fragments and vertebrae 750:bonebeds, which correspond to 1: 4383:, Elsevier, pp. 95–126, 3743:10.1080/02724634.2020.1801704 3179:10.1144/GSL.SP.2006.265.01.04 2706: 1485:Partial skull and postcrania 1433:Neural spine armor fragments 1403:Partial skull and postcrania 1024:, originally identified as a 879: 4317:10.1080/08912963.2012.669128 3937:Gee, Bryan M. (2021-11-08). 3704:10.1080/02724634.2011.582532 3380:10.1080/02724634.2010.501441 3015:"Wichita Falls-Lawton sheet" 2881:University of Texas Bulletin 2787:10.1016/0031-0182(89)90153-3 2423:(indeterminate pollen), and 1767:A micropholid amphibamiform 1368:of the Archer City Formation 1230:of the Archer City Formation 982:of the Archer City Formation 970: 808: 357:, etc.) and early conifers ( 7: 4257:American Journal of Science 4251:Romer, A. S. (1942-03-01). 4177:Carlson, Keith J. (1999). " 4144:Canadian Journal of Zoology 4084:10.1127/0077-7749/2013/0360 3660:Carroll, Robert L. (1964). 2903:Romer, A. S. (1935-11-30). 2635: 2565:Neurodontopteris auriculata 2533:, and the bisaccate pollen 2354:. schneideri, W. piniformis 1218: 883: 10: 4452: 4381:Transformative Paleobotany 4011:10.1016/j.crpv.2013.06.007 3602:"The Family Trematopsidae" 3510:Palaeontologia Electronica 3249:Wardlaw, Bruce R. (2005). 3228:10.1016/j.coal.2013.07.009 3161:Lucas, Spencer G. (2006). 3107:10.1007/s12542-019-00485-8 2962:Review of the Pelycosauria 2769:Martin Sander, P. (1989). 2167:Scales, jaws, and a skull 2090:Anodontacanthus americanus 714: 621:Coleman Junction Limestone 331:, preserving a variety of 4299:Johnson, Gary D. (2012). 3924:American Museum Novitates 3479:. Austin. pp. 47–53. 2628:), while conifer pollen ( 2570:Odontopteris subcrenulata 2376:A. conferta, A. naumannii 2259: 2213: 2202: 2187: 2102:Barbclabornia luederensis 1565:Jaws and other fragments 1340: 1333: 1318: 1126: 1047:Nearly complete skeleton 890: 886: 623:both preserve fossils of 448:. Early reptiles such as 243: 235: 225: 220: 202: 184: 179: 165: 155: 150: 140: 130: 120: 110: 26: 21: 4436:Permian geology of Texas 3471:Hook, Robert W. (1989). 2478:(indeterminate spores), 2065:Scales of indeterminate 888: 648:As with the rest of the 512:History and stratigraphy 379:, etc.) and horsetails ( 4226:Journal of Paleontology 3890:10.1127/njgpa/2018/0700 3809:Journal of Paleontology 3331:Laurin, Michel (1991). 3019:Geologic atlas of Texas 2519:Columinisporites ovalis 2397:Columinisporites ovalis 2060: 604:Santa Anna Branch shale 501:Ectosteorhachis nitidus 300:swamp sediments of the 239:Hentz & Brown, 1987 3999:Comptes Rendus Palevol 3770:The Journal of Geology 3606:The Journal of Geology 2575:Rhachiphyllum schenkii 2560:Sphenopteris germanica 2523:Colatisporites decorus 2420:Colatisporites decorus 2336:Sphenopteris germanica 1205:A large ophiacodontid 1088:Edaphosaurus cruciger. 1086:, often confused with 1082:A common medium-sized 1061: 964:taxa are discredited. 626:Sweetognathus merrilli 609: 433:Edaphosaurus boanerges 4270:10.2475/ajs.240.3.216 3306:10.1130/2006.2399(11) 3298:Wetlands through Time 2696:Paleontology in Texas 2650:Earth sciences portal 1660:Ecolsonia cutlerensis 1060: 833:. They are a type of 563:in the east, through 491:Orthacanthus texensis 451:Protorothyris archeri 256:Archer City Formation 22:Archer City Formation 3849:Neldasaurus wrightae 3845:Chase, J.N. (1965). 3410:Romer, A.S. (1937). 2921:10.1130/GSAB-46-1597 2872:Romer, A.S. (1928). 2429:(seed fern pollen). 2290:Neal Ranch Formation 1641:, the more complete 1050:A small sail-backed 575:counties, as far as 467:Neldasaurus wrightei 439:Ophiacodon uniformis 260:geological formation 3956:10.7717/peerj.12423 2678:Paleontology portal 1914:Jaws and vertebrae 1874:Jaws and vertebrae 1638:Parioxys ferricolus 1632:A partial skeleton 1507:A partial skeleton 1017:A partial skeleton 658:inland marine basin 577:Throckmorton County 520:in the vicinity of 479:Eryops megacephalus 28:Stratigraphic range 4305:Historical Biology 3506:"New eupelycosaur" 2549:Sanzenbacher Ranch 2508:Annularia carinata 2481:Cyclogranisporites 2460:are also present. 2445:Aphlebia erdmannii 1062: 911:Reclassified taxon 868:or (more broadly) 719:In terms of their 670:Ouachita Mountains 666:Arbuckle Mountains 586:, lying above the 544:formations of the 463:Phonerpeton pricei 445:Dimetrodon milleri 427:Apsisaurus witteri 230:Archer City, Texas 3315:978-0-8137-2399-0 2915:(11): 1597–1657. 2606:Annularia spicata 2316:Appalachian Basin 2257: 2256: 2058: 2057: 2010:Partial skeleton 1828: 1827: 1655:."Parioxys" bolli 1354: 1353: 1216: 1215: 968: 967: 959: 947: 946: 943: 936: 929: 922: 917: 912: 907: 818:event in the mid- 780:meandering rivers 662:Wichita Mountains 600:Sedwick Formation 588:Markley Formation 485:Pantylus cordatus 325:meandering rivers 323:crossed by small 302:Markley Formation 262:in north-central 252: 251: 145:Markley Formation 4443: 4422: 4420: 4418: 4402: 4401: 4378: 4369: 4352: 4351: 4335: 4329: 4328: 4296: 4283: 4282: 4272: 4248: 4242: 4241: 4232:(5): 1070–1078. 4221: 4215: 4214: 4174: 4168: 4167: 4150:(6): 1175–1192. 4135: 4129: 4128: 4118: 4105: 4096: 4095: 4063: 4057: 4056: 4024: 4015: 4014: 4005:(7–8): 447–461. 3990: 3979: 3978: 3976: 3958: 3934: 3928: 3927: 3911: 3902: 3901: 3869: 3863: 3862: 3842: 3833: 3832: 3800: 3794: 3793: 3761: 3755: 3754: 3722: 3716: 3715: 3683: 3674: 3673: 3657: 3638: 3637: 3597: 3588: 3587: 3571: 3558: 3557: 3545: 3536: 3535: 3525: 3501: 3495: 3494: 3488: 3480: 3468: 3427: 3426: 3416: 3407: 3392: 3391: 3374:(5): 1628–1631. 3359: 3353: 3352: 3328: 3319: 3318: 3289: 3266: 3265: 3255: 3246: 3240: 3239: 3213: 3204: 3191: 3190: 3158: 3137: 3136: 3126: 3086: 3051: 3050: 3038: 3023: 3022: 3010: 2997: 2996: 2980: 2974: 2973: 2971:10.1130/spe28-p1 2956: 2933: 2932: 2900: 2889: 2888: 2878: 2869: 2860: 2859: 2841: 2832: 2799: 2798: 2766: 2680: 2675: 2674: 2673: 2666: 2661: 2660: 2659: 2652: 2647: 2646: 2645: 2630:Potonieisporites 2578:, among others. 2556:Autunia conferta 2531:Potonieisporites 2487:Punctatisporites 2466:Potonieisporites 2410:Potonieisporites 2307:Autunia conferta 2218: 2183: 2128: 2099:), and teeth of 2054: 2024: 1993: 1960: 1927: 1910:D. sideropelicus 1897: 1835: 1831:Other amphibians 1824: 1811:Skull fragments 1725: 1653:or a dissorophid 1613: 1578: 1548: 1468: 1416: 1361: 1314: 1281: 1223: 1212: 1185: 1161: 1096: 975: 957: 939: 932: 925: 920: 915: 910: 903: 900: 884: 820:Clear Fork Group 644:Paleoenvironment 640:regional stage. 592:Nocona Formation 306:Nocona Formation 248: 213: 211: 210: 195: 193: 192: 135:Nocona Formation 106: 19: 18: 4451: 4450: 4446: 4445: 4444: 4442: 4441: 4440: 4426: 4425: 4416: 4414: 4406: 4405: 4399: 4376: 4370: 4355: 4336: 4332: 4297: 4286: 4249: 4245: 4222: 4218: 4175: 4171: 4156:10.1139/z68-168 4136: 4132: 4116: 4106: 4099: 4064: 4060: 4025: 4018: 3991: 3982: 3935: 3931: 3912: 3905: 3870: 3866: 3843: 3836: 3801: 3797: 3762: 3758: 3737:(4): e1801704. 3723: 3719: 3684: 3677: 3658: 3641: 3598: 3591: 3572: 3561: 3546: 3539: 3502: 3498: 3482: 3481: 3469: 3430: 3414: 3408: 3395: 3360: 3356: 3329: 3322: 3316: 3290: 3269: 3253: 3247: 3243: 3211: 3205: 3194: 3159: 3140: 3087: 3054: 3039: 3026: 3011: 3000: 2981: 2977: 2957: 2936: 2901: 2892: 2876: 2870: 2863: 2839: 2833: 2802: 2767: 2714: 2709: 2676: 2671: 2669: 2662: 2657: 2655: 2648: 2643: 2641: 2638: 2551: 2324: 2262: 2157:Ectosteorhachis 2063: 1833: 1561:E. megacephalus 1359: 1303:protorothyridid 1221: 1192:Stereophallodon 1139:Rare fragments 973: 955: 898: 882: 811: 760: 717: 646: 612: 561:Montague County 530:Llewellyn Price 514: 208: 206: 190: 188: 105: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 43: 35: 17: 12: 11: 5: 4449: 4439: 4438: 4424: 4423: 4404: 4403: 4397: 4353: 4330: 4311:(4): 369–379. 4284: 4263:(3): 216–228. 4243: 4216: 4189:(4): 623–631. 4169: 4130: 4097: 4058: 4039:(2): 222–243. 4016: 3980: 3929: 3903: 3864: 3834: 3815:(3): 561–580. 3795: 3782:10.1086/625101 3776:(8): 925–960. 3756: 3717: 3698:(4): 907–912. 3675: 3639: 3618:10.1086/624952 3612:(2): 149–176. 3589: 3559: 3537: 3496: 3428: 3393: 3354: 3320: 3314: 3267: 3241: 3192: 3138: 3101:(3): 423–451. 3052: 3024: 2998: 2975: 2934: 2890: 2861: 2850:(4): 851–884. 2800: 2711: 2710: 2708: 2705: 2704: 2703: 2698: 2693: 2688: 2682: 2681: 2667: 2653: 2637: 2634: 2550: 2547: 2413:spp. (conifer 2323: 2320: 2261: 2258: 2255: 2254: 2252: 2246: 2243: 2238: 2230: 2229: 2226: 2220: 2219: 2212: 2201: 2198: 2193: 2185: 2184: 2177: 2175:tetrapodomorph 2168: 2165: 2160: 2152: 2151: 2148: 2145: 2142: 2139: 2135: 2134: 2067:palaeoniscoids 2062: 2059: 2056: 2055: 2048: 2046:seymouriamorph 2042: 2039: 2037:S. baylorensis 2034: 2026: 2025: 2018: 2011: 2008: 2003: 1995: 1994: 1987: 1978: 1975: 1970: 1962: 1961: 1954: 1945: 1942: 1937: 1929: 1928: 1921: 1915: 1912: 1907: 1899: 1898: 1891: 1875: 1872: 1870:A. crassidisca 1867: 1859: 1858: 1855: 1852: 1849: 1846: 1842: 1841: 1832: 1829: 1826: 1825: 1818: 1812: 1809: 1804: 1796: 1795: 1793: 1786: 1784: 1779: 1776:Trimerorhachis 1771: 1770: 1768: 1765: 1762: 1757: 1749: 1748: 1746: 1745:A dissorophid 1743: 1740: 1738:R. renascentis 1735: 1727: 1726: 1719: 1713: 1710: 1705: 1697: 1696: 1694: 1685: 1682: 1675: 1667: 1666: 1664: 1633: 1630: 1625: 1615: 1614: 1607: 1602:trimerorhachid 1598: 1593: 1588: 1580: 1579: 1572: 1566: 1563: 1558: 1550: 1549: 1542: 1536: 1533: 1528: 1520: 1519: 1517: 1508: 1505: 1500: 1492: 1491: 1489: 1488:A dissorophid 1486: 1483: 1478: 1470: 1469: 1462: 1461:A dissorophid 1459: 1456: 1451: 1443: 1442: 1440: 1434: 1431: 1426: 1418: 1417: 1410: 1404: 1401: 1396: 1388: 1387: 1384: 1381: 1378: 1375: 1371: 1370: 1358: 1355: 1352: 1351: 1348: 1342: 1341: 1339: 1332: 1329: 1324: 1316: 1315: 1308: 1299: 1296: 1291: 1283: 1282: 1275: 1266: 1263: 1258: 1250: 1249: 1246: 1243: 1240: 1237: 1233: 1232: 1220: 1217: 1214: 1213: 1206: 1203: 1200: 1195: 1187: 1186: 1179: 1172: 1169: 1163: 1162: 1155: 1140: 1137: 1135:O. retroversus 1132: 1124: 1123: 1121: 1114: 1111: 1106: 1098: 1097: 1090: 1080: 1077: 1072: 1064: 1063: 1054: 1052:sphenacodontid 1048: 1045: 1040: 1032: 1031: 1029: 1018: 1015: 1010: 1002: 1001: 998: 995: 992: 989: 985: 984: 972: 969: 966: 965: 950: 948: 945: 944: 937: 930: 923: 918: 913: 908: 892: 891: 889: 887: 881: 878: 810: 807: 784:crevasse splay 774:, and coarser 764:upwards-fining 759: 758:Sandstone beds 756: 716: 713: 701:phyllosilicate 650:Texas red beds 645: 642: 611: 608: 590:and below the 513: 510: 418:type specimens 290:Texas red beds 250: 249: 241: 240: 237: 233: 232: 227: 223: 222: 218: 217: 204: 200: 199: 186: 182: 181: 177: 176: 167: 163: 162: 157: 153: 152: 148: 147: 142: 138: 137: 132: 128: 127: 122: 118: 117: 112: 108: 107: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 24: 23: 15: 9: 6: 4: 3: 2: 4448: 4437: 4434: 4433: 4431: 4413: 4408: 4407: 4400: 4398:9780128130124 4394: 4390: 4386: 4382: 4375: 4368: 4366: 4364: 4362: 4360: 4358: 4350:(9): 283–311. 4349: 4345: 4341: 4334: 4326: 4322: 4318: 4314: 4310: 4306: 4302: 4295: 4293: 4291: 4289: 4280: 4276: 4271: 4266: 4262: 4258: 4254: 4247: 4239: 4235: 4231: 4227: 4220: 4212: 4208: 4204: 4200: 4196: 4192: 4188: 4184: 4180: 4173: 4165: 4161: 4157: 4153: 4149: 4145: 4141: 4134: 4127:(5): 105–159. 4126: 4122: 4115: 4113: 4104: 4102: 4093: 4089: 4085: 4081: 4078:(1): 91–128. 4077: 4073: 4069: 4062: 4054: 4050: 4046: 4042: 4038: 4034: 4030: 4023: 4021: 4012: 4008: 4004: 4000: 3996: 3989: 3987: 3985: 3975: 3970: 3966: 3962: 3957: 3952: 3948: 3944: 3940: 3933: 3925: 3921: 3919: 3910: 3908: 3899: 3895: 3891: 3887: 3883: 3879: 3875: 3868: 3861:(3): 153–225. 3860: 3856: 3852: 3850: 3841: 3839: 3830: 3826: 3822: 3818: 3814: 3810: 3806: 3799: 3791: 3787: 3783: 3779: 3775: 3771: 3767: 3760: 3752: 3748: 3744: 3740: 3736: 3732: 3728: 3721: 3713: 3709: 3705: 3701: 3697: 3693: 3689: 3682: 3680: 3672:(7): 161–250. 3671: 3667: 3663: 3656: 3654: 3652: 3650: 3648: 3646: 3644: 3635: 3631: 3627: 3623: 3619: 3615: 3611: 3607: 3603: 3596: 3594: 3586:(5): 353–407. 3585: 3581: 3577: 3570: 3568: 3566: 3564: 3555: 3551: 3544: 3542: 3533: 3529: 3524: 3519: 3515: 3511: 3507: 3500: 3492: 3486: 3478: 3474: 3467: 3465: 3463: 3461: 3459: 3457: 3455: 3453: 3451: 3449: 3447: 3445: 3443: 3441: 3439: 3437: 3435: 3433: 3424: 3420: 3413: 3406: 3404: 3402: 3400: 3398: 3389: 3385: 3381: 3377: 3373: 3369: 3365: 3358: 3350: 3346: 3342: 3338: 3334: 3327: 3325: 3317: 3311: 3307: 3303: 3299: 3295: 3288: 3286: 3284: 3282: 3280: 3278: 3276: 3274: 3272: 3263: 3259: 3252: 3245: 3237: 3233: 3229: 3225: 3221: 3217: 3210: 3203: 3201: 3199: 3197: 3188: 3184: 3180: 3176: 3172: 3168: 3164: 3157: 3155: 3153: 3151: 3149: 3147: 3145: 3143: 3134: 3130: 3125: 3120: 3116: 3112: 3108: 3104: 3100: 3096: 3092: 3085: 3083: 3081: 3079: 3077: 3075: 3073: 3071: 3069: 3067: 3065: 3063: 3061: 3059: 3057: 3048: 3044: 3037: 3035: 3033: 3031: 3029: 3020: 3016: 3009: 3007: 3005: 3003: 2994: 2990: 2986: 2979: 2972: 2968: 2964: 2963: 2955: 2953: 2951: 2949: 2947: 2945: 2943: 2941: 2939: 2930: 2926: 2922: 2918: 2914: 2910: 2906: 2899: 2897: 2895: 2886: 2882: 2875: 2868: 2866: 2857: 2853: 2849: 2845: 2844:Sedimentology 2838: 2831: 2829: 2827: 2825: 2823: 2821: 2819: 2817: 2815: 2813: 2811: 2809: 2807: 2805: 2796: 2792: 2788: 2784: 2780: 2776: 2772: 2765: 2763: 2761: 2759: 2757: 2755: 2753: 2751: 2749: 2747: 2745: 2743: 2741: 2739: 2737: 2735: 2733: 2731: 2729: 2727: 2725: 2723: 2721: 2719: 2717: 2712: 2702: 2699: 2697: 2694: 2692: 2689: 2687: 2684: 2683: 2679: 2668: 2665: 2654: 2651: 2640: 2633: 2631: 2627: 2626: 2625:Anguisporites 2621: 2620: 2615: 2611: 2610:Sphenophyllum 2607: 2603: 2599: 2595: 2591: 2587: 2586: 2581: 2577: 2576: 2571: 2567: 2566: 2561: 2557: 2546: 2544: 2543: 2538: 2537: 2532: 2528: 2524: 2520: 2516: 2515: 2510: 2509: 2504: 2503:calamitaleans 2500: 2496: 2495:Sphenophyllum 2491: 2489: 2488: 2483: 2482: 2477: 2476: 2472: 2471:Knoxisporites 2467: 2463: 2459: 2458: 2457:Sphenophyllum 2453: 2452: 2447: 2446: 2441: 2440: 2436: 2430: 2428: 2427: 2422: 2421: 2416: 2412: 2411: 2406: 2403: 2399: 2398: 2393: 2389: 2385: 2381: 2377: 2373: 2369: 2365: 2361: 2360: 2355: 2351: 2347: 2346: 2342: 2338: 2337: 2333: 2328: 2319: 2317: 2313: 2312:Dunkard Group 2309: 2308: 2303: 2299: 2295: 2291: 2287: 2283: 2278: 2276: 2272: 2268: 2253: 2251: 2247: 2245:Tooth plates 2244: 2242: 2239: 2237: 2236: 2232: 2231: 2227: 2225: 2222: 2221: 2217: 2210: 2206: 2199: 2197: 2196:O. compressus 2194: 2192: 2191: 2186: 2182: 2178: 2176: 2173: 2172:megalichthyid 2169: 2166: 2164: 2161: 2159: 2158: 2154: 2153: 2149: 2146: 2143: 2140: 2137: 2136: 2133: 2129: 2126: 2124: 2120: 2116: 2115: 2110: 2109: 2104: 2103: 2098: 2097: 2096:Platyacanthus 2092: 2091: 2086: 2085: 2080: 2079: 2074: 2073: 2068: 2053: 2049: 2047: 2043: 2040: 2038: 2035: 2033: 2032: 2028: 2027: 2023: 2019: 2016: 2012: 2009: 2007: 2004: 2002: 2001: 1997: 1996: 1992: 1988: 1986: 1983: 1979: 1976: 1974: 1971: 1969: 1968: 1964: 1963: 1959: 1955: 1953: 1950: 1946: 1943: 1941: 1938: 1936: 1935: 1931: 1930: 1926: 1922: 1920: 1919:diadectomorph 1916: 1913: 1911: 1908: 1906: 1905: 1901: 1900: 1896: 1892: 1889: 1888: 1883: 1880: 1876: 1873: 1871: 1868: 1866: 1865: 1861: 1860: 1856: 1853: 1850: 1847: 1844: 1843: 1840: 1836: 1823: 1819: 1817: 1813: 1810: 1808: 1805: 1803: 1802: 1798: 1797: 1794: 1791: 1787: 1785: 1783: 1780: 1778: 1777: 1773: 1772: 1769: 1766: 1763: 1761: 1758: 1756: 1755: 1751: 1750: 1747: 1744: 1741: 1739: 1736: 1734: 1733: 1729: 1728: 1724: 1720: 1718: 1714: 1711: 1709: 1706: 1704: 1703: 1699: 1698: 1695: 1693: 1692:amphibamiform 1690: 1686: 1683: 1681: 1680: 1676: 1674: 1673: 1669: 1668: 1665: 1662: 1661: 1656: 1652: 1648: 1644: 1640: 1639: 1634: 1631: 1629: 1626: 1624: 1622: 1617: 1616: 1612: 1608: 1606: 1603: 1599: 1597: 1594: 1592: 1589: 1587: 1586: 1582: 1581: 1577: 1573: 1571: 1567: 1564: 1562: 1559: 1557: 1556: 1552: 1551: 1547: 1543: 1541: 1537: 1534: 1532: 1529: 1527: 1526: 1522: 1521: 1518: 1516: 1514: 1509: 1506: 1504: 1501: 1499: 1498: 1494: 1493: 1490: 1487: 1484: 1482: 1479: 1477: 1476: 1472: 1471: 1467: 1463: 1460: 1457: 1455: 1452: 1450: 1449: 1445: 1444: 1441: 1439: 1435: 1432: 1430: 1427: 1425: 1424: 1420: 1419: 1415: 1411: 1409: 1405: 1402: 1400: 1397: 1395: 1394: 1390: 1389: 1385: 1382: 1379: 1376: 1373: 1372: 1369: 1367: 1366:Temnospondyls 1362: 1357:Temnospondyls 1349: 1347: 1344: 1343: 1337: 1330: 1328: 1325: 1323: 1322: 1317: 1313: 1309: 1307: 1304: 1300: 1297: 1295: 1292: 1290: 1289: 1288:Protorothyris 1285: 1284: 1280: 1276: 1274: 1271: 1267: 1264: 1262: 1259: 1257: 1256: 1252: 1251: 1247: 1244: 1241: 1238: 1235: 1234: 1231: 1229: 1224: 1211: 1207: 1204: 1201: 1199: 1198:S. ciscoensis 1196: 1194: 1193: 1189: 1188: 1184: 1180: 1177: 1173: 1170: 1168: 1165: 1164: 1160: 1156: 1153: 1149: 1145: 1144:ophiacodontid 1141: 1138: 1136: 1133: 1131: 1130: 1125: 1122: 1120:edaphosaurid 1119: 1115: 1112: 1110: 1107: 1105: 1104: 1100: 1099: 1095: 1091: 1089: 1085: 1081: 1078: 1076: 1073: 1071: 1070: 1066: 1065: 1059: 1055: 1053: 1049: 1046: 1044: 1041: 1039: 1038: 1034: 1033: 1030: 1027: 1023: 1019: 1016: 1014: 1011: 1009: 1008: 1004: 1003: 999: 996: 993: 990: 987: 986: 983: 981: 976: 963: 954: 951: 949: 942: 938: 935: 931: 928: 924: 919: 914: 909: 906: 902: 901: 897: 894: 893: 885: 877: 875: 871: 867: 863: 859: 855: 851: 848: 844: 840: 836: 832: 827: 825: 821: 817: 816:aridification 806: 805: 804:flash floods. 801: 795: 793: 789: 788:remineralized 785: 781: 777: 773: 769: 765: 755: 753: 749: 745: 741: 738: 734: 730: 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88: 83: 78: 73: 68: 63: 58: 53: 48: 42: 38: 33: 32:Early Permian 29: 25: 20: 4415:. Retrieved 4380: 4347: 4343: 4333: 4308: 4304: 4260: 4256: 4246: 4229: 4225: 4219: 4186: 4182: 4178: 4172: 4147: 4143: 4133: 4124: 4120: 4111: 4075: 4071: 4061: 4036: 4032: 4002: 3998: 3946: 3942: 3932: 3923: 3917: 3881: 3877: 3867: 3858: 3854: 3848: 3812: 3808: 3798: 3773: 3769: 3759: 3734: 3730: 3720: 3695: 3691: 3669: 3665: 3609: 3605: 3583: 3579: 3553: 3549: 3523:10.26879/899 3513: 3509: 3499: 3476: 3422: 3418: 3371: 3367: 3357: 3343:(1): 59–95. 3340: 3336: 3297: 3261: 3257: 3244: 3219: 3215: 3173:(1): 65–93. 3170: 3166: 3098: 3094: 3046: 3042: 3018: 2992: 2988: 2978: 2961: 2912: 2908: 2884: 2880: 2847: 2843: 2778: 2774: 2664:Texas portal 2629: 2623: 2617: 2613: 2609: 2605: 2601: 2597: 2593: 2583: 2579: 2573: 2569: 2563: 2559: 2555: 2552: 2541: 2534: 2530: 2526: 2522: 2518: 2512: 2506: 2498: 2494: 2492: 2485: 2479: 2474: 2470: 2465: 2461: 2455: 2449: 2443: 2438: 2434: 2431: 2424: 2418: 2408: 2395: 2392:palynomorphs 2388:sphenophytes 2380:macrofossils 2375: 2371: 2367: 2363: 2359:Odontopteris 2357: 2353: 2349: 2343: 2334: 2329: 2325: 2305: 2298:marattialean 2279: 2263: 2240: 2233: 2223: 2195: 2190:Orthacanthus 2188: 2162: 2155: 2131: 2112: 2106: 2100: 2094: 2088: 2084:Ctenacanthus 2082: 2076: 2070: 2069:, spines of 2064: 2036: 2029: 2005: 1998: 1972: 1965: 1939: 1932: 1909: 1902: 1885: 1869: 1862: 1838: 1806: 1799: 1781: 1774: 1759: 1752: 1737: 1732:Reiszerpeton 1730: 1716: 1707: 1700: 1677: 1670: 1658: 1654: 1646: 1643:type species 1636: 1627: 1618: 1596: 1590: 1583: 1560: 1553: 1530: 1523: 1511: 1502: 1495: 1481:B. profundum 1480: 1473: 1453: 1446: 1428: 1423:Aspidosaurus 1421: 1398: 1391: 1364: 1345: 1326: 1319: 1293: 1286: 1260: 1253: 1226: 1197: 1190: 1175: 1167:O. uniformis 1166: 1151: 1147: 1134: 1127: 1108: 1101: 1087: 1084:edaphosaurid 1075:E. boanerges 1074: 1069:Edaphosaurus 1067: 1042: 1035: 1012: 1005: 978: 961: 952: 895: 828: 814:up until an 812: 796: 761: 718: 703:minerals in 695:records for 682: 647: 634:index fossil 624: 613: 581: 558: 550:geologic map 526:Alfred Romer 515: 499: 489: 483: 477: 473:Edops craigi 471: 461: 455: 449: 443: 437: 431: 425: 399: 392: 386: 380: 374: 368: 365:marattialean 358: 354:Odontopteris 352: 346: 342:Sphenopteris 340: 318: 255: 253: 221:Type section 4417:17 December 4179:Crossotelos 3884:(1): 1–16. 3516:(3): 1–42. 3258:Permophiles 3222:: 177–186. 2614:Vesicaspora 2598:Asterotheca 2542:. saarensis 2536:Platysaccus 2527:Vesicaspora 2501:) but also 2462:Vesicaspora 2451:Asterotheca 2426:Vesicaspora 2402:sphenophyll 2322:Kola Switch 2224:O. texensis 2119:petalodonts 2114:Gnathorhiza 2072:Xenacanthus 2017:nectridean 2015:urocordylid 2000:Sauropleura 1973:P. cordatus 1949:diplocaulid 1934:Diplocaulus 1807:Z. serratus 1760:T. texensis 1702:Phonerpeton 1689:micropholid 1679:P. cf. mayi 1591:N. wrightae 1585:Neldasaurus 1513:Dissorophus 1475:Brevidorsum 1448:Broiliellus 1438:dissorophid 1399:A. cumminsi 1336:captorhinid 1273:parareptile 1261:B. striatus 1176:Poliosaurus 1148:Theropleura 1103:Lupeosaurus 962:crossed out 941:Morphotaxon 874:groundwater 870:inceptisols 839:water table 800:megaripples 735:with small 721:lithologies 656:between an 638:Wolfcampian 584:Bowie Group 554:North Texas 546:Cisco Group 522:Archer City 402:vertebrates 376:Asterotheca 125:Bowie Group 3949:: e12423. 2707:References 2619:Wilsonites 2594:Pecopteris 2439:jongmansii 2435:Pecopteris 2302:Rotliegend 2294:West Texas 2163:E. nitidus 2123:cladodonts 2041:Vertebrae 1952:nectridean 1944:Vertebrae 1882:embolomere 1816:zatracheid 1790:New Mexico 1672:Pasawioops 1628:"P." bolli 1503:D. angusta 1497:Diploseira 1408:trematopid 1338:eureptile 1294:P. archeri 1270:bolosaurid 1255:Bolosaurus 1129:Ophiacodon 1043:D. milleri 1037:Dimetrodon 1013:A. witteri 1007:Apsisaurus 958:small text 927:Ichnotaxon 880:Paleobiota 794:minerals. 772:scroll-bar 766:stacks of 742:. Fluvial 737:calcareous 729:floodplain 678:transgress 602:, and the 370:Pecopteris 337:Seed ferns 321:floodplain 274:and early 4325:0891-2963 4279:0002-9599 4203:0272-4634 4164:0008-4301 4092:0077-7749 4053:0272-4634 3965:2167-8359 3898:0077-7749 3829:0022-3360 3790:0022-1376 3751:0272-4634 3712:0272-4634 3626:0022-1376 3556:: 97–102. 3532:1094-8074 3485:cite book 3388:0272-4634 3236:0166-5162 3187:0305-8719 3115:0031-0220 2929:0016-7606 2887:: 67–108. 2795:0031-0182 2602:Calamites 2585:Cordaites 2514:Calamites 2475:ruhlandii 2332:seed fern 2235:Sagenodus 2205:xenacanth 2144:Material 2031:Seymouria 2006:S. bairdi 1985:microsaur 1904:Diadectes 1879:archeriid 1851:Material 1801:Zatrachys 1754:Tersomius 1717:Acheloma. 1708:P. pricei 1605:dvinosaur 1568:A common 1531:E. craigi 1454:B. brevis 1380:Material 1346:R. texana 1306:eureptile 1242:Material 1028:reptile. 1022:varanopid 994:Material 980:Synapsids 971:Synapsids 896:Color key 866:aridisols 854:vertisols 809:Paleosols 768:point bar 752:perennial 748:claystone 744:sandstone 709:monsoonal 705:paleosols 687:, with a 617:Sakmarian 496:xenacanth 422:synapsids 394:Calamites 388:Annularia 333:paleosols 329:monsoonal 276:Sakmarian 270:from the 226:Named for 174:siltstone 170:sandstone 151:Lithology 131:Underlies 115:Formation 41:Sakmarian 4430:Category 4112:Archeria 3918:Parioxys 3634:30080836 3425:: 89–97. 3264:: 21–22. 3133:34262228 2989:Breviora 2781:: 1–21. 2636:See also 2590:Riparian 2368:. readii 2250:lungfish 2141:Species 1982:pantylid 1967:Pantylus 1887:Cricotus 1864:Archeria 1848:Species 1764:A skull 1742:A skull 1684:A skull 1651:eryopoid 1647:Parioxys 1621:Parioxys 1393:Acheloma 1377:Species 1331:A skull 1327:R. prima 1239:Species 1228:Reptiles 1219:Reptiles 1142:A large 1116:A small 991:Species 858:chlorite 847:smectite 843:alfisols 831:horizons 733:mudstone 697:hematite 689:latitude 631:conodont 518:bonebeds 424:include 406:bonebeds 404:in pond 272:Asselian 236:Named by 180:Location 160:mudstone 141:Overlies 37:Asselian 4238:1303852 4211:4524033 3974:8582317 3926:(2167). 3124:8276796 2995:: 1–31. 2701:Permian 2580:Walchia 2372:Autunia 2370:), and 2345:Walchia 2341:conifer 2314:of the 2150:Images 2108:Helodus 2087:, and " 2078:Hybodus 1857:Images 1570:eryopid 1540:edopoid 1386:Images 1321:Romeria 1248:Images 1152:Diopeus 1109:L. kayi 1026:diapsid 1000:Images 934:Ootaxon 835:entisol 824:gleying 776:channel 740:nodules 725:fluvial 715:Geology 685:equator 594:of the 516:Fossil 508:fish). 457:Romeria 408:across 367:ferns ( 360:Walchia 348:Autunia 283:Permian 281:of the 268:fossils 203:Country 156:Primary 121:Unit of 39:–early 4395:  4323:  4277:  4236:  4209:  4201:  4162:  4090:  4051:  3971:  3963:  3896:  3827:  3788:  3749:  3710:  3632:  3624:  3530:  3386:  3312:  3234:  3185:  3131:  3121:  3113:  2927:  2793:  2572:, and 2499:thonii 2415:pollen 2405:spores 2282:Bursum 2260:Plants 2241:S. sp. 2200:Teeth 2147:Notes 2138:Genus 2121:, and 1940:D. sp. 1854:Notes 1845:Genus 1782:T. sp. 1555:Eryops 1429:A. sp. 1383:Notes 1374:Genus 1265:A jaw 1245:Notes 1236:Genus 997:Notes 988:Genus 860:- and 792:copper 571:, and 569:Archer 542:Putnam 534:Pueblo 482:, and 442:, and 410:Archer 314:Putnam 286:period 279:stages 212:  194:  185:Region 4377:(PDF) 4234:JSTOR 4207:JSTOR 4117:(PDF) 3943:PeerJ 3630:JSTOR 3415:(PDF) 3254:(PDF) 3212:(PDF) 2877:(PDF) 2840:(PDF) 2384:ferns 2209:shark 2093:" (= 1525:Edops 1118:basal 953:Notes 905:Taxon 573:Young 540:, or 538:Moran 310:Moran 264:Texas 258:is a 197:Texas 166:Other 4419:2021 4393:ISBN 4321:ISSN 4275:ISSN 4199:ISSN 4160:ISSN 4088:ISSN 4049:ISSN 3961:ISSN 3894:ISSN 3825:ISSN 3786:ISSN 3747:ISSN 3708:ISSN 3622:ISSN 3528:ISSN 3491:link 3384:ISSN 3310:ISBN 3232:ISSN 3183:ISSN 3129:PMID 3111:ISSN 3095:PalZ 2925:ISSN 2885:2801 2791:ISSN 2596:and 2497:cf. 2473:cf. 2464:and 2437:cf. 2394:are 2386:and 2330:The 2284:and 2275:USNM 2271:USGS 2061:Fish 1150:and 862:mica 850:clay 727:and 699:and 693:Ξ΄18O 664:and 629:, a 565:Clay 528:and 454:and 414:Clay 412:and 254:The 111:Type 47:Preκž’ 4385:doi 4348:131 4313:doi 4265:doi 4261:240 4191:doi 4152:doi 4080:doi 4076:270 4041:doi 4007:doi 3969:PMC 3951:doi 3886:doi 3882:287 3859:133 3817:doi 3778:doi 3739:doi 3700:doi 3670:131 3614:doi 3584:144 3518:doi 3376:doi 3345:doi 3341:101 3302:doi 3224:doi 3220:119 3175:doi 3171:265 3119:PMC 3103:doi 3047:170 2993:427 2967:doi 2917:doi 2852:doi 2783:doi 2417:), 2407:), 2292:of 2286:Abo 2267:WPA 1877:An 1645:of 1538:An 790:by 610:Age 504:(a 494:(a 339:" ( 312:or 4432:: 4391:, 4379:, 4356:^ 4346:. 4342:. 4319:. 4309:24 4307:. 4303:. 4287:^ 4273:. 4259:. 4255:. 4230:52 4228:. 4205:. 4197:. 4187:19 4185:. 4158:. 4148:46 4146:. 4142:. 4125:13 4123:. 4119:. 4100:^ 4086:. 4074:. 4070:. 4047:. 4037:10 4035:. 4031:. 4019:^ 4003:12 4001:. 3997:. 3983:^ 3967:. 3959:. 3945:. 3941:. 3922:. 3906:^ 3892:. 3880:. 3876:. 3857:. 3853:. 3837:^ 3823:. 3813:80 3811:. 3807:. 3784:. 3774:50 3772:. 3768:. 3745:. 3735:40 3733:. 3729:. 3706:. 3696:31 3694:. 3690:. 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Index

Stratigraphic range
Early Permian
Asselian
Sakmarian
Preκž’
κž’
O
S
D
C
P
T
J
K
Pg
N
Formation
Bowie Group
Nocona Formation
Markley Formation
mudstone
sandstone
siltstone
Texas
United States
Archer City, Texas

geological formation
Texas
fossils

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